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<mods:title id="6F3A97687D400EC55CEB3DD5DD1DE82D">A taxonomic reappraisal of the Smooth Slug Snake Asthenodipsas laevis (Boie, 1827) (Squamata: Pareidae) in Borneo with the description of two new species</mods:title>
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<emphasis id="B93CEAF83616494EFF5FFCECFE5BFCD8" bold="true" box="[151,444,785,812]" italics="true" pageId="3" pageNumber="504">Asthenodipsas stuebingi</emphasis>
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&amp;
<figureCitation id="13732A6F3616494EFECFFCA4FEF0FC87" box="[263,279,857,883]" captionStart="FIGURE 3" captionStartId="7.[151,250,1588,1613]" captionTargetBox="[151,1436,211,1555]" captionTargetId="figure@7.[151,1436,211,1555]" captionTargetPageId="7" captionText="FIGURE 3. A. Dorsum of paratype of Asthenodipsas stuebingi sp. nov. (ZMB 65429). B. Venter of paratype of A. stuebingi sp. nov. (ZMB 65429) (Photographs by Frank Tillack). C. Dorsum of paratype of A. stuebingi sp. nov. (ZMB 65429) in life from Kinabalu Park power station, Kamborangoh Road, Sabah. D. Paratype of A. stuebingi sp. nov. (ZMB 65429) from Kinabalu Park power station, Kamborangoh Road, Sabah in defensive coils. E. Venter of paratype of A. stuebingi sp. nov. (ZMB 65429) in life from Kinabalu Park power station, Kamborangoh Road, Sabah. (Photographs by Andreas Nӧllert). F. Sharp vertebral keel of the holotype of A. stuebingi sp. nov. (SP 04679) as indicated by arrow." httpUri="https://zenodo.org/record/3351078/files/figure.png" pageId="3" pageNumber="504">3</figureCitation>
.
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<taxonomicName id="4C484D693616494EFF5FFC3CFE94FC2D" ID-CoL="CLGK" authorityName="Boie" authorityYear="1827" box="[151,371,961,985]" class="Squamata" family="Pareidae" genus="Amblycephalus" higherTaxonomySource="CoL" kingdom="Animalia" pageId="3" pageNumber="504" phylum="Chordata" rank="species" species="laevis">
<emphasis id="B93CEAF83616494EFF5FFC3CFE94FC2D" box="[151,371,961,985]" italics="true" pageId="3" pageNumber="504">Amblycephalus laevis</emphasis>
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(in part):
<treatmentCitation id="0AE910FB3616494EFE11FC3CFD48FC2D" author="Boulenger, G. A." box="[473,687,961,986]" page="441" pageId="3" pageNumber="504" year="1896">
<bibRefCitation id="EFD94B1B3616494EFE11FC3CFD48FC2D" author="Boulenger, G. A." box="[473,687,961,986]" pageId="3" pageNumber="504" refId="ref15412" refString="Boulenger, G. A. (1896) Catalogue of the Snakes in the British Museum (Natural History). Volume III. Trustees of the British Museum, London, xiv + 727 pp., 15 plates. https: // doi. org / 10.5962 / bhl. title. 8316" type="book" year="1896">Boulenger 1896: 441</bibRefCitation>
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&amp; 442;
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<bibRefCitation id="EFD94B1B3616494EFCCBFC3CFC2EFC2E" author="de Rooij, N." box="[771,969,961,986]" pageId="3" pageNumber="504" refId="ref15605" refString="de Rooij, N. (1917) The Reptiles of the Indo-Australian Archipelago. II. Ophidia. E. J. Brill Ltd., Leiden, 334 pp. https: // doi. org / 10.5962 / bhl. title. 10610" type="book" year="1917">de Rooij 1917: 276</bibRefCitation>
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<emphasis id="B93CEAF83616494EFF5FFC1DFE41FC0C" box="[151,422,992,1016]" italics="true" pageId="3" pageNumber="504">Amblycephalus vertebralis</emphasis>
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(in part):
<treatmentCitation id="0AE910FB3616494EFDC4FC22FD30FC0C" author="Loveridge, A." box="[524,727,991,1016]" page="43" pageId="3" pageNumber="504" year="1938">
<bibRefCitation id="EFD94B1B3616494EFDC4FC22FD30FC0C" author="Loveridge, A." box="[524,727,991,1016]" pageId="3" pageNumber="504" pagination="43 - 46" refId="ref16911" refString="Loveridge, A. (1938) New snakes of the genera Calamaria, Bungarus and Trimeresurus from Mount Kinabalu, North Borneo. Proceedings of the Biological Society of Washington, 51, 43 - 46." type="journal article" year="1938">Loveridge 1938: 43</bibRefCitation>
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.
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<taxonomicName id="4C484D693616494EFF5FFC02FE8CFBE3" authorityName="Quah &amp; Grismer &amp; Lim &amp; Anuar &amp; Imbun" authorityYear="2019" box="[151,363,1023,1047]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="504" phylum="Chordata" rank="species" species="laevis">
<emphasis id="B93CEAF83616494EFF5FFC02FE8CFBE3" box="[151,363,1023,1047]" italics="true" pageId="3" pageNumber="504">Asthenodipsas laevis</emphasis>
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(in part):
<bibRefCitation id="EFD94B1B3616494EFE19FC02FDD1FBE3" author="Das, I." box="[465,566,1022,1047]" pageId="3" pageNumber="504" pagination="3 - 11" refId="ref15508" refString="Das, I. (2006) Crocker Range National Park, Sabah, as a refuge for Borneo's montane herpetofauna. Amphibian and Reptile Conservation, 4 (1), 3 - 11." type="journal article" year="2006">Das 2006</bibRefCitation>
: 007, 2010: 344;
<treatmentCitation id="0AE910FB3616494EFD2DFC02FBC9FBE3" author="Grossmann, W. &amp; Tillack, F." box="[741,1070,1022,1047]" page="188" pageId="3" pageNumber="504" year="2003">
<bibRefCitation id="EFD94B1B3616494EFD2DFC02FBC9FBE3" author="Grossmann, W. &amp; Tillack, F." box="[741,1070,1022,1047]" pageId="3" pageNumber="504" pagination="175 - 90" refId="ref16457" refString="Grossmann, W. &amp; Tillack, F. (2003) On the Taxonomic Status of Asthenodipsas tropidonotus (Van Lidth De Jeude, 1923) and Pareas vertebralis (Boulenger, 1900) (Serpentes: Colubridae: Pareatinae). Russian Journal of Herpetology, 10 (3), 175 - 90." type="journal article" year="2003">Grossmann &amp; Tillack 2003: 188</bibRefCitation>
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;
<treatmentCitation id="0AE910FB3616494EFBF1FC03FAC9FBE3" author="Stuebing, R. B. &amp; Inger, R. F. &amp; Lardner, B." box="[1081,1326,1022,1047]" page="85" pageId="3" pageNumber="504" year="2014">
<bibRefCitation id="EFD94B1B3616494EFBF1FC03FAC9FBE3" author="Stuebing, R. B. &amp; Inger, R. F. &amp; Lardner, B." box="[1081,1326,1022,1047]" pageId="3" pageNumber="504" refId="ref17497" refString="Stuebing, R. B., Inger, R. F. &amp; Lardner, B. (2014) A field guide to the snakes of Borneo (2 nd edition). Natural History Publications (Borneo), Kota Kinabalu, 310 pp." type="book" year="2014">
Stuebing
<emphasis id="B93CEAF83616494EFB51FC02FB21FBE3" box="[1177,1222,1023,1047]" italics="true" pageId="3" pageNumber="504">et al</emphasis>
. 2014: 85
</bibRefCitation>
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.
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</paragraph>
<paragraph id="8BF736EA3616494EFF5FFBE3FCB5FBC1" blockId="3.[151,1436,961,1139]" box="[151,850,1053,1078]" pageId="3" pageNumber="504">
<treatmentCitationGroup id="AB5811C43616494EFF5FFBE3FCB5FBC1" box="[151,850,1053,1078]" pageId="3" pageNumber="504">
<taxonomicName id="4C484D693616494EFF5FFBE3FDB4FBC1" authority=": Malkmus et al. 2002: 342" authorityName="Malkmus" authorityPageNumber="342" authorityYear="2002" box="[151,595,1053,1078]" class="Squamata" family="Pareidae" genus="Internatus" higherTaxonomySource="CoL" kingdom="Animalia" pageId="3" pageNumber="504" phylum="Chordata" rank="species" species="laevis">
<emphasis id="B93CEAF83616494EFF5FFBE3FEA6FBC1" box="[151,321,1053,1077]" italics="true" pageId="3" pageNumber="504">Internatus laevis</emphasis>
:
<treatmentCitation id="0AE910FB3616494EFE86FBE0FDB4FBC1" author="Malkmus, R. &amp; Manthey, U. &amp; Vogel, G. &amp; Hoffmann, P. &amp; Kosuch, J." box="[334,595,1053,1078]" page="342" pageId="3" pageNumber="504" year="2002">
<bibRefCitation id="EFD94B1B3616494EFE86FBE0FDB4FBC1" author="Malkmus, R. &amp; Manthey, U. &amp; Vogel, G. &amp; Hoffmann, P. &amp; Kosuch, J." box="[334,595,1053,1078]" pageId="3" pageNumber="504" refId="ref16983" refString="Malkmus, R., Manthey, U., Vogel, G., Hoffmann, P. &amp; Kosuch, J. (2002) Amphibians and Reptiles of Mount Kinabalu (North- Borneo). A. R. G. Ganter Verlag, Ruggel, 424 pp." type="book" year="2002">
Malkmus
<emphasis id="B93CEAF83616494EFE7CFBE3FE06FBC1" box="[436,481,1053,1077]" italics="true" pageId="3" pageNumber="504">et al</emphasis>
. 2002: 342
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&amp; 343, Figs, 337 &amp; 338.
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<treatmentCitationGroup id="AB5811C43616494EFF5FFBC0FD38FB87" pageId="3" pageNumber="504">
<taxonomicName id="4C484D693616494EFF5FFBC0FE1DFBA0" authority=": Malkmus 1996: 293" authorityName=": Malkmus" authorityPageNumber="293" authorityYear="1996" box="[151,506,1083,1108]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="504" phylum="Chordata" rank="species" species="laevis">
<emphasis id="B93CEAF83616494EFF5FFBC0FEF8FBA0" box="[151,287,1084,1108]" italics="true" pageId="3" pageNumber="504">Pareas laevis</emphasis>
:
<treatmentCitation id="0AE910FB3616494EFEE4FBC6FE1DFBA0" author="Malkmus, R." box="[300,506,1083,1108]" page="293" pageId="3" pageNumber="504" year="1996">
<bibRefCitation id="EFD94B1B3616494EFEE4FBC6FE1DFBA0" author="Malkmus, R." box="[300,506,1083,1108]" pageId="3" pageNumber="504" pagination="277 - 295" refId="ref16949" refString="Malkmus, R. (1996) Herpetologische Beobachtungen am Mount Kinabalu, Nord-Borneo. V. Mitteilungen aus dem Zoologischen Museum in Berlin, 72 (2), 277 - 295." type="journal article" year="1996">Malkmus 1996: 293</bibRefCitation>
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="0AE910FB3616494EFDCCFBC6FCB8FBA0" author="Manthey, U. &amp; Grossmann, W." box="[516,863,1083,1108]" page="377" pageId="3" pageNumber="504" year="1997">
<bibRefCitation id="EFD94B1B3616494EFDCCFBC6FCB8FBA0" author="Manthey, U. &amp; Grossmann, W." box="[516,863,1083,1108]" pageId="3" pageNumber="504" refId="ref17035" refString="Manthey, U. &amp; Grossmann, W. (1997) Amphibien &amp; Reptilien Sudostasiens. Natur und Tier Verlag, Munster, 512 pp." type="book" year="1997">Manthey &amp; Grossmann 1997: 377</bibRefCitation>
</treatmentCitation>
, Abb. 284;
<treatmentCitation id="0AE910FB3616494EFC1AFBC6FB7FFBA0" author="Stuebing, R. B." box="[978,1176,1083,1108]" page="330" pageId="3" pageNumber="504" year="1991">
<bibRefCitation id="EFD94B1B3616494EFC1AFBC6FB7FFBA0" author="Stuebing, R. B." box="[978,1176,1083,1108]" pageId="3" pageNumber="504" pagination="323 - 362" refId="ref17427" refString="Stuebing, R. B. (1991) A checklist of the snakes of Borneo. Raffles Bulletin of Zoology Singapore, 39 (2), 323 - 362." type="journal article" year="1991">Stuebing 1991: 330</bibRefCitation>
</treatmentCitation>
&amp; 331 (in part);
<treatmentCitation id="0AE910FB3616494EFA8AFBC6FE91FB87" author="Stuebing, R. B. &amp; Inger, R. F." page="87" pageId="3" pageNumber="504" year="1999">
<bibRefCitation id="EFD94B1B3616494EFA8AFBC6FE91FB87" author="Stuebing, R. B. &amp; Inger, R. F." pageId="3" pageNumber="504" refId="ref17459" refString="Stuebing, R. B. &amp; Inger, R. F. (1999) A Field Giude to the Snakes of Borneo. Natural History Publications (Borneo), Kota Kinabalu, 254 pp." type="book" year="1999">Stuebing &amp; Inger 1999: 87</bibRefCitation>
</treatmentCitation>
(in part);
<treatmentCitation id="0AE910FB3616494EFE14FBA7FD67FB87" author="Haile, N. S." box="[476,640,1114,1139]" page="766" pageId="3" pageNumber="504" year="1958">
<bibRefCitation id="EFD94B1B3616494EFE14FBA7FD67FB87" author="Haile, N. S." box="[476,640,1114,1139]" pageId="3" pageNumber="504" pagination="743 - 771" refId="ref16514" refString="Haile, N. S. (1958) The snakes of Borneo. Sarawak Museum Journal, Kuching, New Series, 8 (12), 743 - 771." type="journal article" year="1958">Haile 1958: 766</bibRefCitation>
</treatmentCitation>
(in part).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C35265613616494BFF5FFB61FBBAFE93" lastPageId="6" lastPageNumber="507" pageId="3" pageNumber="504" type="materials_examined">
<paragraph id="8BF736EA3616494EFF5FFB61FE2DFB08" blockId="3.[151,1437,1179,2038]" pageId="3" pageNumber="504">
<materialsCitation id="3B203CB73616494EFF5FFB61FE2DFB08" ID-GBIF-Occurrence="2430030365" collectingDate="2009-03-25" collectorName="Adult &amp; Paul Yambun Imbun &amp; Fred Tuh Yit Yu &amp; Safrie" country="Malaysia" county="East" elevation="1859" latitude="5.82324" location="Minduk Sirung" longLatPrecision="1" longitude="116.34724" municipality="Crocker Range Park" pageId="3" pageNumber="504" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
<emphasis id="B93CEAF83616494EFF5FFB61FEECFB42" bold="true" box="[151,267,1180,1206]" pageId="3" pageNumber="504">
<typeStatus id="54F388483616494EFF5FFB61FEE1FB42" box="[151,262,1180,1206]" pageId="3" pageNumber="504" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectorName id="26BD533C3616494EFED9FB61FEB6FB42" box="[273,337,1180,1206]" pageId="3" pageNumber="504">Adult</collectorName>
female, SP 04679 collected by
<collectorName id="26BD533C3616494EFD67FB61FC71FB42" box="[687,918,1180,1206]" pageId="3" pageNumber="504">Paul Yambun Imbun</collectorName>
,
<collectorName id="26BD533C3616494EFC69FB61FBB5FB42" box="[929,1106,1180,1206]" pageId="3" pageNumber="504">Fred Tuh Yit Yu</collectorName>
and
<collectorName id="26BD533C3616494EFB4FFB61FB2DFB42" box="[1159,1226,1180,1206]" pageId="3" pageNumber="504">Safrie</collectorName>
on
<date id="FFF6102A3616494EFB3AFB66FA7BFB42" box="[1266,1436,1179,1206]" pageId="3" pageNumber="504" value="2009-03-25">
<collectingDate id="EFB2E9C23616494EFB3AFB66FA7BFB42" box="[1266,1436,1179,1206]" pageId="3" pageNumber="504" value="2009-03-25">25 March 2009</collectingDate>
</date>
from
<location id="8E9760313616494EFF1FFB43FE63FB2C" LSID="urn:lsid:plazi:treatment:03E187FC3616494BFF5FFCECFBBAFE93:8E9760313616494EFF1FFB43FE63FB2C" box="[215,388,1214,1240]" country="Malaysia" county="East" latitude="5.82324" longLatPrecision="1" longitude="116.34724" municipality="Crocker Range Park" name="Minduk Sirung" pageId="3" pageNumber="504" stateProvince="Sabah">Minduk Sirung</location>
(AlabMahua trail),
<collectingMunicipality id="6B93AC903616494EFDB0FB43FC84FB2C" box="[632,867,1214,1241]" pageId="3" pageNumber="504">Crocker Range Park</collectingMunicipality>
,
<collectingRegion id="498CF8083616494EFCB8FB43FC50FB2C" box="[880,951,1214,1240]" country="Malaysia" name="Sabah" pageId="3" pageNumber="504">Sabah</collectingRegion>
,
<collectingCounty id="62964E663616494EFC0DFB42FC12FB2D" box="[965,1013,1215,1241]" pageId="3" pageNumber="504">East</collectingCounty>
<collectingCountry id="F35F767A3616494EFC37FB43FB81FB2C" box="[1023,1126,1214,1240]" name="Malaysia" pageId="3" pageNumber="504">Malaysia</collectingCountry>
(estimated:
<geoCoordinate id="EE7C502D3616494EFB3CFB42FA99FB2D" box="[1268,1406,1214,1241]" degrees="5.823240" direction="north" orientation="latitude" pageId="3" pageNumber="504" precision="1" value="5.82324">N 5.823240</geoCoordinate>
,
<geoCoordinate id="EE7C502D3616494EFA43FB42FEF9FB0F" degrees="116.347238" direction="east" orientation="longitude" pageId="3" pageNumber="504" precision="1" value="116.34724">E 116.347238</geoCoordinate>
,
<elevation id="0065D1D93616494EFEE1FB1DFE5DFB08" box="[297,442,1248,1276]" metricMagnitude="3" metricUnit="m" metricValue="1.859" pageId="3" pageNumber="504" unit="m" value="1859.0">
<quantity id="4CB09B0F3616494EFEE1FB1DFE99FB08" box="[297,382,1248,1276]" metricMagnitude="3" metricUnit="m" metricValue="1.859" pageId="3" pageNumber="504" unit="m" value="1859.0">1859 m</quantity>
a.s.l.
</elevation>
).
</materialsCitation>
</paragraph>
<paragraph id="8BF736EA3616494EFF0FFAF9FE33FA38" blockId="3.[151,1437,1179,2038]" pageId="3" pageNumber="504">
<materialsCitation id="3B203CB73616494EFF0FFAF9FB19FAB5" ID-GBIF-Occurrence="2430030360" collectingDate="2014-10-26" collectorName="Adult &amp; Siti Azizah Kusop &amp; Junaidi Ais &amp; Adzmie" elevation="1341" latitude="5.988087" location="Kampung Desa Aman" longLatPrecision="1" longitude="116.57016" municipality="Kundasang" pageId="3" pageNumber="504" specimenCount="1" typeStatus="paratype">
<emphasis id="B93CEAF83616494EFF0FFAF9FEA1FAEA" bold="true" box="[199,326,1284,1310]" pageId="3" pageNumber="504">
<typeStatus id="54F388483616494EFF0FFAF9FEA6FAEA" box="[199,321,1284,1310]" pageId="3" pageNumber="504" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectorName id="26BD533C3616494EFE84FAF9FE6BFAEA" box="[332,396,1284,1310]" pageId="3" pageNumber="504">Adult</collectorName>
male (SP 04806) collected by
<collectorName id="26BD533C3616494EFD2EFAF9FC52FAEA" box="[742,949,1284,1310]" pageId="3" pageNumber="504">Siti Azizah Kusop</collectorName>
,
<collectorName id="26BD533C3616494EFC08FAF9FBD9FAEA" box="[960,1086,1284,1310]" pageId="3" pageNumber="504">Junaidi Ais</collectorName>
and R.,
<collectorName id="26BD533C3616494EFB53FAF9FB13FAEA" box="[1179,1268,1284,1310]" pageId="3" pageNumber="504">Adzmie</collectorName>
on
<date id="FFF6102A3616494EFAD7FAF9FF28FAB5" pageId="3" pageNumber="504" value="2014-10-26">
<collectingDate id="EFB2E9C23616494EFAD7FAF9FF28FAB5" pageId="3" pageNumber="504" value="2014-10-26">26 October 2014</collectingDate>
</date>
from
<location id="8E9760313616494EFEDDFADAFDF6FAB5" LSID="urn:lsid:plazi:treatment:03E187FC3616494BFF5FFCECFBBAFE93:8E9760313616494EFEDDFADAFDF6FAB5" box="[277,529,1319,1345]" latitude="5.988087" longLatPrecision="1" longitude="116.57016" municipality="Kundasang" name="Kampung Desa Aman" pageId="3" pageNumber="504">Kampung Desa Aman</location>
,
<collectingMunicipality id="6B93AC903616494EFDD6FADBFD7BFAB4" box="[542,668,1318,1344]" pageId="3" pageNumber="504">Kundasang</collectingMunicipality>
(estimated:
<geoCoordinate id="EE7C502D3616494EFCE0FADBFC4FFAB5" box="[808,936,1318,1345]" degrees="5.988087" direction="north" orientation="latitude" pageId="3" pageNumber="504" precision="1" value="5.988087">N5.988087</geoCoordinate>
,
<geoCoordinate id="EE7C502D3616494EFC7CFADAFBB3FAB5" box="[948,1108,1318,1345]" degrees="116.570162" direction="east" orientation="longitude" pageId="3" pageNumber="504" precision="1" value="116.57016">E 116.570162</geoCoordinate>
,
<elevation id="0065D1D93616494EFBA8FADAFB14FAB5" box="[1120,1267,1318,1345]" metricMagnitude="3" metricUnit="m" metricValue="1.341" pageId="3" pageNumber="504" unit="m" value="1341.0">
<quantity id="4CB09B0F3616494EFBA8FADAFB51FAB5" box="[1120,1206,1319,1345]" metricMagnitude="3" metricUnit="m" metricValue="1.341" pageId="3" pageNumber="504" unit="m" value="1341.0">1341 m</quantity>
a.s.l.
</elevation>
)
</materialsCitation>
.
<materialsCitation id="3B203CB73616494EFAC0FADBFBCFFA73" ID-GBIF-Occurrence="2430030364" collectingDate="1994-08" collectionCode="ZMB" collectorName="Adult &amp; Rudolf Malkmus &amp; Andreas N" country="Malaysia" county="West" elevation="1854" latitude="6.0292" location="Kamborangoh Road" longLatPrecision="7" longitude="116.5473" municipality="Mount Kinabalu" pageId="3" pageNumber="504" specimenCount="1" stateProvince="Sabah" typeStatus="paratype">
<collectorName id="26BD533C3616494EFAC0FADBFAAFFAB4" box="[1288,1352,1318,1344]" pageId="3" pageNumber="504">Adult</collectorName>
female (
<collectionCode id="ED59AE2F3616494EFF57FAB4FF3AFA97" box="[159,221,1353,1379]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="3" pageNumber="504" type="Museum">ZMB</collectionCode>
65429) collected by
<collectorName id="26BD533C3616494EFE07FAB4FD68FA97" box="[463,655,1353,1379]" pageId="3" pageNumber="504">Rudolf Malkmus</collectorName>
and
<collectorName id="26BD533C3616494EFD0FFAB4FCA7FA97" box="[711,832,1353,1379]" pageId="3" pageNumber="504">Andreas N</collectorName>
ӧllert in
<date id="FFF6102A3616494EFC69FAB4FBD5FA97" box="[929,1074,1353,1379]" pageId="3" pageNumber="504" value="1994-08">
<collectingDate id="EFB2E9C23616494EFC69FAB4FBD5FA97" box="[929,1074,1353,1379]" pageId="3" pageNumber="504" value="1994-08">August 1994</collectingDate>
</date>
on
<location id="8E9760313616494EFB97FAB4FAA2FA97" LSID="urn:lsid:plazi:treatment:03E187FC3616494BFF5FFCECFBBAFE93:8E9760313616494EFB97FAB4FAA2FA97" box="[1119,1349,1353,1379]" country="Malaysia" county="West" latitude="6.0292" longLatPrecision="7" longitude="116.5473" municipality="Mount Kinabalu" name="Kamborangoh Road" pageId="3" pageNumber="504" stateProvince="Sabah">Kamborangoh Road</location>
,
<collectingMunicipality id="6B93AC903616494EFA9AFAB7FEE6FA72" pageId="3" pageNumber="504">Mount Kinabalu</collectingMunicipality>
,
<collectingRegion id="498CF8083616494EFEC5FA91FEB3FA72" box="[269,340,1388,1414]" country="Malaysia" name="Sabah" pageId="3" pageNumber="504">Sabah</collectingRegion>
,
<collectingCounty id="62964E663616494EFEA8FA91FE70FA72" box="[352,407,1388,1414]" pageId="3" pageNumber="504">West</collectingCounty>
<collectingCountry id="F35F767A3616494EFE57FA91FDE1FA72" box="[415,518,1388,1414]" name="Malaysia" pageId="3" pageNumber="504">Malaysia</collectingCountry>
(estimated:
<geoCoordinate id="EE7C502D3616494EFD5AFA91FD11FA72" box="[658,758,1388,1414]" degrees="6.0292" direction="north" orientation="latitude" pageId="3" pageNumber="504" precision="5" value="6.0292">N6.0292</geoCoordinate>
,
<geoCoordinate id="EE7C502D3616494EFCCAFA96FC9AFA72" box="[770,893,1387,1414]" degrees="116.5473" direction="east" orientation="longitude" pageId="3" pageNumber="504" precision="5" value="116.5473">E116.5473</geoCoordinate>
,
<elevation id="0065D1D93616494EFC42FA96FBFAFA73" box="[906,1053,1387,1415]" metricMagnitude="3" metricUnit="m" metricValue="1.8539999999999999" pageId="3" pageNumber="504" unit="m" value="1854.0">
<quantity id="4CB09B0F3616494EFC42FA96FC07FA72" box="[906,992,1387,1414]" metricMagnitude="3" metricUnit="m" metricValue="1.8539999999999999" pageId="3" pageNumber="504" unit="m" value="1854.0">1854 m</quantity>
a.s.l.
</elevation>
)
</materialsCitation>
.
<materialsCitation id="3B203CB73616494EFBFBFA91FE28FA38" ID-GBIF-Occurrence="2430030357" collectionCode="MCZ" collectorName="Adult &amp; John A. Griswold Jr. &amp; Kadmayan R" country="Malaysia" elevation="971" latitude="6.029788" location="Kiau" longLatPrecision="1" longitude="116.48973" municipality="West" pageId="3" pageNumber="504" specimenCount="1" stateProvince="Sabah" typeStatus="paratype">
<collectorName id="26BD533C3616494EFBFBFA91FB94FA72" box="[1075,1139,1388,1414]" pageId="3" pageNumber="504">Adult</collectorName>
male (
<collectionCode id="ED59AE2F3616494EFB08FA91FB18FA72" box="[1216,1279,1388,1414]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="3" pageNumber="504" type="Museum">MCZ</collectionCode>
R43591) collected by
<collectorName id="26BD533C3616494EFECAFA73FE02FA5D" box="[258,485,1422,1449]" pageId="3" pageNumber="504">John A. Griswold Jr.</collectorName>
in 1937 from
<collectorName id="26BD533C3616494EFD49FA73FCF1FA5D" box="[641,790,1422,1449]" pageId="3" pageNumber="504">Kadmayan R</collectorName>
, near
<location id="8E9760313616494EFC9FFA73FC77FA5C" LSID="urn:lsid:plazi:treatment:03E187FC3616494BFF5FFCECFBBAFE93:8E9760313616494EFC9FFA73FC77FA5C" box="[855,912,1422,1448]" country="Malaysia" latitude="6.029788" longLatPrecision="1" longitude="116.48973" municipality="West" name="Kiau" pageId="3" pageNumber="504" stateProvince="Sabah">Kiau</location>
,
<collectingRegion id="498CF8083616494EFC52FA73FC06FA5C" box="[922,993,1422,1448]" country="Malaysia" name="Sabah" pageId="3" pageNumber="504">Sabah</collectingRegion>
,
<collectingMunicipality id="6B93AC903616494EFC23FA72FBC5FA5D" box="[1003,1058,1423,1449]" pageId="3" pageNumber="504">West</collectingMunicipality>
<collectingCountry id="F35F767A3616494EFBE1FA73FB77FA5C" box="[1065,1168,1422,1448]" name="Malaysia" pageId="3" pageNumber="504">Malaysia</collectingCountry>
(estimated:
<geoCoordinate id="EE7C502D3616494EFAD0FA73FA7FFA5C" box="[1304,1432,1422,1448]" degrees="6.029788" direction="north" orientation="latitude" pageId="3" pageNumber="504" precision="1" value="6.029788">N6.029788</geoCoordinate>
,
<geoCoordinate id="EE7C502D3616494EFF5FFA4FFED1FA3F" box="[151,310,1457,1484]" degrees="116.489732" direction="east" orientation="longitude" pageId="3" pageNumber="504" precision="1" value="116.48973">E 116.489732</geoCoordinate>
,
<elevation id="0065D1D93616494EFE89FA4CFE22FA38" box="[321,453,1457,1484]" metricMagnitude="2" metricUnit="m" metricValue="9.709999999999999" pageId="3" pageNumber="504" unit="m" value="971.0">
<quantity id="4CB09B0F3616494EFE89FA4CFE6FFA38" box="[321,392,1457,1484]" metricMagnitude="2" metricUnit="m" metricValue="9.709999999999999" pageId="3" pageNumber="504" unit="m" value="971.0">971 m</quantity>
a.s.l.
</elevation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BF736EA3616494EFF0FFA2EFB04F89F" blockId="3.[151,1437,1179,2038]" pageId="3" pageNumber="504">
<emphasis id="B93CEAF83616494EFF0FFA2EFEA5FA1A" bold="true" box="[199,322,1491,1518]" pageId="3" pageNumber="504">Diagnosis.</emphasis>
Morphological analyses of the four specimens from the highlands of
<collectingRegion id="498CF8083616494EFB83FA29FB68FA1A" box="[1099,1167,1492,1518]" country="Malaysia" name="Sabah" pageId="3" pageNumber="504">Sabah</collectingRegion>
confirms the placement of this population in the genus
<taxonomicName id="4C484D693616494EFE3BFA0BFD72F9E4" authorityName="Peters" authorityYear="1864" box="[499,661,1526,1552]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="504" phylum="Chordata" rank="genus">
<emphasis id="B93CEAF83616494EFE3BFA0BFD72F9E4" box="[499,661,1526,1552]" italics="true" pageId="3" pageNumber="504">Asthenodipsas</emphasis>
</taxonomicName>
based on the combination of the following characters: smooth dorsal scales; absence of preoculars and suboculars; one or more supralabials in contact with the eye; a single anterior inframaxillary, followed by two or three pairs of inframaxillaries which are wider than long (
<bibRefCitation id="EFD94B1B3616494EFB61F9C1FF33F98D" author="Grossmann, W. &amp; Tillack, F." pageId="3" pageNumber="504" pagination="175 - 90" refId="ref16457" refString="Grossmann, W. &amp; Tillack, F. (2003) On the Taxonomic Status of Asthenodipsas tropidonotus (Van Lidth De Jeude, 1923) and Pareas vertebralis (Boulenger, 1900) (Serpentes: Colubridae: Pareatinae). Russian Journal of Herpetology, 10 (3), 175 - 90." type="journal article" year="2003">Grossmann &amp; Tillack 2003</bibRefCitation>
).
<taxonomicName id="4C484D693616494EFF20F9A3FE1FF98C" authority="Quah &amp; Grismer &amp; Lim &amp; Anuar &amp; Imbun, 2019" authorityName="Quah &amp; Grismer &amp; Lim &amp; Anuar &amp; Imbun" authorityYear="2019" box="[232,504,1630,1656]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="504" phylum="Chordata" rank="species" species="stuebingi" status="sp. nov.">
<emphasis id="B93CEAF83616494EFF20F9A3FE1FF98C" box="[232,504,1630,1656]" italics="true" pageId="3" pageNumber="504">Asthenodipsas stuebingi</emphasis>
</taxonomicName>
<emphasis id="B93CEAF83616494EFDC8F9A2FDBCF98D" bold="true" box="[512,603,1631,1657]" pageId="3" pageNumber="504">
<taxonomicNameLabel id="A20F57833616494EFDC8F9A2FDBCF98D" box="[512,603,1631,1657]" pageId="3" pageNumber="504" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be differentiated from its congeners by the following combination of characters: a maximum SVL of
<quantity id="4CB09B0F3616494EFDCEF97DFD80F968" box="[518,615,1664,1692]" metricMagnitude="-1" metricUnit="m" metricValue="5.57" pageId="3" pageNumber="504" unit="mm" value="557.0">557 mm</quantity>
; 15/15/15 dorsal scale rows; 165175 ventrals; 3547 subcaudal scales; one or two postoculars; 2(+01)+2 temporals; six supralabials, 3
<superScript id="7C3D9BA23616494EFCA0F95EFC91F945" attach="left" box="[872,886,1699,1713]" fontSize="6" pageId="3" pageNumber="504">rd</superScript>
&amp; 4
<superScript id="7C3D9BA23616494EFC6FF95EFC53F945" attach="left" box="[935,948,1699,1713]" fontSize="6" pageId="3" pageNumber="504">th</superScript>
touching the eye; 46 infralabials, 2
<superScript id="7C3D9BA23616494EFA84F95EFABBF945" attach="left" box="[1356,1372,1699,1713]" fontSize="6" pageId="3" pageNumber="504">nd</superScript>
or 3
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pair in contact; a sharp vertebral keel; dorsum of adults beige to brown and overlain with 3042 irregularly-shaped, rhomboidal dark brown bands beginning posterior to dark patch on the neck that and extends the length of body and tail and onto lateral edges of the ventral scales to form spots, but not encircling body; a narrow, light-coloured vertebral stripe; throat and ventrals beige to yellow and mottled with small, dark spots; head whitish to light-brown with dark speckling on the snout and crown; and iris and pupils black (
<tableCitation id="C6CA03513616494EFC67F8ACFBE1F89F" box="[943,1030,1873,1899]" captionStart="TABLE 4" captionStartId="9.[151,239,461,487]" captionTargetPageId="9" captionText="TABLE 4. Scale counts and selected measurements (mm) of the type series of Asthenodipsas jamilinaisi sp. nov. and A. stuebingi sp. nov. Abbreviations are listed in the Materials and methods." httpUri="http://table.plazi.org/id/DF376662361C4944FF5FFE30FEADFDFF" pageId="3" pageNumber="504" tableUuid="DF376662361C4944FF5FFE30FEADFDFF">Table 4</tableCitation>
,
<tableCitation id="C6CA03513616494EFBDAF8ADFBC7F89F" box="[1042,1056,1872,1899]" captionStart="TABLE 5" captionStartId="15.[151,239,464,490]" captionTargetPageId="15" captionText="TABLE 5. Measurements and scale counts of Asthenodipsas jamilinaisi sp. nov., A. stuebingi sp. nov. and different populations of A. laevis. (*measurements from Grossmann &amp; Tillack 2003; NA = measurement not available). Abbreviations are listed in the Materials and methods." httpUri="http://table.plazi.org/id/DF376662361A4942FF5FFE2DFB05FDFA" pageId="3" pageNumber="504" tableUuid="DF376662361A4942FF5FFE2DFB05FDFA">5</tableCitation>
&amp;
<tableCitation id="C6CA03513616494EFB8CF8ACFBB3F89F" box="[1092,1108,1873,1899]" captionStart="TABLE 6" captionStartId="22.[151,239,701,727]" captionTargetPageId="22" captionText="TABLE 6. Comparison of selected characters between Asthenodipsas jamilinaisi sp. nov., A. stuebingi sp. nov. and different populations of A. laevis with diagnostic characters in bold. Abbreviations are listed in the Materials and Methods. ND = No data." httpUri="http://table.plazi.org/id/DF3766623603495BFF5FFD40FC30FD0F" pageId="3" pageNumber="504" tableUuid="DF3766623603495BFF5FFD40FC30FD0F">6</tableCitation>
;
<figureCitation id="13732A6F3616494EFBA8F8ACFB45F89F" box="[1120,1186,1873,1899]" captionStart="FIGURE 2" captionStartId="6.[151,250,1855,1880]" captionTargetBox="[151,1436,381,1836]" captionTargetId="figure@6.[151,1436,381,1836]" captionTargetPageId="6" captionText="FIGURE 2. A. Dorsum of holotype of Asthenodipsas stuebingi sp. nov. (SP 04679). B. Venter of holotype of A. stuebingi sp. nov. (SP 04679). C. Dorsum of paratype of A. stuebingi sp. nov. (SP 04806). D. Venter of paratype of A. stuebingi sp. nov. (SP 04806). E. Dorsum of paratype of A. stuebingi sp. nov. (MCZ R43591). F. Venter of paratype of A. stuebingi sp. nov. (MCZ R43591)." httpUri="https://zenodo.org/record/3351076/files/figure.png" pageId="3" pageNumber="504">Fig. 2</figureCitation>
&amp;
<figureCitation id="13732A6F3616494EFB0EF8ACFB31F89F" box="[1222,1238,1873,1899]" captionStart="FIGURE 3" captionStartId="7.[151,250,1588,1613]" captionTargetBox="[151,1436,211,1555]" captionTargetId="figure@7.[151,1436,211,1555]" captionTargetPageId="7" captionText="FIGURE 3. A. Dorsum of paratype of Asthenodipsas stuebingi sp. nov. (ZMB 65429). B. Venter of paratype of A. stuebingi sp. nov. (ZMB 65429) (Photographs by Frank Tillack). C. Dorsum of paratype of A. stuebingi sp. nov. (ZMB 65429) in life from Kinabalu Park power station, Kamborangoh Road, Sabah. D. Paratype of A. stuebingi sp. nov. (ZMB 65429) from Kinabalu Park power station, Kamborangoh Road, Sabah in defensive coils. E. Venter of paratype of A. stuebingi sp. nov. (ZMB 65429) in life from Kinabalu Park power station, Kamborangoh Road, Sabah. (Photographs by Andreas Nӧllert). F. Sharp vertebral keel of the holotype of A. stuebingi sp. nov. (SP 04679) as indicated by arrow." httpUri="https://zenodo.org/record/3351078/files/figure.png" pageId="3" pageNumber="504">3</figureCitation>
).
</paragraph>
<paragraph id="8BF736EA36164949FF0FF88EFD9BFE1F" blockId="3.[151,1437,1179,2038]" lastBlockId="4.[151,1437,153,837]" lastPageId="4" lastPageNumber="505" pageId="3" pageNumber="504">
<emphasis id="B93CEAF83616494EFF0FF88EFE01F87A" bold="true" box="[199,486,1907,1934]" pageId="3" pageNumber="504">
Description of
<typeStatus id="54F388483616494EFEB0F889FE06F87A" box="[376,481,1908,1934]" pageId="3" pageNumber="504" type="holotype">holotype</typeStatus>
.
</emphasis>
Adult female SVL
<quantity id="4CB09B0F3616494EFD0CF88EFCC5F87A" box="[708,802,1907,1934]" metricMagnitude="-1" metricUnit="m" metricValue="5.57" pageId="3" pageNumber="504" unit="mm" value="557.0">557 mm</quantity>
and Tal
<quantity id="4CB09B0F3616494EFC4BF889FC31F87A" box="[899,982,1908,1934]" metricMagnitude="-2" metricUnit="m" metricValue="6.2" pageId="3" pageNumber="504" unit="mm" value="62.0">62 mm</quantity>
; rostral as wide as high; head somewhat bulbous and elongate in shape, longer than wide; nasals undivided; internasals shorter than prefrontals; posterior margin of prefrontals contact eye; frontal hexagonal, slightly wider than long; loreals present, longer than wide; supraoculars subpentagonal, approximately half the length and half the width of frontal; preoculars absent; upper and lower postoculars present on right side of head, lower postocular extending to below orbit; single postocular on left side of head, lower margin extending to below orbit; suboculars absent; supralabials 6/6 with 3rd and 4th contacting orbit and 6th elongate; temporals 2(+1)+2/2+2, i.e. on left side of head there are two anterior temporals and two posterior temporals but between lower anterior and lower posterior temporal is an additional scale resulting in two upper temporals and three lower temporals; mental triangular wider than long; anterior inframaxillary pentagonal, in contact with infralabials 13; two pairs of posterior inframaxillaries following anterior inframaxillary, first pair of posterior inframaxillaries slightly oval, rhomboid-shaped, second pair more heaxagonal, elongated; infralabials 6/6 with 3
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pair in medial contact. Body long, bulky, laterally compressed, bearing a prominent keel-shaped vertebral region; dorsal scales smooth, in 15/15/15 rows, vertebrals slightly enlarged; 170 ventrals; 36 divided subcaudals; cloacal scute entire; tail tapering to a point.
</paragraph>
<paragraph id="8BF736EA36114949FF0FFE0EFE08FCB1" blockId="4.[151,1437,153,837]" pageId="4" pageNumber="505">
<emphasis id="B93CEAF836114949FF0FFE0EFD25FDFA" bold="true" box="[199,706,499,526]" pageId="4" pageNumber="505">
Colouration in preservation (
<figureCitation id="13732A6F36114949FDEEFE0EFD54FDFA" box="[550,691,499,526]" captionStart="FIGURE 2" captionStartId="6.[151,250,1855,1880]" captionTargetBox="[151,1436,381,1836]" captionTargetId="figure@6.[151,1436,381,1836]" captionTargetPageId="6" captionText="FIGURE 2. A. Dorsum of holotype of Asthenodipsas stuebingi sp. nov. (SP 04679). B. Venter of holotype of A. stuebingi sp. nov. (SP 04679). C. Dorsum of paratype of A. stuebingi sp. nov. (SP 04806). D. Venter of paratype of A. stuebingi sp. nov. (SP 04806). E. Dorsum of paratype of A. stuebingi sp. nov. (MCZ R43591). F. Venter of paratype of A. stuebingi sp. nov. (MCZ R43591)." httpUri="https://zenodo.org/record/3351076/files/figure.png" pageId="4" pageNumber="505">Fig 2A &amp; B</figureCitation>
).
</emphasis>
The ground colour of the head, body and tail is beige with dark speckling. The speckling is heavier on the dorsum nearer to the vertebral scale rows and lightens up on the flanks. The top of the head is also heavily speckled and the rostral, nasal, loreal, postoculars, supralabials, mental, inframaxillaries and infralabials are mottled with dark spots. On the neck is a dark patch starting from the nape, behind the posterior margin of the jaw, extending to approximately the position of the 11
<superScript id="7C3D9BA236114949FBEDFD80FBD5FD7F" attach="left" box="[1061,1074,637,651]" fontSize="6" pageId="4" pageNumber="505">th</superScript>
ventral. The dark patch starts as a narrow stripe on the 4
<superScript id="7C3D9BA236114949FE68FD5DFE4AFD5A" attach="left" box="[416,429,672,686]" fontSize="6" pageId="4" pageNumber="505">th</superScript>
&amp; 5
<superScript id="7C3D9BA236114949FE17FD5DFE0BFD5A" attach="left" box="[479,492,672,686]" fontSize="6" pageId="4" pageNumber="505">th</superScript>
dorsal scale rows, gradually widens to dorsal scale rows 15, and does not cross the vertebral scales or meet on the ventrals. Starting behind the dark neck patch are 42 dark, rhomboidal bands ranging from 12 dorsal scales in length along the body and tail. The venter is yellow and bears, dark lateral blotches from the extensions of the dorsal bands that meet the ventral scales as well as scattered dark spots between them. The thin vertebral stripe is light-yellow.
</paragraph>
<caption id="DF37666236114949FF5FFC88FDF0FC45" ID-Table-UUID="DF37666236114949FF5FFC88FDF0FC45" httpUri="http://table.plazi.org/id/DF37666236114949FF5FFC88FDF0FC45" pageId="4" pageNumber="505" targetBox="[166,1413,957,1986]" targetIsTable="true" targetPageId="4">
<paragraph id="8BF736EA36114949FF5FFC88FDF0FC45" blockId="4.[151,1436,885,945]" pageId="4" pageNumber="505">
<emphasis id="B93CEAF836114949FF5FFC88FEEDFC79" bold="true" box="[151,266,885,909]" pageId="4" pageNumber="505">TABLE 2.</emphasis>
Summary statistics for
<taxonomicName id="4C484D6936114949FDCFFC88FCEEFC79" authority="Quah &amp; Grismer &amp; Lim &amp; Anuar &amp; Imbun, 2019" authorityName="Quah &amp; Grismer &amp; Lim &amp; Anuar &amp; Imbun" authorityYear="2019" box="[519,777,885,909]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="505" phylum="Chordata" rank="species" species="stuebingi" status="sp. nov.">
<emphasis id="B93CEAF836114949FDCFFC88FCEEFC79" box="[519,777,885,909]" italics="true" pageId="4" pageNumber="505">Asthenodipsas stuebingi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20F578336114949FCC7FC88FC81FC79" box="[783,870,885,909]" pageId="4" pageNumber="505" rank="species">
<emphasis id="B93CEAF836114949FCC7FC88FCB8FC79" bold="true" box="[783,863,885,909]" pageId="4" pageNumber="505">sp. nov</emphasis>
.
</taxonomicNameLabel>
and the different populations of
<taxonomicName id="4C484D6936114949FB0AFC8BFAFAFC79" authorityName=": Malkmus" authorityYear="1996" box="[1218,1309,885,909]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="505" phylum="Chordata" rank="species" species="laevis">
<emphasis id="B93CEAF836114949FB0AFC8BFAFAFC79" box="[1218,1309,885,909]" italics="true" pageId="4" pageNumber="505">A. laevis</emphasis>
</taxonomicName>
, SD = standard deviation and N = sample size.
</paragraph>
</caption>
<paragraph id="8BF736EA36114949FE2DFC40FAEFF836" pageId="4" pageNumber="505">
<table id="F948C44A3611B6B2FF6EFC40FA62F836" box="[166,1413,957,1986]" gridcols="6" gridrows="27" pageId="4" pageNumber="505">
<tr id="357834A83611B6B2FF6EFC40FA62FC22" box="[166,1413,957,982]" gridrow="0" pageId="4" pageNumber="505" rowspan-0="1">
<th id="76A95DD43611B6B2FE2DFC40FD5DFC22" box="[485,698,957,982]" gridcol="1" gridrow="0" pageId="4" pageNumber="505">
<emphasis id="B93CEAF836114949FE2DFC40FD5DFC21" bold="true" box="[485,698,957,982]" pageId="4" pageNumber="505">
<taxonomicName id="4C484D6936114949FE2DFC40FD84FC22" authority="Quah &amp; Grismer &amp; Lim &amp; Anuar &amp; Imbun, 2019" authorityName="Quah &amp; Grismer &amp; Lim &amp; Anuar &amp; Imbun" authorityYear="2019" box="[485,611,957,982]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="505" phylum="Chordata" rank="species" species="stuebingi" status="sp. nov.">
<emphasis id="B93CEAF836114949FE2DFC40FD84FC22" bold="true" box="[485,611,957,982]" italics="true" pageId="4" pageNumber="505">A. stuebingi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20F578336114949FDA1FC40FD5DFC21" box="[617,698,957,981]" pageId="4" pageNumber="505" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<th id="76A95DD43611B6B2FD13FC40FC95FC22" box="[731,882,957,982]" gridcol="2" gridrow="0" pageId="4" pageNumber="505">
<taxonomicName id="4C484D6936114949FD13FC40FCD3FC22" authorityName=": Malkmus" authorityYear="1996" box="[731,820,957,982]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="505" phylum="Chordata" rank="species" species="laevis">
<emphasis id="B93CEAF836114949FD13FC40FCD3FC22" bold="true" box="[731,820,957,982]" italics="true" pageId="4" pageNumber="505">A. laevis</emphasis>
</taxonomicName>
</th>
<th id="76A95DD43611B6B2FC5AFC40FBCEFC22" box="[914,1065,957,982]" gridcol="3" gridrow="0" pageId="4" pageNumber="505">
<taxonomicName id="4C484D6936114949FC5AFC40FC0CFC22" authorityName=": Malkmus" authorityYear="1996" box="[914,1003,957,982]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="505" phylum="Chordata" rank="species" species="laevis">
<emphasis id="B93CEAF836114949FC5AFC40FC0CFC22" bold="true" box="[914,1003,957,982]" italics="true" pageId="4" pageNumber="505">A. laevis</emphasis>
</taxonomicName>
</th>
<th id="76A95DD43611B6B2FB84FC40FB31FC22" box="[1100,1238,957,982]" gridcol="4" gridrow="0" pageId="4" pageNumber="505">
<taxonomicName id="4C484D6936114949FB84FC40FB43FC22" authorityName=": Malkmus" authorityYear="1996" box="[1100,1188,957,982]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="505" phylum="Chordata" rank="species" species="laevis">
<emphasis id="B93CEAF836114949FB84FC40FB43FC22" bold="true" box="[1100,1188,957,982]" italics="true" pageId="4" pageNumber="505">A. laevis</emphasis>
</taxonomicName>
</th>
<th id="76A95DD43611B6B2FB33FC40FA62FC22" box="[1275,1413,957,982]" gridcol="5" gridrow="0" pageId="4" pageNumber="505">
<emphasis id="B93CEAF836114949FB33FC40FAB3FC22" bold="true" box="[1275,1364,957,982]" italics="true" pageId="4" pageNumber="505">A. laevis</emphasis>
</th>
</tr>
<tr id="357834A83611B6B2FF6EFC1CFA62FC0E" box="[166,1413,993,1018]" gridrow="1" pageId="4" pageNumber="505" rowspan-0="1" rowspan-1="1">
<td id="76A95DD43611B6B2FD13FC1CFC95FC0E" box="[731,882,993,1018]" gridcol="2" gridrow="1" pageId="4" pageNumber="505">
<emphasis id="B93CEAF836114949FD13FC1CFCCDFC0D" bold="true" box="[731,810,993,1017]" pageId="4" pageNumber="505">Borneo</emphasis>
</td>
<td id="76A95DD43611B6B2FC5AFC1CFBCEFC0E" box="[914,1065,993,1018]" gridcol="3" gridrow="1" pageId="4" pageNumber="505">
<emphasis id="B93CEAF836114949FC5AFC1CFBF5FC0E" bold="true" box="[914,1042,993,1018]" pageId="4" pageNumber="505">Thai-Malay</emphasis>
</td>
<td id="76A95DD43611B6B2FB84FC1CFB31FC0E" box="[1100,1238,993,1018]" gridcol="4" gridrow="1" pageId="4" pageNumber="505">
<emphasis id="B93CEAF836114949FB84FC1CFB98FC0D" bold="true" box="[1100,1151,993,1017]" pageId="4" pageNumber="505">Java</emphasis>
</td>
<td id="76A95DD43611B6B2FB33FC1CFA62FC0E" box="[1275,1413,993,1018]" gridcol="5" gridrow="1" pageId="4" pageNumber="505">
<emphasis id="B93CEAF836114949FB33FC1CFABEFC0E" bold="true" box="[1275,1369,993,1018]" pageId="4" pageNumber="505">Sumatra</emphasis>
</td>
</tr>
<tr id="357834A83611B6B2FF6EFBF8FA62FBEA" box="[166,1413,1029,1054]" gridrow="2" pageId="4" pageNumber="505" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-4="1" rowspan-5="1">
<td id="76A95DD43611B6B2FC5AFBF8FBCEFBEA" box="[914,1065,1029,1054]" gridcol="3" gridrow="2" pageId="4" pageNumber="505">
<emphasis id="B93CEAF836114949FC5AFBF8FC1BFBEA" bold="true" box="[914,1020,1029,1054]" pageId="4" pageNumber="505">Peninsula</emphasis>
</td>
</tr>
<tr id="357834A83611B6B2FF6EFBD8FA62FBCA" box="[166,1413,1061,1086]" gridrow="3" pageId="4" pageNumber="505" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="76A95DD43611B6B2FF6EFBD8FE2FFBCA" box="[166,456,1061,1086]" gridcol="0" gridrow="3" pageId="4" pageNumber="505">
<emphasis id="B93CEAF836114949FF6EFBD8FE2FFBCA" bold="true" box="[166,456,1061,1086]" pageId="4" pageNumber="505">Posterior dorsal scale rows</emphasis>
</th>
</tr>
<tr id="357834A83611B6B2FF6EFBB6FA62FB90" box="[166,1413,1099,1124]" gridrow="4" pageId="4" pageNumber="505">
<th id="76A95DD43611B6B2FF6EFBB6FE2FFB90" box="[166,456,1099,1124]" gridcol="0" gridrow="4" pageId="4" pageNumber="505">mean (±SD)</th>
<td id="76A95DD43611B6B2FE2DFBB6FD5DFB90" box="[485,698,1099,1124]" gridcol="1" gridrow="4" pageId="4" pageNumber="505">15.00 (±0.00)</td>
<td id="76A95DD43611B6B2FD13FBB6FC95FB90" box="[731,882,1099,1124]" gridcol="2" gridrow="4" pageId="4" pageNumber="505">13.00 (±0.00)</td>
<td id="76A95DD43611B6B2FC5AFBB6FBCEFB90" box="[914,1065,1099,1124]" gridcol="3" gridrow="4" pageId="4" pageNumber="505">13.00 (±0.00)</td>
<td id="76A95DD43611B6B2FB84FBB6FB31FB90" box="[1100,1238,1099,1124]" gridcol="4" gridrow="4" pageId="4" pageNumber="505">13.00 (±0.00)</td>
<td id="76A95DD43611B6B2FB33FBB6FA62FB90" box="[1275,1413,1099,1124]" gridcol="5" gridrow="4" pageId="4" pageNumber="505">13.00 (±0.00)</td>
</tr>
<tr id="357834A83611B6B2FF6EFB8CFA62FB7E" box="[166,1413,1137,1162]" gridrow="5" pageId="4" pageNumber="505">
<th id="76A95DD43611B6B2FF6EFB8CFE2FFB7E" box="[166,456,1137,1162]" gridcol="0" gridrow="5" pageId="4" pageNumber="505">range</th>
<td id="76A95DD43611B6B2FE2DFB8CFD5DFB7E" box="[485,698,1137,1162]" gridcol="1" gridrow="5" pageId="4" pageNumber="505">15</td>
<td id="76A95DD43611B6B2FD13FB8CFC95FB7E" box="[731,882,1137,1162]" gridcol="2" gridrow="5" pageId="4" pageNumber="505">13</td>
<td id="76A95DD43611B6B2FC5AFB8CFBCEFB7E" box="[914,1065,1137,1162]" gridcol="3" gridrow="5" pageId="4" pageNumber="505">13</td>
<td id="76A95DD43611B6B2FB84FB8CFB31FB7E" box="[1100,1238,1137,1162]" gridcol="4" gridrow="5" pageId="4" pageNumber="505">13</td>
<td id="76A95DD43611B6B2FB33FB8CFA62FB7E" box="[1275,1413,1137,1162]" gridcol="5" gridrow="5" pageId="4" pageNumber="505">13</td>
</tr>
<tr id="357834A83611B6B2FF6EFB6AFA62FB44" box="[166,1413,1175,1200]" gridrow="6" pageId="4" pageNumber="505">
<th id="76A95DD43611B6B2FF6EFB6AFE2FFB44" box="[166,456,1175,1200]" gridcol="0" gridrow="6" pageId="4" pageNumber="505">N</th>
<td id="76A95DD43611B6B2FE2DFB6AFD5DFB44" box="[485,698,1175,1200]" gridcol="1" gridrow="6" pageId="4" pageNumber="505">4</td>
<td id="76A95DD43611B6B2FD13FB6AFC95FB44" box="[731,882,1175,1200]" gridcol="2" gridrow="6" pageId="4" pageNumber="505">43</td>
<td id="76A95DD43611B6B2FC5AFB6AFBCEFB44" box="[914,1065,1175,1200]" gridcol="3" gridrow="6" pageId="4" pageNumber="505">9</td>
<td id="76A95DD43611B6B2FB84FB6AFB31FB44" box="[1100,1238,1175,1200]" gridcol="4" gridrow="6" pageId="4" pageNumber="505">11</td>
<td id="76A95DD43611B6B2FB33FB6AFA62FB44" box="[1275,1413,1175,1200]" gridcol="5" gridrow="6" pageId="4" pageNumber="505">5</td>
</tr>
<tr id="357834A83611B6B2FF6EFB40FA62FB21" box="[166,1413,1213,1237]" gridrow="7" pageId="4" pageNumber="505">
<th id="76A95DD43611B6B2FF6EFB40FA62FB21" box="[166,1413,1213,1237]" colspan="6" colspanRight="5" gridcol="0" gridrow="7" pageId="4" pageNumber="505">
<emphasis id="B93CEAF836114949FF6EFB40FEE6FB21" bold="true" box="[166,257,1213,1237]" pageId="4" pageNumber="505">Ventrals</emphasis>
</th>
</tr>
<tr id="357834A83611B6B2FF6EFB1FFA62FB0F" box="[166,1413,1250,1275]" gridrow="8" pageId="4" pageNumber="505">
<th id="76A95DD43611B6B2FF6EFB1FFE2FFB0F" box="[166,456,1250,1275]" gridcol="0" gridrow="8" pageId="4" pageNumber="505">mean (±SD)</th>
<td id="76A95DD43611B6B2FE2DFB1FFD5DFB0F" box="[485,698,1250,1275]" gridcol="1" gridrow="8" pageId="4" pageNumber="505">168.75 (±4.79)</td>
<td id="76A95DD43611B6B2FD13FB1FFC95FB0F" box="[731,882,1250,1275]" gridcol="2" gridrow="8" pageId="4" pageNumber="505">158.16 (±4.46)</td>
<td id="76A95DD43611B6B2FC5AFB1FFBCEFB0F" box="[914,1065,1250,1275]" gridcol="3" gridrow="8" pageId="4" pageNumber="505">157.44 (±7.09)</td>
<td id="76A95DD43611B6B2FB84FB1FFB31FB0F" box="[1100,1238,1250,1275]" gridcol="4" gridrow="8" pageId="4" pageNumber="505">157.64</td>
<td id="76A95DD43611B6B2FB33FB1FFA62FB0F" box="[1275,1413,1250,1275]" gridcol="5" gridrow="8" pageId="4" pageNumber="505">162.20</td>
</tr>
<tr id="357834A83611B6B2FF6EFAFBFA62FAB4" box="[166,1413,1286,1344]" gridrow="9" pageId="4" pageNumber="505">
<th id="76A95DD43611B6B2FF6EFAFBFE2FFAB4" box="[166,456,1286,1344]" gridcol="0" gridrow="9" pageId="4" pageNumber="505">range</th>
<td id="76A95DD43611B6B2FE2DFAFBFD5DFAB4" box="[485,698,1286,1344]" gridcol="1" gridrow="9" pageId="4" pageNumber="505">165175</td>
<td id="76A95DD43611B6B2FD13FAFBFC95FAB4" box="[731,882,1286,1344]" gridcol="2" gridrow="9" pageId="4" pageNumber="505">150167</td>
<td id="76A95DD43611B6B2FC5AFAFBFBCEFAB4" box="[914,1065,1286,1344]" gridcol="3" gridrow="9" pageId="4" pageNumber="505">148173</td>
<td id="76A95DD43611B6B2FB84FAFBFB31FAB4" box="[1100,1238,1286,1344]" gridcol="4" gridrow="9" pageId="4" pageNumber="505">(±6.17) 150165</td>
<td id="76A95DD43611B6B2FB33FAFBFA62FAB4" box="[1275,1413,1286,1344]" gridcol="5" gridrow="9" pageId="4" pageNumber="505">(±2.59) 158165</td>
</tr>
<tr id="357834A83611B6B2FF6EFAB0FA62FA92" box="[166,1413,1357,1382]" gridrow="10" pageId="4" pageNumber="505">
<th id="76A95DD43611B6B2FF6EFAB0FE2FFA92" box="[166,456,1357,1382]" gridcol="0" gridrow="10" pageId="4" pageNumber="505">N</th>
<td id="76A95DD43611B6B2FE2DFAB0FD5DFA92" box="[485,698,1357,1382]" gridcol="1" gridrow="10" pageId="4" pageNumber="505">4</td>
<td id="76A95DD43611B6B2FD13FAB0FC95FA92" box="[731,882,1357,1382]" gridcol="2" gridrow="10" pageId="4" pageNumber="505">43</td>
<td id="76A95DD43611B6B2FC5AFAB0FBCEFA92" box="[914,1065,1357,1382]" gridcol="3" gridrow="10" pageId="4" pageNumber="505">9</td>
<td id="76A95DD43611B6B2FB84FAB0FB31FA92" box="[1100,1238,1357,1382]" gridcol="4" gridrow="10" pageId="4" pageNumber="505">11</td>
<td id="76A95DD43611B6B2FB33FAB0FA62FA92" box="[1275,1413,1357,1382]" gridcol="5" gridrow="10" pageId="4" pageNumber="505">5</td>
</tr>
<tr id="357834A83611B6B2FF6EFA8FFA62FA7F" box="[166,1413,1394,1419]" gridrow="11" pageId="4" pageNumber="505">
<th id="76A95DD43611B6B2FF6EFA8FFA62FA7F" box="[166,1413,1394,1419]" colspan="6" colspanRight="5" gridcol="0" gridrow="11" pageId="4" pageNumber="505">
<emphasis id="B93CEAF836114949FF6EFA8FFEC5FA7F" bold="true" box="[166,290,1394,1419]" pageId="4" pageNumber="505">Subcaudals</emphasis>
</th>
</tr>
<tr id="357834A83611B6B2FF6EFA65FA62FA45" box="[166,1413,1432,1457]" gridrow="12" pageId="4" pageNumber="505">
<th id="76A95DD43611B6B2FF6EFA65FE2FFA45" box="[166,456,1432,1457]" gridcol="0" gridrow="12" pageId="4" pageNumber="505">mean (±SD)</th>
<td id="76A95DD43611B6B2FE2DFA65FD5DFA45" box="[485,698,1432,1457]" gridcol="1" gridrow="12" pageId="4" pageNumber="505">39.50 (±5.45)</td>
<td id="76A95DD43611B6B2FD13FA65FC95FA45" box="[731,882,1432,1457]" gridcol="2" gridrow="12" pageId="4" pageNumber="505">41.53 (±3.95)</td>
<td id="76A95DD43611B6B2FC5AFA65FBCEFA45" box="[914,1065,1432,1457]" gridcol="3" gridrow="12" pageId="4" pageNumber="505">42.89 (±5.11)</td>
<td id="76A95DD43611B6B2FB84FA65FB31FA45" box="[1100,1238,1432,1457]" gridcol="4" gridrow="12" pageId="4" pageNumber="505">39.91 (±3.33)</td>
<td id="76A95DD43611B6B2FB33FA65FA62FA45" box="[1275,1413,1432,1457]" gridcol="5" gridrow="12" pageId="4" pageNumber="505">43.8 (±4.97)</td>
</tr>
<tr id="357834A83611B6B2FF6EFA43FA62FA23" box="[166,1413,1470,1495]" gridrow="13" pageId="4" pageNumber="505">
<th id="76A95DD43611B6B2FF6EFA43FE2FFA23" box="[166,456,1470,1495]" gridcol="0" gridrow="13" pageId="4" pageNumber="505">range</th>
<td id="76A95DD43611B6B2FE2DFA43FD5DFA23" box="[485,698,1470,1495]" gridcol="1" gridrow="13" pageId="4" pageNumber="505">3547</td>
<td id="76A95DD43611B6B2FD13FA43FC95FA23" box="[731,882,1470,1495]" gridcol="2" gridrow="13" pageId="4" pageNumber="505">3449</td>
<td id="76A95DD43611B6B2FC5AFA43FBCEFA23" box="[914,1065,1470,1495]" gridcol="3" gridrow="13" pageId="4" pageNumber="505">3751</td>
<td id="76A95DD43611B6B2FB84FA43FB31FA23" box="[1100,1238,1470,1495]" gridcol="4" gridrow="13" pageId="4" pageNumber="505">3544</td>
<td id="76A95DD43611B6B2FB33FA43FA62FA23" box="[1275,1413,1470,1495]" gridcol="5" gridrow="13" pageId="4" pageNumber="505">3750</td>
</tr>
<tr id="357834A83611B6B2FF6EFA19FA62FA09" box="[166,1413,1508,1533]" gridrow="14" pageId="4" pageNumber="505">
<th id="76A95DD43611B6B2FF6EFA19FE2FFA09" box="[166,456,1508,1533]" gridcol="0" gridrow="14" pageId="4" pageNumber="505">N</th>
<td id="76A95DD43611B6B2FE2DFA19FD5DFA09" box="[485,698,1508,1533]" gridcol="1" gridrow="14" pageId="4" pageNumber="505">4</td>
<td id="76A95DD43611B6B2FD13FA19FC95FA09" box="[731,882,1508,1533]" gridcol="2" gridrow="14" pageId="4" pageNumber="505">43</td>
<td id="76A95DD43611B6B2FC5AFA19FBCEFA09" box="[914,1065,1508,1533]" gridcol="3" gridrow="14" pageId="4" pageNumber="505">9</td>
<td id="76A95DD43611B6B2FB84FA19FB31FA09" box="[1100,1238,1508,1533]" gridcol="4" gridrow="14" pageId="4" pageNumber="505">11</td>
<td id="76A95DD43611B6B2FB33FA19FA62FA09" box="[1275,1413,1508,1533]" gridcol="5" gridrow="14" pageId="4" pageNumber="505">5</td>
</tr>
<tr id="357834A83611B6B2FF6EF9F4FA62F9D6" box="[166,1413,1545,1570]" gridrow="15" pageId="4" pageNumber="505">
<th id="76A95DD43611B6B2FF6EF9F4FA62F9D6" box="[166,1413,1545,1570]" colspan="6" colspanRight="5" gridcol="0" gridrow="15" pageId="4" pageNumber="505">
<emphasis id="B93CEAF836114949FF6EF9F4FEC8F9D6" bold="true" box="[166,303,1545,1570]" pageId="4" pageNumber="505">Supralabials</emphasis>
</th>
</tr>
<tr id="357834A83611B6B2FF6EF9D2FA62F9BC" box="[166,1413,1583,1608]" gridrow="16" pageId="4" pageNumber="505">
<th id="76A95DD43611B6B2FF6EF9D2FE2FF9BC" box="[166,456,1583,1608]" gridcol="0" gridrow="16" pageId="4" pageNumber="505">mean (±SD)</th>
<td id="76A95DD43611B6B2FE2DF9D2FD5DF9BC" box="[485,698,1583,1608]" gridcol="1" gridrow="16" pageId="4" pageNumber="505">6.0 (±0.0)</td>
<td id="76A95DD43611B6B2FD13F9D2FC95F9BC" box="[731,882,1583,1608]" gridcol="2" gridrow="16" pageId="4" pageNumber="505">6.02 (±0.15)</td>
<td id="76A95DD43611B6B2FC5AF9D2FBCEF9BC" box="[914,1065,1583,1608]" gridcol="3" gridrow="16" pageId="4" pageNumber="505">6.00 (±0.00)</td>
<td id="76A95DD43611B6B2FB84F9D2FB31F9BC" box="[1100,1238,1583,1608]" gridcol="4" gridrow="16" pageId="4" pageNumber="505">6.09 (±0.30)</td>
<td id="76A95DD43611B6B2FB33F9D2FA62F9BC" box="[1275,1413,1583,1608]" gridcol="5" gridrow="16" pageId="4" pageNumber="505">6.00 (±0.00)</td>
</tr>
<tr id="357834A83611B6B2FF6EF9A8FA62F99A" box="[166,1413,1621,1646]" gridrow="17" pageId="4" pageNumber="505">
<th id="76A95DD43611B6B2FF6EF9A8FE2FF99A" box="[166,456,1621,1646]" gridcol="0" gridrow="17" pageId="4" pageNumber="505">range</th>
<td id="76A95DD43611B6B2FE2DF9A8FD5DF99A" box="[485,698,1621,1646]" gridcol="1" gridrow="17" pageId="4" pageNumber="505">6.0</td>
<td id="76A95DD43611B6B2FD13F9A8FC95F99A" box="[731,882,1621,1646]" gridcol="2" gridrow="17" pageId="4" pageNumber="505">67</td>
<td id="76A95DD43611B6B2FC5AF9A8FBCEF99A" box="[914,1065,1621,1646]" gridcol="3" gridrow="17" pageId="4" pageNumber="505">6.00</td>
<td id="76A95DD43611B6B2FB84F9A8FB31F99A" box="[1100,1238,1621,1646]" gridcol="4" gridrow="17" pageId="4" pageNumber="505">67</td>
<td id="76A95DD43611B6B2FB33F9A8FA62F99A" box="[1275,1413,1621,1646]" gridcol="5" gridrow="17" pageId="4" pageNumber="505">6.00</td>
</tr>
<tr id="357834A83611B6B2FF6EF986FA62F960" box="[166,1413,1659,1684]" gridrow="18" pageId="4" pageNumber="505">
<th id="76A95DD43611B6B2FF6EF986FE2FF960" box="[166,456,1659,1684]" gridcol="0" gridrow="18" pageId="4" pageNumber="505">N</th>
<td id="76A95DD43611B6B2FE2DF986FD5DF960" box="[485,698,1659,1684]" gridcol="1" gridrow="18" pageId="4" pageNumber="505">4</td>
<td id="76A95DD43611B6B2FD13F986FC95F960" box="[731,882,1659,1684]" gridcol="2" gridrow="18" pageId="4" pageNumber="505">43</td>
<td id="76A95DD43611B6B2FC5AF986FBCEF960" box="[914,1065,1659,1684]" gridcol="3" gridrow="18" pageId="4" pageNumber="505">9</td>
<td id="76A95DD43611B6B2FB84F986FB31F960" box="[1100,1238,1659,1684]" gridcol="4" gridrow="18" pageId="4" pageNumber="505">11</td>
<td id="76A95DD43611B6B2FB33F986FA62F960" box="[1275,1413,1659,1684]" gridcol="5" gridrow="18" pageId="4" pageNumber="505">5</td>
</tr>
<tr id="357834A83611B6B2FF6EF95DFA62F94D" box="[166,1413,1696,1721]" gridrow="19" pageId="4" pageNumber="505">
<th id="76A95DD43611B6B2FF6EF95DFA62F94D" box="[166,1413,1696,1721]" colspan="6" colspanRight="5" gridcol="0" gridrow="19" pageId="4" pageNumber="505">
<emphasis id="B93CEAF836114949FF6EF95DFEC2F94D" bold="true" box="[166,293,1696,1721]" pageId="4" pageNumber="505">Infralabials</emphasis>
</th>
</tr>
<tr id="357834A83611B6B2FF6EF93AFA62F914" box="[166,1413,1735,1760]" gridrow="20" pageId="4" pageNumber="505">
<th id="76A95DD43611B6B2FF6EF93AFE2FF914" box="[166,456,1735,1760]" gridcol="0" gridrow="20" pageId="4" pageNumber="505">mean (±SD)</th>
<td id="76A95DD43611B6B2FE2DF93AFD5DF914" box="[485,698,1735,1760]" gridcol="1" gridrow="20" pageId="4" pageNumber="505">5.00 (±1.15)</td>
<td id="76A95DD43611B6B2FD13F93AFC95F914" box="[731,882,1735,1760]" gridcol="2" gridrow="20" pageId="4" pageNumber="505">5.93 (±0.40)</td>
<td id="76A95DD43611B6B2FC5AF93AFBCEF914" box="[914,1065,1735,1760]" gridcol="3" gridrow="20" pageId="4" pageNumber="505">5.44 (±0.53)</td>
<td id="76A95DD43611B6B2FB84F93AFB31F914" box="[1100,1238,1735,1760]" gridcol="4" gridrow="20" pageId="4" pageNumber="505">5.00 (±0.45)</td>
<td id="76A95DD43611B6B2FB33F93AFA62F914" box="[1275,1413,1735,1760]" gridcol="5" gridrow="20" pageId="4" pageNumber="505">5.2 (±0.45)</td>
</tr>
<tr id="357834A83611B6B2FF6EF911FA62F8F2" box="[166,1413,1772,1798]" gridrow="21" pageId="4" pageNumber="505">
<th id="76A95DD43611B6B2FF6EF911FE2FF8F2" box="[166,456,1772,1798]" gridcol="0" gridrow="21" pageId="4" pageNumber="505">range</th>
<td id="76A95DD43611B6B2FE2DF911FD5DF8F2" box="[485,698,1772,1798]" gridcol="1" gridrow="21" pageId="4" pageNumber="505">46</td>
<td id="76A95DD43611B6B2FD13F911FC95F8F2" box="[731,882,1772,1798]" gridcol="2" gridrow="21" pageId="4" pageNumber="505">57</td>
<td id="76A95DD43611B6B2FC5AF911FBCEF8F2" box="[914,1065,1772,1798]" gridcol="3" gridrow="21" pageId="4" pageNumber="505">56</td>
<td id="76A95DD43611B6B2FB84F911FB31F8F2" box="[1100,1238,1772,1798]" gridcol="4" gridrow="21" pageId="4" pageNumber="505">46</td>
<td id="76A95DD43611B6B2FB33F911FA62F8F2" box="[1275,1413,1772,1798]" gridcol="5" gridrow="21" pageId="4" pageNumber="505">56</td>
</tr>
<tr id="357834A83611B6B2FF6EF8EFFA62F8DF" box="[166,1413,1810,1835]" gridrow="22" pageId="4" pageNumber="505">
<th id="76A95DD43611B6B2FF6EF8EFFE2FF8DF" box="[166,456,1810,1835]" gridcol="0" gridrow="22" pageId="4" pageNumber="505">N</th>
<td id="76A95DD43611B6B2FE2DF8EFFD5DF8DF" box="[485,698,1810,1835]" gridcol="1" gridrow="22" pageId="4" pageNumber="505">4</td>
<td id="76A95DD43611B6B2FD13F8EFFC95F8DF" box="[731,882,1810,1835]" gridcol="2" gridrow="22" pageId="4" pageNumber="505">43</td>
<td id="76A95DD43611B6B2FC5AF8EFFBCEF8DF" box="[914,1065,1810,1835]" gridcol="3" gridrow="22" pageId="4" pageNumber="505">9</td>
<td id="76A95DD43611B6B2FB84F8EFFB31F8DF" box="[1100,1238,1810,1835]" gridcol="4" gridrow="22" pageId="4" pageNumber="505">11</td>
<td id="76A95DD43611B6B2FB33F8EFFA62F8DF" box="[1275,1413,1810,1835]" gridcol="5" gridrow="22" pageId="4" pageNumber="505">5</td>
</tr>
<tr id="357834A83611B6B2FF6EF8C5FA62F8A5" box="[166,1413,1848,1873]" gridrow="23" pageId="4" pageNumber="505">
<th id="76A95DD43611B6B2FF6EF8C5FA62F8A5" box="[166,1413,1848,1873]" colspan="6" colspanRight="5" gridcol="0" gridrow="23" pageId="4" pageNumber="505">
<emphasis id="B93CEAF836114949FF6EF8C5FE59F8A4" bold="true" box="[166,446,1848,1873]" pageId="4" pageNumber="505">Infralabial pair in contact</emphasis>
</th>
</tr>
<tr id="357834A83611B6B2FF6EF8A3FA62F883" box="[166,1413,1886,1911]" gridrow="24" pageId="4" pageNumber="505">
<th id="76A95DD43611B6B2FF6EF8A3FE2FF883" box="[166,456,1886,1911]" gridcol="0" gridrow="24" pageId="4" pageNumber="505">mean (±SD)</th>
<td id="76A95DD43611B6B2FE2DF8A3FD5DF883" box="[485,698,1886,1911]" gridcol="1" gridrow="24" pageId="4" pageNumber="505">2.5 (±0.58)</td>
<td id="76A95DD43611B6B2FD13F8A3FC95F883" box="[731,882,1886,1911]" gridcol="2" gridrow="24" pageId="4" pageNumber="505">2.95 (±0.21)</td>
<td id="76A95DD43611B6B2FC5AF8A3FBCEF883" box="[914,1065,1886,1911]" gridcol="3" gridrow="24" pageId="4" pageNumber="505">3.00 (±0.00)</td>
<td id="76A95DD43611B6B2FB84F8A3FB31F883" box="[1100,1238,1886,1911]" gridcol="4" gridrow="24" pageId="4" pageNumber="505">2.82 (±0.40)</td>
<td id="76A95DD43611B6B2FB33F8A3FA62F883" box="[1275,1413,1886,1911]" gridcol="5" gridrow="24" pageId="4" pageNumber="505">3.00 (±0.00)</td>
</tr>
<tr id="357834A83611B6B2FF6EF879FA62F869" box="[166,1413,1924,1949]" gridrow="25" pageId="4" pageNumber="505">
<th id="76A95DD43611B6B2FF6EF879FE2FF869" box="[166,456,1924,1949]" gridcol="0" gridrow="25" pageId="4" pageNumber="505">range</th>
<td id="76A95DD43611B6B2FE2DF879FD5DF869" box="[485,698,1924,1949]" gridcol="1" gridrow="25" pageId="4" pageNumber="505">23</td>
<td id="76A95DD43611B6B2FD13F879FC95F869" box="[731,882,1924,1949]" gridcol="2" gridrow="25" pageId="4" pageNumber="505">23</td>
<td id="76A95DD43611B6B2FC5AF879FBCEF869" box="[914,1065,1924,1949]" gridcol="3" gridrow="25" pageId="4" pageNumber="505">3.00</td>
<td id="76A95DD43611B6B2FB84F879FB31F869" box="[1100,1238,1924,1949]" gridcol="4" gridrow="25" pageId="4" pageNumber="505">23</td>
<td id="76A95DD43611B6B2FB33F879FA62F869" box="[1275,1413,1924,1949]" gridcol="5" gridrow="25" pageId="4" pageNumber="505">3.00</td>
</tr>
<tr id="357834A83611B6B2FF6EF854FA62F836" box="[166,1413,1961,1986]" gridrow="26" pageId="4" pageNumber="505">
<th id="76A95DD43611B6B2FF6EF854FE2FF836" box="[166,456,1961,1986]" gridcol="0" gridrow="26" pageId="4" pageNumber="505">N</th>
<td id="76A95DD43611B6B2FE2DF854FD5DF836" box="[485,698,1961,1986]" gridcol="1" gridrow="26" pageId="4" pageNumber="505">4</td>
<td id="76A95DD43611B6B2FD13F854FC95F836" box="[731,882,1961,1986]" gridcol="2" gridrow="26" pageId="4" pageNumber="505">43</td>
<td id="76A95DD43611B6B2FC5AF854FBCEF836" box="[914,1065,1961,1986]" gridcol="3" gridrow="26" pageId="4" pageNumber="505">9</td>
<td id="76A95DD43611B6B2FB84F854FB31F836" box="[1100,1238,1961,1986]" gridcol="4" gridrow="26" pageId="4" pageNumber="505">11</td>
<td id="76A95DD43611B6B2FB33F854FA62F836" box="[1275,1413,1961,1986]" gridcol="5" gridrow="26" pageId="4" pageNumber="505">5</td>
</tr>
</table>
</paragraph>
<caption id="DF37666236104948FF5FFF67FA7BFF46" ID-Table-UUID="DF37666236104948FF5FFF67FA7BFF46" box="[151,1436,154,178]" httpUri="http://table.plazi.org/id/DF37666236104948FF5FFF67FA7BFF46" pageId="5" pageNumber="506" targetBox="[166,1334,295,601]" targetIsTable="true" targetPageId="5">
<paragraph id="8BF736EA36104948FF5FFF67FA7BFF46" blockId="5.[151,1436,154,286]" box="[151,1436,154,178]" pageId="5" pageNumber="506">
<emphasis id="B93CEAF836104948FF5FFF67FEE4FF46" bold="true" box="[151,259,154,178]" pageId="5" pageNumber="506">TABLE 3</emphasis>
. Pairwise matrix of statistically significant mean differences of characters among species of the
<taxonomicName id="4C484D6936104948FACAFF67FA7BFF46" authorityName="Peters" authorityYear="1864" box="[1282,1436,154,178]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="506" phylum="Chordata" rank="genus">Asthenodipsas</taxonomicName>
</paragraph>
</caption>
<paragraph id="8BF736EA36104948FF5FFF43FA62FF0E" blockId="5.[151,1436,154,286]" pageId="5" pageNumber="506">
<taxonomicName id="4C484D6936104948FF5FFF43FF32FF22" box="[151,213,190,214]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="laevis">
<emphasis id="B93CEAF836104948FF5FFF43FF32FF22" box="[151,213,190,214]" italics="true" pageId="5" pageNumber="506">laevis</emphasis>
</taxonomicName>
group. Abbreviations are in the Materials and methods.
<emphasis id="B93CEAF836104948FE0AFF1FFBCDFF0E" box="[450,1066,226,250]" italics="true" pageId="5" pageNumber="506">
<taxonomicName id="4C484D6936104948FE0AFF1FFDADFF0E" authorityName="Quah &amp; Grismer &amp; Lim &amp; Anuar &amp; Imbun" authorityYear="2019" box="[450,586,226,250]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="jamilinaisi">A. jamilinaisi</taxonomicName>
<taxonomicName id="4C484D6936104948FDB0FF1FFD16FF0E" authorityName="Quah &amp; Grismer &amp; Lim &amp; Anuar &amp; Imbun" authorityYear="2019" box="[632,753,226,250]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="stuebingi">A. stuebingi</taxonomicName>
<taxonomicName id="4C484D6936104948FC9CFF1FFBCDFF0E" box="[852,1066,226,250]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="laevis">A. laevis A. laevis</taxonomicName>
</emphasis>
Thai-Ma-
<taxonomicName id="4C484D6936104948FB67FF1FFA62FF0E" box="[1199,1413,226,250]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="laevis">
<emphasis id="B93CEAF836104948FB67FF1FFA62FF0E" box="[1199,1413,226,250]" italics="true" pageId="5" pageNumber="506">A. laevis A. laevis</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BF736EA36104948FE0AFEFBFF16FC88" blockId="5.[151,1436,154,286]" lastBlockId="5.[151,1437,649,2044]" pageId="5" pageNumber="506">
<emphasis id="B93CEAF836104948FE0AFEFBFD2FFEEA" bold="true" box="[450,712,262,286]" pageId="5" pageNumber="506">sp. nov. sp. nov.</emphasis>
Borneo lay Peninsula
<collectingRegion id="498CF80836104948FB67FEFAFB3BFEEA" box="[1199,1244,263,286]" country="Indonesia" name="Jawa Timur" pageId="5" pageNumber="506">Java</collectingRegion>
<collectingRegion id="498CF80836104948FAE6FEFBFA65FEEA" box="[1326,1410,262,286]" country="Indonesia" name="Sumatera Utara" pageId="5" pageNumber="506">Sumatra</collectingRegion>
<emphasis id="B93CEAF836104948FF0FFD74FDD3FD50" bold="true" box="[199,564,649,676]" pageId="5" pageNumber="506">
Variation (
<tableCitation id="C6CA035136104948FE8FFD74FE44FD57" box="[327,419,649,676]" captionStart="TABLE 4" captionStartId="9.[151,239,461,487]" captionTargetPageId="9" captionText="TABLE 4. Scale counts and selected measurements (mm) of the type series of Asthenodipsas jamilinaisi sp. nov. and A. stuebingi sp. nov. Abbreviations are listed in the Materials and methods." httpUri="http://table.plazi.org/id/DF376662361C4944FF5FFE30FEADFDFF" pageId="5" pageNumber="506" tableUuid="DF376662361C4944FF5FFE30FEADFDFF">Table 4</tableCitation>
;
<figureCitation id="13732A6F36104948FE7AFD74FE09FD57" box="[434,494,649,676]" captionStart="FIGURE 2" captionStartId="6.[151,250,1855,1880]" captionTargetBox="[151,1436,381,1836]" captionTargetId="figure@6.[151,1436,381,1836]" captionTargetPageId="6" captionText="FIGURE 2. A. Dorsum of holotype of Asthenodipsas stuebingi sp. nov. (SP 04679). B. Venter of holotype of A. stuebingi sp. nov. (SP 04679). C. Dorsum of paratype of A. stuebingi sp. nov. (SP 04806). D. Venter of paratype of A. stuebingi sp. nov. (SP 04806). E. Dorsum of paratype of A. stuebingi sp. nov. (MCZ R43591). F. Venter of paratype of A. stuebingi sp. nov. (MCZ R43591)." httpUri="https://zenodo.org/record/3351076/files/figure.png" pageId="5" pageNumber="506">Fig 2</figureCitation>
&amp;
<figureCitation id="13732A6F36104948FDDDFD74FDC2FD50" box="[533,549,649,676]" captionStart="FIGURE 3" captionStartId="7.[151,250,1588,1613]" captionTargetBox="[151,1436,211,1555]" captionTargetId="figure@7.[151,1436,211,1555]" captionTargetPageId="7" captionText="FIGURE 3. A. Dorsum of paratype of Asthenodipsas stuebingi sp. nov. (ZMB 65429). B. Venter of paratype of A. stuebingi sp. nov. (ZMB 65429) (Photographs by Frank Tillack). C. Dorsum of paratype of A. stuebingi sp. nov. (ZMB 65429) in life from Kinabalu Park power station, Kamborangoh Road, Sabah. D. Paratype of A. stuebingi sp. nov. (ZMB 65429) from Kinabalu Park power station, Kamborangoh Road, Sabah in defensive coils. E. Venter of paratype of A. stuebingi sp. nov. (ZMB 65429) in life from Kinabalu Park power station, Kamborangoh Road, Sabah. (Photographs by Andreas Nӧllert). F. Sharp vertebral keel of the holotype of A. stuebingi sp. nov. (SP 04679) as indicated by arrow." httpUri="https://zenodo.org/record/3351078/files/figure.png" pageId="5" pageNumber="506">3</figureCitation>
).
</emphasis>
The
<typeStatus id="54F3884836104948FDA6FD77FD3EFD50" box="[622,729,650,676]" pageId="5" pageNumber="506" type="paratype">paratypes</typeStatus>
closely resemble the
<typeStatus id="54F3884836104948FC07FD74FBD6FD57" box="[975,1073,649,675]" pageId="5" pageNumber="506" type="holotype">holotype</typeStatus>
in overall external morphology, colouration, and pattern. All
<typeStatus id="54F3884836104948FE1EFD53FDA6FD3C" box="[470,577,686,712]" pageId="5" pageNumber="506" type="paratype">paratypes</typeStatus>
have a slightly darker ground colour than the
<typeStatus id="54F3884836104948FBF7FD50FB46FD33" box="[1087,1185,685,711]" pageId="5" pageNumber="506" type="holotype">holotype</typeStatus>
and a larger dark neck patch. In SP 04806 the markings from the dark neck patch extend further onto the edges of the ventrals, leaving a narrow, light-coloured median stripe on the underside of the neck and the tail is more heavily mottled.
<typeStatus id="54F3884836104948FAE7FD0BFA7BFCE4" box="[1327,1436,758,784]" pageId="5" pageNumber="506" type="paratype">Paratypes</typeStatus>
<collectionCode id="ED59AE2F36104948FF5FFCE7FF33FCC0" box="[151,212,794,820]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="5" pageNumber="506" type="Museum">ZMB</collectionCode>
65429 and
<collectionCode id="ED59AE2F36104948FE95FCE7FE7DFCC0" box="[349,410,794,820]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="5" pageNumber="506" type="Museum">MCZ</collectionCode>
R43591 also have fewer infralabials than the
<typeStatus id="54F3884836104948FC62FCE4FBEBFCC7" box="[938,1036,793,819]" pageId="5" pageNumber="506" type="holotype">holotype</typeStatus>
(45 vs. 6) and the second instead of the third pair are in contact. Variation in scale counts and size measurements within the type series are shown in
<tableCitation id="C6CA035136104948FF5FFC9CFF0AFC88" box="[151,237,865,892]" captionStart="TABLE 4" captionStartId="9.[151,239,461,487]" captionTargetPageId="9" captionText="TABLE 4. Scale counts and selected measurements (mm) of the type series of Asthenodipsas jamilinaisi sp. nov. and A. stuebingi sp. nov. Abbreviations are listed in the Materials and methods." httpUri="http://table.plazi.org/id/DF376662361C4944FF5FFE30FEADFDFF" pageId="5" pageNumber="506" tableUuid="DF376662361C4944FF5FFE30FEADFDFF">Table 4</tableCitation>
.
</paragraph>
<paragraph id="8BF736EA36104948FF6EFEDAFAD1FDAC" pageId="5" pageNumber="506">
<table id="F948C44A3610B6B2FF6EFEDAFAD1FDAD" box="[166,1334,295,601]" gridcols="7" gridrows="8" pageId="5" pageNumber="506">
<tr id="357834A83610B6B2FF6EFEDAFAD1FECB" box="[166,1334,295,319]" gridrow="0" pageId="5" pageNumber="506">
<th id="76A95DD43610B6B2FF6EFEDAFE62FECB" box="[166,389,295,319]" gridcol="0" gridrow="0" pageId="5" pageNumber="506">
<taxonomicName id="4C484D6936104948FF6EFEDAFEC9FECB" authority="Quah &amp; Grismer &amp; Lim &amp; Anuar &amp; Imbun, 2019" authorityName="Quah &amp; Grismer &amp; Lim &amp; Anuar &amp; Imbun" authorityYear="2019" box="[166,302,295,319]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="jamilinaisi" status="sp. nov.">
<emphasis id="B93CEAF836104948FF6EFEDAFEC9FECB" box="[166,302,295,319]" italics="true" pageId="5" pageNumber="506">A. jamilinaisi</emphasis>
</taxonomicName>
<emphasis id="B93CEAF836104948FEFCFEDAFE62FECB" bold="true" box="[308,389,295,319]" pageId="5" pageNumber="506">
<taxonomicNameLabel id="A20F578336104948FEFCFEDAFE62FECB" box="[308,389,295,319]" pageId="5" pageNumber="506" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<th id="76A95DD43610B6B2FE0AFEDAFDDCFECB" box="[450,571,295,319]" gridcol="1" gridrow="0" pageId="5" pageNumber="506">-</th>
<th id="76A95DD43610B6B2FDB0FEDAFCCBFECB" box="[632,812,295,319]" gridcol="2" gridrow="0" pageId="5" pageNumber="506">-</th>
<th id="76A95DD43610B6B2FC9CFEDAFC66FECB" box="[852,897,295,319]" gridcol="3" gridrow="0" pageId="5" pageNumber="506">-</th>
<th id="76A95DD43610B6B2FC1BFEDAFC3CFECB" box="[979,987,295,319]" gridcol="4" gridrow="0" pageId="5" pageNumber="506">-</th>
<th id="76A95DD43610B6B2FB67FEDAFB50FECB" box="[1199,1207,295,319]" gridcol="5" gridrow="0" pageId="5" pageNumber="506">-</th>
<th id="76A95DD43610B6B2FAE6FEDAFAD1FECB" box="[1326,1334,295,319]" gridcol="6" gridrow="0" pageId="5" pageNumber="506">-</th>
</tr>
<tr id="357834A83610B6B2FF6EFEB1FAD1FE91" box="[166,1334,332,357]" gridrow="1" pageId="5" pageNumber="506">
<th id="76A95DD43610B6B2FF6EFEB1FE62FE91" box="[166,389,332,357]" gridcol="0" gridrow="1" pageId="5" pageNumber="506">
<taxonomicName id="4C484D6936104948FF6EFEB0FEF8FE91" authority="Quah &amp; Grismer &amp; Lim &amp; Anuar &amp; Imbun, 2019" authorityName="Quah &amp; Grismer &amp; Lim &amp; Anuar &amp; Imbun" authorityYear="2019" box="[166,287,333,357]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="stuebingi" status="sp. nov.">
<emphasis id="B93CEAF836104948FF6EFEB0FEF8FE91" box="[166,287,333,357]" italics="true" pageId="5" pageNumber="506">A. stuebingi</emphasis>
</taxonomicName>
<emphasis id="B93CEAF836104948FEEDFEB0FE92FE91" bold="true" box="[293,373,333,357]" pageId="5" pageNumber="506">
<taxonomicNameLabel id="A20F578336104948FEEDFEB0FE92FE91" box="[293,373,333,357]" pageId="5" pageNumber="506" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<td id="76A95DD43610B6B2FE0AFEB1FDDCFE91" box="[450,571,332,357]" gridcol="1" gridrow="1" pageId="5" pageNumber="506">SubC</td>
<td id="76A95DD43610B6B2FDB0FEB1FCCBFE91" box="[632,812,332,357]" gridcol="2" gridrow="1" pageId="5" pageNumber="506">-</td>
<td id="76A95DD43610B6B2FC9CFEB1FC66FE91" box="[852,897,332,357]" gridcol="3" gridrow="1" pageId="5" pageNumber="506">-</td>
<td id="76A95DD43610B6B2FC1BFEB1FC3CFE91" box="[979,987,332,357]" gridcol="4" gridrow="1" pageId="5" pageNumber="506">-</td>
<td id="76A95DD43610B6B2FB67FEB1FB50FE91" box="[1199,1207,332,357]" gridcol="5" gridrow="1" pageId="5" pageNumber="506">-</td>
<td id="76A95DD43610B6B2FAE6FEB1FAD1FE91" box="[1326,1334,332,357]" gridcol="6" gridrow="1" pageId="5" pageNumber="506">-</td>
</tr>
<tr id="357834A83610B6B2FF6EFE8FFAD1FE7F" box="[166,1334,370,395]" gridrow="2" pageId="5" pageNumber="506">
<th id="76A95DD43610B6B2FF6EFE8FFE62FE7F" box="[166,389,370,395]" gridcol="0" gridrow="2" pageId="5" pageNumber="506">
<taxonomicName id="4C484D6936104948FF6EFE8EFEA9FE7E" authority="Borneo" authorityName="Borneo" box="[166,334,370,394]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="laevis">
<emphasis id="B93CEAF836104948FF6EFE8EFF1AFE7E" box="[166,253,370,394]" italics="true" pageId="5" pageNumber="506">A. laevis</emphasis>
Borneo
</taxonomicName>
</th>
<td id="76A95DD43610B6B2FE0AFE8FFDDCFE7F" box="[450,571,370,395]" gridcol="1" gridrow="2" pageId="5" pageNumber="506">PDSR, Ven,</td>
<td id="76A95DD43610B6B2FDB0FE8FFCCBFE7F" box="[632,812,370,395]" gridcol="2" gridrow="2" pageId="5" pageNumber="506">PDSR, Ven, InfL,</td>
<td id="76A95DD43610B6B2FC9CFE8FFC66FE7F" box="[852,897,370,395]" gridcol="3" gridrow="2" pageId="5" pageNumber="506">-</td>
<td id="76A95DD43610B6B2FC1BFE8FFC3CFE7F" box="[979,987,370,395]" gridcol="4" gridrow="2" pageId="5" pageNumber="506">-</td>
<td id="76A95DD43610B6B2FB67FE8FFB50FE7F" box="[1199,1207,370,395]" gridcol="5" gridrow="2" pageId="5" pageNumber="506">-</td>
<td id="76A95DD43610B6B2FAE6FE8FFAD1FE7F" box="[1326,1334,370,395]" gridcol="6" gridrow="2" pageId="5" pageNumber="506">-</td>
</tr>
<tr id="357834A83610B6B2FF6EFE6BFAD1FE3B" box="[166,1334,406,463]" gridrow="3" pageId="5" pageNumber="506">
<th id="76A95DD43610B6B2FF6EFE6BFE62FE3B" box="[166,389,406,463]" gridcol="0" gridrow="3" pageId="5" pageNumber="506">
<taxonomicName id="4C484D6936104948FF6EFE4AFE9CFE3B" authority="Thai-Malay" authorityName="Thai-Malay" box="[166,379,438,463]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="laevis">
<emphasis id="B93CEAF836104948FF6EFE4AFF1AFE3B" box="[166,253,439,463]" italics="true" pageId="5" pageNumber="506">A. laevis</emphasis>
Thai-Malay
</taxonomicName>
</th>
<td id="76A95DD43610B6B2FE0AFE6BFDDCFE3B" box="[450,571,406,463]" gridcol="1" gridrow="3" pageId="5" pageNumber="506">SubC PDSR, Ven,</td>
<td id="76A95DD43610B6B2FDB0FE6BFCCBFE3B" box="[632,812,406,463]" gridcol="2" gridrow="3" pageId="5" pageNumber="506">InfL-contact PDSR, Ven, InfL-</td>
<td id="76A95DD43610B6B2FC9CFE6BFC66FE3B" box="[852,897,406,463]" gridcol="3" gridrow="3" pageId="5" pageNumber="506">-</td>
<td id="76A95DD43610B6B2FC1BFE6BFC3CFE3B" box="[979,987,406,463]" gridcol="4" gridrow="3" pageId="5" pageNumber="506">-</td>
<td id="76A95DD43610B6B2FB67FE6BFB50FE3B" box="[1199,1207,406,463]" gridcol="5" gridrow="3" pageId="5" pageNumber="506">-</td>
<td id="76A95DD43610B6B2FAE6FE6BFAD1FE3B" box="[1326,1334,406,463]" gridcol="6" gridrow="3" pageId="5" pageNumber="506">-</td>
</tr>
<tr id="357834A83610B6B2FF6EFE27FAD1FE07" box="[166,1334,474,499]" gridrow="4" pageId="5" pageNumber="506" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<th id="76A95DD43610B6B2FF6EFE27FE62FE07" box="[166,389,474,499]" gridcol="0" gridrow="4" pageId="5" pageNumber="506">Peninsula</th>
<td id="76A95DD43610B6B2FE0AFE27FDDCFE07" box="[450,571,474,499]" gridcol="1" gridrow="4" pageId="5" pageNumber="506">SubC</td>
<td id="76A95DD43610B6B2FDB0FE27FCCBFE07" box="[632,812,474,499]" gridcol="2" gridrow="4" pageId="5" pageNumber="506">contact</td>
</tr>
<tr id="357834A83610B6B2FF6EFE06FAD1FDE0" box="[166,1334,507,532]" gridrow="5" pageId="5" pageNumber="506">
<th id="76A95DD43610B6B2FF6EFE06FE62FDE0" box="[166,389,507,532]" gridcol="0" gridrow="5" pageId="5" pageNumber="506">
<taxonomicName id="4C484D6936104948FF6EFE01FF1AFDE0" box="[166,253,508,532]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="laevis">
<emphasis id="B93CEAF836104948FF6EFE01FF1AFDE0" box="[166,253,508,532]" italics="true" pageId="5" pageNumber="506">A. laevis</emphasis>
</taxonomicName>
Java
</th>
<td id="76A95DD43610B6B2FE0AFE06FDDCFDE0" box="[450,571,507,532]" gridcol="1" gridrow="5" pageId="5" pageNumber="506">PDSR, Ven,</td>
<td id="76A95DD43610B6B2FDB0FE06FCCBFDE0" box="[632,812,507,532]" gridcol="2" gridrow="5" pageId="5" pageNumber="506">PDSR, Ven</td>
<td id="76A95DD43610B6B2FC9CFE06FC66FDE0" box="[852,897,507,532]" gridcol="3" gridrow="5" pageId="5" pageNumber="506">InfL</td>
<td id="76A95DD43610B6B2FC1BFE06FC3CFDE0" box="[979,987,507,532]" gridcol="4" gridrow="5" pageId="5" pageNumber="506">-</td>
<td id="76A95DD43610B6B2FB67FE06FB50FDE0" box="[1199,1207,507,532]" gridcol="5" gridrow="5" pageId="5" pageNumber="506">-</td>
<td id="76A95DD43610B6B2FAE6FE06FAD1FDE0" box="[1326,1334,507,532]" gridcol="6" gridrow="5" pageId="5" pageNumber="506">-</td>
</tr>
<tr id="357834A83610B6B2FF6EFDE2FAD1FDCC" box="[166,1334,543,568]" gridrow="6" pageId="5" pageNumber="506" rowspan-0="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<td id="76A95DD43610B6B2FE0AFDE2FDDCFDCC" box="[450,571,543,568]" gridcol="1" gridrow="6" pageId="5" pageNumber="506">SubC</td>
</tr>
<tr id="357834A83610B6B2FF6EFDBDFAD1FDAD" box="[166,1334,576,601]" gridrow="7" pageId="5" pageNumber="506">
<th id="76A95DD43610B6B2FF6EFDBDFE62FDAD" box="[166,389,576,601]" gridcol="0" gridrow="7" pageId="5" pageNumber="506">
<taxonomicName id="4C484D6936104948FF6EFDBCFF1AFDAC" box="[166,253,576,600]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="laevis">
<emphasis id="B93CEAF836104948FF6EFDBCFF1AFDAC" box="[166,253,576,600]" italics="true" pageId="5" pageNumber="506">A. laevis</emphasis>
</taxonomicName>
Sumatra
</th>
<td id="76A95DD43610B6B2FE0AFDBDFDDCFDAD" box="[450,571,576,601]" gridcol="1" gridrow="7" pageId="5" pageNumber="506">PDSR</td>
<td id="76A95DD43610B6B2FDB0FDBDFCCBFDAD" box="[632,812,576,601]" gridcol="2" gridrow="7" pageId="5" pageNumber="506">PDSR</td>
<td id="76A95DD43610B6B2FC9CFDBDFC66FDAD" box="[852,897,576,601]" gridcol="3" gridrow="7" pageId="5" pageNumber="506">InfL</td>
<td id="76A95DD43610B6B2FC1BFDBDFC3CFDAD" box="[979,987,576,601]" gridcol="4" gridrow="7" pageId="5" pageNumber="506">-</td>
<td id="76A95DD43610B6B2FB67FDBDFB50FDAD" box="[1199,1207,576,601]" gridcol="5" gridrow="7" pageId="5" pageNumber="506">-</td>
<td id="76A95DD43610B6B2FAE6FDBDFAD1FDAD" box="[1326,1334,576,601]" gridcol="6" gridrow="7" pageId="5" pageNumber="506">-</td>
</tr>
</table>
</paragraph>
<paragraph id="8BF736EA36104948FF0FFC78FD21F99B" blockId="5.[151,1437,649,2044]" pageId="5" pageNumber="506">
<emphasis id="B93CEAF836104948FF0FFC78FE85FC54" bold="true" box="[199,354,901,928]" pageId="5" pageNumber="506">Comparison.</emphasis>
<taxonomicName id="4C484D6936104948FEA1FC78FD9DFC6B" authority="Quah &amp; Grismer &amp; Lim &amp; Anuar &amp; Imbun, 2019" authorityName="Quah &amp; Grismer &amp; Lim &amp; Anuar &amp; Imbun" authorityYear="2019" box="[361,634,901,927]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="stuebingi" status="sp. nov.">
<emphasis id="B93CEAF836104948FEA1FC78FD9DFC6B" box="[361,634,901,927]" italics="true" pageId="5" pageNumber="506">Asthenodipsas stuebingi</emphasis>
</taxonomicName>
<emphasis id="B93CEAF836104948FD49FC7BFD3CFC54" bold="true" box="[641,731,902,928]" pageId="5" pageNumber="506">
<taxonomicNameLabel id="A20F578336104948FD49FC7BFD3CFC54" box="[641,731,902,928]" pageId="5" pageNumber="506" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be differentiated from
<taxonomicName id="4C484D6936104948FBC6FC78FB22FC6B" baseAuthorityName="Jablonski &amp; Hegner" baseAuthorityYear="2016" box="[1038,1221,901,927]" class="Reptilia" family="Pareatidae" genus="Aplopeltura" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="boa">
<emphasis id="B93CEAF836104948FBC6FC78FB22FC6B" box="[1038,1221,901,927]" italics="true" pageId="5" pageNumber="506">Aplopeltura boa</emphasis>
</taxonomicName>
by its higher number of mid-dorsal scale rows (15 vs. 13) and divided subcaudals (
<bibRefCitation id="EFD94B1B36104948FCBAFC54FBF1FC30" author="de Rooij, N." box="[882,1046,937,964]" pageId="5" pageNumber="506" refId="ref15605" refString="de Rooij, N. (1917) The Reptiles of the Indo-Australian Archipelago. II. Ophidia. E. J. Brill Ltd., Leiden, 334 pp. https: // doi. org / 10.5962 / bhl. title. 10610" type="book" year="1917">de Rooij 1917</bibRefCitation>
;
<bibRefCitation id="EFD94B1B36104948FBEAFC57FAB2FC30" author="Grossmann, W. &amp; Tillack, F." box="[1058,1365,937,964]" pageId="5" pageNumber="506" pagination="175 - 90" refId="ref16457" refString="Grossmann, W. &amp; Tillack, F. (2003) On the Taxonomic Status of Asthenodipsas tropidonotus (Van Lidth De Jeude, 1923) and Pareas vertebralis (Boulenger, 1900) (Serpentes: Colubridae: Pareatinae). Russian Journal of Herpetology, 10 (3), 175 - 90." type="journal article" year="2003">Grossmann &amp; Tillack 2003</bibRefCitation>
;
<bibRefCitation id="EFD94B1B36104948FAA9FC57FEADFC1C" author="Stuebing, R. B. &amp; Inger, R. F. &amp; Lardner, B." pageId="5" pageNumber="506" refId="ref17497" refString="Stuebing, R. B., Inger, R. F. &amp; Lardner, B. (2014) A field guide to the snakes of Borneo (2 nd edition). Natural History Publications (Borneo), Kota Kinabalu, 310 pp." type="book" year="2014">
Stuebing
<emphasis id="B93CEAF836104948FF07FC33FEE7FC13" box="[207,256,973,999]" italics="true" pageId="5" pageNumber="506">et al</emphasis>
. 2014
</bibRefCitation>
).
<taxonomicName id="4C484D6936104948FE94FC30FD8CFC13" authority="Quah &amp; Grismer &amp; Lim &amp; Anuar &amp; Imbun, 2019" authorityName="Quah &amp; Grismer &amp; Lim &amp; Anuar &amp; Imbun" authorityYear="2019" box="[348,619,973,999]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="stuebingi" status="sp. nov.">
<emphasis id="B93CEAF836104948FE94FC30FD8CFC13" box="[348,619,973,999]" italics="true" pageId="5" pageNumber="506">Asthenodipsas stuebingi</emphasis>
</taxonomicName>
<emphasis id="B93CEAF836104948FDB9FC33FD2DFC1C" bold="true" box="[625,714,974,1000]" pageId="5" pageNumber="506">
<taxonomicNameLabel id="A20F578336104948FDB9FC33FD2DFC1C" box="[625,714,974,1000]" pageId="5" pageNumber="506" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be differentiated from members of the genus
<taxonomicName id="4C484D6936104948FB3AFC33FAA7FC13" box="[1266,1344,974,999]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="506" phylum="Chordata" rank="genus">
<emphasis id="B93CEAF836104948FB3AFC33FAA7FC13" box="[1266,1344,974,999]" italics="true" pageId="5" pageNumber="506">Pareas</emphasis>
</taxonomicName>
by their possession of preocular and subocular scales (absent vs. present), supralabials in contact with orbit (3
<superScript id="7C3D9BA236104948FB33FC0DFAEEFC0A" attach="left" box="[1275,1289,1008,1022]" fontSize="6" pageId="5" pageNumber="506">rd</superScript>
&amp; 4
<superScript id="7C3D9BA236104948FAF1FC0DFAA1FC0A" attach="left" box="[1337,1350,1008,1022]" fontSize="6" pageId="5" pageNumber="506">th</superScript>
contact orbit vs. no supralabials in contact with orbit) and anterior single inframaxillary (present vs. absent) (
<bibRefCitation id="EFD94B1B36104948FB36FBEBFECDFBA0" author="Grossmann, W. &amp; Tillack, F." pageId="5" pageNumber="506" pagination="175 - 90" refId="ref16457" refString="Grossmann, W. &amp; Tillack, F. (2003) On the Taxonomic Status of Asthenodipsas tropidonotus (Van Lidth De Jeude, 1923) and Pareas vertebralis (Boulenger, 1900) (Serpentes: Colubridae: Pareatinae). Russian Journal of Herpetology, 10 (3), 175 - 90." type="journal article" year="2003">Grossmann &amp; Tillack 2003</bibRefCitation>
).
<taxonomicName id="4C484D6936104948FEF6FBC4FD98FBA7" box="[318,639,1081,1107]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="lasgalenensis">
<emphasis id="B93CEAF836104948FEF6FBC4FD98FBA7" box="[318,639,1081,1107]" italics="true" pageId="5" pageNumber="506">Asthenodipsas lasgalenensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C484D6936104948FD45FBC7FCDBFBA7" baseAuthorityName="Lidth De Jeude" baseAuthorityYear="1923" box="[653,828,1081,1108]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="tropidonotus">
<emphasis id="B93CEAF836104948FD45FBC7FCDBFBA7" box="[653,828,1081,1108]" italics="true" pageId="5" pageNumber="506">A. tropidonotus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C484D6936104948FCBCFBC7FBEAFBA7" authorityName="Boulenger" authorityYear="1900" box="[884,1037,1081,1108]" class="Squamata" family="Pareidae" genus="Amblycephalus" higherTaxonomySource="CoL" kingdom="Animalia" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="vertebralis">
<emphasis id="B93CEAF836104948FCBCFBC7FBEAFBA7" box="[884,1037,1081,1108]" italics="true" pageId="5" pageNumber="506">A. vertebralis</emphasis>
</taxonomicName>
can be differentiated from the new species by their higher number subcaudals (5477 vs. 3547), pairs of infralabials in contact (1
<superScript id="7C3D9BA236104948FB03FBA0FB31FB9F" attach="left" box="[1227,1238,1117,1131]" fontSize="6" pageId="5" pageNumber="506">st</superScript>
vs. 3
<superScript id="7C3D9BA236104948FADCFBA1FAC5FB9E" attach="none" box="[1300,1314,1116,1130]" fontSize="6" pageId="5" pageNumber="506">rd</superScript>
) and more pairs of posterior inframaxillaries (three vs. two). In addition,
<taxonomicName id="4C484D6936104948FC9CFB7FFC0AFB6F" authorityName="Boulenger" authorityYear="1900" box="[852,1005,1153,1180]" class="Squamata" family="Pareidae" genus="Amblycephalus" higherTaxonomySource="CoL" kingdom="Animalia" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="vertebralis">
<emphasis id="B93CEAF836104948FC9CFB7FFC0AFB6F" box="[852,1005,1153,1180]" italics="true" pageId="5" pageNumber="506">A. vertebralis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C484D6936104948FBEDFB7FFB34FB6F" baseAuthorityName="Lidth De Jeude" baseAuthorityYear="1923" box="[1061,1235,1153,1180]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="tropidonotus">
<emphasis id="B93CEAF836104948FBEDFB7FFB34FB6F" box="[1061,1235,1153,1180]" italics="true" pageId="5" pageNumber="506">A. tropidonotus</emphasis>
</taxonomicName>
can be differentiated from
<taxonomicName id="4C484D6936104948FEC7FB5BFE7FFB4B" authority="Quah &amp; Grismer &amp; Lim &amp; Anuar &amp; Imbun, 2019" authorityName="Quah &amp; Grismer &amp; Lim &amp; Anuar &amp; Imbun" authorityYear="2019" box="[271,408,1189,1216]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="stuebingi" status="sp. nov.">
<emphasis id="B93CEAF836104948FEC7FB5BFE7FFB4B" box="[271,408,1189,1216]" italics="true" pageId="5" pageNumber="506">A. stuebingi</emphasis>
</taxonomicName>
<emphasis id="B93CEAF836104948FE69FB5BFE1AFB34" bold="true" box="[417,509,1190,1216]" pageId="5" pageNumber="506">
<taxonomicNameLabel id="A20F578336104948FE69FB5BFE1AFB34" box="[417,509,1190,1216]" pageId="5" pageNumber="506" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by their higher number of ventrals (195215 vs. 165175). Adult
<taxonomicName id="4C484D6936104948FB36FB5BFA7BFB4B" box="[1278,1436,1189,1216]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="lasgalensis">
<emphasis id="B93CEAF836104948FB36FB5BFA7BFB4B" box="[1278,1436,1189,1216]" italics="true" pageId="5" pageNumber="506">A. lasgalensis</emphasis>
</taxonomicName>
can also be differentiated from
<taxonomicName id="4C484D6936104948FE3EFB37FD9AFB17" authority="Quah &amp; Grismer &amp; Lim &amp; Anuar &amp; Imbun, 2019" authorityName="Quah &amp; Grismer &amp; Lim &amp; Anuar &amp; Imbun" authorityYear="2019" box="[502,637,1225,1252]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="stuebingi" status="sp. nov.">
<emphasis id="B93CEAF836104948FE3EFB37FD9AFB17" box="[502,637,1225,1252]" italics="true" pageId="5" pageNumber="506">A. stuebingi</emphasis>
</taxonomicName>
<emphasis id="B93CEAF836104948FD4CFB37FD39FB10" bold="true" box="[644,734,1226,1252]" pageId="5" pageNumber="506">
<taxonomicNameLabel id="A20F578336104948FD4CFB37FD39FB10" box="[644,734,1226,1252]" pageId="5" pageNumber="506" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by their dorsal colour pattern that is solid dark brown to black and their whitish labials versus a beige to brown dorsum with dark-bands and a dark patch on the neck in the latter (
<bibRefCitation id="EFD94B1B36104948FF57FAECFE9FFAD8" author="Loredo, A. I. &amp; Wood, Jr. P. L. &amp; Quah, E. S. H. &amp; Shahrul, A. &amp; Greer, L. F. &amp; Ahmad, N. &amp; Grismer, L. L." box="[159,376,1297,1324]" pageId="5" pageNumber="506" pagination="505 - 524" refId="ref16789" refString="Loredo, A. I., Wood, Jr. P. L., Quah, E. S. H., Shahrul, A., Greer, L. F., Ahmad, N. &amp; Grismer, L. L. (2013) Cryptic speciation within Asthenodipsas vertebralis (Boulenger, 1900) (Squamata: Pareatidae), the description of a new species from Peninsular Malaysia, and the resurrection of A. tropidonotus (Lidth de Jude, 1923) from Sumatra: an integrative taxonomic analysis. Zootaxa, 3664 (4), 505 - 524. https: // doi. org / 10.11646 / zootaxa. 3664.4.5" type="journal article" year="2013">
Loredo
<emphasis id="B93CEAF836104948FF32FAEFFECBFADF" box="[250,300,1297,1323]" italics="true" pageId="5" pageNumber="506">et al</emphasis>
. 2013
</bibRefCitation>
). From Bornean populations of
<taxonomicName id="4C484D6936104948FD2BFAEFFC99FADF" authorityName="Peters" authorityYear="1864" box="[739,894,1297,1324]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="malaccana">
<emphasis id="B93CEAF836104948FD2BFAEFFC99FADF" box="[739,894,1297,1324]" italics="true" pageId="5" pageNumber="506">A. malaccana</emphasis>
</taxonomicName>
to which it is most similar in colour pattern,
<taxonomicName id="4C484D6936104948FA4CFAEFFF19FABB" authority="Quah &amp; Grismer &amp; Lim &amp; Anuar &amp; Imbun, 2019" authorityName="Quah &amp; Grismer &amp; Lim &amp; Anuar &amp; Imbun" authorityYear="2019" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="stuebingi" status="sp. nov.">
<emphasis id="B93CEAF836104948FA4CFAEFFF19FABB" italics="true" pageId="5" pageNumber="506">A. stuebingi</emphasis>
</taxonomicName>
<emphasis id="B93CEAF836104948FECEFACBFE86FAA4" bold="true" box="[262,353,1334,1360]" pageId="5" pageNumber="506">
<taxonomicNameLabel id="A20F578336104948FECEFACBFE86FAA4" box="[262,353,1334,1360]" pageId="5" pageNumber="506" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be differentiated by a lower number of supralabials (6 vs. 78) (Chan-ard
<emphasis id="B93CEAF836104948FB2EFACBFAC7FABB" box="[1254,1312,1333,1359]" italics="true" pageId="5" pageNumber="506">et al.</emphasis>
2015;
<bibRefCitation id="EFD94B1B36104948FAB9FACBFF34FA80" author="Das, I." pageId="5" pageNumber="506" refId="ref15544" refString="Das, I. (2010) A Field Guide to the Reptiles of South-East Asia. New Holland Publishers, London, 376 pp." type="book" year="2010">Das 2010</bibRefCitation>
;
<bibRefCitation id="EFD94B1B36104948FF17FAA4FE63FA80" author="de Rooij, N." box="[223,388,1369,1396]" pageId="5" pageNumber="506" refId="ref15605" refString="de Rooij, N. (1917) The Reptiles of the Indo-Australian Archipelago. II. Ophidia. E. J. Brill Ltd., Leiden, 334 pp. https: // doi. org / 10.5962 / bhl. title. 10610" type="book" year="1917">de Rooij 1917</bibRefCitation>
;
<bibRefCitation id="EFD94B1B36104948FE59FAA4FD9AFA80" author="Stuebing, R. B. &amp; Inger, R. F. &amp; Lardner, B." box="[401,637,1369,1396]" pageId="5" pageNumber="506" refId="ref17497" refString="Stuebing, R. B., Inger, R. F. &amp; Lardner, B. (2014) A field guide to the snakes of Borneo (2 nd edition). Natural History Publications (Borneo), Kota Kinabalu, 310 pp." type="book" year="2014">
Stuebing
<emphasis id="B93CEAF836104948FE36FAA7FDD0FA87" box="[510,567,1369,1395]" italics="true" pageId="5" pageNumber="506">et al.</emphasis>
2014
</bibRefCitation>
). From
<taxonomicName id="4C484D6936104948FD1EFAA7FCD0FA87" box="[726,823,1369,1396]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="laevis">
<emphasis id="B93CEAF836104948FD1EFAA7FCD0FA87" box="[726,823,1369,1396]" italics="true" pageId="5" pageNumber="506">A. laevis</emphasis>
</taxonomicName>
to which it was previously confused,
<taxonomicName id="4C484D6936104948FB23FAA7FA94FA87" authority="Quah &amp; Grismer &amp; Lim &amp; Anuar &amp; Imbun, 2019" authorityName="Quah &amp; Grismer &amp; Lim &amp; Anuar &amp; Imbun" authorityYear="2019" box="[1259,1395,1369,1396]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="stuebingi" status="sp. nov.">
<emphasis id="B93CEAF836104948FB23FAA7FA94FA87" box="[1259,1395,1369,1396]" italics="true" pageId="5" pageNumber="506">A. stuebingi</emphasis>
</taxonomicName>
<emphasis id="B93CEAF836104948FAB3FAA7FF2FFA6C" bold="true" pageId="5" pageNumber="506">
<taxonomicNameLabel id="A20F578336104948FAB3FAA7FF2FFA6C" pageId="5" pageNumber="506" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be differentiated by its larger adult length, (Max SVL
<quantity id="4CB09B0F36104948FCAEFA80FC5BFA6C" box="[870,956,1405,1432]" metricMagnitude="-1" metricUnit="m" metricValue="5.57" pageId="5" pageNumber="506" unit="mm" value="557.0">557mm</quantity>
vs.
<quantity id="4CB09B0F36104948FC23FA83FBA2FA6C" box="[1003,1093,1406,1432]" metricMagnitude="-1" metricUnit="m" metricValue="3.7299999999999995" pageId="5" pageNumber="506" unit="mm" value="373.0">373mm</quantity>
), dorsal scales rows (15/15/15 vs. 15/15/13) and sharp vertebral keel (present vs. absent) (
<figureCitation id="13732A6F36104948FCE7FA5CFC6BFA48" box="[815,908,1441,1468]" captionStart="FIGURE 3" captionStartId="7.[151,250,1588,1613]" captionTargetBox="[151,1436,211,1555]" captionTargetId="figure@7.[151,1436,211,1555]" captionTargetPageId="7" captionText="FIGURE 3. A. Dorsum of paratype of Asthenodipsas stuebingi sp. nov. (ZMB 65429). B. Venter of paratype of A. stuebingi sp. nov. (ZMB 65429) (Photographs by Frank Tillack). C. Dorsum of paratype of A. stuebingi sp. nov. (ZMB 65429) in life from Kinabalu Park power station, Kamborangoh Road, Sabah. D. Paratype of A. stuebingi sp. nov. (ZMB 65429) from Kinabalu Park power station, Kamborangoh Road, Sabah in defensive coils. E. Venter of paratype of A. stuebingi sp. nov. (ZMB 65429) in life from Kinabalu Park power station, Kamborangoh Road, Sabah. (Photographs by Andreas Nӧllert). F. Sharp vertebral keel of the holotype of A. stuebingi sp. nov. (SP 04679) as indicated by arrow." httpUri="https://zenodo.org/record/3351078/files/figure.png" pageId="5" pageNumber="506">Figs. 3F</figureCitation>
&amp;
<figureCitation id="13732A6F36104948FC78FA5FFC37FA48" box="[944,976,1442,1468]" captionStart="FIGURE 4" captionStartId="11.[151,250,1896,1921]" captionTargetBox="[181,1406,166,1876]" captionTargetId="figure@11.[181,1406,166,1876]" captionTargetPageId="11" captionText="FIGURE 4. A &amp; B. Dorsum and venter of holotype of Asthenodipsas jamilinaisi sp. nov. (SP 04076). C &amp; D. Dorsum and venter of paratype of A. jamilinaisi sp. nov. (ZRC 2.2742). Photographs by Kelvin Lim. E. Sharp vertebral keel of the holotype of A. jamilinaisi sp. nov. (SP 04679) as indicated by arrow. F. Absence of sharp vertebral keel on A. laevis (SP 04138) from Borneo as indicated by arrow." httpUri="https://zenodo.org/record/3351080/files/figure.png" pageId="5" pageNumber="506">4F</figureCitation>
) (
<tableCitation id="C6CA035136104948FC2EFA5CFB82FA48" box="[998,1125,1441,1468]" captionStart-0="TABLE 2" captionStart-1="TABLE 3" captionStart-2="TABLE 4" captionStart-3="TABLE 5" captionStart-4="TABLE 6" captionStartId-0="4.[151,239,885,909]" captionStartId-1="5.[151,239,154,178]" captionStartId-2="9.[151,239,461,487]" captionStartId-3="15.[151,239,464,490]" captionStartId-4="22.[151,239,701,727]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="9" captionTargetPageId-3="15" captionTargetPageId-4="22" captionText-0="TABLE 2. Summary statistics for Asthenodipsas stuebingi sp. nov. and the different populations of A. laevis, SD = standard deviation and N = sample size." captionText-1="TABLE 3. Pairwise matrix of statistically significant mean differences of characters among species of the Asthenodipsas" captionText-2="TABLE 4. Scale counts and selected measurements (mm) of the type series of Asthenodipsas jamilinaisi sp. nov. and A. stuebingi sp. nov. Abbreviations are listed in the Materials and methods." captionText-3="TABLE 5. Measurements and scale counts of Asthenodipsas jamilinaisi sp. nov., A. stuebingi sp. nov. and different populations of A. laevis. (*measurements from Grossmann &amp; Tillack 2003; NA = measurement not available). Abbreviations are listed in the Materials and methods." captionText-4="TABLE 6. Comparison of selected characters between Asthenodipsas jamilinaisi sp. nov., A. stuebingi sp. nov. and different populations of A. laevis with diagnostic characters in bold. Abbreviations are listed in the Materials and Methods. ND = No data." httpUri-0="http://table.plazi.org/id/DF37666236114949FF5FFC88FDF0FC45" httpUri-1="http://table.plazi.org/id/DF37666236104948FF5FFF67FA7BFF46" httpUri-2="http://table.plazi.org/id/DF376662361C4944FF5FFE30FEADFDFF" httpUri-3="http://table.plazi.org/id/DF376662361A4942FF5FFE2DFB05FDFA" httpUri-4="http://table.plazi.org/id/DF3766623603495BFF5FFD40FC30FD0F" pageId="5" pageNumber="506" tableUuid-0="DF37666236114949FF5FFC88FDF0FC45" tableUuid-1="DF37666236104948FF5FFF67FA7BFF46" tableUuid-2="DF376662361C4944FF5FFE30FEADFDFF" tableUuid-3="DF376662361A4942FF5FFE2DFB05FDFA" tableUuid-4="DF3766623603495BFF5FFD40FC30FD0F">Tables 26</tableCitation>
).
<taxonomicName id="4C484D6936104948FBB1FA5FFAE9FA4F" authority="Quah &amp; Grismer &amp; Lim &amp; Anuar &amp; Imbun, 2019" authorityName="Quah &amp; Grismer &amp; Lim &amp; Anuar &amp; Imbun" authorityYear="2019" box="[1145,1294,1441,1468]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="stuenbingi" status="sp. nov.">
<emphasis id="B93CEAF836104948FBB1FA5FFAE9FA4F" box="[1145,1294,1441,1468]" italics="true" pageId="5" pageNumber="506">A. stuenbingi</emphasis>
</taxonomicName>
<emphasis id="B93CEAF836104948FADCFA5FFA89FA48" bold="true" box="[1300,1390,1442,1468]" pageId="5" pageNumber="506">
<taxonomicNameLabel id="A20F578336104948FADCFA5FFA89FA48" box="[1300,1390,1442,1468]" pageId="5" pageNumber="506" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be differentiated from
<taxonomicName id="4C484D6936104948FE58FA3BFDC1FA2B" authority="Quah &amp; Grismer &amp; Lim &amp; Anuar &amp; Imbun, 2019" authorityName="Quah &amp; Grismer &amp; Lim &amp; Anuar &amp; Imbun" authorityYear="2019" box="[400,550,1477,1504]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="jamilinaisi" status="sp. nov.">
<emphasis id="B93CEAF836104948FE58FA3BFDC1FA2B" box="[400,550,1477,1504]" italics="true" pageId="5" pageNumber="506">A. jamilinaisi</emphasis>
</taxonomicName>
<emphasis id="B93CEAF836104948FDE4FA3BFD62FA14" bold="true" box="[556,645,1478,1504]" pageId="5" pageNumber="506">
<taxonomicNameLabel id="A20F578336104948FDE4FA3BFD62FA14" box="[556,645,1478,1504]" pageId="5" pageNumber="506" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by its lower number of subcaudals (3547 vs. 5253), lower number of ventrals in males (165 vs. 173175), size of vertebral scales (slightly enlarged vs. greatly enlarged), colour pattern (light-coloured head and dorsum with a dark neck patch and distinct bands vs. dark overall colouration of dorsum with muted banding) and body form (robust and stout vs. gracile and laterally compressed) (
<tableCitation id="C6CA035136104948FB61F9CCFACEF9B8" box="[1193,1321,1585,1612]" captionStart-0="TABLE 4" captionStart-1="TABLE 5" captionStart-2="TABLE 6" captionStartId-0="9.[151,239,461,487]" captionStartId-1="15.[151,239,464,490]" captionStartId-2="22.[151,239,701,727]" captionTargetPageId-0="9" captionTargetPageId-1="15" captionTargetPageId-2="22" captionText-0="TABLE 4. Scale counts and selected measurements (mm) of the type series of Asthenodipsas jamilinaisi sp. nov. and A. stuebingi sp. nov. Abbreviations are listed in the Materials and methods." captionText-1="TABLE 5. Measurements and scale counts of Asthenodipsas jamilinaisi sp. nov., A. stuebingi sp. nov. and different populations of A. laevis. (*measurements from Grossmann &amp; Tillack 2003; NA = measurement not available). Abbreviations are listed in the Materials and methods." captionText-2="TABLE 6. Comparison of selected characters between Asthenodipsas jamilinaisi sp. nov., A. stuebingi sp. nov. and different populations of A. laevis with diagnostic characters in bold. Abbreviations are listed in the Materials and Methods. ND = No data." httpUri-0="http://table.plazi.org/id/DF376662361C4944FF5FFE30FEADFDFF" httpUri-1="http://table.plazi.org/id/DF376662361A4942FF5FFE2DFB05FDFA" httpUri-2="http://table.plazi.org/id/DF3766623603495BFF5FFD40FC30FD0F" pageId="5" pageNumber="506" tableUuid-0="DF376662361C4944FF5FFE30FEADFDFF" tableUuid-1="DF376662361A4942FF5FFE2DFB05FDFA" tableUuid-2="DF3766623603495BFF5FFD40FC30FD0F">Tables 46</tableCitation>
). A key to the family
<taxonomicName id="4C484D6936104948FED9F9A8FE96F99B" authorityName="Romer" authorityYear="1956" box="[273,369,1621,1647]" class="Reptilia" family="Pareidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="506" phylum="Chordata" rank="family">Pareidae</taxonomicName>
of Borneo is presented below.
</paragraph>
<paragraph id="8BF736EA36104948FF0FF984FB70F928" blockId="5.[151,1437,649,2044]" pageId="5" pageNumber="506">
<emphasis id="B93CEAF836104948FF0FF984FE59F960" bold="true" box="[199,446,1657,1684]" pageId="5" pageNumber="506">
Distribution (
<figureCitation id="13732A6F36104948FEA1F984FE48F960" box="[361,431,1657,1684]" captionStart="FIGURE 8" captionStartId="21.[151,250,1906,1931]" captionTargetBox="[151,1436,466,1861]" captionTargetId="figure@21.[151,1436,466,1861]" captionTargetPageId="21" captionText="FIGURE 8. Distribution of Asthenodipsas jamilinaisi sp. nov. (yellow), A. stuebingi sp. nov. (red) and reported localities of A. laevis (blue) in Borneo. Star = type locality." httpUri="https://zenodo.org/record/3351088/files/figure.png" pageId="5" pageNumber="506">Fig. 8</figureCitation>
).
</emphasis>
<taxonomicName id="4C484D6936104948FE0EF984FD31F967" authority="Quah &amp; Grismer &amp; Lim &amp; Anuar &amp; Imbun, 2019" authorityName="Quah &amp; Grismer &amp; Lim &amp; Anuar &amp; Imbun" authorityYear="2019" box="[454,726,1657,1683]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="stuebingi" status="sp. nov.">
<emphasis id="B93CEAF836104948FE0EF984FD31F967" box="[454,726,1657,1683]" italics="true" pageId="5" pageNumber="506">Asthenodipsas stuebingi</emphasis>
</taxonomicName>
<emphasis id="B93CEAF836104948FD16F987FCDFF960" bold="true" box="[734,824,1658,1684]" pageId="5" pageNumber="506">
<taxonomicNameLabel id="A20F578336104948FD16F987FCDFF960" box="[734,824,1658,1684]" pageId="5" pageNumber="506" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is only known from the highlands of
<collectingRegion id="498CF80836104948FB2BF984FACDF967" box="[1251,1322,1657,1683]" country="Malaysia" name="Sabah" pageId="5" pageNumber="506">Sabah</collectingRegion>
, East
<collectingCountry id="F35F767A36104948FAA5F987FF3FF943" name="Malaysia" pageId="5" pageNumber="506">Malaysia</collectingCountry>
at Mount Kinabalu, Kundasang and the Crocker Range. It is likely to range further throughout the island and found in similar upland habitats in
<collectingCountry id="F35F767A36104948FDD7F93CFD65F92F" box="[543,642,1729,1755]" name="Malaysia" pageId="5" pageNumber="506">Sarawak</collectingCountry>
,
<collectingCountry id="F35F767A36104948FD46F93CFD12F92F" box="[654,757,1729,1755]" name="Malaysia" pageId="5" pageNumber="506">Malaysia</collectingCountry>
and Kalimantan, Indonesian Borneo.
</paragraph>
<paragraph id="8BF736EA36104948FF0FF91BFC50F8D7" blockId="5.[151,1437,649,2044]" pageId="5" pageNumber="506">
<emphasis id="B93CEAF836104948FF0FF91BFEA9F8F4" bold="true" box="[199,334,1766,1792]" pageId="5" pageNumber="506">Etymology.</emphasis>
The specific epithet
<taxonomicName id="4C484D6936104948FDF1F918FD47F90B" authorityName="Quah &amp; Grismer &amp; Lim &amp; Anuar &amp; Imbun" authorityYear="2019" box="[569,672,1765,1791]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="stuebingi">
<emphasis id="B93CEAF836104948FDF1F918FD47F90B" box="[569,672,1765,1791]" italics="true" pageId="5" pageNumber="506">stuebingi</emphasis>
</taxonomicName>
is a patronym in honour of Robert B. Stuebing for his contributions to the field of herpetology in Borneo, especially on the study of snakes.
</paragraph>
<paragraph id="8BF736EA3610494BFF0FF8D3FBBAFE93" blockId="5.[151,1437,649,2044]" lastBlockId="6.[151,1437,152,359]" lastPageId="6" lastPageNumber="507" pageId="5" pageNumber="506">
<emphasis id="B93CEAF836104948FF0FF8D3FE62F8BC" bold="true" box="[199,389,1837,1864]" pageId="5" pageNumber="506">Natural History</emphasis>
.
<taxonomicName id="4C484D6936104948FE5CF8D0FD44F8B3" authority="Quah &amp; Grismer &amp; Lim &amp; Anuar &amp; Imbun, 2019" authorityName="Quah &amp; Grismer &amp; Lim &amp; Anuar &amp; Imbun" authorityYear="2019" box="[404,675,1837,1863]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="stuebingi" status="sp. nov.">
<emphasis id="B93CEAF836104948FE5CF8D0FD44F8B3" box="[404,675,1837,1863]" italics="true" pageId="5" pageNumber="506">Asthenodipsas stuebingi</emphasis>
</taxonomicName>
<emphasis id="B93CEAF836104948FD62F8D3FCE3F8BC" bold="true" box="[682,772,1838,1864]" pageId="5" pageNumber="506">
<taxonomicNameLabel id="A20F578336104948FD62F8D3FCE3F8BC" box="[682,772,1838,1864]" pageId="5" pageNumber="506" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is a montane species that so far has only been found on the highlands of
<collectingRegion id="498CF80836104948FEE1F8ACFE8AF89F" box="[297,365,1873,1899]" country="Malaysia" name="Sabah" pageId="5" pageNumber="506">Sabah</collectingRegion>
from
<quantity id="4CB09B0F36104948FE7AF8AFFDD8F898" box="[434,575,1874,1900]" metricMagnitude="3" metricUnit="m" metricValue="1.45" metricValueMax="2.0" metricValueMin="0.9" pageId="5" pageNumber="506" unit="m" value="1450.0" valueMax="2000.0" valueMin="900.0">9002000 m</quantity>
a.s.l.. This is in contrast to its congener
<taxonomicName id="4C484D6936104948FBCAF8AFFB85F89F" box="[1026,1122,1873,1900]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="laevis">
<emphasis id="B93CEAF836104948FBCAF8AFFB85F89F" box="[1026,1122,1873,1900]" italics="true" pageId="5" pageNumber="506">A. laevis</emphasis>
</taxonomicName>
that is widely distributed in Borneo and recorded from many lowland localities up to
<quantity id="4CB09B0F36104948FCD1F888FC8AF864" box="[793,877,1909,1936]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="5" pageNumber="506" unit="m" value="1150.0">1150 m</quantity>
in elevation (
<bibRefCitation id="EFD94B1B36104948FBCEF888FB09F864" author="Stuebing, R. B. &amp; Inger, R. F. &amp; Lardner, B." box="[1030,1262,1909,1936]" pageId="5" pageNumber="506" refId="ref17497" refString="Stuebing, R. B., Inger, R. F. &amp; Lardner, B. (2014) A field guide to the snakes of Borneo (2 nd edition). Natural History Publications (Borneo), Kota Kinabalu, 310 pp." type="book" year="2014">
Stuebing
<emphasis id="B93CEAF836104948FBBAF88BFB44F87B" box="[1138,1187,1909,1935]" italics="true" pageId="5" pageNumber="506">et al</emphasis>
. 2014
</bibRefCitation>
). At some locations such as the Crocker Range, both species occur (SP 04476, SP 04570 &amp; SP 04679; see
<tableCitation id="C6CA035136104948FB4EF864FB3DF840" box="[1158,1242,1945,1972]" captionStart="TABLE 5" captionStartId="15.[151,239,464,490]" captionTargetPageId="15" captionText="TABLE 5. Measurements and scale counts of Asthenodipsas jamilinaisi sp. nov., A. stuebingi sp. nov. and different populations of A. laevis. (*measurements from Grossmann &amp; Tillack 2003; NA = measurement not available). Abbreviations are listed in the Materials and methods." httpUri="http://table.plazi.org/id/DF376662361A4942FF5FFE2DFB05FDFA" pageId="5" pageNumber="506" tableUuid="DF376662361A4942FF5FFE2DFB05FDFA">Table 5</tableCitation>
). Similar to other species of slug snakes,
<taxonomicName id="4C484D6936104948FE56F843FDC2F823" authority="Quah &amp; Grismer &amp; Lim &amp; Anuar &amp; Imbun, 2019" authorityName="Quah &amp; Grismer &amp; Lim &amp; Anuar &amp; Imbun" authorityYear="2019" box="[414,549,1981,2008]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="stuebingi" status="sp. nov.">
<emphasis id="B93CEAF836104948FE56F843FDC2F823" box="[414,549,1981,2008]" italics="true" pageId="5" pageNumber="506">A. stuebingi</emphasis>
</taxonomicName>
<emphasis id="B93CEAF836104948FDE4F843FD61F82C" bold="true" box="[556,646,1982,2008]" pageId="5" pageNumber="506">
<taxonomicNameLabel id="A20F578336104948FDE4F843FD61F82C" box="[556,646,1982,2008]" pageId="5" pageNumber="506" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is nocturnal. It is reportedly both terrestrial and arboreal (
<bibRefCitation id="EFD94B1B36104948FADDF840FF10F808" author="Malkmus, R. &amp; Manthey, U. &amp; Vogel, G. &amp; Hoffmann, P. &amp; Kosuch, J." pageId="5" pageNumber="506" refId="ref16983" refString="Malkmus, R., Manthey, U., Vogel, G., Hoffmann, P. &amp; Kosuch, J. (2002) Amphibians and Reptiles of Mount Kinabalu (North- Borneo). A. R. G. Ganter Verlag, Ruggel, 424 pp." type="book" year="2002">
Malkmus
<emphasis id="B93CEAF836104948FA40F843FF4AF80F" italics="true" pageId="5" pageNumber="506">et al</emphasis>
. 2002
</bibRefCitation>
).
<bibRefCitation id="EFD94B1B36104948FEC0F81CFDE7F808" author="Malkmus, R. &amp; Manthey, U. &amp; Vogel, G. &amp; Hoffmann, P. &amp; Kosuch, J." box="[264,512,2017,2044]" pageId="5" pageNumber="506" refId="ref16983" refString="Malkmus, R., Manthey, U., Vogel, G., Hoffmann, P. &amp; Kosuch, J. (2002) Amphibians and Reptiles of Mount Kinabalu (North- Borneo). A. R. G. Ganter Verlag, Ruggel, 424 pp." type="book" year="2002">
Malkmus
<emphasis id="B93CEAF836104948FEB0F81FFE4FF80F" box="[376,424,2017,2043]" italics="true" pageId="5" pageNumber="506">et al</emphasis>
. (2002)
</bibRefCitation>
reported finding a specimen crossing the road at night in rainy weather and another specimen in a bush approximately
<quantity id="4CB09B0F3613494BFDD6FF64FDA6FF47" box="[542,577,153,179]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="507" unit="m" value="1.0">1m</quantity>
above the ground. The
<typeStatus id="54F388483613494BFC84FF64FC4BFF47" box="[844,940,153,179]" pageId="6" pageNumber="507" type="paratype">paratype</typeStatus>
<collectionCode id="ED59AE2F3613494BFC7BFF64FC17FF47" box="[947,1008,153,179]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="6" pageNumber="507" type="Museum">ZMB</collectionCode>
65429 was also collected at night fol- lowing light rains at approximately 22:00 h. A specimen was observed at 09:00 h close the power station of the Kinabalu park headquarters along Kamborangoh road (Andreas N ӧllert
<emphasis id="B93CEAF83613494BFC08FF1FFC1AFF0F" box="[960,1021,225,251]" italics="true" pageId="6" pageNumber="507">in litt</emphasis>
. 2018). When threatened, the snakes roll themselves up into tight coils and remain motionless (
<figureCitation id="13732A6F3613494BFCD3FEF8FC42FEEB" box="[795,933,261,287]" captionStart="FIGURE 3" captionStartId="7.[151,250,1588,1613]" captionTargetBox="[151,1436,211,1555]" captionTargetId="figure@7.[151,1436,211,1555]" captionTargetPageId="7" captionText="FIGURE 3. A. Dorsum of paratype of Asthenodipsas stuebingi sp. nov. (ZMB 65429). B. Venter of paratype of A. stuebingi sp. nov. (ZMB 65429) (Photographs by Frank Tillack). C. Dorsum of paratype of A. stuebingi sp. nov. (ZMB 65429) in life from Kinabalu Park power station, Kamborangoh Road, Sabah. D. Paratype of A. stuebingi sp. nov. (ZMB 65429) from Kinabalu Park power station, Kamborangoh Road, Sabah in defensive coils. E. Venter of paratype of A. stuebingi sp. nov. (ZMB 65429) in life from Kinabalu Park power station, Kamborangoh Road, Sabah. (Photographs by Andreas Nӧllert). F. Sharp vertebral keel of the holotype of A. stuebingi sp. nov. (SP 04679) as indicated by arrow." httpUri="https://zenodo.org/record/3351078/files/figure.png" pageId="6" pageNumber="507">Fig. 3D &amp; E</figureCitation>
) — a behaviour observed in
<taxonomicName id="4C484D693613494BFB2EFEF8FE29FEB0" authority="(Jablonski &amp; Hegner 2016)" authorityName="Jablonski &amp; Hegner" authorityYear="2016" baseAuthorityName="Jablonski &amp; Hegner" baseAuthorityYear="2016" class="Reptilia" family="Pareatidae" genus="Aplopeltura" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="507" phylum="Chordata" rank="species" species="boa">
<emphasis id="B93CEAF83613494BFB2EFEF8FA7BFEEB" box="[1254,1436,261,287]" italics="true" pageId="6" pageNumber="507">Aplopeltura boa</emphasis>
(
<bibRefCitation id="EFD94B1B3613494BFF57FED4FE21FEB0" author="Jablonski, D. &amp; Hegner, D." box="[159,454,297,324]" pageId="6" pageNumber="507" pagination="31 - 32" refId="ref16662" refString="Jablonski, D. &amp; Hegner, D. (2016) Death feigning behaviour in Aplopeltura boa. The Herpetological Bulletin, 136, 31 - 32." type="journal article" year="2016">Jablonski &amp; Hegner 2016</bibRefCitation>
)
</taxonomicName>
.
<taxonomicName id="4C484D693613494BFE11FED4FD0EFEB7" authority="Quah &amp; Grismer &amp; Lim &amp; Anuar &amp; Imbun, 2019" authorityName="Quah &amp; Grismer &amp; Lim &amp; Anuar &amp; Imbun" authorityYear="2019" box="[473,745,297,323]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="507" phylum="Chordata" rank="species" species="stuebingi" status="sp. nov.">
<emphasis id="B93CEAF83613494BFE11FED4FD0EFEB7" box="[473,745,297,323]" italics="true" pageId="6" pageNumber="507">Asthenodipsas stuebingi</emphasis>
</taxonomicName>
<emphasis id="B93CEAF83613494BFD38FED4FCADFEB7" bold="true" box="[752,842,297,323]" pageId="6" pageNumber="507">
<taxonomicNameLabel id="A20F57833613494BFD38FED4FCADFEB7" box="[752,842,297,323]" pageId="6" pageNumber="507" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
presumably feeds on snails or slugs like other mem- bers of the family and nothing is known about the reproductive biology of the species.
</paragraph>
</subSubSection>
</treatment>
</document>