plazi/treatments-xml
2
0
Fork 0
Code Issues Pull requests Releases Activity
treatments-xml/data/03/E8/9C/03E89C04A020000542F2D2CEE5393A39.xml
maintenance bc2564723d added 03
2024-06-21 12:22:17 +02:00

611 lines
78 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="1F061DB9FA9A1D5CBEBE2EE254792047" ID="10.11646/zootaxa.4276.4.9SLASH11349" ID-CLB-Dataset="32840" ID-DOI="10.11646/zootaxa.4276.4.9" ID-GBIF-Dataset="0081d7e4-e694-400c-8683-132166e3e038" ID-ISSN="1175-5326" ID-Zenodo-Dep="808096" ID-ZooBank="CA6D6102-957C-43DB-8278-861564992F4B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1497428755232" checkinUser="plazi" docAuthor="Abdelsalam, Khaled M., Taylor, Paul D. &amp; Dorgham, Mohamed M." docDate="2017" docId="03E89C04A020000542F2D2CEE5393A39" docLanguage="en" docName="zootaxa.4276.4.9SLASH11349.pdf" docOrigin="Zootaxa 4276 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Calyptotheca alexandriensis Abdelsalam &amp; Taylor &amp; Dorgham 2017, n. sp." docType="treatment" docVersion="7" lastPageNumber="587" masterDocId="FFD1E47CA02200004265D650E6383F09" masterDocTitle="A new species of Calyptotheca (Bryozoa: Cheilostomata) from Alexandria, Egypt, southeastern Mediterranean" masterLastPageNumber="590" masterPageNumber="582" pageNumber="584" updateTime="1698440518328" updateUser="ExternalLinkService">
<mods:mods id="64B71E688616A26310374875DB38577A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4A0895DC665CDAC0814FD9440CC9368F">
<mods:title id="B6B8050DD1087B3E7846161567787586">A new species of Calyptotheca (Bryozoa: Cheilostomata) from Alexandria, Egypt, southeastern Mediterranean</mods:title>
</mods:titleInfo>
<mods:name id="E2AB645723B849A8264B0D5FB51068C1" type="personal">
<mods:role id="52018D21918DE2344CAF3ADD9DE431E1">
<mods:roleTerm id="9CDE18F7B97D2527138442F3A42E62F3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9864C3A0E5253E462C3EF20341B44540">Abdelsalam, Khaled M.</mods:namePart>
</mods:name>
<mods:name id="F28DC4F7261D10F57B2337377BC52220" type="personal">
<mods:role id="16D5A69F861D951C4A9E9545E3FB81A5">
<mods:roleTerm id="3DEDD0C604665A2A9BC76F3CA6644FFE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DEE068972938F6D17FB5AA67F7E63FE3">Taylor, Paul D.</mods:namePart>
</mods:name>
<mods:name id="545D220F8E5DA556856559C3F2178D70" type="personal">
<mods:role id="6D26565DD9FBAD8D304E8AA5ABD57A3B">
<mods:roleTerm id="CABE06EC4F3F86E21BF2D5882852788B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="65DE74D6469ECDF8E8FE091A88640106">Dorgham, Mohamed M.</mods:namePart>
</mods:name>
<mods:typeOfResource id="4C589B5295A542D150CB2104C44C7B2C">text</mods:typeOfResource>
<mods:relatedItem id="B0572B5B2CA9057DC452BD13164D767B" type="host">
<mods:titleInfo id="AD3E9DAFF6CC30C8CBE6789E28B4AF23">
<mods:title id="BA7F8FCF5725C362F541D5660C37FE82">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="A139E7BB73FF81767DC1E851F5CE64F5">
<mods:date id="ED71296FDF7F19B80EDC99AAC562CF9E">2017</mods:date>
<mods:detail id="40340080132A624AD31D15CC60F91A30" type="volume">
<mods:number id="9C003F4DA20CF319E24AFE71023F11EC">4276</mods:number>
</mods:detail>
<mods:detail id="C12E7C95FB8EBC7C837307FCAE8EEC51" type="issue">
<mods:number id="F4510789172A7A3F0C06F17333DEB934">4</mods:number>
</mods:detail>
<mods:extent id="14B1218A67960C6C333A31132F2DD540" unit="page">
<mods:start id="BD6B4C38020DD06A895D42A6D354470F">582</mods:start>
<mods:end id="47C9018BFCD60E5662E13FFE61673E61">590</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="E9DC5B01A62E6FA1EF314F5385F2A220">journal article</mods:classification>
<mods:identifier id="D88304375B2B3D5A639E7E5018D2B736" type="CLB-Dataset">32840</mods:identifier>
<mods:identifier id="484D54697459D5BD1B05D41B0F6B6F64" type="DOI">10.11646/zootaxa.4276.4.9</mods:identifier>
<mods:identifier id="64EE91AAF3FE61D44AF1BFA3CB7CDC9B" type="GBIF-Dataset">0081d7e4-e694-400c-8683-132166e3e038</mods:identifier>
<mods:identifier id="DB312FDE60675554774A8AF152DB9A92" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="5DA454BC7AA9AC0A1DF7276766C8BD8D" type="Zenodo-Dep">808096</mods:identifier>
<mods:identifier id="A064096A5099296D2BEB7A04E3B3FA91" type="ZooBank">CA6D6102-957C-43DB-8278-861564992F4B</mods:identifier>
</mods:mods>
<treatment id="03E89C04A020000542F2D2CEE5393A39" ID-DOI="http://doi.org/10.5281/zenodo.6052037" ID-GBIF-Taxon="131878179" ID-Zenodo-Dep="6052037" LSID="urn:lsid:plazi:treatment:03E89C04A020000542F2D2CEE5393A39" httpUri="http://treatment.plazi.org/id/03E89C04A020000542F2D2CEE5393A39" lastPageId="5" lastPageNumber="587" pageId="2" pageNumber="584">
<subSubSection id="C35B7E99A020000242F2D2CEE7333BD0" pageId="2" pageNumber="584" type="nomenclature">
<paragraph id="8BFE2D12A020000242F2D2CEE40E3BB1" blockId="2.[151,566,1182,1241]" box="[151,566,1182,1208]" pageId="2" pageNumber="584">
<heading id="D0B69A7EA020000242F2D2CEE40E3BB1" bold="true" box="[151,566,1182,1208]" fontSize="11" level="1" pageId="2" pageNumber="584" reason="1">
<emphasis id="B935F100A020000242F2D2CEE40E3BB1" bold="true" box="[151,566,1182,1208]" pageId="2" pageNumber="584">
<taxonomicName id="4C415691A020000242F2D2CEE7D43BB1" authority="Abdelsalam &amp; Taylor &amp; Dorgham, 2017" authorityName="Abdelsalam &amp; Taylor &amp; Dorgham" authorityYear="2017" box="[151,492,1182,1208]" class="Gymnolaemata" family="Lanceoporidae" genus="Calyptotheca" kingdom="Animalia" order="Cheilostomatida" pageId="2" pageNumber="584" phylum="Bryozoa" rank="species" species="alexandriensis" status="sp. nov.">
<emphasis id="B935F100A020000242F2D2CEE7D43BB1" bold="true" box="[151,492,1182,1208]" italics="true" pageId="2" pageNumber="584">Calyptotheca alexandriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2064C7BA02000024396D2CEE40E3BB1" box="[499,566,1182,1208]" pageId="2" pageNumber="584" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BFE2D12A020000242F2D290E7333BD0" blockId="2.[151,566,1182,1241]" box="[151,267,1216,1241]" pageId="2" pageNumber="584">
(
<figureCitation id="137A3197A020000242FAD290E73B3BD0" box="[159,259,1216,1241]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="3.[151,250,1311,1333]" captionStartId-1="3.[151,250,1916,1938]" captionStartId-2="4.[151,250,1829,1851]" captionTargetBox-0="[187,1417,818,1260]" captionTargetBox-1="[181,1403,1435,1866]" captionTargetBox-2="[243,1382,449,1784]" captionTargetId-0="figure@3.[151,1436,794,1289]" captionTargetId-1="figure@3.[151,1436,1413,1894]" captionTargetId-2="figure@4.[198,1422,415,1842]" captionTargetPageId-0="3" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 2. Calyptotheca alexandriensis n. sp. A, fresh specimen showing the foliaceous colony-form and deep orange colour; paratype, TBAB BR- 2016 - 2 - 11. B, C, luxuriant growth of colonies on pilings in the Eastern Harbour, Alexandria." captionText-1="FIGURE 3. Calyptotheca alexandriensis n. sp., scanning electron micrographs of the holotype, TBAB BR- 2015 - 2 - 10. A, group of zooids with locations of undeveloped ovicells marked by areas of fewer pseudopores on the frontal shields of the distal zooids. B, primary orifice and suboral avicularium. C, ovicells. Scale bars: A = 500 µm; B = 100 µm; C = 200 µm." captionText-2="FIGURE 4. Calyptotheca alexandriensis n. sp., scanning electron micrographs of the paratype, NHMUK 2016.12.22.1; A – D are of a bleached fragment, E and F of an unbleached fragment. A, group of zooids, some with complete ovicells, others with undeveloped ovicells, and some with orifices covered by closure plates. B, zooid with an undeveloped ovicell and a closure plate between two zooids with complete ovicells. C, oblique view of growing edge showing septulae in transverse vertical walls. D, primary orifice, condyles and suboral avicularium in a zooid viewed obliquely. E, zooid with in-situ operculum and a partly formed ovicell showing the floor of the entooecium. F, oblique view to show the in-situ mandible of a suboral avicularium. Scale bars: A, C = 500 µm; B = 200 µm; D – F = 100 µm." httpUri-0="https://zenodo.org/record/808100/files/figure.png" httpUri-1="https://zenodo.org/record/808102/files/figure.png" httpUri-2="https://zenodo.org/record/808104/files/figure.png" pageId="2" pageNumber="584">Figs 2–4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35B7E99A020000242F2D358E7383A84" pageId="2" pageNumber="584" type="materials_examined">
<paragraph id="8BFE2D12A020000242F2D358E7383A84" blockId="2.[151,1436,1288,2033]" pageId="2" pageNumber="584">
<materialsCitation id="3B29274FA020000242F2D358E7383A84" ID-GBIF-Occurrence="1563874603" collectingDate="2015-12" collectionCode="TBAB, BR-, NHMUK" collectorName="Mediterranean &amp; Mohamed M. &amp; Khaled M." location="Alexandria, Eastern" pageId="2" pageNumber="584" specimenCode="BR-2015-2-10, BR-2016-2-11, NHMUK 2016.12" specimenCount="3" stateProvince="Alexandria, Eastern" typeStatus="holotype">
<emphasis id="B935F100A020000242F2D358E7BA3A28" bold="true" box="[151,386,1288,1313]" pageId="2" pageNumber="584">Material examined.</emphasis>
<typeStatus id="54FA93B0A020000243EED359E7C93A28" box="[395,497,1289,1313]" pageId="2" pageNumber="584" type="holotype">
<emphasis id="B935F100A020000243EED359E7C93A28" box="[395,497,1289,1313]" italics="true" pageId="2" pageNumber="584">Holotype</emphasis>
</typeStatus>
:
<collectionCode id="ED50B5D7A02000024067D359E4753A28" box="[514,589,1289,1313]" pageId="2" pageNumber="584">TBAB</collectionCode>
<specimenCode id="DBE78569A02000024033D358E4C23A28" box="[598,762,1288,1313]" pageId="2" pageNumber="584">
<collectionCode id="ED50B5D7A02000024033D358E4BE3A28" box="[598,646,1288,1313]" pageId="2" pageNumber="584">BR-</collectionCode>
<date id="FFFF0BD2A020000240E2D358E4C23A28" box="[647,762,1288,1313]" pageId="2" pageNumber="584" value="2015-10-02">2015-2-10</date>
</specimenCode>
(
<figureCitation id="137A3197A0200002416ED358E5683A28" box="[779,848,1288,1313]" captionStart="FIGURE 3" captionStartId="3.[151,250,1916,1938]" captionTargetBox="[181,1403,1435,1866]" captionTargetId="figure@3.[151,1436,1413,1894]" captionTargetPageId="3" captionText="FIGURE 3. Calyptotheca alexandriensis n. sp., scanning electron micrographs of the holotype, TBAB BR- 2015 - 2 - 10. A, group of zooids with locations of undeveloped ovicells marked by areas of fewer pseudopores on the frontal shields of the distal zooids. B, primary orifice and suboral avicularium. C, ovicells. Scale bars: A = 500 µm; B = 100 µm; C = 200 µm." httpUri="https://zenodo.org/record/808102/files/figure.png" pageId="2" pageNumber="584">Fig. 3</figureCitation>
),
<collectorName id="26B448C4A0200002410CD358E2373A28" box="[873,1039,1288,1313]" pageId="2" pageNumber="584">Mediterranean</collectorName>
,
<collectingRegion id="4985E3F0A02000024679D358E2C43A29" box="[1052,1276,1288,1313]" isEnumeration="true" pageId="2" pageNumber="584">Alexandria, Eastern</collectingRegion>
Harbour, 0–1 metre depth on metallic marina supports, collected by
<collectorName id="26B448C4A0200002414ED37CE5F23A4C" box="[811,970,1324,1349]" pageId="2" pageNumber="584">Mohamed M.</collectorName>
Dorgham and
<collectorName id="26B448C4A020000246E1D37CE33A3A4C" box="[1156,1282,1324,1349]" pageId="2" pageNumber="584">Khaled M.</collectorName>
Abdelsalam,
<collectingDate id="EFBBF23AA020000242F2D300E76D3A60" box="[151,341,1360,1385]" pageId="2" pageNumber="584" value="2015-12">
<date id="FFFF0BD2A020000242F2D300E76D3A60" box="[151,341,1360,1385]" pageId="2" pageNumber="584" value="2015-12">December 2015</date>
</collectingDate>
.
<typeStatus id="54FA93B0A02000024302D301E7E03A61" box="[359,472,1361,1384]" pageId="2" pageNumber="584" type="paratype">
<emphasis id="B935F100A02000024302D301E7E03A61" box="[359,472,1361,1384]" italics="true" pageId="2" pageNumber="584">Paratypes</emphasis>
</typeStatus>
:
<collectionCode id="ED50B5D7A0200002438AD301E4023A60" box="[495,570,1361,1385]" pageId="2" pageNumber="584">TBAB</collectionCode>
<specimenCode id="DBE78569A0200002402DD300E4D23A60" box="[584,746,1360,1385]" pageId="2" pageNumber="584">
<collectionCode id="ED50B5D7A0200002402DD300E4413A60" box="[584,633,1360,1385]" pageId="2" pageNumber="584">BR-</collectionCode>
<date id="FFFF0BD2A0200002401CD300E4D23A60" box="[633,746,1360,1385]" pageId="2" pageNumber="584" value="2016-11-02">2016-2-11</date>
</specimenCode>
(
<figureCitation id="137A3197A02000024167D300E5073A60" box="[770,831,1360,1385]" captionStart="FIGURE 2" captionStartId="3.[151,250,1311,1333]" captionTargetBox="[187,1417,818,1260]" captionTargetId="figure@3.[151,1436,794,1289]" captionTargetPageId="3" captionText="FIGURE 2. Calyptotheca alexandriensis n. sp. A, fresh specimen showing the foliaceous colony-form and deep orange colour; paratype, TBAB BR- 2016 - 2 - 11. B, C, luxuriant growth of colonies on pilings in the Eastern Harbour, Alexandria." httpUri="https://zenodo.org/record/808100/files/figure.png" pageId="2" pageNumber="584">Fig.2</figureCitation>
A),
<specimenCode id="DBE78569A02000024114D301E2703A60" box="[881,1096,1360,1385]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="2" pageNumber="584">
<collectionCode id="ED50B5D7A02000024114D301E5E33A61" box="[881,987,1361,1384]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="2" pageNumber="584">NHMUK</collectionCode>
2016.12
</specimenCode>
.22.1 (
<figureCitation id="137A3197A020000246F1D300E2D83A60" box="[1172,1248,1360,1385]" captionStart="FIGURE 4" captionStartId="4.[151,250,1829,1851]" captionTargetBox="[243,1382,449,1784]" captionTargetId="figure@4.[198,1422,415,1842]" captionTargetPageId="4" captionText="FIGURE 4. Calyptotheca alexandriensis n. sp., scanning electron micrographs of the paratype, NHMUK 2016.12.22.1; A – D are of a bleached fragment, E and F of an unbleached fragment. A, group of zooids, some with complete ovicells, others with undeveloped ovicells, and some with orifices covered by closure plates. B, zooid with an undeveloped ovicell and a closure plate between two zooids with complete ovicells. C, oblique view of growing edge showing septulae in transverse vertical walls. D, primary orifice, condyles and suboral avicularium in a zooid viewed obliquely. E, zooid with in-situ operculum and a partly formed ovicell showing the floor of the entooecium. F, oblique view to show the in-situ mandible of a suboral avicularium. Scale bars: A, C = 500 µm; B = 200 µm; D – F = 100 µm." httpUri="https://zenodo.org/record/808104/files/figure.png" pageId="2" pageNumber="584">Fig. 4</figureCitation>
), details as for
<typeStatus id="54FA93B0A020000242F2D324E6C53A84" box="[151,253,1396,1421]" pageId="2" pageNumber="584" type="holotype">holotype</typeStatus>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C35B7E99A020000242A2D3C8E5213AB8" box="[199,793,1432,1457]" pageId="2" pageNumber="584" type="etymology">
<paragraph id="8BFE2D12A020000242A2D3C8E5213AB8" blockId="2.[151,1436,1288,2033]" box="[199,793,1432,1457]" pageId="2" pageNumber="584">
<emphasis id="B935F100A020000242A2D3C8E7763AB8" bold="true" box="[199,334,1432,1457]" pageId="2" pageNumber="584">Etymology.</emphasis>
Named for
<collectingRegion id="4985E3F0A020000243B1D3C8E46C3AB8" box="[468,596,1432,1457]" country="Egypt" name="Al Iskandariyah" pageId="2" pageNumber="584">Alexandria</collectingRegion>
, the
<typeStatus id="54FA93B0A020000240E2D3C9E48F3AB8" box="[647,695,1433,1457]" pageId="2" pageNumber="584">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C35B7E99A020000342A2D3EDE4353E49" lastPageId="3" lastPageNumber="585" pageId="2" pageNumber="584" type="description">
<paragraph id="8BFE2D12A020000242A2D3EDE3643948" blockId="2.[151,1436,1288,2033]" pageId="2" pageNumber="584">
<emphasis id="B935F100A020000242A2D3EDE7613ADF" bold="true" box="[199,345,1469,1494]" pageId="2" pageNumber="584">Description.</emphasis>
Colony erect, foliaceous, comprising an open honeycomb of anastomosing bifoliate fronds forming a spheroidal mass (
<figureCitation id="137A3197A0200002438CD3B0E40C3AF0" box="[489,564,1504,1529]" captionStart="FIGURE 2" captionStartId="3.[151,250,1311,1333]" captionTargetBox="[187,1417,818,1260]" captionTargetId="figure@3.[151,1436,794,1289]" captionTargetPageId="3" captionText="FIGURE 2. Calyptotheca alexandriensis n. sp. A, fresh specimen showing the foliaceous colony-form and deep orange colour; paratype, TBAB BR- 2016 - 2 - 11. B, C, luxuriant growth of colonies on pilings in the Eastern Harbour, Alexandria." httpUri="https://zenodo.org/record/808100/files/figure.png" pageId="2" pageNumber="584">Fig. 2</figureCitation>
A). Intracolonial overgrowths occasionally developed. Uniporous septulae present in lateral and transverse vertical walls, situated close to the basal wall (
<figureCitation id="137A3197A02000024675D055E26D3914" box="[1040,1109,1541,1566]" captionStart="FIGURE 4" captionStartId="4.[151,250,1829,1851]" captionTargetBox="[243,1382,449,1784]" captionTargetId="figure@4.[198,1422,415,1842]" captionTargetPageId="4" captionText="FIGURE 4. Calyptotheca alexandriensis n. sp., scanning electron micrographs of the paratype, NHMUK 2016.12.22.1; A – D are of a bleached fragment, E and F of an unbleached fragment. A, group of zooids, some with complete ovicells, others with undeveloped ovicells, and some with orifices covered by closure plates. B, zooid with an undeveloped ovicell and a closure plate between two zooids with complete ovicells. C, oblique view of growing edge showing septulae in transverse vertical walls. D, primary orifice, condyles and suboral avicularium in a zooid viewed obliquely. E, zooid with in-situ operculum and a partly formed ovicell showing the floor of the entooecium. F, oblique view to show the in-situ mandible of a suboral avicularium. Scale bars: A, C = 500 µm; B = 200 µm; D – F = 100 µm." httpUri="https://zenodo.org/record/808104/files/figure.png" pageId="2" pageNumber="584">Fig. 4</figureCitation>
C). Colour vivid deep orange to red when alive (
<figureCitation id="137A3197A0200002430DD078E7923948" box="[360,426,1576,1601]" captionStart="FIGURE 2" captionStartId="3.[151,250,1311,1333]" captionTargetBox="[187,1417,818,1260]" captionTargetId="figure@3.[151,1436,794,1289]" captionTargetPageId="3" captionText="FIGURE 2. Calyptotheca alexandriensis n. sp. A, fresh specimen showing the foliaceous colony-form and deep orange colour; paratype, TBAB BR- 2016 - 2 - 11. B, C, luxuriant growth of colonies on pilings in the Eastern Harbour, Alexandria." httpUri="https://zenodo.org/record/808100/files/figure.png" pageId="2" pageNumber="584">Fig. 2</figureCitation>
A–C), fading to pale brown in spirit or dry. Ancestrula and early astogeny unknown.
</paragraph>
<paragraph id="8BFE2D12A020000242A2D01DE2A938C7" blockId="2.[151,1436,1288,2033]" pageId="2" pageNumber="584">
Autozooids subrectangular, about 1.75 x longer than wide (
<figureCitation id="137A3197A0200002410BD01DE58E396C" box="[878,950,1613,1638]" captionStart="FIGURE 3" captionStartId="3.[151,250,1916,1938]" captionTargetBox="[181,1403,1435,1866]" captionTargetId="figure@3.[151,1436,1413,1894]" captionTargetPageId="3" captionText="FIGURE 3. Calyptotheca alexandriensis n. sp., scanning electron micrographs of the holotype, TBAB BR- 2015 - 2 - 10. A, group of zooids with locations of undeveloped ovicells marked by areas of fewer pseudopores on the frontal shields of the distal zooids. B, primary orifice and suboral avicularium. C, ovicells. Scale bars: A = 500 µm; B = 100 µm; C = 200 µm." httpUri="https://zenodo.org/record/808102/files/figure.png" pageId="2" pageNumber="584">Figs 3</figureCitation>
A, 4A). Frontal shield moderately convex, densely pseudoporous, pseudopores subcircular, about 10–35 µm in diameter, decreasing in size from the centre to the edge of the frontal shield, sunken, calcification around the pseudopores in the form of a polygonal network with low pustules often present at the corners; marginal areolar pores not clearly distinct from pseudopores; zooidal boundaries well defined, marked by thin but well-defined salient walls. Primary orifice broader than long (
<figureCitation id="137A3197A02000024759D08DE3BA39FC" box="[1340,1410,1757,1782]" captionStart="FIGURE 3" captionStartId="3.[151,250,1916,1938]" captionTargetBox="[181,1403,1435,1866]" captionTargetId="figure@3.[151,1436,1413,1894]" captionTargetPageId="3" captionText="FIGURE 3. Calyptotheca alexandriensis n. sp., scanning electron micrographs of the holotype, TBAB BR- 2015 - 2 - 10. A, group of zooids with locations of undeveloped ovicells marked by areas of fewer pseudopores on the frontal shields of the distal zooids. B, primary orifice and suboral avicularium. C, ovicells. Scale bars: A = 500 µm; B = 100 µm; C = 200 µm." httpUri="https://zenodo.org/record/808102/files/figure.png" pageId="2" pageNumber="584">Figs 3</figureCitation>
B, 4D), inclined to face distofrontally, with a broad, U-shaped sinus divided from the semielliptical poster by a pair of rounded, non-serrated condyles. Closure plates cover some orifices, imperforate and with an irregularly granular surface (
<figureCitation id="137A3197A0200002429DD118E7063868" box="[248,318,1864,1889]" captionStart="FIGURE 4" captionStartId="4.[151,250,1829,1851]" captionTargetBox="[243,1382,449,1784]" captionTargetId="figure@4.[198,1422,415,1842]" captionTargetPageId="4" captionText="FIGURE 4. Calyptotheca alexandriensis n. sp., scanning electron micrographs of the paratype, NHMUK 2016.12.22.1; A – D are of a bleached fragment, E and F of an unbleached fragment. A, group of zooids, some with complete ovicells, others with undeveloped ovicells, and some with orifices covered by closure plates. B, zooid with an undeveloped ovicell and a closure plate between two zooids with complete ovicells. C, oblique view of growing edge showing septulae in transverse vertical walls. D, primary orifice, condyles and suboral avicularium in a zooid viewed obliquely. E, zooid with in-situ operculum and a partly formed ovicell showing the floor of the entooecium. F, oblique view to show the in-situ mandible of a suboral avicularium. Scale bars: A, C = 500 µm; B = 200 µm; D – F = 100 µm." httpUri="https://zenodo.org/record/808104/files/figure.png" pageId="2" pageNumber="584">Fig. 4</figureCitation>
B). Ovicells present in a high proportion of zooids (
<figureCitation id="137A3197A020000241EED118E5EB3868" box="[907,979,1864,1889]" captionStart="FIGURE 3" captionStartId="3.[151,250,1916,1938]" captionTargetBox="[181,1403,1435,1866]" captionTargetId="figure@3.[151,1436,1413,1894]" captionTargetPageId="3" captionText="FIGURE 3. Calyptotheca alexandriensis n. sp., scanning electron micrographs of the holotype, TBAB BR- 2015 - 2 - 10. A, group of zooids with locations of undeveloped ovicells marked by areas of fewer pseudopores on the frontal shields of the distal zooids. B, primary orifice and suboral avicularium. C, ovicells. Scale bars: A = 500 µm; B = 100 µm; C = 200 µm." httpUri="https://zenodo.org/record/808102/files/figure.png" pageId="2" pageNumber="584">Figs 3</figureCitation>
C, 4A, B), hyperstomial, bulbous, wider than long, ectooecium uncalcified, endooecial calcification resembling frontal shield but with smaller, more closely spaced pseudopores; floor of incompletely developed ovicells with a scattering of 10–20 irregularly arranged pores of variable size (
<figureCitation id="137A3197A02000024334D1E5E7AE38C4" box="[337,406,1973,1998]" captionStart="FIGURE 3" captionStartId="3.[151,250,1916,1938]" captionTargetBox="[181,1403,1435,1866]" captionTargetId="figure@3.[151,1436,1413,1894]" captionTargetPageId="3" captionText="FIGURE 3. Calyptotheca alexandriensis n. sp., scanning electron micrographs of the holotype, TBAB BR- 2015 - 2 - 10. A, group of zooids with locations of undeveloped ovicells marked by areas of fewer pseudopores on the frontal shields of the distal zooids. B, primary orifice and suboral avicularium. C, ovicells. Scale bars: A = 500 µm; B = 100 µm; C = 200 µm." httpUri="https://zenodo.org/record/808102/files/figure.png" pageId="2" pageNumber="584">Figs 3</figureCitation>
A, 4A, B). Ovicellate zooids without apparent orificial dimorphism.
</paragraph>
<paragraph id="8BFE2D12A020000342A2D188E27A3E14" blockId="2.[151,1436,1288,2033]" lastBlockId="3.[151,1436,151,320]" lastPageId="3" lastPageNumber="585" pageId="2" pageNumber="584">
Avicularia suboral (
<figureCitation id="137A3197A020000243CDD188E7CA38F8" box="[424,498,2008,2033]" captionStart="FIGURE 3" captionStartId="3.[151,250,1916,1938]" captionTargetBox="[181,1403,1435,1866]" captionTargetId="figure@3.[151,1436,1413,1894]" captionTargetPageId="3" captionText="FIGURE 3. Calyptotheca alexandriensis n. sp., scanning electron micrographs of the holotype, TBAB BR- 2015 - 2 - 10. A, group of zooids with locations of undeveloped ovicells marked by areas of fewer pseudopores on the frontal shields of the distal zooids. B, primary orifice and suboral avicularium. C, ovicells. Scale bars: A = 500 µm; B = 100 µm; C = 200 µm." httpUri="https://zenodo.org/record/808102/files/figure.png" pageId="2" pageNumber="584">Figs 3</figureCitation>
B, 4D, F), present in the majority of, possibly all, zooids, oriented predominantly laterally but slightly distally, all avicularia on the same side of a frond apparently oriented in the same direction, either facing left or right; plane of avicularium inclined at a steep angle to the frontal surface of the colony; small in size, about
<quantity id="4CB980F7A02100034370D6B0E78F3FFE" box="[277,439,224,248]" metricMagnitude="-5" metricUnit="m" metricValue="8.499999999999998" metricValueMax="9.0" metricValueMin="8.0" pageId="3" pageNumber="585" unit="mm" value="0.08499999999999999" valueMax="0.09" valueMin="0.08">0.08–0.09 mm</quantity>
long; opesia semicircular; pivotal bar complete, more or less straight, longer than opesia; rostrum arch-shaped, with a hook-like distal end. Vicarious avicularia not observed.
</paragraph>
<paragraph id="8BFE2D12A021000342A2D777E4353E49" blockId="3.[151,1436,151,320]" box="[199,525,295,320]" pageId="3" pageNumber="585">
<emphasis id="B935F100A021000342A2D777E7453E49" bold="true" box="[199,381,295,320]" pageId="3" pageNumber="585">Measurements.</emphasis>
See
<tableCitation id="C6C318A9A021000343D1D777E43F3E49" box="[436,519,295,320]" captionStart="TABLE 1" captionStartId="3.[151,239,368,391]" captionText="TABLE 1. Measurements (in mm) of Calyptotheca alexandriensis n. sp." pageId="3" pageNumber="585">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
<caption id="DF3E7D9AA021000342F2D720E5983E8E" box="[151,928,367,391]" pageId="3" pageNumber="585">
<paragraph id="8BFE2D12A021000342F2D720E5983E8E" blockId="3.[151,928,367,391]" box="[151,928,367,391]" pageId="3" pageNumber="585">
<emphasis id="B935F100A021000342F2D720E7323E8E" bold="true" box="[151,266,368,391]" pageId="3" pageNumber="585">TABLE 1.</emphasis>
Measurements (in mm) of
<taxonomicName id="4C415691A02100034048D73FE5643E8E" authority="Abdelsalam &amp; Taylor &amp; Dorgham, 2017" authorityName="Abdelsalam &amp; Taylor &amp; Dorgham" authorityYear="2017" box="[557,860,367,391]" class="Gymnolaemata" family="Lanceoporidae" genus="Calyptotheca" kingdom="Animalia" order="Cheilostomatida" pageId="3" pageNumber="585" phylum="Bryozoa" rank="species" species="alexandriensis" status="sp. nov.">
<emphasis id="B935F100A02100034048D73FE5643E8E" box="[557,860,367,391]" italics="true" pageId="3" pageNumber="585">Calyptotheca alexandriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2064C7BA02100034101D720E5983E8E" box="[868,928,368,391]" pageId="3" pageNumber="585" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BFE2D12A021000343F0D7F3E4C43DE1" pageId="3" pageNumber="585">
<table id="F941DFB2A021FFFF42FAD7F2E2A93DE1" box="[159,1169,418,744]" gridcols="5" gridrows="8" pageId="3" pageNumber="585">
<tr id="35712F50A021FFFF42FAD7F2E2A93EB1" box="[159,1169,418,440]" gridrow="0" pageId="3" pageNumber="585" rowspan-0="1">
<th id="76A0462CA021FFFF4332D7F2E46F3EB1" box="[343,599,418,440]" gridcol="1" gridrow="0" pageId="3" pageNumber="585">Mean</th>
<th id="76A0462CA021FFFF403BD7F2E4C43EB1" box="[606,764,418,440]" gridcol="2" gridrow="0" pageId="3" pageNumber="585">Range</th>
<th id="76A0462CA021FFFF41E7D7F2E5963EB1" box="[898,942,418,440]" gridcol="3" gridrow="0" pageId="3" pageNumber="585">SD</th>
<th id="76A0462CA021FFFF461DD7F2E2A93EB1" box="[1144,1169,418,440]" gridcol="4" gridrow="0" pageId="3" pageNumber="585">N</th>
</tr>
<tr id="35712F50A021FFFF42FAD79CE2A93EEB" box="[159,1169,460,482]" gridrow="1" pageId="3" pageNumber="585">
<th id="76A0462CA021FFFF42FAD79CE7683EEB" box="[159,336,460,482]" gridcol="0" gridrow="1" pageId="3" pageNumber="585">Zooid length</th>
<td id="76A0462CA021FFFF4332D79CE46F3EEB" box="[343,599,460,482]" gridcol="1" gridrow="1" pageId="3" pageNumber="585">0.56</td>
<td id="76A0462CA021FFFF403BD79CE4C43EEB" box="[606,764,460,482]" gridcol="2" gridrow="1" pageId="3" pageNumber="585">0.46–0.65</td>
<td id="76A0462CA021FFFF41E7D79CE5963EEB" box="[898,942,460,482]" gridcol="3" gridrow="1" pageId="3" pageNumber="585">0.06</td>
<td id="76A0462CA021FFFF461DD79CE2A93EEB" box="[1144,1169,460,482]" gridcol="4" gridrow="1" pageId="3" pageNumber="585">12</td>
</tr>
<tr id="35712F50A021FFFF42FAD7A5E2A93D02" box="[159,1169,501,523]" gridrow="2" pageId="3" pageNumber="585">
<th id="76A0462CA021FFFF42FAD7A5E7683D02" box="[159,336,501,523]" gridcol="0" gridrow="2" pageId="3" pageNumber="585">Zooid width</th>
<td id="76A0462CA021FFFF4332D7A5E46F3D02" box="[343,599,501,523]" gridcol="1" gridrow="2" pageId="3" pageNumber="585">0.32</td>
<td id="76A0462CA021FFFF403BD7A5E4C43D02" box="[606,764,501,523]" gridcol="2" gridrow="2" pageId="3" pageNumber="585">0.27–0.35</td>
<td id="76A0462CA021FFFF41E7D7A5E5963D02" box="[898,942,501,523]" gridcol="3" gridrow="2" pageId="3" pageNumber="585">0.03</td>
<td id="76A0462CA021FFFF461DD7A5E2A93D02" box="[1144,1169,501,523]" gridcol="4" gridrow="2" pageId="3" pageNumber="585">12</td>
</tr>
<tr id="35712F50A021FFFF42FAD44EE2A93D3D" box="[159,1169,542,564]" gridrow="3" pageId="3" pageNumber="585">
<th id="76A0462CA021FFFF42FAD44EE7683D3D" box="[159,336,542,564]" gridcol="0" gridrow="3" pageId="3" pageNumber="585">Orifice length</th>
<td id="76A0462CA021FFFF4332D44EE46F3D3D" box="[343,599,542,564]" gridcol="1" gridrow="3" pageId="3" pageNumber="585">0.13</td>
<td id="76A0462CA021FFFF403BD44EE4C43D3D" box="[606,764,542,564]" gridcol="2" gridrow="3" pageId="3" pageNumber="585">0.11–0.14</td>
<td id="76A0462CA021FFFF41E7D44EE5963D3D" box="[898,942,542,564]" gridcol="3" gridrow="3" pageId="3" pageNumber="585">0.01</td>
<td id="76A0462CA021FFFF461DD44EE2A93D3D" box="[1144,1169,542,564]" gridcol="4" gridrow="3" pageId="3" pageNumber="585">10</td>
</tr>
<tr id="35712F50A021FFFF42FAD418E2A93D57" box="[159,1169,584,606]" gridrow="4" pageId="3" pageNumber="585">
<th id="76A0462CA021FFFF42FAD418E7683D57" box="[159,336,584,606]" gridcol="0" gridrow="4" pageId="3" pageNumber="585">Orifice width</th>
<td id="76A0462CA021FFFF4332D418E46F3D57" box="[343,599,584,606]" gridcol="1" gridrow="4" pageId="3" pageNumber="585">0.17</td>
<td id="76A0462CA021FFFF403BD418E4C43D57" box="[606,764,584,606]" gridcol="2" gridrow="4" pageId="3" pageNumber="585">0.17–0.18</td>
<td id="76A0462CA021FFFF41E7D418E5963D57" box="[898,942,584,606]" gridcol="3" gridrow="4" pageId="3" pageNumber="585">0.01</td>
<td id="76A0462CA021FFFF461DD418E2A93D57" box="[1144,1169,584,606]" gridcol="4" gridrow="4" pageId="3" pageNumber="585">10</td>
</tr>
<tr id="35712F50A021FFFF42FAD421E2A93D8E" box="[159,1169,625,647]" gridrow="5" pageId="3" pageNumber="585">
<th id="76A0462CA021FFFF42FAD421E7683D8E" box="[159,336,625,647]" gridcol="0" gridrow="5" pageId="3" pageNumber="585">Ovicell length</th>
<td id="76A0462CA021FFFF4332D421E46F3D8E" box="[343,599,625,647]" gridcol="1" gridrow="5" pageId="3" pageNumber="585">0.26</td>
<td id="76A0462CA021FFFF403BD421E4C43D8E" box="[606,764,625,647]" gridcol="2" gridrow="5" pageId="3" pageNumber="585">0.25–0.27</td>
<td id="76A0462CA021FFFF41E7D421E5963D8E" box="[898,942,625,647]" gridcol="3" gridrow="5" pageId="3" pageNumber="585">0.01</td>
<td id="76A0462CA021FFFF461DD421E2A93D8E" box="[1144,1169,625,647]" gridcol="4" gridrow="5" pageId="3" pageNumber="585">10</td>
</tr>
<tr id="35712F50A021FFFF42FAD4CAE2A93DB9" box="[159,1169,666,688]" gridrow="6" pageId="3" pageNumber="585">
<th id="76A0462CA021FFFF42FAD4CAE7683DB9" box="[159,336,666,688]" gridcol="0" gridrow="6" pageId="3" pageNumber="585">Ovicell width</th>
<td id="76A0462CA021FFFF4332D4CAE46F3DB9" box="[343,599,666,688]" gridcol="1" gridrow="6" pageId="3" pageNumber="585">0.35</td>
<td id="76A0462CA021FFFF403BD4CAE4C43DB9" box="[606,764,666,688]" gridcol="2" gridrow="6" pageId="3" pageNumber="585">0.33–0.37</td>
<td id="76A0462CA021FFFF41E7D4CAE5963DB9" box="[898,942,666,688]" gridcol="3" gridrow="6" pageId="3" pageNumber="585">0.02</td>
<td id="76A0462CA021FFFF461DD4CAE2A93DB9" box="[1144,1169,666,688]" gridcol="4" gridrow="6" pageId="3" pageNumber="585">10</td>
</tr>
<tr id="35712F50A021FFFF42FAD481E2A93DE1" box="[159,1169,721,744]" gridrow="7" pageId="3" pageNumber="585" rowspan-3="1" rowspan-4="1">
<th id="76A0462CA021FFFF42FAD481E7683DE1" box="[159,336,721,744]" gridcol="0" gridrow="7" pageId="3" pageNumber="585">SD, standard</th>
<td id="76A0462CA021FFFF4332D481E46F3DE1" box="[343,599,721,744]" gridcol="1" gridrow="7" pageId="3" pageNumber="585">deviation; N, number of</td>
<td id="76A0462CA021FFFF403BD481E4C43DE1" box="[606,764,721,744]" gridcol="2" gridrow="7" pageId="3" pageNumber="585">determinations</td>
</tr>
</table>
</paragraph>
<caption id="DF3E7D9AA021000342F2D34FE3213A5D" httpUri="https://zenodo.org/record/808100/files/figure.png" pageId="3" pageNumber="585" targetBox="[187,1417,818,1260]" targetPageId="3">
<paragraph id="8BFE2D12A021000342F2D34FE3213A5D" blockId="3.[151,1436,1311,1364]" pageId="3" pageNumber="585">
<emphasis id="B935F100A021000342F2D34FE72A3A3D" bold="true" box="[151,274,1311,1333]" pageId="3" pageNumber="585">FIGURE 2.</emphasis>
<taxonomicName id="4C415691A0210003437DD34FE4003A3C" authority="Abdelsalam &amp; Taylor &amp; Dorgham, 2017" authorityName="Abdelsalam &amp; Taylor &amp; Dorgham" authorityYear="2017" box="[280,568,1311,1333]" class="Gymnolaemata" family="Lanceoporidae" genus="Calyptotheca" kingdom="Animalia" order="Cheilostomatida" pageId="3" pageNumber="585" phylum="Bryozoa" rank="species" species="alexandriensis" status="sp. nov.">
<emphasis id="B935F100A0210003437DD34FE4003A3C" box="[280,568,1311,1333]" italics="true" pageId="3" pageNumber="585">Calyptotheca alexandriensis</emphasis>
</taxonomicName>
<emphasis id="B935F100A0210003405BD34FE4B63A3D" bold="true" box="[574,654,1311,1332]" pageId="3" pageNumber="585">
<taxonomicNameLabel id="A2064C7BA0210003405BD34FE44E3A3D" box="[574,630,1311,1332]" pageId="3" pageNumber="585" rank="species">n. sp.</taxonomicNameLabel>
A
</emphasis>
, fresh specimen showing the foliaceous colony-form and deep orange colour; paratype, TBAB BR-2016-2-11. B, C, luxuriant growth of colonies on pilings in the Eastern Harbour, Alexandria.
</paragraph>
</caption>
<caption id="DF3E7D9AA021000342F2D12CE31D38D9" httpUri="https://zenodo.org/record/808102/files/figure.png" pageId="3" pageNumber="585" targetBox="[181,1403,1435,1866]" targetPageId="3">
<paragraph id="8BFE2D12A021000342F2D12CE31D38D9" blockId="3.[151,1436,1916,2000]" pageId="3" pageNumber="585">
<emphasis id="B935F100A021000342F2D12CE72E3898" bold="true" box="[151,278,1916,1938]" pageId="3" pageNumber="585">FIGURE 3.</emphasis>
<taxonomicName id="4C415691A02100034345D12CE47D389B" authority="Abdelsalam &amp; Taylor &amp; Dorgham, 2017" authorityName="Abdelsalam &amp; Taylor &amp; Dorgham" authorityYear="2017" box="[288,581,1916,1938]" class="Gymnolaemata" family="Lanceoporidae" genus="Calyptotheca" kingdom="Animalia" order="Cheilostomatida" pageId="3" pageNumber="585" phylum="Bryozoa" rank="species" species="alexandriensis" status="sp. nov.">
<emphasis id="B935F100A02100034345D12CE47D389B" box="[288,581,1916,1938]" italics="true" pageId="3" pageNumber="585">Calyptotheca alexandriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2064C7BA0210003402BD12DE4B2389B" box="[590,650,1917,1938]" pageId="3" pageNumber="585" rank="species">
<emphasis id="B935F100A0210003402BD12DE4B2389B" bold="true" box="[590,650,1917,1938]" pageId="3" pageNumber="585">n. sp.</emphasis>
</taxonomicNameLabel>
, scanning electron micrographs of the holotype, TBAB BR-2015-2-10.
<emphasis id="B935F100A021000347E6D12CE3AD3898" bold="true" box="[1411,1429,1916,1937]" pageId="3" pageNumber="585">A</emphasis>
, group of zooids with locations of undeveloped ovicells marked by areas of fewer pseudopores on the frontal shields of the distal zooids.
<emphasis id="B935F100A02100034280D1EAE6CE38C6" bold="true" box="[229,246,1978,1999]" pageId="3" pageNumber="585">B</emphasis>
, primary orifice and suboral avicularium.
<emphasis id="B935F100A021000340C6D1EAE48D38D9" bold="true" box="[675,693,1978,2000]" pageId="3" pageNumber="585">C</emphasis>
, ovicells. Scale bars:
<emphasis id="B935F100A021000341F7D1EAE59C38C6" bold="true" box="[914,932,1978,1999]" pageId="3" pageNumber="585">A</emphasis>
= 500 µm;
<emphasis id="B935F100A0210003467ED1EAE21438C6" bold="true" box="[1051,1068,1978,1999]" pageId="3" pageNumber="585">B</emphasis>
= 100 µm;
<emphasis id="B935F100A021000346C6D1EAE28D38D9" bold="true" box="[1187,1205,1978,2000]" pageId="3" pageNumber="585">C</emphasis>
= 200 µm.
</paragraph>
</caption>
<subSubSection id="C35B7E99A026000542A2D6C7E4D63A04" lastPageId="5" lastPageNumber="587" pageId="4" pageNumber="586" type="discussion">
<paragraph id="8BFE2D12A026000442A2D6C7E4903E81" blockId="4.[151,1436,151,392]" pageId="4" pageNumber="586">
<emphasis id="B935F100A026000442A2D6C7E7033FB9" bold="true" box="[199,315,151,176]" pageId="4" pageNumber="586">Remarks.</emphasis>
The shape of the primary orifice, with a broad sinus and condyles, evenly pseudoporous frontal shield, presence of adventitious avicularia and ovicell morphology permit this bryozoan to be assigned to
<taxonomicName id="4C415691A026000442F2D6B0E7143FF1" authorityName="Harmer" authorityYear="1957" box="[151,300,224,248]" class="Gymnolaemata" family="Lanceoporidae" genus="Calyptotheca" kingdom="Animalia" order="Cheilostomatida" pageId="4" pageNumber="586" phylum="Bryozoa" rank="genus">
<emphasis id="B935F100A026000442F2D6B0E7143FF1" box="[151,300,224,248]" italics="true" pageId="4" pageNumber="586">Calyptotheca</emphasis>
</taxonomicName>
. As noted above,
<taxonomicName id="4C415691A02600044074D6B0E49E3FF1" authorityName="Harmer" authorityYear="1957" box="[529,678,224,248]" class="Gymnolaemata" family="Lanceoporidae" genus="Calyptotheca" kingdom="Animalia" order="Cheilostomatida" pageId="4" pageNumber="586" phylum="Bryozoa" rank="genus">
<emphasis id="B935F100A02600044074D6B0E49E3FF1" box="[529,678,224,248]" italics="true" pageId="4" pageNumber="586">Calyptotheca</emphasis>
</taxonomicName>
is a speciose genus (
<bibRefCitation id="EFD050E3A026000441D7D68FE2BE3FF1" author="Bock" box="[946,1158,223,248]" pageId="4" pageNumber="586" refString="Bock, P., Reverter Gil, O. &amp; Hayward, P. (2017) Calyptotheca Harmer, 1957. In: Bock, P. &amp; Gordon, D. (Eds.), World List of Bryozoa. Accessed through: World Register of Marine Species. Available from: http: // marinespecies. org / aphia. php? p = taxdetails &amp; id = 110934 (accessed 11 April 2017)" type="journal article" year="2017">
Bock
<emphasis id="B935F100A02600044199D6B1E2043FF1" box="[1020,1084,224,248]" italics="true" pageId="4" pageNumber="586">et al.</emphasis>
2017
</bibRefCitation>
). There is considerable variability in the shape, disposition and orientation of the adventitious avicularia.
<bibRefCitation id="EFD050E3A02600044629D754E3A43E15" author="Cumming" box="[1100,1436,260,285]" pageId="4" pageNumber="586" refString="Cumming, R. L. &amp; Tilbrook, K. J. (2014) Six species of Calyptotheca (Bryozoa, Cheilostomata, Lanceoporidae) from the Gulf of Carpentaria and northern Australia, with description of a new species. Zootaxa, 3827 (2), 147 - 169." type="journal article" year="2014">Cumming &amp; Tilbrook (2014)</bibRefCitation>
identified a subgroup of species having suboral avicularia, like that seen in the new species from
<collectingCountry id="F3566D82A026000446BBD778E31D3E49" box="[1246,1317,296,320]" name="Egypt" pageId="4" pageNumber="586">Egypt</collectingCountry>
. However, in most of these species the avicularium is oriented parallel to the length of the autozooid, whereas in
<taxonomicName id="4C415691A026000447E7D71CE7023E81" authority="Abdelsalam &amp; Taylor &amp; Dorgham, 2017" authorityName="Abdelsalam &amp; Taylor &amp; Dorgham" authorityYear="2017" class="Gymnolaemata" family="Lanceoporidae" genus="Calyptotheca" kingdom="Animalia" order="Cheilostomatida" pageId="4" pageNumber="586" phylum="Bryozoa" rank="species" species="alexandriensis" status="sp. nov.">
<emphasis id="B935F100A026000447E7D71CE7023E81" italics="true" pageId="4" pageNumber="586">C. alexandriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2064C7BA02600044324D73FE7B83E81" box="[321,384,367,392]" pageId="4" pageNumber="586" rank="species">
<emphasis id="B935F100A02600044324D73FE7B83E81" bold="true" box="[321,384,367,392]" pageId="4" pageNumber="586">n. sp.</emphasis>
</taxonomicNameLabel>
it is oriented transversely.
</paragraph>
<caption id="DF3E7D9AA026000442F2D175E56738FB" httpUri="https://zenodo.org/record/808104/files/figure.png" pageId="4" pageNumber="586" targetBox="[243,1382,449,1784]" targetPageId="4">
<paragraph id="8BFE2D12A026000442F2D175E56738FB" blockId="4.[151,1436,1828,2035]" pageId="4" pageNumber="586">
<emphasis id="B935F100A026000442F2D175E72B3833" bold="true" box="[151,275,1829,1851]" pageId="4" pageNumber="586">FIGURE 4.</emphasis>
<taxonomicName id="4C415691A0260004437ED174E4053833" authority="Abdelsalam &amp; Taylor &amp; Dorgham, 2017" authorityName="Abdelsalam &amp; Taylor &amp; Dorgham" authorityYear="2017" box="[283,573,1828,1850]" class="Gymnolaemata" family="Lanceoporidae" genus="Calyptotheca" kingdom="Animalia" order="Cheilostomatida" pageId="4" pageNumber="586" phylum="Bryozoa" rank="species" species="alexandriensis" status="sp. nov.">
<emphasis id="B935F100A0260004437ED174E4053833" box="[283,573,1828,1850]" italics="true" pageId="4" pageNumber="586">Calyptotheca alexandriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2064C7BA02600044026D175E4453833" box="[579,637,1829,1850]" pageId="4" pageNumber="586" rank="species">
<emphasis id="B935F100A02600044026D175E4453833" bold="true" box="[579,637,1829,1850]" pageId="4" pageNumber="586">n. sp.</emphasis>
</taxonomicNameLabel>
, scanning electron micrographs of the paratype, NHMUK 2016.12.22.1;
<emphasis id="B935F100A0260004470ED175E3A43833" bold="true" box="[1387,1436,1829,1850]" pageId="4" pageNumber="586">A–D</emphasis>
are of a bleached fragment,
<emphasis id="B935F100A026000443D1D114E7FD3850" bold="true" box="[436,453,1860,1881]" pageId="4" pageNumber="586">E</emphasis>
and
<emphasis id="B935F100A0260004439DD114E43F3850" bold="true" box="[504,519,1860,1881]" pageId="4" pageNumber="586">F</emphasis>
of an unbleached fragment.
<emphasis id="B935F100A02600044149D113E57D3851" bold="true" box="[812,837,1859,1880]" pageId="4" pageNumber="586">A,</emphasis>
group of zooids, some with complete ovicells, others with undeveloped ovicells, and some with orifices covered by closure plates.
<emphasis id="B935F100A026000441E3D132E5AF387E" bold="true" box="[902,919,1890,1911]" pageId="4" pageNumber="586">B</emphasis>
, zooid with an undeveloped ovicell and a closure plate between two zooids with complete ovicells.
<emphasis id="B935F100A026000440C6D1D1E48D389E" bold="true" box="[675,693,1921,1943]" pageId="4" pageNumber="586">C</emphasis>
, oblique view of growing edge showing septulae in transverse vertical walls.
<emphasis id="B935F100A026000442BCD1F0E6D338BC" bold="true" box="[217,235,1952,1973]" pageId="4" pageNumber="586">D</emphasis>
, primary orifice, condyles and suboral avicularium in a zooid viewed obliquely.
<emphasis id="B935F100A02600044679D1F0E21538BC" bold="true" box="[1052,1069,1952,1973]" pageId="4" pageNumber="586">E</emphasis>
, zooid with in-situ operculum and a partly formed ovicell showing the floor of the entooecium.
<emphasis id="B935F100A02600044147D1EEE51738DA" bold="true" box="[802,815,1982,2003]" pageId="4" pageNumber="586">F</emphasis>
, oblique view to show the in-situ mandible of a suboral avicularium. Scale bars:
<emphasis id="B935F100A026000443F4D18DE79B38FB" bold="true" box="[401,419,2013,2034]" pageId="4" pageNumber="586">A</emphasis>
,
<emphasis id="B935F100A026000443D5D18DE7FA38FA" bold="true" box="[432,450,2013,2035]" pageId="4" pageNumber="586">C</emphasis>
= 500 µm;
<emphasis id="B935F100A0260004405CD18DE47238FB" bold="true" box="[569,586,2013,2034]" pageId="4" pageNumber="586">B</emphasis>
= 200 µm;
<emphasis id="B935F100A026000440A4D18DE4D738FB" bold="true" box="[705,751,2013,2034]" pageId="4" pageNumber="586">D–F</emphasis>
= 100 µm.
</paragraph>
</caption>
<paragraph id="8BFE2D12A027000542A2D6C7E7B83D11" blockId="5.[151,1436,151,1328]" pageId="5" pageNumber="587">
Only two species of
<taxonomicName id="4C415691A027000543D6D6C8E4703FB9" authorityName="Harmer" authorityYear="1957" box="[435,584,152,176]" class="Gymnolaemata" family="Lanceoporidae" genus="Calyptotheca" kingdom="Animalia" order="Cheilostomatida" pageId="5" pageNumber="587" phylum="Bryozoa" rank="genus">
<emphasis id="B935F100A027000543D6D6C8E4703FB9" box="[435,584,152,176]" italics="true" pageId="5" pageNumber="587">Calyptotheca</emphasis>
</taxonomicName>
have been recorded previously from the Mediterranean Sea according to a recent review of bryozoans from the Mediterranean Sea (Rosso &amp; Di Martino 2016).
<bibRefCitation id="EFD050E3A02700054609D6ECE31E3FDD" author="Hayward" box="[1132,1318,188,213]" pageId="5" pageNumber="587" refString="Hayward, P. J. (1974) Studies on the cheilostome bryozoan fauna of the Aegean island of Chios. Journal of Natural History, 8, 369 - 402." type="journal article" year="1974">Hayward (1974)</bibRefCitation>
originally described
<taxonomicName id="4C415691A0270005436ED6B0E7CE3FF1" authorityName="Hayward" authorityYear="1974" box="[267,502,224,248]" class="Gymnolaemata" family="Lanceoporidae" genus="Calyptotheca" kingdom="Animalia" order="Cheilostomatida" pageId="5" pageNumber="587" phylum="Bryozoa" rank="species" species="rugosa">
<emphasis id="B935F100A0270005436ED6B0E7CE3FF1" box="[267,502,224,248]" italics="true" pageId="5" pageNumber="587">Calyptotheca rugosa</emphasis>
</taxonomicName>
from Chios in
<collectingCountry id="F3566D82A027000540C3D6B0E4C13FF1" box="[678,761,224,248]" name="Greece" pageId="5" pageNumber="587">Greece</collectingCountry>
. This species is encrusting, unlike
<taxonomicName id="4C415691A027000546E3D6B0E3733FF1" authority="Abdelsalam &amp; Taylor &amp; Dorgham, 2017" authorityName="Abdelsalam &amp; Taylor &amp; Dorgham" authorityYear="2017" box="[1158,1355,224,248]" class="Gymnolaemata" family="Lanceoporidae" genus="Calyptotheca" kingdom="Animalia" order="Cheilostomatida" pageId="5" pageNumber="587" phylum="Bryozoa" rank="species" species="alexandriensis" status="sp. nov.">
<emphasis id="B935F100A027000546E3D6B0E3733FF1" box="[1158,1355,224,248]" italics="true" pageId="5" pageNumber="587">C. alexandriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2064C7BA02700054731D68FE3AC3FF1" box="[1364,1428,223,248]" pageId="5" pageNumber="587" rank="species">
<emphasis id="B935F100A02700054731D68FE3AC3FF1" bold="true" box="[1364,1428,223,248]" pageId="5" pageNumber="587">n. sp.</emphasis>
</taxonomicNameLabel>
, and also differs from
<taxonomicName id="4C415691A027000543F4D754E4603E15" authority="Abdelsalam &amp; Taylor &amp; Dorgham, 2017" authorityName="Abdelsalam &amp; Taylor &amp; Dorgham" authorityYear="2017" box="[401,600,260,284]" class="Gymnolaemata" family="Lanceoporidae" genus="Calyptotheca" kingdom="Animalia" order="Cheilostomatida" pageId="5" pageNumber="587" phylum="Bryozoa" rank="species" species="alexandriensis" status="sp. nov.">
<emphasis id="B935F100A027000543F4D754E4603E15" box="[401,600,260,284]" italics="true" pageId="5" pageNumber="587">C. alexandriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2064C7BA02700054004D754E49A3E14" box="[609,674,260,285]" pageId="5" pageNumber="587" rank="species">
<emphasis id="B935F100A02700054004D754E49A3E14" bold="true" box="[609,674,260,285]" pageId="5" pageNumber="587">n. sp.</emphasis>
</taxonomicNameLabel>
in having rare, rounded suboral avicularia and partially immersed ovicells. A second species of
<taxonomicName id="4C415691A0270005438ED778E4B83E49" authorityName="Harmer" authorityYear="1957" box="[491,640,296,320]" class="Gymnolaemata" family="Lanceoporidae" genus="Calyptotheca" kingdom="Animalia" order="Cheilostomatida" pageId="5" pageNumber="587" phylum="Bryozoa" rank="genus">
<emphasis id="B935F100A0270005438ED778E4B83E49" box="[491,640,296,320]" italics="true" pageId="5" pageNumber="587">Calyptotheca</emphasis>
</taxonomicName>
, found in the Strait of
<collectingCountry id="F3566D82A027000541ECD777E5D63E49" box="[905,1006,295,320]" name="Gibraltar" pageId="5" pageNumber="587">Gibraltar</collectingCountry>
as well as the Atlantic (Azores), was described as
<taxonomicName id="4C415691A02700054351D71CE4C93E6C" authority="Osburn, 1952" authorityName="Osburn" authorityYear="1952" box="[308,753,332,357]" class="Gymnolaemata" family="Lanceoporidae" genus="Calyptotheca" kingdom="Animalia" order="Cheilostomatida" pageId="5" pageNumber="587" phylum="Bryozoa" rank="species" species="obscura">
<emphasis id="B935F100A02700054351D71CE4093E6D" box="[308,561,332,356]" italics="true" pageId="5" pageNumber="587">Calyptotheca obscura</emphasis>
(
<bibRefCitation id="EFD050E3A02700054022D71CE4D03E6C" author="Osburn" box="[583,744,332,357]" pageId="5" pageNumber="587" refString="Osburn, R. C. (1952) Bryozoa of the Pacific coast of America, Part 2, Cheilostomata-Ascophora. Reports of the Allan Hancock Pacific Expeditions, 14, 271 - 611." type="journal article" year="1952">Osburn, 1952</bibRefCitation>
)
</taxonomicName>
by
<bibRefCitation id="EFD050E3A02700054144D71CE20A3E6D" box="[801,1074,332,357]" pageId="5" pageNumber="587" refString="Harmelin, J. G., Lopez de la Cuadra, C. &amp; Garcia-Gomez, J. C. (1989) Description et variabilite de Calyptotheca obscura n. sp. (Bryozoa, Cheilostomata). Bulletin du Museum National d'Histoire Naturelle, 11, 295 - 305." type="journal article">
Harmelin
<emphasis id="B935F100A027000541FCD71DE5E03E6D" box="[921,984,332,356]" italics="true" pageId="5" pageNumber="587">et al.</emphasis>
(1989)
</bibRefCitation>
. This species has transversely oriented adventitious avicularia of similar morphology to those of
<taxonomicName id="4C415691A027000541A1D720E2A83E81" authority="Abdelsalam &amp; Taylor &amp; Dorgham, 2017" authorityName="Abdelsalam &amp; Taylor &amp; Dorgham" authorityYear="2017" box="[964,1168,368,392]" class="Gymnolaemata" family="Lanceoporidae" genus="Calyptotheca" kingdom="Animalia" order="Cheilostomatida" pageId="5" pageNumber="587" phylum="Bryozoa" rank="species" species="alexandriensis" status="sp. nov.">
<emphasis id="B935F100A027000541A1D720E2A83E81" box="[964,1168,368,392]" italics="true" pageId="5" pageNumber="587">C. alexandriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2064C7BA027000546FAD73FE2DD3E81" box="[1183,1253,367,392]" pageId="5" pageNumber="587" rank="species">
<emphasis id="B935F100A027000546FAD73FE2DD3E81" bold="true" box="[1183,1253,367,392]" pageId="5" pageNumber="587">n. sp.</emphasis>
</taxonomicNameLabel>
but, unlike
<taxonomicName id="4C415691A027000547E7D720E7023EA5" authority="Abdelsalam &amp; Taylor &amp; Dorgham, 2017" authorityName="Abdelsalam &amp; Taylor &amp; Dorgham" authorityYear="2017" class="Gymnolaemata" family="Lanceoporidae" genus="Calyptotheca" kingdom="Animalia" order="Cheilostomatida" pageId="5" pageNumber="587" phylum="Bryozoa" rank="species" species="alexandriensis" status="sp. nov.">
<emphasis id="B935F100A027000547E7D720E7023EA5" italics="true" pageId="5" pageNumber="587">C. alexandriensis</emphasis>
</taxonomicName>
<emphasis id="B935F100A02700054326D7C4E7B33EA4" bold="true" box="[323,395,404,429]" pageId="5" pageNumber="587">
<taxonomicNameLabel id="A2064C7BA02700054326D7C4E7BC3EA4" box="[323,388,404,429]" pageId="5" pageNumber="587" rank="species">n. sp.</taxonomicNameLabel>
,
</emphasis>
the avicularia are positioned around the edges of the autozooids rather than suborally. The colour of
<taxonomicName id="4C415691A02700054363D7E8E7BA3ED9" authorityName="Osburn" authorityYear="1952" box="[262,386,440,464]" class="Gymnolaemata" family="Lanceoporidae" genus="Calyptotheca" kingdom="Animalia" order="Cheilostomatida" pageId="5" pageNumber="587" phylum="Bryozoa" rank="species" species="obscura">
<emphasis id="B935F100A02700054363D7E8E7BA3ED9" box="[262,386,440,464]" italics="true" pageId="5" pageNumber="587">C. obscura</emphasis>
</taxonomicName>
is dark red whereas that of
<taxonomicName id="4C415691A027000540A4D7E8E5BE3ED9" authority="Abdelsalam &amp; Taylor &amp; Dorgham, 2017" authorityName="Abdelsalam &amp; Taylor &amp; Dorgham" authorityYear="2017" box="[705,902,440,464]" class="Gymnolaemata" family="Lanceoporidae" genus="Calyptotheca" kingdom="Animalia" order="Cheilostomatida" pageId="5" pageNumber="587" phylum="Bryozoa" rank="species" species="alexandriensis" status="sp. nov.">
<emphasis id="B935F100A027000540A4D7E8E5BE3ED9" box="[705,902,440,464]" italics="true" pageId="5" pageNumber="587">C. alexandriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2064C7BA027000541EAD7E7E5F73ED9" box="[911,975,439,464]" pageId="5" pageNumber="587" rank="species">
<emphasis id="B935F100A027000541EAD7E7E5F73ED9" bold="true" box="[911,975,439,464]" pageId="5" pageNumber="587">n. sp.</emphasis>
</taxonomicNameLabel>
is very dark orange, and
<taxonomicName id="4C415691A02700054697D7E8E3563ED9" authorityName="Osburn" authorityYear="1952" box="[1266,1390,440,464]" class="Gymnolaemata" family="Lanceoporidae" genus="Calyptotheca" kingdom="Animalia" order="Cheilostomatida" pageId="5" pageNumber="587" phylum="Bryozoa" rank="species" species="obscura">
<emphasis id="B935F100A02700054697D7E8E3563ED9" box="[1266,1390,440,464]" italics="true" pageId="5" pageNumber="587">C. obscura</emphasis>
</taxonomicName>
has encrusting colonies and denticulate condyles contrasting with the erect colonies and smooth condyles of
<taxonomicName id="4C415691A027000547E7D78CE7033D11" authority="Abdelsalam &amp; Taylor &amp; Dorgham, 2017" authorityName="Abdelsalam &amp; Taylor &amp; Dorgham" authorityYear="2017" class="Gymnolaemata" family="Lanceoporidae" genus="Calyptotheca" kingdom="Animalia" order="Cheilostomatida" pageId="5" pageNumber="587" phylum="Bryozoa" rank="species" species="alexandriensis" status="sp. nov.">
<emphasis id="B935F100A027000547E7D78CE7033D11" italics="true" pageId="5" pageNumber="587">C. alexandriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2064C7BA02700054324D7AFE7B83D11" box="[321,384,511,536]" pageId="5" pageNumber="587" rank="species">
<emphasis id="B935F100A02700054324D7AFE7B83D11" bold="true" box="[321,384,511,536]" pageId="5" pageNumber="587">n. sp.</emphasis>
</taxonomicNameLabel>
</paragraph>
<paragraph id="8BFE2D12A027000542A2D475E4813C55" blockId="5.[151,1436,151,1328]" pageId="5" pageNumber="587">
<bibRefCitation id="EFD050E3A027000542A2D475E74E3D35" author="Dumont" box="[199,374,548,572]" pageId="5" pageNumber="587" refString="Dumont, J. P. C. (1981) A report on the cheilostome Bryozoa of the Sudanese Red Sea. Journal of Natural History, 15, 623 - 637." type="journal article" year="1981">Dumont (1981)</bibRefCitation>
recorded five species of
<taxonomicName id="4C415691A027000540FCD474E5163D35" authorityName="Harmer" authorityYear="1957" box="[665,814,548,572]" class="Gymnolaemata" family="Lanceoporidae" genus="Calyptotheca" kingdom="Animalia" order="Cheilostomatida" pageId="5" pageNumber="587" phylum="Bryozoa" rank="genus">
<emphasis id="B935F100A027000540FCD474E5163D35" box="[665,814,548,572]" italics="true" pageId="5" pageNumber="587">Calyptotheca</emphasis>
</taxonomicName>
from the
<collectingRegion id="4985E3F0A027000541C7D474E23D3D35" box="[930,1029,548,573]" country="Egypt" name="Al Bahr al Ahmar" pageId="5" pageNumber="587">Red Sea</collectingRegion>
:
<taxonomicName id="4C415691A02700054674D474E7053D69" authority="Canu &amp; Bassler, 1929" authorityName="Canu &amp; Bassler" authorityYear="1929" class="Gymnolaemata" family="Lanceoporidae" genus="Calyptotheca" kingdom="Animalia" order="Cheilostomatida" pageId="5" pageNumber="587" phylum="Bryozoa" rank="species" species="acutirostris">
<emphasis id="B935F100A02700054674D474E3083D35" box="[1041,1328,548,572]" italics="true" pageId="5" pageNumber="587">Calyptotheca acutirostris</emphasis>
(
<bibRefCitation id="EFD050E3A02700054725D474E70D3D69" author="Canu" pageId="5" pageNumber="587" refString="Canu, F. &amp; Bassler, R. S. (1929) Bryozoa of the Philippine region. United States National Museum Bulletin, 100, 1 - 685, 94 pls." type="journal article" year="1929">Canu &amp; Bassler, 1929</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C415691A02700054328D418E5603D69" authority="Dumont, 1981" authorityName="Dumont" authorityYear="1981" box="[333,856,584,608]" class="Gymnolaemata" family="Lanceoporidae" genus="Calyptotheca" kingdom="Animalia" order="Cheilostomatida" pageId="5" pageNumber="587" phylum="Bryozoa" rank="species" species="heteroavicularia">
<emphasis id="B935F100A02700054328D418E49F3D69" box="[333,679,584,608]" italics="true" pageId="5" pageNumber="587">Calyptotheca heteroavicularia</emphasis>
<bibRefCitation id="EFD050E3A027000540D4D418E5603D69" author="Dumont" box="[689,856,584,608]" pageId="5" pageNumber="587" refString="Dumont, J. P. C. (1981) A report on the cheilostome Bryozoa of the Sudanese Red Sea. Journal of Natural History, 15, 623 - 637." type="journal article" year="1981">Dumont, 1981</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C415691A0270005410DD418E2CF3D69" authority="Dumont, 1981" authorityName="Dumont" authorityYear="1981" box="[872,1271,584,608]" class="Gymnolaemata" family="Lanceoporidae" genus="Calyptotheca" kingdom="Animalia" order="Cheilostomatida" pageId="5" pageNumber="587" phylum="Bryozoa" rank="species" species="nigra">
<emphasis id="B935F100A0270005410DD418E27D3D56" box="[872,1093,584,608]" italics="true" pageId="5" pageNumber="587">Calyptotheca nigra</emphasis>
<bibRefCitation id="EFD050E3A0270005462AD418E2CF3D69" author="Dumont" box="[1103,1271,584,608]" pageId="5" pageNumber="587" refString="Dumont, J. P. C. (1981) A report on the cheilostome Bryozoa of the Sudanese Red Sea. Journal of Natural History, 15, 623 - 637." type="journal article" year="1981">Dumont, 1981</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C415691A02700054763D418E7863D8D" authority="Dumont, 1981" authorityName="Dumont" authorityYear="1981" class="Gymnolaemata" family="Lanceoporidae" genus="Calyptotheca" kingdom="Animalia" order="Cheilostomatida" pageId="5" pageNumber="587" phylum="Bryozoa" rank="species" species="sudanensis">
<emphasis id="B935F100A02700054763D418E72A3D8D" italics="true" pageId="5" pageNumber="587">Calyptotheca sudanensis</emphasis>
<bibRefCitation id="EFD050E3A0270005437CD43DE7863D8D" author="Dumont" box="[281,446,620,644]" pageId="5" pageNumber="587" refString="Dumont, J. P. C. (1981) A report on the cheilostome Bryozoa of the Sudanese Red Sea. Journal of Natural History, 15, 623 - 637." type="journal article" year="1981">Dumont, 1981</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C415691A0270005439ED43CE5FA3D8D" authority="Waters, 1909" authorityName="Waters" authorityYear="1909" box="[507,962,620,644]" class="Gymnolaemata" family="Lanceoporidae" genus="Calyptotheca" kingdom="Animalia" order="Cheilostomatida" pageId="5" pageNumber="587" phylum="Bryozoa" rank="species" species="wasinensis">
<emphasis id="B935F100A0270005439ED43CE5283D8D" box="[507,784,620,644]" italics="true" pageId="5" pageNumber="587">Calyptotheca wasinensis</emphasis>
(
<bibRefCitation id="EFD050E3A02700054145D43CE5813D8D" author="Waters" box="[800,953,620,644]" pageId="5" pageNumber="587" refString="Waters, A. W. (1909) Reports on the marine biology of the Sudanese Red Sea. 12. The Bryozoa. Part 1. Cheilostomata. Journal of the Linnean Society (Zoology) London, 31, 123 - 181." type="journal article" year="1909">Waters, 1909</bibRefCitation>
)
</taxonomicName>
. All of the species from the
<collectingRegion id="4985E3F0A02700054767D43CE3673D8D" box="[1282,1375,620,645]" country="Egypt" name="Al Bahr al Ahmar" pageId="5" pageNumber="587">Red Sea</collectingRegion>
have encrusting colonies in contrast to the erect colonies of
<taxonomicName id="4C415691A02700054144D4C0E5D13DA1" authority="Abdelsalam &amp; Taylor &amp; Dorgham, 2017" authorityName="Abdelsalam &amp; Taylor &amp; Dorgham" authorityYear="2017" box="[801,1001,656,680]" class="Gymnolaemata" family="Lanceoporidae" genus="Calyptotheca" kingdom="Animalia" order="Cheilostomatida" pageId="5" pageNumber="587" phylum="Bryozoa" rank="species" species="alexandriensis" status="sp. nov.">
<emphasis id="B935F100A02700054144D4C0E5D13DA1" box="[801,1001,656,680]" italics="true" pageId="5" pageNumber="587">C. alexandriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2064C7BA02700054190D4DFE2003DA1" box="[1013,1080,655,680]" pageId="5" pageNumber="587" rank="species">
<emphasis id="B935F100A02700054190D4DFE2003DA1" bold="true" box="[1013,1080,655,680]" pageId="5" pageNumber="587">n. sp.</emphasis>
</taxonomicNameLabel>
. Adventitious avicularia in
<taxonomicName id="4C415691A027000547E7D4C0E72F3DC5" authorityName="Canu &amp; Bassler" authorityYear="1929" class="Gymnolaemata" family="Lanceoporidae" genus="Calyptotheca" kingdom="Animalia" order="Cheilostomatida" pageId="5" pageNumber="587" phylum="Bryozoa" rank="species" species="acutirostris">
<emphasis id="B935F100A027000547E7D4C0E72F3DC5" italics="true" pageId="5" pageNumber="587">C. acutirostris</emphasis>
</taxonomicName>
are located proximolaterally of the orifice and directed distally, those of
<taxonomicName id="4C415691A02700054624D4E4E3233DC5" authorityName="Dumont" authorityYear="1981" box="[1089,1307,692,716]" class="Gymnolaemata" family="Lanceoporidae" genus="Calyptotheca" kingdom="Animalia" order="Cheilostomatida" pageId="5" pageNumber="587" phylum="Bryozoa" rank="species" species="heteroavicularia">
<emphasis id="B935F100A02700054624D4E4E3233DC5" box="[1089,1307,692,716]" italics="true" pageId="5" pageNumber="587">C. heteroavicularia</emphasis>
</taxonomicName>
are suboral and directed proximally,
<taxonomicName id="4C415691A027000543CBD488E4343DE6" authorityName="Dumont" authorityYear="1981" box="[430,524,728,751]" class="Gymnolaemata" family="Lanceoporidae" genus="Calyptotheca" kingdom="Animalia" order="Cheilostomatida" pageId="5" pageNumber="587" phylum="Bryozoa" rank="species" species="nigra">
<emphasis id="B935F100A027000543CBD488E4343DE6" box="[430,524,728,751]" italics="true" pageId="5" pageNumber="587">C. nigra</emphasis>
</taxonomicName>
lacks adventitious avicularia, and in
<taxonomicName id="4C415691A027000541CED488E27C3DE6" authorityName="Waters" authorityYear="1909" box="[939,1092,728,751]" class="Gymnolaemata" family="Lanceoporidae" genus="Calyptotheca" kingdom="Animalia" order="Cheilostomatida" pageId="5" pageNumber="587" phylum="Bryozoa" rank="species" species="wasinensis">
<emphasis id="B935F100A027000541CED488E27C3DE6" box="[939,1092,728,751]" italics="true" pageId="5" pageNumber="587">C. wasinensis</emphasis>
</taxonomicName>
the adventitious avicularia are scattered around the edges of the autozooids. Only those of
<taxonomicName id="4C415691A0270005414BD4ACE5F23C1D" authorityName="Dumont" authorityYear="1981" box="[814,970,764,788]" class="Gymnolaemata" family="Lanceoporidae" genus="Calyptotheca" kingdom="Animalia" order="Cheilostomatida" pageId="5" pageNumber="587" phylum="Bryozoa" rank="species" species="sudanensis">
<emphasis id="B935F100A0270005414BD4ACE5F23C1D" box="[814,970,764,788]" italics="true" pageId="5" pageNumber="587">C. sudanensis</emphasis>
</taxonomicName>
are suboral and directed laterally, as in
<taxonomicName id="4C415691A027000547E7D4ACE7033C31" authority="Abdelsalam &amp; Taylor &amp; Dorgham, 2017" authorityName="Abdelsalam &amp; Taylor &amp; Dorgham" authorityYear="2017" class="Gymnolaemata" family="Lanceoporidae" genus="Calyptotheca" kingdom="Animalia" order="Cheilostomatida" pageId="5" pageNumber="587" phylum="Bryozoa" rank="species" species="alexandriensis" status="sp. nov.">
<emphasis id="B935F100A027000547E7D4ACE7033C31" italics="true" pageId="5" pageNumber="587">C. alexandriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2064C7BA02700054321D54FE7BC3C31" box="[324,388,799,824]" pageId="5" pageNumber="587" rank="species">
<emphasis id="B935F100A02700054321D54FE7BC3C31" bold="true" box="[324,388,799,824]" pageId="5" pageNumber="587">n. sp.</emphasis>
</taxonomicNameLabel>
, but this reef-associate has small, brown- or green-coloured colonies quite unlike the large deep orange colonies of
<taxonomicName id="4C415691A027000543CDD514E4553C55" authority="Abdelsalam &amp; Taylor &amp; Dorgham, 2017" authorityName="Abdelsalam &amp; Taylor &amp; Dorgham" authorityYear="2017" box="[424,621,836,860]" class="Gymnolaemata" family="Lanceoporidae" genus="Calyptotheca" kingdom="Animalia" order="Cheilostomatida" pageId="5" pageNumber="587" phylum="Bryozoa" rank="species" species="alexandriensis" status="sp. nov.">
<emphasis id="B935F100A027000543CDD514E4553C55" box="[424,621,836,860]" italics="true" pageId="5" pageNumber="587">C. alexandriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2064C7BA02700054011D514E48A3C54" box="[628,690,836,861]" pageId="5" pageNumber="587" rank="species">
<emphasis id="B935F100A02700054011D514E48A3C54" bold="true" box="[628,690,836,861]" pageId="5" pageNumber="587">n. sp.</emphasis>
</taxonomicNameLabel>
.
</paragraph>
<paragraph id="8BFE2D12A027000542A2D538E7723B74" blockId="5.[151,1436,151,1328]" pageId="5" pageNumber="587">
A website devoted to the bryozoans of the
<collectingRegion id="4985E3F0A027000540C3D537E5333C89" box="[678,779,871,896]" country="Israel" name="Northern" pageId="5" pageNumber="587">Northern</collectingRegion>
Bay of Safaga in the
<collectingRegion id="4985E3F0A0270005419BD537E2623C89" box="[1022,1114,871,896]" country="Egypt" name="Al Bahr al Ahmar" pageId="5" pageNumber="587">Red Sea</collectingRegion>
contains SEM images of six species of
<taxonomicName id="4C415691A02700054378D5DCE78A3CAD" authorityName="Harmer" authorityYear="1957" box="[285,434,908,932]" class="Gymnolaemata" family="Lanceoporidae" genus="Calyptotheca" kingdom="Animalia" order="Cheilostomatida" pageId="5" pageNumber="587" phylum="Bryozoa" rank="genus">
<emphasis id="B935F100A02700054378D5DCE78A3CAD" box="[285,434,908,932]" italics="true" pageId="5" pageNumber="587">Calyptotheca</emphasis>
</taxonomicName>
:
<taxonomicName id="4C415691A027000543ACD5DCE4643CAD" box="[457,604,908,932]" class="Gymnolaemata" family="Lanceoporidae" genus="Calyptotheca" kingdom="Animalia" order="Cheilostomatida" pageId="5" pageNumber="587" phylum="Bryozoa" rank="species" species="wasiensis">
<emphasis id="B935F100A027000543ACD5DCE4643CAD" box="[457,604,908,932]" italics="true" pageId="5" pageNumber="587">C. wasiensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C415691A02700054016D5DCE4EF3CAD" authorityName="Dumont" authorityYear="1981" box="[627,727,908,932]" class="Gymnolaemata" family="Lanceoporidae" genus="Calyptotheca" kingdom="Animalia" order="Cheilostomatida" pageId="5" pageNumber="587" phylum="Bryozoa" rank="species" species="nigra">
<emphasis id="B935F100A02700054016D5DCE4EF3CAD" box="[627,727,908,932]" italics="true" pageId="5" pageNumber="587">C. nigra</emphasis>
</taxonomicName>
,
<taxonomicName id="4C415691A0270005408BD5DCE5A93CAD" authorityName="Dumont" authorityYear="1981" box="[750,913,908,932]" class="Gymnolaemata" family="Lanceoporidae" genus="Calyptotheca" kingdom="Animalia" order="Cheilostomatida" pageId="5" pageNumber="587" phylum="Bryozoa" rank="species" species="sudanensis">
<emphasis id="B935F100A0270005408BD5DCE5A93CAD" box="[750,913,908,932]" italics="true" pageId="5" pageNumber="587">C. sudanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C415691A027000541C2D5DCE25F3CAD" box="[935,1127,908,933]" class="Gymnolaemata" family="Lanceoporidae" genus="Calyptotheca" kingdom="Animalia" order="Cheilostomatida" pageId="5" pageNumber="587" phylum="Bryozoa" rank="species" species="wasiensis">
<emphasis id="B935F100A027000541C2D5DCE5823CAA" box="[935,954,908,931]" italics="true" pageId="5" pageNumber="587">C</emphasis>
. cf.
<emphasis id="B935F100A0270005419ED5DDE25F3CAD" box="[1019,1127,909,932]" italics="true" pageId="5" pageNumber="587">wasiensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C415691A02700054619D5DCE32A3CAD" authorityName="Harmer" authorityYear="1957" box="[1148,1298,908,932]" class="Gymnolaemata" family="Lanceoporidae" genus="Calyptotheca" kingdom="Animalia" order="Cheilostomatida" pageId="5" pageNumber="587" phylum="Bryozoa" rank="genus">
<emphasis id="B935F100A02700054619D5DCE32A3CAD" box="[1148,1298,908,932]" italics="true" pageId="5" pageNumber="587">Calyptotheca</emphasis>
</taxonomicName>
sp. 1, and
<taxonomicName id="4C415691A027000542F2D5E0E7153CC1" authorityName="Harmer" authorityYear="1957" box="[151,301,944,968]" class="Gymnolaemata" family="Lanceoporidae" genus="Calyptotheca" kingdom="Animalia" order="Cheilostomatida" pageId="5" pageNumber="587" phylum="Bryozoa" rank="genus">
<emphasis id="B935F100A027000542F2D5E0E7153CC1" box="[151,301,944,968]" italics="true" pageId="5" pageNumber="587">Calyptotheca</emphasis>
</taxonomicName>
sp. 2 (Ostrovsky
<emphasis id="B935F100A02700054064D5E1E4053CC1" box="[513,573,944,968]" italics="true" pageId="5" pageNumber="587">et al.</emphasis>
, accessed
<date id="FFFF0BD2A027000540DED5E0E54A3CC1" box="[699,882,943,968]" pageId="5" pageNumber="587" value="2017-03-20">20 March 2017</date>
). Images of the four specimens identified as
<taxonomicName id="4C415691A027000547E7D5E0E72A3CE5" authorityName="Dumont" authorityYear="1981" class="Gymnolaemata" family="Lanceoporidae" genus="Calyptotheca" kingdom="Animalia" order="Cheilostomatida" pageId="5" pageNumber="587" phylum="Bryozoa" rank="species" species="sudanensis">
<emphasis id="B935F100A027000547E7D5E0E72A3CE5" italics="true" pageId="5" pageNumber="587">C. sudanensis</emphasis>
</taxonomicName>
show the closest resemblance to
<taxonomicName id="4C415691A027000540FCD584E5593CE5" authority="Abdelsalam &amp; Taylor &amp; Dorgham, 2017" authorityName="Abdelsalam &amp; Taylor &amp; Dorgham" authorityYear="2017" box="[665,865,980,1004]" class="Gymnolaemata" family="Lanceoporidae" genus="Calyptotheca" kingdom="Animalia" order="Cheilostomatida" pageId="5" pageNumber="587" phylum="Bryozoa" rank="species" species="alexandriensis" status="sp. nov.">
<emphasis id="B935F100A027000540FCD584E5593CE5" box="[665,865,980,1004]" italics="true" pageId="5" pageNumber="587">C. alexandriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2064C7BA0270005410ED584E5943CE4" box="[875,940,980,1005]" pageId="5" pageNumber="587" rank="species">
<emphasis id="B935F100A0270005410ED584E5943CE4" bold="true" box="[875,940,980,1005]" pageId="5" pageNumber="587">n. sp.</emphasis>
</taxonomicNameLabel>
, particularly with respect to the transverse suboral avicularium, although this appears to be smaller and restricted to one side of the sinus in the Safaga material. In addition, the floor of the ovicells appears to lack the large pores seen in
<taxonomicName id="4C415691A0270005463FD24CE3273B3D" authority="Abdelsalam &amp; Taylor &amp; Dorgham, 2017" authorityName="Abdelsalam &amp; Taylor &amp; Dorgham" authorityYear="2017" box="[1114,1311,1052,1076]" class="Gymnolaemata" family="Lanceoporidae" genus="Calyptotheca" kingdom="Animalia" order="Cheilostomatida" pageId="5" pageNumber="587" phylum="Bryozoa" rank="species" species="alexandriensis" status="sp. nov.">
<emphasis id="B935F100A0270005463FD24CE3273B3D" box="[1114,1311,1052,1076]" italics="true" pageId="5" pageNumber="587">C. alexandriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2064C7BA0270005474DD24CE3503B3C" box="[1320,1384,1052,1077]" pageId="5" pageNumber="587" rank="species">
<emphasis id="B935F100A0270005474DD24CE3503B3C" bold="true" box="[1320,1384,1052,1077]" pageId="5" pageNumber="587">n. sp.</emphasis>
</taxonomicNameLabel>
The lack of descriptive text means that colony-form and colour cannot be compared between the material from Safaga and
<collectingRegion id="4985E3F0A027000542A3D234E77E3B74" box="[198,326,1124,1149]" country="Egypt" name="Al Iskandariyah" pageId="5" pageNumber="587">Alexandria</collectingRegion>
.
</paragraph>
<paragraph id="8BFE2D12A027000542A2D2D8E4D63A04" blockId="5.[151,1436,151,1328]" pageId="5" pageNumber="587">
In possessing a transversely oriented suboral avicularium,
<taxonomicName id="4C415691A0270005411FD2D8E27B3BA9" authority="Abdelsalam &amp; Taylor &amp; Dorgham, 2017" authorityName="Abdelsalam &amp; Taylor &amp; Dorgham" authorityYear="2017" box="[890,1091,1160,1184]" class="Gymnolaemata" family="Lanceoporidae" genus="Calyptotheca" kingdom="Animalia" order="Cheilostomatida" pageId="5" pageNumber="587" phylum="Bryozoa" rank="species" species="alexandriensis" status="sp. nov.">
<emphasis id="B935F100A0270005411FD2D8E27B3BA9" box="[890,1091,1160,1184]" italics="true" pageId="5" pageNumber="587">C. alexandriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2064C7BA02700054635D2D7E2AC3BA9" box="[1104,1172,1159,1184]" pageId="5" pageNumber="587" rank="species">
<emphasis id="B935F100A02700054635D2D7E2AC3BA9" bold="true" box="[1104,1172,1159,1184]" pageId="5" pageNumber="587">n. sp.</emphasis>
</taxonomicNameLabel>
resembles the species
<taxonomicName id="4C415691A027000542F2D2FCE4823BCD" authority="MacGillivray, 1879" authorityName="MacGillivray" authorityYear="1879" box="[151,698,1196,1221]" class="Gymnolaemata" family="Lanceoporidae" genus="Calyptotheca" kingdom="Animalia" order="Cheilostomatida" pageId="5" pageNumber="587" phylum="Bryozoa" rank="species" species="subimmersa">
<emphasis id="B935F100A027000542F2D2FCE7863BCD" box="[151,446,1196,1220]" italics="true" pageId="5" pageNumber="587">Calyptotheca subimmersa</emphasis>
(
<bibRefCitation id="EFD050E3A027000543B5D2FCE4893BCD" author="MacGillivray" box="[464,689,1196,1221]" pageId="5" pageNumber="587" refString="MacGillivray, P. H. (1879) Polyzoa. In: McCoy, F. (Ed.), Prodromus of the Zoology of Victoria; or, Figures and descriptions of the living species of all classes of the Victorian indigenous animals. Vol. I. Decade IV. Government Printer, Melbourne, pp. 21 - 40, pls. 35 - 39." type="book chapter" year="1879">MacGillivray, 1879</bibRefCitation>
)
</taxonomicName>
from
<collectingCountry id="F3566D82A02700054160D2FCE5573BCC" box="[773,879,1196,1221]" name="Australia" pageId="5" pageNumber="587">Australia</collectingCountry>
. However, the avicularium of
<taxonomicName id="4C415691A027000546B6D2FCE3463BCD" authorityName="MacGillivray" authorityYear="1879" box="[1235,1406,1196,1220]" class="Gymnolaemata" family="Lanceoporidae" genus="Calyptotheca" kingdom="Animalia" order="Cheilostomatida" pageId="5" pageNumber="587" phylum="Bryozoa" rank="species" species="subimmersa">
<emphasis id="B935F100A027000546B6D2FCE3463BCD" box="[1235,1406,1196,1220]" italics="true" pageId="5" pageNumber="587">C. subimmersa</emphasis>
</taxonomicName>
is smaller than that of
<taxonomicName id="4C415691A02700054312D280E4033BE1" authority="Abdelsalam &amp; Taylor &amp; Dorgham, 2017" authorityName="Abdelsalam &amp; Taylor &amp; Dorgham" authorityYear="2017" box="[375,571,1232,1256]" class="Gymnolaemata" family="Lanceoporidae" genus="Calyptotheca" kingdom="Animalia" order="Cheilostomatida" pageId="5" pageNumber="587" phylum="Bryozoa" rank="species" species="alexandriensis" status="sp. nov.">
<emphasis id="B935F100A02700054312D280E4033BE1" box="[375,571,1232,1256]" italics="true" pageId="5" pageNumber="587">C. alexandriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2064C7BA02700054027D29FE4B93BE1" box="[578,641,1231,1256]" pageId="5" pageNumber="587" rank="species">
<emphasis id="B935F100A02700054027D29FE4B93BE1" bold="true" box="[578,641,1231,1256]" pageId="5" pageNumber="587">n. sp.</emphasis>
</taxonomicNameLabel>
, rounded rather than arch-shaped, and opens in the plane of the frontal surface of the zooid instead of being steeply inclined.
</paragraph>
</subSubSection>
<subSubSection id="C35B7E99A027000542A2D347E5393A39" box="[199,769,1303,1328]" pageId="5" pageNumber="587" type="distribution">
<paragraph id="8BFE2D12A027000542A2D347E5393A39" blockId="5.[151,1436,151,1328]" box="[199,769,1303,1328]" pageId="5" pageNumber="587">
<emphasis id="B935F100A027000542A2D347E7583A39" bold="true" box="[199,352,1303,1328]" pageId="5" pageNumber="587">Distribution.</emphasis>
<collectingRegion id="4985E3F0A02700054302D348E7833A26" box="[359,443,1304,1327]" country="American Samoa" name="Eastern" pageId="5" pageNumber="587">Eastern</collectingRegion>
Harbour,
<collectingRegion id="4985E3F0A02700054049D347E4933A39" box="[556,683,1303,1328]" country="Egypt" name="Al Iskandariyah" pageId="5" pageNumber="587">Alexandria</collectingRegion>
,
<collectingCountry id="F3566D82A027000540D3D348E4C53A39" box="[694,765,1304,1328]" name="Egypt" pageId="5" pageNumber="587">Egypt</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
Reference in a new issue View git blame Copy permalink