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<document id="D85772C3D5842687ABF393EF77B5AEFD" ID-DOI="10.5281/zenodo.191476" ID-GBIF-Dataset="8603d4d1-efc8-44bb-84ae-76d3d8dd8161" ID-ISSN="1175-5326" ID-Zenodo-Dep="191476" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461128727612" checkinUser="plazi" docAuthor="Passos, Paulo, Mueses-Cisneros, Jonh Jairo, Lynch, John D. &amp; Fernandes, Ronaldo" docDate="2009" docId="03D087ECAA54775BFF11B28E160E23D6" docLanguage="en" docName="zt02293p034.pdf" docOrigin="Zootaxa 2293" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Atractus iridescens Peracca 1896" docType="treatment" docVersion="8" lastPageNumber="15" masterDocId="FFE9FF94AA5E7755FF86B32815112046" masterDocTitle="Pacific lowland snakes of the genus Atractus (Serpentes: Dipsadidae), with description of three new species" masterLastPageNumber="34" masterPageNumber="1" pageNumber="11" updateTime="1698597252759" updateUser="plazi">
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<mods:title id="04712E0F18783DBF8F4D7902582F5C41">Pacific lowland snakes of the genus Atractus (Serpentes: Dipsadidae), with description of three new species</mods:title>
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<mods:namePart id="C42A0281BF821FE78F77B023F162F450">Passos, Paulo</mods:namePart>
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<mods:namePart id="C2FCBEF11053673CA71D9948E454D9A9">Mueses-Cisneros, Jonh Jairo</mods:namePart>
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<mods:namePart id="B7C96263462018FAC0621B96E911A611">Lynch, John D.</mods:namePart>
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<treatment id="03D087ECAA54775BFF11B28E160E23D6" ID-DOI="http://doi.org/10.5281/zenodo.6214659" ID-GBIF-Taxon="119626676" ID-Zenodo-Dep="6214659" LSID="urn:lsid:plazi:treatment:03D087ECAA54775BFF11B28E160E23D6" httpUri="http://treatment.plazi.org/id/03D087ECAA54775BFF11B28E160E23D6" lastPageId="14" lastPageNumber="15" pageId="10" pageNumber="11">
<subSubSection id="C3636571AA54775FFF11B28E141521A5" pageId="10" pageNumber="11" type="nomenclature">
<paragraph id="8BC636FAAA54775FFF11B28E17202186" blockId="10.[151,561,422,483]" box="[151,561,422,448]" pageId="10" pageNumber="11">
<heading id="D08E8196AA54775FFF11B28E17202186" bold="true" box="[151,561,422,448]" fontSize="11" level="1" pageId="10" pageNumber="11" reason="1">
<taxonomicName id="4C794D79AA54775FFF11B28E17202186" authority="Peracca, 1896" authorityName="Peracca" authorityYear="1896" box="[151,561,422,448]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="iridescens">
<emphasis id="B90DEAE8AA54775FFF11B28E17202186" bold="true" box="[151,561,422,448]" pageId="10" pageNumber="11">
<emphasis id="B90DEAE8AA54775FFF11B28E146C2186" bold="true" box="[151,381,422,448]" italics="true" pageId="10" pageNumber="11">Atractus iridescens</emphasis>
<bibRefCitation id="EFE84B0BAA54775FFE02B28E17202186" author="Peracca" box="[388,561,422,448]" pageId="10" pageNumber="33" refString="Peracca, M. G. (1896) Sopra alcuni ofidii nuovi o poco noti dell'America meridionale. Bolletin Museo di Torino, 11, 1 - 12." type="journal article" year="1896">Peracca, 1896</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BC636FAAA54775FFF11B2E1141521A5" blockId="10.[151,561,422,483]" box="[151,260,457,483]" pageId="10" pageNumber="11">
<figureCitation id="13422A7FAA54775FFF11B2E115F921A5" box="[151,232,457,483]" captionStart="FIGURE 7" captionStartId="10.[151,254,1400,1424]" captionTargetBox="[203,1360,803,1374]" captionTargetId="figure@10.[189,1397,796,1376]" captionTargetPageId="10" captionText="FIGURE 7. Dorsal (A) and ventral (B) views of Atractus iridescens (ICN 10902). SVL 353 mm, CL 50 mm." httpUri="https://zenodo.org/record/191483/files/figure.png" pageId="10" pageNumber="11">Figs. 7</figureCitation>
,
<figureCitation id="13422A7FAA54775FFF73B2E1141521A5" box="[245,260,457,483]" captionStart="FIGURE 8" captionStartId="11.[151,254,1965,1989]" captionTargetBox="[174,1417,501,1964]" captionTargetId="figure@11.[168,1428,491,2012]" captionTargetPageId="11" captionText="FIGURE 8. Dorsal (A) and lateral (B) views of the head of Atractus iridescens (ICN 10902). Scale = 5 mm." httpUri="https://zenodo.org/record/191484/files/figure.png" pageId="10" pageNumber="11">8</figureCitation>
</paragraph>
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<paragraph id="8BC636FAAA54775FFF11B13E168A220B" blockId="10.[151,999,534,589]" pageId="10" pageNumber="11">
<taxonomicName id="4C794D79AA54775FFF11B13E1712226B" authority="Peracca, 1896" authorityName="Peracca" authorityYear="1896" box="[151,515,534,557]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="iridescens">
<emphasis id="B90DEAE8AA54775FFF11B13E1475226B" box="[151,356,534,557]" italics="true" pageId="10" pageNumber="11">Atractus iridescens</emphasis>
<bibRefCitation id="EFE84B0BAA54775FFEEDB13E1712226B" author="Peracca" box="[363,515,534,557]" pageId="10" pageNumber="33" refString="Peracca, M. G. (1896) Sopra alcuni ofidii nuovi o poco noti dell'America meridionale. Bolletin Museo di Torino, 11, 1 - 12." type="journal article" year="1896">Peracca, 1896</bibRefCitation>
</taxonomicName>
; Boll. Mus. Zool. Anat. Comp. Torino 252:2.
<taxonomicName id="4C794D79AA54775FFF11B11E1475220B" box="[151,356,566,589]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="iridescens">
<emphasis id="B90DEAE8AA54775FFF11B11E1475220B" box="[151,356,566,589]" italics="true" pageId="10" pageNumber="11">Atractus iridescens</emphasis>
</taxonomicName>
<bibRefCitation id="EFE84B0BAA54775FFE06B11E1723220B" author="Boulenger" box="[384,562,566,589]" pageId="10" pageNumber="31" refString="Boulenger, G. A. (1913) A collection of batrachians and reptiles made by Dr. H. G. F. Spurrell, F. Z. S., in the Choco. Colombia. Proceedings Zollogical Society, London, 1913, 1019 - 1038." type="proceedings paper" year="1913">Boulenger, 1913</bibRefCitation>
; Proc. Zool. Soc. London 4:1035.
</paragraph>
</subSubSection>
<subSubSection id="C3636571AA54775FFF11B155113922A2" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="8BC636FAAA54775FFF11B155113922A2" blockId="10.[151,1437,637,740]" pageId="10" pageNumber="11">
<emphasis id="B90DEAE8AA54775FFF11B155140622D1" bold="true" box="[151,279,637,663]" pageId="10" pageNumber="11">
<typeStatus id="54C28858AA54775FFF11B155140322D1" box="[151,274,637,663]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
:
</emphasis>
Adult male, from “
<collectingCountry id="F36E766AAA54775FFD80B155176322D1" box="[518,626,637,663]" name="United States of America" pageId="10" pageNumber="11">América</collectingCountry>
Meridionali” (= South
<collectingCountry id="F36E766AAA54775FFC15B15516EF22D1" box="[915,1022,637,663]" name="United States of America" pageId="10" pageNumber="11">America</collectingCountry>
) originally deposited at Museo di Zoologia dell Università degli Studi di Torino, Turin
<collectionCode id="ED68AE3FAA54775FFCB7B18C169722F8" box="[817,902,676,702]" httpUri="http://grbio.org/cool/yjj8-is06" name="Museo di Zoologia, Instituto di Zoologia e Anatomia Comparata Universita di Torino" pageId="10" pageNumber="11">MZUT</collectionCode>
R1830, now housed at Museo Regionali di Scienze Naturali, Turin (
<bibRefCitation id="EFE84B0BAA54775FFE31B1E217FE22A2" author="Andreone" box="[439,751,714,740]" pageId="10" pageNumber="31" refString="Andreone, F. &amp; Gavetti, E. (2007) The Life and Herpetological Contributions of Mario Ciacinto Peracca (1861 - 1923). Society of the Study of Amphibians and Reptiles Press, Villanova, Pennsylvania, U. S. A., 570 pp." type="book chapter" year="2007">Andreone &amp; Gavetti 2007</bibRefCitation>
), (photographs examined).
</paragraph>
</subSubSection>
<caption id="DF066672AA54775FFF11B650103725C9" box="[151,1318,1400,1424]" httpUri="https://zenodo.org/record/191483/files/figure.png" pageId="10" pageNumber="11" targetBox="[203,1360,803,1374]" targetPageId="10">
<paragraph id="8BC636FAAA54775FFF11B650103725C9" blockId="10.[151,1318,1400,1424]" box="[151,1318,1400,1424]" pageId="10" pageNumber="11">
<emphasis id="B90DEAE8AA54775FFF11B650140925D6" bold="true" box="[151,280,1400,1424]" pageId="10" pageNumber="11">FIGURE 7.</emphasis>
Dorsal (A) and ventral (B) views of
<taxonomicName id="4C794D79AA54775FFD24B650166125C9" box="[674,880,1400,1423]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="iridescens">
<emphasis id="B90DEAE8AA54775FFD24B650166125C9" box="[674,880,1400,1423]" italics="true" pageId="10" pageNumber="11">Atractus iridescens</emphasis>
</taxonomicName>
(ICN 10902). SVL 353 mm, CL 50 mm.
</paragraph>
</caption>
<subSubSection id="C3636571AA54775EFF40B6C014F82186" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="diagnosis">
<paragraph id="8BC636FAAA54775FFF40B6C016FE2771" blockId="10.[151,1437,1512,2002]" pageId="10" pageNumber="11">
<emphasis id="B90DEAE8AA54775FFF40B6C014592644" bold="true" box="[198,328,1512,1538]" pageId="10" pageNumber="11">Diagnosis:</emphasis>
<taxonomicName id="4C794D79AA54775FFED0B6C1172C2644" box="[342,573,1513,1538]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="iridescens">
<emphasis id="B90DEAE8AA54775FFED0B6C1172C2644" box="[342,573,1513,1538]" italics="true" pageId="10" pageNumber="11">Atractus iridescens</emphasis>
</taxonomicName>
is distinguished from all congeners by combination of the following characters: (1) 17/17/17 smooth dorsal scale rows; (2) two postoculars; (3) loreal long; (4) temporals 1+2; (5) seven supralabials, third and fourth contacting orbit; (6) seven infralabials, first three or four contacting chinshields; (7) six or seven maxillary teeth; (8) generally four gular scale rows; (9) three preventrals; (10) 135144 ventrals in females,
<quantity id="4C819B1FAA54775FFD85B5AB179F26DB" box="[515,654,1667,1693]" metricMagnitude="0" metricUnit="m" metricValue="3.6195" metricValueMax="3.81" metricValueMin="3.429" pageId="10" pageNumber="11" unit="in" value="142.5" valueMax="150.0" valueMin="135.0">135150 in</quantity>
males; (11) 2537 subcaudals in females,
<quantity id="4C819B1FAA54775FFB1AB5AB101926DB" box="[1180,1288,1667,1693]" metricMagnitude="-1" metricUnit="m" metricValue="9.017" metricValueMax="9.652" metricValueMin="8.382" pageId="10" pageNumber="11" unit="in" value="35.5" valueMax="38.0" valueMin="33.0">3338 in</quantity>
males; (12) dorsum reddish brown scattered with small and irregular dark blotches; (13) venter beige with a median line comprised of small black dots; (14) moderate body size, single subadult female with
<quantity id="4C819B1FAA54775FFB27B5F8101426AC" box="[1185,1285,1744,1770]" metricMagnitude="-1" metricUnit="m" metricValue="1.7" pageId="10" pageNumber="11" unit="mm" value="170.0">170 mm</quantity>
SVL, males reaching
<quantity id="4C819B1FAA54775FFE8FB5DF147E2757" box="[265,367,1783,1809]" metricMagnitude="-1" metricUnit="m" metricValue="3.53" pageId="10" pageNumber="11" unit="mm" value="353.0">353 mm</quantity>
SVL; (15) long tail in females (16.417.6% SVL) and males (14.218.9% SVL); (16) hemipenis moderately bilobed, slightly semicapitate, and non-calyculate.
</paragraph>
<paragraph id="8BC636FAAA54775FFF40B46C108C2794" blockId="10.[151,1437,1512,2002]" pageId="10" pageNumber="11">
<emphasis id="B90DEAE8AA54775FFF40B46C14672718" bold="true" box="[198,374,1860,1886]" pageId="10" pageNumber="11">Comparisons:</emphasis>
<taxonomicName id="4C794D79AA54775FFE04B46C1779271B" box="[386,616,1860,1885]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="iridescens">
<emphasis id="B90DEAE8AA54775FFE04B46C1779271B" box="[386,616,1860,1885]" italics="true" pageId="10" pageNumber="11">Atractus iridescens</emphasis>
</taxonomicName>
is distinguished from all congeners, except
<taxonomicName id="4C794D79AA54775FFB0CB46C11EE271B" box="[1162,1279,1860,1885]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="typhon">
<emphasis id="B90DEAE8AA54775FFB0CB46C11EE271B" box="[1162,1279,1860,1885]" italics="true" pageId="10" pageNumber="11">A. typhon</emphasis>
</taxonomicName>
by having a lateral projection on the base of the hemipenis.
<taxonomicName id="4C794D79AA54775FFD47B44316B227C2" box="[705,931,1899,1924]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="iridescens">
<emphasis id="B90DEAE8AA54775FFD47B44316B227C2" box="[705,931,1899,1924]" italics="true" pageId="10" pageNumber="11">Atractus iridescens</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C794D79AA54775FFBBBB44311BF27C2" box="[1085,1198,1899,1924]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="typhon">
<emphasis id="B90DEAE8AA54775FFBBBB44311BF27C2" box="[1085,1198,1899,1924]" italics="true" pageId="10" pageNumber="11">A. typhon</emphasis>
</taxonomicName>
by having 17 dorsal scale rows at midbody, 135150 ventrals in males, dorsum with irregular dark blotches, venter with median black stripe (vs. 15 dorsals, 156163 ventrals, banded dorsum, venter with quadriculate dark bars).
</paragraph>
<paragraph id="8BC636FAAA55775EFF11B3B014F82186" blockId="11.[151,1437,152,448]" pageId="11" pageNumber="12">
Additionally,
<taxonomicName id="4C794D79AA55775EFEBDB3B014C520F7" box="[315,468,152,177]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="iridescens">
<emphasis id="B90DEAE8AA55775EFEBDB3B014C520F7" box="[315,468,152,177]" italics="true" pageId="11" pageNumber="12">A. iridescens</emphasis>
</taxonomicName>
shares only with
<taxonomicName id="4C794D79AA55775EFD2FB3B0163820F7" box="[681,809,152,177]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="echidna">
<emphasis id="B90DEAE8AA55775EFD2FB3B0163820F7" box="[681,809,152,177]" italics="true" pageId="11" pageNumber="12">A. echidna</emphasis>
</taxonomicName>
(hemipenis unknown) 36 subcaudals in males, three preventrals, six to eight maxillary teeth, dorsum brown with irregular dark blotches.
<taxonomicName id="4C794D79AA55775EFB1EB3971091209E" box="[1176,1408,191,216]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="iridescens">
<emphasis id="B90DEAE8AA55775EFB1EB3971091209E" box="[1176,1408,191,216]" italics="true" pageId="11" pageNumber="12">Atractus iridescens</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="4C794D79AA55775EFE01B3CD171D20B8" box="[391,524,229,254]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="echidna">
<emphasis id="B90DEAE8AA55775EFE01B3CD171D20B8" box="[391,524,229,254]" italics="true" pageId="11" pageNumber="12">A. echidna</emphasis>
</taxonomicName>
by having 17 dorsal scale rows, 135150 ventrals in males, venter with a median black stripe anteriorly and entirely black posteriorly (vs. 15 dorsals, 127 ventrals in the single male, venter cream with irregular brown spots posteriorly); from
<emphasis id="B90DEAE8AA55775EFCDBB21A1199210D" box="[861,1160,306,331]" italics="true" pageId="11" pageNumber="12">
<taxonomicName id="4C794D79AA55775EFCDBB21A111A210D" box="[861,1035,306,331]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="boulengerii">A. boulengerii</taxonomicName>
,
<taxonomicName id="4C794D79AA55775EFB9EB21A1193210D" box="[1048,1154,306,331]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="clarki">A. clarki</taxonomicName>
,
</emphasis>
and
<taxonomicName id="4C794D79AA55775EFB45B21A1067210D" box="[1219,1398,306,331]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="multicinctus">
<emphasis id="B90DEAE8AA55775EFB45B21A1067210D" box="[1219,1398,306,331]" italics="true" pageId="11" pageNumber="12">A. multicinctus</emphasis>
</taxonomicName>
by having venter heavily marked with black dots, sometimes concentrated on the center of each ventral scale forming a conspicuous midline (vs. venter immaculate creamish white, occasionally with a few disperse black dots posteriorly in
<taxonomicName id="4C794D79AA55775EFEF5B28F14C92186" box="[371,472,423,448]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="clarki">
<emphasis id="B90DEAE8AA55775EFEF5B28F14C92186" box="[371,472,423,448]" italics="true" pageId="11" pageNumber="12">A. clarki</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<caption id="DF066672AA55775EFF11B485100C2782" box="[151,1309,1965,1989]" httpUri="https://zenodo.org/record/191484/files/figure.png" pageId="11" pageNumber="12" targetBox="[174,1417,501,1964]" targetPageId="11">
<paragraph id="8BC636FAAA55775EFF11B485100C2782" blockId="11.[151,1435,1965,2026]" box="[151,1309,1965,1989]" pageId="11" pageNumber="12">
<emphasis id="B90DEAE8AA55775EFF11B48514082783" bold="true" box="[151,281,1965,1989]" pageId="11" pageNumber="12">FIGURE 8.</emphasis>
Dorsal (A) and lateral (B) views of the head of
<taxonomicName id="4C794D79AA55775EFC9EB48616F42783" box="[792,997,1966,1989]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="iridescens">
<emphasis id="B90DEAE8AA55775EFC9EB48616F42783" box="[792,997,1966,1989]" italics="true" pageId="11" pageNumber="12">Atractus iridescens</emphasis>
</taxonomicName>
(ICN 10902). Scale = 5 mm.
</paragraph>
</caption>
<subSubSection id="C3636571AA55775BFF40B4F817C6210A" lastPageId="14" lastPageNumber="15" pageId="11" pageNumber="12" type="description">
<paragraph id="8BC636FAAA557759FF40B4F8143722EE" blockId="11.[151,1435,1965,2026]" lastBlockId="12.[151,1437,152,1956]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<emphasis id="B90DEAE8AA55775EFF40B4F8147027AC" bold="true" box="[198,353,2000,2026]" pageId="11" pageNumber="12">Description:</emphasis>
Head about twice as long as wide, flattened in lateral view, rounded in dorsal view; snout truncate in lateral view, rounded in dorsal view; cervical constriction indistinct; rostral subtriangular in frontal view, broader than high, poorly visible in dorsal view; internasal as long as wide; internasal suture linear or sinistral with respect to prefrontal suture; prefrontal as long as wide; supraocular sub-trapezoidal, about twice as long as wide; frontal sub-triangular, as long as wide; parietal about twice as long as wide; nasal divided; nostril restricted to prenasal; prenasal twice as high as long; postnasal as high as long; loreal long, contacting second and third supralabials; pupil round; two postoculars; upper postocular slightly higher and longer than lower; temporals 1+2; first temporal twice as long as high; upper posterior temporal elongate, three times longer than wide; seven supralabials, third and fourth contacting orbit; first two supralabials of similar size and smaller than third; sixth higher and seventh longer than remaining supralabials; symphisial semicircular, about three times broader than long; seven infralabials, first three or four contacting chinshields; first pair of infralabials in contact behind symphisial, preventing symphisial/chinshields contact; chinshields about three times longer than wide; four gular scales rows; three or four preventrals; generally 17/17/17 smooth dorsal scale rows; dorsals lacking apical pits, supra-anal tubercles, and keels; caudal spine moderate, conical, and acuminated.
</paragraph>
<paragraph id="8BC636FAAA527759FF40B19D17932305" blockId="12.[151,1437,152,1956]" pageId="12" pageNumber="13">
<emphasis id="B90DEAE8AA527759FF40B19D14802289" bold="true" box="[198,401,693,719]" pageId="12" pageNumber="13">Maxillary arch:</emphasis>
Arched in dorsal view, with four or five prediastemal and two postdiastemal teeth; prediastemal teeth large, moderately spaced, of similar size, curved posteriorly, angular in cross section, robust at base, narower at apices; maxillary diastema moderately long; postdiastemal teeth half size of the last prediastemal tooth; lateral process absent.
</paragraph>
<paragraph id="8BC636FAAA527759FF40B067173125E8" blockId="12.[151,1437,152,1956]" pageId="12" pageNumber="13">
<emphasis id="B90DEAE8AA527759FF40B06714F6232F" bold="true" box="[198,487,847,873]" pageId="12" pageNumber="13">Colour in preservative:</emphasis>
Dorsum of head reddish brown, occasionally with small beige spots at the orbital region; head background reddish brown to dorsal margin of supralabials, except for black postorbital and creamy temporal stripes; temporal region and posterior portion of seventh supralabial creamish white, forming lower portion of temporal stripe; supralabials mostly cream, with invasion of brown pigment on posterior suture of supralabials; fifth and sixth supralabials predominantly dark brown, constituting the descending portion of postocular stripe; infralabials, chinshields, gulars, and preventrals creamish white with diffuse dark brown dots; ventral ground colour beige with three series of dark brown spots disposed linearly, forming ventral lines; two small paraventral stripes restricted to lateral or latero-medial region of ventrals; large median stripe increasing in size posteriorly; ventral lines collapsed on posterior regions of belly, covering ventral scales entirely; tail uniformly black; dorsal ground colour reddish brown with about 30 irregular black blotches; dark blotches with creamy borders and disposed linearly, extending from third to sixth dorsal scale rows; blotches (two or three scales long and wide) generally clearly distinct from dorsal ground colour and paired on paravertebral region; blotches occasionally collapsed at vertebral region on the first third of body; posterior blotches decreasing in size (one or two scales long and wide) and not connected to opposite one above vertebral region; blotches occasionally weakly distinct from dorsal ground colour, forming almost reticulate pattern.
</paragraph>
<paragraph id="8BC636FAAA527759FF40B69216AF27E2" blockId="12.[151,1437,152,1956]" pageId="12" pageNumber="13">
<emphasis id="B90DEAE8AA527759FF40B69214FF2592" bold="true" box="[198,494,1466,1492]" pageId="12" pageNumber="13">Hemipenis morphology</emphasis>
(everted organs
<emphasis id="B90DEAE8AA527759FD3AB69317DA2592" box="[700,715,1467,1492]" italics="true" pageId="12" pageNumber="13">n</emphasis>
= 4): Organ moderately bilobed, slightly semicapitate, and non-calyculate; lobes attenuate, centrifugally oriented, covered with disperse spinules, and slightly smaller than hemipenial body; apices of lobes curved and oriented medially, forming well defined distal clasps; tips of lobes covered with disperse papillae; hemipenial body with intrasulcar region thick (capitular area) on both sides of organ; weakly defined capitular groove with inverted “V” shape, delimiting capitate area on the sulcate side, and with median retraction on the asulcate side of hemipenis; capitular region delimited on the sulcate side by large hooked spines arranged in diagonal series; intrasulcar region with six moderate hooked spines; median spines decreasing in size toward proximal portion of asulcate side of organ;
<emphasis id="B90DEAE8AA527759FB38B5E1108D26A4" box="[1214,1436,1737,1762]" italics="true" pageId="12" pageNumber="13">sulcus spermaticus</emphasis>
bifurcates on basal portion of hemipenis;
<emphasis id="B90DEAE8AA527759FD0EB5D81678274F" box="[648,873,1776,1801]" italics="true" pageId="12" pageNumber="13">sulcus spermaticus</emphasis>
branches centrifugally oriented, running to the tips of lobes; margins of
<emphasis id="B90DEAE8AA527759FE38B43F178F2776" box="[446,670,1815,1840]" italics="true" pageId="12" pageNumber="13">sulcus spermaticus</emphasis>
stout and narrow; hemipenial body mostly smooth, with a series of moderate spines on asulcate side of organ immediately below large spines, which delimit the capitular region of hemipenis; basal portion of hemipenis body with a large lobular projection; proximal region of hemipenis with longitudinal plicae and disperse spinules (
<figureCitation id="13422A7FAA527759FCBBB4A2169227E2" box="[829,899,1930,1956]" captionStart="FIGURE 9" captionStartId="13.[151,255,1719,1743]" captionTargetBox="[188,1418,185,1689]" captionTargetId="figure@13.[163,1425,160,1709]" captionTargetPageId="13" captionText="FIGURE 9. Sulcate and asulcate sides of the hemipenis of (A) Atractus iridescens (ICN 10902), (B) detailed view of lateral expansion of the hemipenis of Atractus iridesnces, (C) sulcate and asulcate sides of the hemipenis of Atractus typhon sp. nov. (ICN 10901). Scale = 5 mm." httpUri="https://zenodo.org/record/191485/files/figure.png" pageId="12" pageNumber="13">Fig. 9</figureCitation>
a, b).
</paragraph>
<caption id="DF066672AA537758FF11B59F17782755" httpUri="https://zenodo.org/record/191485/files/figure.png" pageId="13" pageNumber="14" targetBox="[188,1418,185,1689]" targetPageId="13">
<paragraph id="8BC636FAAA537758FF11B59F17782755" blockId="13.[151,1437,1719,1812]" pageId="13" pageNumber="14">
<emphasis id="B90DEAE8AA537758FF11B59F140D2689" bold="true" box="[151,284,1719,1743]" pageId="13" pageNumber="14">FIGURE 9.</emphasis>
Sulcate and asulcate sides of the hemipenis of (A)
<taxonomicName id="4C794D79AA537758FCCBB59F110E2688" box="[845,1055,1719,1742]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="iridescens">
<emphasis id="B90DEAE8AA537758FCCBB59F110E2688" box="[845,1055,1719,1742]" italics="true" pageId="13" pageNumber="14">Atractus iridescens</emphasis>
</taxonomicName>
(ICN 10902), (B) detailed view of lateral expansion of the hemipenis of
<taxonomicName id="4C794D79AA537758FDB0B5F2161826B7" box="[566,777,1754,1777]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="iridesnces">
<emphasis id="B90DEAE8AA537758FDB0B5F2161826B7" box="[566,777,1754,1777]" italics="true" pageId="13" pageNumber="14">Atractus iridesnces</emphasis>
</taxonomicName>
, (C) sulcate and asulcate sides of the hemipenis of
<taxonomicName id="4C794D79AA537758FAC6B5F215F12752" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="typhon" status="sp. nov.">
<emphasis id="B90DEAE8AA537758FAC6B5F215F12752" italics="true" pageId="13" pageNumber="14">Atractus typhon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E5793AA537758FF60B5D414232755" box="[230,306,1788,1811]" pageId="13" pageNumber="14" rank="species">sp.nov.</taxonomicNameLabel>
(ICN 10901). Scale = 5mm.
</paragraph>
</caption>
<paragraph id="8BC636FAAA53775BFF40B44417C6210A" blockId="13.[151,1436,1900,2004]" lastBlockId="14.[151,1437,152,912]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<emphasis id="B90DEAE8AA537758FF40B444145227C0" bold="true" box="[198,323,1900,1926]" pageId="13" pageNumber="14">Variation:</emphasis>
Largest male
<quantity id="4C819B1FAA537758FE76B444174227C0" box="[496,595,1900,1926]" metricMagnitude="-1" metricUnit="m" metricValue="3.53" pageId="13" pageNumber="14" unit="mm" value="353.0">353 mm</quantity>
SVL,
<quantity id="4C819B1FAA537758FD22B44417E927C0" box="[676,760,1900,1926]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="14" unit="mm" value="50.0">50 mm</quantity>
CL, largest female
<quantity id="4C819B1FAA537758FC63B444115927C0" box="[997,1096,1900,1926]" metricMagnitude="-1" metricUnit="m" metricValue="1.7" pageId="13" pageNumber="14" unit="mm" value="170.0">170 mm</quantity>
SVL,
<quantity id="4C819B1FAA537758FB1EB44411FC27C0" box="[1176,1261,1900,1926]" metricMagnitude="-2" metricUnit="m" metricValue="3.8" pageId="13" pageNumber="14" unit="mm" value="38.0">38 mm</quantity>
CL; tail 14.2 18.9% (
<emphasis id="B90DEAE8AA537758FF77B4BB15EF27EA" box="[241,254,1939,1964]" italics="true" pageId="13" pageNumber="14"></emphasis>
= 17.7; SD = 2.5;
<emphasis id="B90DEAE8AA537758FE5EB4BB14F627EA" box="[472,487,1939,1964]" italics="true" pageId="13" pageNumber="14">n</emphasis>
= 4) SVL in males, 16.417.6% (
<emphasis id="B90DEAE8AA537758FCF4B4BB169027EA" box="[882,897,1939,1964]" italics="true" pageId="13" pageNumber="14">n</emphasis>
= 2) SVL in females; 135150 (
<emphasis id="B90DEAE8AA537758FB7FB4BB101427EA" box="[1273,1285,1939,1964]" italics="true" pageId="13" pageNumber="14"></emphasis>
= 137.8; SD = 3.3;
<emphasis id="B90DEAE8AA537758FF65B49215E32795" box="[227,242,1978,2003]" italics="true" pageId="13" pageNumber="14">n</emphasis>
= 5) ventrals in males, 134144 (
<emphasis id="B90DEAE8AA537758FDFAB49217982795" box="[636,649,1978,2003]" italics="true" pageId="13" pageNumber="14"></emphasis>
= 139; SD = 4.6;
<emphasis id="B90DEAE8AA537758FCD9B492167F2795" box="[863,878,1978,2003]" italics="true" pageId="13" pageNumber="14">n</emphasis>
= 3) in females; 3343 (
<emphasis id="B90DEAE8AA537758FB0DB49211892795" box="[1163,1176,1978,2003]" italics="true" pageId="13" pageNumber="14"></emphasis>
= 37.7; SD = 1.6;
<emphasis id="B90DEAE8AA537758FAF3B49210952795" box="[1397,1412,1978,2003]" italics="true" pageId="13" pageNumber="14">n</emphasis>
= 5) subcaudals in males, 2537 (
<emphasis id="B90DEAE8AA50775BFD8CB3B0170920F7" box="[522,536,152,177]" italics="true" pageId="14" pageNumber="15">n</emphasis>
= 2) in females; 3 (
<emphasis id="B90DEAE8AA50775BFD79B3B0161F20F7" box="[767,782,152,177]" italics="true" pageId="14" pageNumber="15">n</emphasis>
= 1 side) or 4 (
<emphasis id="B90DEAE8AA50775BFC44B3B016C120F7" box="[962,976,152,177]" italics="true" pageId="14" pageNumber="15">n</emphasis>
= 15 sides) first infralabials contacting chinshields; 3 (
<emphasis id="B90DEAE8AA50775BFECFB3971446209E" box="[329,343,191,216]" italics="true" pageId="14" pageNumber="15">n</emphasis>
= 3 side) or 4 (
<emphasis id="B90DEAE8AA50775BFD8FB3971709209E" box="[521,536,191,216]" italics="true" pageId="14" pageNumber="15">n</emphasis>
= 10 sides) gular scale rows; 3 (
<emphasis id="B90DEAE8AA50775BFC17B39716B1209E" box="[913,928,191,216]" italics="true" pageId="14" pageNumber="15">n</emphasis>
= 5) or 4 (
<emphasis id="B90DEAE8AA50775BFB9AB397113A209E" box="[1052,1067,191,216]" italics="true" pageId="14" pageNumber="15">n</emphasis>
= 2) preventrals; 811 (
<emphasis id="B90DEAE8AA50775BFAC5B3971041209E" box="[1347,1360,191,216]" italics="true" pageId="14" pageNumber="15"></emphasis>
= 8.8; SD = 0.7;
<emphasis id="B90DEAE8AA50775BFE89B3CD140F20B8" box="[271,286,229,254]" italics="true" pageId="14" pageNumber="15">n</emphasis>
= 12 sides) dorsal scale rows at the level of second subcaudal;
<quantity id="4C819B1FAA50775BFB87B3CD119F20B9" box="[1025,1166,229,255]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.6" metricValueMin="4.4" pageId="14" pageNumber="15" unit="mm" value="5.5" valueMax="6.6" valueMin="4.4">4.46.6 mm</quantity>
(
<emphasis id="B90DEAE8AA50775BFB1AB3CD11B820B8" box="[1180,1193,229,254]" italics="true" pageId="14" pageNumber="15"></emphasis>
= 5.4; SD = 1.2;
<emphasis id="B90DEAE8AA50775BFAF3B3CD109520B8" box="[1397,1412,229,254]" italics="true" pageId="14" pageNumber="15">n</emphasis>
= 6) body diameter; 6 (
<emphasis id="B90DEAE8AA50775BFE08B224148C2163" box="[398,413,268,293]" italics="true" pageId="14" pageNumber="15">n</emphasis>
= 4 sides) or 7 (
<emphasis id="B90DEAE8AA50775BFDDBB224177D2163" box="[605,620,268,293]" italics="true" pageId="14" pageNumber="15">n</emphasis>
= 4 sides) maxillary teeth; 5 (
<emphasis id="B90DEAE8AA50775BFC4AB22416CA2163" box="[972,987,268,293]" italics="true" pageId="14" pageNumber="15">n</emphasis>
= 2 sides) palatine teeth; 10 (
<emphasis id="B90DEAE8AA50775BFAB0B22410542163" box="[1334,1349,268,293]" italics="true" pageId="14" pageNumber="15">n</emphasis>
= 1) or 11 (
<emphasis id="B90DEAE8AA50775BFF42B21A15C2210D" box="[196,211,306,331]" italics="true" pageId="14" pageNumber="15">n</emphasis>
=1) pterigoyd teeth; 8 (
<emphasis id="B90DEAE8AA50775BFE6CB21A14E9210D" box="[490,504,306,331]" italics="true" pageId="14" pageNumber="15">n</emphasis>
= 1) dentary teeth.
</paragraph>
</subSubSection>
<subSubSection id="C3636571AA50775BFF40B27016E02186" pageId="14" pageNumber="15" type="distribution">
<paragraph id="8BC636FAAA50775BFF40B27016E02186" blockId="14.[151,1437,152,912]" pageId="14" pageNumber="15">
<emphasis id="B90DEAE8AA50775BFF40B27014782134" bold="true" box="[198,361,344,370]" pageId="14" pageNumber="15">Distribution:</emphasis>
Pacific lowlands of
<collectingCountry id="F36E766AAA50775BFDDDB27117DE2135" box="[603,719,345,371]" name="Colombia" pageId="14" pageNumber="15">Colombia</collectingCountry>
from Nuqui (
<geoCoordinate id="EE4D503DAA50775BFCF7B27116C62135" box="[881,983,345,371]" direction="north" orientation="latitude" pageId="14" pageNumber="15" precision="925" value="5.7166667">05º43N</geoCoordinate>
,
<geoCoordinate id="EE4D503DAA50775BFC65B271115C2135" box="[995,1101,345,371]" direction="west" orientation="longitude" pageId="14" pageNumber="15" precision="925" value="-77.26667">77º16W</geoCoordinate>
) west of Condoto (
<geoCoordinate id="EE4D503DAA50775BFAB7B27110862135" box="[1329,1431,345,371]" direction="north" orientation="latitude" pageId="14" pageNumber="15" precision="925" value="5.133333">05º08N</geoCoordinate>
,
<geoCoordinate id="EE4D503DAA50775BFF11B2A8141321DC" box="[151,258,384,410]" direction="west" orientation="longitude" pageId="14" pageNumber="15" precision="925" value="-76.65">76º39W</geoCoordinate>
) in the department of Chocó south to Barbacoas (
<geoCoordinate id="EE4D503DAA50775BFCD1B2A816AE21DC" box="[855,959,384,410]" direction="north" orientation="latitude" pageId="14" pageNumber="15" precision="925" value="1.65">01º39N</geoCoordinate>
,
<geoCoordinate id="EE4D503DAA50775BFC4AB2A8112621DC" box="[972,1079,384,410]" direction="west" orientation="longitude" pageId="14" pageNumber="15" precision="925" value="-78.166664">78º10W</geoCoordinate>
) in the department of Nariño.
<taxonomicName id="4C794D79AA50775BFF11B28F14692186" box="[151,376,423,448]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="iridescens">
<emphasis id="B90DEAE8AA50775BFF11B28F14692186" box="[151,376,423,448]" italics="true" pageId="14" pageNumber="15">Atractus iridescens</emphasis>
</taxonomicName>
inhabits rainforest at elevations of
<quantity id="4C819B1FAA50775BFC93B28E169A2186" box="[789,907,422,448]" metricMagnitude="2" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="14" pageNumber="15" unit="m" value="100.0" valueMax="150.0" valueMin="50.0">50150 m</quantity>
(
<figureCitation id="13422A7FAA50775BFC1DB28E16F22186" box="[923,995,422,448]" captionStart="FIGURE 6" captionStartId="9.[151,254,1797,1821]" captionTargetBox="[153,1428,904,1773]" captionTargetId="figure@9.[151,1436,890,1812]" captionTargetPageId="9" captionText="FIGURE 6. Geographical distribution of the Atractus paucidens species group." httpUri="https://zenodo.org/record/191482/files/figure.png" pageId="14" pageNumber="15">Fig. 6</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3636571AA50775BFF40B2E5160E23D6" pageId="14" pageNumber="15" type="discussion">
<paragraph id="8BC636FAAA50775BFF40B2E5160E23D6" blockId="14.[151,1437,152,912]" pageId="14" pageNumber="15">
<emphasis id="B90DEAE8AA50775BFF40B2E5145321A1" bold="true" box="[198,322,461,487]" pageId="14" pageNumber="15">Remarks:</emphasis>
<bibRefCitation id="EFE84B0BAA50775BFECDB2E514EC21A1" author="Peracca" box="[331,509,461,487]" pageId="14" pageNumber="33" refString="Peracca, M. G. (1896) Sopra alcuni ofidii nuovi o poco noti dell'America meridionale. Bolletin Museo di Torino, 11, 1 - 12." type="journal article" year="1896">Peracca (1896)</bibRefCitation>
described
<taxonomicName id="4C794D79AA50775BFD06B2E5160B21A0" box="[640,794,461,486]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="iridescens">
<emphasis id="B90DEAE8AA50775BFD06B2E5160B21A0" box="[640,794,461,486]" italics="true" pageId="14" pageNumber="15">A. iridescens</emphasis>
</taxonomicName>
based on a specimen from “
<collectingCountry id="F36E766AAA50775BFBF6B2E511C921A1" box="[1136,1240,461,487]" name="United States of America" pageId="14" pageNumber="15">America</collectingCountry>
Meridionali” (= South
<collectingCountry id="F36E766AAA50775BFF62B2DC145A2248" box="[228,331,500,526]" name="United States of America" pageId="14" pageNumber="15">America</collectingCountry>
) without precise collection data.
<bibRefCitation id="EFE84B0BAA50775BFD52B2DC16B72248" author="Boulenger" box="[724,934,500,526]" pageId="14" pageNumber="31" refString="Boulenger, G. A. (1913) A collection of batrachians and reptiles made by Dr. H. G. F. Spurrell, F. Z. S., in the Choco. Colombia. Proceedings Zollogical Society, London, 1913, 1019 - 1038." type="proceedings paper" year="1913">Boulenger (1913)</bibRefCitation>
associated the name with three specimens collected at Peña Lisa, Condoto in the department of Chocó,
<collectingCountry id="F36E766AAA50775BFCEFB13216F02272" box="[873,993,538,564]" name="Colombia" pageId="14" pageNumber="15">Colombia</collectingCountry>
. Analyses of data and photographs of the
<typeStatus id="54C28858AA50775BFF40B1691422221D" box="[198,307,577,603]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C794D79AA50775BFED9B1691713221C" box="[351,514,577,602]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="iridescens">
<emphasis id="B90DEAE8AA50775BFED9B1691713221C" box="[351,514,577,602]" italics="true" pageId="14" pageNumber="15">A. iridescens</emphasis>
</taxonomicName>
corroborate Boulengers identification. We report herein three additional specimens extending the species distribution about
<quantity id="4C819B1FAA50775BFD7FB140164322C4" box="[761,850,616,642]" metricMagnitude="5" metricUnit="m" metricValue="5.0" pageId="14" pageNumber="15" unit="km" value="500.0">500 km</quantity>
south. We found another specimen (CSJ 563) that may be referable to
<taxonomicName id="4C794D79AA50775BFE03B1A7170E22EE" box="[389,543,655,680]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="iridescens">
<emphasis id="B90DEAE8AA50775BFE03B1A7170E22EE" box="[389,543,655,680]" italics="true" pageId="14" pageNumber="15">A. iridescens</emphasis>
</taxonomicName>
, but it is labelled as coming from “
<collectingCountry id="F36E766AAA50775BFC45B1A6115522EE" box="[963,1092,654,680]" name="Saint Pierre and Miquelon" pageId="14" pageNumber="15">San Pedro</collectingCountry>
”, a municipality on the other side of the Andes. There are at least five localities named
<collectingCountry id="F36E766AAA50775BFC06B19D11102289" box="[896,1025,693,719]" name="Saint Pierre and Miquelon" pageId="14" pageNumber="15">San Pedro</collectingCountry>
in the department of Antioquia,
<collectingCountry id="F36E766AAA50775BFF11B1F4141F22B0" box="[151,270,732,758]" name="Colombia" pageId="14" pageNumber="15">Colombia</collectingCountry>
, and it is very likely that the specimen came from
<collectingCountry id="F36E766AAA50775BFCE4B1F416CB22B0" box="[866,986,732,758]" name="Saint Pierre and Miquelon" pageId="14" pageNumber="15">San Pedro</collectingCountry>
de Urabá (
<geoCoordinate id="EE4D503DAA50775BFBD8B1F411D422B0" box="[1118,1221,732,758]" direction="north" orientation="latitude" pageId="14" pageNumber="15" precision="925" value="8.283334">08º17N</geoCoordinate>
,
<geoCoordinate id="EE4D503DAA50775BFB57B1F4102B22B0" box="[1233,1338,732,758]" direction="west" orientation="longitude" pageId="14" pageNumber="15" precision="925" value="-76.38333">76º23W</geoCoordinate>
, ca.
<quantity id="4C819B1FAA50775BFAF6B1F415A3235A" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="15" unit="m" value="200.0">200 m</quantity>
), on the northern portion of the Pacific lowlands of
<collectingCountry id="F36E766AAA50775BFCB0B02A16A3235A" box="[822,946,770,796]" name="Colombia" pageId="14" pageNumber="15">Colombia</collectingCountry>
. If this record would be considered, the species range would extend
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north from Nuqui. However, the colour pattern of this specimen is poorly preserved and its identity cannot be properly established, thus we prefer not to include this doubtful record of
<taxonomicName id="4C794D79AA50775BFF11B05F143F23D6" box="[151,302,887,912]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="iridescens">
<emphasis id="B90DEAE8AA50775BFF11B05F143F23D6" box="[151,302,887,912]" italics="true" pageId="14" pageNumber="15">A. iridescens</emphasis>
</taxonomicName>
until further specimens become available.
</paragraph>
</subSubSection>
</treatment>
</document>