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<document id="94248B373F651C73D8D6273EE2527CD8" ID-CLB-Dataset="5365" ID-DOI="10.11646/zootaxa.3812.1.1" ID-GBIF-Dataset="fd5ba21e-09ce-4ac6-b84f-56632ed93917" ID-ISSN="1175-5326" ID-Zenodo-Dep="4919079" ID-ZooBank="7847D731-9F66-4856-A79F-9435FED25B1D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623268636583" checkinUser="felipe" docAuthor="Murányi, Dávid, Gamboa, Maribet &amp; Orci, Kirill Márk" docDate="2014" docId="03F99336FF2CFFC81BE4586AD770FBAC" docLanguage="en" docName="zt03812p082.pdf" docOrigin="Zootaxa 3812 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Zwicknia acuta Muranyi &amp; Orci 2014, sp. n." docType="treatment" docVersion="8" lastPageNumber="40" masterDocId="FFC0EB4EFF08FFEF1B735B43D539FFDD" masterDocTitle="Zwicknia gen. n., a new genus for the Capnia bifrons species group, with descriptions of three new species based on morphology, drumming signals and molecular genetics, and a synopsis of the West Palaearctic and Nearctic genera of Capniidae (Plecoptera)" masterLastPageNumber="82" masterPageNumber="1" pageNumber="37" updateTime="1699007786927" updateUser="ExternalLinkService">
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<mods:title id="A1C0D4F6DE423B1C7C54B92C49AA75E3">Zwicknia gen. n., a new genus for the Capnia bifrons species group, with descriptions of three new species based on morphology, drumming signals and molecular genetics, and a synopsis of the West Palaearctic and Nearctic genera of Capniidae (Plecoptera)</mods:title>
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<mods:namePart id="E28A8A3730DE36FAE2CAFE61B6F25F40">Orci, Kirill Márk</mods:namePart>
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<taxonomicName id="4C5059A3FF2CFFCB1BE4586AD711FC9E" ID-CoL="7GGKR" authority="Muranyi &amp; Orci" authorityName="Muranyi &amp; Orci" authorityYear="2014" box="[151,552,809,835]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="acuta" status="sp. nov.">
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Murányi &amp; Orci
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(
<figureCitation id="136B3EA5FF2CFFCB1BEC580FD439FCB8" box="[159,256,844,869]" captionStart="FIGURES 58–61" captionStartId="31.[151,264,1086,1109]" captionTargetBox="[151,1436,193,1065]" captionTargetId="figure-492@31.[151,1436,193,1065]" captionTargetPageId="31" captionText="FIGURES 58–61. Micropterous wings of Zwicknia Murányi, gen. n. males—58: Z. bifrons (Newman, 1838); 59: Z. acuta Murányi &amp; Orci, sp. n.; 60: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 61: Z. sevanica (Zhiltzova, 1964)—scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919108" httpUri="https://zenodo.org/record/4919108/files/figure.png" pageId="36" pageNumber="37">Figs. 59</figureCitation>
, 
<figureCitation id="136B3EA5FF2CFFCB1A7D580FD415FCB8" box="[270,300,844,869]" captionStart="FIGURES 62–65" captionStartId="33.[151,264,1508,1531]" captionTargetBox="[151,1436,231,1487]" captionTargetId="figure-310@33.[151,1436,193,1487]" captionTargetPageId="33" captionText="FIGURES 62–65. Female genitalia of Zwicknia Murányi, gen. n.—62–63: Z. bifrons (Newman, 1838); 64: Z. acuta Murányi &amp; Orci, sp. n.; 65: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 62: terminalia, ventral view; 63: inner genital organs, dorsal view; 64–65: subgenital plates, ventral view (gc: genital cavity; go: genital opening; Sg: subgenital plate)—scales 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919112" httpUri="https://zenodo.org/record/4919112/files/figure.png" pageId="36" pageNumber="37">64</figureCitation>
, 
<figureCitation id="136B3EA5FF2CFFCB1A4F580FD488FCB8" box="[316,433,844,869]" captionStart="FIGURES 66–71" captionStartId="34.[151,264,1663,1686]" captionTargetBox="[151,1436,326,1642]" captionTargetId="figure-254@34.[151,1436,326,1642]" captionTargetPageId="34" captionText="FIGURES 66–71. Larva of Zwicknia Murányi, gen. n.—66: Z. bifrons (Newman, 1838); 67-69, 71: Z. acuta Murányi &amp; Orci, sp. n.; 70: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 66: head, dorso-lateral view; 67: mouthparts, ventral view; 68: pro- and mesothorax, ventral view; 69: meso- and metathorax, ventral view; 70: Tergite 5, dorsal view; 71: female terminalia, ventral view (Es: ecdysial suture; Etn: eutrochantin; Gl: glossa; Lb: labrum; Lp: labial palp; Me: mentum; MeFs: mesothoracic furcasternum; MeFsa: mesothoracic furcasternal arm; MeFsp: mesothoracic furcasternal pit; MeTn: mesothoracic trochantin; MFs: metathoracic furcasternum; Mp: maxillary palp; MTn: metathoracic trochantin; Oc: frontal ocellus; Pc: pedicell; PFs: prothoracic furcasternum; Pg: paraglossa; Pp: paraproct; St 8: Sternite 8)—scales 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919114" httpUri="https://zenodo.org/record/4919114/files/figure.png" pageId="36" pageNumber="37">67–69, 71</figureCitation>
, 
<figureCitation id="136B3EA5FF2CFFCB1AB3580FD4E7FCBE" box="[448,478,844,867]" captionStart="FIGURES 72–77" captionStartId="35.[151,264,1828,1851]" captionTargetBox="[151,1436,483,1807]" captionTargetId="figure-202@35.[151,1436,483,1807]" captionTargetPageId="35" captionText="FIGURES 72–77. Larva of Zwicknia Murányi, gen. n.—72, 74, 76: Z. bifrons (Newman, 1838); 73, 75: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 77: Z. acuta Murányi &amp; Orci, sp. n.; 72-73: hind femur, lateral view; 74-75: male terminalia, lateral view; 76-77: 11-13 cercal segments, lateral view (Pp: paraproct; Tg 10: tergite 10)— scales 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919116" httpUri="https://zenodo.org/record/4919116/files/figure.png" pageId="36" pageNumber="37">77</figureCitation>
, 
<figureCitation id="136B3EA5FF2CFFCB1A9E580FD70CFCB8" box="[493,565,844,869]" captionStart="FIGURES 78–81" captionStartId="38.[151,264,1941,1964]" captionTargetBox="[226,1366,264,1921]" captionTargetId="figure-99@38.[213,1381,249,1934]" captionTargetPageId="38" captionText="FIGURES 78–81. Male terminalia of Zwicknia acuta Murányi &amp; Orci, sp. n.; paratype, Slovakia, Banskobystrický Region, Krupinská Planina, Pôtor—78: dorsal view; 79: ventral view; 80: lateral view; 81: caudal view (Fp: fusion plate; Pp: paraproct)—scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919118" httpUri="https://zenodo.org/record/4919118/files/figure.png" pageId="36" pageNumber="37">78–81</figureCitation>
, 
<figureCitation id="136B3EA5FF2CFFCB1937580FD790FCB8" box="[580,681,844,869]" captionStart="FIGURES 106–109" captionStartId="54.[151,264,1087,1110]" captionTargetBox="[151,1436,193,1067]" captionTargetId="figure-252@54.[151,1436,193,1067]" captionTargetPageId="54" captionText="FIGURES 106–109. Lateral view of male epiproct of Zwicknia Murányi, gen. n.—106: Z. bifrons (Newman, 1838); 107: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 108–109: Z. acuta Murányi &amp; Orci, sp. n.—scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919136" httpUri="https://zenodo.org/record/4919136/files/figure.png" pageId="36" pageNumber="37">108–109</figureCitation>
, 
<figureCitation id="136B3EA5FF2CFFCB19C4580FD7DAFCB8" box="[695,739,844,869]" captionStart="FIGURES 114–119" captionStartId="56.[151,265,1540,1563]" captionTargetBox="[151,1436,193,1520]" captionTargetId="figure-203@56.[151,1436,193,1520]" captionTargetPageId="56" captionText="FIGURES 114–119. Dorsal view of male epiproct of Zwicknia Murányi, gen. n.—114: Z. bifrons (Newman, 1838); 115, 117: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 116: Z. kovacsi Murányi &amp; Gamboa, sp. n.; 118: Z. sevanica (Zhiltzova, 1964); 119: Z. acuta Murányi &amp; Orci, sp. n.—scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919140" httpUri="https://zenodo.org/record/4919140/files/figure.png" pageId="36" pageNumber="37">119</figureCitation>
, 
<figureCitation id="136B3EA5FF2CFFCB1982580FD624FCB8" box="[753,797,844,869]" captionStart="FIGURES 120–123" captionStartId="57.[151,264,1761,1784]" captionTargetBox="[165,1401,468,1695]" captionTargetId="figure-157@57.[151,1436,457,1720]" captionTargetPageId="57" captionText="FIGURES 120–123. Male terminalia of Zwicknia Murányi, gen. n., dorso-caudal view—120: Z. bifrons (Newman, 1838); 121: Z. acuta Murányi &amp; Orci, sp. n.; 122: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 123: Z. kovacsi Murányi &amp; Gamboa, sp. n.—scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919142" httpUri="https://zenodo.org/record/4919142/files/figure.png" pageId="36" pageNumber="37">121</figureCitation>
, 
<figureCitation id="136B3EA5FF2CFFCB185F580FD6A8FCB8" box="[812,913,844,869]" captionStart="FIGURES 127–147" captionStartId="59.[151,264,1277,1300]" captionTargetBox="[154,1418,219,1236]" captionTargetId="figure-173@59.[151,1436,193,1256]" captionTargetPageId="59" captionText="FIGURES 127–147. Male epiproct of Zwicknia Murányi, gen. n.—127–130 Z. acuta Murányi &amp; Orci, sp. n.; 131–132: Z. kovacsi Murányi &amp; Gamboa, sp. n.; 133–138: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 139–143: Z. bifrons (Newman, 1838); 144–145: Z. sevanica (Zhiltzova, 1964); 146: Z. tuberculata (Zhiltzova, 1964); 147: Z. turkestanica turkestanica (Kimmins, 1950); 127–131, 133, 135, 137, 139–147: lateral view; 132, 134, 136, 138: dorsal view (127: Slovakia, Krupinská Planina, Pôtor; 128: Hungary, Zemplén Mts., Kishuta; 129: Serbia, Fruska Gora, Vrdnik; 130: Serbia, Zlatibor Mts., Crni Rzav; 131–132: Romania, Rodna Mts., Borşa–Staţiunea Borşa; 133–134, 137–138: Croatia, Ivanščica Mts., Stari Golubovec; 135–136: Hungary, Mecsek Mts., Komló-Zobákpuszta; 139: Hungary, Börzsöny Mts., Szokolya-Királyrét; 140: Hungary, Mátra Mts., Gyöngyössolymos; 141: Hungary, Bükk Mts., Nagyvisnyó; 142: Hungary, Bakony Mts., Bakonybél; 143: Serbia, Maljen Mts., Brajkovići; 144–145: Turkey, Akyarma Pass; 146: Kazakhstan, Karatau Mts., Bajaldyrtal; 147: Kazakhstan, Tarbagatai Mts., Ayagos)—scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919146" httpUri="https://zenodo.org/record/4919146/files/figure.png" pageId="36" pageNumber="37">127–130</figureCitation>
, 
<figureCitation id="136B3EA5FF2CFFCB18EC580FD13AFCB8" box="[927,1027,844,869]" captionStart="FIGURES 148–164" captionStartId="60.[151,264,1223,1246]" captionTargetBox="[166,1423,196,1203]" captionTargetId="figure-337@60.[151,1436,181,1215]" captionTargetPageId="60" captionText="FIGURES 148–164. Humps of male Tergite 9 of Zwicknia Murányi, gen. n., caudal view—148–151: Z. acuta Murányi &amp; Orci, sp. n.; 152–156: Z. bifrons (Newman, 1838); 157–159: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 160: Z. kovacsi Murányi &amp; Gamboa, sp. n.; 161–162: Z. sevanica (Zhiltzova, 1964); 163: Z. tuberculata (Zhiltzova, 1964); 164: Z. turkestanica turkestanica (Kimmins, 1950) (148: Slovakia, Krupinská Planina, Pôtor; 149: Hungary, Zemplén Mts., Kishuta; 150: Serbia, Fruska Gora, Vrdnik; 151: Serbia, Zlatibor Mts., Crni Rzav; 152: Hungary, Börzsöny Mts., Szokolya-Királyrét; 153: Hungary, Mátra Mts., Gyöngyössolymos; 154: Hungary, Bükk Mts., Nagyvisnyó; 155: Hungary, Bakony Mts., Bakonybél; 156: Serbia, Maljen Mts., Brajkovići; 157, 159: Croatia, Ivanščica Mts., Stari Golubovec; 158: Hungary, Mecsek Mts., Komló-Zobákpuszta; 160: Romania, Rodna Mts., Borşa-Staţiunea Borşa; 161–162: Turkey, Akyarma Pass; 163: Kazakhstan, Karatau Mts., Bajaldyrtal; 164: Kazakhstan, Tarbagatai Mts., Ayagos)—scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919148" httpUri="https://zenodo.org/record/4919148/files/figure.png" pageId="36" pageNumber="37">148–151</figureCitation>
, 
<figureCitation id="136B3EA5FF2CFFCB1F61580FD107FCB8" box="[1042,1086,844,869]" captionStart="FIGURES 165–168" captionStartId="61.[151,264,1330,1353]" captionTargetBox="[162,1428,857,1282]" captionTargetId="figure-628@61.[151,1436,850,1309]" captionTargetPageId="61" captionText="FIGURES 165–168. Slow oscillograms of male calls of four Zwicknia Murányi, gen. n. species, showing call sequences versus sporadic call production—165: two call sequence of Z. bifrons (Newman, 1838), &quot;//&quot; indicates where 2 minutes 23 seconds of silence was cut out from the recording; 166: three call sequences of Z. acuta Murányi &amp; Orci, sp. n.; 167: sporadic calls of Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 168: sporadic call production in Z. kovacsi Murányi &amp; Gamboa, sp. n.—time scale is the same for the four oscillograms; for detailed data on the animals and recording circumstances, see Table 9." figureDoi="http://doi.org/10.5281/zenodo.4919154" httpUri="https://zenodo.org/record/4919154/files/figure.png" pageId="36" pageNumber="37">166</figureCitation>
, 
<figureCitation id="136B3EA5FF2CFFCB1F3E580FD18BFCB8" box="[1101,1202,844,869]" captionStart="FIGURES 169–184" captionStartId="62.[151,264,1845,1868]" captionTargetBox="[201,1412,198,1820]" captionTargetId="figure-140@62.[168,1412,183,1822]" captionTargetPageId="62" captionText="FIGURES 169–184. Male drumming calls of Zwicknia Murányi, gen. n.—169–173: Z. bifrons (Newman, 1838); 174–178: Z. acuta Murányi &amp; Orci, sp. n.; 179–182: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 183–184: Z. kovacsi Murányi &amp; Gamboa, sp. n. (see detailed data on the animals and recording circumstances in Table 9)—Oscillograms in Figs. 169–179, 181, 183 have the same time axis resolution (time scale bar is presented in Fig. 169 for all of them), an other time scale is used in the faster oscillograms of Figs. 180, 182, 184 (time scale bar for them can be found in Fig. 184), those are zoomed sections of Figs. 179, 181, 183, respectively." figureDoi="http://doi.org/10.5281/zenodo.4919156" httpUri="https://zenodo.org/record/4919156/files/figure.png" pageId="36" pageNumber="37">174–178</figureCitation>
, 
<figureCitation id="136B3EA5FF2CFFCB1FB3580FD01CFCB8" box="[1216,1317,844,869]" captionStart="FIGURES 185–189" captionStartId="63.[151,264,1012,1035]" captionTargetBox="[166,1430,178,988]" captionTargetId="figure-458@63.[151,1436,178,1004]" captionTargetPageId="63" captionText="FIGURES 185–189. Male-female drumming duets of three Zwicknia Murányi, gen. n. species.—185: Z. bifrons (Newman, 1838) from Hungary, Pilis Mts., Bükkös Stream, recorded at 16.9 °C; 186–187: Z. acuta Murányi &amp; Orci, sp. n. from Slovakia, Krupinská Planina, Stará Rieka, recorded at 18.2 °C; 188: call-answer duet in Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n. from Hungary, Mecsek Mts., Völgységi Stream, recorded at 18.2 °C; 189: call-answer-reply duet in Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n. from Hungary, Mecsek Mts., Völgységi Stream, recorded at 17.9 °C—arrows with a gender sign point to the beginning of a section of the oscillogram containing the beats of that gender." figureDoi="http://doi.org/10.5281/zenodo.4919158" httpUri="https://zenodo.org/record/4919158/files/figure.png" pageId="36" pageNumber="37">186–187</figureCitation>
, 
<figureCitation id="136B3EA5FF2CFFCB1E40580FD0A1FCB8" box="[1331,1432,844,869]" captionStart-0="FIGURE 190" captionStart-1="FIGURE 191" captionStart-2="FIGURE 192" captionStart-3="FIGURES 193–195" captionStartId-0="64.[151,250,1125,1147]" captionStartId-1="65.[151,250,1357,1379]" captionStartId-2="66.[151,249,1180,1202]" captionStartId-3="67.[151,264,1829,1852]" captionTargetBox-0="[163,1430,185,1044]" captionTargetBox-1="[164,1416,208,1328]" captionTargetBox-2="[156,1424,193,1155]" captionTargetBox-3="[199,1367,231,1794]" captionTargetId-0="figure-392@64.[151,1436,177,1058]" captionTargetId-1="figure-231@65.[151,1436,193,1336]" captionTargetId-2="figure-240@66.[151,1433,185,1171]" captionTargetId-3="figure-90@67.[199,1367,231,1810]" captionTargetPageId-0="64" captionTargetPageId-1="65" captionTargetPageId-2="66" captionTargetPageId-3="67" captionText-0="FIGURE 190. Inter-beat interval variation during male calls in the four examined Zwicknia Murányi, gen. n. species.—Box plot colors: white—Z. bifrons (Newman, 1838), red—Z. acuta Murányi &amp; Orci, sp. n., blue—Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n. In Z. kovacsi Murányi &amp; Gamboa, sp. n. we have drumming samples only from two specimens therefore in the case of that species two simple mark series are used to show inter-beat intervals in their calls (circles for specimen 1, and triangles for specimen 2). i1, i2, i3 and so on are labels containing the serial number of inter-beat intervals counting them from the beginning of each call. Box plots show the duration values measured in 7 specimens in Z. bifrons and Z. acuta, and 5 specimens in Z. rupprechti. Vertical axis has a logarithmic scale. Only calls recorded at an ambient air temperature of 17–19 oC were used. (see the measurement data in Appendix Tables 1–4)." captionText-1="FIGURE 191. A scatter plot showing the clear separation of male calls of Zwicknia Murányi, gen. n. along two drumming signal characters—Z. bifrons (Newman, 1838): triangles; Z. acuta Murányi &amp; Orci, sp. n.: circles; Z. kovacsi Murányi &amp; Gamboa, sp. n.: +; Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.: X—Each mark represents the mean values of the characters for a given specimen. Note that both axes have logarithmic scale. Air temperature varied between 15–21 oC in every species (for details see Tables 6–8)." captionText-2="FIGURE 192. Phylogenetic tree of cox1 including Zwicknia Murányi, gen. n. with our accession codes (300941-301003), and GenBank closest species (with alphanumeric accession codes) as outgroup. MP and ML had same topology. Numbers on braches are ML/MP bootstrap values&gt;50%. Species names right to the tree refer to morphological species. P-Distance table is showed." captionText-3="FIGURES 193–195. Type localities of the new species described in Zwicknia Murányi, gen. n.—193: Zwicknia acuta Murányi &amp; Orci, sp. n.: Stará rieka, Slovakia, Banskobystrický Region, Krupinská Planina, Pôtor; 194: Zwicknia kovacsi Murányi &amp; Gamboa, sp. n.: Cimpoies Stream, Romania, Maramureş County, Rodna Mts., Borşa-Staţiunea Borşa; 195: Zwicknia rupprechti Murányi, Orci &amp; Gamboa, sp. n.: Völgységi Stream, Hungary, Baranya County, Mecsek Mts., Komló-Zobákpuszta." figureDoi-0="http://doi.org/10.5281/zenodo.4919160" figureDoi-1="http://doi.org/10.5281/zenodo.4919162" figureDoi-2="http://doi.org/10.5281/zenodo.4919164" figureDoi-3="http://doi.org/10.5281/zenodo.4919166" httpUri-0="https://zenodo.org/record/4919160/files/figure.png" httpUri-1="https://zenodo.org/record/4919162/files/figure.png" httpUri-2="https://zenodo.org/record/4919164/files/figure.png" httpUri-3="https://zenodo.org/record/4919166/files/figure.png" pageId="36" pageNumber="37">190–193</figureCitation>
, 
<figureCitation id="136B3EA5FF2CFFCB1BE4582CD5C4FC55" box="[151,253,879,904]" captionStart="FIGURES 196–197" captionStartId="68.[151,264,1908,1931]" captionTargetBox="[188,1393,187,1898]" captionTargetId="figure-93@68.[188,1393,187,1898]" captionTargetPageId="68" captionText="FIGURES 196–197. Distribution of Zwicknia Murányi, gen. n.—196: global distribution; 197: studied populations of the Carpathian Basin and the northern Balkans—in Fig. 196, white dot on Z. bifrons symbols indicate populations studied by Rupprecht (1997), dotted lines indicate limits of s.l. Z. bifrons literature reports; in Fig. 197, white dot on symbols indicate populations where drumming signals were studied by us, white star indicate type localities." figureDoi="http://doi.org/10.5281/zenodo.4919168" httpUri="https://zenodo.org/record/4919168/files/figure.png" pageId="36" pageNumber="37">196–197</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34A71ABFF2CFFCB1BE458F2D43FFBFE" pageId="36" pageNumber="37" type="reference_group">
<paragraph id="8BEF2220FF2CFFCB1BE458F2D4A3FC3B" blockId="36.[151,1437,945,1059]" pageId="36" pageNumber="37">
<taxonomicName id="4C5059A3FF2CFFCB1BE458F2D495FC1A" ID-CoL="5X9KQ" authority="(Pictet, 1833)" authorityName="sensu Morton" authorityYear="1896" baseAuthorityName="Pictet" baseAuthorityYear="1833" box="[151,428,945,967]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="nigra">
<emphasis id="B924FE32FF2CFFCB1BE458F2D426FC1A" box="[151,287,945,967]" italics="true" pageId="36" pageNumber="37">Capnia nigra</emphasis>
(
<bibRefCitation id="EFC15FD1FF2CFFCB1A5F58F2D49FFC1A" author="Pictet, F. J." box="[300,422,945,967]" pageId="36" pageNumber="37" pagination="44 - 65" refId="ref44601" refString="Pictet, F. J. (1833) Memoire sur les Metamorphoses des Perles. Annales des Sciences Naturales, 28, 44 - 65." type="journal article" year="1833">Pictet, 1833</bibRefCitation>
)
</taxonomicName>
—figures in 
<bibRefCitation id="EFC15FD1FF2CFFCB194358F2D7FCFC1A" author="Klapalek, F." box="[560,709,945,967]" pageId="36" pageNumber="37" pagination="683 - 738" refId="ref43152" refString="Klapalek, F. (1896) Uber die Geschlechtstheile der Plecopteren, mit besonderer Rucksicht auf die Morphologie der Genitalanhange. Sitzungsberichten der kaiserlichen Akademie der Wissenschaften in Wien, 105, 683 - 738." type="journal article" year="1896">Klapálek 1896</bibRefCitation>
and 
<bibRefCitation id="EFC15FD1FF2CFFCB198458F1D645FC1A" author="Despax, R." box="[759,892,945,967]" pageId="36" pageNumber="37" pagination="1 - 280" refId="ref41539" refString="Despax, R. (1951) Plecopteres. Faune de France, 55, 1 - 280." type="journal article" year="1951">Despax 1951</bibRefCitation>
(enumerated under the general synonymies) possibly referring to 
<emphasis id="B924FE32FF2CFFCB1A335892D4A3FC3B" box="[320,410,977,998]" italics="true" pageId="36" pageNumber="37">
<taxonomicName id="4C5059A3FF2CFFCB1A335892D4AFFC3B" ID-CoL="7GGKR" authorityName="Muranyi &amp; Orci" authorityYear="2014" box="[320,406,977,998]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="acuta">Z. acuta</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8BEF2220FF2CFFCB1BE458ADD43FFBFE" blockId="36.[151,1437,945,1059]" pageId="36" pageNumber="37">
<taxonomicName id="4C5059A3FF2CFFCB1BE458ADD4E4FBD9" ID-CoL="QQPJ" authority="(Newman, 1838)" authorityName="Capnor" baseAuthorityName="Newman" baseAuthorityYear="1838" box="[151,477,1006,1028]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="bifrons">
<emphasis id="B924FE32FF2CFFCB1BE458ADD416FBD9" box="[151,303,1006,1028]" italics="true" pageId="36" pageNumber="37">Capnia bifrons</emphasis>
(
<bibRefCitation id="EFC15FD1FF2CFFCB1A4F58ADD4EFFBD9" author="Newman, E." box="[316,470,1006,1028]" pageId="36" pageNumber="37" pagination="372 - 402" refId="ref44180" refString="Newman, E. (1838) Entomological notes. The Entomological Magazine, 5 (4), 372 - 402." type="journal article" year="1838">Newman, 1838</bibRefCitation>
)
</taxonomicName>
—figures in 
<bibRefCitation id="EFC15FD1FF2CFFCB191358ADD7D2FBD9" author="Winkler, O." box="[608,747,1006,1028]" pageId="36" pageNumber="37" pagination="1 - 95" refId="ref46090" refString="Winkler, O. (1957) Plecoptera Slovenska (Faunisticko-systematicka studia). Biologicke Prace, 3 (7), 1 - 95." type="journal article" year="1957">Winkler 1957</bibRefCitation>
(enumerated under the general synonymies) possibly referring to 
<emphasis id="B924FE32FF2CFFCB1EF458ACD43FFBFE" italics="true" pageId="36" pageNumber="37">
<taxonomicName id="4C5059A3FF2CFFCB1EF458ACD43AFBFE" ID-CoL="7GGKR" authorityName="Muranyi &amp; Orci" authorityYear="2014" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="acuta">Z. acuta</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C34A71ABFF2CFFCB1BE45F0AD4BCFA7C" pageId="36" pageNumber="37" type="diagnosis">
<paragraph id="8BEF2220FF2CFFCB1BE45F0AD4BCFA7C" blockId="36.[151,1437,1097,2005]" pageId="36" pageNumber="37">
<emphasis id="B924FE32FF2CFFCB1BE45F0AD428FBBF" bold="true" box="[151,273,1097,1122]" pageId="36" pageNumber="37">Diagnosis.</emphasis>
Male epiproct: Ep-scl narrow and pointed in dorsal view, tip straight, long and acute in lateral view; ventral membranous section ends far before the base in lateral view, apical spines stout, distributed also on the apex of Ep-scl. Process of male Tg 9: high, perpendicularly elevated, 2× wider than Ep-scl, rectangular and with indenting sides caudally. Males produce sequences of (2)-3-4-(6) drumming calls (
<figureCitation id="136B3EA5FF2CFFCB1F305FF0D196FB11" box="[1091,1199,1203,1228]" captionStart="FIGURES 165–168" captionStartId="61.[151,264,1330,1353]" captionTargetBox="[162,1428,857,1282]" captionTargetId="figure-628@61.[151,1436,850,1309]" captionTargetPageId="61" captionText="FIGURES 165–168. Slow oscillograms of male calls of four Zwicknia Murányi, gen. n. species, showing call sequences versus sporadic call production—165: two call sequence of Z. bifrons (Newman, 1838), &quot;//&quot; indicates where 2 minutes 23 seconds of silence was cut out from the recording; 166: three call sequences of Z. acuta Murányi &amp; Orci, sp. n.; 167: sporadic calls of Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 168: sporadic call production in Z. kovacsi Murányi &amp; Gamboa, sp. n.—time scale is the same for the four oscillograms; for detailed data on the animals and recording circumstances, see Table 9." figureDoi="http://doi.org/10.5281/zenodo.4919154" httpUri="https://zenodo.org/record/4919154/files/figure.png" pageId="36" pageNumber="37">Figs. 166</figureCitation>
, 
<figureCitation id="136B3EA5FF2CFFCB1FCE5FF0D01DFB11" box="[1213,1316,1203,1228]" captionStart="FIGURES 169–184" captionStartId="62.[151,264,1845,1868]" captionTargetBox="[201,1412,198,1820]" captionTargetId="figure-140@62.[168,1412,183,1822]" captionTargetPageId="62" captionText="FIGURES 169–184. Male drumming calls of Zwicknia Murányi, gen. n.—169–173: Z. bifrons (Newman, 1838); 174–178: Z. acuta Murányi &amp; Orci, sp. n.; 179–182: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 183–184: Z. kovacsi Murányi &amp; Gamboa, sp. n. (see detailed data on the animals and recording circumstances in Table 9)—Oscillograms in Figs. 169–179, 181, 183 have the same time axis resolution (time scale bar is presented in Fig. 169 for all of them), an other time scale is used in the faster oscillograms of Figs. 180, 182, 184 (time scale bar for them can be found in Fig. 184), those are zoomed sections of Figs. 179, 181, 183, respectively." figureDoi="http://doi.org/10.5281/zenodo.4919156" httpUri="https://zenodo.org/record/4919156/files/figure.png" pageId="36" pageNumber="37">174–178</figureCitation>
). Calls are monophasic beat series in which beats are produced with gradually increasing inter-beat intervals (
<figureCitation id="136B3EA5FF2CFFCB1F905F95D005FB32" box="[1251,1340,1238,1263]" captionStart="FIGURE 190" captionStartId="64.[151,250,1125,1147]" captionTargetBox="[163,1430,185,1044]" captionTargetId="figure-392@64.[151,1436,177,1058]" captionTargetPageId="64" captionText="FIGURE 190. Inter-beat interval variation during male calls in the four examined Zwicknia Murányi, gen. n. species.—Box plot colors: white—Z. bifrons (Newman, 1838), red—Z. acuta Murányi &amp; Orci, sp. n., blue—Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n. In Z. kovacsi Murányi &amp; Gamboa, sp. n. we have drumming samples only from two specimens therefore in the case of that species two simple mark series are used to show inter-beat intervals in their calls (circles for specimen 1, and triangles for specimen 2). i1, i2, i3 and so on are labels containing the serial number of inter-beat intervals counting them from the beginning of each call. Box plots show the duration values measured in 7 specimens in Z. bifrons and Z. acuta, and 5 specimens in Z. rupprechti. Vertical axis has a logarithmic scale. Only calls recorded at an ambient air temperature of 17–19 oC were used. (see the measurement data in Appendix Tables 1–4)." figureDoi="http://doi.org/10.5281/zenodo.4919160" httpUri="https://zenodo.org/record/4919160/files/figure.png" pageId="36" pageNumber="37">Fig 190</figureCitation>
, red box plots). Inter beat intervals vary between 70–130 ms. Call duration varies between 350–750 ms and calls contain 4–10 beats. Inter call intervals increase during call sequence and vary between 350–3250 ms. All these parameters refer to signals produced at ambient air temperature between 15–21 
<superScript id="7C258F68FF2CFFCB18C65E7CD685FA90" attach="right" box="[949,956,1343,1357]" fontSize="6" pageId="36" pageNumber="37">o</superScript>
C. Male female drumming duet is a long sequence of male call—female answer alternations, where female answers may overlap the terminal part of male calls (
<figureCitation id="136B3EA5FF2CFFCB1BAA5ECBD441FA7C" box="[217,376,1416,1441]" captionStart="FIGURES 185–189" captionStartId="63.[151,264,1012,1035]" captionTargetBox="[166,1430,178,988]" captionTargetId="figure-458@63.[151,1436,178,1004]" captionTargetPageId="63" captionText="FIGURES 185–189. Male-female drumming duets of three Zwicknia Murányi, gen. n. species.—185: Z. bifrons (Newman, 1838) from Hungary, Pilis Mts., Bükkös Stream, recorded at 16.9 °C; 186–187: Z. acuta Murányi &amp; Orci, sp. n. from Slovakia, Krupinská Planina, Stará Rieka, recorded at 18.2 °C; 188: call-answer duet in Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n. from Hungary, Mecsek Mts., Völgységi Stream, recorded at 18.2 °C; 189: call-answer-reply duet in Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n. from Hungary, Mecsek Mts., Völgységi Stream, recorded at 17.9 °C—arrows with a gender sign point to the beginning of a section of the oscillogram containing the beats of that gender." figureDoi="http://doi.org/10.5281/zenodo.4919158" httpUri="https://zenodo.org/record/4919158/files/figure.png" pageId="36" pageNumber="37">Figs 186–187</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34A71ABFF2CFFCA1BB45EEFD6E2FBB7" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37" type="materials_examined">
<paragraph id="8BEF2220FF2CFFCA1BB45EEFD7B5FD39" blockId="36.[151,1437,1097,2005]" lastBlockId="37.[151,1437,151,2015]" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37">
<emphasis id="B924FE32FF2CFFCB1BB45EEFD44CFA18" bold="true" box="[199,373,1452,1477]" pageId="36" pageNumber="37">Type material.</emphasis>
<typeStatus id="54EB9C82FF2CFFCB1A0E5EEFD4DCFA18" box="[381,485,1452,1477]" pageId="36" pageNumber="37" type="holotype">Holotype</typeStatus>
male: 
<collectingCountry id="F34762B0FF2CFFCB19405EEFD7FAFA18" box="[563,707,1452,1477]" name="Slovakia" pageId="36" pageNumber="37">SLOVAKIA</collectingCountry>
: 
<collectingRegion id="4994ECC2FF2CFFCB19BD5EEFD6DCFA18" box="[718,997,1452,1477]" country="Slovakia" name="Banskobystricky" pageId="36" pageNumber="37">Banskobystrický Region</collectingRegion>
, Krupinská Planina, Pôtor, Stará Rieka Stream, 
<geoCoordinate id="EE6444E7FF2CFFCB1B8A5E93D4AEFA35" box="[249,407,1487,1512]" degrees="48" direction="north" minutes="13.731" orientation="latitude" pageId="36" pageNumber="37" precision="1" value="48.22885">N 48°13.731’</geoCoordinate>
<geoCoordinate id="EE6444E7FF2CFFCB1AD05E93D779FA35" box="[419,576,1487,1512]" degrees="19" direction="east" minutes="24.946" orientation="longitude" pageId="36" pageNumber="37" precision="1" value="19.415766">E 19°24.946’</geoCoordinate>
, 
<quantity id="4CA88FC5FF2CFFCB19215E8CD7A7FA3A" box="[594,670,1487,1512]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="36" pageNumber="37" unit="m" value="200.0">200 m</quantity>
a.s.l., 16.03.201 0, leg. D. 
<taxonomicName id="4C5059A3FF2CFFCB18935E8CD17AFA35" authority="Murányi &amp; Gamboa &amp; Orci, 2014" authorityName="Murányi &amp; Gamboa &amp; Orci" authorityYear="2014" box="[992,1091,1487,1512]" genus="Muranyi" pageId="36" pageNumber="69" rank="genus" status="gen. nov.">Murányi</taxonomicName>
, K. Orci (
<collectionCode id="ED41BAE5FF2CFFCB1FB25E8CD024FA35" box="[1217,1309,1487,1512]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="36" pageNumber="37">HNHM</collectionCode>
: PLP3882; drumming recorded as 2010/No.1). 
<typeStatus id="54EB9C82FF2CFFCB195E5EB0D7A7F9D6" box="[557,670,1523,1547]" pageId="36" pageNumber="37" type="paratype">Paratypes</typeStatus>
: same locality and date: 1f (
<collectionCode id="ED41BAE5FF2CFFCB18AD5EB1D103F9D6" box="[990,1082,1522,1547]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="36" pageNumber="37">HNHM</collectionCode>
: PLP3883; drumming recorded as 2010/No.1, pair of the 
<typeStatus id="54EB9C82FF2CFFCB1AB95D56D708F9F3" box="[458,561,1557,1582]" pageId="36" pageNumber="37" type="holotype">holotype</typeStatus>
), 
<quantity id="4CA88FC5FF2CFFCB19345D56D752F9F3" box="[583,619,1557,1582]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="36" pageNumber="37" unit="m" value="1.0">1m</quantity>
(
<collectionCode id="ED41BAE5FF2CFFCB190D5D56D7E3F9F3" box="[638,730,1557,1582]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="36" pageNumber="37">HNHM</collectionCode>
: PLP3884; drumming recorded as 2010/No.2), 
<quantity id="4CA88FC5FF2CFFCB1E755D56D013F9F3" box="[1286,1322,1557,1582]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="36" pageNumber="37" unit="m" value="1.0">1m</quantity>
(
<collectionCode id="ED41BAE5FF2CFFCB1E4C5D55D0A0F9F0" box="[1343,1433,1558,1581]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="36" pageNumber="37">HNHM</collectionCode>
: PLP3885; drumming recorded as 2010/No.3), 
<quantity id="4CA88FC5FF2CFFCB19EF5D7BD7F9F98C" box="[668,704,1592,1617]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="36" pageNumber="37" unit="m" value="1.0">1m</quantity>
(
<collectionCode id="ED41BAE5FF2CFFCB19BD5D7BD613F98C" box="[718,810,1592,1617]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="36" pageNumber="37">HNHM</collectionCode>
: PLP3886; drumming recorded as 2010/No.4, 
<figureCitation id="136B3EA5FF2CFFCB1E5C5D7BD0B6F98C" box="[1327,1423,1592,1617]" captionStart="FIGURES 169–184" captionStartId="62.[151,264,1845,1868]" captionTargetBox="[201,1412,198,1820]" captionTargetId="figure-140@62.[168,1412,183,1822]" captionTargetPageId="62" captionText="FIGURES 169–184. Male drumming calls of Zwicknia Murányi, gen. n.—169–173: Z. bifrons (Newman, 1838); 174–178: Z. acuta Murányi &amp; Orci, sp. n.; 179–182: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 183–184: Z. kovacsi Murányi &amp; Gamboa, sp. n. (see detailed data on the animals and recording circumstances in Table 9)—Oscillograms in Figs. 169–179, 181, 183 have the same time axis resolution (time scale bar is presented in Fig. 169 for all of them), an other time scale is used in the faster oscillograms of Figs. 180, 182, 184 (time scale bar for them can be found in Fig. 184), those are zoomed sections of Figs. 179, 181, 183, respectively." figureDoi="http://doi.org/10.5281/zenodo.4919156" httpUri="https://zenodo.org/record/4919156/files/figure.png" pageId="36" pageNumber="37">Fig. 175</figureCitation>
), 
<quantity id="4CA88FC5FF2CFFCB1BE45D1FD583F9A8" box="[151,186,1628,1653]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="36" pageNumber="37" unit="m" value="1.0">1m</quantity>
(
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: PLP3887; drumming recorded as 2010/No.5), 
<quantity id="4CA88FC5FF2CFFCB18325D1FD65CF9A8" box="[833,869,1628,1653]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="36" pageNumber="37" unit="m" value="1.0">1m</quantity>
(
<collectionCode id="ED41BAE5FF2CFFCB18065D1FD6E9F9A8" box="[885,976,1628,1653]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="36" pageNumber="37">HNHM</collectionCode>
: PLP3888; drumming recorded as 2010/ No.6), 
<quantity id="4CA88FC5FF2CFFCB1B955D3CD422F945" box="[230,283,1663,1688]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="36" pageNumber="37" unit="m" value="15.0">15m</quantity>
, 1f larva (
<collectionCode id="ED41BAE5FF2CFFCB1AFC5D3CD4D3F945" box="[399,490,1663,1688]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="36" pageNumber="37">HNHM</collectionCode>
: PLP3372), 
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(
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); 24.03.200 6, leg. L. Dányi, J. Kontschán, D. 
<taxonomicName id="4C5059A3FF2CFFCB1E7B5D3CD054F945" authority="Murányi &amp; Gamboa &amp; Orci, 2014" authorityName="Murányi &amp; Gamboa &amp; Orci" authorityYear="2014" box="[1288,1389,1663,1688]" genus="Muranyi" pageId="36" pageNumber="69" rank="genus" status="gen. nov.">Murányi</taxonomicName>
: 
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2f, 1f larva (
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: PLP1832; 
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, 
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terminalia, 
<specimenCount id="9D56E9A9FF2CFFCB18E35DE0D129F966" box="[912,1040,1698,1723]" pageId="36" pageNumber="37" type="female">one female</specimenCount>
and the larva prepared for SEM, specimens used for drawings and photos 
<figureCitation id="136B3EA5FF2CFFCB19005D86D7EBF903" box="[627,722,1733,1758]" captionStart="FIGURES 58–61" captionStartId="31.[151,264,1086,1109]" captionTargetBox="[151,1436,193,1065]" captionTargetId="figure-492@31.[151,1436,193,1065]" captionTargetPageId="31" captionText="FIGURES 58–61. Micropterous wings of Zwicknia Murányi, gen. n. males—58: Z. bifrons (Newman, 1838); 59: Z. acuta Murányi &amp; Orci, sp. n.; 60: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 61: Z. sevanica (Zhiltzova, 1964)—scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919108" httpUri="https://zenodo.org/record/4919108/files/figure.png" pageId="36" pageNumber="37">Figs. 59</figureCitation>
, 
<figureCitation id="136B3EA5FF2CFFCB19935D86D7C7F903" box="[736,766,1733,1758]" captionStart="FIGURES 62–65" captionStartId="33.[151,264,1508,1531]" captionTargetBox="[151,1436,231,1487]" captionTargetId="figure-310@33.[151,1436,193,1487]" captionTargetPageId="33" captionText="FIGURES 62–65. Female genitalia of Zwicknia Murányi, gen. n.—62–63: Z. bifrons (Newman, 1838); 64: Z. acuta Murányi &amp; Orci, sp. n.; 65: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 62: terminalia, ventral view; 63: inner genital organs, dorsal view; 64–65: subgenital plates, ventral view (gc: genital cavity; go: genital opening; Sg: subgenital plate)—scales 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919112" httpUri="https://zenodo.org/record/4919112/files/figure.png" pageId="36" pageNumber="37">64</figureCitation>
, 
<figureCitation id="136B3EA5FF2CFFCB187E5D86D6B8F903" box="[781,897,1733,1758]" captionStart="FIGURES 66–71" captionStartId="34.[151,264,1663,1686]" captionTargetBox="[151,1436,326,1642]" captionTargetId="figure-254@34.[151,1436,326,1642]" captionTargetPageId="34" captionText="FIGURES 66–71. Larva of Zwicknia Murányi, gen. n.—66: Z. bifrons (Newman, 1838); 67-69, 71: Z. acuta Murányi &amp; Orci, sp. n.; 70: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 66: head, dorso-lateral view; 67: mouthparts, ventral view; 68: pro- and mesothorax, ventral view; 69: meso- and metathorax, ventral view; 70: Tergite 5, dorsal view; 71: female terminalia, ventral view (Es: ecdysial suture; Etn: eutrochantin; Gl: glossa; Lb: labrum; Lp: labial palp; Me: mentum; MeFs: mesothoracic furcasternum; MeFsa: mesothoracic furcasternal arm; MeFsp: mesothoracic furcasternal pit; MeTn: mesothoracic trochantin; MFs: metathoracic furcasternum; Mp: maxillary palp; MTn: metathoracic trochantin; Oc: frontal ocellus; Pc: pedicell; PFs: prothoracic furcasternum; Pg: paraglossa; Pp: paraproct; St 8: Sternite 8)—scales 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919114" httpUri="https://zenodo.org/record/4919114/files/figure.png" pageId="36" pageNumber="37">67–69, 71</figureCitation>
, 
<figureCitation id="136B3EA5FF2CFFCB18E35D85D696F900" box="[912,943,1734,1757]" captionStart="FIGURES 72–77" captionStartId="35.[151,264,1828,1851]" captionTargetBox="[151,1436,483,1807]" captionTargetId="figure-202@35.[151,1436,483,1807]" captionTargetPageId="35" captionText="FIGURES 72–77. Larva of Zwicknia Murányi, gen. n.—72, 74, 76: Z. bifrons (Newman, 1838); 73, 75: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 77: Z. acuta Murányi &amp; Orci, sp. n.; 72-73: hind femur, lateral view; 74-75: male terminalia, lateral view; 76-77: 11-13 cercal segments, lateral view (Pp: paraproct; Tg 10: tergite 10)— scales 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919116" httpUri="https://zenodo.org/record/4919116/files/figure.png" pageId="36" pageNumber="37">77</figureCitation>
, 
<figureCitation id="136B3EA5FF2CFFCB18CE5D86D13CF903" box="[957,1029,1733,1758]" captionStart="FIGURES 78–81" captionStartId="38.[151,264,1941,1964]" captionTargetBox="[226,1366,264,1921]" captionTargetId="figure-99@38.[213,1381,249,1934]" captionTargetPageId="38" captionText="FIGURES 78–81. Male terminalia of Zwicknia acuta Murányi &amp; Orci, sp. n.; paratype, Slovakia, Banskobystrický Region, Krupinská Planina, Pôtor—78: dorsal view; 79: ventral view; 80: lateral view; 81: caudal view (Fp: fusion plate; Pp: paraproct)—scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919118" httpUri="https://zenodo.org/record/4919118/files/figure.png" pageId="36" pageNumber="37">78–81</figureCitation>
, 
<figureCitation id="136B3EA5FF2CFFCB1F605D86D141F903" box="[1043,1144,1733,1758]" captionStart="FIGURES 106–109" captionStartId="54.[151,264,1087,1110]" captionTargetBox="[151,1436,193,1067]" captionTargetId="figure-252@54.[151,1436,193,1067]" captionTargetPageId="54" captionText="FIGURES 106–109. Lateral view of male epiproct of Zwicknia Murányi, gen. n.—106: Z. bifrons (Newman, 1838); 107: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 108–109: Z. acuta Murányi &amp; Orci, sp. n.—scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919136" httpUri="https://zenodo.org/record/4919136/files/figure.png" pageId="36" pageNumber="37">108–109</figureCitation>
, 
<figureCitation id="136B3EA5FF2CFFCB1FF55D86D18BF903" box="[1158,1202,1733,1758]" captionStart="FIGURES 114–119" captionStartId="56.[151,265,1540,1563]" captionTargetBox="[151,1436,193,1520]" captionTargetId="figure-203@56.[151,1436,193,1520]" captionTargetPageId="56" captionText="FIGURES 114–119. Dorsal view of male epiproct of Zwicknia Murányi, gen. n.—114: Z. bifrons (Newman, 1838); 115, 117: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 116: Z. kovacsi Murányi &amp; Gamboa, sp. n.; 118: Z. sevanica (Zhiltzova, 1964); 119: Z. acuta Murányi &amp; Orci, sp. n.—scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919140" httpUri="https://zenodo.org/record/4919140/files/figure.png" pageId="36" pageNumber="37">119</figureCitation>
, 
<figureCitation id="136B3EA5FF2CFFCB1FCC5D86D1D2F903" box="[1215,1259,1733,1758]" captionStart="FIGURES 120–123" captionStartId="57.[151,264,1761,1784]" captionTargetBox="[165,1401,468,1695]" captionTargetId="figure-157@57.[151,1436,457,1720]" captionTargetPageId="57" captionText="FIGURES 120–123. Male terminalia of Zwicknia Murányi, gen. n., dorso-caudal view—120: Z. bifrons (Newman, 1838); 121: Z. acuta Murányi &amp; Orci, sp. n.; 122: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 123: Z. kovacsi Murányi &amp; Gamboa, sp. n.—scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919142" httpUri="https://zenodo.org/record/4919142/files/figure.png" pageId="36" pageNumber="37">121</figureCitation>
, 
<figureCitation id="136B3EA5FF2CFFCB1F895D86D01FF903" box="[1274,1318,1733,1758]" captionStart="FIGURES 127–147" captionStartId="59.[151,264,1277,1300]" captionTargetBox="[154,1418,219,1236]" captionTargetId="figure-173@59.[151,1436,193,1256]" captionTargetPageId="59" captionText="FIGURES 127–147. Male epiproct of Zwicknia Murányi, gen. n.—127–130 Z. acuta Murányi &amp; Orci, sp. n.; 131–132: Z. kovacsi Murányi &amp; Gamboa, sp. n.; 133–138: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 139–143: Z. bifrons (Newman, 1838); 144–145: Z. sevanica (Zhiltzova, 1964); 146: Z. tuberculata (Zhiltzova, 1964); 147: Z. turkestanica turkestanica (Kimmins, 1950); 127–131, 133, 135, 137, 139–147: lateral view; 132, 134, 136, 138: dorsal view (127: Slovakia, Krupinská Planina, Pôtor; 128: Hungary, Zemplén Mts., Kishuta; 129: Serbia, Fruska Gora, Vrdnik; 130: Serbia, Zlatibor Mts., Crni Rzav; 131–132: Romania, Rodna Mts., Borşa–Staţiunea Borşa; 133–134, 137–138: Croatia, Ivanščica Mts., Stari Golubovec; 135–136: Hungary, Mecsek Mts., Komló-Zobákpuszta; 139: Hungary, Börzsöny Mts., Szokolya-Királyrét; 140: Hungary, Mátra Mts., Gyöngyössolymos; 141: Hungary, Bükk Mts., Nagyvisnyó; 142: Hungary, Bakony Mts., Bakonybél; 143: Serbia, Maljen Mts., Brajkovići; 144–145: Turkey, Akyarma Pass; 146: Kazakhstan, Karatau Mts., Bajaldyrtal; 147: Kazakhstan, Tarbagatai Mts., Ayagos)—scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919146" httpUri="https://zenodo.org/record/4919146/files/figure.png" pageId="36" pageNumber="37">127</figureCitation>
, 
<figureCitation id="136B3EA5FF2CFFCB1E475D86D058F903" box="[1332,1377,1733,1758]" captionStart="FIGURES 148–164" captionStartId="60.[151,264,1223,1246]" captionTargetBox="[166,1423,196,1203]" captionTargetId="figure-337@60.[151,1436,181,1215]" captionTargetPageId="60" captionText="FIGURES 148–164. Humps of male Tergite 9 of Zwicknia Murányi, gen. n., caudal view—148–151: Z. acuta Murányi &amp; Orci, sp. n.; 152–156: Z. bifrons (Newman, 1838); 157–159: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 160: Z. kovacsi Murányi &amp; Gamboa, sp. n.; 161–162: Z. sevanica (Zhiltzova, 1964); 163: Z. tuberculata (Zhiltzova, 1964); 164: Z. turkestanica turkestanica (Kimmins, 1950) (148: Slovakia, Krupinská Planina, Pôtor; 149: Hungary, Zemplén Mts., Kishuta; 150: Serbia, Fruska Gora, Vrdnik; 151: Serbia, Zlatibor Mts., Crni Rzav; 152: Hungary, Börzsöny Mts., Szokolya-Királyrét; 153: Hungary, Mátra Mts., Gyöngyössolymos; 154: Hungary, Bükk Mts., Nagyvisnyó; 155: Hungary, Bakony Mts., Bakonybél; 156: Serbia, Maljen Mts., Brajkovići; 157, 159: Croatia, Ivanščica Mts., Stari Golubovec; 158: Hungary, Mecsek Mts., Komló-Zobákpuszta; 160: Romania, Rodna Mts., Borşa-Staţiunea Borşa; 161–162: Turkey, Akyarma Pass; 163: Kazakhstan, Karatau Mts., Bajaldyrtal; 164: Kazakhstan, Tarbagatai Mts., Ayagos)—scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919148" httpUri="https://zenodo.org/record/4919148/files/figure.png" pageId="36" pageNumber="37">148</figureCitation>
), 
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(
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); 23.03.200 9, leg. N. Hordós, D. 
<taxonomicName id="4C5059A3FF2CFFCB18775DABD651F8DC" authority="Murányi &amp; Gamboa &amp; Orci, 2014" authorityName="Murányi &amp; Gamboa &amp; Orci" authorityYear="2014" box="[772,872,1768,1793]" genus="Muranyi" pageId="36" pageNumber="69" rank="genus" status="gen. nov.">Murányi</taxonomicName>
, J. Papp, Zs. Ujvári: 
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2f, 1f larva (
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: PLP3044); 23.03.201 1, leg. K. Orci: 
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: PLP3811; used for molecular studies as 300985, drumming recorded as 2011/No.1), 
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: PLP3812; used for molecular studies as 300982, drumming recorded as 2011/No.2), 
<quantity id="4CA88FC5FF2CFFCB1A5E5C11D468F8B6" box="[301,337,1874,1899]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="36" pageNumber="37" unit="m" value="1.0">1m</quantity>
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<collectionCode id="ED41BAE5FF2CFFCB1A1A5C11D4FDF8B6" box="[361,452,1874,1899]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="36" pageNumber="37">HNHM</collectionCode>
: PLP3813; drumming recorded as 2011/No.3), 
<quantity id="4CA88FC5FF2CFFCB1F7E5C11D108F8B6" box="[1037,1073,1874,1899]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="36" pageNumber="37" unit="m" value="1.0">1m</quantity>
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: PLP3814; used for molecular studies as 300992, drumming recorded as 2011/No.4), 
<quantity id="4CA88FC5FF2CFFCB18EC5C36D6FAF853" box="[927,963,1909,1934]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="36" pageNumber="37" unit="m" value="1.0">1m</quantity>
(
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: PLP3815; used for molecular studies as 301003, drumming recorded as 2011/No.5), 
<quantity id="4CA88FC5FF2CFFCB18735CDBD61DF86C" box="[768,804,1944,1969]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="36" pageNumber="37" unit="m" value="1.0">1m</quantity>
(
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: PLP3816; drumming recorded as 2011/No.6), 
<quantity id="4CA88FC5FF2CFFCB1BE45CFFD583F808" box="[151,186,1980,2005]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="36" pageNumber="37" unit="m" value="1.0">1m</quantity>
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<collectionCode id="ED41BAE5FF2CFFCB1BBF5CFFD411F808" box="[204,296,1980,2005]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="36" pageNumber="37">HNHM</collectionCode>
: PLP3808; used for molecular studies as 300983, drumming recorded as 2011/No.7), 
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, 08.03.201 1, leg. 
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used for drawings 
<figureCitation id="136B3EA5FF2DFFCA1AD55A42D728FEC7" box="[422,529,257,282]" captionStart="FIGURES 127–147" captionStartId="59.[151,264,1277,1300]" captionTargetBox="[154,1418,219,1236]" captionTargetId="figure-173@59.[151,1436,193,1256]" captionTargetPageId="59" captionText="FIGURES 127–147. Male epiproct of Zwicknia Murányi, gen. n.—127–130 Z. acuta Murányi &amp; Orci, sp. n.; 131–132: Z. kovacsi Murányi &amp; Gamboa, sp. n.; 133–138: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 139–143: Z. bifrons (Newman, 1838); 144–145: Z. sevanica (Zhiltzova, 1964); 146: Z. tuberculata (Zhiltzova, 1964); 147: Z. turkestanica turkestanica (Kimmins, 1950); 127–131, 133, 135, 137, 139–147: lateral view; 132, 134, 136, 138: dorsal view (127: Slovakia, Krupinská Planina, Pôtor; 128: Hungary, Zemplén Mts., Kishuta; 129: Serbia, Fruska Gora, Vrdnik; 130: Serbia, Zlatibor Mts., Crni Rzav; 131–132: Romania, Rodna Mts., Borşa–Staţiunea Borşa; 133–134, 137–138: Croatia, Ivanščica Mts., Stari Golubovec; 135–136: Hungary, Mecsek Mts., Komló-Zobákpuszta; 139: Hungary, Börzsöny Mts., Szokolya-Királyrét; 140: Hungary, Mátra Mts., Gyöngyössolymos; 141: Hungary, Bükk Mts., Nagyvisnyó; 142: Hungary, Bakony Mts., Bakonybél; 143: Serbia, Maljen Mts., Brajkovići; 144–145: Turkey, Akyarma Pass; 146: Kazakhstan, Karatau Mts., Bajaldyrtal; 147: Kazakhstan, Tarbagatai Mts., Ayagos)—scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919146" httpUri="https://zenodo.org/record/4919146/files/figure.png" pageId="37" pageNumber="38">Figs. 128</figureCitation>
, 
<figureCitation id="136B3EA5FF2DFFCA196D5A42D772FEC7" box="[542,587,257,282]" captionStart="FIGURES 148–164" captionStartId="60.[151,264,1223,1246]" captionTargetBox="[166,1423,196,1203]" captionTargetId="figure-337@60.[151,1436,181,1215]" captionTargetPageId="60" captionText="FIGURES 148–164. Humps of male Tergite 9 of Zwicknia Murányi, gen. n., caudal view—148–151: Z. acuta Murányi &amp; Orci, sp. n.; 152–156: Z. bifrons (Newman, 1838); 157–159: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 160: Z. kovacsi Murányi &amp; Gamboa, sp. n.; 161–162: Z. sevanica (Zhiltzova, 1964); 163: Z. tuberculata (Zhiltzova, 1964); 164: Z. turkestanica turkestanica (Kimmins, 1950) (148: Slovakia, Krupinská Planina, Pôtor; 149: Hungary, Zemplén Mts., Kishuta; 150: Serbia, Fruska Gora, Vrdnik; 151: Serbia, Zlatibor Mts., Crni Rzav; 152: Hungary, Börzsöny Mts., Szokolya-Királyrét; 153: Hungary, Mátra Mts., Gyöngyössolymos; 154: Hungary, Bükk Mts., Nagyvisnyó; 155: Hungary, Bakony Mts., Bakonybél; 156: Serbia, Maljen Mts., Brajkovići; 157, 159: Croatia, Ivanščica Mts., Stari Golubovec; 158: Hungary, Mecsek Mts., Komló-Zobákpuszta; 160: Romania, Rodna Mts., Borşa-Staţiunea Borşa; 161–162: Turkey, Akyarma Pass; 163: Kazakhstan, Karatau Mts., Bajaldyrtal; 164: Kazakhstan, Tarbagatai Mts., Ayagos)—scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919148" httpUri="https://zenodo.org/record/4919148/files/figure.png" pageId="37" pageNumber="38">149</figureCitation>
), 
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(
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: PLH1258; used for molecular studies as 300989, drumming recorded as 2011/No.1), 
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: PLH1300; used for molecular studies as 30100 and 30101, drumming not recorded), 
<quantity id="4CA88FC5FF2DFFCA1A625A04D40CFEBD" box="[273,309,327,352]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="37" pageNumber="38" unit="m" value="1.0">1m</quantity>
(only bioacoustic study, specimen escaped; drumming recorded as 2011/No.4, 
<figureCitation id="136B3EA5FF2DFFCA1FCA5A04D022FEBD" box="[1209,1307,327,352]" captionStart="FIGURES 169–184" captionStartId="62.[151,264,1845,1868]" captionTargetBox="[201,1412,198,1820]" captionTargetId="figure-140@62.[168,1412,183,1822]" captionTargetPageId="62" captionText="FIGURES 169–184. Male drumming calls of Zwicknia Murányi, gen. n.—169–173: Z. bifrons (Newman, 1838); 174–178: Z. acuta Murányi &amp; Orci, sp. n.; 179–182: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 183–184: Z. kovacsi Murányi &amp; Gamboa, sp. n. (see detailed data on the animals and recording circumstances in Table 9)—Oscillograms in Figs. 169–179, 181, 183 have the same time axis resolution (time scale bar is presented in Fig. 169 for all of them), an other time scale is used in the faster oscillograms of Figs. 180, 182, 184 (time scale bar for them can be found in Fig. 184), those are zoomed sections of Figs. 179, 181, 183, respectively." figureDoi="http://doi.org/10.5281/zenodo.4919156" httpUri="https://zenodo.org/record/4919156/files/figure.png" pageId="37" pageNumber="38">Fig. 174</figureCitation>
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, 15.03.201 1, leg. 
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: PLP3818; used for drawings 
<figureCitation id="136B3EA5FF2DFFCA1F3E5ACDD183FE7A" box="[1101,1210,398,423]" captionStart="FIGURES 127–147" captionStartId="59.[151,264,1277,1300]" captionTargetBox="[154,1418,219,1236]" captionTargetId="figure-173@59.[151,1436,193,1256]" captionTargetPageId="59" captionText="FIGURES 127–147. Male epiproct of Zwicknia Murányi, gen. n.—127–130 Z. acuta Murányi &amp; Orci, sp. n.; 131–132: Z. kovacsi Murányi &amp; Gamboa, sp. n.; 133–138: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 139–143: Z. bifrons (Newman, 1838); 144–145: Z. sevanica (Zhiltzova, 1964); 146: Z. tuberculata (Zhiltzova, 1964); 147: Z. turkestanica turkestanica (Kimmins, 1950); 127–131, 133, 135, 137, 139–147: lateral view; 132, 134, 136, 138: dorsal view (127: Slovakia, Krupinská Planina, Pôtor; 128: Hungary, Zemplén Mts., Kishuta; 129: Serbia, Fruska Gora, Vrdnik; 130: Serbia, Zlatibor Mts., Crni Rzav; 131–132: Romania, Rodna Mts., Borşa–Staţiunea Borşa; 133–134, 137–138: Croatia, Ivanščica Mts., Stari Golubovec; 135–136: Hungary, Mecsek Mts., Komló-Zobákpuszta; 139: Hungary, Börzsöny Mts., Szokolya-Királyrét; 140: Hungary, Mátra Mts., Gyöngyössolymos; 141: Hungary, Bükk Mts., Nagyvisnyó; 142: Hungary, Bakony Mts., Bakonybél; 143: Serbia, Maljen Mts., Brajkovići; 144–145: Turkey, Akyarma Pass; 146: Kazakhstan, Karatau Mts., Bajaldyrtal; 147: Kazakhstan, Tarbagatai Mts., Ayagos)—scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919146" httpUri="https://zenodo.org/record/4919146/files/figure.png" pageId="37" pageNumber="38">Figs. 129</figureCitation>
, 
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, for molecular studies as 300996, drumming recorded as 2011/No.1, 
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: PLP3889; used for molecular studies as 300962); Braničevo District, Homoljske Planina, Krepoljin, stream N of the village, 
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, 
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, 16.03.201 1, leg. 
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, 
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, 17.03.201 1, leg. 
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. Kovács, G. Magos, D. 
<taxonomicName id="4C5059A3FF2DFFCA19A45922D605FDA7" authority="Murányi &amp; Gamboa &amp; Orci, 2014" authorityName="Murányi &amp; Gamboa &amp; Orci" authorityYear="2014" box="[727,828,609,634]" genus="Muranyi" pageId="37" pageNumber="69" rank="genus" status="gen. nov.">Murányi</taxonomicName>
: 
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(
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: PLP3582; used for drawings 
<figureCitation id="136B3EA5FF2DFFCA1E5F5922D0AEFDA7" box="[1324,1431,609,634]" captionStart="FIGURES 127–147" captionStartId="59.[151,264,1277,1300]" captionTargetBox="[154,1418,219,1236]" captionTargetId="figure-173@59.[151,1436,193,1256]" captionTargetPageId="59" captionText="FIGURES 127–147. Male epiproct of Zwicknia Murányi, gen. n.—127–130 Z. acuta Murányi &amp; Orci, sp. n.; 131–132: Z. kovacsi Murányi &amp; Gamboa, sp. n.; 133–138: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 139–143: Z. bifrons (Newman, 1838); 144–145: Z. sevanica (Zhiltzova, 1964); 146: Z. tuberculata (Zhiltzova, 1964); 147: Z. turkestanica turkestanica (Kimmins, 1950); 127–131, 133, 135, 137, 139–147: lateral view; 132, 134, 136, 138: dorsal view (127: Slovakia, Krupinská Planina, Pôtor; 128: Hungary, Zemplén Mts., Kishuta; 129: Serbia, Fruska Gora, Vrdnik; 130: Serbia, Zlatibor Mts., Crni Rzav; 131–132: Romania, Rodna Mts., Borşa–Staţiunea Borşa; 133–134, 137–138: Croatia, Ivanščica Mts., Stari Golubovec; 135–136: Hungary, Mecsek Mts., Komló-Zobákpuszta; 139: Hungary, Börzsöny Mts., Szokolya-Királyrét; 140: Hungary, Mátra Mts., Gyöngyössolymos; 141: Hungary, Bükk Mts., Nagyvisnyó; 142: Hungary, Bakony Mts., Bakonybél; 143: Serbia, Maljen Mts., Brajkovići; 144–145: Turkey, Akyarma Pass; 146: Kazakhstan, Karatau Mts., Bajaldyrtal; 147: Kazakhstan, Tarbagatai Mts., Ayagos)—scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919146" httpUri="https://zenodo.org/record/4919146/files/figure.png" pageId="37" pageNumber="38">Figs. 130</figureCitation>
, 
<figureCitation id="136B3EA5FF2DFFCA1BE459C7D5FDFD40" box="[151,196,644,669]" captionStart="FIGURES 148–164" captionStartId="60.[151,264,1223,1246]" captionTargetBox="[166,1423,196,1203]" captionTargetId="figure-337@60.[151,1436,181,1215]" captionTargetPageId="60" captionText="FIGURES 148–164. Humps of male Tergite 9 of Zwicknia Murányi, gen. n., caudal view—148–151: Z. acuta Murányi &amp; Orci, sp. n.; 152–156: Z. bifrons (Newman, 1838); 157–159: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 160: Z. kovacsi Murányi &amp; Gamboa, sp. n.; 161–162: Z. sevanica (Zhiltzova, 1964); 163: Z. tuberculata (Zhiltzova, 1964); 164: Z. turkestanica turkestanica (Kimmins, 1950) (148: Slovakia, Krupinská Planina, Pôtor; 149: Hungary, Zemplén Mts., Kishuta; 150: Serbia, Fruska Gora, Vrdnik; 151: Serbia, Zlatibor Mts., Crni Rzav; 152: Hungary, Börzsöny Mts., Szokolya-Királyrét; 153: Hungary, Mátra Mts., Gyöngyössolymos; 154: Hungary, Bükk Mts., Nagyvisnyó; 155: Hungary, Bakony Mts., Bakonybél; 156: Serbia, Maljen Mts., Brajkovići; 157, 159: Croatia, Ivanščica Mts., Stari Golubovec; 158: Hungary, Mecsek Mts., Komló-Zobákpuszta; 160: Romania, Rodna Mts., Borşa-Staţiunea Borşa; 161–162: Turkey, Akyarma Pass; 163: Kazakhstan, Karatau Mts., Bajaldyrtal; 164: Kazakhstan, Tarbagatai Mts., Ayagos)—scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919148" httpUri="https://zenodo.org/record/4919148/files/figure.png" pageId="37" pageNumber="38">151</figureCitation>
), 
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(
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: PLP3819; used for molecular studies as 300994, drumming recorded as 2011/No.1), 1f (
<collectionCode id="ED41BAE5FF2DFFCA1BEC59E4D5C2FD1D" box="[159,251,679,704]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="37" pageNumber="38">HNHM</collectionCode>
: PLP3890; used for molecular studies as 300952), 
<quantity id="4CA88FC5FF2DFFCA181859E4D6B6FD1D" box="[875,911,679,704]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="37" pageNumber="38" unit="m" value="1.0">1m</quantity>
(only bioacoustic study, specimen escaped; drumming recorded as 2011/No.3, 
<figureCitation id="136B3EA5FF2DFFCA196D5988D746FD39" box="[542,639,715,740]" captionStart="FIGURES 169–184" captionStartId="62.[151,264,1845,1868]" captionTargetBox="[201,1412,198,1820]" captionTargetId="figure-140@62.[168,1412,183,1822]" captionTargetPageId="62" captionText="FIGURES 169–184. Male drumming calls of Zwicknia Murányi, gen. n.—169–173: Z. bifrons (Newman, 1838); 174–178: Z. acuta Murányi &amp; Orci, sp. n.; 179–182: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 183–184: Z. kovacsi Murányi &amp; Gamboa, sp. n. (see detailed data on the animals and recording circumstances in Table 9)—Oscillograms in Figs. 169–179, 181, 183 have the same time axis resolution (time scale bar is presented in Fig. 169 for all of them), an other time scale is used in the faster oscillograms of Figs. 180, 182, 184 (time scale bar for them can be found in Fig. 184), those are zoomed sections of Figs. 179, 181, 183, respectively." figureDoi="http://doi.org/10.5281/zenodo.4919156" httpUri="https://zenodo.org/record/4919156/files/figure.png" pageId="37" pageNumber="38">Fig. 178</figureCitation>
).
</paragraph>
<paragraph id="8BEF2220FF2DFFCA1BB459ADD6E2FBB7" blockId="37.[151,1437,151,2015]" pageId="37" pageNumber="38">
<emphasis id="B924FE32FF2DFFCA1BB459ADD62CFCDA" bold="true" box="[199,789,750,775]" pageId="37" pageNumber="38">Other material—Records based on morphology:</emphasis>
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: 
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, im 
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, 18.03.197 1, leg. 
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: 
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(
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; 
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only as ‘ 
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, 1861, leg. 
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: 
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(
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: 
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, 
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, 15.01.195 1, leg. 
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: 
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<collectingRegion id="4994ECC2FF2DFFCA1FD85877D0A1FC90" box="[1195,1432,820,845]" country="Germany" name="Sachsen-Anhalt" pageId="37" pageNumber="38">Saxony-Anhalt State</collectingRegion>
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, 15.03.200 5, 
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: 
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1f (
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: 
<collectingRegion id="4994ECC2FF2DFFCA1F455814D1D5FCAD" box="[1078,1260,855,880]" country="Hungary" name="Nograd" pageId="37" pageNumber="38">Nógrád County</collectingRegion>
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at the gorge of 
<collectorName id="26A547F6FF2DFFCA190C5838D626FC49" box="[639,799,891,916]" pageId="37" pageNumber="38">Párizs Stream</collectorName>
, 03.04.200 5, leg. 
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<collectorName id="26A547F6FF2DFFCA1F7F5838D15DFC49" box="[1036,1124,891,916]" pageId="37" pageNumber="38">Kovács</collectorName>
: 
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(
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)
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; Borsod-Abaúj- 
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, 14.03.200 4, leg. D. 
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, 
<collectorName id="26A547F6FF2DFFCA1F8658DCD057FC6A" box="[1269,1390,926,951]" pageId="37" pageNumber="38">Zs. Sóvári</collectorName>
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(
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<collectingMunicipality id="6B8BB85AFF2DFFCA18415882D6B5FC07" box="[818,908,961,986]" pageId="37" pageNumber="38">Kishuta</collectingMunicipality>
, 
<location id="8E8F74FBFF2DFFCA18EA5881D164FC07" LSID="urn:lsid:plazi:treatment:03F99336FF2CFFC81BE4586AD770FBAC:8E8F74FBFF2DFFCA18EA5881D164FC07" box="[921,1117,961,986]" country="Hungary" county="Zemplen Mts." municipality="Kishuta" name="Kemence Stream" pageId="37" pageNumber="38" stateProvince="Borsod-Abauj-Zemplen">Kemence Stream</location>
, 
<quantity id="4CA88FC5FF2DFFCA1F195882D18DFC04" box="[1130,1204,961,986]" metricMagnitude="2" metricUnit="m" metricValue="1.75" pageId="37" pageNumber="38" unit="m" value="175.0">
<elevation id="007DC513FF2DFFCA1F195882D18DFC04" box="[1130,1204,961,986]" metricMagnitude="2" metricUnit="m" metricValue="1.75" pageId="37" pageNumber="38" unit="m" value="175.0">175 m</elevation>
</quantity>
, 14.03.200 4, leg. D. 
<taxonomicName id="4C5059A3FF2DFFCA1BE458A7D5C3FC20" authority="Murányi &amp; Gamboa &amp; Orci, 2014" authorityName="Murányi &amp; Gamboa &amp; Orci" authorityYear="2014" box="[151,250,996,1021]" genus="Muranyi" pageId="37" pageNumber="69" rank="genus" status="gen. nov.">Murányi</taxonomicName>
, 
<collectorName id="26A547F6FF2DFFCA1A7958A6D4BFFC20" box="[266,390,996,1021]" pageId="37" pageNumber="38">Zs. Sóvári</collectorName>
: 1f larva (
<collectionCode id="ED41BAE5FF2DFFCA197758A7D759FC20" box="[516,608,996,1021]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="37" pageNumber="38">HNHM</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B38287DFF2DFFCA190B58A7D07DFBFD" ID-GBIF-Occurrence="3325453350" collectionCode="HNHM" collectorName="L. Danyi &amp; J. Kontschan" country="Slovakia" county="Banskobistricky Region" elevation="470" location="Horny Tisovnik" municipality="Javorie Mts." pageId="37" pageNumber="38" specimenCount="1" stateProvince="Borsod-Abauj-Zemplen County">
<collectingCountry id="F34762B0FF2DFFCA190B58A7D631FC20" box="[632,776,996,1021]" name="Slovakia" pageId="37" pageNumber="38">SLOVAKIA</collectingCountry>
: 
<collectingCounty id="628E5AACFF2DFFCA186458A7D114FC20" box="[791,1069,996,1021]" pageId="37" pageNumber="38">Banskobistrický Region</collectingCounty>
, 
<collectingMunicipality id="6B8BB85AFF2DFFCA1F4E58A7D1F6FC21" box="[1085,1231,996,1021]" pageId="37" pageNumber="38">Javorie Mts.</collectingMunicipality>
, 
<location id="8E8F74FBFF2DFFCA1FAC58A7D0A1FC20" LSID="urn:lsid:plazi:treatment:03F99336FF2CFFC81BE4586AD770FBAC:8E8F74FBFF2DFFCA1FAC58A7D0A1FC20" box="[1247,1432,996,1021]" country="Slovakia" county="Banskobistricky Region" municipality="Javorie Mts." name="Horny Tisovnik" pageId="37" pageNumber="38" stateProvince="Borsod-Abauj-Zemplen County">Horný Tisovník</location>
, 
<location id="8E8F74FBFF2DFFCA1BE45F44D5C6FBFD" LSID="urn:lsid:plazi:treatment:03F99336FF2CFFC81BE4586AD770FBAC:8E8F74FBFF2DFFCA1BE45F44D5C6FBFD" box="[151,255,1031,1056]" country="Slovakia" county="Banskobistricky Region" municipality="Javorie Mts." name="Kostolne" pageId="37" pageNumber="38" stateProvince="Borsod-Abauj-Zemplen County">Kostolné</location>
, 
<location id="8E8F74FBFF2DFFCA1A785F44D4FEFBFD" LSID="urn:lsid:plazi:treatment:03F99336FF2CFFC81BE4586AD770FBAC:8E8F74FBFF2DFFCA1A785F44D4FEFBFD" box="[267,455,1031,1056]" country="Slovakia" county="Banskobistricky Region" municipality="Javorie Mts." name="Tisovnik Stream" pageId="37" pageNumber="38" stateProvince="Borsod-Abauj-Zemplen County">Tisovník Stream</location>
, 
<quantity id="4CA88FC5FF2DFFCA1AA05F44D724FBC2" box="[467,541,1031,1056]" metricMagnitude="2" metricUnit="m" metricValue="4.7" pageId="37" pageNumber="38" unit="m" value="470.0">
<elevation id="007DC513FF2DFFCA1AA05F44D724FBC2" box="[467,541,1031,1056]" metricMagnitude="2" metricUnit="m" metricValue="4.7" pageId="37" pageNumber="38" unit="m" value="470.0">470 m</elevation>
</quantity>
, 24.03.200 6, leg. 
<collectorName id="26A547F6FF2DFFCA19945F4BD677FBFD" box="[743,846,1031,1056]" pageId="37" pageNumber="38">L. Dányi</collectorName>
, 
<collectorName id="26A547F6FF2DFFCA18295F4BD6D6FBFD" box="[858,1007,1031,1056]" pageId="37" pageNumber="38">J. Kontschán</collectorName>
, D. 
<taxonomicName id="4C5059A3FF2DFFCA1F6E5F44D1BBFBFD" authority="Murányi &amp; Gamboa &amp; Orci, 2014" authorityName="Murányi &amp; Gamboa &amp; Orci" authorityYear="2014" box="[1053,1154,1031,1056]" genus="Muranyi" pageId="37" pageNumber="69" rank="genus" status="gen. nov.">Murányi</taxonomicName>
: 
<quantity id="4CA88FC5FF2DFFCA1FFE5F44D18BFBFD" box="[1165,1202,1031,1056]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="37" pageNumber="38" unit="m" value="8.0">
<elevation id="007DC513FF2DFFCA1FFE5F44D18BFBFD" box="[1165,1202,1031,1056]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="37" pageNumber="38" unit="m" value="8.0">8m</elevation>
</quantity>
1f (
<collectionCode id="ED41BAE5FF2DFFCA1F935F44D005FBFD" box="[1248,1340,1031,1056]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="37" pageNumber="38">HNHM</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B38287DFF2DFFCA1E235F44D6EEFBB7" ID-GBIF-Occurrence="3325453304" collectionCode="WNHM" country="Slovakia" pageId="37" pageNumber="38" specimenCount="20" specimenCount-male="20" stateProvince="Borsod-Abauj-Zemplen County">
further 
<specimenCount id="9D56E9A9FF2DFFCA1BE45F6ED5C4FB9B" box="[151,253,1069,1094]" pageId="37" pageNumber="38" type="male">20 males</specimenCount>
present in the 
<collectionCode id="ED41BAE5FF2DFFCA1AD95F6DD730FB98" box="[426,521,1070,1093]" country="0" httpUri="http://grbio.org/cool/ie1n-abad" name="Oklahoma Baptist University, Webster Natural History Museum" pageId="37" pageNumber="38">WNHM</collectionCode>
, collected during second half of the 19 
<superScript id="7C258F68FF2DFFCA18BA5F69D6ECFBE5" attach="left" box="[969,981,1066,1080]" fontSize="6" pageId="37" pageNumber="38">th</superScript>
century, but without any reliable labels (some are possibly from 
<collectorName id="26A547F6FF2DFFCA1ADE5F12D4CCFBB7" box="[429,501,1105,1130]" pageId="37" pageNumber="38">Mainz</collectorName>
or 
<collectorName id="26A547F6FF2DFFCA19695F11D754FBB7" box="[538,621,1106,1130]" pageId="37" pageNumber="38">Bergen</collectorName>
, collected by 
<collectorName id="26A547F6FF2DFFCA18745F11D656FBB4" box="[775,879,1106,1129]" pageId="37" pageNumber="38">F. Brauer</collectorName>
in 1865)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34A71ABFF2DFFC81BB45F37D481FCA5" lastPageId="39" lastPageNumber="40" pageId="37" pageNumber="38" type="description">
<paragraph id="8BEF2220FF2DFFCA1BB45F37D1F0FB0F" blockId="37.[151,1437,151,2015]" pageId="37" pageNumber="38">
<emphasis id="B924FE32FF2DFFCA1BB45F37D461FB50" bold="true" box="[199,344,1140,1165]" pageId="37" pageNumber="38">Description.</emphasis>
Head, thorax, appendages and basal segments of the abdomen generotypic. Males micropterous, females macropterous. Body length: 
<typeStatus id="54EB9C82FF2DFFCA19335FD4D79BFB6D" box="[576,674,1175,1200]" pageId="37" pageNumber="38" type="holotype">holotype</typeStatus>
7.5 mm, male 
<typeStatus id="54EB9C82FF2DFFCA18245FDBD6FBFB6D" box="[855,962,1176,1200]" pageId="37" pageNumber="38" type="paratype">paratypes</typeStatus>
6.5–8.0, female 
<typeStatus id="54EB9C82FF2DFFCA1FF45FDBD1CBFB6D" box="[1159,1266,1176,1200]" pageId="37" pageNumber="38" type="paratype">paratypes</typeStatus>
8.0–10.5 mm; forewing length: 
<typeStatus id="54EB9C82FF2DFFCA1A2B5FF9D483FB0E" box="[344,442,1210,1235]" pageId="37" pageNumber="38" type="holotype">holotype</typeStatus>
1.6 mm, male 
<typeStatus id="54EB9C82FF2DFFCA19115FF8D7F4FB0E" box="[610,717,1211,1235]" pageId="37" pageNumber="38" type="paratype">paratypes</typeStatus>
1.3–1.7 mm, female 
<typeStatus id="54EB9C82FF2DFFCA18CF5FF8D11EFB0E" box="[956,1063,1211,1235]" pageId="37" pageNumber="38" type="paratype">paratypes</typeStatus>
8.5–11.0 mm.
</paragraph>
<paragraph id="8BEF2220FF2DFFCA1BB45F9ED078F9CD" blockId="37.[151,1437,151,2015]" pageId="37" pageNumber="38">
Male terminalia (
<figureCitation id="136B3EA5FF2DFFCA1AFE5F9ED72EFB2B" box="[397,535,1245,1270]" captionStart="FIGURES 78–81" captionStartId="38.[151,264,1941,1964]" captionTargetBox="[226,1366,264,1921]" captionTargetId="figure-99@38.[213,1381,249,1934]" captionTargetPageId="38" captionText="FIGURES 78–81. Male terminalia of Zwicknia acuta Murányi &amp; Orci, sp. n.; paratype, Slovakia, Banskobystrický Region, Krupinská Planina, Pôtor—78: dorsal view; 79: ventral view; 80: lateral view; 81: caudal view (Fp: fusion plate; Pp: paraproct)—scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919118" httpUri="https://zenodo.org/record/4919118/files/figure.png" pageId="37" pageNumber="38">Figs. 78–81</figureCitation>
, 
<figureCitation id="136B3EA5FF2DFFCA19565F9ED76AFB2B" box="[549,595,1245,1270]" captionStart="FIGURES 120–123" captionStartId="57.[151,264,1761,1784]" captionTargetBox="[165,1401,468,1695]" captionTargetId="figure-157@57.[151,1436,457,1720]" captionTargetPageId="57" captionText="FIGURES 120–123. Male terminalia of Zwicknia Murányi, gen. n., dorso-caudal view—120: Z. bifrons (Newman, 1838); 121: Z. acuta Murányi &amp; Orci, sp. n.; 122: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 123: Z. kovacsi Murányi &amp; Gamboa, sp. n.—scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919142" httpUri="https://zenodo.org/record/4919142/files/figure.png" pageId="37" pageNumber="38">121</figureCitation>
): Process of Tg 9 high, perpendicularly elevated, apex 2× wider than the medial section of Ep-scl; rectangular in shape, apex bearing two small hump-like tips; sides indented in caudal view, forming the membranous portion narrowest medially (
<figureCitation id="136B3EA5FF2DFFCA18325E67D6D1FAE0" box="[833,1000,1316,1341]" captionStart="FIGURES 148–164" captionStartId="60.[151,264,1223,1246]" captionTargetBox="[166,1423,196,1203]" captionTargetId="figure-337@60.[151,1436,181,1215]" captionTargetPageId="60" captionText="FIGURES 148–164. Humps of male Tergite 9 of Zwicknia Murányi, gen. n., caudal view—148–151: Z. acuta Murányi &amp; Orci, sp. n.; 152–156: Z. bifrons (Newman, 1838); 157–159: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 160: Z. kovacsi Murányi &amp; Gamboa, sp. n.; 161–162: Z. sevanica (Zhiltzova, 1964); 163: Z. tuberculata (Zhiltzova, 1964); 164: Z. turkestanica turkestanica (Kimmins, 1950) (148: Slovakia, Krupinská Planina, Pôtor; 149: Hungary, Zemplén Mts., Kishuta; 150: Serbia, Fruska Gora, Vrdnik; 151: Serbia, Zlatibor Mts., Crni Rzav; 152: Hungary, Börzsöny Mts., Szokolya-Királyrét; 153: Hungary, Mátra Mts., Gyöngyössolymos; 154: Hungary, Bükk Mts., Nagyvisnyó; 155: Hungary, Bakony Mts., Bakonybél; 156: Serbia, Maljen Mts., Brajkovići; 157, 159: Croatia, Ivanščica Mts., Stari Golubovec; 158: Hungary, Mecsek Mts., Komló-Zobákpuszta; 160: Romania, Rodna Mts., Borşa-Staţiunea Borşa; 161–162: Turkey, Akyarma Pass; 163: Kazakhstan, Karatau Mts., Bajaldyrtal; 164: Kazakhstan, Tarbagatai Mts., Ayagos)—scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919148" httpUri="https://zenodo.org/record/4919148/files/figure.png" pageId="37" pageNumber="38">Figs. 148–151</figureCitation>
). Tg 10, B-scl and Lb-scl generotypic. Ep-scl narrow and pointed in dorsal view, medially not swollen, its medial width ½ to ⅓ of basal width; tip straight and acute in lateral view, divided section long. Ventral membranous part between the division of Ep-scl ends far before the base in lateral view; apical spines stout, distributed not only on the membranous part but extend to the Ep-scl (
<figureCitation id="136B3EA5FF2DFFCA1B865EF2D499FA17" box="[245,416,1457,1482]" captionStart="FIGURES 106–109" captionStartId="54.[151,264,1087,1110]" captionTargetBox="[151,1436,193,1067]" captionTargetId="figure-252@54.[151,1436,193,1067]" captionTargetPageId="54" captionText="FIGURES 106–109. Lateral view of male epiproct of Zwicknia Murányi, gen. n.—106: Z. bifrons (Newman, 1838); 107: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 108–109: Z. acuta Murányi &amp; Orci, sp. n.—scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919136" httpUri="https://zenodo.org/record/4919136/files/figure.png" pageId="37" pageNumber="38">Figs. 108–109</figureCitation>
, 
<figureCitation id="136B3EA5FF2DFFCA1AC05EF2D4E6FA17" box="[435,479,1457,1482]" captionStart="FIGURES 114–119" captionStartId="56.[151,265,1540,1563]" captionTargetBox="[151,1436,193,1520]" captionTargetId="figure-203@56.[151,1436,193,1520]" captionTargetPageId="56" captionText="FIGURES 114–119. Dorsal view of male epiproct of Zwicknia Murányi, gen. n.—114: Z. bifrons (Newman, 1838); 115, 117: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 116: Z. kovacsi Murányi &amp; Gamboa, sp. n.; 118: Z. sevanica (Zhiltzova, 1964); 119: Z. acuta Murányi &amp; Orci, sp. n.—scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919140" httpUri="https://zenodo.org/record/4919140/files/figure.png" pageId="37" pageNumber="38">119</figureCitation>
, 
<figureCitation id="136B3EA5FF2DFFCA1A825EF2D761FA17" box="[497,600,1457,1482]" captionStart="FIGURES 127–147" captionStartId="59.[151,264,1277,1300]" captionTargetBox="[154,1418,219,1236]" captionTargetId="figure-173@59.[151,1436,193,1256]" captionTargetPageId="59" captionText="FIGURES 127–147. Male epiproct of Zwicknia Murányi, gen. n.—127–130 Z. acuta Murányi &amp; Orci, sp. n.; 131–132: Z. kovacsi Murányi &amp; Gamboa, sp. n.; 133–138: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 139–143: Z. bifrons (Newman, 1838); 144–145: Z. sevanica (Zhiltzova, 1964); 146: Z. tuberculata (Zhiltzova, 1964); 147: Z. turkestanica turkestanica (Kimmins, 1950); 127–131, 133, 135, 137, 139–147: lateral view; 132, 134, 136, 138: dorsal view (127: Slovakia, Krupinská Planina, Pôtor; 128: Hungary, Zemplén Mts., Kishuta; 129: Serbia, Fruska Gora, Vrdnik; 130: Serbia, Zlatibor Mts., Crni Rzav; 131–132: Romania, Rodna Mts., Borşa–Staţiunea Borşa; 133–134, 137–138: Croatia, Ivanščica Mts., Stari Golubovec; 135–136: Hungary, Mecsek Mts., Komló-Zobákpuszta; 139: Hungary, Börzsöny Mts., Szokolya-Királyrét; 140: Hungary, Mátra Mts., Gyöngyössolymos; 141: Hungary, Bükk Mts., Nagyvisnyó; 142: Hungary, Bakony Mts., Bakonybél; 143: Serbia, Maljen Mts., Brajkovići; 144–145: Turkey, Akyarma Pass; 146: Kazakhstan, Karatau Mts., Bajaldyrtal; 147: Kazakhstan, Tarbagatai Mts., Ayagos)—scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919146" httpUri="https://zenodo.org/record/4919146/files/figure.png" pageId="37" pageNumber="38">127–130</figureCitation>
). I-scl generotypic, Ec long and rarely everted on the non in-copula specimens. St 9 slightly projecting medially, vesicle medium sized to large, 
<figureCitation id="136B3EA5FF2DFFCA18885E97D175FA30" box="[1019,1100,1492,1517]" captionStart="FIGURES 78–81" captionStartId="38.[151,264,1941,1964]" captionTargetBox="[226,1366,264,1921]" captionTargetId="figure-99@38.[213,1381,249,1934]" captionTargetPageId="38" captionText="FIGURES 78–81. Male terminalia of Zwicknia acuta Murányi &amp; Orci, sp. n.; paratype, Slovakia, Banskobystrický Region, Krupinská Planina, Pôtor—78: dorsal view; 79: ventral view; 80: lateral view; 81: caudal view (Fp: fusion plate; Pp: paraproct)—scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919118" httpUri="https://zenodo.org/record/4919118/files/figure.png" pageId="37" pageNumber="38">Fig. 79</figureCitation>
illustrates the smallest of the range. Sg rounded with pronounced triangular shape, tip usually incised. Pp, Fp, Rp and cerci generotypic.
</paragraph>
<paragraph id="8BEF2220FF2DFFCA1BB45D59D47EF9A7" blockId="37.[151,1437,151,2015]" pageId="37" pageNumber="38">
Female subgenital plate (
<figureCitation id="136B3EA5FF2DFFCA1A825D59D772F9EE" box="[497,587,1562,1587]" captionStart="FIGURES 62–65" captionStartId="33.[151,264,1508,1531]" captionTargetBox="[151,1436,231,1487]" captionTargetId="figure-310@33.[151,1436,193,1487]" captionTargetPageId="33" captionText="FIGURES 62–65. Female genitalia of Zwicknia Murányi, gen. n.—62–63: Z. bifrons (Newman, 1838); 64: Z. acuta Murányi &amp; Orci, sp. n.; 65: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 62: terminalia, ventral view; 63: inner genital organs, dorsal view; 64–65: subgenital plates, ventral view (gc: genital cavity; go: genital opening; Sg: subgenital plate)—scales 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919112" httpUri="https://zenodo.org/record/4919112/files/figure.png" pageId="37" pageNumber="38">Fig. 64</figureCitation>
): Less rectangular than usual, posterior margin rounded and sometimes slightly overhanging the segment. Antero-lateral recess usually distinct, the plate is entirely brown; lateral sclerites relatively large.
</paragraph>
<paragraph id="8BEF2220FF2DFFC91BB45DC7D0A0FF09" blockId="37.[151,1437,151,2015]" lastBlockId="38.[151,1436,151,212]" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38">
Drumming: Males produce (2)-3-4-(6) drumming calls in a sequence (
<figureCitation id="136B3EA5FF2DFFCA18A75DC7D10CF940" box="[980,1077,1668,1693]" captionStart="FIGURES 165–168" captionStartId="61.[151,264,1330,1353]" captionTargetBox="[162,1428,857,1282]" captionTargetId="figure-628@61.[151,1436,850,1309]" captionTargetPageId="61" captionText="FIGURES 165–168. Slow oscillograms of male calls of four Zwicknia Murányi, gen. n. species, showing call sequences versus sporadic call production—165: two call sequence of Z. bifrons (Newman, 1838), &quot;//&quot; indicates where 2 minutes 23 seconds of silence was cut out from the recording; 166: three call sequences of Z. acuta Murányi &amp; Orci, sp. n.; 167: sporadic calls of Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 168: sporadic call production in Z. kovacsi Murányi &amp; Gamboa, sp. n.—time scale is the same for the four oscillograms; for detailed data on the animals and recording circumstances, see Table 9." figureDoi="http://doi.org/10.5281/zenodo.4919154" httpUri="https://zenodo.org/record/4919154/files/figure.png" pageId="37" pageNumber="38">Fig. 166</figureCitation>
). Calls in a sequence are similar to each other. However, the first call is generally shorter and of lower amplitude than the following ones. Calls are monophasic signals consisting of one beat group with inter beat intervals increasing gradually toward the end of beat group (
<figureCitation id="136B3EA5FF2DFFCA1B965DAED479F8DB" box="[229,320,1773,1798]" captionStart="FIGURE 190" captionStartId="64.[151,250,1125,1147]" captionTargetBox="[163,1430,185,1044]" captionTargetId="figure-392@64.[151,1436,177,1058]" captionTargetPageId="64" captionText="FIGURE 190. Inter-beat interval variation during male calls in the four examined Zwicknia Murányi, gen. n. species.—Box plot colors: white—Z. bifrons (Newman, 1838), red—Z. acuta Murányi &amp; Orci, sp. n., blue—Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n. In Z. kovacsi Murányi &amp; Gamboa, sp. n. we have drumming samples only from two specimens therefore in the case of that species two simple mark series are used to show inter-beat intervals in their calls (circles for specimen 1, and triangles for specimen 2). i1, i2, i3 and so on are labels containing the serial number of inter-beat intervals counting them from the beginning of each call. Box plots show the duration values measured in 7 specimens in Z. bifrons and Z. acuta, and 5 specimens in Z. rupprechti. Vertical axis has a logarithmic scale. Only calls recorded at an ambient air temperature of 17–19 oC were used. (see the measurement data in Appendix Tables 1–4)." figureDoi="http://doi.org/10.5281/zenodo.4919160" httpUri="https://zenodo.org/record/4919160/files/figure.png" pageId="37" pageNumber="38">Fig 190</figureCitation>
: red box plots, Appendix 
<tableCitation id="C6D2179BFF2DFFCA19295DAED792F8DB" box="[602,683,1773,1798]" captionStart="TABLE 1" captionStartId="7.[147,247,602,628]" captionTargetBox="[167,1849,720,1573]" captionText="TABLE 1. Ventral sclerites of valid extant Capniidae genera. — Capniella not included because of lacking information; in case of two character states the first one refers to the type species; for abbreviations of sclerites see Figs. 53-55." httpUri="http://table.plazi.org/id/DF2F72A8FF0FFFE81BE05919D188FD46" pageId="37" pageNumber="38" tableUuid="DF2F72A8FF0FFFE81BE05919D188FD46">Table 1</tableCitation>
). The amplitude envelope of calls are of the following main 
<typeStatus id="54EB9C82FF2DFFCA1E345DADD0BFF8DB" box="[1351,1414,1774,1798]" pageId="37" pageNumber="38">types</typeStatus>
: a full length crescending (e.g. see the second call in 
<figureCitation id="136B3EA5FF2DFFCA19BE5C57D617F8F0" box="[717,814,1812,1837]" captionStart="FIGURES 169–184" captionStartId="62.[151,264,1845,1868]" captionTargetBox="[201,1412,198,1820]" captionTargetId="figure-140@62.[168,1412,183,1822]" captionTargetPageId="62" captionText="FIGURES 169–184. Male drumming calls of Zwicknia Murányi, gen. n.—169–173: Z. bifrons (Newman, 1838); 174–178: Z. acuta Murányi &amp; Orci, sp. n.; 179–182: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 183–184: Z. kovacsi Murányi &amp; Gamboa, sp. n. (see detailed data on the animals and recording circumstances in Table 9)—Oscillograms in Figs. 169–179, 181, 183 have the same time axis resolution (time scale bar is presented in Fig. 169 for all of them), an other time scale is used in the faster oscillograms of Figs. 180, 182, 184 (time scale bar for them can be found in Fig. 184), those are zoomed sections of Figs. 179, 181, 183, respectively." figureDoi="http://doi.org/10.5281/zenodo.4919156" httpUri="https://zenodo.org/record/4919156/files/figure.png" pageId="37" pageNumber="38">Fig. 175</figureCitation>
), an initial crescendo+nearly constant main part (see 4 
<superScript id="7C258F68FF2DFFCA1EE35C53D0A5F8C3" attach="none" box="[1424,1436,1808,1822]" fontSize="6" pageId="37" pageNumber="38">th</superScript>
call in 
<figureCitation id="136B3EA5FF2DFFCA1B905C79D479F88E" box="[227,320,1850,1875]" captionStart="FIGURES 169–184" captionStartId="62.[151,264,1845,1868]" captionTargetBox="[201,1412,198,1820]" captionTargetId="figure-140@62.[168,1412,183,1822]" captionTargetPageId="62" captionText="FIGURES 169–184. Male drumming calls of Zwicknia Murányi, gen. n.—169–173: Z. bifrons (Newman, 1838); 174–178: Z. acuta Murányi &amp; Orci, sp. n.; 179–182: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 183–184: Z. kovacsi Murányi &amp; Gamboa, sp. n. (see detailed data on the animals and recording circumstances in Table 9)—Oscillograms in Figs. 169–179, 181, 183 have the same time axis resolution (time scale bar is presented in Fig. 169 for all of them), an other time scale is used in the faster oscillograms of Figs. 180, 182, 184 (time scale bar for them can be found in Fig. 184), those are zoomed sections of Figs. 179, 181, 183, respectively." figureDoi="http://doi.org/10.5281/zenodo.4919156" httpUri="https://zenodo.org/record/4919156/files/figure.png" pageId="37" pageNumber="38">Fig. 176</figureCitation>
) or a crescending first half+decrescending second half pattern (e.g. 4 
<superScript id="7C258F68FF2DFFCA1F325C75D174F899" attach="left" box="[1089,1101,1846,1860]" fontSize="6" pageId="37" pageNumber="38">th</superScript>
call in 
<figureCitation id="136B3EA5FF2DFFCA1FD35C79D039F88E" box="[1184,1280,1850,1875]" captionStart="FIGURES 169–184" captionStartId="62.[151,264,1845,1868]" captionTargetBox="[201,1412,198,1820]" captionTargetId="figure-140@62.[168,1412,183,1822]" captionTargetPageId="62" captionText="FIGURES 169–184. Male drumming calls of Zwicknia Murányi, gen. n.—169–173: Z. bifrons (Newman, 1838); 174–178: Z. acuta Murányi &amp; Orci, sp. n.; 179–182: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 183–184: Z. kovacsi Murányi &amp; Gamboa, sp. n. (see detailed data on the animals and recording circumstances in Table 9)—Oscillograms in Figs. 169–179, 181, 183 have the same time axis resolution (time scale bar is presented in Fig. 169 for all of them), an other time scale is used in the faster oscillograms of Figs. 180, 182, 184 (time scale bar for them can be found in Fig. 184), those are zoomed sections of Figs. 179, 181, 183, respectively." figureDoi="http://doi.org/10.5281/zenodo.4919156" httpUri="https://zenodo.org/record/4919156/files/figure.png" pageId="37" pageNumber="38">Fig. 177</figureCitation>
). Calls follow each other with increasing inter-call intervals. See 
<figureCitation id="136B3EA5FF2DFFCA198A5C1ED699F8AB" box="[761,928,1885,1910]" captionStart="FIGURES 169–184" captionStartId="62.[151,264,1845,1868]" captionTargetBox="[201,1412,198,1820]" captionTargetId="figure-140@62.[168,1412,183,1822]" captionTargetPageId="62" captionText="FIGURES 169–184. Male drumming calls of Zwicknia Murányi, gen. n.—169–173: Z. bifrons (Newman, 1838); 174–178: Z. acuta Murányi &amp; Orci, sp. n.; 179–182: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 183–184: Z. kovacsi Murányi &amp; Gamboa, sp. n. (see detailed data on the animals and recording circumstances in Table 9)—Oscillograms in Figs. 169–179, 181, 183 have the same time axis resolution (time scale bar is presented in Fig. 169 for all of them), an other time scale is used in the faster oscillograms of Figs. 180, 182, 184 (time scale bar for them can be found in Fig. 184), those are zoomed sections of Figs. 179, 181, 183, respectively." figureDoi="http://doi.org/10.5281/zenodo.4919156" httpUri="https://zenodo.org/record/4919156/files/figure.png" pageId="37" pageNumber="38">Figs. 174–178</figureCitation>
for the oscillographic pattern of the male drumming calls of this species. Descriptive statistics of five drumming characters are presented in 
<tableCitation id="C6D2179BFF2DFFCA1F935CC3D00FF844" box="[1248,1334,1920,1945]" captionStart="TABLE 6" captionStartId="43.[151,241,728,751]" captionTargetPageId="43" captionText="TABLE 6. Descriptive statistics for the examined rhythmic characters of the male drumming calls of Zwicknia acuta Murányi &amp; Orci, sp. n., and also for the ambient air temperature during the sound recordings.—&quot;n&quot; is the number of male specimens examined. In each character data points where mean values obtained for each examined male on the basis of 1-10 signal measurements per specimen. Number of examined specimens per population: Hungary: Zemplén Mts—4; Slovakia: Krupinská Planina—10; Serbia: Fruska Gora—4, Homoljske Planina—1, Zlatibor Mts—4." httpUri="http://table.plazi.org/id/DF2F72A8FF23FFC41BE4599BD1F4FCA7" pageId="37" pageNumber="38" tableUuid="DF2F72A8FF23FFC41BE4599BD1F4FCA7">Table 6</tableCitation>
. See 
<figureCitation id="136B3EA5FF2DFFCA1E035CC3D5F8F861" captionStart="FIGURE 190" captionStartId="64.[151,250,1125,1147]" captionTargetBox="[163,1430,185,1044]" captionTargetId="figure-392@64.[151,1436,177,1058]" captionTargetPageId="64" captionText="FIGURE 190. Inter-beat interval variation during male calls in the four examined Zwicknia Murányi, gen. n. species.—Box plot colors: white—Z. bifrons (Newman, 1838), red—Z. acuta Murányi &amp; Orci, sp. n., blue—Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n. In Z. kovacsi Murányi &amp; Gamboa, sp. n. we have drumming samples only from two specimens therefore in the case of that species two simple mark series are used to show inter-beat intervals in their calls (circles for specimen 1, and triangles for specimen 2). i1, i2, i3 and so on are labels containing the serial number of inter-beat intervals counting them from the beginning of each call. Box plots show the duration values measured in 7 specimens in Z. bifrons and Z. acuta, and 5 specimens in Z. rupprechti. Vertical axis has a logarithmic scale. Only calls recorded at an ambient air temperature of 17–19 oC were used. (see the measurement data in Appendix Tables 1–4)." figureDoi="http://doi.org/10.5281/zenodo.4919160" httpUri="https://zenodo.org/record/4919160/files/figure.png" pageId="37" pageNumber="38">Fig. 190</figureCitation>
for a box plot series showing the variation of inter-beat intervals within a call in relation to the other three 
<taxonomicName id="4C5059A3FF2DFFCA1BE45C85D45BF802" box="[151,354,1990,2015]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="undetermined">Zwicknia species</taxonomicName>
, where drumming signals are characterized in this study. Appendix 
<tableCitation id="C6D2179BFF2DFFCA1FD65C85D039F802" box="[1189,1280,1990,2015]" captionStart="TABLE 1" captionStartId="7.[147,247,602,628]" captionTargetBox="[167,1849,720,1573]" captionText="TABLE 1. Ventral sclerites of valid extant Capniidae genera. — Capniella not included because of lacking information; in case of two character states the first one refers to the type species; for abbreviations of sclerites see Figs. 53-55." httpUri="http://table.plazi.org/id/DF2F72A8FF0FFFE81BE05919D188FD46" pageId="37" pageNumber="38" tableUuid="DF2F72A8FF0FFFE81BE05919D188FD46">Table 1</tableCitation>
contains the measurement data of inter-beat intervals of this species. The male female drumming duet is a long sequence of male call—female answer, where the female answers sometimes overlaps the terminal part of male calls (
<figureCitation id="136B3EA5FF2EFFC91F9A5BF8D0B7FF09" box="[1257,1422,187,212]" captionStart="FIGURES 185–189" captionStartId="63.[151,264,1012,1035]" captionTargetBox="[166,1430,178,988]" captionTargetId="figure-458@63.[151,1436,178,1004]" captionTargetPageId="63" captionText="FIGURES 185–189. Male-female drumming duets of three Zwicknia Murányi, gen. n. species.—185: Z. bifrons (Newman, 1838) from Hungary, Pilis Mts., Bükkös Stream, recorded at 16.9 °C; 186–187: Z. acuta Murányi &amp; Orci, sp. n. from Slovakia, Krupinská Planina, Stará Rieka, recorded at 18.2 °C; 188: call-answer duet in Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n. from Hungary, Mecsek Mts., Völgységi Stream, recorded at 18.2 °C; 189: call-answer-reply duet in Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n. from Hungary, Mecsek Mts., Völgységi Stream, recorded at 17.9 °C—arrows with a gender sign point to the beginning of a section of the oscillogram containing the beats of that gender." figureDoi="http://doi.org/10.5281/zenodo.4919158" httpUri="https://zenodo.org/record/4919158/files/figure.png" pageId="38" pageNumber="39">Figs. 186–187</figureCitation>
).
</paragraph>
<caption id="DF2F72A8FF2EFFC91BE45CD6D4AAF834" ID-DOI="http://doi.org/10.5281/zenodo.4919118" ID-Zenodo-Dep="4919118" httpUri="https://zenodo.org/record/4919118/files/figure.png" pageId="38" pageNumber="39" startId="38.[151,264,1941,1964]" targetBox="[226,1366,264,1921]" targetPageId="38">
<paragraph id="8BEF2220FF2EFFC91BE45CD6D4AAF834" blockId="38.[151,1436,1941,2026]" pageId="38" pageNumber="39">
<emphasis id="B924FE32FF2EFFC91BE45CD6D46FF876" bold="true" box="[151,342,1941,1964]" pageId="38" pageNumber="39">FIGURES 78–81.</emphasis>
Male terminalia of 
<taxonomicName id="4C5059A3FF2EFFC919555CD5D64EF871" authority="Muranyi &amp; Orci" authorityName="Muranyi &amp; Orci" authorityYear="2014" box="[550,887,1942,1964]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="acuta" status="sp. nov.">
<emphasis id="B924FE32FF2EFFC919555CD5D7FAF871" box="[550,707,1942,1964]" italics="true" pageId="38" pageNumber="39">Zwicknia acuta</emphasis>
Murányi &amp; Orci
</taxonomicName>
, 
<emphasis id="B924FE32FF2EFFC918F05CD4D684F871" bold="true" box="[899,957,1943,1964]" pageId="38" pageNumber="39">
<taxonomicNameLabel id="A2174349FF2EFFC918F05CD4D684F871" box="[899,957,1943,1964]" pageId="38" pageNumber="39" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; paratype, Slovakia, Banskobystrický Region, Krupinská Planina, Pôtor 
<emphasis id="B924FE32FF2EFFC91AD05CF5D480F816" box="[419,441,1974,1995]" italics="true" pageId="38" pageNumber="39">—</emphasis>
78: dorsal view; 79: ventral view; 80: lateral view; 81: caudal view (Fp: fusion plate; Pp: paraproct)—scale 1 mm.
</paragraph>
</caption>
<paragraph id="8BEF2220FF2FFFC81BB45BD4D642FEE2" blockId="39.[151,1437,151,1137]" pageId="39" pageNumber="40">
Genetics: Three different haplotypes were found corresponding well to the geographical distribution (
<collectingCountry id="F34762B0FF2FFFC81E405BD4D0A1FF6D" box="[1331,1432,151,176]" name="Hungary" pageId="39" pageNumber="40">Hungary</collectingCountry>
, 
<collectingCountry id="F34762B0FF2FFFC81BE45BFFD5DBFF08" box="[151,226,188,213]" name="Serbia" pageId="39" pageNumber="40">Serbia</collectingCountry>
, and 
<collectingCountry id="F34762B0FF2FFFC81A6D5BFFD4BCFF08" box="[286,389,188,213]" name="Slovakia" pageId="39" pageNumber="40">Slovakia</collectingCountry>
). The 
<collectingCountry id="F34762B0FF2FFFC81ABE5BFFD716FF08" box="[461,559,188,213]" name="Slovakia" pageId="39" pageNumber="40">Slovakia</collectingCountry>
haplotypes clustered at the same node as the northern Hungarian population, resulting in 0.4% nucleotide divergence and two diagnostic characters. We found that the specimens from Maljen Mts. of 
<collectingCountry id="F34762B0FF2FFFC81B8E5A47D47DFEC0" box="[253,324,260,285]" name="Serbia" pageId="39" pageNumber="40">Serbia</collectingCountry>
included individuals of both 
<taxonomicName id="4C5059A3FF2FFFC819C05A46D61CFEC0" authorityName="Capnor" baseAuthorityName="Newman" baseAuthorityYear="1838" box="[691,805,260,285]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="bifrons">
<emphasis id="B924FE32FF2FFFC819C05A46D61CFEC0" box="[691,805,260,285]" italics="true" pageId="39" pageNumber="40">Z. bifrons</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C5059A3FF2FFFC818185A46D6F7FEC1" authorityName="Muranyi &amp; Orci" authorityYear="2014" box="[875,974,261,284]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="acuta">
<emphasis id="B924FE32FF2FFFC818185A46D6F7FEC1" box="[875,974,261,284]" italics="true" pageId="39" pageNumber="40">Z. acuta</emphasis>
</taxonomicName>
(
<figureCitation id="136B3EA5FF2FFFC818965A47D177FEC0" box="[997,1102,260,285]" captionStart="FIGURE 192" captionStartId="66.[151,249,1180,1202]" captionTargetBox="[156,1424,193,1155]" captionTargetId="figure-240@66.[151,1433,185,1171]" captionTargetPageId="66" captionText="FIGURE 192. Phylogenetic tree of cox1 including Zwicknia Murányi, gen. n. with our accession codes (300941-301003), and GenBank closest species (with alphanumeric accession codes) as outgroup. MP and ML had same topology. Numbers on braches are ML/MP bootstrap values&gt;50%. Species names right to the tree refer to morphological species. P-Distance table is showed." figureDoi="http://doi.org/10.5281/zenodo.4919164" httpUri="https://zenodo.org/record/4919164/files/figure.png" pageId="39" pageNumber="40">Fig. 192</figureCitation>
). However, despite that the morphology and mating calls correspond to 
<taxonomicName id="4C5059A3FF2FFFC819F65A6AD7C8FE9D" authorityName="Capnor" baseAuthorityName="Newman" baseAuthorityYear="1838" box="[645,753,295,320]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="bifrons">
<emphasis id="B924FE32FF2FFFC819F65A6AD7C8FE9D" box="[645,753,295,320]" italics="true" pageId="39" pageNumber="40">Z. bifrons</emphasis>
</taxonomicName>
in all cases.
</paragraph>
<paragraph id="8BEF2220FF2FFFC81BB45A0FD481FCA5" blockId="39.[151,1437,151,1137]" pageId="39" pageNumber="40">
<emphasis id="B924FE32FF2FFFC81BB45A0FD405FEBE" bold="true" box="[199,316,332,357]" pageId="39" pageNumber="40">Affinities.</emphasis>
This species is morphologically very close to 
<taxonomicName id="4C5059A3FF2FFFC818255A0ED6FDFEB8" authorityName="Capnor" baseAuthorityName="Newman" baseAuthorityYear="1838" box="[854,964,332,357]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="bifrons">
<emphasis id="B924FE32FF2FFFC818255A0ED6FDFEB8" box="[854,964,332,357]" italics="true" pageId="39" pageNumber="40">Z. bifrons</emphasis>
</taxonomicName>
. One of the Serbian populations (Maljen Mts.) may be hybrids, as they share mitochondrial haplotypes (
<figureCitation id="136B3EA5FF2FFFC818165A2CD6F1FE55" box="[869,968,367,392]" captionStart="FIGURE 192" captionStartId="66.[151,249,1180,1202]" captionTargetBox="[156,1424,193,1155]" captionTargetId="figure-240@66.[151,1433,185,1171]" captionTargetPageId="66" captionText="FIGURE 192. Phylogenetic tree of cox1 including Zwicknia Murányi, gen. n. with our accession codes (300941-301003), and GenBank closest species (with alphanumeric accession codes) as outgroup. MP and ML had same topology. Numbers on braches are ML/MP bootstrap values&gt;50%. Species names right to the tree refer to morphological species. P-Distance table is showed." figureDoi="http://doi.org/10.5281/zenodo.4919164" httpUri="https://zenodo.org/record/4919164/files/figure.png" pageId="39" pageNumber="40">Fig. 192</figureCitation>
). Individuals have drumming signals and morphology similar to 
<taxonomicName id="4C5059A3FF2FFFC81AEF5AD0D730FE76" authorityName="Capnor" baseAuthorityName="Newman" baseAuthorityYear="1838" box="[412,521,402,427]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="bifrons">
<emphasis id="B924FE32FF2FFFC81AEF5AD0D730FE76" box="[412,521,402,427]" italics="true" pageId="39" pageNumber="40">Z. bifrons</emphasis>
</taxonomicName>
and illustrations are provided (
<figureCitation id="136B3EA5FF2FFFC8181E5AD1D6E3FE76" box="[877,986,402,427]" captionStart="FIGURES 127–147" captionStartId="59.[151,264,1277,1300]" captionTargetBox="[154,1418,219,1236]" captionTargetId="figure-173@59.[151,1436,193,1256]" captionTargetPageId="59" captionText="FIGURES 127–147. Male epiproct of Zwicknia Murányi, gen. n.—127–130 Z. acuta Murányi &amp; Orci, sp. n.; 131–132: Z. kovacsi Murányi &amp; Gamboa, sp. n.; 133–138: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 139–143: Z. bifrons (Newman, 1838); 144–145: Z. sevanica (Zhiltzova, 1964); 146: Z. tuberculata (Zhiltzova, 1964); 147: Z. turkestanica turkestanica (Kimmins, 1950); 127–131, 133, 135, 137, 139–147: lateral view; 132, 134, 136, 138: dorsal view (127: Slovakia, Krupinská Planina, Pôtor; 128: Hungary, Zemplén Mts., Kishuta; 129: Serbia, Fruska Gora, Vrdnik; 130: Serbia, Zlatibor Mts., Crni Rzav; 131–132: Romania, Rodna Mts., Borşa–Staţiunea Borşa; 133–134, 137–138: Croatia, Ivanščica Mts., Stari Golubovec; 135–136: Hungary, Mecsek Mts., Komló-Zobákpuszta; 139: Hungary, Börzsöny Mts., Szokolya-Királyrét; 140: Hungary, Mátra Mts., Gyöngyössolymos; 141: Hungary, Bükk Mts., Nagyvisnyó; 142: Hungary, Bakony Mts., Bakonybél; 143: Serbia, Maljen Mts., Brajkovići; 144–145: Turkey, Akyarma Pass; 146: Kazakhstan, Karatau Mts., Bajaldyrtal; 147: Kazakhstan, Tarbagatai Mts., Ayagos)—scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919146" httpUri="https://zenodo.org/record/4919146/files/figure.png" pageId="39" pageNumber="40">Figs. 143</figureCitation>
, 
<figureCitation id="136B3EA5FF2FFFC818955AD1D12AFE76" box="[998,1043,402,427]" captionStart="FIGURES 148–164" captionStartId="60.[151,264,1223,1246]" captionTargetBox="[166,1423,196,1203]" captionTargetId="figure-337@60.[151,1436,181,1215]" captionTargetPageId="60" captionText="FIGURES 148–164. Humps of male Tergite 9 of Zwicknia Murányi, gen. n., caudal view—148–151: Z. acuta Murányi &amp; Orci, sp. n.; 152–156: Z. bifrons (Newman, 1838); 157–159: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 160: Z. kovacsi Murányi &amp; Gamboa, sp. n.; 161–162: Z. sevanica (Zhiltzova, 1964); 163: Z. tuberculata (Zhiltzova, 1964); 164: Z. turkestanica turkestanica (Kimmins, 1950) (148: Slovakia, Krupinská Planina, Pôtor; 149: Hungary, Zemplén Mts., Kishuta; 150: Serbia, Fruska Gora, Vrdnik; 151: Serbia, Zlatibor Mts., Crni Rzav; 152: Hungary, Börzsöny Mts., Szokolya-Királyrét; 153: Hungary, Mátra Mts., Gyöngyössolymos; 154: Hungary, Bükk Mts., Nagyvisnyó; 155: Hungary, Bakony Mts., Bakonybél; 156: Serbia, Maljen Mts., Brajkovići; 157, 159: Croatia, Ivanščica Mts., Stari Golubovec; 158: Hungary, Mecsek Mts., Komló-Zobákpuszta; 160: Romania, Rodna Mts., Borşa-Staţiunea Borşa; 161–162: Turkey, Akyarma Pass; 163: Kazakhstan, Karatau Mts., Bajaldyrtal; 164: Kazakhstan, Tarbagatai Mts., Ayagos)—scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919148" httpUri="https://zenodo.org/record/4919148/files/figure.png" pageId="39" pageNumber="40">156</figureCitation>
, 
<figureCitation id="136B3EA5FF2FFFC81F535AD1D177FE76" box="[1056,1102,402,427]" captionStart="FIGURES 169–184" captionStartId="62.[151,264,1845,1868]" captionTargetBox="[201,1412,198,1820]" captionTargetId="figure-140@62.[168,1412,183,1822]" captionTargetPageId="62" captionText="FIGURES 169–184. Male drumming calls of Zwicknia Murányi, gen. n.—169–173: Z. bifrons (Newman, 1838); 174–178: Z. acuta Murányi &amp; Orci, sp. n.; 179–182: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 183–184: Z. kovacsi Murányi &amp; Gamboa, sp. n. (see detailed data on the animals and recording circumstances in Table 9)—Oscillograms in Figs. 169–179, 181, 183 have the same time axis resolution (time scale bar is presented in Fig. 169 for all of them), an other time scale is used in the faster oscillograms of Figs. 180, 182, 184 (time scale bar for them can be found in Fig. 184), those are zoomed sections of Figs. 179, 181, 183, respectively." figureDoi="http://doi.org/10.5281/zenodo.4919156" httpUri="https://zenodo.org/record/4919156/files/figure.png" pageId="39" pageNumber="40">173</figureCitation>
). However, nearby Slovakian and Hungarian populations not sympatric with 
<taxonomicName id="4C5059A3FF2FFFC819BE5AF4D679FE12" authorityName="Capnor" baseAuthorityName="Newman" baseAuthorityYear="1838" box="[717,832,438,463]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="bifrons">
<emphasis id="B924FE32FF2FFFC819BE5AF4D679FE12" box="[717,832,438,463]" italics="true" pageId="39" pageNumber="40">Z. bifrons</emphasis>
</taxonomicName>
populations and other Serbian populations exhibit differences in both morphology and mating calls. Morphologically, the males differ by having narrow and pointed Ep-scl with an acute and straight apex, instead of wide and blunt Ep-scl and upcurved tip; ventral membranous section terminate far before the base instead of nearly reaching it, and Tg 9 process with indented sides caudally instead of consistently narrow sides. Males of 
<taxonomicName id="4C5059A3FF2FFFC819DA5906D63FFD81" authorityName="Muranyi &amp; Orci" authorityYear="2014" box="[681,774,581,604]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="acuta">
<emphasis id="B924FE32FF2FFFC819DA5906D63FFD81" box="[681,774,581,604]" italics="true" pageId="39" pageNumber="40">Z. acuta</emphasis>
</taxonomicName>
are easily distinguishable from other 
<taxonomicName id="4C5059A3FF2FFFC81FC45907D04EFD80" box="[1207,1399,580,605]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B924FE32FF2FFFC81FC45907D024FD80" box="[1207,1309,580,605]" italics="true" pageId="39" pageNumber="40">Zwicknia</emphasis>
species
</taxonomicName>
on the basis of acute Ep-scl tip and very high, rectangular process of Tg 9. Females are difficult to separate with certainty and the larvae are morphologically indistinguishable. The drumming signals of 
<taxonomicName id="4C5059A3FF2FFFC81FEC59CFD1C4FD7E" authorityName="Muranyi &amp; Orci" authorityYear="2014" box="[1183,1277,652,675]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="acuta">
<emphasis id="B924FE32FF2FFFC81FEC59CFD1C4FD7E" box="[1183,1277,652,675]" italics="true" pageId="39" pageNumber="40">Z. acuta</emphasis>
</taxonomicName>
clearly differ from those of the other three 
<taxonomicName id="4C5059A3FF2FFFC81A9859EDD795FD1A" box="[491,684,686,711]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B924FE32FF2FFFC81A9859EDD76BFD1A" box="[491,594,686,711]" italics="true" pageId="39" pageNumber="40">Zwicknia</emphasis>
species
</taxonomicName>
analysed (
<figureCitation id="136B3EA5FF2FFFC8185A59EDD6E8FD1A" box="[809,977,686,711]" captionStart-0="FIGURES 165–168" captionStart-1="FIGURES 169–184" captionStartId-0="61.[151,264,1330,1353]" captionStartId-1="62.[151,264,1845,1868]" captionTargetBox-0="[162,1428,857,1282]" captionTargetBox-1="[201,1412,198,1820]" captionTargetId-0="figure-628@61.[151,1436,850,1309]" captionTargetId-1="figure-140@62.[168,1412,183,1822]" captionTargetPageId-0="61" captionTargetPageId-1="62" captionText-0="FIGURES 165–168. Slow oscillograms of male calls of four Zwicknia Murányi, gen. n. species, showing call sequences versus sporadic call production—165: two call sequence of Z. bifrons (Newman, 1838), &quot;//&quot; indicates where 2 minutes 23 seconds of silence was cut out from the recording; 166: three call sequences of Z. acuta Murányi &amp; Orci, sp. n.; 167: sporadic calls of Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 168: sporadic call production in Z. kovacsi Murányi &amp; Gamboa, sp. n.—time scale is the same for the four oscillograms; for detailed data on the animals and recording circumstances, see Table 9." captionText-1="FIGURES 169–184. Male drumming calls of Zwicknia Murányi, gen. n.—169–173: Z. bifrons (Newman, 1838); 174–178: Z. acuta Murányi &amp; Orci, sp. n.; 179–182: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 183–184: Z. kovacsi Murányi &amp; Gamboa, sp. n. (see detailed data on the animals and recording circumstances in Table 9)—Oscillograms in Figs. 169–179, 181, 183 have the same time axis resolution (time scale bar is presented in Fig. 169 for all of them), an other time scale is used in the faster oscillograms of Figs. 180, 182, 184 (time scale bar for them can be found in Fig. 184), those are zoomed sections of Figs. 179, 181, 183, respectively." figureDoi-0="http://doi.org/10.5281/zenodo.4919154" figureDoi-1="http://doi.org/10.5281/zenodo.4919156" httpUri-0="https://zenodo.org/record/4919154/files/figure.png" httpUri-1="https://zenodo.org/record/4919156/files/figure.png" pageId="39" pageNumber="40">Figs. 165–184</figureCitation>
). Male drumming calls are shorter (
<figureCitation id="136B3EA5FF2FFFC81E1C59EDD5FAFD36" captionStart="FIGURE 191" captionStartId="65.[151,250,1357,1379]" captionTargetBox="[164,1416,208,1328]" captionTargetId="figure-231@65.[151,1436,193,1336]" captionTargetPageId="65" captionText="FIGURE 191. A scatter plot showing the clear separation of male calls of Zwicknia Murányi, gen. n. along two drumming signal characters—Z. bifrons (Newman, 1838): triangles; Z. acuta Murányi &amp; Orci, sp. n.: circles; Z. kovacsi Murányi &amp; Gamboa, sp. n.: +; Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.: X—Each mark represents the mean values of the characters for a given specimen. Note that both axes have logarithmic scale. Air temperature varied between 15–21 oC in every species (for details see Tables 6–8)." figureDoi="http://doi.org/10.5281/zenodo.4919162" httpUri="https://zenodo.org/record/4919162/files/figure.png" pageId="39" pageNumber="40">Fig. 191</figureCitation>
, 
<tableCitation id="C6D2179BFF2FFFC81BA35991D475FD36" box="[208,332,722,747]" captionStart-0="TABLE 6" captionStart-1="TABLE 7" captionStart-2="TABLE 8" captionStartId-0="43.[151,241,728,751]" captionStartId-1="48.[151,241,152,175]" captionStartId-2="65.[151,239,1605,1628]" captionTargetPageId-0="43" captionTargetPageId-1="48" captionTargetPageId-2="65" captionText-0="TABLE 6. Descriptive statistics for the examined rhythmic characters of the male drumming calls of Zwicknia acuta Murányi &amp; Orci, sp. n., and also for the ambient air temperature during the sound recordings.—&quot;n&quot; is the number of male specimens examined. In each character data points where mean values obtained for each examined male on the basis of 1-10 signal measurements per specimen. Number of examined specimens per population: Hungary: Zemplén Mts—4; Slovakia: Krupinská Planina—10; Serbia: Fruska Gora—4, Homoljske Planina—1, Zlatibor Mts—4." captionText-1="TABLE 7. Descriptive statistics for the examined rhythmic characters of the male drumming calls of Zwicknia bifrons (Newman, 1838), and also for the ambient air temperature during the sound recordings.—&quot;n&quot; is the number of male specimens examined. In each character data points where mean values obtained for each examined male on the basis of 1-10 signal measurements per specimen. Number of examined specimens per population: Hungary: Pilis Mts— 7, Bakony Mts—2, Börzsöny Mts—4, Mátra Mts—5;Serbia: Maljen Mts—7." captionText-2="TABLE 8. Descriptive statistics for the examined rhythmic characters of the male drumming calls of Zwicknia rupprechti Murányi, Orci &amp; Gamboa, sp. n., and also for the ambient air temperature during the sound recordings.—&quot;n&quot; is the number of male specimens examined. In each character data points where mean values obtained for each examined male on the basis of 1-10 signal measurements per specimen. Number of examined specimens per population: Hungary: Mecsek Mts—7; Croatia: Ivanščica Mts—3." httpUri-0="http://table.plazi.org/id/DF2F72A8FF23FFC41BE4599BD1F4FCA7" pageId="39" pageNumber="40" tableUuid-0="DF2F72A8FF23FFC41BE4599BD1F4FCA7">Tables 6–8</tableCitation>
) and contain generally a fewer number of beats than in 
<taxonomicName id="4C5059A3FF2FFFC818BB5990D10DFD36" authorityName="Capnor" baseAuthorityName="Newman" baseAuthorityYear="1838" box="[968,1076,722,747]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="bifrons">
<emphasis id="B924FE32FF2FFFC818BB5990D10DFD36" box="[968,1076,722,747]" italics="true" pageId="39" pageNumber="40">Z. bifrons</emphasis>
</taxonomicName>
, but are much longer than in 
<taxonomicName id="4C5059A3FF2FFFC81EF65990D5D2FCD3" authorityName="Muranyi &amp; Gamboa" authorityYear="2014" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="kovacsi">
<emphasis id="B924FE32FF2FFFC81EF65990D5D2FCD3" italics="true" pageId="39" pageNumber="40">Z. kovacsi</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C5059A3FF2FFFC81A5759B5D481FCD3" authorityName="Muranyi, Orci &amp; Gamboa" authorityYear="2014" box="[292,440,757,782]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="rupprechti">
<emphasis id="B924FE32FF2FFFC81A5759B5D481FCD3" box="[292,440,757,782]" italics="true" pageId="39" pageNumber="40">Z. rupprechti</emphasis>
</taxonomicName>
. The number of calls in a call sequence is generally higher, and calls are repeated with shorter inter call intervals than in 
<taxonomicName id="4C5059A3FF2FFFC819635859D742FCEF" authorityName="Capnor" baseAuthorityName="Newman" baseAuthorityYear="1838" box="[528,635,793,818]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="bifrons">
<emphasis id="B924FE32FF2FFFC819635859D742FCEF" box="[528,635,793,818]" italics="true" pageId="39" pageNumber="40">Z. bifrons</emphasis>
</taxonomicName>
(
<figureCitation id="136B3EA5FF2FFFC819F8585AD616FCEF" box="[651,815,793,818]" captionStart="FIGURES 165–168" captionStartId="61.[151,264,1330,1353]" captionTargetBox="[162,1428,857,1282]" captionTargetId="figure-628@61.[151,1436,850,1309]" captionTargetPageId="61" captionText="FIGURES 165–168. Slow oscillograms of male calls of four Zwicknia Murányi, gen. n. species, showing call sequences versus sporadic call production—165: two call sequence of Z. bifrons (Newman, 1838), &quot;//&quot; indicates where 2 minutes 23 seconds of silence was cut out from the recording; 166: three call sequences of Z. acuta Murányi &amp; Orci, sp. n.; 167: sporadic calls of Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 168: sporadic call production in Z. kovacsi Murányi &amp; Gamboa, sp. n.—time scale is the same for the four oscillograms; for detailed data on the animals and recording circumstances, see Table 9." figureDoi="http://doi.org/10.5281/zenodo.4919154" httpUri="https://zenodo.org/record/4919154/files/figure.png" pageId="39" pageNumber="40">Figs. 165–166</figureCitation>
, 
<tableCitation id="C6D2179BFF2FFFC81849585AD68EFCEF" box="[826,951,793,818]" captionStart-0="TABLE 6" captionStart-1="TABLE 7" captionStartId-0="43.[151,241,728,751]" captionStartId-1="48.[151,241,152,175]" captionTargetPageId-0="43" captionTargetPageId-1="48" captionText-0="TABLE 6. Descriptive statistics for the examined rhythmic characters of the male drumming calls of Zwicknia acuta Murányi &amp; Orci, sp. n., and also for the ambient air temperature during the sound recordings.—&quot;n&quot; is the number of male specimens examined. In each character data points where mean values obtained for each examined male on the basis of 1-10 signal measurements per specimen. Number of examined specimens per population: Hungary: Zemplén Mts—4; Slovakia: Krupinská Planina—10; Serbia: Fruska Gora—4, Homoljske Planina—1, Zlatibor Mts—4." captionText-1="TABLE 7. Descriptive statistics for the examined rhythmic characters of the male drumming calls of Zwicknia bifrons (Newman, 1838), and also for the ambient air temperature during the sound recordings.—&quot;n&quot; is the number of male specimens examined. In each character data points where mean values obtained for each examined male on the basis of 1-10 signal measurements per specimen. Number of examined specimens per population: Hungary: Pilis Mts— 7, Bakony Mts—2, Börzsöny Mts—4, Mátra Mts—5;Serbia: Maljen Mts—7." httpUri-0="http://table.plazi.org/id/DF2F72A8FF23FFC41BE4599BD1F4FCA7" pageId="39" pageNumber="40" tableUuid-0="DF2F72A8FF23FFC41BE4599BD1F4FCA7">Tables 6–7</tableCitation>
). Molecular diagnostics of this species also yielded differences among the other species with a node of four diagnostic characters as compared to 
<taxonomicName id="4C5059A3FF2FFFC81E72587ED055FC88" authorityName="Capnor" baseAuthorityName="Newman" baseAuthorityYear="1838" box="[1281,1388,828,853]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="bifrons">
<emphasis id="B924FE32FF2FFFC81E72587ED055FC88" box="[1281,1388,828,853]" italics="true" pageId="39" pageNumber="40">Z. bifrons</emphasis>
</taxonomicName>
and 1–5% of total divergence.
</paragraph>
</subSubSection>
<subSubSection id="C34A71ABFF2FFFC81BB458C0D437FBF7" pageId="39" pageNumber="40" type="distribution">
<paragraph id="8BEF2220FF2FFFC81BB458C0D437FBF7" blockId="39.[151,1437,151,1137]" pageId="39" pageNumber="40">
<emphasis id="B924FE32FF2FFFC81BB458C0D4C0FC41" bold="true" box="[199,505,899,924]" pageId="39" pageNumber="40">Distribution and ecology.</emphasis>
The species is found in southern 
<collectingCountry id="F34762B0FF2FFFC818E258C0D6CFFC41" box="[913,1014,899,924]" name="Slovakia" pageId="39" pageNumber="40">Slovakia</collectingCountry>
, northeastern 
<collectingCountry id="F34762B0FF2FFFC81FEA58C7D1C5FC41" box="[1177,1276,900,924]" name="Hungary" pageId="39" pageNumber="40">Hungary</collectingCountry>
, northern and central 
<collectingCountry id="F34762B0FF2FFFC81B9E58E5D401FC62" box="[237,312,934,959]" name="Serbia" pageId="39" pageNumber="40">Serbia</collectingCountry>
. Specimens from 
<collectingCountry id="F34762B0FF2FFFC8197958E5D764FC62" box="[522,605,934,959]" name="Austria" pageId="39" pageNumber="40">Austria</collectingCountry>
and 
<collectingCountry id="F34762B0FF2FFFC819E958E5D63BFC62" box="[666,770,934,959]" name="Germany" pageId="39" pageNumber="40">Germany</collectingCountry>
are attributed to this species on the basis of morphology (
<figureCitation id="136B3EA5FF2FFFC81BEC5889D47EFC3E" box="[159,327,970,995]" captionStart="FIGURES 196–197" captionStartId="68.[151,264,1908,1931]" captionTargetBox="[188,1393,187,1898]" captionTargetId="figure-93@68.[188,1393,187,1898]" captionTargetPageId="68" captionText="FIGURES 196–197. Distribution of Zwicknia Murányi, gen. n.—196: global distribution; 197: studied populations of the Carpathian Basin and the northern Balkans—in Fig. 196, white dot on Z. bifrons symbols indicate populations studied by Rupprecht (1997), dotted lines indicate limits of s.l. Z. bifrons literature reports; in Fig. 197, white dot on symbols indicate populations where drumming signals were studied by us, white star indicate type localities." figureDoi="http://doi.org/10.5281/zenodo.4919168" httpUri="https://zenodo.org/record/4919168/files/figure.png" pageId="39" pageNumber="40">Figs. 196–197</figureCitation>
). Adults were found in March emerging from slow or moderately fast flowing, large to mediumsized streams, mostly in alder (
<taxonomicName id="4C5059A3FF2FFFC81A8558AED632FBDB" authority="Gaertin." authorityName="Gaertin." box="[502,779,1005,1030]" class="Magnoliopsida" family="Betulaceae" genus="Alnus" kingdom="Plantae" order="Fagales" pageId="39" pageNumber="40" phylum="Tracheophyta" rank="species" species="glutinosa">
<emphasis id="B924FE32FF2FFFC81A8558AED79CFBDB" box="[502,677,1005,1030]" italics="true" pageId="39" pageNumber="40">Alnus glutinosa</emphasis>
Gaertin.
</taxonomicName>
) forests, but also in open grasslands between 200– 
<quantity id="4CA88FC5FF2FFFC81E3B58AED0A5FBDB" box="[1352,1436,1005,1030]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="39" pageNumber="40" unit="m" value="1.0">1,000m</quantity>
(
<figureCitation id="136B3EA5FF2FFFC81BEC5F52D43BFBF7" box="[159,258,1041,1066]" captionStart="FIGURES 193–195" captionStartId="67.[151,264,1829,1852]" captionTargetBox="[199,1367,231,1794]" captionTargetId="figure-90@67.[199,1367,231,1810]" captionTargetPageId="67" captionText="FIGURES 193–195. Type localities of the new species described in Zwicknia Murányi, gen. n.—193: Zwicknia acuta Murányi &amp; Orci, sp. n.: Stará rieka, Slovakia, Banskobystrický Region, Krupinská Planina, Pôtor; 194: Zwicknia kovacsi Murányi &amp; Gamboa, sp. n.: Cimpoies Stream, Romania, Maramureş County, Rodna Mts., Borşa-Staţiunea Borşa; 195: Zwicknia rupprechti Murányi, Orci &amp; Gamboa, sp. n.: Völgységi Stream, Hungary, Baranya County, Mecsek Mts., Komló-Zobákpuszta." figureDoi="http://doi.org/10.5281/zenodo.4919166" httpUri="https://zenodo.org/record/4919166/files/figure.png" pageId="39" pageNumber="40">Fig. 193</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34A71ABFF2FFFC81BB45F77D770FBAC" pageId="39" pageNumber="40" type="etymology">
<paragraph id="8BEF2220FF2FFFC81BB45F77D770FBAC" blockId="39.[151,1437,151,1137]" pageId="39" pageNumber="40">
<emphasis id="B924FE32FF2FFFC81BB45F77D477FB90" bold="true" box="[199,334,1076,1101]" pageId="39" pageNumber="40">Etymology.</emphasis>
The name 
<taxonomicName id="4C5059A3FF2FFFC81AB95F76D731FB91" authorityName="Muranyi &amp; Orci" authorityYear="2014" box="[458,520,1077,1100]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="acuta">
<emphasis id="B924FE32FF2FFFC81AB95F76D731FB91" box="[458,520,1077,1100]" italics="true" pageId="39" pageNumber="40">acuta</emphasis>
</taxonomicName>
(from the latin word acutus, meaning acute) refers to the acute tip of the epiproct. Used as an adjective, gender feminine.
</paragraph>
</subSubSection>
</treatment>
</document>