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<document id="AC1901EA25A27A10F5FD07C362FAB083" ID-DOI="10.5281/zenodo.5393502" ID-ISSN="1638-9387" ID-Zenodo-Dep="5393502" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630318937109" checkinUser="marcus" docAuthor="Golovatch, Sergei I., Geoffroy, Jean-Jacques &amp; Mauriès, Jean-Paul" docDate="2006" docId="03FEF24BA9706A45FCACFD591B6DC465" docLanguage="en" docName="Zoosystema.2006.28.1.75-92.pdf" docOrigin="Zoosystema 28 (1)" docStyle="DocumentStyle:2A9C6F9EA5F90D98170D9E0D9CA29375.4:Zoosystema.2000-2008.journal_article.type1" docStyleId="2A9C6F9EA5F90D98170D9E0D9CA29375" docStyleName="Zoosystema.2000-2008.journal_article.type1" docStyleVersion="4" docTitle="Nepalella grandis Golovatch &amp; Geoffroy &amp; Mauriès 2006, n. sp." docType="treatment" docVersion="3" lastPageNumber="83" masterDocId="FFC78A33A9756A4DFF9CFFAB1F53C162" masterDocTitle="Four new Chordeumatida (Diplopoda) from caves in China" masterLastPageNumber="92" masterPageNumber="75" pageNumber="80" updateTime="1699225294555" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="A8BBD9E165A244EFACF75FEF1830D22C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="94922AA94C2605461CE271C758562306">
<mods:title id="5A8857EF4BE6DCA703C83B1A1C411E15">Four new Chordeumatida (Diplopoda) from caves in China</mods:title>
</mods:titleInfo>
<mods:name id="BBC6E546AC07CF643AE781BDE3717869" type="personal">
<mods:role id="D4CBB0983821B2D0ED1221011F07B4F7">
<mods:roleTerm id="0FE50CF6A75888706415639C5D7EB290">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="38D2934E090CE64D5AF2BF4D9CEE6CC1">Golovatch, Sergei I.</mods:namePart>
<mods:affiliation id="5B7FD7FA060227C1DD1CF4AE6DB8B79F">Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071 (Russia) sgol @ orc. ru</mods:affiliation>
<mods:nameIdentifier id="963E75240D0A6578DD17AAF41BE1F5D0" type="email">sgol@orc.ru</mods:nameIdentifier>
</mods:name>
<mods:name id="0DD8037D927F9E86D7191C8CE979A75D" type="personal">
<mods:role id="C053ABB8622B0FBE06964EE080CE370D">
<mods:roleTerm id="0967F4C9A36371B9EFC99CAA29599B59">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="49E053EB9772B92E40B10FA5F6993747">Geoffroy, Jean-Jacques</mods:namePart>
<mods:affiliation id="4976CCCB4A9D008D5715237F1C48E6B0">Muséum national d’Histoire naturelle, Département Écologie et Gestion de la Biodiversité, USM 306 Réseaux trophiques du Sol, 4 avenue du Petit Château, F- 91800 Brunoy (France) geoffroy @ mnhn. fr</mods:affiliation>
<mods:nameIdentifier id="98D5A179685A75020B741156CFA448BB" type="email">geoffroy@mnhn.fr</mods:nameIdentifier>
</mods:name>
<mods:name id="6FE3AAF023837C4EBB08B03AA1B82A9A" type="personal">
<mods:role id="42767458ABA3CC18ED3C4C5F0AD56BD4">
<mods:roleTerm id="8C0C9C52F8A2D6B15B2B940FF3BED49F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="213950F3CE67C45C9226DE80651C1E1E">Mauriès, Jean-Paul</mods:namePart>
<mods:affiliation id="ABD8F43DBA4E2B9FE43E9E182A45DB82">Muséum national d’Histoire naturelle, Département Systématique et Évolution, USM 602 Taxonomie et Collections, case postale 53, 61 rue Buffon, F- 75231 Paris cedex 05 (France) collmill @ mnhn. fr</mods:affiliation>
<mods:nameIdentifier id="D53F1839746ADE2DDC5BC4D320810A67" type="email">collmill@mnhn.fr</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="07EB0B057EFD6B6DF55D3EC52035CFE3">text</mods:typeOfResource>
<mods:relatedItem id="A07D1C7816B26CB0E6466BEFE9B84F22" type="host">
<mods:titleInfo id="DBF4A02E865D4A8F6F80AFE943AF9593">
<mods:title id="CF797018B969EE1937A6C31385364FED">Zoosystema</mods:title>
</mods:titleInfo>
<mods:part id="3C2B5CEF7834BBB7F06B6E501A3C5A9C">
<mods:date id="1FBE8B1F11F748EDAE8FBF732CC84759">2006</mods:date>
<mods:detail id="78E934045803797119F591F48212A0FF" type="volume">
<mods:number id="D4E1A493B66B132DDD447C8CC6B40DDD">28</mods:number>
</mods:detail>
<mods:detail id="E16B9BD2E66D642A86BAD1C0F94B701F" type="issue">
<mods:number id="6D3A5CA62F23487EDDFE66955F57CB02">1</mods:number>
</mods:detail>
<mods:extent id="F6B2DFE56960B9B18D343FE3742F02EE" unit="page">
<mods:start id="273594BB03B61442DD911BFF8E5085C0">75</mods:start>
<mods:end id="5F9206BC14066B2D32300BC18B7687BE">92</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="C85192431C2794A8DDF6F30EB023C912">journal article</mods:classification>
<mods:identifier id="2ED708EF8775167FEBADD6730B98C6CE" type="DOI">10.5281/zenodo.5393502</mods:identifier>
<mods:identifier id="56B7C9660F59BBAA71BD507E1D69D1BC" type="ISSN">1638-9387</mods:identifier>
<mods:identifier id="4C9856A425266821151C66D62E1CC491" type="Zenodo-Dep">5393502</mods:identifier>
</mods:mods>
<treatment id="03FEF24BA9706A45FCACFD591B6DC465" LSID="urn:lsid:plazi:treatment:03FEF24BA9706A45FCACFD591B6DC465" httpUri="http://treatment.plazi.org/id/03FEF24BA9706A45FCACFD591B6DC465" lastPageId="8" lastPageNumber="83" pageId="5" pageNumber="80">
<subSubSection id="C34D10D6A9706A48FCACFD591B7DC26F" box="[816,1070,754,781]" pageId="5" pageNumber="80" type="nomenclature">
<paragraph id="8BE8435DA9706A48FCACFD591B7DC26F" blockId="5.[816,1070,754,812]" box="[816,1070,754,781]" pageId="5" pageNumber="80">
<heading id="D0A0F431A9706A48FCACFD591B7DC26F" box="[816,1070,754,781]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="80" reason="2">
<taxonomicName id="4C5738DEA9706A48FCACFD591CBEC26F" ID-CoL="46WVR" authority="Golovatch &amp; Geoffroy &amp; Mauriès, 2006" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès" authorityYear="2006" box="[816,1005,754,781]" class="Diplopoda" family="Lankasomatidae" genus="Nepalella" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="80" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
<emphasis id="B9239F4FA9706A48FCACFD591CBEC26F" bold="true" box="[816,1005,754,781]" italics="true" pageId="5" pageNumber="80">Nepalella grandis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2102234A9706A48FC6FFD581B7DC26F" box="[1011,1070,755,781]" pageId="5" pageNumber="80" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34D10D6A9706A48FCC2FCB91B53C24E" box="[862,1024,786,812]" pageId="5" pageNumber="80" type="description">
<paragraph id="8BE8435DA9706A48FCC2FCB91B53C24E" blockId="5.[816,1070,754,812]" box="[862,1024,786,812]" pageId="5" pageNumber="80">
(
<figureCitation id="136C5FD8A9706A48FCF9FCB91CF1C24E" box="[869,930,786,812]" captionStart="FIG" captionStartId="3.[112,123,1411,1428]" captionTargetBox="[127,1195,217,1368]" captionTargetId="figure-93@3.[89,1232,156,1420]" captionTargetPageId="3" captionText="FIG. 1. — A-C, Nepalella marmorata n. sp., paratype ♂; D-F, N. grandis n. sp., paratype ♂; G, N. grandis n. sp., paratype ♀; A, D, anterior body end, dorsal view; B, E, midbody segments, dorsal view; C, F, same, lateral view; G, vulva, front view. Scale bars: A-F, 2.0 mm; G, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.5393504" httpUri="https://zenodo.org/record/5393504/files/figure.png" pageId="5" pageNumber="80">Figs 1</figureCitation>
D-G; 4)
</paragraph>
</subSubSection>
<subSubSection id="C34D10D6A9706A48FD39FCE71C01C285" pageId="5" pageNumber="80" type="materials_examined">
<paragraph id="8BE8435DA9706A48FD39FCE71C01C285" blockId="5.[677,1210,843,999]" pageId="5" pageNumber="80">
<materialsCitation id="3B3F4900A9706A48FD39FCE71BE6C2D1" collectingDate="1999-08-18" collectionCode="MNHN" collectorName="J. &amp; B. Lips" country="China" county="Zheng Xiong County" location="Nos." municipality="Cave Bai Yin Dong" pageId="5" pageNumber="80" specimenCode="DB42" specimenCount="2" specimenCount-male="2" stateProvince="Yunnan" typeStatus="holotype">
TYPE MATERIAL. —
<typeStatus id="54ECFDFFA9706A48FCECFCE01C80C201" box="[880,979,843,867]" pageId="5" pageNumber="80" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F34003CDA9706A48FC42FCE01B4CC201" box="[990,1055,843,867]" name="China" pageId="5" pageNumber="80">China</collectingCountry>
,
<collectingRegion id="49938DBFA9706A48FBB7FCE71BE5C201" box="[1067,1206,844,867]" country="China" name="Yunnan" pageId="5" pageNumber="80">Yunnan Prov.</collectingRegion>
,
<collectingCounty id="62893BD1A9706A48FD39FCCE1C2EC21C" box="[677,893,869,894]" pageId="5" pageNumber="80">Zheng Xiong County</collectingCounty>
,
<collectingMunicipality id="6B8CD927A9706A48FC16FCCD1B03C21C" box="[906,1104,869,894]" pageId="5" pageNumber="80">Cave Bai Yin Dong</collectingMunicipality>
,
<location id="8E881586A9706A48FBC1FCCD1BDFC21F" LSID="urn:lsid:plazi:treatment:03FEF24BA9706A45FCACFD591B6DC465:8E881586A9706A48FBC1FCCD1BDFC21F" box="[1117,1164,870,893]" country="China" county="Zheng Xiong County" municipality="Cave Bai Yin Dong" name="Nos." pageId="5" pageNumber="80" stateProvince="Yunnan">Nos.</location>
523 &amp; 524,
<date id="FFE9659DA9706A48FD6BFC2B1C2CC2FA" box="[759,895,896,920]" pageId="5" pageNumber="80" value="1999-08-18">
<collectingDate id="EFAD9C75A9706A48FD6BFC2B1C2CC2FA" box="[759,895,896,920]" pageId="5" pageNumber="80" value="1999-08-18">18.VIII.1999</collectingDate>
</date>
, leg.
<collectorName id="26A2268BA9706A48FC2AFC2B1B66C2FA" box="[950,1077,896,920]" pageId="5" pageNumber="80">J. &amp; B. Lips</collectorName>
,
<specimenCount id="9D5188D4A9706A48FBDDFC281B06C2F5" box="[1089,1109,899,919]" pageId="5" pageNumber="80" type="male">♂</specimenCount>
(
<collectionCode id="ED46DB98A9706A48FBF8FC2B1BEAC2FA" box="[1124,1209,896,920]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="80">MNHN</collectionCode>
<specimenCode id="DBF1EB26A9706A48FD39FC301DB7C2D1" box="[677,740,923,947]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="80">DB42</specimenCode>
).
<typeStatus id="54ECFDFFA9706A48FD68FC301C00C2D1" box="[756,851,923,947]" pageId="5" pageNumber="80" type="paratype">Paratypes</typeStatus>
: same locality,
<specimenCount id="9D5188D4A9706A48FC7CFC301B5AC2D0" box="[992,1033,923,947]" pageId="5" pageNumber="80" type="male">1 ♂</specimenCount>
(
<collectionCode id="ED46DB98A9706A48FB89FC301B3AC2D1" box="[1045,1129,923,947]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="80">MNHN</collectionCode>
<specimenCode id="DBF1EB26A9706A48FBF3FC301BFDC2D1" box="[1135,1198,923,947]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="80">DB42</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B3F4900A9706A48FD39FC1E1CD0C2AF" box="[677,899,949,973]" collectingDate="1999-08-18" collectionCode="MNHN" collectorName="J. &amp; B. Lips" country="China" county="Zheng Xiong County" isTailEnumeration="true" location="Nos." municipality="Cave Bai Yin Dong" pageId="5" pageNumber="80" specimenCode="DB42" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="holotype">
<specimenCount id="9D5188D4A9706A48FD39FC1E1D9FC2AE" box="[677,716,949,973]" pageId="5" pageNumber="80" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED46DB98A9706A48FD40FC1E1C61C2AF" box="[732,818,949,973]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="80">MNHN</collectionCode>
<specimenCode id="DBF1EB26A9706A48FCA7FC1E1C2FC2AF" box="[827,892,949,973]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="80">DB42</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B3F4900A9706A48FC0CFC1E1B3CC2AF" box="[912,1135,949,973]" collectionCode="MNHN" country="China" pageId="5" pageNumber="80" specimenCode="DB42" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="holotype">
<specimenCount id="9D5188D4A9706A48FC0CFC1E1CEBC2AE" box="[912,952,949,973]" pageId="5" pageNumber="80" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED46DB98A9706A48FC54FC1E1B4DC2AF" box="[968,1054,949,973]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="80">MNHN</collectionCode>
<specimenCode id="DBF1EB26A9706A48FBBAFC1E1B3BC2AF" box="[1062,1128,949,973]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="80">DB42</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B3F4900A9706A48FBE0FC1E1C1DC285" collectionCode="MNHN" country="China" pageId="5" pageNumber="80" specimenCode="DB42" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
6 juvs (
<collectionCode id="ED46DB98A9706A48FD30FC641C53C285" box="[684,768,975,999]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="80">MNHN</collectionCode>
<specimenCode id="DBF1EB26A9706A48FC9AFC641C15C285" box="[774,838,975,999]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="80">DB42</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34D10D6A9706A48FD39FC541BEBC575" box="[677,1208,1023,1047]" pageId="5" pageNumber="80" type="etymology">
<paragraph id="8BE8435DA9706A48FD39FC541BEBC575" blockId="5.[677,1208,1023,1047]" box="[677,1208,1023,1047]" pageId="5" pageNumber="80">ETYMOLOGY. — To emphasize the particularly large size.</paragraph>
</subSubSection>
<subSubSection id="C34D10D6A9706A48FD39FB841B4EC5D2" pageId="5" pageNumber="80" type="diagnosis">
<paragraph id="8BE8435DA9706A48FD39FB841B4EC5D2" blockId="5.[677,1210,1071,1200]" pageId="5" pageNumber="80">
DIAGNOSIS. — Differs from congeners by the particularly large size, as well as certain details of leg and gonopod structure. The unusually strong colpocoxites, surmounted by a pseudosegmented head-like structure, also make
<taxonomicName id="4C5738DEA9706A48FD39FB321C54C5D2" authority="Golovatch &amp; Geoffroy &amp; Mauriès, 2006" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès" authorityYear="2006" box="[677,775,1176,1200]" class="Diplopoda" family="Lankasomatidae" genus="Nepalella" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="80" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
<emphasis id="B9239F4FA9706A48FD39FB321C54C5D2" box="[677,775,1176,1200]" italics="true" pageId="5" pageNumber="80">N. grandis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2102234A9706A48FC91FB321C10C5D2" box="[781,835,1177,1200]" pageId="5" pageNumber="80" rank="species">n. sp.</taxonomicNameLabel>
easily distinguishable.
</paragraph>
</subSubSection>
<subSubSection id="C34D10D6A9706A45FD39FB651C99C012" lastPageId="8" lastPageNumber="83" pageId="5" pageNumber="80" type="description">
<paragraph id="8BE8435DA9706A48FD39FB651C66C584" blockId="5.[677,1210,1230,1700]" box="[677,821,1230,1256]" pageId="5" pageNumber="80">DESCRIPTION</paragraph>
<paragraph id="8BE8435DA9706A48FD39FB461DBAC425" blockId="5.[677,1210,1230,1700]" pageId="5" pageNumber="80">
Length
<emphasis id="B9239F4FA9706A48FD64FB451C5AC465" box="[760,777,1262,1287]" italics="true" pageId="5" pageNumber="80">c.</emphasis>
40 (
<specimenCount id="9D5188D4A9706A48FCA6FB5A1C03C465" box="[826,848,1265,1287]" pageId="5" pageNumber="80" type="male">♂</specimenCount>
) to
<quantity id="4CAFEEB8A9706A48FCE1FB461C98C46A" box="[893,971,1261,1288]" metricMagnitude="-2" metricUnit="m" metricValue="4.2" pageId="5" pageNumber="80" unit="mm" value="42.0">42 mm</quantity>
(
<specimenCount id="9D5188D4A9706A48FC47FB5A1CBDC465" box="[987,1006,1265,1287]" pageId="5" pageNumber="80" type="female">♀</specimenCount>
), maximum width 2.8 (
<specimenCount id="9D5188D4A9706A48FD46FABB1DA3C444" box="[730,752,1296,1318]" pageId="5" pageNumber="80" type="male">♂</specimenCount>
) to 2.9-3.0 mm (
<specimenCount id="9D5188D4A9706A48FC26FABB1C9EC444" box="[954,973,1296,1318]" pageId="5" pageNumber="80" type="female">♀</specimenCount>
). Coloration entirely pallid.
</paragraph>
<paragraph id="8BE8435DA9706A48FD27FAE71B6FC4C4" blockId="5.[677,1210,1230,1700]" pageId="5" pageNumber="80">Head width = segment 6 = 20 &gt;&gt; collum&gt; segment 2&gt; 3&gt; 5 &lt;7; after segment 20 or 21, body very gradually tapering toward telson.</paragraph>
<paragraph id="8BE8435DA9706A48FD27FA001C09C7C6" blockId="5.[677,1210,1230,1700]" pageId="5" pageNumber="80">
Body with 30 segments. Head densely setose, clypeolabral region slightly convex. Eye patches triangular, each composed of 10 (
<specimenCount id="9D5188D4A9706A48FC71FA451B53C766" box="[1005,1024,1518,1540]" pageId="5" pageNumber="80" type="female">♀</specimenCount>
) or 15 (
<specimenCount id="9D5188D4A9706A48FBC9FA451B38C766" box="[1109,1131,1518,1540]" pageId="5" pageNumber="80" type="male">♂</specimenCount>
) rather convex ocelli. Antennae extremely long and slender (
<figureCitation id="136C5FD8A9706A48FD30F9811C55C726" box="[684,774,1578,1604]" captionStart="FIG" captionStartId="3.[112,123,1411,1428]" captionTargetBox="[127,1195,217,1368]" captionTargetId="figure-93@3.[89,1232,156,1420]" captionTargetPageId="3" captionText="FIG. 1. — A-C, Nepalella marmorata n. sp., paratype ♂; D-F, N. grandis n. sp., paratype ♂; G, N. grandis n. sp., paratype ♀; A, D, anterior body end, dorsal view; B, E, midbody segments, dorsal view; C, F, same, lateral view; G, vulva, front view. Scale bars: A-F, 2.0 mm; G, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.5393504" httpUri="https://zenodo.org/record/5393504/files/figure.png" pageId="5" pageNumber="80">Figs 1D</figureCitation>
;
<figureCitation id="136C5FD8A9706A48FC89F9811C6BC726" box="[789,824,1578,1604]" captionStart="FIG" captionStartId="7.[113,124,1458,1475]" captionTargetBox="[46,1277,131,1453]" captionTargetId="figure-88@7.[46,1277,131,1453]" captionTargetPageId="7" captionText="FIG. 4. — Nepalella grandis n. sp., paratype ♂: A, antenna; B, leg 7, caudal view; C, leg 10, front view; D, leg 11, front view; E, leg 12, front view; F, anterior gonopods (leg-pair 8), caudal view; G, H, posterior gonopods (leg-pair 9), front and caudal views, respectively. Scale bar: A-E, 0.8 mm; F-H, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.5393512" httpUri="https://zenodo.org/record/5393512/files/figure.png" pageId="5" pageNumber="80">4A</figureCitation>
), reaching beyond body segment 8 dorsally; antennomere 7 with a peculiar flattened seta dorsally in distal third. Gnathochilarium without promentum.
</paragraph>
<caption id="DF2813D5A9736A4BFFEDFA671F8EC491" ID-DOI="http://doi.org/10.5281/zenodo.5393508" ID-Zenodo-Dep="5393508" httpUri="https://zenodo.org/record/5393508/files/figure.png" pageId="6" pageNumber="81" startId="6.[113,124,1484,1501]" targetBox="[176,1108,220,1435]" targetPageId="6">
<paragraph id="8BE8435DA9736A4BFFEDFA671F8EC491" blockId="6.[113,1209,1484,1523]" pageId="6" pageNumber="81">
FIG. 3. —
<taxonomicName id="4C5738DEA9736A4BFF5EFA671E3EC4BF" authority="Golovatch &amp; Geoffroy &amp; Mauriès, 2006" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès" authorityYear="2006" box="[194,365,1484,1501]" class="Diplopoda" family="Lankasomatidae" genus="Nepalella" kingdom="Animalia" order="Chordeumatida" pageId="6" pageNumber="81" phylum="Arthropoda" rank="species" species="marmorata" status="sp. nov.">
<emphasis id="B9239F4FA9736A4BFF5EFA671E3EC4BF" box="[194,365,1484,1501]" italics="true" pageId="6" pageNumber="81">Nepalella marmorata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2102234A9736A4BFEEEFA671EF6C4BF" box="[370,421,1484,1501]" pageId="6" pageNumber="81" rank="species">n. sp.</taxonomicNameLabel>
, paratype ♂, posterior gonopods (leg-pair 9), front (
<emphasis id="B9239F4FA9736A4BFCCDFA671C0DC4BF" bold="true" box="[849,862,1484,1501]" pageId="6" pageNumber="81">A</emphasis>
) and caudal (
<emphasis id="B9239F4FA9736A4BFC52FA671C88C4BF" bold="true" box="[974,987,1484,1501]" pageId="6" pageNumber="81">B</emphasis>
) views, respectively. Scale bar: 0.4 mm.
</paragraph>
</caption>
<paragraph id="8BE8435DA9736A4BFF14F9811C98C7C1" blockId="6.[113,646,1578,1699]" lastBlockId="6.[677,1210,1578,1699]" pageId="6" pageNumber="81">
Collum usual, obcordate in shape, with rudimentary paraterga (
<figureCitation id="136C5FD8A9736A4BFED6F9E11EF2C706" box="[330,417,1610,1636]" captionStart="FIG" captionStartId="3.[112,123,1411,1428]" captionTargetBox="[127,1195,217,1368]" captionTargetId="figure-93@3.[89,1232,156,1420]" captionTargetPageId="3" captionText="FIG. 1. — A-C, Nepalella marmorata n. sp., paratype ♂; D-F, N. grandis n. sp., paratype ♂; G, N. grandis n. sp., paratype ♀; A, D, anterior body end, dorsal view; B, E, midbody segments, dorsal view; C, F, same, lateral view; G, vulva, front view. Scale bars: A-F, 2.0 mm; G, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.5393504" httpUri="https://zenodo.org/record/5393504/files/figure.png" pageId="6" pageNumber="81">Fig. 1A</figureCitation>
). Tegument smooth, shining, translucid, only prozona distinctly and densely striolate transversely. Metatergal setation 3 + 3, typical, macrochaetae medium-sized, rather thick but pointed, positioned on minute knobs; stricture between pro- and metazona very shallow, inconspicuous (
<figureCitation id="136C5FD8A9736A4BFCD2F9221CDAC7C1" box="[846,905,1673,1699]" captionStart="FIG" captionStartId="3.[112,123,1411,1428]" captionTargetBox="[127,1195,217,1368]" captionTargetId="figure-93@3.[89,1232,156,1420]" captionTargetPageId="3" captionText="FIG. 1. — A-C, Nepalella marmorata n. sp., paratype ♂; D-F, N. grandis n. sp., paratype ♂; G, N. grandis n. sp., paratype ♀; A, D, anterior body end, dorsal view; B, E, midbody segments, dorsal view; C, F, same, lateral view; G, vulva, front view. Scale bars: A-F, 2.0 mm; G, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.5393504" httpUri="https://zenodo.org/record/5393504/files/figure.png" pageId="6" pageNumber="81">Fig. 1</figureCitation>
D-F).
</paragraph>
<caption id="DF2813D5A9726A4AFFEDFA191EFEC492" ID-DOI="http://doi.org/10.5281/zenodo.5393512" ID-Zenodo-Dep="5393512" httpUri="https://zenodo.org/record/5393512/files/figure.png" pageId="7" pageNumber="82" startId="7.[113,124,1458,1475]" targetBox="[46,1277,131,1453]" targetPageId="7">
<paragraph id="8BE8435DA9726A4AFFEDFA191EFEC492" blockId="7.[113,1211,1458,1520]" pageId="7" pageNumber="82">
FIG. 4. —
<taxonomicName id="4C5738DEA9726A4AFF5CFA191E1CC4A1" authority="Golovatch &amp; Geoffroy &amp; Mauriès, 2006" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès" authorityYear="2006" box="[192,335,1458,1475]" class="Diplopoda" family="Lankasomatidae" genus="Nepalella" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="82" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
<emphasis id="B9239F4FA9726A4AFF5CFA191E1CC4A1" box="[192,335,1458,1475]" italics="true" pageId="7" pageNumber="82">Nepalella grandis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2102234A9726A4AFEC8FA191ED5C4A1" box="[340,390,1458,1475]" pageId="7" pageNumber="82" rank="species">n. sp.</taxonomicNameLabel>
, paratype ♂:
<emphasis id="B9239F4FA9726A4AFE6FFA191D53C4A1" bold="true" box="[499,512,1458,1475]" pageId="7" pageNumber="82">A</emphasis>
, antenna;
<emphasis id="B9239F4FA9726A4AFDCBFA191D37C4A1" bold="true" box="[599,612,1458,1475]" pageId="7" pageNumber="82">B</emphasis>
, leg 7, caudal view;
<emphasis id="B9239F4FA9726A4AFC91FA191C48C4A1" bold="true" box="[781,795,1458,1475]" pageId="7" pageNumber="82">C</emphasis>
, leg 10, front view;
<emphasis id="B9239F4FA9726A4AFC20FA191C99C4A1" bold="true" box="[956,970,1458,1475]" pageId="7" pageNumber="82">D</emphasis>
, leg 11, front view;
<emphasis id="B9239F4FA9726A4AFBF0FA191B2BC4A1" bold="true" box="[1132,1144,1458,1475]" pageId="7" pageNumber="82">E</emphasis>
, leg 12, front view;
<emphasis id="B9239F4FA9726A4AFF51FA631F8BC4BB" bold="true" box="[205,216,1480,1497]" pageId="7" pageNumber="82">F</emphasis>
, anterior gonopods (leg-pair 8), caudal view;
<emphasis id="B9239F4FA9726A4AFDCCFA631D0DC4BB" bold="true" box="[592,606,1480,1497]" pageId="7" pageNumber="82">G</emphasis>
,
<emphasis id="B9239F4FA9726A4AFDF4FA631D25C4BB" bold="true" box="[616,630,1480,1497]" pageId="7" pageNumber="82">H</emphasis>
, posterior gonopods (leg-pair 9), front and caudal views, respectively. Scale bar: A-E, 0.8 mm; F-H, 0.4 mm.
</paragraph>
</caption>
<paragraph id="8BE8435DA9726A4AFF14F9801E11C7C1" blockId="7.[113,647,1578,1700]" pageId="7" pageNumber="82">
Paraterga small dorsolateral swellings/shoulders, not set off by peritremata, rather regularly rounded in dorsal view, only more abruptly so after caudal corner (
<figureCitation id="136C5FD8A9726A4AFF59F9221E53C7C1" box="[197,256,1673,1699]" captionStart="FIG" captionStartId="3.[112,123,1411,1428]" captionTargetBox="[127,1195,217,1368]" captionTargetId="figure-93@3.[89,1232,156,1420]" captionTargetPageId="3" captionText="FIG. 1. — A-C, Nepalella marmorata n. sp., paratype ♂; D-F, N. grandis n. sp., paratype ♂; G, N. grandis n. sp., paratype ♀; A, D, anterior body end, dorsal view; B, E, midbody segments, dorsal view; C, F, same, lateral view; G, vulva, front view. Scale bars: A-F, 2.0 mm; G, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.5393504" httpUri="https://zenodo.org/record/5393504/files/figure.png" pageId="7" pageNumber="82">Fig. 1</figureCitation>
D-F).
</paragraph>
<paragraph id="8BE8435DA9726A45FD27F9811EFAC012" blockId="7.[677,1210,1578,1700]" lastBlockId="8.[113,647,216,1288]" lastPageId="8" lastPageNumber="83" pageId="7" pageNumber="82">CIX (15) = 0.46; MIX (15) = 2.08; MA (15) ≈ 165°; PIX impossible to evaluate as paraterga too poorly developed. Axial suture distinct, usual; segment 7 only very slightly broader than adjacent ones, conspicuous due to a clearly visible and frontally projecting anterior lobe of pleurite, latter without ventral apophyses. Limbus barely visible, microdentate, traceable only near pleurites ventrolaterally.</paragraph>
<paragraph id="8BE8435DA97D6A45FF14FEDC1D1DC24E" blockId="8.[113,647,216,1288]" pageId="8" pageNumber="83">
Legs very long and slender, slightly longer in males than in females,
<emphasis id="B9239F4FA97D6A45FEB6FE3C1E69C0D2" box="[298,314,407,432]" italics="true" pageId="8" pageNumber="83">c.</emphasis>
1.8 times as long as midbody height. Legs 1 and 2 slightly reduced as usual, tarsi with usual ventral brushes, but without papillae; male coxa 2 perforated by gonopore orifice. All tarsi slightly bulging immediately at base, male prefemora microgranulate ventrally (
<figureCitation id="136C5FD8A97D6A45FE16FD9F1E8DC32C" box="[394,478,564,590]" captionStart="FIG" captionStartId="7.[113,124,1458,1475]" captionTargetBox="[46,1277,131,1453]" captionTargetId="figure-88@7.[46,1277,131,1453]" captionTargetPageId="7" captionText="FIG. 4. — Nepalella grandis n. sp., paratype ♂: A, antenna; B, leg 7, caudal view; C, leg 10, front view; D, leg 11, front view; E, leg 12, front view; F, anterior gonopods (leg-pair 8), caudal view; G, H, posterior gonopods (leg-pair 9), front and caudal views, respectively. Scale bar: A-E, 0.8 mm; F-H, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.5393512" httpUri="https://zenodo.org/record/5393512/files/figure.png" pageId="8" pageNumber="83">Fig. 4E</figureCitation>
). Male legs 3-7 somewhat incrassate, pairs 3 and 4 especially so, each conspicuously papillate over caudal face of distoventral third (
<figureCitation id="136C5FD8A97D6A45FE94FD381E0FC3CF" box="[264,348,659,685]" captionStart="FIG" captionStartId="7.[113,124,1458,1475]" captionTargetBox="[46,1277,131,1453]" captionTargetId="figure-88@7.[46,1277,131,1453]" captionTargetPageId="7" captionText="FIG. 4. — Nepalella grandis n. sp., paratype ♂: A, antenna; B, leg 7, caudal view; C, leg 10, front view; D, leg 11, front view; E, leg 12, front view; F, anterior gonopods (leg-pair 8), caudal view; G, H, posterior gonopods (leg-pair 9), front and caudal views, respectively. Scale bar: A-E, 0.8 mm; F-H, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.5393512" httpUri="https://zenodo.org/record/5393512/files/figure.png" pageId="8" pageNumber="83">Fig. 4B</figureCitation>
). Male legs 10 and 11 with coxal glands; prefemur 10 conspicuously enlarged at base (
<figureCitation id="136C5FD8A97D6A45FF5FFD781E47C38F" box="[195,276,723,749]" captionStart="FIG" captionStartId="7.[113,124,1458,1475]" captionTargetBox="[46,1277,131,1453]" captionTargetId="figure-88@7.[46,1277,131,1453]" captionTargetPageId="7" captionText="FIG. 4. — Nepalella grandis n. sp., paratype ♂: A, antenna; B, leg 7, caudal view; C, leg 10, front view; D, leg 11, front view; E, leg 12, front view; F, anterior gonopods (leg-pair 8), caudal view; G, H, posterior gonopods (leg-pair 9), front and caudal views, respectively. Scale bar: A-E, 0.8 mm; F-H, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.5393512" httpUri="https://zenodo.org/record/5393512/files/figure.png" pageId="8" pageNumber="83">Fig. 4C</figureCitation>
); prefemur 11 with a strong, microgranulate, parabasal process ventrally (
<figureCitation id="136C5FD8A97D6A45FD81FD591D29C26F" box="[541,634,754,781]" captionStart="FIG" captionStartId="7.[113,124,1458,1475]" captionTargetBox="[46,1277,131,1453]" captionTargetId="figure-88@7.[46,1277,131,1453]" captionTargetPageId="7" captionText="FIG. 4. — Nepalella grandis n. sp., paratype ♂: A, antenna; B, leg 7, caudal view; C, leg 10, front view; D, leg 11, front view; E, leg 12, front view; F, anterior gonopods (leg-pair 8), caudal view; G, H, posterior gonopods (leg-pair 9), front and caudal views, respectively. Scale bar: A-E, 0.8 mm; F-H, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.5393512" httpUri="https://zenodo.org/record/5393512/files/figure.png" pageId="8" pageNumber="83">Fig. 4D</figureCitation>
). Claws invariably long, sabre-shaped, simple.
</paragraph>
<paragraph id="8BE8435DA97D6A45FF14FC991D33C2E9" blockId="8.[113,647,216,1288]" pageId="8" pageNumber="83">
Anterior gonopods (leg-pair 8) as usual very strongly reduced, sternum with a median protuberance and two lateral lobules, coxites spike-like (
<figureCitation id="136C5FD8A97D6A45FD9FFCDA1D00C2E9" box="[515,595,881,907]" captionStart="FIG" captionStartId="7.[113,124,1458,1475]" captionTargetBox="[46,1277,131,1453]" captionTargetId="figure-88@7.[46,1277,131,1453]" captionTargetPageId="7" captionText="FIG. 4. — Nepalella grandis n. sp., paratype ♂: A, antenna; B, leg 7, caudal view; C, leg 10, front view; D, leg 11, front view; E, leg 12, front view; F, anterior gonopods (leg-pair 8), caudal view; G, H, posterior gonopods (leg-pair 9), front and caudal views, respectively. Scale bar: A-E, 0.8 mm; F-H, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.5393512" httpUri="https://zenodo.org/record/5393512/files/figure.png" pageId="8" pageNumber="83">Fig. 4F</figureCitation>
).
</paragraph>
<paragraph id="8BE8435DA97D6A45FF14FC3A1B6DC073" blockId="8.[113,647,216,1288]" lastBlockId="8.[677,1210,216,368]" pageId="8" pageNumber="83">
Posterior gonopods (leg-pair 9) hypertophied (
<figureCitation id="136C5FD8A97D6A45FFE5FC1B1FA2C2A9" box="[121,241,944,971]" captionStart="FIG" captionStartId="7.[113,124,1458,1475]" captionTargetBox="[46,1277,131,1453]" captionTargetId="figure-88@7.[46,1277,131,1453]" captionTargetPageId="7" captionText="FIG. 4. — Nepalella grandis n. sp., paratype ♂: A, antenna; B, leg 7, caudal view; C, leg 10, front view; D, leg 11, front view; E, leg 12, front view; F, anterior gonopods (leg-pair 8), caudal view; G, H, posterior gonopods (leg-pair 9), front and caudal views, respectively. Scale bar: A-E, 0.8 mm; F-H, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.5393512" httpUri="https://zenodo.org/record/5393512/files/figure.png" pageId="8" pageNumber="83">Fig. 4G, H</figureCitation>
), with two ear-shaped folds at base on anterior face of colpocoxites and two large flagella beset with simple pilosity. Bases of coxite with two small, roundish bulges beset with minute plumose setae on caudal face. Colpocoxites especially prominent, higher than telopodites, distal one-third gently curved and directed distocaudally, delimited at base by a transverse sulcus on front face, with a relatively small, recurved, spoon-shaped process at base of caudal face, and an inner, gently ribbed lobe with a minute solenomere at tip.Telopoditomere 1 more strongly setose on front face, telopoditomere 2 with two vestigial segments apically.
</paragraph>
<paragraph id="8BE8435DA97D6A45FD27FEBC1C99C012" blockId="8.[677,1210,216,368]" pageId="8" pageNumber="83">
Vulva (
<figureCitation id="136C5FD8A97D6A45FC9BFEBC1C08C053" box="[775,859,279,305]" captionStart="FIG" captionStartId="3.[112,123,1411,1428]" captionTargetBox="[127,1195,217,1368]" captionTargetId="figure-93@3.[89,1232,156,1420]" captionTargetPageId="3" captionText="FIG. 1. — A-C, Nepalella marmorata n. sp., paratype ♂; D-F, N. grandis n. sp., paratype ♂; G, N. grandis n. sp., paratype ♀; A, D, anterior body end, dorsal view; B, E, midbody segments, dorsal view; C, F, same, lateral view; G, vulva, front view. Scale bars: A-F, 2.0 mm; G, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.5393504" httpUri="https://zenodo.org/record/5393504/files/figure.png" pageId="8" pageNumber="83">Fig. 1G</figureCitation>
) rather short, densely pilose, receptacle process on operculum small but evident, curved, digitiform, slender.
</paragraph>
</subSubSection>
<subSubSection id="C34D10D6A97D6A45FD39FE3D1B6DC465" pageId="8" pageNumber="83" type="discussion">
<paragraph id="8BE8435DA97D6A45FD39FE3D1C5AC0CC" blockId="8.[677,1210,406,1288]" box="[677,777,406,432]" pageId="8" pageNumber="83">REMARKS</paragraph>
<paragraph id="8BE8435DA97D6A45FD39FE1E1C17C24E" blockId="8.[677,1210,406,1288]" pageId="8" pageNumber="83">
<taxonomicName id="4C5738DEA97D6A45FD39FE1E1C04C0AD" authority="Golovatch &amp; Geoffroy &amp; Mauriès, 2006" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès" authorityYear="2006" box="[677,855,437,463]" class="Diplopoda" family="Lankasomatidae" genus="Nepalella" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="83" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
<emphasis id="B9239F4FA97D6A45FD39FE1E1C04C0AD" box="[677,855,437,463]" italics="true" pageId="8" pageNumber="83">Nepalella grandis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2102234A97D6A45FCC2FE1D1CCAC0B2" box="[862,921,438,464]" pageId="8" pageNumber="83" rank="species">n. sp.</taxonomicNameLabel>
is amongst the largest species in the order
<taxonomicName id="4C5738DEA97D6A45FCFAFE7E1B4AC08D" authorityName="Pocock" authorityYear="1894" box="[870,1049,469,495]" class="Diplopoda" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="83" phylum="Arthropoda" rank="order">Chordeumatida</taxonomicName>
, and certainly the largest in
<taxonomicName id="4C5738DEA97D6A45FCAFFE5E1CC7C36D" authorityName="S HEAR" authorityYear="1979" box="[819,916,501,527]" class="Diplopoda" family="Lankasomatidae" genus="Nepalella" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis id="B9239F4FA97D6A45FCAFFE5E1CC7C36D" box="[819,916,501,527]" italics="true" pageId="8" pageNumber="83">Nepalella</emphasis>
</taxonomicName>
. This can also be associated with cave gigantism, as this animal is definitely troglobitic, showing such typical troglomorphisms as unpigmented ocelli and tegument, extremely long legs and antennae. The number of ocelli per eye patch is also reduced, especially in the female. However, because very long and slender antennae are characteristic of many, often definitely epigeic congeners, their value as a troglomorphic trait is highly limited.
</paragraph>
<paragraph id="8BE8435DA97D6A45FD27FC991B6DC465" blockId="8.[677,1210,406,1288]" pageId="8" pageNumber="83">
This genus is well defined (
<bibRefCitation id="EFC63EACA97D6A45FC40FC991B0AC22E" author="SHEAR W. A." box="[988,1113,818,844]" pageId="8" pageNumber="83" pagination="63 - 72" refId="ref8455" refString="SHEAR W. A. 2002. - Five new chordeumatidan millipeds from China: new species of Vieteuma (Kashmireumatidae) and Nepalella (Megalotylidae). Proceedings of the California Academy of Sciences 53 (6): 63 - 72." type="journal article" year="2002">Shear 2002</bibRefCitation>
) and has hitherto been known to comprise 20 species or subspecies from
<collectingCountry id="F34003CDA97D6A45FCD0FCDA1CD8C2E9" box="[844,907,881,907]" name="Nepal" pageId="8" pageNumber="83">Nepal</collectingCountry>
(10),
<collectingCountry id="F34003CDA97D6A45FC5BFCDA1B77C2E9" box="[967,1060,881,907]" name="Thailand" pageId="8" pageNumber="83">Thailand</collectingCountry>
(2),
<collectingCountry id="F34003CDA97D6A45FBCFFCD91BE9C2EE" box="[1107,1210,882,908]" name="Myanmar" pageId="8" pageNumber="83">Myanmar</collectingCountry>
(2),
<collectingCountry id="F34003CDA97D6A45FD4CFC3A1C7EC2C9" box="[720,813,913,939]" name="Vietnam" pageId="8" pageNumber="83">Vietnam</collectingCountry>
(1) and southern
<collectingCountry id="F34003CDA97D6A45FC74FC3A1B78C2C9" box="[1000,1067,913,939]" name="China" pageId="8" pageNumber="83">China</collectingCountry>
(5). The only previous key dealt with 14 of these (
<bibRefCitation id="EFC63EACA97D6A45FB8AFC1B1BFEC2A9" author="MAURIES J. - P." box="[1046,1197,944,971]" pageId="8" pageNumber="83" pagination="3 - 49" refId="ref8278" refString="MAURIES J. - P. 1988. - Myriapodes du Nepal. II. Diplopodes craspedosomides nouveaux de l'Himalaya et de la region indo-malaise (Craspedosomidea et Chordeumidea). Revue suisse de Zoologie 96 (1): 3 - 49." type="journal article" year="1988">Mauriès 1988</bibRefCitation>
). With the addition of a further two congeners from Chinese caves,
<taxonomicName id="4C5738DEA97D6A45FCD0FC5B1CFDC568" authorityName="S HEAR" authorityYear="1979" box="[844,942,1008,1034]" class="Diplopoda" family="Lankasomatidae" genus="Nepalella" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis id="B9239F4FA97D6A45FCD0FC5B1CFDC568" box="[844,942,1008,1034]" italics="true" pageId="8" pageNumber="83">Nepalella</emphasis>
</taxonomicName>
can now be regarded as one of the most speciose genera among the oriental
<taxonomicName id="4C5738DEA97D6A45FD39FB841C00C52B" authorityName="Pocock" authorityYear="1894" box="[677,851,1071,1097]" class="Diplopoda" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="83" phylum="Arthropoda" rank="order">Chordeumatida</taxonomicName>
. Considering all available descriptions (
<bibRefCitation id="EFC63EACA97D6A45FD75FBE41C3AC50B" author="SHEAR W. A." box="[745,873,1103,1129]" pageId="8" pageNumber="83" pagination="115 - 130" refId="ref8357" refString="SHEAR W. A. 1979. - Diplopoda from the Nepal Himalayas. Chordeumida with comments on the Asian chordeumid fauna. Senckenbergiana biologica 60 (1 - 2): 115 - 130." type="journal article" year="1979">Shear 1979</bibRefCitation>
,
<bibRefCitation id="EFC63EACA97D6A45FCE9FBE41CE3C50B" author="SHEAR W. A." box="[885,944,1103,1129]" pageId="8" pageNumber="83" pagination="229 - 240" refId="ref8393" refString="SHEAR W. A. 1987. - Chordeumatid Diplopoda from the Nepal Himalayas, II. Courier Forschungsinstitut Senckenberg 93: 229 - 240." type="journal article" year="1987">1987</bibRefCitation>
,
<bibRefCitation id="EFC63EACA97D6A45FC20FBE41CA4C50B" author="SHEAR W. A." box="[956,1015,1103,1129]" pageId="8" pageNumber="83" pagination="1 - 10" refId="ref8419" refString="SHEAR W. A. 1999. - A new troglobitic milliped of the genus Nepalella from China (Diplopoda, Chordeumatida, Megalotylidae). Myriapodologica 6 (1): 1 - 10." type="journal article" year="1999">1999</bibRefCitation>
,
<bibRefCitation id="EFC63EACA97D6A45FB9FFBE41B6DC50B" author="SHEAR W. A." box="[1027,1086,1103,1129]" pageId="8" pageNumber="83" pagination="63 - 72" refId="ref8455" refString="SHEAR W. A. 2002. - Five new chordeumatidan millipeds from China: new species of Vieteuma (Kashmireumatidae) and Nepalella (Megalotylidae). Proceedings of the California Academy of Sciences 53 (6): 63 - 72." type="journal article" year="2002">2002</bibRefCitation>
;
<bibRefCitation id="EFC63EACA97D6A45FBD5FBE41DB3C5EB" author="GOLOVATCH S. I." pageId="8" pageNumber="83" pagination="123 - 127" refId="ref7969" refString="GOLOVATCH S. I. 1983. - Contributions to the millipede fauna of Vietnam (Diplopoda) I. Chordeumatida. Acta Zoologica Academiae Scientiarum Hungaricae 29 (1 - 3): 123 - 127." type="journal article" year="1983">Golovatch 1983</bibRefCitation>
;
<bibRefCitation id="EFC63EACA97D6A45FD71FBC51CD8C5EB" author="MAURIES J. - P." box="[749,907,1134,1161]" pageId="8" pageNumber="83" pagination="3 - 49" refId="ref8278" refString="MAURIES J. - P. 1988. - Myriapodes du Nepal. II. Diplopodes craspedosomides nouveaux de l'Himalaya et de la region indo-malaise (Craspedosomidea et Chordeumidea). Revue suisse de Zoologie 96 (1): 3 - 49." type="journal article" year="1988">Mauriès 1988</bibRefCitation>
), the following refined and updated key to
<taxonomicName id="4C5738DEA97D6A45FCD0FB251CAEC5CA" box="[844,1021,1166,1192]" class="Diplopoda" family="Lankasomatidae" genus="Nepalella" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="83" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9239F4FA97D6A45FCD0FB251CFEC5CA" box="[844,941,1166,1192]" italics="true" pageId="8" pageNumber="83">Nepalella</emphasis>
species
</taxonomicName>
is proposed. This will hopefully encourage further work on this remarkable genus, in which many other new species undoubtedly remain to be discovered.
</paragraph>
</subSubSection>
</treatment>
</document>
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