425 lines
61 KiB
XML
425 lines
61 KiB
XML
<document id="978AF9220AFABF64ADAFF8248D1C64A6" ID-DOI="10.11646/zootaxa.5397.3.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="10468850" ID-ZooBank="975A734B-749B-4599-8C1E-7D5B0EA56175" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1704708069737" checkinUser="plazi" docAuthor="Anisyutkin, Leonid N., Vasilenko, Dmitry V. & Perkovsky, Evgeny E." docDate="2024" docId="03E48782FFACE90182CAF969FCDEF83D" docLanguage="en" docName="zootaxa.5397.3.2.pdf" docOrigin="Zootaxa 5397 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5397.3.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ectobius danekrae Anisyutkin et Perkovsky 2024, sp. nov." docType="treatment" docVersion="3" lastPageNumber="346" masterDocId="FFDDFFFAFFADE905825DFFC2FFF7FF89" masterDocTitle="A new amber representative of the genus Ectobius Stephens, 1835 (Blattodea: Ectobiidae) from Storebaelt (Denmark)" masterLastPageNumber="350" masterPageNumber="342" pageNumber="343" updateTime="1704709129728" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="D6CBD6F69447F1F69561AF99F3F73A6C">A new amber representative of the genus Ectobius Stephens, 1835 (Blattodea: Ectobiidae) from Storebaelt (Denmark)</mods:title>
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<mods:namePart id="49F0925DABA600183C184A939955413D">Anisyutkin, Leonid N.</mods:namePart>
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<mods:nameIdentifier id="F3B0951490C75A38A8E7598E6D90B650" type="ORCID">0000-0002-7587-9078</mods:nameIdentifier>
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<mods:nameIdentifier id="E09CE6F0A4B55E8E358A48A5CA131C27" type="email">nisyutkin@zin.ru</mods:nameIdentifier>
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<mods:name id="43CE9A6387E917F71E4F26677BCD3942" type="personal">
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<mods:roleTerm id="D3372975ED9A1941E6F6E797C0C8FA6B">Author</mods:roleTerm>
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<mods:namePart id="0D6D057B60EA2E881C7D8857C783F8E0">Vasilenko, Dmitry V.</mods:namePart>
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<mods:affiliation id="1AB85D467B9CB98749678B0163B84A7E">Cherepovets State University, Lunacharsky prospect 5, 162600, Cherepovets, Russia. & Borissiak Paleontological Institute, Russian Academy of Sciences, Moscow, 117647 Russia.</mods:affiliation>
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<mods:roleTerm id="193F5E81E011A7336A49E773C007341E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="75F75C0DD61A1FB909A08D07CC3AC003">Perkovsky, Evgeny E.</mods:namePart>
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<mods:affiliation id="38E27521F37B3D5C15063BDA8531761C">Natural History Museum of Denmark, Universitetsparken 15, Copenhagen, 2100 Denmark.</mods:affiliation>
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<mods:title id="9392B47768E0E2BD135C464A7C9BFC47">Zootaxa</mods:title>
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<mods:part id="7A2B17FFC351F0EA824B0F2149B158EC">
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<mods:date id="22615ECF7BDA12BFC193B2E018F73D9F">2024</mods:date>
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<mods:number id="60AD02777C4D8B0AF73B896CB34873D4">2024-01-05</mods:number>
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<mods:number id="0884137CE8E54A24675A71286D45090A">5397</mods:number>
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<mods:url id="E295100CB9CCF010990A4E61BB34919E">http://dx.doi.org/10.11646/zootaxa.5397.3.2</mods:url>
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<mods:classification id="A374B4C91D1C0B6D00946D3AB4C2AA0E">journal article</mods:classification>
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<treatment id="03E48782FFACE90182CAF969FCDEF83D" ID-DOI="http://doi.org/10.5281/zenodo.10468759" ID-Zenodo-Dep="10468759" LSID="urn:lsid:plazi:treatment:03E48782FFACE90182CAF969FCDEF83D" httpUri="http://treatment.plazi.org/id/03E48782FFACE90182CAF969FCDEF83D" lastPageId="4" lastPageNumber="346" pageId="1" pageNumber="343">
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<subSubSection id="C357651FFFACE90482CAF969FCE8F94F" box="[151,799,1707,1734]" pageId="1" pageNumber="343" type="nomenclature">
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<paragraph id="8BF23694FFACE90482CAF969FCE8F94F" blockId="1.[151,799,1707,1770]" box="[151,799,1707,1734]" pageId="1" pageNumber="343">
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<heading id="D0BA81F8FFACE90482CAF969FCE8F94F" bold="true" box="[151,799,1707,1734]" fontSize="11" level="1" pageId="1" pageNumber="343" reason="1">
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<emphasis id="B939EA86FFACE90482CAF969FCE8F94F" bold="true" box="[151,799,1707,1734]" pageId="1" pageNumber="343">
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<taxonomicName id="4C4D4D17FFACE90482CAF969FD43F94F" authority="Anisyutkin et Perkovsky" authorityName="Anisyutkin et Perkovsky" authorityYear="2024" box="[151,692,1707,1734]" class="Insecta" family="Ectobiidae" genus="Ectobius" kingdom="Animalia" order="Blattodea" pageId="1" pageNumber="343" phylum="Arthropoda" rank="species" species="danekrae" status="sp. nov.">
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<emphasis id="B939EA86FFACE90482CAF969FE8FF94F" bold="true" box="[151,376,1707,1734]" italics="true" pageId="1" pageNumber="343">Ectobius danekrae</emphasis>
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Anisyutkin et Perkovsky
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</taxonomicName>
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,
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<taxonomicNameLabel id="A20A57FDFFACE904809CF96EFCE8F94F" box="[705,799,1708,1734]" pageId="1" pageNumber="343" rank="species">sp. nov.</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C357651FFFACE90482CAF912FE8CF963" box="[151,379,1744,1770]" pageId="1" pageNumber="343" type="description">
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<paragraph id="8BF23694FFACE90482CAF912FE8CF963" blockId="1.[151,799,1707,1770]" box="[151,379,1744,1770]" pageId="1" pageNumber="343">
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(
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<figureCitation id="13762A11FFACE90482C2F912FED5F963" box="[159,290,1744,1770]" captionStart="FIGURE 1" captionStartId="2.[152,255,1679,1703]" captionTargetBox="[159,1428,189,1649]" captionTargetId="figure-21@2.[151,1436,179,1653]" captionTargetPageId="2" captionText="FIGURE 1. Ectobius danekrae Anisyutkin et Perkovsky, sp. nov., male holotype.A, B—general view of inclusion: from above (A) and below (B); C, D—general view from above (C) and below (D); E—abdominal apex from below." figureDoi="http://doi.org/10.5281/zenodo.10468856" httpUri="https://zenodo.org/record/10468856/files/figure.png" pageId="1" pageNumber="343">Figs. 1A–E</figureCitation>
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,
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<figureCitation id="13762A11FFACE9048373F912FE86F963" box="[302,369,1744,1770]" captionStart="FIGURE 2" captionStartId="4.[152,255,1443,1467]" captionTargetBox="[159,1427,189,1412]" captionTargetId="figure-21@4.[151,1435,181,1420]" captionTargetPageId="4" captionText="FIGURE 2. Ectobius danekrae Anisyutkin et Perkovsky, sp. nov., male holotype. A—outlines of body from above; B— abdominal apex and part of leg from below." figureDoi="http://doi.org/10.5281/zenodo.10468862" httpUri="https://zenodo.org/record/10468862/files/figure.png" pageId="1" pageNumber="343">2A, B</figureCitation>
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)
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C357651FFFACE90482CAF8DAFD4BF8DF" pageId="1" pageNumber="343" type="etymology">
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<paragraph id="8BF23694FFACE90482CAF8DAFD4BF8DF" blockId="1.[151,1437,1816,2023]" pageId="1" pageNumber="343">
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<emphasis id="B939EA86FFACE90482CAF8DAFEEFF8BB" bold="true" box="[151,280,1816,1842]" pageId="1" pageNumber="343">Etymology</emphasis>
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. Inspired by the Danish Statute, the term
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<taxonomicName id="4C4D4D17FFACE90480AFF8DAFC96F8BB" authorityName="Anisyutkin et Perkovsky" authorityYear="2024" box="[754,865,1816,1842]" class="Insecta" family="Ectobiidae" genus="Ectobius" kingdom="Animalia" order="Blattodea" pageId="1" pageNumber="343" phylum="Arthropoda" rank="species" species="danekrae">“danekrae</taxonomicName>
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” was introduced in 1989 as a term for earth-found natural objects of unique natural historical value.
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</paragraph>
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</subSubSection>
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<subSubSection id="C357651FFFACE904829AF8A2FB72F86F" pageId="1" pageNumber="343" type="materials_examined">
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<paragraph id="8BF23694FFACE904829AF8A2FDD6F84A" blockId="1.[151,1437,1816,2023]" pageId="1" pageNumber="343">
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||
<emphasis id="B939EA86FFACE904829AF8A2FED8F8F3" bold="true" box="[199,303,1888,1914]" pageId="1" pageNumber="343">Material</emphasis>
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.
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<materialsCitation id="3B253CC9FFACE9048363F8A2FDD6F84A" collectionCode="ZMUC" country="Denmark" county="Storebaelt" location="Danish" municipality="Musholm Bugt" pageId="1" pageNumber="343" specimenCode="ZMUC 902245" specimenCount="1" stateProvince="Sjaelland" typeStatus="holotype">
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<typeStatus id="54F68836FFACE9048363F8A2FE5EF8F3" box="[318,425,1888,1914]" pageId="1" pageNumber="343" type="holotype">Holotype</typeStatus>
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, male, northwestern
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<collectingRegion id="4989F876FFACE90480CEF8A2FD1AF8F3" box="[659,749,1888,1914]" country="Denmark" name="Sjaelland" pageId="1" pageNumber="343">Zealand</collectingRegion>
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||
(
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<collectingCountry id="F35A7604FFACE90480A0F8A2FC9CF8F3" box="[765,875,1888,1914]" name="Denmark" pageId="1" pageNumber="343">Denmark</collectingCountry>
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),
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<location id="8E92604FFFACE9048123F8A2FC3AF8F3" LSID="urn:lsid:plazi:treatment:03E48782FFACE90182CAF969FCDEF83D:8E92604FFFACE9048123F8A2FC3AF8F3" box="[894,973,1888,1914]" country="Denmark" county="Storebaelt" municipality="Musholm Bugt" name="Danish" pageId="1" pageNumber="343" stateProvince="Sjaelland">Danish</location>
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amber, late
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<location id="8E92604FFFACE9048604F8A3FB59F8F2" LSID="urn:lsid:plazi:treatment:03E48782FFACE90182CAF969FCDEF83D:8E92604FFFACE9048604F8A3FB59F8F2" box="[1113,1198,1889,1915]" country="Denmark" county="Storebaelt" municipality="Musholm Bugt" name="Eocene" pageId="1" pageNumber="343" stateProvince="Sjaelland">Eocene</location>
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, with label “NHMD-624757 ex-DK 176 (1);
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<specimenCode id="DBEB9EEFFFACE904839EF846FD81F816" box="[451,630,1924,1951]" collectionCode="ZMUC" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="1" pageNumber="343">ZMUC 902245</specimenCode>
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<collectingMunicipality id="6B96ACEEFFACE90480D9F846FCC0F816" box="[644,823,1924,1951]" pageId="1" pageNumber="343">Musholm Bugt</collectingMunicipality>
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,
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<collectingCounty id="62934E18FFACE9048117F846FC42F817" box="[842,949,1924,1950]" pageId="1" pageNumber="343">Storebaelt</collectingCounty>
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<collectingCountry id="F35A7604FFACE904819EF846FBD9F817" box="[963,1070,1924,1950]" name="Denmark" pageId="1" pageNumber="343">Denmark</collectingCountry>
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,
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<collectingRegion id="4989F876FFACE904861CF846FAF2F817" box="[1089,1285,1924,1950]" country="Denmark" name="Sjaelland" pageId="1" pageNumber="343">Region Sjaelland</collectingRegion>
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, Kalundborg Kommune. Leg. Jørgensen, Benny.
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</materialsCitation>
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</paragraph>
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<paragraph id="8BF23694FFACE904829AF80FFB72F86F" blockId="1.[151,1437,1816,2023]" box="[199,1157,1996,2023]" pageId="1" pageNumber="343">
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<materialsCitation id="3B253CC9FFACE904829AF80FFB8DF86F" box="[199,1146,1996,2023]" collectingDate="1966-06-07" country="Denmark" location="DENMARK" pageId="1" pageNumber="343" specimenCount="1">
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Det: Andersen, Nils,
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<date id="FFF31054FFACE90483EEF80EFDC5F86F" box="[435,562,1996,2022]" pageId="1" pageNumber="343" value="1966-06-07">
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<collectingDate id="EFB7E9BCFFACE90483EEF80EFDC5F86F" box="[435,562,1996,2022]" pageId="1" pageNumber="343" value="1966-06-07">07/06/1966</collectingDate>
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</date>
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NATURAL HISTORY MUSEUM OF
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<collectingCountry id="F35A7604FFACE90481B1F80EFB8DF86F" box="[1004,1146,1996,2022]" name="Denmark" pageId="1" pageNumber="343">DENMARK</collectingCountry>
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</materialsCitation>
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”
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</paragraph>
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</subSubSection>
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<caption id="DF32661CFFAFE90782C5F94DFB49F942" ID-DOI="http://doi.org/10.5281/zenodo.10468856" ID-Zenodo-Dep="10468856" httpUri="https://zenodo.org/record/10468856/files/figure.png" pageId="2" pageNumber="344" startId="2.[152,255,1679,1703]" targetBox="[159,1428,189,1649]" targetPageId="2" targetType="figure">
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<paragraph id="8BF23694FFAFE90782C5F94DFB49F942" blockId="2.[151,1436,1679,1739]" pageId="2" pageNumber="344">
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||
<emphasis id="B939EA86FFAFE90782C5F94DFEE5F92E" bold="true" box="[152,274,1679,1703]" pageId="2" pageNumber="344">FIGURE 1</emphasis>
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||
.
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||
<taxonomicName id="4C4D4D17FFAFE9078340F94DFD20F92E" authority="Anisyutkin et Perkovsky" authorityName="Anisyutkin et Perkovsky" authorityYear="2024" box="[285,727,1679,1703]" class="Insecta" family="Ectobiidae" genus="Ectobius" kingdom="Animalia" order="Blattodea" pageId="2" pageNumber="344" phylum="Arthropoda" rank="species" species="danekrae" status="sp. nov.">
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||
<emphasis id="B939EA86FFAFE9078340F94DFE2EF92E" box="[285,473,1679,1703]" italics="true" pageId="2" pageNumber="344">Ectobius danekrae</emphasis>
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||
Anisyutkin et Perkovsky
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</taxonomicName>
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||
,
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||
<emphasis id="B939EA86FFAFE90780BDF94DFCC7F92E" bold="true" box="[736,816,1679,1703]" pageId="2" pageNumber="344">
|
||
<taxonomicNameLabel id="A20A57FDFFAFE90780BDF94DFCC7F92E" box="[736,816,1679,1703]" pageId="2" pageNumber="344" rank="species">sp. nov.</taxonomicNameLabel>
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||
</emphasis>
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||
, male holotype. A, B—general view of inclusion: from above (A) and below (B); C, D—general view from above (C) and below (D); E—abdominal apex from below.
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||
</paragraph>
|
||
</caption>
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||
<subSubSection id="C357651FFFAFE901829AF93BFCDEF83D" lastPageId="4" lastPageNumber="346" pageId="2" pageNumber="344" type="description">
|
||
<paragraph id="8BF23694FFAFE907829AF93BFB91F8D2" blockId="2.[151,1437,1785,2028]" pageId="2" pageNumber="344">
|
||
<emphasis id="B939EA86FFAFE907829AF93BFE10F89A" bold="true" box="[199,487,1785,1811]" pageId="2" pageNumber="344">Description of inclusion</emphasis>
|
||
(
|
||
<figureCitation id="13762A11FFAFE90783A4F93BFD80F89A" box="[505,631,1785,1811]" captionStart="FIGURE 1" captionStartId="2.[152,255,1679,1703]" captionTargetBox="[159,1428,189,1649]" captionTargetId="figure-21@2.[151,1436,179,1653]" captionTargetPageId="2" captionText="FIGURE 1. Ectobius danekrae Anisyutkin et Perkovsky, sp. nov., male holotype.A, B—general view of inclusion: from above (A) and below (B); C, D—general view from above (C) and below (D); E—abdominal apex from below." figureDoi="http://doi.org/10.5281/zenodo.10468856" httpUri="https://zenodo.org/record/10468856/files/figure.png" pageId="2" pageNumber="344">Fig. 1A, B</figureCitation>
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||
). Cylindrical, very clear yellow amber piece, brought to light by sand pumping in Storebaelt (Bonde
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<emphasis id="B939EA86FFAFE90783A5F8DCFDD8F8BE" box="[504,559,1821,1847]" italics="true" pageId="2" pageNumber="344">at al</emphasis>
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||
. 2008). The sample is oblong,
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||
<quantity id="4CB59B71FFAFE90781C2F8DFFC06F8B1" box="[927,1009,1821,1848]" metricMagnitude="-2" metricUnit="m" metricValue="3.7" pageId="2" pageNumber="344" unit="mm" value="37.0">37 mm</quantity>
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long. The amber contains an almost completely preserved adult male cockroach with mouthparts, wings, legs and genitalia.
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</paragraph>
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<paragraph id="8BF23694FFAFE907829AF8A7FA98F8F6" blockId="2.[151,1437,1785,2028]" box="[199,1391,1893,1920]" pageId="2" pageNumber="344">
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||
Syninclusion:
|
||
<taxonomicName id="4C4D4D17FFAFE9078335F8A7FE16F8F6" box="[360,481,1893,1919]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="344" phylum="Arthropoda" rank="order">Hemiptera</taxonomicName>
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,
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||
<taxonomicName id="4C4D4D17FFAFE90783B1F8A7FD5AF8F6" box="[492,685,1893,1919]" class="Insecta" family="Drepanosiphidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="344" phylum="Arthropoda" rank="family">Drepanosiphidae</taxonomicName>
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||
,
|
||
<taxonomicName id="4C4D4D17FFAFE90780E5F8A7FB70F8F6" authority="Heie, 1967" authorityName="Heie" authorityYear="1967" box="[696,1159,1893,1919]" class="Insecta" family="Aphididae" genus="Palaeophyllaphis" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="344" phylum="Arthropoda" rank="species" species="longirostris">
|
||
<emphasis id="B939EA86FFAFE90780E5F8A7FBF2F8F6" box="[696,1029,1893,1919]" italics="true" pageId="2" pageNumber="344">Palaeophyllaphis longirostris</emphasis>
|
||
<bibRefCitation id="EFDC4B65FFAFE9078651F8A7FB70F8F6" author="Heie, O. E." box="[1036,1159,1893,1919]" pageId="2" pageNumber="344" pagination="1 - 274" refId="ref4823" refString="Heie, O. E. (1967) Studies on fossil aphids (Homoptera: Aphidoidea), especially in the Copenhagen collection of fossils in Baltic amber. Spolia zoologica Musei Hauniensis, 26, 1 - 274." type="journal article" year="1967">Heie, 1967</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFDC4B65FFAFE90786C8F8A7FA94F8F6" author="Bonde, N. C. & Hald, N. & Andersen, S. & Jakobsen, S. L." box="[1173,1379,1893,1920]" pageId="2" pageNumber="344" refId="ref4449" refString="Bonde, N. C., Hald, N., Andersen, S. & Jakobsen, S. L. (2008) Danekrae: de bedste danske fossiler. Gyldendal, Copenhagen, 226 p." type="book" year="2008">
|
||
Bonde
|
||
<emphasis id="B939EA86FFAFE90786BAF8A4FAEFF8F6" box="[1255,1304,1893,1919]" italics="true" pageId="2" pageNumber="344">et al</emphasis>
|
||
. 2008
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF23694FFAFE906829AF84BFD6CFE94" blockId="2.[151,1437,1785,2028]" lastBlockId="3.[151,1437,151,2013]" lastPageId="3" lastPageNumber="345" pageId="2" pageNumber="344">
|
||
<emphasis id="B939EA86FFAFE907829AF84BFEA6F82A" bold="true" box="[199,337,1929,1955]" pageId="2" pageNumber="344">Description</emphasis>
|
||
. General color, as far as can be judged from the cockroach inclusion, light reddish (yellowish?) (
|
||
<figureCitation id="13762A11FFAFE90782C2F86FFED3F84E" box="[159,292,1965,1991]" captionStart="FIGURE 1" captionStartId="2.[152,255,1679,1703]" captionTargetBox="[159,1428,189,1649]" captionTargetId="figure-21@2.[151,1436,179,1653]" captionTargetPageId="2" captionText="FIGURE 1. Ectobius danekrae Anisyutkin et Perkovsky, sp. nov., male holotype.A, B—general view of inclusion: from above (A) and below (B); C, D—general view from above (C) and below (D); E—abdominal apex from below." figureDoi="http://doi.org/10.5281/zenodo.10468856" httpUri="https://zenodo.org/record/10468856/files/figure.png" pageId="2" pageNumber="344">Figs. 1C–E</figureCitation>
|
||
); facial part of head with two blackish spots: larger spot between eyes and smaller one between and below antennal sockets; antennae at least partly blackish; maxillary palps, especially apical segment, probably darker as compared with general color; disk of pronotum with large hexagon-like black spot; lateral parts of pronotum and costal field of tegmina lighter, yellowish; tegmina with darker space between veins; legs (with exception of blackish spots at distal ends of femora and proximal ends of tibiae), abdomen from below (with exception of slightly darkened median part) and cerci light reddish.
|
||
</paragraph>
|
||
<paragraph id="8BF23694FFAEE906829AFEE5FB06FEC8" blockId="3.[151,1437,151,2013]" box="[199,1265,295,321]" pageId="3" pageNumber="345">Dense pubescence visible on antennae, tarsi and cerci. Surfaces smooth, punctuation indistinct.</paragraph>
|
||
<paragraph id="8BF23694FFAEE906829AFE89FAB4FAB8" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="345">
|
||
Head protruding weakly from under pronotum, subtriangular in shape (
|
||
<figureCitation id="13762A11FFAEE9068183FE89FB95FEEC" box="[990,1122,331,357]" captionStart="FIGURE 1" captionStartId="2.[152,255,1679,1703]" captionTargetBox="[159,1428,189,1649]" captionTargetId="figure-21@2.[151,1436,179,1653]" captionTargetPageId="2" captionText="FIGURE 1. Ectobius danekrae Anisyutkin et Perkovsky, sp. nov., male holotype.A, B—general view of inclusion: from above (A) and below (B); C, D—general view from above (C) and below (D); E—abdominal apex from below." figureDoi="http://doi.org/10.5281/zenodo.10468856" httpUri="https://zenodo.org/record/10468856/files/figure.png" pageId="3" pageNumber="345">Figs. 1C, D</figureCitation>
|
||
,
|
||
<figureCitation id="13762A11FFAEE9068631FE89FB65FEEC" box="[1132,1170,331,357]" captionStart="FIGURE 2" captionStartId="4.[152,255,1443,1467]" captionTargetBox="[159,1427,189,1412]" captionTargetId="figure-21@4.[151,1435,181,1420]" captionTargetPageId="4" captionText="FIGURE 2. Ectobius danekrae Anisyutkin et Perkovsky, sp. nov., male holotype. A—outlines of body from above; B— abdominal apex and part of leg from below." figureDoi="http://doi.org/10.5281/zenodo.10468862" httpUri="https://zenodo.org/record/10468862/files/figure.png" pageId="3" pageNumber="345">2A</figureCitation>
|
||
); eyes large, interocular space wide; antennae long (
|
||
<figureCitation id="13762A11FFAEE9068392FEADFDA3FE00" box="[463,596,367,393]" captionStart="FIGURE 1" captionStartId="2.[152,255,1679,1703]" captionTargetBox="[159,1428,189,1649]" captionTargetId="figure-21@2.[151,1436,179,1653]" captionTargetPageId="2" captionText="FIGURE 1. Ectobius danekrae Anisyutkin et Perkovsky, sp. nov., male holotype.A, B—general view of inclusion: from above (A) and below (B); C, D—general view from above (C) and below (D); E—abdominal apex from below." figureDoi="http://doi.org/10.5281/zenodo.10468856" httpUri="https://zenodo.org/record/10468856/files/figure.png" pageId="3" pageNumber="345">Figs. 1C, D</figureCitation>
|
||
,
|
||
<figureCitation id="13762A11FFAEE906803DFEADFD72FE00" box="[608,645,367,393]" captionStart="FIGURE 2" captionStartId="4.[152,255,1443,1467]" captionTargetBox="[159,1427,189,1412]" captionTargetId="figure-21@4.[151,1435,181,1420]" captionTargetPageId="4" captionText="FIGURE 2. Ectobius danekrae Anisyutkin et Perkovsky, sp. nov., male holotype. A—outlines of body from above; B— abdominal apex and part of leg from below." figureDoi="http://doi.org/10.5281/zenodo.10468862" httpUri="https://zenodo.org/record/10468862/files/figure.png" pageId="3" pageNumber="345">2A</figureCitation>
|
||
), with more than 40 antennomeres, about as long as or slightly shorter then body length; scapus largest and widest; pedicel short, conical in shape, thinner than scapus and wider than segments of flagellum; 1st flagellomere elongated, more than twice as long as wide, succeeding ~ 10 antennomeres short and transverse, subsequent segments gradually become longer, about twice as long as wide. Maxillary palps long, 3rd segment elongated, 4th elongated conical and 5th – subtriangular in shape; 3rd and 4th segments probably subequal in length, 5 – largest (unfortunately, it is impossible to accurately measure the size of maxillary palps segments on inclusion). Pronotum transverse (
|
||
<figureCitation id="13762A11FFAEE90680F5FD85FCFBFDE8" box="[680,780,583,609]" captionStart="FIGURE 1" captionStartId="2.[152,255,1679,1703]" captionTargetBox="[159,1428,189,1649]" captionTargetId="figure-21@2.[151,1436,179,1653]" captionTargetPageId="2" captionText="FIGURE 1. Ectobius danekrae Anisyutkin et Perkovsky, sp. nov., male holotype.A, B—general view of inclusion: from above (A) and below (B); C, D—general view from above (C) and below (D); E—abdominal apex from below." figureDoi="http://doi.org/10.5281/zenodo.10468856" httpUri="https://zenodo.org/record/10468856/files/figure.png" pageId="3" pageNumber="345">Figs. 1C</figureCitation>
|
||
,
|
||
<figureCitation id="13762A11FFAEE9068147FD85FCB7FDE8" box="[794,832,583,609]" captionStart="FIGURE 2" captionStartId="4.[152,255,1443,1467]" captionTargetBox="[159,1427,189,1412]" captionTargetId="figure-21@4.[151,1435,181,1420]" captionTargetPageId="4" captionText="FIGURE 2. Ectobius danekrae Anisyutkin et Perkovsky, sp. nov., male holotype. A—outlines of body from above; B— abdominal apex and part of leg from below." figureDoi="http://doi.org/10.5281/zenodo.10468862" httpUri="https://zenodo.org/record/10468862/files/figure.png" pageId="3" pageNumber="345">2A</figureCitation>
|
||
), gradually widened caudally, anterior margin nearly straight, lateral – widely rounded, posterior – weakly protruded caudally. Scutellum distinct, in shape of isosceles triangle (
|
||
<figureCitation id="13762A11FFAEE906835DFD4DFE92FD20" box="[256,357,655,681]" captionStart="FIGURE 1" captionStartId="2.[152,255,1679,1703]" captionTargetBox="[159,1428,189,1649]" captionTargetId="figure-21@2.[151,1436,179,1653]" captionTargetPageId="2" captionText="FIGURE 1. Ectobius danekrae Anisyutkin et Perkovsky, sp. nov., male holotype.A, B—general view of inclusion: from above (A) and below (B); C, D—general view from above (C) and below (D); E—abdominal apex from below." figureDoi="http://doi.org/10.5281/zenodo.10468856" httpUri="https://zenodo.org/record/10468856/files/figure.png" pageId="3" pageNumber="345">Figs. 1C</figureCitation>
|
||
,
|
||
<figureCitation id="13762A11FFAEE9068328FD4DFE6CFD20" box="[373,411,655,681]" captionStart="FIGURE 2" captionStartId="4.[152,255,1443,1467]" captionTargetBox="[159,1427,189,1412]" captionTargetId="figure-21@4.[151,1435,181,1420]" captionTargetPageId="4" captionText="FIGURE 2. Ectobius danekrae Anisyutkin et Perkovsky, sp. nov., male holotype. A—outlines of body from above; B— abdominal apex and part of leg from below." figureDoi="http://doi.org/10.5281/zenodo.10468862" httpUri="https://zenodo.org/record/10468862/files/figure.png" pageId="3" pageNumber="345">2A</figureCitation>
|
||
). Tegmina and wings fully developed (
|
||
<figureCitation id="13762A11FFAEE9068138FD4DFC3DFD20" box="[869,970,655,681]" captionStart="FIGURE 1" captionStartId="2.[152,255,1679,1703]" captionTargetBox="[159,1428,189,1649]" captionTargetId="figure-21@2.[151,1436,179,1653]" captionTargetPageId="2" captionText="FIGURE 1. Ectobius danekrae Anisyutkin et Perkovsky, sp. nov., male holotype.A, B—general view of inclusion: from above (A) and below (B); C, D—general view from above (C) and below (D); E—abdominal apex from below." figureDoi="http://doi.org/10.5281/zenodo.10468856" httpUri="https://zenodo.org/record/10468856/files/figure.png" pageId="3" pageNumber="345">Figs. 1C</figureCitation>
|
||
,
|
||
<figureCitation id="13762A11FFAEE9068186FD4DFBF7FD20" box="[987,1024,655,681]" captionStart="FIGURE 2" captionStartId="4.[152,255,1443,1467]" captionTargetBox="[159,1427,189,1412]" captionTargetId="figure-21@4.[151,1435,181,1420]" captionTargetPageId="4" captionText="FIGURE 2. Ectobius danekrae Anisyutkin et Perkovsky, sp. nov., male holotype. A—outlines of body from above; B— abdominal apex and part of leg from below." figureDoi="http://doi.org/10.5281/zenodo.10468862" httpUri="https://zenodo.org/record/10468862/files/figure.png" pageId="3" pageNumber="345">2A</figureCitation>
|
||
), surpassed abdominal apex (forma
|
||
<emphasis id="B939EA86FFAEE90682CAFD71FE97FD45" box="[151,352,691,716]" italics="true" pageId="3" pageNumber="345">macroptera sensu</emphasis>
|
||
<bibRefCitation id="EFDC4B65FFAEE906833AFD71FDDDFD44" author="Anisyutkin, L. N." box="[359,554,691,717]" pageId="3" pageNumber="345" pagination="664 - 674" refId="ref3951" refString="Anisyutkin, L. N. (2022) New data on the systematics and distribution of the cockroach genera Placoblatta Bey-Bienko, 1969 and Anisolampra Bey-Bienko, 1969 (Dictyoptera, Blaberidae: Epilamprinae). Entomological Review, 102 (5), 664 - 674. https: // doi. org / 10.1134 / S 0013873822050098" type="journal article" year="2022">Anisyutkin 2022</bibRefCitation>
|
||
). Tegmina elongated, moderately coriaceous, with distinct venation. Sc about as half as tegmina length, forked apically. Costal field strongly elongated and narrow. R roundly curved, with more than 20 regularly arranged anterior rami. M indiscernible. CuA fused with R in proximal part, R+CuA with more than 17 simple and regularly arranged posterior rami. CuP widely curved, but sharply curved at the point of confluence with the posterior margin of tegmen. Anal veins 7–8, these veins partly subobsolete, elongated parallel to posterior margin of tegmen. Structure of wings indistinguishable, but anal field large. Legs with well developed armament (
|
||
<figureCitation id="13762A11FFAEE9068347FC49FE53FC2C" box="[282,420,907,933]" captionStart="FIGURE 1" captionStartId="2.[152,255,1679,1703]" captionTargetBox="[159,1428,189,1649]" captionTargetId="figure-21@2.[151,1436,179,1653]" captionTargetPageId="2" captionText="FIGURE 1. Ectobius danekrae Anisyutkin et Perkovsky, sp. nov., male holotype.A, B—general view of inclusion: from above (A) and below (B); C, D—general view from above (C) and below (D); E—abdominal apex from below." figureDoi="http://doi.org/10.5281/zenodo.10468856" httpUri="https://zenodo.org/record/10468856/files/figure.png" pageId="3" pageNumber="345">Figs. 1D, E</figureCitation>
|
||
,
|
||
<figureCitation id="13762A11FFAEE90683EEFC49FE20FC2C" box="[435,471,907,933]" captionStart="FIGURE 2" captionStartId="4.[152,255,1443,1467]" captionTargetBox="[159,1427,189,1412]" captionTargetId="figure-21@4.[151,1435,181,1420]" captionTargetPageId="4" captionText="FIGURE 2. Ectobius danekrae Anisyutkin et Perkovsky, sp. nov., male holotype. A—outlines of body from above; B— abdominal apex and part of leg from below." figureDoi="http://doi.org/10.5281/zenodo.10468862" httpUri="https://zenodo.org/record/10468862/files/figure.png" pageId="3" pageNumber="345">2B</figureCitation>
|
||
). Anterior margin of fore femora armed probably according to
|
||
<typeStatus id="54F68836FFAEE90686E1FC49FB1BFC2C" box="[1212,1260,907,933]" pageId="3" pageNumber="345">type</typeStatus>
|
||
B (
|
||
<emphasis id="B939EA86FFAEE9068740FC49FAACFC2D" box="[1309,1371,907,932]" italics="true" pageId="3" pageNumber="345">sensu</emphasis>
|
||
<bibRefCitation id="EFDC4B65FFAEE906873BFC49FEDDFC40" author="Bey-Bienko, G. Ya." pageId="3" pageNumber="345" refId="ref4280" refString="Bey-Bienko, G. Ya. (1950) Nasekomye Tarakanovye [Cockroach insects]. Fauna of the USSR. New Series 40. Nauka Publishers, Leningrad, Moscow, 343 pp. [in Russian]" type="book" year="1950">Bey-Bienko 1950</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFDC4B65FFAEE906836BFC6DFE47FC40" author="Roth, L. M." box="[310,432,943,969]" pageId="3" pageNumber="345" pagination="1 - 186" refId="ref5926" refString="Roth, L. M. (2003) Systematics and phylogeny of cockroaches (Dictyoptera: Blattaria). Oriental Insects, 37, 1 - 186. http: // doi. org / 10.1080 / 00305316.2003.10417344" type="journal article" year="2003">Roth 2003</bibRefCitation>
|
||
), with at least one large apical spine (
|
||
<figureCitation id="13762A11FFAEE906810BFC6DFC58FC40" box="[854,943,943,969]" captionStart="FIGURE 1" captionStartId="2.[152,255,1679,1703]" captionTargetBox="[159,1428,189,1649]" captionTargetId="figure-21@2.[151,1436,179,1653]" captionTargetPageId="2" captionText="FIGURE 1. Ectobius danekrae Anisyutkin et Perkovsky, sp. nov., male holotype.A, B—general view of inclusion: from above (A) and below (B); C, D—general view from above (C) and below (D); E—abdominal apex from below." figureDoi="http://doi.org/10.5281/zenodo.10468856" httpUri="https://zenodo.org/record/10468856/files/figure.png" pageId="3" pageNumber="345">Fig. 1D</figureCitation>
|
||
). Fore tibia not thickened distally. Structure of hind tarsi (
|
||
<figureCitation id="13762A11FFAEE9068372FC11FE7FFC64" box="[303,392,979,1005]" captionStart="FIGURE 1" captionStartId="2.[152,255,1679,1703]" captionTargetBox="[159,1428,189,1649]" captionTargetId="figure-21@2.[151,1436,179,1653]" captionTargetPageId="2" captionText="FIGURE 1. Ectobius danekrae Anisyutkin et Perkovsky, sp. nov., male holotype.A, B—general view of inclusion: from above (A) and below (B); C, D—general view from above (C) and below (D); E—abdominal apex from below." figureDoi="http://doi.org/10.5281/zenodo.10468856" httpUri="https://zenodo.org/record/10468856/files/figure.png" pageId="3" pageNumber="345">Fig. 1D</figureCitation>
|
||
): metatarsus slightly longer than other segments combined; 2nd and 5th segments elongated, subequal in length, 3rd segment slightly shorter than 2nd; 4th segment – very short; all segments with distinct apical pulvillae; metatarsus and probably second segment with two rows (rows poorly visible on inclusion) of spines along lower margin, other tarsal segments (as far as can be judged from the inclusion) without spines; claws weakly asymmetrical or symmetrical; arolium about half of claw length. Fore and mid tarsi similar to hind tarsi, but segments comparatively shorter. Abdominal sternites without visible specializations (
|
||
<figureCitation id="13762A11FFAEE9068638FB45FB3DFB28" box="[1125,1226,1159,1185]" captionStart="FIGURE 1" captionStartId="2.[152,255,1679,1703]" captionTargetBox="[159,1428,189,1649]" captionTargetId="figure-21@2.[151,1436,179,1653]" captionTargetPageId="2" captionText="FIGURE 1. Ectobius danekrae Anisyutkin et Perkovsky, sp. nov., male holotype.A, B—general view of inclusion: from above (A) and below (B); C, D—general view from above (C) and below (D); E—abdominal apex from below." figureDoi="http://doi.org/10.5281/zenodo.10468856" httpUri="https://zenodo.org/record/10468856/files/figure.png" pageId="3" pageNumber="345">Figs. 1D</figureCitation>
|
||
,
|
||
<figureCitation id="13762A11FFAEE9068684FB45FB0AFB28" box="[1241,1277,1159,1185]" captionStart="FIGURE 2" captionStartId="4.[152,255,1443,1467]" captionTargetBox="[159,1427,189,1412]" captionTargetId="figure-21@4.[151,1435,181,1420]" captionTargetPageId="4" captionText="FIGURE 2. Ectobius danekrae Anisyutkin et Perkovsky, sp. nov., male holotype. A—outlines of body from above; B— abdominal apex and part of leg from below." figureDoi="http://doi.org/10.5281/zenodo.10468862" httpUri="https://zenodo.org/record/10468862/files/figure.png" pageId="3" pageNumber="345">2B</figureCitation>
|
||
). Cercus long and flattened, about twice as long as hypandrium, with more than 7 distinct segments (
|
||
<figureCitation id="13762A11FFAEE9068634FB69FB3EFB4C" box="[1129,1225,1195,1221]" captionStart="FIGURE 1" captionStartId="2.[152,255,1679,1703]" captionTargetBox="[159,1428,189,1649]" captionTargetId="figure-21@2.[151,1436,179,1653]" captionTargetPageId="2" captionText="FIGURE 1. Ectobius danekrae Anisyutkin et Perkovsky, sp. nov., male holotype.A, B—general view of inclusion: from above (A) and below (B); C, D—general view from above (C) and below (D); E—abdominal apex from below." figureDoi="http://doi.org/10.5281/zenodo.10468856" httpUri="https://zenodo.org/record/10468856/files/figure.png" pageId="3" pageNumber="345">Figs. 1E</figureCitation>
|
||
,
|
||
<figureCitation id="13762A11FFAEE906868BFB69FB0DFB4C" box="[1238,1274,1195,1221]" captionStart="FIGURE 2" captionStartId="4.[152,255,1443,1467]" captionTargetBox="[159,1427,189,1412]" captionTargetId="figure-21@4.[151,1435,181,1420]" captionTargetPageId="4" captionText="FIGURE 2. Ectobius danekrae Anisyutkin et Perkovsky, sp. nov., male holotype. A—outlines of body from above; B— abdominal apex and part of leg from below." figureDoi="http://doi.org/10.5281/zenodo.10468862" httpUri="https://zenodo.org/record/10468862/files/figure.png" pageId="3" pageNumber="345">2B</figureCitation>
|
||
). Hypandrium elongated (
|
||
<figureCitation id="13762A11FFAEE906834BFB0DFE8EFB60" box="[278,377,1231,1257]" captionStart="FIGURE 1" captionStartId="2.[152,255,1679,1703]" captionTargetBox="[159,1428,189,1649]" captionTargetId="figure-21@2.[151,1436,179,1653]" captionTargetPageId="2" captionText="FIGURE 1. Ectobius danekrae Anisyutkin et Perkovsky, sp. nov., male holotype.A, B—general view of inclusion: from above (A) and below (B); C, D—general view from above (C) and below (D); E—abdominal apex from below." figureDoi="http://doi.org/10.5281/zenodo.10468856" httpUri="https://zenodo.org/record/10468856/files/figure.png" pageId="3" pageNumber="345">Figs. 1E</figureCitation>
|
||
,
|
||
<figureCitation id="13762A11FFAEE90683D4FB0DFE5BFB60" box="[393,428,1231,1257]" captionStart="FIGURE 2" captionStartId="4.[152,255,1443,1467]" captionTargetBox="[159,1427,189,1412]" captionTargetId="figure-21@4.[151,1435,181,1420]" captionTargetPageId="4" captionText="FIGURE 2. Ectobius danekrae Anisyutkin et Perkovsky, sp. nov., male holotype. A—outlines of body from above; B— abdominal apex and part of leg from below." figureDoi="http://doi.org/10.5281/zenodo.10468862" httpUri="https://zenodo.org/record/10468862/files/figure.png" pageId="3" pageNumber="345">2B</figureCitation>
|
||
), subtriangular in shape, slightly asymmetrical, left caudolateral side seems to be more rounded than right one; left stylus large, evidently subcylindrical, outgrowth slightly narrowed towards the apex, with several chaetae or sensillae at apex; right stylus short, in shape of transverse asymmetrical outgrowth.
|
||
</paragraph>
|
||
<paragraph id="8BF23694FFAEE906829AFAF9FDB3FA48" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="345">Measurements (mm). Total body length 16.1* (as preserved, from vertex to tegmen apex); head length 1.7*; head width 1.5*; pronotum length 1.5*; pronotum width 2.5*; tegmen length <5.5; tegmen width 2.2*; costal field length 3; anal field length 2.5; anal field width 1.2. Dimensions marked with * are approximate due to inclusion distortion and cockroach deformation.</paragraph>
|
||
<paragraph id="8BF23694FFAEE906829AFA09FE66FA6C" blockId="3.[151,1437,151,2013]" box="[199,401,1483,1509]" pageId="3" pageNumber="345">Female unknown.</paragraph>
|
||
<paragraph id="8BF23694FFAEE906829AFA2CFE68F8C4" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="345">
|
||
<emphasis id="B939EA86FFAEE906829AFA2CFEACF981" bold="true" box="[199,347,1518,1544]" pageId="3" pageNumber="345">Comparison</emphasis>
|
||
. The new species readily differs from the two known from Baltic amber species (
|
||
<taxonomicName id="4C4D4D17FFAEE906877CFA2DFA6AF980" baseAuthorityName="Germar & Berendt" baseAuthorityYear="1856" box="[1313,1437,1519,1545]" class="Insecta" family="Ectobiidae" genus="Ectobius" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="345" phylum="Arthropoda" rank="species" species="balticus">
|
||
<emphasis id="B939EA86FFAEE906877CFA2DFAC5F981" box="[1313,1330,1519,1544]" italics="true" pageId="3" pageNumber="345">E</emphasis>
|
||
.
|
||
<emphasis id="B939EA86FFAEE9068719FA2DFA6AF980" box="[1348,1437,1519,1545]" italics="true" pageId="3" pageNumber="345">balticus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4D4D17FFAEE9068294F9D1FEB1F9A4" authorityName="Shelford" authorityYear="1910" box="[201,326,1555,1581]" class="Insecta" family="Ectobiidae" genus="Ectobius" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="345" phylum="Arthropoda" rank="species" species="inclusus">
|
||
<emphasis id="B939EA86FFAEE9068294F9D1FF2DF9A5" box="[201,218,1555,1580]" italics="true" pageId="3" pageNumber="345">E</emphasis>
|
||
.
|
||
<emphasis id="B939EA86FFAEE90682B7F9D1FEB1F9A4" box="[234,326,1555,1581]" italics="true" pageId="3" pageNumber="345">inclusus</emphasis>
|
||
</taxonomicName>
|
||
) and extant species of the genus by presence of small right stylus. Additionally,
|
||
<taxonomicName id="4C4D4D17FFAEE90686B5F9D1FA86F9A4" authority="Anisyutkin & Vasilenko & Perkovsky, 2024" authorityName="Anisyutkin & Vasilenko & Perkovsky" authorityYear="2024" box="[1256,1393,1555,1581]" class="Insecta" family="Ectobiidae" genus="Ectobius" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="345" phylum="Arthropoda" rank="species" species="danekrae" status="sp. nov.">
|
||
<emphasis id="B939EA86FFAEE90686B5F9D1FB0EF9A5" box="[1256,1273,1555,1580]" italics="true" pageId="3" pageNumber="345">E</emphasis>
|
||
.
|
||
<emphasis id="B939EA86FFAEE9068754F9D1FA86F9A4" box="[1289,1393,1555,1581]" italics="true" pageId="3" pageNumber="345">danekrae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B939EA86FFAEE9068726F9D1FF3FF9D8" bold="true" pageId="3" pageNumber="345">
|
||
<taxonomicNameLabel id="A20A57FDFFAEE9068726F9D1FF3FF9D8" pageId="3" pageNumber="345" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="4C4D4D17FFAEE906833CF9F5FE2CF9D8" baseAuthorityName="Germar & Berendt" baseAuthorityYear="1856" box="[353,475,1591,1617]" class="Insecta" family="Ectobiidae" genus="Ectobius" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="345" phylum="Arthropoda" rank="species" species="balticus">
|
||
<emphasis id="B939EA86FFAEE906833CF9F5FE85F9D9" box="[353,370,1591,1616]" italics="true" pageId="3" pageNumber="345">E</emphasis>
|
||
.
|
||
<emphasis id="B939EA86FFAEE90683DFF9F5FE2CF9D8" box="[386,475,1591,1617]" italics="true" pageId="3" pageNumber="345">balticus</emphasis>
|
||
</taxonomicName>
|
||
in comparatively light colored abdomen (dark, nearly black in
|
||
<taxonomicName id="4C4D4D17FFAEE90686F3F9F5FADFF9D8" baseAuthorityName="Germar & Berendt" baseAuthorityYear="1856" box="[1198,1320,1591,1617]" class="Insecta" family="Ectobiidae" genus="Ectobius" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="345" phylum="Arthropoda" rank="species" species="balticus">
|
||
<emphasis id="B939EA86FFAEE90686F3F9F5FB48F9D9" box="[1198,1215,1591,1616]" italics="true" pageId="3" pageNumber="345">E</emphasis>
|
||
.
|
||
<emphasis id="B939EA86FFAEE9068692F9F5FADFF9D8" box="[1231,1320,1591,1617]" italics="true" pageId="3" pageNumber="345">balticus</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<bibRefCitation id="EFDC4B65FFAEE906871AF9F5FEA8F9FC" author="Germar, E. F. & Berendt, G. C." pageId="3" pageNumber="345" pagination="1 - 125" refId="ref4664" refString="Germar, E. F. & Berendt, G. C. (1856) Die im Bernstein befindlichen Hemipteren und Orthopteren der Vorwelt. In: Berendt, G. C. (1845 - 1856) Die im Bernstein befindlichen organischen Reste der Vorwelt. Bd. 2. Abth. I. Berlin, pp. 1 - 125. https: // doi. org / 10.5962 / bhl. title. 51864" type="book chapter" year="1856">Germar and Berendt 1856</bibRefCitation>
|
||
and data of authors) and from
|
||
<taxonomicName id="4C4D4D17FFAEE90680E0F999FCCEF9FC" authorityName="Shelford" authorityYear="1910" box="[701,825,1627,1653]" class="Insecta" family="Ectobiidae" genus="Ectobius" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="345" phylum="Arthropoda" rank="species" species="inclusus">
|
||
<emphasis id="B939EA86FFAEE90680E0F999FD39F9FD" box="[701,718,1627,1652]" italics="true" pageId="3" pageNumber="345">E</emphasis>
|
||
.
|
||
<emphasis id="B939EA86FFAEE9068080F999FCCEF9FC" box="[733,825,1627,1653]" italics="true" pageId="3" pageNumber="345">inclusus</emphasis>
|
||
</taxonomicName>
|
||
by comparatively short left stylus (left stylus strongly elongated in
|
||
<taxonomicName id="4C4D4D17FFAEE9068376F9BDFE50F910" authorityName="Shelford" authorityYear="1910" box="[299,423,1663,1689]" class="Insecta" family="Ectobiidae" genus="Ectobius" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="345" phylum="Arthropoda" rank="species" species="inclusus">
|
||
<emphasis id="B939EA86FFAEE9068376F9BDFECBF911" box="[299,316,1663,1688]" italics="true" pageId="3" pageNumber="345">E</emphasis>
|
||
.
|
||
<emphasis id="B939EA86FFAEE9068316F9BDFE50F910" box="[331,423,1663,1689]" italics="true" pageId="3" pageNumber="345">inclusus</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<bibRefCitation id="EFDC4B65FFAEE906839BF9BDFD9CF910" author="Shelford, R." box="[454,619,1663,1689]" pageId="3" pageNumber="345" pagination="336 - 355" refId="ref6078" refString="Shelford, R. (1910) On a collection of Blattidae preserved in amber, from Prussia. Zoological Journal of the Linnean Society, 30, 336 - 355. https: // doi. org / 10.1111 / j. 1096 - 642.1910. tb 02140. x" type="journal article" year="1910">Shelford 1910</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C4D4D17FFAEE90680DDF9BDFCA4F910" authority="Anisyutkin & Vasilenko & Perkovsky, 2024" authorityName="Anisyutkin & Vasilenko & Perkovsky" authorityYear="2024" box="[640,851,1663,1689]" class="Insecta" family="Ectobiidae" genus="Ectobius" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="345" phylum="Arthropoda" rank="species" species="danekrae" status="sp. nov.">
|
||
<emphasis id="B939EA86FFAEE90680DDF9BDFCA4F910" box="[640,851,1663,1689]" italics="true" pageId="3" pageNumber="345">Ectobius danekrae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B939EA86FFAEE9068106F9BDFC41F910" bold="true" box="[859,950,1663,1689]" pageId="3" pageNumber="345">
|
||
<taxonomicNameLabel id="A20A57FDFFAEE9068106F9BDFC41F910" box="[859,950,1663,1689]" pageId="3" pageNumber="345" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
readily differs from described from Green River Formation
|
||
<taxonomicName id="4C4D4D17FFAEE9068307F961FE4BF934" box="[346,444,1699,1725]" class="Insecta" family="Ectobiidae" genus="Ectobius" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="345" phylum="Arthropoda" rank="species" species="kohlsi">
|
||
<emphasis id="B939EA86FFAEE9068307F961FE9CF935" box="[346,363,1699,1724]" italics="true" pageId="3" pageNumber="345">E</emphasis>
|
||
.
|
||
<emphasis id="B939EA86FFAEE9068324F961FE4BF934" box="[377,444,1699,1725]" italics="true" pageId="3" pageNumber="345">kohlsi</emphasis>
|
||
</taxonomicName>
|
||
by uniformly coloured disc of pronotum and tegmina (pronotum and tegmina distinctly punctate in
|
||
<taxonomicName id="4C4D4D17FFAEE9068347F905FE8BF968" box="[282,380,1735,1761]" class="Insecta" family="Ectobiidae" genus="Ectobius" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="345" phylum="Arthropoda" rank="species" species="kohlsi">
|
||
<emphasis id="B939EA86FFAEE9068347F905FEDCF969" box="[282,299,1735,1760]" italics="true" pageId="3" pageNumber="345">E</emphasis>
|
||
.
|
||
<emphasis id="B939EA86FFAEE9068364F905FE8BF968" box="[313,380,1735,1761]" italics="true" pageId="3" pageNumber="345">kohlsi</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<bibRefCitation id="EFDC4B65FFAEE90683CAF905FD75F968" author="Vrsansky, P. & Oruzinsky, R. & Barna, P. & Vidlicka, L. & Labandeira, C." box="[407,642,1735,1761]" pageId="3" pageNumber="345" pagination="28 - 36" refId="ref6890" refString="Vrsansky, P., Oruzinsky, R., Barna, P., Vidlicka, L. & Labandeira, C. (2014) Native Ectobius (Blattaria: Ectobiidae) from the early Eocene Green River Formation of Colorado and its reintroduction to North America 49 million years later. Annals of the Entomological Society of America, 107 (1), 28 - 36. https: // doi. org / 10.1603 / an 13042" type="journal article" year="2014">
|
||
Vršanský
|
||
<emphasis id="B939EA86FFAEE906805BF905FDC0F968" box="[518,567,1735,1761]" italics="true" pageId="3" pageNumber="345">et al</emphasis>
|
||
. 2014
|
||
</bibRefCitation>
|
||
). It is impossible to reliably distinguish the new species from all extant species (not all species of the genus have known males and the structure of styli), but those and
|
||
<taxonomicName id="4C4D4D17FFAEE90686B4F929FA85F88C" authority="Anisyutkin & Vasilenko & Perkovsky, 2024" authorityName="Anisyutkin & Vasilenko & Perkovsky" authorityYear="2024" box="[1257,1394,1771,1797]" class="Insecta" family="Ectobiidae" genus="Ectobius" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="345" phylum="Arthropoda" rank="species" species="danekrae" status="sp. nov.">
|
||
<emphasis id="B939EA86FFAEE90686B4F929FB0DF88D" box="[1257,1274,1771,1796]" italics="true" pageId="3" pageNumber="345">E</emphasis>
|
||
.
|
||
<emphasis id="B939EA86FFAEE9068757F929FA85F88C" box="[1290,1394,1771,1797]" italics="true" pageId="3" pageNumber="345">danekrae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20A57FDFFAEE9068726F929FF3DF8A0" pageId="3" pageNumber="345" rank="species">
|
||
<emphasis id="B939EA86FFAEE9068726F929FA62F88C" bold="true" box="[1403,1429,1771,1797]" pageId="3" pageNumber="345">sp</emphasis>
|
||
.
|
||
<emphasis id="B939EA86FFAEE90682CAF8CDFF34F8A0" bold="true" box="[151,195,1807,1833]" pageId="3" pageNumber="345">nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
are separated by more than 36 million years (
|
||
<bibRefCitation id="EFDC4B65FFAEE906808CF8CDFC72F8A0" author="Iakovleva, A. I." box="[721,901,1807,1833]" pageId="3" pageNumber="345" pagination="32 - 48" refId="ref4951" refString="Iakovleva, A. I. (2017) Detalization of Eocene dinocyst zonation for Eastern Peritethys. Bulletin of the Moscow Society of Naturalists. Geological Series, 92 (2), 32 - 48. [in Russian, with English summary]" type="journal article" year="2017">Iakovleva 2017</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFDC4B65FFAEE90681CCF8CDFB74F8A0" author="Iakovleva, A. I. & Mychko, E. V. & Aleksandrova, G. N." box="[913,1155,1807,1833]" pageId="3" pageNumber="345" pagination="77 - 78" refId="ref4997" refString="Iakovleva, A. I., Mychko, E. V. & Aleksandrova, G. N. (2021) New dinocyst data from the " Primorsky " amber quarry (Kaliningrad Region): clarification of the deposits' age as the base for a revision of paleogeographic reconstructions of the Danish- Polish strait in the late Paleogene. In: Golubev, V. K. & Nazarov, V. N. (Eds.), Paleostrat- 2021. Annual Meeting of the Paleontological section of the Moscow Society of Naturalists and the Moscow Branch of the Russian Paleontological Society, Russian Academy of Sciences, (Moscow, January 25 - 26, 2021). Abstracts. A. A. Borisyak Paleontological Institute, Russian Academy of Sciences, Moscow, pp. 77 - 78. [in Russian]" type="book chapter" year="2021">
|
||
Iakovleva
|
||
<emphasis id="B939EA86FFAEE906865AF8CDFBCEF8A0" box="[1031,1081,1807,1833]" italics="true" pageId="3" pageNumber="345">et al</emphasis>
|
||
. 2021
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFDC4B65FFAEE90686D2F8CDFB3EF8A0" author="Iakovleva, A. I. & Aleksandrova, G. N. & Mychko, E. V." box="[1167,1225,1807,1833]" pageId="3" pageNumber="345" pagination="1 - 40" refId="ref5147" refString="Iakovleva, A. I., Aleksandrova, G. N. & Mychko, E. V. (2022) Late Eocene (Priabonian) dinoflagellate cysts from Primorsky quarry, southeast Baltic coast, Kaliningrad Oblast, Russia. Palynology, 46, 1 - 40. https: // doi. org / 10.1080 / 01916122.2021.1980743" type="journal article" year="2022">2022</bibRefCitation>
|
||
) and are too much to imply conspecificity.
|
||
</paragraph>
|
||
<paragraph id="8BF23694FFAEE906829AF895FEE8F854" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="345">
|
||
The new species is somewhat similar to
|
||
<taxonomicName id="4C4D4D17FFAEE90680DFF895FCECF8F8" authorityName="Linnaeus" authorityYear="1758" box="[642,795,1879,1905]" class="Insecta" family="Ectobiidae" genus="Ectobius" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="345" phylum="Arthropoda" rank="species" species="lapponicus">
|
||
<emphasis id="B939EA86FFAEE90680DFF895FD64F8F9" box="[642,659,1879,1904]" italics="true" pageId="3" pageNumber="345">E</emphasis>
|
||
.
|
||
<emphasis id="B939EA86FFAEE90680FDF895FCECF8F8" box="[672,795,1879,1905]" italics="true" pageId="3" pageNumber="345">lapponicus</emphasis>
|
||
</taxonomicName>
|
||
in pattern of coloration. But the presence of such a unique character as the presence of a right stylus, only slightly asymmetrical tarsal claws and small arolia does not allow it to be attributed to any of the known species groups, including
|
||
<taxonomicName id="4C4D4D17FFAEE906812FF85DFBA1F830" authorityName="Linnaeus" authorityYear="1758" box="[882,1110,1951,1977]" class="Insecta" family="Ectobiidae" genus="Ectobius" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="345" phylum="Arthropoda" rank="species" species="lapponicus">
|
||
<emphasis id="B939EA86FFAEE906812FF85DFBA1F830" box="[882,1110,1951,1977]" italics="true" pageId="3" pageNumber="345">Ectobius lapponicus</emphasis>
|
||
</taxonomicName>
|
||
species group (see
|
||
<bibRefCitation id="EFDC4B65FFAEE9068769F85DFEE4F854" author="Vrsansky, P. & Oruzinsky, R. & Barna, P. & Vidlicka, L. & Labandeira, C." pageId="3" pageNumber="345" pagination="28 - 36" refId="ref6890" refString="Vrsansky, P., Oruzinsky, R., Barna, P., Vidlicka, L. & Labandeira, C. (2014) Native Ectobius (Blattaria: Ectobiidae) from the early Eocene Green River Formation of Colorado and its reintroduction to North America 49 million years later. Annals of the Entomological Society of America, 107 (1), 28 - 36. https: // doi. org / 10.1603 / an 13042" type="journal article" year="2014">
|
||
Vršanský
|
||
<emphasis id="B939EA86FFAEE90682CAF801FF3FF854" box="[151,200,1987,2013]" italics="true" pageId="3" pageNumber="345">et al</emphasis>
|
||
. 2014
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF32661CFFA9E90182C5FA61FDA4FA69" ID-DOI="http://doi.org/10.5281/zenodo.10468862" ID-Zenodo-Dep="10468862" httpUri="https://zenodo.org/record/10468862/files/figure.png" pageId="4" pageNumber="346" startId="4.[152,255,1443,1467]" targetBox="[159,1427,189,1412]" targetPageId="4" targetType="figure">
|
||
<paragraph id="8BF23694FFA9E90182C5FA61FDA4FA69" blockId="4.[151,1436,1443,1504]" pageId="4" pageNumber="346">
|
||
<emphasis id="B939EA86FFA9E90182C5FA61FEE1FA35" bold="true" box="[152,278,1443,1468]" pageId="4" pageNumber="346">FIGURE 2</emphasis>
|
||
.
|
||
<taxonomicName id="4C4D4D17FFA9E901837BFA66FD05FA35" authority="Anisyutkin et Perkovsky" authorityName="Anisyutkin et Perkovsky" authorityYear="2024" box="[294,754,1444,1468]" class="Insecta" family="Ectobiidae" genus="Ectobius" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="346" phylum="Arthropoda" rank="species" species="danekrae" status="sp. nov.">
|
||
<emphasis id="B939EA86FFA9E901837BFA66FE10FA35" box="[294,487,1444,1468]" italics="true" pageId="4" pageNumber="346">Ectobius danekrae</emphasis>
|
||
Anisyutkin et Perkovsky
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B939EA86FFA9E901815DFA66FCA3FA35" bold="true" box="[768,852,1444,1468]" pageId="4" pageNumber="346">
|
||
<taxonomicNameLabel id="A20A57FDFFA9E901815DFA66FCA3FA35" box="[768,852,1444,1468]" pageId="4" pageNumber="346" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male holotype. A—outlines of body from above; B— abdominal apex and part of leg from below.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF23694FFA9E901829AF9F0FE39F955" blockId="4.[151,1437,1586,1972]" pageId="4" pageNumber="346">
|
||
<emphasis id="B939EA86FFA9E901829AF9F0FEFDF9C5" bold="true" box="[199,266,1586,1612]" pageId="4" pageNumber="346">Notes</emphasis>
|
||
. A variation in the coloration of pronotum of
|
||
<taxonomicName id="4C4D4D17FFA9E9018177F9F1FC51F9C5" baseAuthorityName="Germar & Berendt" baseAuthorityYear="1856" box="[810,934,1586,1612]" class="Insecta" family="Ectobiidae" genus="Ectobius" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="346" phylum="Arthropoda" rank="species" species="balticus">
|
||
<emphasis id="B939EA86FFA9E9018177F9F1FCCCF9C5" box="[810,827,1587,1612]" italics="true" pageId="4" pageNumber="346">E</emphasis>
|
||
.
|
||
<emphasis id="B939EA86FFA9E9018110F9F0FC51F9C5" box="[845,934,1586,1612]" italics="true" pageId="4" pageNumber="346">balticus</emphasis>
|
||
</taxonomicName>
|
||
was mentioned by
|
||
<bibRefCitation id="EFDC4B65FFA9E90186D0F9F0FAB1F9C5" author="Shelford, R." box="[1165,1350,1586,1612]" pageId="4" pageNumber="346" pagination="336 - 355" refId="ref6078" refString="Shelford, R. (1910) On a collection of Blattidae preserved in amber, from Prussia. Zoological Journal of the Linnean Society, 30, 336 - 355. https: // doi. org / 10.1111 / j. 1096 - 642.1910. tb 02140. x" type="journal article" year="1910">Shelford (1910)</bibRefCitation>
|
||
. Single known specimen of
|
||
<taxonomicName id="4C4D4D17FFA9E9018321F995FDF3F9F9" authority="Anisyutkin & Vasilenko & Perkovsky, 2024" authorityName="Anisyutkin & Vasilenko & Perkovsky" authorityYear="2024" box="[380,516,1622,1648]" class="Insecta" family="Ectobiidae" genus="Ectobius" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="346" phylum="Arthropoda" rank="species" species="danekrae" status="sp. nov.">
|
||
<emphasis id="B939EA86FFA9E9018321F995FE7AF9F9" box="[380,397,1623,1648]" italics="true" pageId="4" pageNumber="346">E</emphasis>
|
||
.
|
||
<emphasis id="B939EA86FFA9E90183C1F994FDF3F9F9" box="[412,516,1622,1648]" italics="true" pageId="4" pageNumber="346">danekrae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B939EA86FFA9E9018051F994FD91F9F9" bold="true" box="[524,614,1622,1648]" pageId="4" pageNumber="346">
|
||
<taxonomicNameLabel id="A20A57FDFFA9E9018051F994FD91F9F9" box="[524,614,1622,1648]" pageId="4" pageNumber="346" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is characterized by large dark spot on pronotum and comparatively light lower part of abdomen, as compared with fossil
|
||
<taxonomicName id="4C4D4D17FFA9E90180EEF9B9FCDCF91D" baseAuthorityName="Germar & Berendt" baseAuthorityYear="1856" box="[691,811,1658,1684]" class="Insecta" family="Ectobiidae" genus="Ectobius" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="346" phylum="Arthropoda" rank="species" species="balticus">
|
||
<emphasis id="B939EA86FFA9E90180EEF9B9FD33F91D" box="[691,708,1659,1684]" italics="true" pageId="4" pageNumber="346">E</emphasis>
|
||
.
|
||
<emphasis id="B939EA86FFA9E901808FF9B8FCDCF91D" box="[722,811,1658,1684]" italics="true" pageId="4" pageNumber="346">balticus</emphasis>
|
||
</taxonomicName>
|
||
and extant
|
||
<taxonomicName id="4C4D4D17FFA9E90181F0F9B9FB0AF91C" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[941,1277,1658,1685]" class="Insecta" family="Ectobiidae" genus="Ectobius" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="346" phylum="Arthropoda" rank="species" species="lapponicus">
|
||
<emphasis id="B939EA86FFA9E90181F0F9B9FC49F91D" box="[941,958,1659,1684]" italics="true" pageId="4" pageNumber="346">E</emphasis>
|
||
.
|
||
<emphasis id="B939EA86FFA9E9018191F9B8FBB0F91D" box="[972,1095,1658,1684]" italics="true" pageId="4" pageNumber="346">lapponicus</emphasis>
|
||
<bibRefCitation id="EFDC4B65FFA9E9018613F9B8FB0AF91C" author="Linnaeus, C." box="[1102,1277,1658,1685]" pageId="4" pageNumber="346" refId="ref5325" refString="Linnaeus, C. (1758) Systema natura per regna tria naturae secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Editio decima, reformata. Tomus I. Salvii, Holmiae, 824 pp. https: // doi. org / 10.5962 / bhl. title. 542" type="book" year="1758">Linnaeus, 1758</bibRefCitation>
|
||
</taxonomicName>
|
||
. The presence of a dark spot on pronotum and a light abdomen makes it unlikely that the specimen of
|
||
<taxonomicName id="4C4D4D17FFA9E90186CFF95DFAECF931" authority="Anisyutkin & Vasilenko & Perkovsky, 2024" authorityName="Anisyutkin & Vasilenko & Perkovsky" authorityYear="2024" box="[1170,1307,1694,1720]" class="Insecta" family="Ectobiidae" genus="Ectobius" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="346" phylum="Arthropoda" rank="species" species="danekrae" status="sp. nov.">
|
||
<emphasis id="B939EA86FFA9E90186CFF95DFB54F931" box="[1170,1187,1695,1720]" italics="true" pageId="4" pageNumber="346">E</emphasis>
|
||
.
|
||
<emphasis id="B939EA86FFA9E90186EEF95CFAECF931" box="[1203,1307,1694,1720]" italics="true" pageId="4" pageNumber="346">danekrae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B939EA86FFA9E9018779F95CFA77F931" bold="true" box="[1316,1408,1694,1720]" pageId="4" pageNumber="346">
|
||
<taxonomicNameLabel id="A20A57FDFFA9E9018779F95CFA77F931" box="[1316,1408,1694,1720]" pageId="4" pageNumber="346" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is represented by a light form.
|
||
</paragraph>
|
||
<paragraph id="8BF23694FFA9E901829AF924FDF3F8AD" blockId="4.[151,1437,1586,1972]" pageId="4" pageNumber="346">Additionally, species from the Baltic amber need revision, and it is highly probably that in Baltic amber more species are awaiting description.</paragraph>
|
||
<paragraph id="8BF23694FFA9E901829AF8ECFCDEF83D" blockId="4.[151,1437,1586,1972]" pageId="4" pageNumber="346">
|
||
All known to the authors representatives of the genus
|
||
<taxonomicName id="4C4D4D17FFA9E901816AF8ECFC6EF8C1" authorityName="Stephens" authorityYear="1835" box="[823,921,1838,1864]" class="Insecta" family="Ectobiidae" genus="Ectobius" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="346" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B939EA86FFA9E901816AF8ECFC6EF8C1" box="[823,921,1838,1864]" italics="true" pageId="4" pageNumber="346">Ectobius</emphasis>
|
||
</taxonomicName>
|
||
have only one left stylus (
|
||
<bibRefCitation id="EFDC4B65FFA9E9018690F8ECFAA7F8C1" author="Bohn, H." box="[1229,1360,1838,1864]" pageId="4" pageNumber="346" pagination="317 - 342" refId="ref4319" refString="Bohn, H. (1989) Revision of the sylvestris group of Ectobius Stephens in Europe (Blattaria: Blattellidae). Entomologica scandinavica, 20, 317 - 342. https: // doi. org / 10.1163 / 187631289 x 00357" type="journal article" year="1989">Bohn 1989</bibRefCitation>
|
||
). Bey-Bienko pointed out that “hypandrium with only one clear left stylus, right stylus absent or occasionally presented as an extremely small, poorly visible tubercle” (
|
||
<bibRefCitation id="EFDC4B65FFA9E90180C4F8B4FC92F818" author="Bey-Bienko, G. Ya." box="[665,869,1910,1937]" pageId="4" pageNumber="346" refId="ref4280" refString="Bey-Bienko, G. Ya. (1950) Nasekomye Tarakanovye [Cockroach insects]. Fauna of the USSR. New Series 40. Nauka Publishers, Leningrad, Moscow, 343 pp. [in Russian]" type="book" year="1950">Bey-Bienko 1950</bibRefCitation>
|
||
, p. 186). Unfortunately, Bey-Bienko not specified species with “extremely small, poorly visible” right stylus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |