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<document id="A024FE3371C41AB5C8B7ABD5C66D0764" ID-CLB-Dataset="110553" ID-DOI="10.11646/palaeoentomology.5.2.8" ID-GBIF-Dataset="2df5b4ce-f1af-4788-b366-133a450a4233" ID-ISSN="2624-2834" ID-Zenodo-Dep="6530481" ID-ZooBank="671C89F0-463F-4FBA-927E-80A8FE7656C0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1652083590456" checkinUser="plazi" docAuthor="Azar, Dany &amp; Nel, André" docDate="2022" docId="7534F3340656FFF536D519D0FA00FD85" docLanguage="en" docName="palaeoentomology.5.2.8.pdf" docOrigin="Palaeoentomology 5 (2)" docSource="http://zoobank.org/671c89f0-463f-4fba-927e-80a8fe7656c0" docStyle="DocumentStyle:4EE6FD780FF4521D541D0D0600705D0E.4:Palaeoentomology.2018-.journal_article" docStyleId="4EE6FD780FF4521D541D0D0600705D0E" docStyleName="Palaeoentomology.2018-.journal_article" docStyleVersion="4" docTitle="Lebanosmylus leae Azar &amp; Nel 2022, sp. nov." docType="treatment" docUuid="2006186C-A70E-44CD-984A-8D86E7EC7518" docUuidSource="ZooBank" docVersion="6" lastPageNumber="156" masterDocId="890D8B4C0657FFF4365D1A79FFA2FFFC" masterDocTitle="The youngest and first Lebanese representative of the family Saucrosmylidae (Insecta, Neuroptera) from the Cenomanian" masterLastPageNumber="160" masterPageNumber="155" pageNumber="156" updateTime="1699329911539" updateUser="ExternalLinkService">
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<mods:title id="4E1F119E014716601B98F587A5543752">The youngest and first Lebanese representative of the family Saucrosmylidae (Insecta, Neuroptera) from the Cenomanian</mods:title>
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<mods:namePart id="11410818F79F4F25CC1F6373310F1BB3">Azar, Dany</mods:namePart>
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<mods:namePart id="9441D743A3F1656B506F723BDEF56501">Nel, André</mods:namePart>
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<mods:title id="ABBFDF7640E623B6D6FB965A4412FABD">Palaeoentomology</mods:title>
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<mods:part id="0064CCE3E026F934D52F2B08D431F5D6">
<mods:date id="8F0E3CC1064D3FB5EDA86124465087A8">2022</mods:date>
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<treatment id="7534F3340656FFF536D519D0FA00FD85" ID-DOI="http://doi.org/10.5281/zenodo.6950141" ID-GBIF-Taxon="198140664" ID-Zenodo-Dep="6950141" LSID="urn:lsid:zoobank.org:act:2006186C-A70E-44CD-984A-8D86E7EC7518" httpUri="http://treatment.plazi.org/id/7534F3340656FFF536D519D0FA00FD85" lastPageNumber="156" pageId="1" pageNumber="156">
<subSubSection id="B58711A90656FFF536D519D0FE67FC3F" box="[136,453,937,964]" pageId="1" pageNumber="156" type="nomenclature">
<paragraph id="FD2242220656FFF536D519D0FE67FC3F" blockId="1.[136,777,937,1073]" box="[136,453,937,964]" pageId="1" pageNumber="156">
<heading id="A66AF54E0656FFF536D519D0FE67FC3F" bold="true" box="[136,453,937,964]" fontSize="10" level="2" pageId="1" pageNumber="156" reason="3">
<emphasis id="CFE99E300656FFF536D519D0FE67FC3F" bold="true" box="[136,453,937,964]" pageId="1" pageNumber="156">
<taxonomicName id="3A9D39A10656FFF536D519D0FEC7FC38" authority="Azar &amp; Nel, 2022" authorityName="Azar &amp; Nel" authorityYear="2022" box="[136,357,937,964]" class="Insecta" family="Saucrosmylidae" genus="Lebanosmylus" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="156" phylum="Arthropoda" rank="species" species="leae" status="sp. nov.">
<emphasis id="CFE99E300656FFF536D519D0FEC7FC38" bold="true" box="[136,357,937,964]" italics="true" pageId="1" pageNumber="156">Lebanosmylus leae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D4DA234B0656FFF5373619D0FE67FC3F" box="[363,453,937,963]" pageId="1" pageNumber="156" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="B58711A90656FFF536D519B4FEE3FBCD" pageId="1" pageNumber="156" type="description">
<paragraph id="FD2242220656FFF536D519B4FEE3FBCD" blockId="1.[136,777,937,1073]" pageId="1" pageNumber="156">
(
<figureCitation id="65A65EA70656FFF536CD19B4FF75FC14" box="[144,215,973,1000]" captionStart="FIGURE 1" captionStartId="2.[136,235,1777,1801]" captionTargetBox="[324,1264,202,1730]" captionTargetId="figure-108@2.[324,1264,202,1754]" captionTargetPageId="2" captionText="FIGURE 1. Lebanosmylus leae gen. et sp. nov., holotype, wing. A, Photograph. B, Photograph with inverted colors. C, Line drawing, terminology of venation after Liu et al. (2013). Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.6530484" httpUri="https://zenodo.org/record/6530484/files/figure.png" pageId="1" pageNumber="156">Figs 1</figureCitation>
,
<figureCitation id="65A65EA70656FFF536B819B7FF56FC14" box="[229,244,974,1000]" captionStart="FIGURE 2" captionStartId="3.[136,235,1518,1542]" captionTargetBox="[325,1263,191,1466]" captionTargetId="figure-253@3.[325,1263,191,1495]" captionTargetPageId="3" captionText="FIGURE 2. Lebanosmylus leae gen. et sp. nov., holotype, wing counterpart. A, Photograph. B, Photograph with inverted colors. C, Line drawing. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.6530488" httpUri="https://zenodo.org/record/6530488/files/figure.png" pageId="1" pageNumber="156">2</figureCitation>
)
<uri id="890C4E200656FFF536D5198BFEE3FBCD" pageId="1" pageNumber="156">
urn:lsid:zoobank.org:act:
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</paragraph>
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<subSubSection id="B58711A90656FFF536D51E26FDE3FB1A" pageId="1" pageNumber="156" type="materials_examined">
<paragraph id="FD2242220656FFF536D51E26FDE3FB1A" blockId="1.[136,778,1119,2093]" pageId="1" pageNumber="156">
<emphasis id="CFE99E300656FFF536D51E26FF55FB85" bold="true" box="[136,247,1119,1145]" pageId="1" pageNumber="156">Material.</emphasis>
<materialsCitation id="4DF5487F0656FFF536A11E26FDE3FB1A" ID-GBIF-Occurrence="3874233301" collectorName="Sciences II" country="Lebanon" location="Faculty of Sciences" pageId="1" pageNumber="156" specimenCount="1" typeStatus="holotype">
<typeStatus id="2226FC800656FFF536A11E26FEC6FB85" box="[252,356,1119,1145]" pageId="1" pageNumber="156" type="holotype">Holotype</typeStatus>
(part and counterpart of an incomplete wing), Hussein Ibrahim collection, will be deposited in the Natural History Museum of the Lebanese University,
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Faculty of
<collectorName id="506827F40656FFF5375C1EB5FEC1FB1A" box="[257,355,1228,1254]" pageId="1" pageNumber="156">Sciences</collectorName>
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II, Fanar,
<collectingCountry id="858A02B20656FFF537851EB5FD9CFB1A" box="[472,574,1228,1254]" name="Lebanon" pageId="1" pageNumber="156">Lebanon</collectingCountry>
.
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</paragraph>
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<subSubSection id="B58711A90656FFF536E01E88FDD5FAD3" pageId="1" pageNumber="156" type="etymology">
<paragraph id="FD2242220656FFF536E01E88FDD5FAD3" blockId="1.[136,778,1119,2093]" pageId="1" pageNumber="156">
<emphasis id="CFE99E300656FFF536E01E88FEE6FAF7" bold="true" box="[189,324,1265,1291]" pageId="1" pageNumber="156">Etymology.</emphasis>
Named after Lea Ibrahim, daughter of Hussein Ibrahim, discoverer of the material.
</paragraph>
</subSubSection>
<subSubSection id="B58711A90656FFF536E01F40FE5FFAA8" box="[189,509,1337,1364]" pageId="1" pageNumber="156" type="diagnosis">
<paragraph id="FD2242220656FFF536E01F40FE5FFAA8" blockId="1.[136,778,1119,2093]" box="[189,509,1337,1364]" pageId="1" pageNumber="156">
<emphasis id="CFE99E300656FFF536E01F40FE9AFAAF" bold="true" box="[189,312,1337,1363]" pageId="1" pageNumber="156">Diagnosis.</emphasis>
As for the genus.
</paragraph>
</subSubSection>
<subSubSection id="B58711A90656FFF536E01F27FE38FA1A" pageId="1" pageNumber="156" type="materials_examined">
<paragraph id="FD2242220656FFF536E01F27FE38FA1A" blockId="1.[136,778,1119,2093]" pageId="1" pageNumber="156">
<emphasis id="CFE99E300656FFF536E01F27FD84FA84" bold="true" box="[189,550,1374,1400]" pageId="1" pageNumber="156">
<typeStatus id="2226FC800656FFF536E01F27FF5AFA84" box="[189,248,1374,1400]" pageId="1" pageNumber="156">Type</typeStatus>
locality and horizon.
</emphasis>
Mid-Cenomanian, Late Cretaceous, in marine lithographic limestone of the famous Hjoula fossil fishes outcrop (
<bibRefCitation id="990C3FD30656FFF534641FDEFF61FA1A" author="Maksoud, S. &amp; Azar, D." pageId="1" pageNumber="156" pagination="223 - 227" refId="ref3341" refString="Maksoud, S. &amp; Azar, D. (2021) Hjoula: A remarkable mid- Cenomanian Lebanese fossil fish Lagerstatte now also promising for fossil insects. Palaeoentomology, 4 (3), 223 - 227. https: // doi. org / 10.11646 / palaeoentomology. 4.3.8" type="journal article" year="2021">Maksoud &amp; Azar, 2021</bibRefCitation>
;
<bibRefCitation id="990C3FD30656FFF536901FB5FE2CFA1A" author="Azar, D. &amp; Maksoud, S. &amp; Huang, D. Y. &amp; Nel, A." box="[205,398,1483,1510]" pageId="1" pageNumber="156" pagination="78 - 89" refId="ref2638" refString="Azar, D., Maksoud, S., Huang, D. Y. &amp; Nel, A. (2019) First Lebanese dragonflies (Insecta: Odonata, Aeshnoptera, Cavilabiata) from the Arabo-African mid-Cretaceous paleocontinent. Cretaceous Research, 93, 78 - 89. https: // doi. org / 10.1016 / j. cretres. 2018.08.025" type="journal article" year="2019">
Azar
<emphasis id="CFE99E300656FFF537561FB5FE9EFA19" box="[267,316,1483,1509]" italics="true" pageId="1" pageNumber="156">et al</emphasis>
., 2019
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="B58711A90656FFF536E01F89FA00FD85" pageId="1" pageNumber="156" type="description">
<paragraph id="FD2242220656FFF536E01F89FA00FD85" blockId="1.[136,778,1119,2093]" lastBlockId="1.[810,1452,171,633]" pageId="1" pageNumber="156">
<emphasis id="CFE99E300656FFF536E01F89FEEDF9F6" bold="true" box="[189,335,1520,1546]" pageId="1" pageNumber="156">Description.</emphasis>
An incomplete wing, with base, fragments of mid-part, and apex missing, preserved part
<quantity id="3A65EFC70656FFF536D51C40FF50F9AF" box="[136,242,1593,1619]" metricMagnitude="-2" metricUnit="m" metricValue="4.02" pageId="1" pageNumber="156" unit="mm" value="40.2">40.2 mm</quantity>
long, wing very broad, 20.0 mm wide, with a broad cubital area (possibly a hind wing); wing dark brown, with three transverse hyaline zones (somewhat similar to those in the hind wing of
<taxonomicName id="3A9D39A10656FFF534541CDFFCA0F93C" box="[521,770,1702,1728]" class="Insecta" family="Osmylidae" genus="Bellinympha" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="156" phylum="Arthropoda" rank="species" species="filicifolia">
<emphasis id="CFE99E300656FFF534541CDFFCA0F93C" box="[521,770,1702,1728]" italics="true" pageId="1" pageNumber="156">Bellinympha filicifolia</emphasis>
</taxonomicName>
; see
<bibRefCitation id="990C3FD30656FFF536EE1CB2FEDCF919" author="Wang, Y. J. &amp; Liu, Z. Q. &amp; Wang, X. &amp; Shih, C. K. &amp; Zhao, Y. Y. &amp; Engel, M. S. &amp; Ren, D." box="[179,382,1738,1765]" pageId="1" pageNumber="156" pagination="16212 - 16215" refId="ref3744" refString="Wang, Y. J., Liu, Z. Q., Wang, X., Shih, C. K., Zhao, Y. Y., Engel, M. S. &amp; Ren, D. (2010) Ancient pinnate leaf mimesis among lacewings. Proceedings of the National Academy of Science of the United States of America, 107, 16212 - 16215. https: // doi. org / 10.1073 / pnas. 1006460107" type="journal article" year="2010">
Wang
<emphasis id="CFE99E300656FFF536A61CB2FE8FF918" box="[251,301,1738,1764]" italics="true" pageId="1" pageNumber="156">et al</emphasis>
., 2010
</bibRefCitation>
), and no dark spots; costal area
<quantity id="3A65EFC70656FFF534BA1CB2FF16F8F5" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="156" unit="mm" value="1.7">1.7 mm</quantity>
wide, rather narrow (hind wing?), with numerous simple and short veinlets; ScP and R straight, ScP ending into RA at
<emphasis id="CFE99E300656FFF537531D40FE8AF8AE" box="[270,296,1849,1874]" italics="true" pageId="1" pageNumber="156">ca</emphasis>
.
<quantity id="3A65EFC70656FFF537671D41FE00F8AE" box="[314,418,1848,1874]" metricMagnitude="-2" metricUnit="m" metricValue="1.3199999999999998" pageId="1" pageNumber="156" unit="mm" value="13.2">13.2 mm</quantity>
of wing apex; RA(+ScP) with numerous elongate anterior veinlets, some forked; no presectorial crossveins visible; area between RP and RA quite broad, broadest medially and distally narrowed near point of fusion of ScP with RA, with faint traces of at least two rows of irregular cells; base of RP virtually close to wing base; first posterior branch of RP (MA
<emphasis id="CFE99E300656FFF534DC126AFD1DF7D0" box="[641,703,2067,2092]" italics="true" pageId="1" pageNumber="156">sensu</emphasis>
<bibRefCitation id="990C3FD30656FFF5349A126BFC2CFF39" author="Liu, Q. &amp; Zhang, H. C. &amp; Wang, B. &amp; Fang, Y. &amp; Zheng, D. R. &amp; Zhang, Q. &amp; Jarzembowski, E. A." box="[711,910,171,2092]" pageId="1" pageNumber="156" pagination="387 - 391" refId="ref3165" refString="Liu, Q., Zhang, H. C., Wang, B., Fang, Y., Zheng, D. R., Zhang, Q. &amp; Jarzembowski, E. A. (2013) A new genus of Saucrosmylinae (Insecta, Neuroptera) from the Middle Jurassic of Daohugou, Inner Mongolia, China. Zootaxa, 3736 (4), 387 - 391. https: // doi. org / 10.11646 / zootaxa. 3736.4.6" type="journal article" year="2013">
Liu
<emphasis id="CFE99E300656FFF534A8126AFCE2FF39" box="[757,832,171,2092]" italics="true" pageId="1" pageNumber="156">et al</emphasis>
., 2013
</bibRefCitation>
) with a long stem and numerous distal branches, covering a very broad area; base of second branch of RP close to that of first branch, second branch straight and simple; eight more distal posterior branches of RP covering a broad area; pattern of crossveins between them poorly visible but no anterior Banksian fold; anteriormost branch of M (MP1
<emphasis id="CFE99E300656FFF532121BFFFB2FFE63" box="[1103,1165,390,415]" italics="true" pageId="1" pageNumber="156">sensu</emphasis>
<bibRefCitation id="990C3FD30656FFF532C51BFCFAEDFE63" author="Liu, Q. &amp; Zhang, H. C. &amp; Wang, B. &amp; Fang, Y. &amp; Zheng, D. R. &amp; Zhang, Q. &amp; Jarzembowski, E. A." box="[1176,1359,389,415]" pageId="1" pageNumber="156" pagination="387 - 391" refId="ref3165" refString="Liu, Q., Zhang, H. C., Wang, B., Fang, Y., Zheng, D. R., Zhang, Q. &amp; Jarzembowski, E. A. (2013) A new genus of Saucrosmylinae (Insecta, Neuroptera) from the Middle Jurassic of Daohugou, Inner Mongolia, China. Zootaxa, 3736 (4), 387 - 391. https: // doi. org / 10.11646 / zootaxa. 3736.4.6" type="journal article" year="2013">
Liu
<emphasis id="CFE99E300656FFF532941BFFFB5FFE63" box="[1225,1277,389,415]" italics="true" pageId="1" pageNumber="156">et al</emphasis>
., 2013
</bibRefCitation>
) simple and curved; posterior-most branch of M (MP2
<emphasis id="CFE99E300656FFF533631BD3FADEFE3F" box="[1342,1404,426,451]" italics="true" pageId="1" pageNumber="156">sensu</emphasis>
<bibRefCitation id="990C3FD30656FFF533D91BD0FC0CFE14" author="Liu, Q. &amp; Zhang, H. C. &amp; Wang, B. &amp; Fang, Y. &amp; Zheng, D. R. &amp; Zhang, Q. &amp; Jarzembowski, E. A." pageId="1" pageNumber="156" pagination="387 - 391" refId="ref3165" refString="Liu, Q., Zhang, H. C., Wang, B., Fang, Y., Zheng, D. R., Zhang, Q. &amp; Jarzembowski, E. A. (2013) A new genus of Saucrosmylinae (Insecta, Neuroptera) from the Middle Jurassic of Daohugou, Inner Mongolia, China. Zootaxa, 3736 (4), 387 - 391. https: // doi. org / 10.11646 / zootaxa. 3736.4.6" type="journal article" year="2013">
Liu
<emphasis id="CFE99E300656FFF535771BB7FCFFFE14" box="[810,861,462,488]" italics="true" pageId="1" pageNumber="156">et al</emphasis>
., 2013
</bibRefCitation>
) separated from CuA (no oblique vein O), with three or four posterior branches; CuA covering a very broad area with numerous elongate posterior branches (at least six or seven preserved); CuP and anal veins not preserved; posterior margin of wing sigmoidally curved.
</paragraph>
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</treatment>
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