202 lines
20 KiB
XML
202 lines
20 KiB
XML
<document id="189E6389C706A0C87EC7508C821C7406" ID-CLB-Dataset="302762" ID-DOI="10.17161/jom.v0i25.4659" ID-GBIF-Dataset="a954f134-084d-41e5-8a41-61af66929a3b" ID-ISSN="2325-4467" ID-Zenodo-Dep="13736871" ID-ZooBank="urn:lsid:zoobank.org:pub:9AFCD1E3-42B6-4CB3-8321-35C60579289C" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="guilherme" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1722892427188" checkinUser="felipe" docAuthor="Engel, Michael S., Ortega-Blanco, Jaime, Nascimbene, Paul C. & Singh, Hukam" docDate="2013" docId="03E487BF7579FFC3FE97F99DFDED21A3" docLanguage="en" docName="JMelittology.2013.25.1-12.pdf" docOrigin="Journal of Melittology 2013 (25)" docSource="https://doi.org/10.17161/jom.v0i25.4659" docStyle="DocumentStyle:20C606B715423E98DFED429F4976A416.4:JMelittology.2013-.journal_article.0cover" docStyleId="20C606B715423E98DFED429F4976A416" docStyleName="JMelittology.2013-.journal_article.0cover" docStyleVersion="4" docTitle="Melikertes (Paramelikertes) Engel, Ortega-Blanco, Nascimbene & Singh, 2013, new subgenus" docType="treatment" docVersion="4" lastPageNumber="5" masterDocId="FFDDFFC7757AFFC7FFCFFFBDFF8D2740" masterDocTitle="The bees of Early Eocene Cambay amber (Hymenoptera: Apidae)" masterLastPageNumber="12" masterPageNumber="1" pageNumber="4" updateTime="1725892524702" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-4.0">
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<mods:titleInfo id="EB97DB654DECE136E03FEB927E9C360B">
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<mods:title id="21710A33A4AAC1905228E05A9B657B5E">The bees of Early Eocene Cambay amber (Hymenoptera: Apidae)</mods:title>
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<mods:namePart id="879DADBEA65C3D277A9441914C5EB446">Engel, Michael S.</mods:namePart>
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<mods:affiliation id="3D70D0068B7E3827374ED0530BBA1DAF">Division of Entomology, Natural History Museum, and Department of Ecology & Evolutionary Biology, 1501 Crestline Drive - Suite 140, University of Kansas, Lawrence, Kansas 66045, USA (msengel @ ku. edu; jaimeortega @ ku. edu). & Division of Invertebrate Zoology, American Museum of Natural History, Central Park West</mods:affiliation>
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<mods:namePart id="E53DEAA814EADB7F0D9F0F95C9B0790A">Ortega-Blanco, Jaime</mods:namePart>
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<mods:affiliation id="38526B6D12BAAEBA9F63BB55B262C04B">Division of Entomology, Natural History Museum, and Department of Ecology & Evolutionary Biology, 1501 Crestline Drive - Suite 140, University of Kansas, Lawrence, Kansas 66045, USA (msengel @ ku. edu; jaimeortega @ ku. edu).</mods:affiliation>
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<mods:namePart id="FE6821E713538022EF6A5958C936E48A">Nascimbene, Paul C.</mods:namePart>
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<mods:affiliation id="3F3F1A6FB170ED8EBEECF4201332E363">Division of Invertebrate Zoology, American Museum of Natural History, Central Park West</mods:affiliation>
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<mods:namePart id="123C4889C19DE9378D4FB645A4D0BE74">Singh, Hukam</mods:namePart>
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<mods:affiliation id="F4180FF0C111C6EB8F4E1102B9BE385B">Birbal Sahni Institute of Palaeobotany, 53 University Road, Lucknow 226 007, India (hukams @ gmail. com).</mods:affiliation>
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<mods:title id="B64662A951A2F37D26C540CEF47BCB36">Journal of Melittology</mods:title>
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<mods:date id="2FCDEAE9156339BA4FF0696480AF7F25">2013</mods:date>
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<mods:number id="4052823A8195AC95A83EE0E09B1F19FE">2013-12-17</mods:number>
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<treatment id="03E487BF7579FFC3FE97F99DFDED21A3" ID-DOI="http://doi.org/10.5281/zenodo.13736879" ID-Zenodo-Dep="13736879" LSID="urn:lsid:plazi:treatment:03E487BF7579FFC3FE97F99DFDED21A3" httpUri="http://treatment.plazi.org/id/03E487BF7579FFC3FE97F99DFDED21A3" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
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<subSubSection id="C35765227579FFC4FE97F99DFBC82118" pageId="3" pageNumber="4" type="nomenclature">
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<paragraph id="8BF236A97579FFC4FE97F99DFC552178" blockId="3.[235,1093,1536,1624]" box="[344,984,1568,1592]" pageId="3" pageNumber="4">
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<heading id="D0BA81C57579FFC4FE97F99DFC552178" box="[344,984,1568,1592]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="4" reason="2">
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<taxonomicName id="4C4D4D2A7579FFC4FE97F99DFD812178" authorityName="Engel & Ortega-Blanco & Nascimbene & Singh" authorityYear="2013" box="[344,524,1568,1592]" class="Insecta" family="Apidae" genus="Melikertes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="subGenus" status="subgen. nov." subGenus="Paramelikertes">
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<emphasis id="B939EABB7579FFC4FE97F99DFD812178" bold="true" box="[344,524,1568,1592]" italics="true" pageId="3" pageNumber="4">Paramelikertes</emphasis>
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</taxonomicName>
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Engel & Ortega-Blanco,
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<taxonomicNameLabel id="A20A57C07579FFC4FCE1F99CFC552178" box="[814,984,1568,1592]" pageId="3" pageNumber="4" rank="subGenus">new subgenus</taxonomicNameLabel>
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</heading>
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</paragraph>
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ZooBank:
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</uri>
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</subSubSection>
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<subSubSection id="C35765227579FFC4FF1FF93CFF7621F8" pageId="3" pageNumber="4" type="materials_examined">
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<paragraph id="8BF236A97579FFC4FF1FF93CFF7621F8" blockId="3.[160,1168,1664,1784]" pageId="3" pageNumber="4">
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<materialsCitation id="3B253CF47579FFC4FF1FF93CFF7A21F8" collectionCode="SPECIES" pageId="3" pageNumber="4" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54F6880B7579FFC4FF1FF93CFE8821D8" box="[208,261,1665,1688]" pageId="3" pageNumber="4">TYPE</typeStatus>
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<collectionCode id="ED5CAE6C7579FFC4FEDEF93DFEE821D8" box="[273,357,1664,1688]" pageId="3" pageNumber="4">SPECIES</collectionCode>
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:
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<taxonomicName id="4C4D4D2A7579FFC4FEBCF93DFBC021D8" authority="Engel & Ortega-Blanco" authorityName="Engel & Ortega-Blanco & Nascimbene & Singh" authorityYear="2013" box="[371,1101,1664,1688]" class="Insecta" family="Apidae" genus="Melikertes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="gujaratensis" status="sp. nov." subGenus="Paramelikertes">
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<emphasis id="B939EABB7579FFC4FEBCF93DFE6C21D8" box="[371,481,1664,1688]" italics="true" pageId="3" pageNumber="4">Melikertes</emphasis>
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(
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<emphasis id="B939EABB7579FFC4FE39F93DFD1D21D8" box="[502,656,1664,1688]" italics="true" pageId="3" pageNumber="4">Paramelikertes</emphasis>
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)
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<emphasis id="B939EABB7579FFC4FD6BF93CFCA821D8" box="[676,805,1665,1688]" italics="true" pageId="3" pageNumber="4">gujaratensis</emphasis>
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Engel & Ortega-Blanco
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</taxonomicName>
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,
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<taxonomicNameLabel id="A20A57C07579FFC4FB92F93CFF7A21F8" pageId="3" pageNumber="4" rank="species">new species</taxonomicNameLabel>
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||
</materialsCitation>
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.
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C35765227579FFC3FF1FF97DFD0E2164" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="diagnosis">
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<paragraph id="8BF236A97579FFC3FF1FF97DFD0E2164" blockId="3.[160,1168,1664,1784]" lastBlockId="4.[160,1168,1388,1764]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
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DIAGNOSIS: This new group can be differentiated from
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<taxonomicName id="4C4D4D2A7579FFC4FCB0F97DFC7D2198" authorityName="Engel" authorityYear="1998" box="[895,1008,1728,1752]" class="Insecta" family="Apidae" genus="Melikertes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus" sensu="stricto">
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<emphasis id="B939EABB7579FFC4FCB0F97DFC7D2198" box="[895,1008,1728,1752]" italics="true" pageId="3" pageNumber="4">Melikertes</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20A57C07579FFC4FC33F97CFBBC2198" box="[1020,1073,1729,1752]" pageId="3" pageNumber="4" sensu="stricto">
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<emphasis id="B939EABB7579FFC4FC33F97CFBA62198" box="[1020,1067,1729,1752]" italics="true" pageId="3" pageNumber="4">s.str</emphasis>
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.
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</taxonomicNameLabel>
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(
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<bibRefCitation id="EFDC4B587579FFC4FB8AF97DFF6921B8" author="Engel, M. S." pageId="3" pageNumber="4" pagination="1 - 192" refId="ref3854" refString="Engel, M. S. 2001 a. A monograph of the Baltic amber bees and evolution of the Apoidea (Hymenoptera). Bulletin of the American Museum of Natural History 259: 1 - 192." type="journal article" year="2001">Engel, 2001a</bibRefCitation>
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) by the presence of only two submarginal cells (
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<figureCitation id="13762A2C7579FFC4FCF5F95DFC0621B8" box="[826,907,1760,1784]" captionStart="Figures 1–2" captionStartId="3.[160,241,1254,1276]" captionTargetBox="[160,1168,224,1249]" captionTargetId="figure-123@3.[160,1168,224,1249]" captionTargetPageId="3" captionText="Figures 1–2. Photographs of holotype worker (AMNH Tad-41-A) of Melikertes (Paramelikertes) gujaratensis, new subgenus and species. 1. Entire specimen as preserved in dorsal oblique view. 2. Detail of forewing." figureDoi="http://doi.org/10.5281/zenodo.13736875" httpUri="https://zenodo.org/record/13736875/files/figure.png" pageId="3" pageNumber="4">Figs. 2</figureCitation>
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,
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<figureCitation id="13762A2C7579FFC4FC55F95DFC2521B8" box="[922,936,1760,1784]" captionStart="Figure 3" captionStartId="4.[160,231,1267,1289]" captionTargetBox="[194,1164,242,1243]" captionTargetId="figure-161@4.[160,1168,225,1262]" captionTargetPageId="4" captionText="Figure 3. Holotype worker (AMNH Tad-41-A) of Melikertes (Paramelikertes) gujaratensis, new subgenus and species." figureDoi="http://doi.org/10.5281/zenodo.13736877" httpUri="https://zenodo.org/record/13736877/files/figure.png" pageId="3" pageNumber="4">3</figureCitation>
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), the confluence of 2rs-m with 2m-cu, the more elongate and narrow marginal cell, the marginal cell apex acutely pointed on the wing margin, the more densely setose metasoma, and the more strongly developed and dense stout bristles of the mid- and hind legs. Like
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<taxonomicName id="4C4D4D2A757EFFC3FBD0FA11FB1D2284" authorityName="Engel" authorityYear="1998" box="[1055,1168,1452,1476]" class="Insecta" family="Apidae" genus="Melikertes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
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<emphasis id="B939EABB757EFFC3FBD0FA11FB1D2284" box="[1055,1168,1452,1476]" italics="true" pageId="4" pageNumber="5">Melikertes</emphasis>
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</taxonomicName>
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the malar space is apparently short, the clypeus lacks the clypeal protrusion typical of
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<taxonomicName id="4C4D4D2A757EFFC3FF6FFA51FEEC2144" authority="Engel" authorityName="Engel" authorityYear="2001" box="[160,353,1516,1540]" class="Insecta" family="Apidae" genus="Succinapis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
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<emphasis id="B939EABB757EFFC3FF6FFA51FE982143" box="[160,277,1516,1539]" italics="true" pageId="4" pageNumber="5">Succinapis</emphasis>
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Engel
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</taxonomicName>
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, the mesoscutellum apparently does not project over the metanotum, and the metasomal terga are not banded.
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</paragraph>
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</subSubSection>
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||
<caption id="DF326621757EFFC3FF6FFB4EFE012266" ID-DOI="http://doi.org/10.5281/zenodo.13736877" ID-Zenodo-Dep="13736877" httpUri="https://zenodo.org/record/13736877/files/figure.png" pageId="4" pageNumber="5" startId="4.[160,231,1267,1289]" targetBox="[194,1164,242,1243]" targetPageId="4" targetType="figure">
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<paragraph id="8BF236A9757EFFC3FF6FFB4EFE012266" blockId="4.[160,1136,1267,1318]" pageId="4" pageNumber="5">
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||
<emphasis id="B939EABB757EFFC3FF6FFB4EFF722248" bold="true" box="[160,255,1267,1289]" pageId="4" pageNumber="5">Figure 3.</emphasis>
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Holotype worker (AMNH Tad-41-A) of
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<taxonomicName id="4C4D4D2A757EFFC3FD7EFB4EFBBA2249" authorityName="Engel & Ortega-Blanco & Nascimbene & Singh" authorityYear="2013" box="[689,1079,1267,1289]" class="Insecta" family="Apidae" genus="Melikertes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="gujaratensis" status="new subgenus and species" subGenus="Paramelikertes">
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||
<emphasis id="B939EABB757EFFC3FD7EFB4EFC9B2248" box="[689,790,1267,1288]" italics="true" pageId="4" pageNumber="5">Melikertes</emphasis>
|
||
(
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||
<emphasis id="B939EABB757EFFC3FCEBFB4EFC3F2248" box="[804,946,1267,1288]" italics="true" pageId="4" pageNumber="5">Paramelikertes</emphasis>
|
||
)
|
||
<emphasis id="B939EABB757EFFC3FC0FFB49FBBA2249" box="[960,1079,1268,1289]" italics="true" pageId="4" pageNumber="5">gujaratensis</emphasis>
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</taxonomicName>
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||
,
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<taxonomicNameLabel id="A20A57C0757EFFC3FB8CFB49FE052266" pageId="4" pageNumber="5" rank="species">new subgenus and species</taxonomicNameLabel>
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||
.
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</paragraph>
|
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</caption>
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<subSubSection id="C3576522757EFFC3FF1FF991FBD32124" pageId="4" pageNumber="5" type="etymology">
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<paragraph id="8BF236A9757EFFC3FF1FF991FBD32124" blockId="4.[160,1168,1388,1764]" pageId="4" pageNumber="5">
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ETYMOLOGY: The new subgeneric name is a combination of the Greek prefix
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<emphasis id="B939EABB757EFFC3FBAEF991FB1D2103" box="[1121,1168,1580,1603]" italics="true" pageId="4" pageNumber="5">para</emphasis>
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(meaning, “near”) and
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<taxonomicName id="4C4D4D2A757EFFC3FE7FF9F1FDAC2124" authorityName="Engel" authorityYear="1998" box="[432,545,1612,1636]" class="Insecta" family="Apidae" genus="Melikertes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
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<emphasis id="B939EABB757EFFC3FE7FF9F1FDAC2124" box="[432,545,1612,1636]" italics="true" pageId="4" pageNumber="5">Melikertes</emphasis>
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</taxonomicName>
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,
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<typeStatus id="54F6880B757EFFC3FDE1F9F1FDEF2124" box="[558,610,1612,1636]" pageId="4" pageNumber="5">type</typeStatus>
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genus of the tribe. The name is masculine.
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</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3576522757EFFC3FF1FF9D1FDED21A3" pageId="4" pageNumber="5" type="discussion">
|
||
<paragraph id="8BF236A9757EFFC3FF1FF9D1FDED21A3" blockId="4.[160,1168,1388,1764]" pageId="4" pageNumber="5">
|
||
COMMENTS: The genus
|
||
<taxonomicName id="4C4D4D2A757EFFC3FE10F9D1FD1721C4" authority="Engel" authorityName="Engel" authorityYear="1998" box="[479,666,1644,1668]" class="Insecta" family="Apidae" genus="Melikertes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B939EABB757EFFC3FE10F9D1FDDD21C4" box="[479,592,1644,1668]" italics="true" pageId="4" pageNumber="5">Melikertes</emphasis>
|
||
Engel
|
||
</taxonomicName>
|
||
was previously known from three species in Baltic amber –
|
||
<taxonomicName id="4C4D4D2A757EFFC3FEBAF931FD3F21E4" authority="(Menge)" baseAuthorityName="Menge" baseAuthorityYear="1856" box="[373,690,1676,1700]" class="Insecta" family="Apidae" genus="Melikertes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="proavus">
|
||
<emphasis id="B939EABB757EFFC3FEBAF931FDC921E3" box="[373,580,1676,1700]" italics="true" pageId="4" pageNumber="5">Melikertes proavus</emphasis>
|
||
(Menge)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4D4D2A757EFFC3FD70F931FC3221E4" authority="(Engel)" baseAuthorityName="Engel" baseAuthorityYear="1998" box="[703,959,1676,1700]" class="Insecta" family="Apidae" genus="Melikertes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="stilbonotus">
|
||
<emphasis id="B939EABB757EFFC3FD70F931FD5521E3" box="[703,728,1676,1699]" italics="true" pageId="4" pageNumber="5">M</emphasis>
|
||
.
|
||
<emphasis id="B939EABB757EFFC3FD27F931FCD221E4" box="[744,863,1676,1700]" italics="true" pageId="4" pageNumber="5">stilbonotus</emphasis>
|
||
(Engel)
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C4D4D2A757EFFC3FBCDF931FF6E2184" authority="Engel" authorityName="Engel" class="Insecta" family="Apidae" genus="Melikertes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="clypeatus">
|
||
<emphasis id="B939EABB757EFFC3FBCDF931FB9621E3" box="[1026,1051,1676,1699]" italics="true" pageId="4" pageNumber="5">M</emphasis>
|
||
.
|
||
<emphasis id="B939EABB757EFFC3FBE4F931FB1D21E4" box="[1067,1168,1676,1700]" italics="true" pageId="4" pageNumber="5">clypeatus</emphasis>
|
||
Engel
|
||
</taxonomicName>
|
||
(
|
||
<tableCitation id="C6CF0312757EFFC3FF3CF911FEC52184" box="[243,328,1708,1732]" captionStart="Table 1" captionStartId="1.[160,221,225,246]" captionText="Table 1. Currently described Eocene amber Apinae (all are preserved in Baltic amber except the two new species from Cambay amber reported herein and Exebotrigona velteni Engel & Michener in Fushan amber)." pageId="4" pageNumber="5">Table 1</tableCitation>
|
||
). Two new species are added here, one in the new subgenus, and the remainder classified in
|
||
<taxonomicName id="4C4D4D2A757EFFC3FE7CF971FDA921A4" authorityName="Engel" authorityYear="1998" box="[435,548,1740,1764]" class="Insecta" family="Apidae" genus="Melikertes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus" sensu="stricto">
|
||
<emphasis id="B939EABB757EFFC3FE7CF971FDA921A4" box="[435,548,1740,1764]" italics="true" pageId="4" pageNumber="5">Melikertes</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20A57C0757EFFC3FDE5F971FDED21A3" box="[554,608,1740,1763]" pageId="4" pageNumber="5" sensu="stricto">
|
||
<emphasis id="B939EABB757EFFC3FDE5F971FDD421A3" box="[554,601,1740,1763]" italics="true" pageId="4" pageNumber="5">s.str</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |