262 lines
42 KiB
XML
262 lines
42 KiB
XML
<document id="DC14802BB04282C37752857628ED11F3" ID-DOI="10.11646/zootaxa.5297.3.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="8004996" ID-ZooBank="ABF87F77-6C54-4460-919E-78D67CFAB176" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1685959583509" checkinUser="plazi" docAuthor="Mikhaljova, Elena V." docDate="2023" docId="03E48797B827F71064CF1D3664C559C4" docLanguage="en" docName="zootaxa.5297.3.3.pdf" docOrigin="Zootaxa 5297 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5297.3.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nepalmatoiulus simultaneus Mikhaljova 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="387" masterDocId="FFDDFFEFB822F71764581F216535584D" masterDocTitle="Descriptions of the new species of the genus Nepalmatoiulus Mauriès, 1983 from China, with notes on the closely coexisting species (Diplopoda, Julida, Julidae)" masterLastPageNumber="392" masterPageNumber="380" pageNumber="385" updateTime="1685960003826" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="1207AB13104DF8A12045B6BE957DA517">Descriptions of the new species of the genus Nepalmatoiulus Mauriès, 1983 from China, with notes on the closely coexisting species (Diplopoda, Julida, Julidae)</mods:title>
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<mods:namePart id="4A44851E6ED1F858AD0C58DD4ABFA119">Mikhaljova, Elena V.</mods:namePart>
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<mods:title id="6B23C1D346E1C3FF7EA9C07FB146FE94">Zootaxa</mods:title>
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<mods:part id="C77F1F4F8F8BA6A106EF6BBE825A223A">
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<mods:date id="1A0241A99727D6E41C143302DC6A17EE">2023</mods:date>
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<mods:number id="1A0AF0A687125EEBC22C71B3A5821384">2023-06-02</mods:number>
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<mods:number id="7C93F527AFB4747AB42EC589948477E8">5297</mods:number>
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<mods:url id="9A4D82E9FC6C4410060E5A695CEF0106">http://dx.doi.org/10.11646/zootaxa.5297.3.3</mods:url>
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<mods:classification id="264F7D64F78E84301641696603224371">journal article</mods:classification>
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<mods:identifier id="D3318C2CA066497940B30D1568A5E43E" type="DOI">10.11646/zootaxa.5297.3.3</mods:identifier>
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<mods:identifier id="A26450422712876EB60F94CCFE676B3A" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03E48797B827F71064CF1D3664C559C4" ID-DOI="http://doi.org/10.5281/zenodo.8005004" ID-Zenodo-Dep="8005004" LSID="urn:lsid:plazi:treatment:03E48797B827F71064CF1D3664C559C4" httpUri="http://treatment.plazi.org/id/03E48797B827F71064CF1D3664C559C4" lastPageId="7" lastPageNumber="387" pageId="5" pageNumber="385">
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<subSubSection id="C357650AB827F71264CF1D3667635A7F" box="[151,598,535,562]" pageId="5" pageNumber="385" type="nomenclature">
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<paragraph id="8BF23681B827F71264CF1D3667635A7F" blockId="5.[151,598,535,599]" box="[151,598,535,562]" pageId="5" pageNumber="385">
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<heading id="D0BA81EDB827F71264CF1D3667635A7F" bold="true" box="[151,598,535,562]" fontSize="11" level="1" pageId="5" pageNumber="385" reason="1">
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<emphasis id="B939EA93B827F71264CF1D3667635A7F" bold="true" box="[151,598,535,562]" pageId="5" pageNumber="385">
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<taxonomicName id="4C4D4D02B827F71264CF1D3664C75A7F" authority="Mikhaljova, 2023" authorityName="Mikhaljova" authorityYear="2023" box="[151,498,535,562]" class="Diplopoda" family="Julidae" genus="Nepalmatoiulus" kingdom="Animalia" order="Julida" pageId="5" pageNumber="385" phylum="Arthropoda" rank="species" species="simultaneus" status="sp. nov.">
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<emphasis id="B939EA93B827F71264CF1D3664C75A7F" bold="true" box="[151,498,535,562]" italics="true" pageId="5" pageNumber="385">Nepalmatoiulus simultaneus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20A57E8B827F71265A11D3967635A7F" box="[505,598,536,562]" pageId="5" pageNumber="385" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C357650AB827F71264CF1D1D640B5A1B" box="[151,318,572,599]" pageId="5" pageNumber="385" type="description">
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<paragraph id="8BF23681B827F71264CF1D1D640B5A1B" blockId="5.[151,598,535,599]" box="[151,318,572,599]" pageId="5" pageNumber="385">
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<figureCitation id="13762A04B827F71264CF1D1D64225A1A" box="[151,279,572,599]" captionStart-0="FIGURES 15–20" captionStart-1="FIGURES 21–26" captionStartId-0="6.[152,270,1615,1639]" captionStartId-1="7.[152,270,1857,1881]" captionTargetBox-0="[158,1430,181,1580]" captionTargetBox-1="[151,1436,417,1834]" captionTargetId-0="figure-15@6.[158,1430,181,1591]" captionTargetId-1="figure-122@7.[151,1436,417,1834]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURES 15–20. Nepalmatoiulus simultaneus sp. nov., male (Figs 15–18, 20, FSCB) and female (Fig. 19, ZMUM) paratypes. 15. Leg pair 1, anterior view. 16. Penis, caudal view. 17. Leg pair 2, anterior view. 18. Claw 2, anterior view. 19. Left vulva, lateral view. 20. Opisthomere, anteromesal view. Abbreviation: d, outgrowth of mesomeral process. Scales: 10 μm (Fig. 18), 20 μm (Figs 15–16), 100 μm (Figs 17, 19), 0.5 mm (Fig. 20)." captionText-1="FIGURES 21–26. Nepalmatoiulus simultaneus sp. nov., male holotype (ZMUM) and male paratype (FSCB). 21. Left promere of male-paratype, caudal view. 22. Left promere of male-holotype, caudal view. 23. Opisthomere, mesal view. 24. Distal part of left promere of male-paratype, caudal view. 25. Distal part of left promere of male-holotype, caudal view. 26. Velum, solenomere and accessory membrane. Abbreviations: a, protrusion; d, outgrowth of mesomeral process; m, accessory membrane. Scales: 10 μm (Fig. 24), 20 μm (Figs 25–26), 100 μm (Figs 21–23)." figureDoi-0="http://doi.org/10.5281/zenodo.8005008" figureDoi-1="http://doi.org/10.5281/zenodo.8005010" httpUri-0="https://zenodo.org/record/8005008/files/figure.png" httpUri-1="https://zenodo.org/record/8005010/files/figure.png" pageId="5" pageNumber="385">Figs 15–26</figureCitation>
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,
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<figureCitation id="13762A04B827F712657A1D1D640B5A1B" box="[290,318,572,598]" captionStart="FIGURE 39" captionStartId="12.[152,255,852,876]" captionTargetBox="[225,1362,187,822]" captionTargetId="figure-14@12.[219,1369,181,828]" captionTargetPageId="12" captionText="FIGURE 39. Size relations of coexisting Nepalmatoiulus alternus sp. nov. (continuous lines, square - ♁, diamond - ♀) and Nepalmatoiulus simultaneus sp. nov. (stippled lines, heart - ♁, oval - ♀). H, midbody vertical diameter (mm), p.s., number of podous rings." figureDoi="http://doi.org/10.5281/zenodo.8005020" httpUri="https://zenodo.org/record/8005020/files/figure.png" pageId="5" pageNumber="385">39</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C357650AB827F71264CF1DA567995B47" pageId="5" pageNumber="385" type="materials_examined">
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<paragraph id="8BF23681B827F71264CF1DA567995B47" blockId="5.[151,1437,644,2038]" pageId="5" pageNumber="385">
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<emphasis id="B939EA93B827F71264CF1DA564B25AD3" bold="true" box="[151,391,644,670]" pageId="5" pageNumber="385">Material examined.</emphasis>
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<materialsCitation id="3B253CDCB827F71265CD1DA567DB5AAB" collectingDate="2014-05-06" collectionCode="ZMUM" collectorName="Laojunshan E & I. Belousov & Belousov, I." country="China" elevation="4075" latitude="26.882778" location="Laotzyunshan Mt." longLatPrecision="20" longitude="99.62889" pageId="5" pageNumber="385" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
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<typeStatus id="54F68823B827F71265CD1DA567375AD3" box="[405,514,644,670]" pageId="5" pageNumber="385">Holotype</typeStatus>
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: male (dissected) (
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<collectionCode id="ED5CAE44B827F71266BD1DA566775AD3" box="[741,834,644,670]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="5" pageNumber="385">ZMUM</collectionCode>
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),
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<collectingCountry id="F35A7611B827F71267031DA566945AD3" box="[859,929,644,670]" name="China" pageId="5" pageNumber="385">China</collectingCountry>
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,
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<collectingRegion id="4989F863B827F71267EA1DA4614E5AD3" box="[946,1147,644,671]" country="China" name="Yunnan" pageId="5" pageNumber="385">Yunnan Province</collectingRegion>
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,
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<collectorName id="26B85357B827F71260D51DA5601B5AD2" box="[1165,1326,644,671]" pageId="5" pageNumber="385">Laojunshan E</collectorName>
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slope of
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<location id="8E92605AB827F71264CF1D8964535A8E" LSID="urn:lsid:plazi:treatment:03E48797B827F71064CF1D3664C559C4:8E92605AB827F71264CF1D8964535A8E" box="[151,358,680,707]" country="China" latitude="26.882778" longLatPrecision="20" longitude="99.62889" name="Laotzyunshan Mt." pageId="5" pageNumber="385" stateProvince="Yunnan">Laotzyunshan Mt.</location>
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,
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<geoCoordinate id="EE795046B827F712652A1D89673E5A8E" box="[370,523,680,707]" degrees="26" direction="north" minutes="52" orientation="latitude" pageId="5" pageNumber="385" precision="15" seconds="58" value="26.882778">26°52´58´´ N</geoCoordinate>
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,
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<geoCoordinate id="EE795046B827F712664F1D8967995A8E" box="[535,684,680,707]" degrees="99" direction="east" minutes="37" orientation="longitude" pageId="5" pageNumber="385" precision="15" seconds="44" value="99.62889">99°37´44´´ E</geoCoordinate>
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, H =
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<quantity id="4CB59B64B827F71266B31D8966775A8E" box="[747,834,680,707]" metricMagnitude="3" metricUnit="m" metricValue="4.075" pageId="5" pageNumber="385" unit="m" value="4075.0">
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<elevation id="0060D1B2B827F71266B31D8966775A8E" box="[747,834,680,707]" metricMagnitude="3" metricUnit="m" metricValue="4.075" pageId="5" pageNumber="385" unit="m" value="4075.0">4075 m</elevation>
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</quantity>
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, crooked
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<taxonomicName id="4C4D4D02B827F71267E81D8961605A8F" box="[944,1109,680,706]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="385" phylum="Tracheophyta" rank="genus">
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||
<emphasis id="B939EA93B827F71267E81D8961605A8F" box="[944,1109,680,706]" italics="true" pageId="5" pageNumber="385">Rhododendron</emphasis>
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</taxonomicName>
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forest, wet litter + numerous streams, sifter,
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<date id="FFF31041B827F71265181DED64F45AAA" box="[320,449,716,743]" pageId="5" pageNumber="385" value="2014-05-06">
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||
<collectingDate id="EFB7E9A9B827F71265181DED64F45AAA" box="[320,449,716,743]" pageId="5" pageNumber="385" value="2014-05-06">05.06.2014</collectingDate>
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</date>
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, leg.
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<collectorName id="26B85357B827F71265A21DEC67495AAB" box="[506,636,716,743]" pageId="5" pageNumber="385">I. Belousov</collectorName>
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, I. Kabak.
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</materialsCitation>
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<materialsCitation id="3B253CDCB827F71266AB1DEC61A75AAB" box="[755,1170,716,743]" collectionCode="FSCB" country="China" pageId="5" pageNumber="385" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
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<typeStatus id="54F68823B827F71266AB1DEC66515AAA" box="[755,868,717,743]" pageId="5" pageNumber="385">Paratypes</typeStatus>
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:
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<specimenCount id="9D4BFD08B827F71267361DED66865AAB" box="[878,947,716,742]" pageId="5" pageNumber="385" type="male">1male</specimenCount>
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(dissected) (
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<collectionCode id="ED5CAE44B827F71260191DED61BE5AAB" box="[1089,1163,716,742]" pageId="5" pageNumber="385">FSCB</collectionCode>
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)
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</materialsCitation>
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,
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<materialsCitation id="3B253CDCB827F71260C51DED679D5B47" collectingDate="2014-05-06" collectionCode="ZMUM" collectorName="Laojunshan E & I. Belousov & Belousov, I." country="China" elevation="4075" latitude="26.882778" location="Laotzyunshan Mt." longLatPrecision="20" longitude="99.62889" pageId="5" pageNumber="385" specimenCount="3" specimenCount-female="3" stateProvince="Yunnan" typeStatus="paratype">
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<specimenCount id="9D4BFD08B827F71260C51DED603D5AAB" box="[1181,1288,716,742]" pageId="5" pageNumber="385" type="female">3 females</specimenCount>
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(one of them dissected) (
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<collectionCode id="ED5CAE44B827F712654E1DD164465B47" box="[278,371,752,778]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="5" pageNumber="385">ZMUM</collectionCode>
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), same data as for holotype
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C357650AB827F712649F1C35672E5BF3" pageId="5" pageNumber="385" type="diagnosis">
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||
<paragraph id="8BF23681B827F712649F1C35672E5BF3" blockId="5.[151,1437,644,2038]" pageId="5" pageNumber="385">
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||
<emphasis id="B939EA93B827F712649F1C3564775B63" bold="true" box="[199,322,788,814]" pageId="5" pageNumber="385">Diagnosis.</emphasis>
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Differs from congeners mainly by the presence of the strongly papillate cone-shaped subapical outgrowth (
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<emphasis id="B939EA93B827F71265461C19641B5B1F" bold="true" box="[286,302,824,850]" pageId="5" pageNumber="385">d</emphasis>
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||
in
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<figureCitation id="13762A04B827F712650F1C1964855B1F" box="[343,432,824,850]" captionStart="FIGURES 15–20" captionStartId="6.[152,270,1615,1639]" captionTargetBox="[158,1430,181,1580]" captionTargetId="figure-15@6.[158,1430,181,1591]" captionTargetPageId="6" captionText="FIGURES 15–20. Nepalmatoiulus simultaneus sp. nov., male (Figs 15–18, 20, FSCB) and female (Fig. 19, ZMUM) paratypes. 15. Leg pair 1, anterior view. 16. Penis, caudal view. 17. Leg pair 2, anterior view. 18. Claw 2, anterior view. 19. Left vulva, lateral view. 20. Opisthomere, anteromesal view. Abbreviation: d, outgrowth of mesomeral process. Scales: 10 μm (Fig. 18), 20 μm (Figs 15–16), 100 μm (Figs 17, 19), 0.5 mm (Fig. 20)." figureDoi="http://doi.org/10.5281/zenodo.8005008" httpUri="https://zenodo.org/record/8005008/files/figure.png" pageId="5" pageNumber="385">Figs 20</figureCitation>
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,
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<figureCitation id="13762A04B827F71265E61C1964EE5B1F" box="[446,475,824,850]" captionStart="FIGURES 21–26" captionStartId="7.[152,270,1857,1881]" captionTargetBox="[151,1436,417,1834]" captionTargetId="figure-122@7.[151,1436,417,1834]" captionTargetPageId="7" captionText="FIGURES 21–26. Nepalmatoiulus simultaneus sp. nov., male holotype (ZMUM) and male paratype (FSCB). 21. Left promere of male-paratype, caudal view. 22. Left promere of male-holotype, caudal view. 23. Opisthomere, mesal view. 24. Distal part of left promere of male-paratype, caudal view. 25. Distal part of left promere of male-holotype, caudal view. 26. Velum, solenomere and accessory membrane. Abbreviations: a, protrusion; d, outgrowth of mesomeral process; m, accessory membrane. Scales: 10 μm (Fig. 24), 20 μm (Figs 25–26), 100 μm (Figs 21–23)." figureDoi="http://doi.org/10.5281/zenodo.8005010" httpUri="https://zenodo.org/record/8005010/files/figure.png" pageId="5" pageNumber="385">23</figureCitation>
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) of the posterior gonopod mesomeral process and the absence of the velar process coupled with the apically obliquely rounded anterior gonopod and subrectangular penis. Particularly similar to
|
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<taxonomicName id="4C4D4D02B827F71264CF1CA164D65BD7" authorityName="Mikhaljova" authorityYear="2020" box="[151,483,896,922]" class="Diplopoda" family="Julidae" genus="Nepalmatoiulus" kingdom="Animalia" order="Julida" pageId="5" pageNumber="385" phylum="Arthropoda" rank="species" species="lanpingensis">
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||
<emphasis id="B939EA93B827F71264CF1CA164D65BD7" box="[151,483,896,922]" italics="true" pageId="5" pageNumber="385">Nepalmatoiulus lanpingensis</emphasis>
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</taxonomicName>
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and
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||
<taxonomicName id="4C4D4D02B827F712667C1CA1660B5BD7" authority="Mikhaljova, 2023" authorityName="Mikhaljova" authorityYear="2023" box="[548,830,896,922]" class="Diplopoda" family="Julidae" genus="Nepalmatoiulus" kingdom="Animalia" order="Julida" pageId="5" pageNumber="385" phylum="Arthropoda" rank="species" species="alternus" status="sp. nov.">
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||
<emphasis id="B939EA93B827F712667C1CA1660B5BD7" box="[548,830,896,922]" italics="true" pageId="5" pageNumber="385">Nepalmatoiulus alternus</emphasis>
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</taxonomicName>
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||
<emphasis id="B939EA93B827F71267121CA1669C5BD7" bold="true" box="[842,937,896,922]" pageId="5" pageNumber="385">
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||
<taxonomicNameLabel id="A20A57E8B827F71267121CA1669C5BD7" box="[842,937,896,922]" pageId="5" pageNumber="385" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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, but differs from these species by certain characters (see Discussion below).
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C357650AB827F710649F1CE9674B590C" lastPageId="7" lastPageNumber="387" pageId="5" pageNumber="385" type="description">
|
||
<paragraph id="8BF23681B827F712649F1CE966165C3F" blockId="5.[151,1437,644,2038]" pageId="5" pageNumber="385">
|
||
<emphasis id="B939EA93B827F712649F1CE9646C5BAF" bold="true" box="[199,345,968,994]" pageId="5" pageNumber="385">Description.</emphasis>
|
||
<emphasis id="B939EA93B827F71265381CE964AF5BAF" box="[352,410,968,994]" italics="true" pageId="5" pageNumber="385">Male</emphasis>
|
||
. Length in alcohol 27.0–30.0 mm, midbody vertical diameter 2.0– 2.2 mm, with 56(–1) (in a male
|
||
<typeStatus id="54F68823B827F71264B21CCD64655C4B" box="[234,336,1004,1030]" pageId="5" pageNumber="385">holotype</typeStatus>
|
||
), 48(–1) rings, excluding telson. Coloration in alcohol from dark brown-grey to brown-gray with broad light longitudinal dorsal strip formed by lighter dorso-anterior metazonital portions. Venter (including basal parts of legs) brown. Legs dorsally and antennae dark brown. Occiput, forehead and collum from black brown to dark brown, ventral parts of head light brown. Eyes black.
|
||
</paragraph>
|
||
<paragraph id="8BF23681B827F712649F1B5D670F5D6B" blockId="5.[151,1437,644,2038]" pageId="5" pageNumber="385">
|
||
Head smooth and sparsely setose, 2 epicranial setae, 4 supralabral setae; not less than 12 labral setae in the
|
||
<typeStatus id="54F68823B827F71264CF1B8065CE5CF6" box="[151,251,1185,1211]" pageId="5" pageNumber="385">paratype</typeStatus>
|
||
. Eye patches almost oval, composed of not less than 43 ommatidia.Antennae medium-sized, rather slender and clavate. Gnathochilarium with no less than 8 nonapical stipital setae; lamellae linguales each with not less than 6 (in the
|
||
<typeStatus id="54F68823B827F712655A1BC964515D4F" box="[258,356,1256,1282]" pageId="5" pageNumber="385">paratype</typeStatus>
|
||
) setae; mandibular stipites each with subtriangular rounded smooth lobe; apical portion of the lobe without well demarcated border.
|
||
</paragraph>
|
||
<paragraph id="8BF23681B827F712649F1A11661A5EFF" blockId="5.[151,1437,644,2038]" pageId="5" pageNumber="385">
|
||
Collum laterally with several distinct longer lower striae at posterior margin, not reaching to anterior margin, dorsally with short striae at posterior margin only. Length of the collum striae gradually decreasing toward dorsum. A transverse row of sparse setae at hind edge of collum. Body rings circular.Anterior body half with smooth prozona. Prozona of posterior body half generally smooth, but with barely noticeable, very shallow obliquely located striae. Metazona with dense, regular, longitudinal striae reaching hind margin (16 or 17 striae in an approximate square with sides equal to metazonital length of a dorsal side of midbody ring). Metazonite gradually growing somewhat narrower toward body venter. Limbus straight, smooth (of
|
||
<typeStatus id="54F68823B827F7126711192866B45E6E" box="[841,897,1545,1571]" pageId="5" pageNumber="385">Type</typeStatus>
|
||
<quantity id="4CB59B64B827F71267D31929668F5E6F" box="[907,954,1544,1570]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="5" pageNumber="385" unit="in" value="1.0">1 in</quantity>
|
||
<bibRefCitation id="EFDC4B70B827F712679D1929615D5E6F" author="Enghoff, H." box="[965,1128,1544,1570]" pageId="5" pageNumber="385" pagination="241 - 331" refId="ref5882" refString="Enghoff, H. (1987) Revision of Nepalmatoiulus Mauries, 1983 - a Southeast Asiatic genus of millipedes (Diplopoda: Julida: Julidae). Courier Forschung-Institut Senckenberg, 93, 241 - 331." type="journal article" year="1987">Enghoff 1987</bibRefCitation>
|
||
). A transverse row of thin setae at hind edge of metazonites, setae gradually growing longer and denser toward telson. Ozopores small, lying immediately behind suture dividing pro- and metazona. Caudal dorsal projection of telson straight and long, covered with setae and carrying at tip a claw-shaped process curved dorsally. Preanal ring setose; anal valves densely setose distally; subanal scale densely setose only in posterior half.
|
||
</paragraph>
|
||
<paragraph id="8BF23681B827F712649F199C644C5FBB" blockId="5.[151,1437,644,2038]" pageId="5" pageNumber="385">
|
||
Legs relatively short and slender. Very delicately serrate ventral pads on postfemur and tibia, starting from legs 2 (
|
||
<figureCitation id="13762A04B827F71264ED19C1643C5EB7" box="[181,265,1760,1786]" captionStart="FIGURES 15–20" captionStartId="6.[152,270,1615,1639]" captionTargetBox="[158,1430,181,1580]" captionTargetId="figure-15@6.[158,1430,181,1591]" captionTargetPageId="6" captionText="FIGURES 15–20. Nepalmatoiulus simultaneus sp. nov., male (Figs 15–18, 20, FSCB) and female (Fig. 19, ZMUM) paratypes. 15. Leg pair 1, anterior view. 16. Penis, caudal view. 17. Leg pair 2, anterior view. 18. Claw 2, anterior view. 19. Left vulva, lateral view. 20. Opisthomere, anteromesal view. Abbreviation: d, outgrowth of mesomeral process. Scales: 10 μm (Fig. 18), 20 μm (Figs 15–16), 100 μm (Figs 17, 19), 0.5 mm (Fig. 20)." figureDoi="http://doi.org/10.5281/zenodo.8005008" httpUri="https://zenodo.org/record/8005008/files/figure.png" pageId="5" pageNumber="385">Fig. 17</figureCitation>
|
||
). The ventral pads gradually decrease towards telson; tibia of most hind legs with ventral pads on distal part only. Claw of all legs at base with a long (somewhat longer than claw) setiform accessory claw ventrally (
|
||
<figureCitation id="13762A04B827F7126137182565835F0F" captionStart="FIGURES 15–20" captionStartId="6.[152,270,1615,1639]" captionTargetBox="[158,1430,181,1580]" captionTargetId="figure-15@6.[158,1430,181,1591]" captionTargetPageId="6" captionText="FIGURES 15–20. Nepalmatoiulus simultaneus sp. nov., male (Figs 15–18, 20, FSCB) and female (Fig. 19, ZMUM) paratypes. 15. Leg pair 1, anterior view. 16. Penis, caudal view. 17. Leg pair 2, anterior view. 18. Claw 2, anterior view. 19. Left vulva, lateral view. 20. Opisthomere, anteromesal view. Abbreviation: d, outgrowth of mesomeral process. Scales: 10 μm (Fig. 18), 20 μm (Figs 15–16), 100 μm (Figs 17, 19), 0.5 mm (Fig. 20)." figureDoi="http://doi.org/10.5281/zenodo.8005008" httpUri="https://zenodo.org/record/8005008/files/figure.png" pageId="5" pageNumber="385">Fig. 18</figureCitation>
|
||
). Leg pair 1 forming hook, distal segment not in close contact with the basal segments (“Open hook”
|
||
<typeStatus id="54F68823B827F7126115180860485F0E" box="[1357,1405,1833,1859]" pageId="5" pageNumber="385">type</typeStatus>
|
||
in
|
||
<bibRefCitation id="EFDC4B70B827F71264CF186D64005F2B" author="Enghoff, H." box="[151,309,1868,1894]" pageId="5" pageNumber="385" pagination="241 - 331" refId="ref5882" refString="Enghoff, H. (1987) Revision of Nepalmatoiulus Mauries, 1983 - a Southeast Asiatic genus of millipedes (Diplopoda: Julida: Julidae). Courier Forschung-Institut Senckenberg, 93, 241 - 331." type="journal article" year="1987">Enghoff 1987</bibRefCitation>
|
||
). Leg pair 1: postfemur with slightly scaly-rugose ventral surface; tip ventrally slightly rugosity; coxa with one seta; distal segment laterally with small seta, without tarsal remnant (
|
||
<figureCitation id="13762A04B827F712605D1851616C5FC6" box="[1029,1113,1904,1931]" captionStart="FIGURES 15–20" captionStartId="6.[152,270,1615,1639]" captionTargetBox="[158,1430,181,1580]" captionTargetId="figure-15@6.[158,1430,181,1591]" captionTargetPageId="6" captionText="FIGURES 15–20. Nepalmatoiulus simultaneus sp. nov., male (Figs 15–18, 20, FSCB) and female (Fig. 19, ZMUM) paratypes. 15. Leg pair 1, anterior view. 16. Penis, caudal view. 17. Leg pair 2, anterior view. 18. Claw 2, anterior view. 19. Left vulva, lateral view. 20. Opisthomere, anteromesal view. Abbreviation: d, outgrowth of mesomeral process. Scales: 10 μm (Fig. 18), 20 μm (Figs 15–16), 100 μm (Figs 17, 19), 0.5 mm (Fig. 20)." figureDoi="http://doi.org/10.5281/zenodo.8005008" httpUri="https://zenodo.org/record/8005008/files/figure.png" pageId="5" pageNumber="385">Fig. 15</figureCitation>
|
||
). Coxa 2 with one mesapical oral seta, a gland opening positioned in central and axial position (
|
||
<figureCitation id="13762A04B827F71267F418B561345FE3" box="[940,1025,1940,1966]" captionStart="FIGURES 15–20" captionStartId="6.[152,270,1615,1639]" captionTargetBox="[158,1430,181,1580]" captionTargetId="figure-15@6.[158,1430,181,1591]" captionTargetPageId="6" captionText="FIGURES 15–20. Nepalmatoiulus simultaneus sp. nov., male (Figs 15–18, 20, FSCB) and female (Fig. 19, ZMUM) paratypes. 15. Leg pair 1, anterior view. 16. Penis, caudal view. 17. Leg pair 2, anterior view. 18. Claw 2, anterior view. 19. Left vulva, lateral view. 20. Opisthomere, anteromesal view. Abbreviation: d, outgrowth of mesomeral process. Scales: 10 μm (Fig. 18), 20 μm (Figs 15–16), 100 μm (Figs 17, 19), 0.5 mm (Fig. 20)." figureDoi="http://doi.org/10.5281/zenodo.8005008" httpUri="https://zenodo.org/record/8005008/files/figure.png" pageId="5" pageNumber="385">Fig. 17</figureCitation>
|
||
) according to
|
||
<bibRefCitation id="EFDC4B70B827F71260F718B560655FE3" author="Enghoff, H." box="[1199,1360,1940,1966]" pageId="5" pageNumber="385" pagination="241 - 331" refId="ref5882" refString="Enghoff, H. (1987) Revision of Nepalmatoiulus Mauries, 1983 - a Southeast Asiatic genus of millipedes (Diplopoda: Julida: Julidae). Courier Forschung-Institut Senckenberg, 93, 241 - 331." type="journal article" year="1987">Enghoff 1987</bibRefCitation>
|
||
. Penis subrectangular, about 2.2 longer than wide (
|
||
<figureCitation id="13762A04B827F71266D3189967EA5F9F" box="[651,735,1976,2002]" captionStart="FIGURES 15–20" captionStartId="6.[152,270,1615,1639]" captionTargetBox="[158,1430,181,1580]" captionTargetId="figure-15@6.[158,1430,181,1591]" captionTargetPageId="6" captionText="FIGURES 15–20. Nepalmatoiulus simultaneus sp. nov., male (Figs 15–18, 20, FSCB) and female (Fig. 19, ZMUM) paratypes. 15. Leg pair 1, anterior view. 16. Penis, caudal view. 17. Leg pair 2, anterior view. 18. Claw 2, anterior view. 19. Left vulva, lateral view. 20. Opisthomere, anteromesal view. Abbreviation: d, outgrowth of mesomeral process. Scales: 10 μm (Fig. 18), 20 μm (Figs 15–16), 100 μm (Figs 17, 19), 0.5 mm (Fig. 20)." figureDoi="http://doi.org/10.5281/zenodo.8005008" httpUri="https://zenodo.org/record/8005008/files/figure.png" pageId="5" pageNumber="385">Fig. 16</figureCitation>
|
||
). Ventral margin of body ring VII with lobes similar to figure
|
||
<quantity id="4CB59B64B827F71264CF18FD65E55FBB" box="[151,208,2012,2038]" metricMagnitude="-1" metricUnit="m" metricValue="8.128" pageId="5" pageNumber="385" unit="in" value="32.0">32 in</quantity>
|
||
<bibRefCitation id="EFDC4B70B827F712648F18FD64415FBB" author="Enghoff, H." box="[215,372,2012,2038]" pageId="5" pageNumber="385" pagination="241 - 331" refId="ref5882" refString="Enghoff, H. (1987) Revision of Nepalmatoiulus Mauries, 1983 - a Southeast Asiatic genus of millipedes (Diplopoda: Julida: Julidae). Courier Forschung-Institut Senckenberg, 93, 241 - 331." type="journal article" year="1987">Enghoff 1987</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF326609B824F71164C0196E66365E99" ID-DOI="http://doi.org/10.5281/zenodo.8005008" ID-Zenodo-Dep="8005008" httpUri="https://zenodo.org/record/8005008/files/figure.png" pageId="6" pageNumber="386" startId="6.[152,270,1615,1639]" targetBox="[158,1430,181,1580]" targetPageId="6" targetType="figure">
|
||
<paragraph id="8BF23681B824F71164C0196E66365E99" blockId="6.[151,1437,1615,1748]" pageId="6" pageNumber="386">
|
||
<emphasis id="B939EA93B824F71164C0196E646C5E2A" bold="true" box="[152,345,1615,1639]" pageId="6" pageNumber="386">FIGURES 15–20.</emphasis>
|
||
<taxonomicName id="4C4D4D02B824F7116506196E67485E2A" authority="Mikhaljova, 2023" authorityName="Mikhaljova" authorityYear="2023" box="[350,637,1615,1639]" class="Diplopoda" family="Julidae" genus="Nepalmatoiulus" kingdom="Animalia" order="Julida" pageId="6" pageNumber="386" phylum="Arthropoda" rank="species" species="simultaneus" status="sp. nov.">
|
||
<emphasis id="B939EA93B824F7116506196E67485E2A" box="[350,637,1615,1639]" italics="true" pageId="6" pageNumber="386">Nepalmatoiulus simultaneus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B939EA93B824F71166DA197167E75E25" bold="true" box="[642,722,1616,1640]" pageId="6" pageNumber="386">
|
||
<taxonomicNameLabel id="A20A57E8B824F71166DA197167E75E25" box="[642,722,1616,1640]" pageId="6" pageNumber="386" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male (Figs 15–18, 20, FSCB) and female (Fig. 19, ZMUM) paratypes.
|
||
<emphasis id="B939EA93B824F71164CF195265855EC6" bold="true" box="[151,176,1651,1675]" pageId="6" pageNumber="386">15</emphasis>
|
||
. Leg pair 1, anterior view.
|
||
<emphasis id="B939EA93B824F7116593195264D15EC6" bold="true" box="[459,484,1651,1675]" pageId="6" pageNumber="386">16</emphasis>
|
||
. Penis, caudal view.
|
||
<emphasis id="B939EA93B824F71166E6195267E25EC6" bold="true" box="[702,727,1651,1675]" pageId="6" pageNumber="386">17</emphasis>
|
||
. Leg pair 2, anterior view.
|
||
<emphasis id="B939EA93B824F71167AA1952613E5EC6" bold="true" box="[1010,1035,1651,1675]" pageId="6" pageNumber="386">18</emphasis>
|
||
. Claw 2, anterior view.
|
||
<emphasis id="B939EA93B824F711615D1952602B5EC6" bold="true" box="[1285,1310,1651,1675]" pageId="6" pageNumber="386">19</emphasis>
|
||
. Left vulva, lateral view.
|
||
<emphasis id="B939EA93B824F711654319B664015EE2" bold="true" box="[283,308,1687,1711]" pageId="6" pageNumber="386">20</emphasis>
|
||
. Opisthomere, anteromesal view.
|
||
<emphasis id="B939EA93B824F71166C819B666085EFD" bold="true" box="[656,829,1687,1712]" pageId="6" pageNumber="386">Abbreviation: d</emphasis>
|
||
, outgrowth of mesomeral process. Scales: 10 μm (Fig. 18), 20 μm (Figs 15–16), 100 μm (Figs 17, 19), 0.5 mm (Fig. 20).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF23681B824F710649F1823666158B4" blockId="6.[151,1436,1794,2036]" lastBlockId="7.[151,1437,150,393]" lastPageId="7" lastPageNumber="387" pageId="6" pageNumber="386">
|
||
Gonopods slightly protruding. Anterior gonopod flattened, with subparallel margins, distally slightly expanded, apically relatively obliquely rounded, in caudal view subapically excavated for accommodation of mesomeral process, distal margins of the excavation papillate (especially mesal margin), rugose margin of apical excavation somewhat produced into low short blunt caudad protrusion (
|
||
<emphasis id="B939EA93B824F7116712184F666D5FC5" bold="true" box="[842,856,1902,1928]" pageId="6" pageNumber="386">a</emphasis>
|
||
), subcentrally with knoblike flattened rudimentary telopodite (
|
||
<figureCitation id="13762A04B824F711654E18B364DB5FE1" box="[278,494,1937,1964]" captionStart="FIGURES 21–26" captionStartId="7.[152,270,1857,1881]" captionTargetBox="[151,1436,417,1834]" captionTargetId="figure-122@7.[151,1436,417,1834]" captionTargetPageId="7" captionText="FIGURES 21–26. Nepalmatoiulus simultaneus sp. nov., male holotype (ZMUM) and male paratype (FSCB). 21. Left promere of male-paratype, caudal view. 22. Left promere of male-holotype, caudal view. 23. Opisthomere, mesal view. 24. Distal part of left promere of male-paratype, caudal view. 25. Distal part of left promere of male-holotype, caudal view. 26. Velum, solenomere and accessory membrane. Abbreviations: a, protrusion; d, outgrowth of mesomeral process; m, accessory membrane. Scales: 10 μm (Fig. 24), 20 μm (Figs 25–26), 100 μm (Figs 21–23)." figureDoi="http://doi.org/10.5281/zenodo.8005010" httpUri="https://zenodo.org/record/8005010/files/figure.png" pageId="6" pageNumber="386">Figs 21–22, 24–25</figureCitation>
|
||
). In addition, anterior gonopod apex shows some variation. Thus, male
|
||
<typeStatus id="54F68823B824F711615718B3605A5FE1" box="[1295,1391,1938,1964]" pageId="6" pageNumber="386">paratype</typeStatus>
|
||
has a more rounded anterior gonopod apex and a more extensive papillary field (
|
||
<figureCitation id="13762A04B824F71167B31897615F5F9D" box="[1003,1130,1974,2000]" captionStart="FIGURES 21–26" captionStartId="7.[152,270,1857,1881]" captionTargetBox="[151,1436,417,1834]" captionTargetId="figure-122@7.[151,1436,417,1834]" captionTargetPageId="7" captionText="FIGURES 21–26. Nepalmatoiulus simultaneus sp. nov., male holotype (ZMUM) and male paratype (FSCB). 21. Left promere of male-paratype, caudal view. 22. Left promere of male-holotype, caudal view. 23. Opisthomere, mesal view. 24. Distal part of left promere of male-paratype, caudal view. 25. Distal part of left promere of male-holotype, caudal view. 26. Velum, solenomere and accessory membrane. Abbreviations: a, protrusion; d, outgrowth of mesomeral process; m, accessory membrane. Scales: 10 μm (Fig. 24), 20 μm (Figs 25–26), 100 μm (Figs 21–23)." figureDoi="http://doi.org/10.5281/zenodo.8005010" httpUri="https://zenodo.org/record/8005010/files/figure.png" pageId="6" pageNumber="386">Figs 21, 24</figureCitation>
|
||
) than that of male
|
||
<typeStatus id="54F68823B824F7116162189760A95F9D" box="[1338,1436,1974,2000]" pageId="6" pageNumber="386">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="13762A04B824F71164C718FB64165FB9" box="[159,291,2009,2036]" captionStart="FIGURES 21–26" captionStartId="7.[152,270,1857,1881]" captionTargetBox="[151,1436,417,1834]" captionTargetId="figure-122@7.[151,1436,417,1834]" captionTargetPageId="7" captionText="FIGURES 21–26. Nepalmatoiulus simultaneus sp. nov., male holotype (ZMUM) and male paratype (FSCB). 21. Left promere of male-paratype, caudal view. 22. Left promere of male-holotype, caudal view. 23. Opisthomere, mesal view. 24. Distal part of left promere of male-paratype, caudal view. 25. Distal part of left promere of male-holotype, caudal view. 26. Velum, solenomere and accessory membrane. Abbreviations: a, protrusion; d, outgrowth of mesomeral process; m, accessory membrane. Scales: 10 μm (Fig. 24), 20 μm (Figs 25–26), 100 μm (Figs 21–23)." figureDoi="http://doi.org/10.5281/zenodo.8005010" httpUri="https://zenodo.org/record/8005010/files/figure.png" pageId="6" pageNumber="386">Figs 22, 25</figureCitation>
|
||
). Flagellum slender, short, caudally covered with cuticular spikes in the distal part. Opisthomere with apically hook-shaped mesomeral process and subapical, cone-shaped, apically strongly papillate outgrowth (
|
||
<emphasis id="B939EA93B825F71061DB1FB760A958FD" bold="true" box="[1411,1436,150,176]" pageId="7" pageNumber="387">d)</emphasis>
|
||
(
|
||
<figureCitation id="13762A04B825F71064C71F9A65C35898" box="[159,246,187,213]" captionStart="FIGURES 15–20" captionStartId="6.[152,270,1615,1639]" captionTargetBox="[158,1430,181,1580]" captionTargetId="figure-15@6.[158,1430,181,1591]" captionTargetPageId="6" captionText="FIGURES 15–20. Nepalmatoiulus simultaneus sp. nov., male (Figs 15–18, 20, FSCB) and female (Fig. 19, ZMUM) paratypes. 15. Leg pair 1, anterior view. 16. Penis, caudal view. 17. Leg pair 2, anterior view. 18. Claw 2, anterior view. 19. Left vulva, lateral view. 20. Opisthomere, anteromesal view. Abbreviation: d, outgrowth of mesomeral process. Scales: 10 μm (Fig. 18), 20 μm (Figs 15–16), 100 μm (Figs 17, 19), 0.5 mm (Fig. 20)." figureDoi="http://doi.org/10.5281/zenodo.8005008" httpUri="https://zenodo.org/record/8005008/files/figure.png" pageId="7" pageNumber="387">Figs 20</figureCitation>
|
||
,
|
||
<figureCitation id="13762A04B825F71065591F9A64145898" box="[257,289,187,213]" captionStart="FIGURES 21–26" captionStartId="7.[152,270,1857,1881]" captionTargetBox="[151,1436,417,1834]" captionTargetId="figure-122@7.[151,1436,417,1834]" captionTargetPageId="7" captionText="FIGURES 21–26. Nepalmatoiulus simultaneus sp. nov., male holotype (ZMUM) and male paratype (FSCB). 21. Left promere of male-paratype, caudal view. 22. Left promere of male-holotype, caudal view. 23. Opisthomere, mesal view. 24. Distal part of left promere of male-paratype, caudal view. 25. Distal part of left promere of male-holotype, caudal view. 26. Velum, solenomere and accessory membrane. Abbreviations: a, protrusion; d, outgrowth of mesomeral process; m, accessory membrane. Scales: 10 μm (Fig. 24), 20 μm (Figs 25–26), 100 μm (Figs 21–23)." figureDoi="http://doi.org/10.5281/zenodo.8005010" httpUri="https://zenodo.org/record/8005010/files/figure.png" pageId="7" pageNumber="387">23</figureCitation>
|
||
). Membranous velum with not very sloped smooth margin. Margin of accessory membrane (
|
||
<emphasis id="B939EA93B825F71061721F9A60745898" bold="true" box="[1322,1345,187,213]" pageId="7" pageNumber="387">m</emphasis>
|
||
) serrate (
|
||
<figureCitation id="13762A04B825F71064C71FFE641758B4" box="[159,290,223,249]" captionStart="FIGURES 21–26" captionStartId="7.[152,270,1857,1881]" captionTargetBox="[151,1436,417,1834]" captionTargetId="figure-122@7.[151,1436,417,1834]" captionTargetPageId="7" captionText="FIGURES 21–26. Nepalmatoiulus simultaneus sp. nov., male holotype (ZMUM) and male paratype (FSCB). 21. Left promere of male-paratype, caudal view. 22. Left promere of male-holotype, caudal view. 23. Opisthomere, mesal view. 24. Distal part of left promere of male-paratype, caudal view. 25. Distal part of left promere of male-holotype, caudal view. 26. Velum, solenomere and accessory membrane. Abbreviations: a, protrusion; d, outgrowth of mesomeral process; m, accessory membrane. Scales: 10 μm (Fig. 24), 20 μm (Figs 25–26), 100 μm (Figs 21–23)." figureDoi="http://doi.org/10.5281/zenodo.8005010" httpUri="https://zenodo.org/record/8005010/files/figure.png" pageId="7" pageNumber="387">Figs 23, 26</figureCitation>
|
||
). Long and slender solenomere spinose anteriorly.
|
||
</paragraph>
|
||
<paragraph id="8BF23681B825F710649F1E22674B590C" blockId="7.[151,1437,150,393]" pageId="7" pageNumber="387">
|
||
<emphasis id="B939EA93B825F710649F1E22642E5950" box="[199,283,259,285]" italics="true" pageId="7" pageNumber="387">Female</emphasis>
|
||
. Length in alcohol 30.0–35.0 mm, midbody vertical diameter 3.2– 3.5 mm, with 58(–1), 51(–1), 47(–1) rings, excluding telson. Vulva as in
|
||
<figureCitation id="13762A04B825F71066701E06674F590C" box="[552,634,295,321]" captionStart="FIGURES 15–20" captionStartId="6.[152,270,1615,1639]" captionTargetBox="[158,1430,181,1580]" captionTargetId="figure-15@6.[158,1430,181,1591]" captionTargetPageId="6" captionText="FIGURES 15–20. Nepalmatoiulus simultaneus sp. nov., male (Figs 15–18, 20, FSCB) and female (Fig. 19, ZMUM) paratypes. 15. Leg pair 1, anterior view. 16. Penis, caudal view. 17. Leg pair 2, anterior view. 18. Claw 2, anterior view. 19. Left vulva, lateral view. 20. Opisthomere, anteromesal view. Abbreviation: d, outgrowth of mesomeral process. Scales: 10 μm (Fig. 18), 20 μm (Figs 15–16), 100 μm (Figs 17, 19), 0.5 mm (Fig. 20)." figureDoi="http://doi.org/10.5281/zenodo.8005008" httpUri="https://zenodo.org/record/8005008/files/figure.png" pageId="7" pageNumber="387">Fig. 19</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C357650AB825F710649F1E6A64C559C4" pageId="7" pageNumber="387" type="etymology">
|
||
<paragraph id="8BF23681B825F710649F1E6A64C559C4" blockId="7.[151,1437,150,393]" pageId="7" pageNumber="387">
|
||
<emphasis id="B939EA93B825F710649F1E6A647B5928" bold="true" box="[199,334,331,357]" pageId="7" pageNumber="387">Etymology.</emphasis>
|
||
The species epithet refers to the simultaneous finding of specimens of
|
||
<taxonomicName id="4C4D4D02B825F710603F1E6A60385928" authority="Mikhaljova, 2023" authorityName="Mikhaljova" authorityYear="2023" box="[1127,1293,331,357]" class="Diplopoda" family="Julidae" genus="Nepalmatoiulus" kingdom="Animalia" order="Julida" pageId="7" pageNumber="387" phylum="Arthropoda" rank="species" species="simultaneus" status="sp. nov.">
|
||
<emphasis id="B939EA93B825F710603F1E6A60385928" box="[1127,1293,331,357]" italics="true" pageId="7" pageNumber="387">N. simultaneus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B939EA93B825F710614C1E6A60585928" bold="true" box="[1300,1389,331,357]" pageId="7" pageNumber="387">
|
||
<taxonomicNameLabel id="A20A57E8B825F710614C1E6A60585928" box="[1300,1389,331,357]" pageId="7" pageNumber="387" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C4D4D02B825F71064CF1E4E642159C4" authority="Mikhaljova, 2023" authorityName="Mikhaljova" authorityYear="2023" box="[151,276,367,393]" class="Diplopoda" family="Julidae" genus="Nepalmatoiulus" kingdom="Animalia" order="Julida" pageId="7" pageNumber="387" phylum="Arthropoda" rank="species" species="alternus" status="sp. nov.">
|
||
<emphasis id="B939EA93B825F71064CF1E4E642159C4" box="[151,276,367,393]" italics="true" pageId="7" pageNumber="387">N. alternus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B939EA93B825F71065431E4E644159C4" bold="true" box="[283,372,367,393]" pageId="7" pageNumber="387">
|
||
<taxonomicNameLabel id="A20A57E8B825F71065431E4E644159C4" box="[283,372,367,393]" pageId="7" pageNumber="387" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Adjective.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |