treatments-xml/data/05/4F/DB/054FDB70FE85FE9CB49C1C0C7BB3231D.xml
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<document ID-DOI="10.7717/peerj.11514" ID-GBIF-Dataset="e56413f9-7dc9-4ebc-9085-008cb38b4dc1" ID-ISSN="2167-8359" ID-PMC="PMC8254503" ID-PubMed="34249486" ID-Zenodo-Dep="5102026" approvalRequired="1" approvalRequired_for_document="1" checkinTime="1626269878614" checkinUser="diego" docAuthor="Prebus, Matthew M." docDate="2021" docId="054FDB70FE85FE9CB49C1C0C7BB3231D" docLanguage="en" docName="PeerJ.9.e11514.pdf.imf" docOrigin="PeerJ (e 11514) 9" docSource="http://dx.doi.org/10.7717/peerj.11514" docStyle="DocumentStyle:B6346867C9D49AB2DD3A1D607021D2E9:PeerJ.2021-.journal_article" docStyleId="B6346867C9D49AB2DD3A1D607021D2E9" docStyleName="PeerJ.2021-.journal_article" docTitle="Temnothorax terricola" docType="treatment" docVersion="4" lastPageNumber="326" masterDocId="F976A308FFC7FFDAB56E1A1D79322757" masterDocTitle="Taxonomic revision of the Temnothorax salvini clade (Hymenoptera: Formicidae), with a key to the clades of New World Temnothorax" masterLastPageNumber="462" masterPageNumber="1" pageNumber="323" updateTime="1668138851842" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Taxonomic revision of the Temnothorax salvini clade (Hymenoptera: Formicidae), with a key to the clades of New World Temnothorax</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Prebus, Matthew M.</mods:namePart>
<mods:affiliation>School of Life Sciences, Arizona State University, Tempe, Arizona, United States &amp; Department of Entomology &amp; Nematology, University of California, Davis, Davis, California, United States</mods:affiliation>
<mods:nameIdentifier type="email">mprebus@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>PeerJ</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="series">
<mods:title>e 11514</mods:title>
</mods:detail>
<mods:detail type="pubDate">
<mods:number>2021-06-30</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>9</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>462</mods:end>
</mods:extent>
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<mods:location>
<mods:url>http://dx.doi.org/10.7717/peerj.11514</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.7717/peerj.11514</mods:identifier>
<mods:identifier type="GBIF-Dataset">e56413f9-7dc9-4ebc-9085-008cb38b4dc1</mods:identifier>
<mods:identifier type="ISSN">2167-8359</mods:identifier>
<mods:identifier type="PMC">PMC8254503</mods:identifier>
<mods:identifier type="PubMed">34249486</mods:identifier>
<mods:identifier type="Zenodo-Dep">5102026</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.5102612" ID-GBIF-Taxon="183542640" ID-Zenodo-Dep="5102612" LSID="urn:lsid:plazi:treatment:054FDB70FE85FE9CB49C1C0C7BB3231D" httpUri="http://treatment.plazi.org/id/054FDB70FE85FE9CB49C1C0C7BB3231D" lastPageId="326" lastPageNumber="326" pageId="322" pageNumber="323">
<subSubSection box="[498,945,1553,1580]" pageId="322" pageNumber="323" type="nomenclature">
<paragraph blockId="322.[498,977,1553,1620]" box="[498,945,1553,1580]" pageId="322" pageNumber="323">
<heading box="[498,945,1553,1580]" level="2" pageId="322" pageNumber="323">
<taxonomicName authority="(Mann, 1920)" baseAuthorityName="Mann" baseAuthorityYear="1920" box="[498,945,1553,1580]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="322" pageNumber="323" phylum="Arthropoda" rank="species" species="terricola">
<emphasis bold="true" box="[498,945,1553,1580]" pageId="322" pageNumber="323">
<emphasis bold="true" box="[498,770,1553,1580]" italics="true" pageId="322" pageNumber="323">Temnothorax terricola</emphasis>
(
<bibRefCitation author="Mann WM" box="[789,935,1553,1580]" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="322" pageNumber="323" pagination="403 - 439" part="42" refId="ref271797" refString="Mann WM. 1920. Additions to the ant fauna of the West Indies and Central America. Bulletin of the American Museum of Natural History 42: 403 - 439." title="Additions to the ant fauna of the West Indies and Central America" type="journal article" year="1920">
<emphasis bold="true" box="[789,935,1553,1580]" italics="true" pageId="322" pageNumber="323">Mann, 1920</emphasis>
</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[498,977,1593,1620]" pageId="322" pageNumber="323" type="distribution">
<paragraph blockId="322.[498,977,1553,1620]" box="[498,977,1593,1620]" pageId="322" pageNumber="323">
Distribution:
<figureCitation box="[657,766,1593,1619]" captionStart="Figure 133" captionStartId="258.[116,181,1036,1058]" captionTargetBox="[90,1541,234,1023]" captionTargetId="figure-184@258.[90,1524,234,1008]" captionTargetPageId="258" captionText="Figure 133 Geographical and elevational distribution of the pulchellus group. (A) Temnothorax agavicola sp. nov. (B) T. albispinus. (C) T. bahoruco sp. nov. (D) T. balaclava sp. nov. (E) T. ciferrii (F) T. flavidulus (G) T. harlequina sp. nov. (H) T. hippolytus sp. nov. (I) T. laticrus sp. nov. (J) T. magnabulla sp. nov. (K) T. pulchellus (L) T. terricola (M) T. torrei (N) T. wettereri sp. nov. (O) T. wilsoni sp. nov. Colored scale to the left of each map represents elevation in meters. Points in black represent the species named in each subfigure, while points in red represent other members of the species group. Type localities are represented by triangles; if present, types of synonyms are named; non-type localities are represented by circles. Bounding box in inset map shows location of main map. Full-size DOI: 10.7717/peerj.11514/fig-133" figureDoi="http://doi.org/10.5281/zenodo.5102387" httpUri="https://zenodo.org/record/5102387/files/figure.png" pageId="322" pageNumber="323">Fig. 133L</figureCitation>
; worker:
<figureCitation box="[878,971,1593,1619]" captionStart="Figure 146" captionStartId="322.[524,589,1066,1088]" captionTargetBox="[498,1541,234,1053]" captionTargetId="figure-220@322.[498,1295,234,1037]" captionTargetPageId="322" captionText="Figure 146 Temnothorax terricola lectotype worker (USNMENT00922871). (A) Profile view. (B) Full-face view. (C) Dorsal view. Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-146" figureDoi="http://doi.org/10.5281/zenodo.5102428" httpUri="https://zenodo.org/record/5102428/files/figure.png" pageId="322" pageNumber="323">Fig. 146</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="322" pageNumber="323" type="reference_group">
<paragraph blockId="322.[498,1537,1656,1921]" pageId="322" pageNumber="323">
<treatmentCitationGroup pageId="322" pageNumber="323">
<treatmentCitation author="Mann WM" box="[498,1260,1656,1682]" page="423" pageId="322" pageNumber="323" year="1920">
<taxonomicName authority=": Mann, 1920: 423" authorityName=": Mann" authorityPageNumber="423" authorityYear="1920" box="[498,1128,1656,1682]" class="Insecta" family="Formicidae" genus="Macromischa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="322" pageNumber="323" phylum="Arthropoda" rank="species" species="terricola" subGenus="Antillaemyrmex">
<emphasis box="[498,971,1656,1682]" italics="true" pageId="322" pageNumber="323">Macromischa (Antillaemyrmex) terricola</emphasis>
:
<bibRefCitation author="Mann WM" box="[987,1187,1656,1682]" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="322" pageNumber="323" pagination="403 - 439" part="42" refId="ref271797" refString="Mann WM. 1920. Additions to the ant fauna of the West Indies and Central America. Bulletin of the American Museum of Natural History 42: 403 - 439." title="Additions to the ant fauna of the West Indies and Central America" type="journal article" year="1920">
<emphasis box="[987,1128,1656,1682]" italics="true" pageId="322" pageNumber="323">Mann, 1920</emphasis>
: 423
</bibRefCitation>
</taxonomicName>
, fig. 7
</treatmentCitation>
.
<typeStatus box="[1273,1365,1656,1682]" pageId="322" pageNumber="323">Syntype</typeStatus>
workers and gyne.
<location LSID="urn:lsid:plazi:treatment:054FDB70FE85FE9CB49C1C0C7BB3231D:88393CBDFE85FE98B7561CBD7BA521ED" box="[568,663,1696,1722]" country="Cuba" municipality="One" name="Baracoa" pageId="322" pageNumber="323">Baracoa</location>
,
<collectingCountry box="[676,741,1696,1722]" name="Cuba" pageId="322" pageNumber="323">Cuba</collectingCountry>
.
<collectingMunicipality box="[753,803,1696,1722]" pageId="322" pageNumber="323">One</collectingMunicipality>
<typeStatus box="[811,900,1696,1722]" pageId="322" pageNumber="323">syntype</typeStatus>
worker here designated
<emphasis bold="true" box="[1192,1301,1696,1722]" pageId="322" pageNumber="323">
<typeStatus box="[1192,1301,1696,1722]" pageId="322" pageNumber="323">lectotype</typeStatus>
</emphasis>
.
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="322.[498,1537,1656,1921]" pageId="322" pageNumber="323">
<treatmentCitationGroup pageId="322" pageNumber="323">
<taxonomicName authority="(Mann)" baseAuthorityName="Mann" baseAuthorityYear="1920" box="[498,890,1735,1762]" class="Insecta" family="Formicidae" genus="Antillaemyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="322" pageNumber="323" phylum="Arthropoda" rank="species" species="terricola">
<emphasis box="[498,788,1735,1761]" italics="true" pageId="322" pageNumber="323">Antillaemyrmex terricola</emphasis>
(Mann)
</taxonomicName>
:
<treatmentCitation author="Wheeler WM" box="[902,1113,1735,1762]" page="32" pageId="322" pageNumber="323" year="1931">
<bibRefCitation author="Wheeler WM" box="[902,1113,1735,1762]" journalOrPublisher="Bulletin of the Museum of Comparative Zoology" pageId="322" pageNumber="323" pagination="1 - 34" part="72" refId="ref273428" refString="Wheeler WM. 1931. New and little-known ants of the genera Macromischa, Creosomyrmex and Antillaemyrmex. Bulletin of the Museum of Comparative Zoology 72: 1 - 34." title="New and little-known ants of the genera Macromischa, Creosomyrmex and Antillaemyrmex" type="journal article" year="1931">
<emphasis box="[902,1068,1735,1762]" italics="true" pageId="322" pageNumber="323">
<author box="[902,999,1735,1761]" fns="*" inm="true" npo="*" pageId="322" pageNumber="323">Wheeler</author>
,
<year box="[1013,1068,1736,1762]" pageId="322" pageNumber="323">1931</year>
</emphasis>
: 32
</bibRefCitation>
</treatmentCitation>
. First combination in
<taxonomicName authorityName="Mann" authorityYear="1920" box="[498,683,1775,1801]" class="Insecta" family="Formicidae" genus="Antillaemyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="322" pageNumber="323" phylum="Arthropoda" rank="genus">
<emphasis box="[498,683,1775,1801]" italics="true" pageId="322" pageNumber="323">Antillaemyrmex</emphasis>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="322.[498,1537,1656,1921]" pageId="322" pageNumber="323">
<treatmentCitationGroup pageId="322" pageNumber="323">
<taxonomicName authority="(Mann)" baseAuthorityName="Mann" baseAuthorityYear="1920" box="[498,855,1815,1841]" class="Insecta" family="Formicidae" genus="Leptothorax" kingdom="Animalia" order="Hymenoptera" pageId="322" pageNumber="323" phylum="Arthropoda" rank="species" species="terricolus">
<emphasis box="[498,753,1815,1841]" italics="true" pageId="322" pageNumber="323">Leptothorax terricolus</emphasis>
(Mann)
</taxonomicName>
:
<treatmentCitation author="Baroni Urbani C." box="[867,1168,1815,1842]" page="518" pageId="322" pageNumber="323" year="1978">
<bibRefCitation author="Baroni Urbani C." box="[867,1168,1815,1842]" journalOrPublisher="Entomologica Basiliensia" pageId="322" pageNumber="323" pagination="395 - 618" part="3" refId="ref269776" refString="Baroni Urbani C. 1978. Materiali per una revisione dei Leptothorax neotropicali appartenenti al sottogenere Macromischa Roger, n. comb. (Hymenoptera: Formicidae). Entomologica Basiliensia 3: 395 - 618." title="Materiali per una revisione dei Leptothorax neotropicali appartenenti al sottogenere Macromischa Roger, n. comb. (Hymenoptera: Formicidae)" type="journal article" year="1978">
<emphasis box="[867,1108,1815,1841]" italics="true" pageId="322" pageNumber="323">
<author box="[867,1040,1815,1841]" case="TC" fns="*" inm="true" npo="*" pageId="322" pageNumber="323">Baroni Urbani</author>
,
<year box="[1053,1108,1815,1841]" pageId="322" pageNumber="323">1978</year>
</emphasis>
: 518
</bibRefCitation>
</treatmentCitation>
. First combination in
<taxonomicName authority="Mayr" box="[498,638,1855,1881]" class="Insecta" family="Formicidae" genus="Leptothorax" kingdom="Animalia" order="Hymenoptera" pageId="322" pageNumber="323" phylum="Arthropoda" rank="genus">
<emphasis box="[498,638,1855,1881]" italics="true" pageId="322" pageNumber="323">Leptothorax</emphasis>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="322.[498,1537,1656,1921]" box="[530,1537,1895,1921]" pageId="322" pageNumber="323">
<treatmentCitationGroup box="[530,1537,1895,1921]" pageId="322" pageNumber="323">
<taxonomicName authority="(Mann)" baseAuthorityName="Mann" baseAuthorityYear="1920" box="[530,893,1895,1921]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="322" pageNumber="323" phylum="Arthropoda" rank="species" species="terricola">
<emphasis box="[530,791,1895,1921]" italics="true" pageId="322" pageNumber="323">Temnothorax terricola</emphasis>
(Mann)
</taxonomicName>
:
<treatmentCitation author="Bolton B." box="[906,1111,1895,1921]" page="272" pageId="322" pageNumber="323" year="2003">
<bibRefCitation author="Bolton B." box="[906,1111,1895,1921]" journalOrPublisher="Memoirs of the American Entomological Institute" pageId="322" pageNumber="323" pagination="370" part="71" refId="ref270076" refString="Bolton B. 2003. Synopsis and Classification of Formicidae. Memoirs of the American Entomological Institute 71: 370." title="Synopsis and Classification of Formicidae" type="journal article" year="2003">
<emphasis box="[906,1051,1895,1921]" italics="true" pageId="322" pageNumber="323">
<collectingRegion box="[906,984,1895,1921]" country="United Kingdom" name="Bolton" pageId="322" pageNumber="323">
<author box="[906,984,1895,1921]" fns="*" inm="true" npo="*" pageId="322" pageNumber="323">Bolton</author>
</collectingRegion>
,
<year box="[996,1051,1895,1921]" pageId="322" pageNumber="323">2003</year>
</emphasis>
: 272
</bibRefCitation>
</treatmentCitation>
. First combination in
<taxonomicName authorityName="Mayr" authorityYear="1861" box="[1374,1531,1895,1921]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="322" pageNumber="323" phylum="Arthropoda" rank="genus">
<emphasis box="[1374,1531,1895,1921]" italics="true" pageId="322" pageNumber="323">Temnothorax</emphasis>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="323" pageNumber="324" type="materials_examined">
<paragraph blockId="323.[498,1542,234,1935]" pageId="323" pageNumber="324">
<emphasis bold="true" box="[530,834,234,260]" pageId="323" pageNumber="324">Type material examined:</emphasis>
<materialsCitation ID-GBIF-Occurrence="3321861739" collectionCode="USNM" collectorName="W. M. Mann &amp; N. M. No." country="Cuba" location="Guantanamo" pageId="323" pageNumber="324" specimenCount="1" stateProvince="Guantanamo" typeStatus="lectotype">
<emphasis box="[843,1069,234,260]" italics="true" pageId="323" pageNumber="324">
<typeStatus box="[843,951,235,260]" pageId="323" pageNumber="324">Lectotype</typeStatus>
worker of
</emphasis>
<taxonomicName authorityName=": Mann" authorityYear="1920" class="Insecta" family="Formicidae" genus="Macromischa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="323" pageNumber="324" phylum="Arthropoda" rank="species" species="terricola" subGenus="Antillaemyrmex">Macromischa (Antillaemyrmex) terricola</taxonomicName>
:
<collectingCountry box="[610,691,274,300]" name="Cuba" pageId="323" pageNumber="324">CUBA</collectingCountry>
:
<collectingRegion box="[703,859,274,300]" country="Cuba" name="Guantanamo" pageId="323" pageNumber="324">Guantanamo</collectingRegion>
: Baracoa, 1918,
<collectorName box="[1050,1198,275,301]" pageId="323" pageNumber="324">W.M. Mann</collectorName>
, Cotype U.S.
<collectorName box="[1350,1463,274,301]" pageId="323" pageNumber="324">N.M. No.</collectorName>
54153 (USNMENT00922871, top specimen, furthest from pin) [
<collectionCode box="[1175,1267,313,340]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="323" pageNumber="324" type="Museum">USNM</collectionCode>
].
</materialsCitation>
</paragraph>
<paragraph blockId="323.[498,1542,234,1935]" pageId="323" pageNumber="324">
<materialsCitation ID-GBIF-Occurrence="3321861326" collectionCode="USNM" collectorName="W. M. Mann &amp; N. M. No." country="Cuba" location="Guantanamo" pageId="323" pageNumber="324" specimenCount="13" specimenCount-worker="13" stateProvince="Guantanamo" typeStatus="paralectotype">
<emphasis box="[530,904,354,380]" italics="true" pageId="323" pageNumber="324">
<typeStatus box="[530,684,354,380]" pageId="323" pageNumber="324">Paralectotype</typeStatus>
workers and gyne:
</emphasis>
same pin as
<typeStatus box="[1060,1165,354,380]" pageId="323" pageNumber="324">lectotype</typeStatus>
(USNMENT00922871, remaining specimens on pin) [
<collectionCode box="[859,950,393,420]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="323" pageNumber="324" type="Museum">USNM</collectionCode>
];
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3321861333" collectionCode="USNM" collectorName="W. M. Mann &amp; N. M. No." country="Cuba" location="Guantanamo" pageId="323" pageNumber="324" specimenCount="13" specimenCount-worker="13" stateProvince="Guantanamo" typeStatus="paralectotype">
same data as lectotype: 1 gyne, head and mesosoma missing,
<specimenCount box="[730,845,433,460]" pageId="323" pageNumber="324" type="worker">3 workers</specimenCount>
, head of
<specimenCount box="[947,1061,433,460]" pageId="323" pageNumber="324" type="worker">2 workers</specimenCount>
missing (USNMENT00922870) [
<collectionCode box="[1446,1535,433,460]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="323" pageNumber="324" type="Museum">USNM</collectionCode>
]
<specimenCount box="[498,616,473,499]" pageId="323" pageNumber="324" type="worker">4 workers</specimenCount>
, head of
<specimenCount box="[725,830,473,499]" pageId="323" pageNumber="324" type="worker">1 worker</specimenCount>
missing (USNMENT00532116) [
<collectionCode box="[1223,1312,473,500]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="323" pageNumber="324" type="Museum">USNM</collectionCode>
]
<specimenCount box="[1327,1443,473,499]" pageId="323" pageNumber="324" type="worker">3 workers</specimenCount>
(USNMENT00922871) [
<collectionCode box="[784,876,512,539]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="323" pageNumber="324" type="Museum">USNM</collectionCode>
]
</materialsCitation>
.
</paragraph>
<paragraph blockId="323.[498,1542,234,1935]" pageId="323" pageNumber="324">
<emphasis bold="true" box="[530,888,553,579]" pageId="323" pageNumber="324">Non-type material examined:</emphasis>
<materialsCitation ID-GBIF-Occurrence="3321861724" collectingDate="2001-08-24" collectionCode="UCDC" collectorName="P. S. Ward &amp; Same" country="Cuba" elevation="500" latitude="20.183332" location="Alto de Cotilla" longLatPrecision="1" longitude="-74.48333" pageId="323" pageNumber="324" specimenCount="5" specimenCount-worker="5" stateProvince="Guantanamo">
<collectingCountry box="[897,978,553,579]" name="Cuba" pageId="323" pageNumber="324">CUBA</collectingCountry>
:
<collectingRegion box="[989,1145,553,579]" country="Cuba" name="Guantanamo" pageId="323" pageNumber="324">Guantanamo</collectingRegion>
:
<quantity box="[1157,1218,553,579]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="323" pageNumber="324" unit="km" value="1.0">1 km</quantity>
NE
<location LSID="urn:lsid:plazi:treatment:054FDB70FE85FE9CB49C1C0C7BB3231D:88393CBDFE84FE99B19418347C9B2514" box="[1274,1449,553,579]" country="Cuba" latitude="20.183332" longLatPrecision="1" longitude="-74.48333" name="Alto de Cotilla" pageId="323" pageNumber="324" stateProvince="Guantanamo">Alto de Cotilla</location>
,
<geoCoordinate box="[498,650,588,619]" degrees="20.183333" direction="north" orientation="latitude" pageId="323" pageNumber="324" precision="1" value="20.183332">
20.183333
<emphasis box="[617,627,588,606]" italics="true" pageId="323" pageNumber="324">
<superScript attach="right" box="[617,627,588,606]" fontSize="8" pageId="323" pageNumber="324">°</superScript>
</emphasis>
N
</geoCoordinate>
<geoCoordinate box="[656,816,588,619]" degrees="74.483333" direction="west" orientation="longitude" pageId="323" pageNumber="324" precision="1" value="-74.48333">
74.483333
<emphasis box="[775,785,588,606]" italics="true" pageId="323" pageNumber="324">
<superScript attach="right" box="[775,785,588,606]" fontSize="8" pageId="323" pageNumber="324">°</superScript>
</emphasis>
W
</geoCoordinate>
,
<quantity box="[827,901,593,620]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="323" pageNumber="324" unit="m" value="500.0">
<elevation box="[827,901,593,620]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="323" pageNumber="324" unit="m" value="500.0">500 m</elevation>
</quantity>
,
<date box="[912,1095,593,619]" pageId="323" pageNumber="324" value="2001-08-24">
<collectingDate box="[912,1095,593,619]" pageId="323" pageNumber="324" value="2001-08-24">24 August 2001</collectingDate>
</date>
,
<collectorName box="[1106,1224,593,620]" pageId="323" pageNumber="324">P.S. Ward</collectorName>
#14453-6, pine forest, sifted litter (leaf mold, rotten wood),
<specimenCount box="[867,972,633,659]" pageId="323" pageNumber="324" type="worker">1 worker</specimenCount>
(CASENT0712509) [
<collectionCode box="[1223,1311,632,659]" country="USA" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="323" pageNumber="324" type="Museum">UCDC</collectionCode>
]
<specimenCount box="[1326,1431,633,659]" pageId="323" pageNumber="324" type="worker">1 worker</specimenCount>
(CASENT0758707) [
<collectionCode box="[741,829,672,699]" country="USA" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="323" pageNumber="324" type="Museum">UCDC</collectionCode>
]
<specimenCount box="[844,959,673,699]" pageId="323" pageNumber="324" type="worker">3 workers</specimenCount>
(CASENT0758300) [
<collectionCode box="[1210,1300,672,699]" country="USA" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="323" pageNumber="324" type="Museum">UCDC</collectionCode>
].
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3321861483" collectingDate="2001-08-24" collectionCode="UCDC" collectorName="P. S. Ward &amp; Same" country="Cuba" elevation="500" latitude="20.183332" location="Alto de Cotilla" longLatPrecision="1" longitude="-74.48333" pageId="323" pageNumber="324" specimenCount="5" specimenCount-worker="5" stateProvince="Guantanamo">
<collectorName box="[1320,1382,673,699]" pageId="323" pageNumber="324">Same</collectorName>
data as previous, except
</materialsCitation>
:
<materialsCitation ID-GBIF-Occurrence="3321861633" collectingDate="2001-08-23" collectionCode="UCDC" collectorName="P. S. Ward" country="Cuba" elevation="20" latitude="20.55" location="La Vega de Taco" longLatPrecision="555" longitude="-74.71667" pageId="323" pageNumber="324" specimenCount="2" specimenCount-worker="2" stateProvince="Guantanamo">
<quantity box="[702,763,712,738]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="323" pageNumber="324" unit="km" value="6.0">6 km</quantity>
NW
<location LSID="urn:lsid:plazi:treatment:054FDB70FE85FE9CB49C1C0C7BB3231D:88393CBDFE84FE99B65018D47D3525B4" box="[830,1031,712,739]" country="Cuba" latitude="20.55" longLatPrecision="555" longitude="-74.71667" name="La Vega de Taco" pageId="323" pageNumber="324" stateProvince="Guantanamo">La Vega de Taco</location>
,
<geoCoordinate box="[1044,1140,708,739]" degrees="20.55" direction="north" orientation="latitude" pageId="323" pageNumber="324" precision="555" value="20.55">
20.55
<emphasis box="[1106,1116,708,726]" italics="true" pageId="323" pageNumber="324">
<superScript attach="right" box="[1106,1116,708,726]" fontSize="8" pageId="323" pageNumber="324">°</superScript>
</emphasis>
N
</geoCoordinate>
<geoCoordinate box="[1148,1308,708,739]" degrees="74.716667" direction="west" orientation="longitude" pageId="323" pageNumber="324" precision="1" value="-74.71667">
74.716667
<emphasis box="[1266,1276,708,726]" italics="true" pageId="323" pageNumber="324">
<superScript attach="right" box="[1266,1276,708,726]" fontSize="8" pageId="323" pageNumber="324">°</superScript>
</emphasis>
W
</geoCoordinate>
,
<quantity box="[1321,1383,712,739]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="323" pageNumber="324" unit="m" value="20.0">
<elevation box="[1321,1383,712,739]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="323" pageNumber="324" unit="m" value="20.0">20 m</elevation>
</quantity>
,
<date pageId="323" pageNumber="324" value="2001-08-23">
<collectingDate pageId="323" pageNumber="324" value="2001-08-23">23 August 2001</collectingDate>
</date>
,
<collectorName box="[568,689,752,778]" pageId="323" pageNumber="324">P.S. Ward</collectorName>
#14441-1, rainforest edge, ground foragers,
<specimenCount box="[1193,1308,752,778]" pageId="323" pageNumber="324" type="worker">2 workers</specimenCount>
(CASENT0915980) [
<collectionCode box="[505,595,792,819]" country="USA" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="323" pageNumber="324" type="Museum">UCDC</collectionCode>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection box="[530,1184,832,858]" pageId="323" pageNumber="324" type="distribution">
<paragraph blockId="323.[498,1542,234,1935]" box="[530,1184,832,858]" pageId="323" pageNumber="324">
<emphasis bold="true" box="[530,755,832,858]" pageId="323" pageNumber="324">Geographic range:</emphasis>
Low elevations of
<collectingCountry box="[978,1040,832,858]" name="Cuba" pageId="323" pageNumber="324">Cuba</collectingCountry>
(
<figureCitation box="[1058,1167,832,858]" captionStart="Figure 133" captionStartId="258.[116,181,1036,1058]" captionTargetBox="[90,1541,234,1023]" captionTargetId="figure-184@258.[90,1524,234,1008]" captionTargetPageId="258" captionText="Figure 133 Geographical and elevational distribution of the pulchellus group. (A) Temnothorax agavicola sp. nov. (B) T. albispinus. (C) T. bahoruco sp. nov. (D) T. balaclava sp. nov. (E) T. ciferrii (F) T. flavidulus (G) T. harlequina sp. nov. (H) T. hippolytus sp. nov. (I) T. laticrus sp. nov. (J) T. magnabulla sp. nov. (K) T. pulchellus (L) T. terricola (M) T. torrei (N) T. wettereri sp. nov. (O) T. wilsoni sp. nov. Colored scale to the left of each map represents elevation in meters. Points in black represent the species named in each subfigure, while points in red represent other members of the species group. Type localities are represented by triangles; if present, types of synonyms are named; non-type localities are represented by circles. Bounding box in inset map shows location of main map. Full-size DOI: 10.7717/peerj.11514/fig-133" figureDoi="http://doi.org/10.5281/zenodo.5102387" httpUri="https://zenodo.org/record/5102387/files/figure.png" pageId="323" pageNumber="324">Fig. 133L</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="323" pageNumber="324" type="diagnosis">
<paragraph blockId="323.[498,1542,234,1935]" pageId="323" pageNumber="324">
<emphasis bold="true" box="[530,752,872,898]" pageId="323" pageNumber="324">Worker diagnosis:</emphasis>
<taxonomicName baseAuthorityName="Mann" baseAuthorityYear="1920" box="[759,1017,872,898]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="323" pageNumber="324" phylum="Arthropoda" rank="species" species="terricola">
<emphasis box="[759,1017,872,898]" italics="true" pageId="323" pageNumber="324">Temnothorax terricola</emphasis>
</taxonomicName>
can be separated from all other species in the
<taxonomicName baseAuthorityName="Forel" baseAuthorityYear="1899" box="[498,573,912,938]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="323" pageNumber="324" phylum="Arthropoda" rank="species" species="salvini">
<emphasis box="[498,573,912,938]" italics="true" pageId="323" pageNumber="324">salvini</emphasis>
</taxonomicName>
clade by the following character combination: head in full face view blocky; promesonotal suture shallowly impressed; metanotal groove not impressed; propodeum not depressed below the level of the promesonotum; propodeum bearing standing setae dorsally; propodeal spines about as long as the propodeal declivity, and directed posterodorsally; hind femora weakly incrassate; petiolar node rounded: dorsal face transitioning to posterior face through a broad curve; petiolar node with only two erect setae dorsally; postpetiole broad: more than 2.2 times the width of the petiole; dorsum of head smooth and shining; setae on head, mesosoma, waist segments and gaster erect, short, sparse and blunt (never long and tapering); head, mesosoma, waist segments, and gaster are uniformly testaceous; mandibles and extremities light brown.
</paragraph>
<paragraph blockId="323.[498,1542,234,1935]" pageId="323" pageNumber="324">
<emphasis bold="true" box="[530,716,1310,1336]" pageId="323" pageNumber="324">Similar species:</emphasis>
<emphasis box="[723,1132,1310,1336]" italics="true" pageId="323" pageNumber="324">
<taxonomicName baseAuthorityName="Menozzi &amp; Russo" baseAuthorityYear="1930" box="[723,964,1310,1336]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="323" pageNumber="324" phylum="Arthropoda" rank="species" species="ciferrii">Temnothorax ciferrii</taxonomicName>
,
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[975,1132,1310,1336]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="323" pageNumber="324" phylum="Arthropoda" rank="species" species="harlequina" status="sp. nov.">T. harlequina</taxonomicName>
</emphasis>
<taxonomicNameLabel box="[1138,1229,1311,1337]" pageId="323" pageNumber="324" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1240,1389,1310,1336]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="323" pageNumber="324" phylum="Arthropoda" rank="species" species="hippolytus" status="sp. nov.">
<emphasis box="[1240,1389,1310,1336]" italics="true" pageId="323" pageNumber="324">T. hippolytus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1396,1485,1311,1337]" pageId="323" pageNumber="324" rank="species">sp. nov.</taxonomicNameLabel>
, and
<emphasis box="[498,865,1350,1376]" italics="true" pageId="323" pageNumber="324">
<taxonomicName baseAuthorityName="Aguayo" baseAuthorityYear="1931" box="[498,595,1351,1376]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="323" pageNumber="324" phylum="Arthropoda" rank="species" species="torrei">T. torrei</taxonomicName>
.
<taxonomicName baseAuthorityName="Mann" baseAuthorityYear="1920" box="[606,865,1350,1376]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="323" pageNumber="324" phylum="Arthropoda" rank="species" species="terricola">Temnothorax terricola</taxonomicName>
</emphasis>
can be distinguished from
<taxonomicName baseAuthorityName="Menozzi &amp; Russo" baseAuthorityYear="1930" box="[1181,1287,1350,1376]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="323" pageNumber="324" phylum="Arthropoda" rank="species" species="ciferrii">
<emphasis box="[1181,1287,1350,1376]" italics="true" pageId="323" pageNumber="324">T. ciferrii</emphasis>
</taxonomicName>
and
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1346,1503,1350,1376]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="323" pageNumber="324" phylum="Arthropoda" rank="species" species="harlequina" status="sp. nov.">
<emphasis box="[1346,1503,1350,1376]" italics="true" pageId="323" pageNumber="324">T. harlequina</emphasis>
</taxonomicName>
<taxonomicNameLabel pageId="323" pageNumber="324" rank="species">sp. nov.</taxonomicNameLabel>
by the setae count of two on the dorsum of the petiolar node, versus four.
<taxonomicName baseAuthorityName="Mann" baseAuthorityYear="1920" box="[498,759,1430,1456]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="323" pageNumber="324" phylum="Arthropoda" rank="species" species="terricola">
<emphasis box="[498,759,1430,1456]" italics="true" pageId="323" pageNumber="324">Temnothorax terricola</emphasis>
</taxonomicName>
is closely related to
<taxonomicName baseAuthorityName="Aguayo" baseAuthorityYear="1931" box="[1002,1097,1430,1455]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="323" pageNumber="324" phylum="Arthropoda" rank="species" species="torrei">
<emphasis box="[1002,1097,1430,1455]" italics="true" pageId="323" pageNumber="324">T. torrei</emphasis>
</taxonomicName>
and
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1159,1310,1430,1456]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="323" pageNumber="324" phylum="Arthropoda" rank="species" species="hippolytus" status="sp. nov.">
<emphasis box="[1159,1310,1430,1456]" italics="true" pageId="323" pageNumber="324">T. hippolytus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1319,1412,1430,1456]" pageId="323" pageNumber="324" rank="species">sp. nov.</taxonomicNameLabel>
, but the smooth and shining head sculpture can be used to separate
<taxonomicName baseAuthorityName="Mann" baseAuthorityYear="1920" box="[1205,1332,1470,1496]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="323" pageNumber="324" phylum="Arthropoda" rank="species" species="terricola">
<emphasis box="[1205,1332,1470,1496]" italics="true" pageId="323" pageNumber="324">T. terricola</emphasis>
</taxonomicName>
from
<taxonomicName baseAuthorityName="Aguayo" baseAuthorityYear="1931" box="[1407,1503,1470,1495]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="323" pageNumber="324" phylum="Arthropoda" rank="species" species="torrei">
<emphasis box="[1407,1503,1470,1495]" italics="true" pageId="323" pageNumber="324">T. torrei</emphasis>
</taxonomicName>
, in which the head sculpture is at least somewhat areolate.
<taxonomicName baseAuthorityName="Mann" baseAuthorityYear="1920" box="[1154,1415,1509,1535]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="323" pageNumber="324" phylum="Arthropoda" rank="species" species="terricola">
<emphasis box="[1154,1415,1509,1535]" italics="true" pageId="323" pageNumber="324">Temnothorax terricola</emphasis>
</taxonomicName>
can be differentiated from
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[728,878,1549,1575]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="323" pageNumber="324" phylum="Arthropoda" rank="species" species="hippolytus" status="sp. nov.">
<emphasis box="[728,878,1549,1575]" italics="true" pageId="323" pageNumber="324">T. hippolytus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[886,977,1550,1576]" pageId="323" pageNumber="324" rank="species">sp. nov.</taxonomicNameLabel>
by the boxy head shape (ovular in
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1391,1541,1549,1575]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="323" pageNumber="324" phylum="Arthropoda" rank="species" species="hippolytus" status="sp. nov.">
<emphasis box="[1391,1541,1549,1575]" italics="true" pageId="323" pageNumber="324">T. hippolytus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[498,593,1590,1616]" pageId="323" pageNumber="324" rank="species">sp. nov.</taxonomicNameLabel>
), the shallowly impressed promesonotal suture, and the less angulate transition between the dorsal and posterior faces of the petiolar node.
</paragraph>
</subSubSection>
<subSubSection lastPageId="325" lastPageNumber="326" pageId="323" pageNumber="324" type="description">
<paragraph blockId="323.[498,1542,234,1935]" lastBlockId="324.[498,1542,234,1894]" lastPageId="324" lastPageNumber="325" pageId="323" pageNumber="324">
<emphasis bold="true" box="[530,918,1668,1695]" pageId="323" pageNumber="324">
Worker measurements (
<emphasis bold="true" box="[824,840,1669,1695]" italics="true" pageId="323" pageNumber="324">n</emphasis>
= 14):
</emphasis>
SL =
<geoCoordinate box="[985,1047,1669,1695]" degrees="0.349" direction="north" orientation="latitude" pageId="323" pageNumber="324" precision="55" value="0.349">0.349</geoCoordinate>
<geoCoordinate box="[1048,1124,1669,1695]" degrees="0.401" direction="west" orientation="longitude" pageId="323" pageNumber="324" precision="55" value="-0.401">0.401</geoCoordinate>
(0.380); FRS =
<geoCoordinate box="[1306,1368,1669,1695]" degrees="0.130" direction="north" orientation="latitude" pageId="323" pageNumber="324" precision="55" value="0.13">0.130</geoCoordinate>
<geoCoordinate box="[1368,1445,1669,1695]" degrees="0.164" direction="west" orientation="longitude" pageId="323" pageNumber="324" precision="55" value="-0.164">0.164</geoCoordinate>
(0.145); CW =
<geoCoordinate box="[580,642,1709,1735]" degrees="0.401" direction="north" orientation="latitude" pageId="323" pageNumber="324" precision="55" value="0.401">0.401</geoCoordinate>
<geoCoordinate box="[642,719,1709,1735]" degrees="0.471" direction="west" orientation="longitude" pageId="323" pageNumber="324" precision="55" value="-0.471">0.471</geoCoordinate>
(0.450); CWb =
<geoCoordinate box="[922,984,1708,1734]" degrees="0.360" direction="north" orientation="latitude" pageId="323" pageNumber="324" precision="55" value="0.36">0.360</geoCoordinate>
<geoCoordinate box="[984,1061,1709,1735]" degrees="0.441" direction="west" orientation="longitude" pageId="323" pageNumber="324" precision="55" value="-0.441">0.441</geoCoordinate>
(0.416); PoOC =
<geoCoordinate box="[1274,1336,1709,1735]" degrees="0.213" direction="north" orientation="latitude" pageId="323" pageNumber="324" precision="55" value="0.213">0.213</geoCoordinate>
<geoCoordinate box="[1336,1413,1709,1735]" degrees="0.243" direction="west" orientation="longitude" pageId="323" pageNumber="324" precision="55" value="-0.243">0.243</geoCoordinate>
(0.229); CL =
<geoCoordinate box="[567,629,1749,1775]" degrees="0.471" direction="north" orientation="latitude" pageId="323" pageNumber="324" precision="55" value="0.471">0.471</geoCoordinate>
<geoCoordinate box="[630,706,1748,1775]" degrees="0.551" direction="west" orientation="longitude" pageId="323" pageNumber="324" precision="55" value="-0.551">0.551</geoCoordinate>
(0.514); EL =
<geoCoordinate box="[879,941,1749,1775]" degrees="0.099" direction="north" orientation="latitude" pageId="323" pageNumber="324" precision="55" value="0.099">0.099</geoCoordinate>
<geoCoordinate box="[942,1019,1748,1775]" degrees="0.126" direction="west" orientation="longitude" pageId="323" pageNumber="324" precision="55" value="-0.126">0.126</geoCoordinate>
(0.110); EW =
<geoCoordinate box="[1204,1266,1748,1774]" degrees="0.069" direction="north" orientation="latitude" pageId="323" pageNumber="324" precision="55" value="0.069">0.069</geoCoordinate>
<geoCoordinate box="[1266,1343,1749,1775]" degrees="0.101" direction="west" orientation="longitude" pageId="323" pageNumber="324" precision="55" value="-0.101">0.101</geoCoordinate>
(0.084); MD =
<geoCoordinate box="[580,642,1788,1814]" degrees="0.094" direction="north" orientation="latitude" pageId="323" pageNumber="324" precision="55" value="0.094">0.094</geoCoordinate>
<geoCoordinate box="[642,719,1788,1814]" degrees="0.122" direction="west" orientation="longitude" pageId="323" pageNumber="324" precision="55" value="-0.122">0.122</geoCoordinate>
(0.108); WL =
<geoCoordinate box="[904,966,1788,1814]" degrees="0.493" direction="north" orientation="latitude" pageId="323" pageNumber="324" precision="55" value="0.493">0.493</geoCoordinate>
<geoCoordinate box="[966,1043,1788,1815]" degrees="0.597" direction="west" orientation="longitude" pageId="323" pageNumber="324" precision="55" value="-0.597">0.597</geoCoordinate>
(0.552); SPST =
<geoCoordinate box="[1245,1307,1788,1814]" degrees="0.147" direction="north" orientation="latitude" pageId="323" pageNumber="324" precision="55" value="0.147">0.147</geoCoordinate>
<geoCoordinate box="[1308,1385,1788,1814]" degrees="0.204" direction="west" orientation="longitude" pageId="323" pageNumber="324" precision="55" value="-0.204">0.204</geoCoordinate>
(0.176); MPST =
<geoCoordinate box="[607,669,1828,1854]" degrees="0.161" direction="north" orientation="latitude" pageId="323" pageNumber="324" precision="55" value="0.161">0.161</geoCoordinate>
<geoCoordinate box="[669,746,1828,1854]" degrees="0.220" direction="west" orientation="longitude" pageId="323" pageNumber="324" precision="55" value="-0.22">0.220</geoCoordinate>
(0.182); PEL =
<geoCoordinate box="[935,997,1828,1854]" degrees="0.200" direction="north" orientation="latitude" pageId="323" pageNumber="324" precision="55" value="0.2">0.200</geoCoordinate>
<geoCoordinate box="[997,1074,1828,1855]" degrees="0.251" direction="west" orientation="longitude" pageId="323" pageNumber="324" precision="55" value="-0.251">0.251</geoCoordinate>
(0.220); NOL =
<geoCoordinate box="[1274,1336,1828,1854]" degrees="0.130" direction="north" orientation="latitude" pageId="323" pageNumber="324" precision="55" value="0.13">0.130</geoCoordinate>
<geoCoordinate box="[1336,1413,1828,1854]" degrees="0.164" direction="west" orientation="longitude" pageId="323" pageNumber="324" precision="55" value="-0.164">0.164</geoCoordinate>
(0.145); NOH =
<geoCoordinate box="[598,660,1868,1894]" degrees="0.074" direction="north" orientation="latitude" pageId="323" pageNumber="324" precision="55" value="0.074">0.074</geoCoordinate>
<geoCoordinate box="[660,737,1868,1894]" degrees="0.099" direction="west" orientation="longitude" pageId="323" pageNumber="324" precision="55" value="-0.099">0.099</geoCoordinate>
(0.090); PEH =
<geoCoordinate box="[933,995,1868,1894]" degrees="0.140" direction="north" orientation="latitude" pageId="323" pageNumber="324" precision="55" value="0.14">0.140</geoCoordinate>
<geoCoordinate box="[995,1072,1868,1894]" degrees="0.170" direction="west" orientation="longitude" pageId="323" pageNumber="324" precision="55" value="-0.17">0.170</geoCoordinate>
(0.155); PPL =
<geoCoordinate box="[1262,1324,1868,1894]" degrees="0.129" direction="north" orientation="latitude" pageId="323" pageNumber="324" precision="55" value="0.129">0.129</geoCoordinate>
<geoCoordinate box="[1324,1401,1868,1894]" degrees="0.160" direction="west" orientation="longitude" pageId="323" pageNumber="324" precision="55" value="-0.16">0.160</geoCoordinate>
(0.141); PPH =
<geoCoordinate box="[588,650,1908,1934]" degrees="0.136" direction="north" orientation="latitude" pageId="323" pageNumber="324" precision="55" value="0.136">0.136</geoCoordinate>
<geoCoordinate box="[650,727,1908,1935]" degrees="0.175" direction="west" orientation="longitude" pageId="323" pageNumber="324" precision="55" value="-0.175">0.175</geoCoordinate>
(0.157); PW =
<geoCoordinate box="[912,974,1908,1934]" degrees="0.268" direction="north" orientation="latitude" pageId="323" pageNumber="324" precision="55" value="0.268">0.268</geoCoordinate>
<geoCoordinate box="[975,1052,1908,1934]" degrees="0.306" direction="west" orientation="longitude" pageId="323" pageNumber="324" precision="55" value="-0.306">0.306</geoCoordinate>
(0.290); SBPA =
<geoCoordinate box="[1260,1322,1908,1934]" degrees="0.116" direction="north" orientation="latitude" pageId="323" pageNumber="324" precision="55" value="0.116">0.116</geoCoordinate>
<geoCoordinate box="[1322,1399,1908,1934]" degrees="0.147" direction="west" orientation="longitude" pageId="323" pageNumber="324" precision="55" value="-0.147">0.147</geoCoordinate>
(0.133); SPTI =
<geoCoordinate box="[590,652,234,260]" degrees="0.147" direction="north" orientation="latitude" pageId="324" pageNumber="325" precision="55" value="0.147">0.147</geoCoordinate>
<geoCoordinate box="[653,730,234,260]" degrees="0.188" direction="west" orientation="longitude" pageId="324" pageNumber="325" precision="55" value="-0.188">0.188</geoCoordinate>
(0.165); PEW =
<geoCoordinate box="[932,994,234,260]" degrees="0.099" direction="north" orientation="latitude" pageId="324" pageNumber="325" precision="55" value="0.099">0.099</geoCoordinate>
<geoCoordinate box="[994,1071,234,260]" degrees="0.123" direction="west" orientation="longitude" pageId="324" pageNumber="325" precision="55" value="-0.123">0.123</geoCoordinate>
(0.110); PNW =
<geoCoordinate box="[1278,1340,234,260]" degrees="0.096" direction="north" orientation="latitude" pageId="324" pageNumber="325" precision="55" value="0.096">0.096</geoCoordinate>
<geoCoordinate box="[1340,1417,234,260]" degrees="0.120" direction="west" orientation="longitude" pageId="324" pageNumber="325" precision="55" value="-0.12">0.120</geoCoordinate>
(0.109); PPW =
<geoCoordinate box="[590,652,274,300]" degrees="0.236" direction="north" orientation="latitude" pageId="324" pageNumber="325" precision="55" value="0.236">0.236</geoCoordinate>
<geoCoordinate box="[653,730,274,300]" degrees="0.277" direction="west" orientation="longitude" pageId="324" pageNumber="325" precision="55" value="-0.277">0.277</geoCoordinate>
(0.258); HFL =
<geoCoordinate box="[917,979,274,300]" degrees="0.310" direction="north" orientation="latitude" pageId="324" pageNumber="325" precision="55" value="0.31">0.310</geoCoordinate>
<geoCoordinate box="[979,1056,274,300]" degrees="0.370" direction="west" orientation="longitude" pageId="324" pageNumber="325" precision="55" value="-0.37">0.370</geoCoordinate>
(0.348); HFWmax =
<geoCoordinate box="[1306,1368,274,300]" degrees="0.089" direction="north" orientation="latitude" pageId="324" pageNumber="325" precision="55" value="0.089">0.089</geoCoordinate>
<geoCoordinate box="[1368,1445,274,300]" degrees="0.131" direction="west" orientation="longitude" pageId="324" pageNumber="325" precision="55" value="-0.131">0.131</geoCoordinate>
(0.108); HFWmin =
<geoCoordinate box="[645,707,314,340]" degrees="0.036" direction="north" orientation="latitude" pageId="324" pageNumber="325" precision="55" value="0.036">0.036</geoCoordinate>
<geoCoordinate box="[708,785,314,341]" degrees="0.053" direction="west" orientation="longitude" pageId="324" pageNumber="325" precision="55" value="-0.053">0.053</geoCoordinate>
(0.044); CS =
<geoCoordinate box="[958,1020,314,341]" degrees="0.596" direction="north" orientation="latitude" pageId="324" pageNumber="325" precision="55" value="0.596">0.596</geoCoordinate>
<geoCoordinate box="[1021,1098,314,340]" degrees="0.702" direction="west" orientation="longitude" pageId="324" pageNumber="325" precision="55" value="-0.702">0.702</geoCoordinate>
(0.673); ES =
<geoCoordinate box="[1269,1331,314,340]" degrees="0.134" direction="north" orientation="latitude" pageId="324" pageNumber="325" precision="55" value="0.134">0.134</geoCoordinate>
<geoCoordinate box="[1331,1408,314,340]" degrees="0.177" direction="west" orientation="longitude" pageId="324" pageNumber="325" precision="55" value="-0.177">0.177</geoCoordinate>
(0.152); SI = 8799 (92); OI = 2125 (23); CI = 7684 (81); WLI = 129137 (133); SBI = 2934 (32); PSI = 2736 (32); PWI = 215255 (235); PLI = 128187 (157); NI = 140201 (162); PNWI = 89111 (100); NLI = 6077 (66); FI = 202285 (248).
</paragraph>
<paragraph blockId="324.[498,1542,234,1894]" pageId="324" pageNumber="325">
<emphasis bold="true" box="[530,775,473,499]" pageId="324" pageNumber="325">Worker description:</emphasis>
In full-face view, head subquadrate, longer than broad (CI 7684). Mandibles weakly striate, shining, and armed with five teeth: the apical-most well developed, followed by a smaller preapical tooth, which is followed by three equally developed smaller teeth. Anterior clypeal margin entire and evenly rounded. Antennal scapes short: when fully retracted, failing to reach the posterior margin of the head capsule by about the maximum width of the scape (SI 8799). Antennae 12-segmented; antennal club of three segments, with the apical-most segment one and a half times longer than the preceding two in combination. Frontal carinae short, extending past the antennal toruli by one and a half times the maximum width of the antennal scape. Compound eyes weakly protruding past the lateral margins of the head capsule. Lateral margin of head very weakly convex, nearly flat, giving the head a boxy appearance. Posterior head margin flat to very slightly concave, rounding evenly into the lateral margins.
</paragraph>
<paragraph blockId="324.[498,1542,234,1894]" pageId="324" pageNumber="325">
In profile view, compound eyes ovular and moderately large (OI 2125), with 9 ommatidia in longest row. Pronotal declivity distinct, with neck and anterior face of pronotum forming a rounded ~130
<emphasis box="[908,918,1027,1045]" italics="true" pageId="324" pageNumber="325">
<superScript attach="left" box="[908,918,1027,1045]" fontSize="8" pageId="324" pageNumber="325">°</superScript>
</emphasis>
angle; anterior and dorsal faces of pronotum forming a rounded ~140
<emphasis box="[686,696,1067,1085]" italics="true" pageId="324" pageNumber="325">
<superScript attach="left" box="[686,696,1067,1085]" fontSize="8" pageId="324" pageNumber="325">°</superScript>
</emphasis>
angle. Mesosoma evenly, but weakly, convex dorsally from where it joins the pronotal declivity to the propodeal spines, nearly flat. Promesonotal suture extending from the posterior margin of the procoxal insertion only to the mesothoracic spiracle, which is well developed. Metanotal groove nearly entirely absent: only visible as a faint disruption in the surface sculpture between meso- and metacoxal insertions to the minute metathoracic spiracle, which is nearly indistinguishable against the ground sculpture. Propodeal spiracle weakly developed, directed posterolaterally, and separated from the propodeal declivity by about three and a half spiracle diameters. Propodeal spines well developed, moderately long (PSI 2736), about as long as the propodeal declivity, tapering evenly from the base, very weakly downcurved, and acute. Propodeal declivity straight and flat, forming a rounded ~100
<emphasis box="[991,1001,1465,1483]" italics="true" pageId="324" pageNumber="325">
<superScript attach="left" box="[991,1001,1465,1483]" fontSize="8" pageId="324" pageNumber="325">°</superScript>
</emphasis>
angle with the base of the propodeal spines. Propodeal lobes rounded and very weakly developed. Metapleural gland bulla small, extending from the metacoxal insertion halfway to the propodeal spiracle. Petiole moderately long (PLI 128187), without tubercles anterodorsally. Subpetiolar process in the form of a weakly developed, acute, triangular tooth; ventral margin of petiole strongly concave posterior to it. Petiolar peduncle short, comprising about a quarter of the total length of the petiole. Petiolar node robust: grading evenly into the petiolar peduncle anteriorly, anterior face concave; anterior face forming a rounded, ~90
<emphasis box="[1331,1341,1744,1762]" italics="true" pageId="324" pageNumber="325">
<superScript attach="left" box="[1331,1341,1744,1762]" fontSize="8" pageId="324" pageNumber="325">°</superScript>
</emphasis>
angle with the dorsal face, which is evenly, strongly convex; dorsal face rounding evenly into the posterior face, which forms a ~110
<emphasis box="[796,806,1824,1842]" italics="true" pageId="324" pageNumber="325">
<superScript attach="left" box="[796,806,1824,1842]" fontSize="8" pageId="324" pageNumber="325">°</superScript>
</emphasis>
angle with the caudal cylinder. Postpetiole evenly rounded anterodorsally, before flattening posterodorsally; weakly lobed ventrally.
</paragraph>
<paragraph blockId="325.[498,1542,234,1934]" pageId="325" pageNumber="326">In dorsal view, humeri weakly developed: rounded and only slightly wider than the rest of the mesosoma; mesothoracic spiracles weakly protruding past the lateral margins of the mesosoma, visible as slight angles where the pronotum meets the mesonotum. Metanotal groove absent: mesonotum and propodeum completely fused and converging evenly to the bases of the propodeal spines. Propodeal spines broadly approximated basally and diverging apically, their apices separated from each other about their length; negative space between them a “U”. Petiolar peduncle with spiracles barely protruding past the lateral margins; not constricted anterior to them. Petiolar node ovular, subtly narrowed anteriorly, slightly wider than the peduncle, and evenly grading into the caudal cylinder, which is the same width as the node. Postpetiole moderately to very broad (PWI 215255) and campaniform, articulating with nearly the entire anterior margin of the gaster, leaving small angulate margins on each side exposed. Anterior margin of the postpetiole flat, with corners evenly rounding into the lateral margins, which evenly diverge to the angulate posterior corners; posterior margin broadly concave. Metafemur weakly to moderately incrassate (FI 202285).</paragraph>
<paragraph blockId="325.[498,1542,234,1934]" pageId="325" pageNumber="326">Sculpture: median lobe of clypeus compressed between the closely approximated frontal lobes. Median clypeal carina present, flanked two additional equally developed carinae over smooth and shining ground sculpture. Antennal scapes shining through weak sculpture. Cephalic dorsum predominantly smooth and shining, with weak costulate sculpture medial to the frontal carinae, faint areolate sculpture surrounding the compound eyes and between the eyes and the mandibular insertions. Pronotal neck areolate. Mesosoma with lateral surfaces and dorsum of propodeum shining through weak areolate sculpture; dorsum of promesonotum and propodeal declivity smooth and shining. Femora shining through very weak areolate sculpture. Petiole smooth and shining ventrally, with shallow areolate sculpture surrounding the base of the petiolar node, which is mostly smooth and shining dorsally. Dorsal surface of postpetiole smooth and shining, with shallow areolate sculpture on the posterior quarter areolate. Gaster smooth and shining, without spectral iridescence on the first tergite. Surface of the first gastral sternite smooth and shining.</paragraph>
<paragraph blockId="325.[498,1542,234,1934]" pageId="325" pageNumber="326">Setae: antennal scapes and funiculi with short, adpressed pilosity. Dorsum of head, mesosoma, waist segments and gaster with short, erect, blunt-tipped setae, some of which are nearly clavate; the longest setae are roughly the width of the compound eye. The head bears ~32, mesosoma ~16, petiole 2, postpetiole ~10, and first gastral tergite ~26 setae. Sparse, adpressed pubescence present on the entire body, but only apparent on the darker colored integument.</paragraph>
<paragraph blockId="325.[498,1542,234,1934]" pageId="325" pageNumber="326">Color: head, mesosoma, waist segments, and gaster are uniformly testaceous. Mandibles and extremities are light brown.</paragraph>
<paragraph blockId="325.[498,1542,234,1934]" box="[530,914,1708,1735]" pageId="325" pageNumber="326">
<emphasis bold="true" box="[530,601,1708,1734]" pageId="325" pageNumber="326">Gyne:</emphasis>
Too damaged to describe.
</paragraph>
<paragraph blockId="325.[498,1542,234,1934]" box="[530,730,1748,1775]" pageId="325" pageNumber="326">
<emphasis bold="true" box="[530,597,1749,1775]" pageId="325" pageNumber="326">Male:</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="325" pageNumber="326" type="etymology">
<paragraph blockId="325.[498,1542,234,1934]" pageId="325" pageNumber="326">
<emphasis bold="true" box="[530,668,1788,1814]" pageId="325" pageNumber="326">Etymology:</emphasis>
Behavioural, from the Latin terra (= earth) + colere (= dwelling), in reference to the nesting habitat of this species.
</paragraph>
</subSubSection>
<subSubSection lastPageId="326" lastPageNumber="327" pageId="325" pageNumber="326" type="discussion">
<paragraph blockId="325.[498,1542,234,1934]" lastBlockId="326.[498,1542,234,1098]" lastPageId="326" lastPageNumber="327" pageId="325" pageNumber="326">
<emphasis bold="true" box="[530,670,1868,1894]" pageId="325" pageNumber="326">Comments:</emphasis>
<taxonomicName baseAuthorityName="Mann" baseAuthorityYear="1920" box="[679,940,1868,1894]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="325" pageNumber="326" phylum="Arthropoda" rank="species" species="terricola">
<emphasis box="[679,940,1868,1894]" italics="true" pageId="325" pageNumber="326">Temnothorax terricola</emphasis>
</taxonomicName>
is mostly known from the
<collectingRegion box="[1264,1499,1868,1894]" country="Cuba" name="Guantanamo" pageId="325" pageNumber="326">Guantanamo region</collectingRegion>
in eastern
<collectingCountry box="[588,653,1908,1934]" name="Cuba" pageId="325" pageNumber="326">Cuba</collectingCountry>
. The
<typeStatus box="[715,765,1908,1934]" pageId="325" pageNumber="326">type</typeStatus>
series was collected from under a stone in low elevation pine forest near Baracoa. Two more series were collected by Phil Ward: stray ground foragers were collected at bread crumb baits on the edge of disturbed rainforest just north of Baracoa, and several more workers were recovered from Winkler leaf litter extraction in mid-elevation mixed
<taxonomicName box="[750,918,354,380]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="326" pageNumber="327" phylum="Tracheophyta" rank="species" species="cubensis">
<emphasis box="[750,918,354,380]" italics="true" pageId="326" pageNumber="327">Pinus cubensis</emphasis>
</taxonomicName>
and broadleaf evergreen forest. This species was reported from the Nipe-Sagua-Baracoa mountain range in
<collectingCountry box="[1193,1255,394,420]" name="Cuba" pageId="326" pageNumber="327">Cuba</collectingCountry>
in a survey by
<bibRefCitation author="Ferrer EP &amp; Triana JL" journalOrPublisher="Boletin de la SEA" pageId="326" pageNumber="327" pagination="121 - 136" part="35" refId="ref270872" refString="Ferrer EP, Triana JL. 2004. Biodiversidad del orden Hymenoptera en los macizos montanosos de Cuba oriental. Boletin de la SEA 35: 121 - 136." title="Biodiversidad del orden Hymenoptera en los macizos montanosos de Cuba oriental" type="journal article" year="2004">
<emphasis italics="true" pageId="326" pageNumber="327">Ferrer &amp; Triana (2004)</emphasis>
</bibRefCitation>
, which is consistent with the previous collections; I have not been able to personally examine these specimens, however. This species, like other members of the
<taxonomicName baseAuthorityName="Emery" baseAuthorityYear="1894" box="[498,611,513,539]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="326" pageNumber="327" phylum="Arthropoda" rank="species" species="pulchellus">
<emphasis box="[498,611,513,539]" italics="true" pageId="326" pageNumber="327">pulchellus</emphasis>
</taxonomicName>
group, is probably ground or leaf litter nesting. The original description by
<bibRefCitation author="Mann WM" box="[498,649,553,579]" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="326" pageNumber="327" pagination="403 - 439" part="42" refId="ref271797" refString="Mann WM. 1920. Additions to the ant fauna of the West Indies and Central America. Bulletin of the American Museum of Natural History 42: 403 - 439." title="Additions to the ant fauna of the West Indies and Central America" type="journal article" year="1920">
<emphasis box="[498,649,553,579]" italics="true" pageId="326" pageNumber="327">Mann (1920)</emphasis>
</bibRefCitation>
notes that this species has a dark brown to black head and gaster. Perhaps the intensity of the integumental coloration in the
<typeStatus box="[1052,1102,593,619]" pageId="326" pageNumber="327">type</typeStatus>
series has faded with time, but all of the specimens from the
<typeStatus box="[784,834,633,659]" pageId="326" pageNumber="327">type</typeStatus>
series that I have observed are dark to medium brown. The two series collected by Phil Ward are substantially lighter, with the head and gaster medium brown, while the rest of the body is light brown; the lateral surface of the pronotum is also slightly less densely sculptured than the
<typeStatus box="[1159,1209,753,779]" pageId="326" pageNumber="327">type</typeStatus>
series, shining through weak areolate sculpture. This species has long been suspected to have close relationships to other members of the
<taxonomicName baseAuthorityName="Forel" baseAuthorityYear="1899" box="[759,832,832,858]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="326" pageNumber="327" phylum="Arthropoda" rank="species" species="salvini">
<emphasis box="[759,832,832,858]" italics="true" pageId="326" pageNumber="327">salvini</emphasis>
</taxonomicName>
clade, specifically members of the
<taxonomicName baseAuthorityName="Emery" baseAuthorityYear="1894" box="[1235,1346,832,858]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="326" pageNumber="327" phylum="Arthropoda" rank="species" species="pulchellus">
<emphasis box="[1235,1346,832,858]" italics="true" pageId="326" pageNumber="327">pulchellus</emphasis>
</taxonomicName>
group:
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1908" box="[498,641,872,898]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="326" pageNumber="327" phylum="Arthropoda" rank="species" species="albispinus">
<emphasis box="[498,641,872,898]" italics="true" pageId="326" pageNumber="327">T. albispinus</emphasis>
</taxonomicName>
,
<emphasis box="[653,908,872,898]" italics="true" pageId="326" pageNumber="327">
<taxonomicName baseAuthorityName="Menozzi &amp; Russo" baseAuthorityYear="1930" box="[653,759,872,898]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="326" pageNumber="327" phylum="Arthropoda" rank="species" species="ciferrii">T. ciferrii</taxonomicName>
,
<taxonomicName box="[768,908,872,898]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="326" pageNumber="327" phylum="Arthropoda" rank="species" species="flavidulus">T. flavidulus</taxonomicName>
</emphasis>
, and
<taxonomicName baseAuthorityName="Emery" baseAuthorityYear="1894" box="[970,1109,872,898]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="326" pageNumber="327" phylum="Arthropoda" rank="species" species="pulchellus">
<emphasis box="[970,1109,872,898]" italics="true" pageId="326" pageNumber="327">T. pulchellus</emphasis>
</taxonomicName>
. In the first formal morphology-based phylogenetic work including this species,
<bibRefCitation author="Fontenla Rizo JL" box="[978,1217,912,938]" journalOrPublisher="Avicennia" pageId="326" pageNumber="327" pagination="35 - 44" part="13" refId="ref270970" refString="Fontenla Rizo JL. 2000. Definition, phylogenetic relationships and morphological species groups of taxon &quot; Macromischa &quot; (Hymenoptera: Formicidae: Leptothorax). Avicennia 13: 35 - 44." title="Definition, phylogenetic relationships and morphological species groups of taxon &quot; Macromischa &quot; (Hymenoptera: Formicidae: Leptothorax)" type="journal article" year="2000">
<emphasis box="[978,1217,912,938]" italics="true" pageId="326" pageNumber="327">Fontenla Rizo (2000)</emphasis>
</bibRefCitation>
, while not including all of the taxa in Baroni Urbani s
<taxonomicName baseAuthorityName="Emery" baseAuthorityYear="1894" box="[835,951,951,977]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="326" pageNumber="327" phylum="Arthropoda" rank="species" species="pulchellus">
<emphasis box="[835,951,951,977]" italics="true" pageId="326" pageNumber="327">pulchellus</emphasis>
</taxonomicName>
group, corroborated these relationships to some degree, with the caveat that
<taxonomicName baseAuthorityName="Mann" baseAuthorityYear="1920" box="[827,956,991,1017]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="326" pageNumber="327" phylum="Arthropoda" rank="species" species="terricola">
<emphasis box="[827,956,991,1017]" italics="true" pageId="326" pageNumber="327">T. terricola</emphasis>
</taxonomicName>
and
<taxonomicName baseAuthorityName="Aguayo" baseAuthorityYear="1931" box="[1014,1110,992,1017]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="326" pageNumber="327" phylum="Arthropoda" rank="species" species="torrei">
<emphasis box="[1014,1110,992,1017]" italics="true" pageId="326" pageNumber="327">T. torrei</emphasis>
</taxonomicName>
fell outside of this group. Combined molecular and morphological analyses (Prebus, in prep.) mostly agree with
<bibRefCitation author="Fontenla Rizo JL" journalOrPublisher="Avicennia" pageId="326" pageNumber="327" pagination="35 - 44" part="13" refId="ref270970" refString="Fontenla Rizo JL. 2000. Definition, phylogenetic relationships and morphological species groups of taxon &quot; Macromischa &quot; (Hymenoptera: Formicidae: Leptothorax). Avicennia 13: 35 - 44." title="Definition, phylogenetic relationships and morphological species groups of taxon &quot; Macromischa &quot; (Hymenoptera: Formicidae: Leptothorax)" type="journal article" year="2000">
<emphasis italics="true" pageId="326" pageNumber="327">Fontenla Rizo (2000)</emphasis>
</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>