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<document id="1DB69B6A3263DB8E334772EF7A9AA8A6" ID-DOI="10.1111/j.1096-3642.2010.00616.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5757493" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630446332507" checkinUser="valdenar" docAuthor="Meijer, Hanneke J. M. &amp; Due, Rokus Awe" docDate="2010" docId="03E48781E55EFFE8FC5DF91F0104C401" docLanguage="en" docName="j.1096-3642.2010.00616.x.pdf" docOrigin="Zoological Journal of the Linnean Society 160 (4)" docSource="http://dx.doi.org/10.1111/j.1096-3642.2010.00616.x" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Leptoptilos robustus Meijer &amp; Due 2010, SP. NOV." docType="treatment" docVersion="3" lastPageNumber="716" masterDocId="FFDDFFF9E559FFE1FFFEFF850066C201" masterDocTitle="A new species of giant marabou stork (Aves: Ciconiiformes) from the Pleistocene of Liang Bua, Flores (Indonesia)" masterLastPageNumber="724" masterPageNumber="707" pageNumber="714" updateTime="1699273657469" updateUser="plazi">
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<mods:title id="71FC2747EB3D1352EB55F239D60187D2">A new species of giant marabou stork (Aves: Ciconiiformes) from the Pleistocene of Liang Bua, Flores (Indonesia)</mods:title>
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<mods:namePart id="B081A8D7D6BBE9DA3024E3057E9628A5">Meijer, Hanneke J. M.</mods:namePart>
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<mods:namePart id="E7BBFC9B6C84EB1F55DAB5E723638661">Due, Rokus Awe</mods:namePart>
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<mods:title id="21D6BB65B3D623C5A8C184E93F7B2F54">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="C003C7FCD9E287CA29424FD5C2784B5B">2010</mods:date>
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<mods:number id="52D3CF404DF38C690FB05017AAD3AA6F">2010-11-24</mods:number>
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<treatment id="03E48781E55EFFE8FC5DF91F0104C401" LSID="urn:lsid:plazi:treatment:03E48781E55EFFE8FC5DF91F0104C401" httpUri="http://treatment.plazi.org/id/03E48781E55EFFE8FC5DF91F0104C401" lastPageId="9" lastPageNumber="716" pageId="7" pageNumber="714">
<subSubSection id="C357651CE55EFFE6FC5DF91F055EC4AF" box="[931,1336,1689,1711]" pageId="7" pageNumber="714" type="nomenclature">
<paragraph id="8BF23697E55EFFE6FC5DF91F055EC4AF" blockId="7.[931,1336,1689,1711]" box="[931,1336,1689,1711]" pageId="7" pageNumber="714">
<heading id="D0BA81FBE55EFFE6FC5DF91F055EC4AF" box="[931,1336,1689,1711]" centered="true" fontSize="9" level="2" pageId="7" pageNumber="714" reason="4">
<taxonomicName id="4C4D4D14E55EFFE6FC5DF91F04AFC4AF" authority="Meijer &amp; Due, 2010" authorityName="Meijer &amp; Due" authorityYear="2010" box="[931,1225,1690,1711]" class="Aves" family="Ciconiidae" genus="Leptoptilos" kingdom="Animalia" order="Ciconiiformes" pageId="7" pageNumber="720" phylum="Chordata" rank="species" species="robustus" status="sp. nov.">LEPTOPTILOS ROBUSTUS</taxonomicName>
<emphasis id="B939EA85E55EFFE6FB2DF918055EC4AF" bold="true" box="[1235,1336,1689,1710]" pageId="7" pageNumber="714">
<taxonomicNameLabel id="A20A57FEE55EFFE6FB2DF918055EC4AF" box="[1235,1336,1689,1710]" pageId="7" pageNumber="714" rank="species">SP. NOV.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C357651CE55EFFE8FCC4F9450104C401" lastPageId="9" lastPageNumber="716" pageId="7" pageNumber="714" type="materials_examined">
<paragraph id="8BF23697E55EFFE6FCC4F94505C7C56F" blockId="7.[826,1441,1728,1903]" pageId="7" pageNumber="714">
<emphasis id="B939EA85E55EFFE6FCC4F94503C2C4D4" box="[826,932,1728,1749]" italics="true" pageId="7" pageNumber="714">
<typeStatus id="54F68835E55EFFE6FCC4F94503F8C4D4" box="[826,926,1728,1749]" pageId="7" pageNumber="714" type="holotype">Holotype</typeStatus>
:
</emphasis>
Associated partial skeleton with wing and leg bones (
<figureCitation id="13762A12E55EFFE6FC46F95B0399C4F5" box="[952,1023,1758,1780]" captionStart="Figure 3" captionStartId="4.[144,223,1052,1071]" captionTargetBox="[399,1167,197,1022]" captionTargetId="figure-350@4.[399,1167,197,1022]" captionTargetPageId="4" captionText="Figure 3. Leptoptilos robustus sp. nov., proximal left carpometacarpus (LBA-XI-01). A, ventral view; B, dorsal view. Abbreviations: fccd, fovea carpalis caudalis; fi, fossa infratrochlearis; fs, fossa supratrochlearis; pp, processus pisiformis; tc, trochlea carpalis. Scale bar = 50 mm. Photographs by E. Kruidenier." figureDoi="http://doi.org/10.5281/zenodo.5757501" httpUri="https://zenodo.org/record/5757501/files/figure.png" pageId="7" pageNumber="714">Figs 3</figureCitation>
,
<figureCitation id="13762A12E55EFFE6FBF0F95B047BC4F5" box="[1038,1053,1758,1780]" captionStart="Figure 5" captionStartId="6.[144,223,1156,1175]" captionTargetBox="[399,1167,196,1126]" captionTargetId="figure-319@6.[399,1167,196,1126]" captionTargetPageId="6" captionText="Figure 5. Leptoptilos robustus sp. nov., distal part of left ulna (LBA-XI-02). A, ventral view; B, dorsal view. Abbreviations: cdu, condylus dorsalis ulnae; cvu, condylus ventralis ulnae; dr, depression radialis; f, foramen; itc, incisura tuberculi carpalis; pr, papillae remigales; si, sulcus intercondylaris; tc, tuberculum carpale. Scale bar = 50 mm. Photographs by E. Kruidenier." figureDoi="http://doi.org/10.5281/zenodo.5757505" httpUri="https://zenodo.org/record/5757505/files/figure.png" pageId="7" pageNumber="714">5</figureCitation>
,
<figureCitation id="13762A12E55EFFE6FBD3F95A045BC4F5" box="[1069,1085,1759,1780]" captionStart="Figure 7" captionStartId="8.[144,223,1310,1329]" captionTargetBox="[399,1167,194,1281]" captionTargetId="figure-163@8.[399,1167,194,1281]" captionTargetPageId="8" captionText="Figure 7. Leptoptilos robustus sp. nov., left femur (LBA-XI-03). A, caudal view; B, cranial view. Abbreviations: cl, condylus lateralis; cm, condylus medialis; ct, crista trochanteris; faa, facies articularis antitrochanterica; fcd, facies caudalis; fcr, facies cranialis; fp, fossa poplitea; ii, incisura intercondylaris; licd, linea intermuscularis caudalis; licr, linea intermuscularis cranialis; pf, pneumatic foramen; sp, sulcus patellaris; tf, trochanter femoris; tfi, trochlea fibularis. Scale bar = 50 mm. Photographs by E. Kruidenier." figureDoi="http://doi.org/10.5281/zenodo.5757509" httpUri="https://zenodo.org/record/5757509/files/figure.png" pageId="7" pageNumber="714">7</figureCitation>
,
<figureCitation id="13762A12E55EFFE6FBB5F95A043CC4F4" box="[1099,1114,1759,1781]" captionStart="Figure 9" captionStartId="10.[144,223,1273,1292]" captionTargetBox="[399,1167,195,1243]" captionTargetId="figure-235@10.[399,1167,195,1243]" captionTargetPageId="10" captionText="Figure 9. Leptoptilos robustus sp. nov., distal left tibiotarsus (LBA-XI-04). A, anterior view of specimen; B, anterior view of distal fragment; C, posterior view of distal fragment. Abbreviations: cs, location of natural fracture at which cross-section is visible; dce, distal opening of canalis extensorius; fcd, facies caudalis; fcr, facies cranialis; gm, gnawing marks; pce, proximal opening of canalis extensorius; pst, pons supratendineus; se, sulcus extensorius; tub, tubercle laterodistal of the pons supratendineus. Scale bars = 50 mm. Photographs by E. Kruidenier." figureDoi="http://doi.org/10.5281/zenodo.5757513" httpUri="https://zenodo.org/record/5757513/files/figure.png" pageId="7" pageNumber="714">9</figureCitation>
), consisting of the proximal half of a left carpometacarpus (preliminary registration nr LBA-XI-01) and the distal part of a left ulna (LBA-XI-02), an almost complete left femur (LBA-XI- 03), and the distal part of a left tibiotarsus (LBA-XI-
</paragraph>
<caption id="DF32661FE551FFE9FF6EFA9B020CC7A6" ID-DOI="http://doi.org/10.5281/zenodo.5757509" ID-Zenodo-Dep="5757509" httpUri="https://zenodo.org/record/5757509/files/figure.png" pageId="8" pageNumber="715" startId="8.[144,223,1310,1329]" targetBox="[399,1167,194,1281]" targetPageId="8">
<paragraph id="8BF23697E551FFE9FF6EFA9B020CC7A6" blockId="8.[144,1421,1310,1447]" pageId="8" pageNumber="715">
<emphasis id="B939EA85E551FFE9FF6EFA9B023CC733" bold="true" box="[144,602,1310,1330]" pageId="8" pageNumber="715">
Figure 7.
<taxonomicName id="4C4D4D14E551FFE9FEFAFA9B0193C730" authority="Meijer &amp; Due, 2010" authorityName="Meijer &amp; Due" authorityYear="2010" box="[260,501,1310,1329]" class="Aves" family="Ciconiidae" genus="Leptoptilos" kingdom="Animalia" order="Ciconiiformes" pageId="8" pageNumber="715" phylum="Chordata" rank="species" species="robustus" status="sp. nov.">
<emphasis id="B939EA85E551FFE9FEFAFA9B0193C730" bold="true" box="[260,501,1310,1329]" italics="true" pageId="8" pageNumber="715">Leptoptilos robustus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20A57FEE551FFE9FE01FA9A023CC733" box="[511,602,1311,1330]" pageId="8" pageNumber="715" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, left femur (LBA-XI-03). A, caudal view; B, cranial view. Abbreviations: cl, condylus lateralis; cm, condylus medialis; ct, crista trochanteris; faa, facies articularis antitrochanterica; fcd, facies caudalis; fcr, facies cranialis; fp, fossa poplitea; ii, incisura intercondylaris; licd, linea intermuscularis caudalis; licr, linea intermuscularis cranialis; pf, pneumatic foramen; sp, sulcus patellaris; tf, trochanter femoris; tfi, trochlea fibularis. Scale bar = 50 mm. Photographs by E. Kruidenier.
</paragraph>
</caption>
<paragraph id="8BF23697E551FFE9FF6EFA520257C40A" blockId="8.[144,758,1495,1547]" pageId="8" pageNumber="715">
04). The bones are stored at the National Centre for Archaeology in
<collectingRegion id="4989F875E551FFE9FEB8FA7001C6C40A" box="[326,416,1525,1547]" country="Indonesia" name="Jakarta Raya" pageId="8" pageNumber="715">Jakarta</collectingRegion>
(
<collectingCountry id="F35A7607E551FFE9FE4FFA700243C40A" box="[433,549,1525,1547]" name="Indonesia" pageId="8" pageNumber="715">Indonesia</collectingCountry>
).
</paragraph>
<paragraph id="8BF23697E551FFE9FF6EF9CF01F1C49C" blockId="8.[144,758,1610,1693]" pageId="8" pageNumber="715">
<emphasis id="B939EA85E551FFE9FF6EF9CF016BC45E" box="[144,269,1610,1631]" italics="true" pageId="8" pageNumber="715">Etymology:</emphasis>
From the Latin
<taxonomicName id="4C4D4D14E551FFE9FE30F9CF0256C45E" authorityName="Meijer &amp; Due" authorityYear="2010" box="[462,560,1610,1631]" class="Aves" family="Ciconiidae" genus="Leptoptilos" kingdom="Animalia" order="Ciconiiformes" pageId="8" pageNumber="715" phylum="Chordata" rank="species" species="robustus">
<emphasis id="B939EA85E551FFE9FE30F9CF0256C45E" box="[462,560,1610,1631]" italics="true" pageId="8" pageNumber="715">robustus</emphasis>
</taxonomicName>
meaning strong, robust, and referring to the large tibiotarsus and the thickness of its cortex.
</paragraph>
<paragraph id="8BF23697E551FFE9FF6EF95502ACC502" blockId="8.[144,758,1743,1796]" pageId="8" pageNumber="715">
<emphasis id="B939EA85E551FFE9FF6EF955014FC4E5" box="[144,297,1743,1765]" italics="true" pageId="8" pageNumber="715">Type locality:</emphasis>
<materialsCitation id="3B253CCAE551FFE9FEC8F94A02A0C502" country="Indonesia" latitude="-8.530666" location="Liang Bua cave" longLatPrecision="1" longitude="120.44358" municipality="Manggarai Province" pageId="8" pageNumber="715" specimenCount="1">
<location id="8E92604CE551FFE9FEC8F94A0192C4E4" LSID="urn:lsid:plazi:treatment:03E48781E55EFFE8FC5DF91F0104C401:8E92604CE551FFE9FEC8F94A0192C4E4" box="[310,500,1743,1765]" country="Indonesia" latitude="-8.530666" longLatPrecision="1" longitude="120.44358" municipality="Manggarai Province" name="Liang Bua cave" pageId="8" pageNumber="715">Liang Bua cave</location>
,
<collectingMunicipality id="6B96ACEDE551FFE9FDFAF94A0295C4E4" box="[516,755,1743,1765]" pageId="8" pageNumber="715">Manggarai Province</collectingMunicipality>
,
<collectingCountry id="F35A7607E551FFE9FF6EF96B00BCC505" box="[144,218,1774,1796]" name="Indonesia" pageId="8" pageNumber="715">Flores</collectingCountry>
,
<collectingCountry id="F35A7607E551FFE9FF19F96B0131C505" box="[231,343,1774,1796]" name="Indonesia" pageId="8" pageNumber="715">Indonesia</collectingCountry>
at
<geoCoordinate id="EE795050E551FFE9FE7CF96B0272C502" box="[386,532,1772,1796]" degrees="08" direction="south" minutes="31" orientation="latitude" pageId="8" pageNumber="715" precision="1" seconds="50.4" value="-8.530666">08°3150.4″S</geoCoordinate>
,
<geoCoordinate id="EE795050E551FFE9FDDDF96B02A0C502" box="[547,710,1772,1796]" degrees="120" direction="east" minutes="26" orientation="longitude" pageId="8" pageNumber="715" precision="1" seconds="36.9" value="120.44358">120°2636.9″E</geoCoordinate>
</materialsCitation>
.
</paragraph>
<paragraph id="8BF23697E551FFE9FF6EF8BE0449C40A" blockId="8.[144,758,1850,1903]" lastBlockId="8.[806,1421,1495,1547]" pageId="8" pageNumber="715">
<emphasis id="B939EA85E551FFE9FF6EF8BE01ECC551" box="[144,394,1850,1872]" italics="true" pageId="8" pageNumber="715">Type horizon and age:</emphasis>
Sector XI at Liang Bua (
<figureCitation id="13762A12E551FFE9FD53F8BF028BC551" box="[685,749,1850,1872]" captionStart="Figure 1" captionStartId="1.[826,905,1544,1563]" captionTargetBox="[826,1440,198,1515]" captionTargetId="figure-462@1.[826,1442,196,1515]" captionTargetPageId="1" captionText="Figure 1. Map of South-East Asia, showing (A) the location of Flores within Indonesia and (B) the location of Liang Bua Cave in western Flores. (C) is a map of the excavated sectors within Liang Bua. Sector VII/XI is located against the eastern wall of the cave. A and B adapted from van den Hoek Ostende et al. (2007), C is taken from Morwood et al. (2005) and reprinted with permission." figureDoi="http://doi.org/10.5281/zenodo.5757497" httpUri="https://zenodo.org/record/5757497/files/figure.png" pageId="8" pageNumber="715">Fig. 1</figureCitation>
) at a depth of
<quantity id="4CB59B72E551FFE9FEC6F8DC01D9C56F" box="[312,447,1881,1903]" metricMagnitude="0" metricUnit="m" metricValue="4.475" metricValueMax="4.7" metricValueMin="4.25" pageId="8" pageNumber="715" unit="cm" value="447.5" valueMax="470.0" valueMin="425.0">425470 cm</quantity>
in layers of brown clayey silts (Layer O in
<figureCitation id="13762A12E551FFE9FC0AFA520453C7EC" box="[1012,1077,1495,1517]" captionStart="Figure 2" captionStartId="2.[144,223,1380,1399]" captionTargetBox="[146,1422,198,1347]" captionTargetId="figure-223@2.[144,1424,196,1350]" captionTargetPageId="2" captionText="Figure 2. Stratigraphical sequence of the Sector VII/XI trench. Remains from Leptoptilos robustus sp. nov. were retrieved from Sector XI, spits 4347, which correspond to Layers OP. The white tuffaceous layer is indicated by Na-d and has been dated at 1117 kya. From Morwood et al. (2005), reprinted with permission." figureDoi="http://doi.org/10.5281/zenodo.5757499" httpUri="https://zenodo.org/record/5757499/files/figure.png" pageId="8" pageNumber="715">Fig. 2</figureCitation>
) and Late Pleistocene in age (
<bibRefCitation id="EFDC4B66E551FFE9FCD3FA700442C40A" author="Morwood MJ &amp; Brown P &amp; Jatmiko &amp; Sutikna T &amp; Wahyu Saptomo E &amp; Westaway KE &amp; Due AR &amp; Roberts RG &amp; Maeda T &amp; Wasisto S &amp; Djubiantono T" box="[813,1060,1525,1547]" pageId="8" pageNumber="715" pagination="1012 - 1017" refId="ref8980" refString="Morwood MJ, Brown P, Jatmiko, Sutikna T, Wahyu Saptomo E, Westaway KE, Due AR, Roberts RG, Maeda T, Wasisto S, Djubiantono T. 2005. Further evidence for small-bodied hominins from the Late Pleistocene of Flores, Indonesia. Nature 437: 1012 - 1017." type="journal article" year="2005">
Morwood
<emphasis id="B939EA85E551FFE9FC5FFA7303BEC40B" box="[929,984,1525,1547]" italics="true" pageId="8" pageNumber="715">et al.</emphasis>
, 2005
</bibRefCitation>
).
</paragraph>
<paragraph id="8BF23697E551FFE8FCD8F9C302C4C673" blockId="8.[806,1421,1605,1903]" lastBlockId="9.[164,779,902,1536]" lastPageId="9" lastPageNumber="716" pageId="8" pageNumber="715">
<emphasis id="B939EA85E551FFE9FCD8F9C303FBC45A" box="[806,925,1606,1627]" italics="true" pageId="8" pageNumber="715">Diagnosis:</emphasis>
A large species of extinct
<taxonomicName id="4C4D4D14E551FFE9FAEEF9C005EBC45B" authorityName="LESSON" authorityYear="1831" box="[1296,1421,1605,1626]" class="Aves" family="Ciconiidae" genus="Leptoptilos" kingdom="Animalia" order="Ciconiiformes" pageId="8" pageNumber="715" phylum="Chordata" rank="genus">
<emphasis id="B939EA85E551FFE9FAEEF9C005EBC45B" box="[1296,1421,1605,1626]" italics="true" pageId="8" pageNumber="715">Leptoptilos</emphasis>
</taxonomicName>
resembling
<taxonomicName id="4C4D4D14E551FFE9FC4DF9E0044EC478" baseAuthorityName="RMNH" baseAuthorityYear="1491" box="[947,1064,1636,1658]" class="Aves" family="Ciconiidae" genus="Leptoptilos" kingdom="Animalia" order="Ciconiiformes" pageId="8" pageNumber="715" phylum="Chordata" rank="species" species="dubius">
<emphasis id="B939EA85E551FFE9FC4DF9E0044EC478" box="[947,1064,1636,1658]" italics="true" pageId="8" pageNumber="715">L. dubius</emphasis>
</taxonomicName>
in dimensions of the femur, carpometacarpus, and ulna, but with a tibiotarsus wider and deeper than any living
<taxonomicName id="4C4D4D14E551FFE9FB4FF9240548C4B7" authorityName="LESSON" authorityYear="1831" box="[1201,1326,1697,1718]" class="Aves" family="Ciconiidae" genus="Leptoptilos" kingdom="Animalia" order="Ciconiiformes" pageId="8" pageNumber="715" phylum="Chordata" rank="genus">
<emphasis id="B939EA85E551FFE9FB4FF9240548C4B7" box="[1201,1326,1697,1718]" italics="true" pageId="8" pageNumber="715">Leptoptilos</emphasis>
</taxonomicName>
and yet smaller than
<taxonomicName id="4C4D4D14E551FFE9FC2FF945043AC4D4" baseAuthorityName="Milne-Edwards" baseAuthorityYear="1868" box="[977,1116,1728,1749]" class="Aves" family="Ciconiidae" genus="Leptoptilos" kingdom="Animalia" order="Ciconiiformes" pageId="8" pageNumber="715" phylum="Chordata" rank="species" species="falconeri">
<emphasis id="B939EA85E551FFE9FC2FF945043AC4D4" box="[977,1116,1728,1749]" italics="true" pageId="8" pageNumber="715">L. falconeri</emphasis>
</taxonomicName>
, and with the following unique combination of characters: pneumatized carpometacarpus with a distinct foramen in the fossa infratrochlearis; femur with well-pronounced muscle scars on proximocranial surface; linea intermuscularis cranialis and linea intermuscularis caudalis with a more lateral and medial location on the shaft, respectively, than extant
<taxonomicName id="4C4D4D14E550FFE8FE01FC2102EEC1BB" authorityName="KAHL" authorityYear="1971" box="[511,648,932,954]" class="Aves" family="Ciconiidae" kingdom="Animalia" order="Ciconiiformes" pageId="9" pageNumber="716" phylum="Chordata" rank="tribe" tribe="Leptoptilini">Leptoptilini</taxonomicName>
; condylus ventralis ulnae pointed, elevated, and projecting distally; tuberculum carpale rectangular in ventral view; foramen in incisura tuberculi carpalis; tibiotarsus with straight shaft and bone wall thicker than any species of
<taxonomicName id="4C4D4D14E550FFE8FED2FBB801CFC653" authorityName="LESSON" authorityYear="1831" box="[300,425,1085,1106]" class="Aves" family="Ciconiidae" genus="Leptoptilos" kingdom="Animalia" order="Ciconiiformes" pageId="9" pageNumber="716" phylum="Chordata" rank="genus">
<emphasis id="B939EA85E550FFE8FED2FBB801CFC653" box="[300,425,1085,1106]" italics="true" pageId="9" pageNumber="716">Leptoptilos</emphasis>
</taxonomicName>
; sulcus extensorius shallow, narrow, and located on medial half of bone.
</paragraph>
<caption id="DF32661FE550FFE8FF5AFC880263C13C" ID-DOI="http://doi.org/10.5281/zenodo.5757511" ID-Zenodo-Dep="5757511" httpUri="https://zenodo.org/record/5757511/files/figure.png" pageId="9" pageNumber="716" startId="9.[164,243,781,800]" targetBox="[325,1281,201,750]" targetPageId="9">
<paragraph id="8BF23697E550FFE8FF5AFC880263C13C" blockId="9.[164,1441,781,830]" pageId="9" pageNumber="716">
<emphasis id="B939EA85E550FFE8FF5AFC88016AC121" bold="true" box="[164,268,781,800]" pageId="9" pageNumber="716">Figure 8.</emphasis>
Graph showing the minimum width and minimum depth of the shaft of the femur of
<emphasis id="B939EA85E550FFE8FB4CFC88009BC13C" bold="true" pageId="9" pageNumber="716">
<taxonomicName id="4C4D4D14E550FFE8FB4CFC8805C7C121" authority="Meijer &amp; Due, 2010" authorityName="Meijer &amp; Due" authorityYear="2010" box="[1202,1441,781,800]" class="Aves" family="Ciconiidae" genus="Leptoptilos" kingdom="Animalia" order="Ciconiiformes" pageId="9" pageNumber="716" phylum="Chordata" rank="species" species="robustus" status="sp. nov.">
<emphasis id="B939EA85E550FFE8FB4CFC8805C7C121" bold="true" box="[1202,1441,781,800]" italics="true" pageId="9" pageNumber="716">Leptoptilos robustus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20A57FEE550FFE8FF5AFCAF009BC13C" box="[164,253,810,829]" pageId="9" pageNumber="716" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and extant
<taxonomicName id="4C4D4D14E550FFE8FE7CFCAF0267C13C" authorityName="KAHL" authorityYear="1971" box="[386,513,810,829]" class="Aves" family="Ciconiidae" kingdom="Animalia" order="Ciconiiformes" pageId="9" pageNumber="716" phylum="Chordata" rank="tribe" tribe="Leptoptilini">Leptoptilini</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BF23697E550FFE8FF42FBFF0104C401" blockId="9.[164,779,902,1536]" pageId="9" pageNumber="716">
Apomorphies for the genus
<taxonomicName id="4C4D4D14E550FFE8FDDFFBFF02F8C68E" authorityName="LESSON" authorityYear="1831" box="[545,670,1146,1167]" class="Aves" family="Ciconiidae" genus="Leptoptilos" kingdom="Animalia" order="Ciconiiformes" pageId="9" pageNumber="716" phylum="Chordata" rank="genus">
<emphasis id="B939EA85E550FFE8FDDFFBFF02F8C68E" box="[545,670,1146,1167]" italics="true" pageId="9" pageNumber="716">Leptoptilos</emphasis>
</taxonomicName>
are the pneumatized carpometacarpus with a distinct foramen in the fossa infratrochlearis and the pointed condylus ventralis ulnae. Autapomorphic characters for
<taxonomicName id="4C4D4D14E550FFE8FF30FB700134C70B" authority="Meijer &amp; Due, 2010" authorityName="Meijer &amp; Due" authorityYear="2010" box="[206,338,1269,1290]" class="Aves" family="Ciconiidae" genus="Leptoptilos" kingdom="Animalia" order="Ciconiiformes" pageId="9" pageNumber="716" phylum="Chordata" rank="species" species="robustus" status="sp. nov.">
<emphasis id="B939EA85E550FFE8FF30FB700134C70B" box="[206,338,1269,1290]" italics="true" pageId="9" pageNumber="716">L. robustus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20A57FEE550FFE8FEA3FB7001D3C70B" box="[349,437,1269,1290]" pageId="9" pageNumber="716" rank="species">sp. nov.</taxonomicNameLabel>
are its large size, the thickening of the cortical bone wall of the tibiotarsus, the absence of a rotation in the tibiotarsal shaft, the elevated and distal projection of the pointed condylus ventralis ulnae, pronounced muscle scars on the proximocranial surface of the femur, and a more lateral and medial location of the linea intermuscularis cranialis and linea intermuscularis caudalis on the femur shaft.
</paragraph>
</subSubSection>
</treatment>
</document>