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<document id="646D1CCD8D2A23CE56F3464C1865673C" ID-DOI="10.5281/zenodo.186646" ID-GBIF-Dataset="e80d7f6c-b367-43d7-80a8-1403082fb549" ID-ISSN="1175-5326" ID-Zenodo-Dep="186646" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461054379412" checkinUser="plazi" docAuthor="Wu, Min &amp; Zheng, Wei" docDate="2009" docId="03E18793FF98FF89FF16FF0473A37210" docLanguage="en" docName="zt02053p031.pdf" docOrigin="Zootaxa 2053" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Pupinidius obrutschewi subsp. obrutschewi Moellendorff 1901" docType="treatment" docVersion="7" lastPageNumber="16" masterDocId="FFD8FFEBFF95FF86FF81FF937359701A" masterDocTitle="A review of Chinese Pupinidius Moellendorff (Gastropoda, Stylommatophora: Enidae), with the description of a new species" masterLastPageNumber="31" masterPageNumber="1" pageNumber="14" updateTime="1698580698258" updateUser="plazi">
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<mods:title id="FB1E309747C3B11C060A173FDEEA7EE7">A review of Chinese Pupinidius Moellendorff (Gastropoda, Stylommatophora: Enidae), with the description of a new species</mods:title>
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<mods:namePart id="7753FE4A7EF89BDABBC58EB57E1CA6C6">Wu, Min</mods:namePart>
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<mods:date id="C1BE083DA1766BF1BEB4936DBB34A559">2009</mods:date>
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<subSubSection id="C352650EFF98FF8BFF16FF04702A70AB" box="[151,883,151,177]" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="8BF73685FF98FF8BFF16FF04702A70AB" blockId="13.[151,883,151,212]" box="[151,883,151,177]" pageId="13" pageNumber="14">
<heading id="D0BF81E9FF98FF8BFF16FF04702A70AB" bold="true" box="[151,883,151,177]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
<emphasis id="B93CEA97FF98FF8BFF16FF04702A70AB" bold="true" box="[151,883,151,177]" pageId="13" pageNumber="14">
9.
<taxonomicName id="4C484D06FF98FF8BFF35FF04702A70AB" ID-CoL="7V2B4" authority="Moellendorff, 1901" authorityName="Moellendorff" authorityYear="1901" box="[180,883,151,177]" class="Gastropoda" family="Enidae" genus="Pupinidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="14" phylum="Mollusca" rank="subSpecies" species="obrutschewi" subSpecies="obrutschewi">
<emphasis id="B93CEA97FF98FF8BFF35FF04713270AB" bold="true" box="[180,619,151,177]" italics="true" pageId="13" pageNumber="14">Pupinidius obrutschewi obrutschewi</emphasis>
(
<bibRefCitation id="EFD94B74FF98FF8BFDFBFF04703270AB" author="Moellendorff" box="[634,875,151,177]" pageId="13" pageNumber="31" refString="Moellendorff, O. F. (1901) Binnen-Mollusken aus Westchina und Centralasien. II. Annuaire du Musee Zoologique de l'Academie Imperiale des St. - Petersburg, 6, 299 - 404, Taf. XII - XVII." type="journal article" year="1901">Moellendorff, 1901</bibRefCitation>
)
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C352650EFF98FF8BFF16FF29728770CE" box="[151,478,186,212]" pageId="13" pageNumber="14" type="description">
<paragraph id="8BF73685FF98FF8BFF16FF29728770CE" blockId="13.[151,883,151,212]" box="[151,478,186,212]" pageId="13" pageNumber="14">
<figureCitation id="13732A00FF98FF8BFF16FF2973B970CE" box="[151,224,186,212]" captionStart="FIGURE 1" captionStartId="3.[151,255,1609,1633]" captionTargetBox="[161,1429,545,1583]" captionTargetId="figure@3.[151,1436,527,1584]" captionTargetPageId="3" captionText="FIGURE 1. Distribution map of Pupinidius spp. Red dot, Pupinidius anocamptus, P. nanpingensis nanpingensis and P. nanpingensis ambigua; red circle, P. porrectus; black dot, P. pupinella altispirus; black circle, P. chrysalis; blue dot, P. obrutschewi contractus and P. pupinella pupinella; blue circle: P. latilabrum; green dot, P. melinostoma, P. nanpingensis nanpingensis and P. wenxian sp. nov.; yellow dot, P. obrutschewi obrutschewi and P. streptaxis; yellow circle, P. porrecta pygmaea. P. pupinidius and P. gregorii have not been mapped here due to the unrecognizable locality names. Although the original description of the type locality of P. chrysalis is not accurate and geographically wrong, mapping of the species is possible here based only on the recognized information therein." httpUri="https://zenodo.org/record/186647/files/figure.png" pageId="13" pageNumber="14">Figs 1</figureCitation>
,
<figureCitation id="13732A00FF98FF8BFF6FFF2973A570CE" box="[238,252,186,212]" captionStart="FIGURE 4. A D" captionStartId="11.[151,257,1870,1894]" captionTargetBox="[220,1366,192,1846]" captionTargetId="figure@11.[220,1367,192,1846]" captionTargetPageId="11" captionText="FIGURE 4. A D, Pupinidius nanpingensis nanpingensis (Moellendorff, 1901). A B, respectively HBUMM- 05497 - specimen 7 &amp; 8; C, Holotype, SMF- 41994; D, type specimen, ZI RAS-N 193. E G, Pupinidius obrutschewi obrutschewi (Moellendorff, 1901). E, type specimen, ZI RAS-N 512; F, HBUMM- 05672 - specimen 12; G, Lectotype, SMF- 41982." httpUri="https://zenodo.org/record/186650/files/figure.png" pageId="13" pageNumber="14">4</figureCitation>
EG, 5AB, 8E, 13
</paragraph>
</subSubSection>
<subSubSection id="C352650EFF98FF8BFF16FE9270167182" pageId="13" pageNumber="14" type="reference_group">
<paragraph id="8BF73685FF98FF8BFF16FE9270487102" blockId="13.[151,1436,257,408]" box="[151,785,257,280]" pageId="13" pageNumber="14">
<treatmentCitationGroup id="AB5811ABFF98FF8BFF16FE9270487102" box="[151,785,257,280]" pageId="13" pageNumber="14">
<taxonomicName id="4C484D06FF98FF8BFF16FE9271087102" ID-CoL="7QFKZ" authority="Sturany, 1900: 29" authorityName="Sturany" authorityPageNumber="29" authorityYear="1900" box="[151,593,257,280]" class="Gastropoda" family="Enidae" genus="Buliminus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="obrutschewi">
<emphasis id="B93CEA97FF98FF8BFF16FE9272D57102" box="[151,396,257,280]" italics="true" pageId="13" pageNumber="14">Buliminus obrutschewi</emphasis>
<treatmentCitation id="0AE91094FF98FF8BFE12FE9271087102" author="Sturany" box="[403,593,257,280]" page="29" pageId="13" pageNumber="14" year="1900">
<bibRefCitation id="EFD94B74FF98FF8BFE12FE9271737102" author="Sturany" box="[403,554,257,280]" pageId="13" pageNumber="31" refString="Sturany, R. (1900) Obrutschew's Mollusken-Ausbbeute aus Hochasien. Denkschriften (Akademie der Wissenschaften in Wien, Mathematisch-Naturwissenschaftliche Klasse), 70, 17 - 48, 4 Taf." type="journal article" year="1900">Sturany, 1900</bibRefCitation>
: 29
</treatmentCitation>
</taxonomicName>
, pl. 3, figs 2628.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF73685FF98FF8BFF16FEB270757122" blockId="13.[151,1436,257,408]" box="[151,812,289,312]" pageId="13" pageNumber="14">
<treatmentCitationGroup id="AB5811ABFF98FF8BFF16FEB270757122" box="[151,812,289,312]" pageId="13" pageNumber="14">
<taxonomicName id="4C484D06FF98FF8BFF16FEB270717122" ID-CoL="7QFKZ" authority="Moellendorff, 1901: 342" authorityName="Moellendorff" authorityPageNumber="342" authorityYear="1901" box="[151,808,289,312]" class="Gastropoda" family="Enidae" genus="Buliminus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="obrutschewi" subGenus="Pupinidius">
<emphasis id="B93CEA97FF98FF8BFF16FEB2725A7122" box="[151,259,289,312]" italics="true" pageId="13" pageNumber="14">Buliminus</emphasis>
(
<emphasis id="B93CEA97FF98FF8BFE93FEB272DC7122" box="[274,389,289,312]" italics="true" pageId="13" pageNumber="14">Pupinidius</emphasis>
)
<emphasis id="B93CEA97FF98FF8BFE14FEB2714E7122" box="[405,535,289,312]" italics="true" pageId="13" pageNumber="14">obrutschewi</emphasis>
<treatmentCitation id="0AE91094FF98FF8BFD9FFEB270717122" author="Moellendorff" box="[542,808,289,312]" page="342" pageId="13" pageNumber="14" year="1901">
<bibRefCitation id="EFD94B74FF98FF8BFD9FFEB271AB7122" author="Moellendorff" box="[542,754,289,312]" pageId="13" pageNumber="31" refString="Moellendorff, O. F. (1901) Binnen-Mollusken aus Westchina und Centralasien. II. Annuaire du Musee Zoologique de l'Academie Imperiale des St. - Petersburg, 6, 299 - 404, Taf. XII - XVII." type="journal article" year="1901">Moellendorff, 1901</bibRefCitation>
: 342
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF73685FF98FF8BFF16FED270F97162" blockId="13.[151,1436,257,408]" pageId="13" pageNumber="14">
<treatmentCitationGroup id="AB5811ABFF98FF8BFF16FED270F97162" pageId="13" pageNumber="14">
<taxonomicName id="4C484D06FF98FF8BFF16FED270C27142" authority="Moellendorff, 1901: 342" authorityName="Moellendorff" authorityPageNumber="342" authorityYear="1901" box="[151,923,321,344]" class="Gastropoda" family="Enidae" genus="Buliminus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="14" phylum="Mollusca" rank="subSpecies" species="obrutschewi" subGenus="Pupinidius" subSpecies="eurystoma">
<emphasis id="B93CEA97FF98FF8BFF16FED2725A7142" box="[151,259,321,344]" italics="true" pageId="13" pageNumber="14">Buliminus</emphasis>
(
<emphasis id="B93CEA97FF98FF8BFE93FED272DC7142" box="[274,389,321,344]" italics="true" pageId="13" pageNumber="14">Pupinidius</emphasis>
)
<emphasis id="B93CEA97FF98FF8BFE14FED271D57142" box="[405,652,321,344]" italics="true" pageId="13" pageNumber="14">obrutschewi eurystoma</emphasis>
<treatmentCitation id="0AE91094FF98FF8BFD12FED270C27142" author="Moellendorff" box="[659,923,321,344]" page="342" pageId="13" pageNumber="14" year="1901">
<bibRefCitation id="EFD94B74FF98FF8BFD12FED2703E7142" author="Moellendorff" box="[659,871,321,344]" pageId="13" pageNumber="31" refString="Moellendorff, O. F. (1901) Binnen-Mollusken aus Westchina und Centralasien. II. Annuaire du Musee Zoologique de l'Academie Imperiale des St. - Petersburg, 6, 299 - 404, Taf. XII - XVII." type="journal article" year="1901">Moellendorff, 1901</bibRefCitation>
: 342
</treatmentCitation>
</taxonomicName>
, pl. 8, figs 23, 24;
<treatmentCitation id="0AE91094FF98FF8BFBE1FED276167142" author="Wiegmann" box="[1120,1359,321,344]" page="246" pageId="13" pageNumber="14" year="1901">
<bibRefCitation id="EFD94B74FF98FF8BFBE1FED276437142" author="Wiegmann" box="[1120,1306,321,344]" pageId="13" pageNumber="31" refString="Wiegmann, F. (1901) Binnen-Mollusken aus Westchina und Centralasien. Zootomische Untersuchungen. II. Die Buliminiden. Annuaire du Musee Zoologique de l'Academie Imperiale des St. - Petersburg, 2, 220 - 297." type="journal article" year="1901">Wiegmann, 1901</bibRefCitation>
: 246
</treatmentCitation>
, pl. 10, figs 4044;
<treatmentCitation id="0AE91094FF98FF8BFEC7FEF271537162" author="Kobelt" box="[326,522,353,376]" page="846" pageId="13" pageNumber="14" year="1902">
<bibRefCitation id="EFD94B74FF98FF8BFEC7FEF2728C7162" author="Kobelt" box="[326,469,353,376]" pageId="13" pageNumber="31" refString="Kobelt, W. (1902) Die Familie Buliminidae. Systematisches Conchylien-Cabinet, 397 - 1051, 133 pls." type="book chapter" year="1902">Kobelt, 1902</bibRefCitation>
: 846
</treatmentCitation>
, pl. 108, figs 1, 2, pl. 120, figs 2327.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF73685FF98FF8BFF16FE1270167182" blockId="13.[151,1436,257,408]" box="[151,847,385,408]" pageId="13" pageNumber="14">
<treatmentCitationGroup id="AB5811ABFF98FF8BFF16FE1270167182" box="[151,847,385,408]" pageId="13" pageNumber="14">
<taxonomicName id="4C484D06FF98FF8BFF16FE1271E37182" ID-CoL="7V2B4" authority="Yen, 1939: 83" authorityName="Yen" authorityPageNumber="83" authorityYear="1939" box="[151,698,385,408]" class="Gastropoda" family="Enidae" genus="Pupinidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="14" phylum="Mollusca" rank="subSpecies" species="obrutschewi" subSpecies="obrutschewi">
<emphasis id="B93CEA97FF98FF8BFF16FE1271457182" box="[151,540,385,408]" italics="true" pageId="13" pageNumber="14">Pupinidius obrutschewi obrutschewi</emphasis>
<treatmentCitation id="0AE91094FF98FF8BFDA2FE1271E37182" author="Yen" box="[547,698,385,408]" page="83" pageId="13" pageNumber="14" year="1939">
<bibRefCitation id="EFD94B74FF98FF8BFDA2FE1271CA7182" author="Yen" box="[547,659,385,408]" pageId="13" pageNumber="31" refString="Yen, T. C. (1939) Die chinesischen Land-und Sußwasser-Gastropoden des Natur-Museums Senckenberg. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 444, 1 - 234, pls. 1 - 16." type="journal article" year="1939">Yen, 1939</bibRefCitation>
: 83
</treatmentCitation>
</taxonomicName>
, pl. 7, fig. 46.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C352650EFF98FF8BFF16FE2D77B5734C" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="8BF73685FF98FF8BFF16FE2D77B5734C" blockId="13.[151,1437,446,2038]" pageId="13" pageNumber="14">
<emphasis id="B93CEA97FF98FF8BFF16FE2D72D271C2" bold="true" box="[151,395,446,472]" pageId="13" pageNumber="14">Material examined.</emphasis>
HBUMM-05672 (20 out of 36 mature shells were measured; specimen-1 for dissection):
<quantity id="4CB09B60FF98FF8BFF16FE7773A971E4" box="[151,240,484,510]" metricMagnitude="3" metricUnit="m" metricValue="1.233" pageId="13" pageNumber="14" unit="m" value="1233.0">1233 m</quantity>
a.s.l., 33°41΄47.3΄ N, 104°29΄10.3΄ E, eastern bank of Lianghekou, Gansu,
<date id="FFF61045FF98FF8BFBE6FE77778271E4" box="[1127,1243,484,510]" pageId="13" pageNumber="14" value="2006-10-02">2006-X-2</date>
, limestone, coll. J.Liu &amp; W.Zheng; HBUMM-05678 (30 fmss):
<quantity id="4CB09B60FF98FF8BFD3EFD987041723F" box="[703,792,523,549]" metricMagnitude="3" metricUnit="m" metricValue="1.286" pageId="13" pageNumber="14" unit="m" value="1286.0">1286 m</quantity>
a.s.l., 33°41΄38.5΄ N, 104°28΄48.6΄ E, western bank of Lianghekou, Gansu,
<date id="FFF61045FF98FF8BFE10FDA271517251" box="[401,520,561,587]" pageId="13" pageNumber="14" value="2006-10-02">2006-X-2</date>
, other stone replacing limestone, coll. J.Liu &amp; W.Zheng (measurement: ewh 1.5001.6691.875, wh 6.5007.0447.500, h 18.721.023.3, diam. maj. 10.111.111.8, aph 9.09.710.9, apw 6.47.58.0, h/diam. maj. 1.621.892.12, h/aph 1.992.162.33);
<collectingCountry id="F35F7615FF98FF8BFC70FDEE7754728D" box="[1009,1037,637,663]" name="Serbia" pageId="13" pageNumber="14">ZI</collectingCountry>
RAS-N512:
<typeStatus id="54F38827FF98FF8BFB2AFDEE7787728D" box="[1195,1246,637,663]" pageId="13" pageNumber="14">type</typeStatus>
material,
<date id="FFF61045FF98FF8BFAD6FDEE73BA72A7" pageId="13" pageNumber="14" value="1885-06-22">1885- VI-22</date>
, coll. Potanin. SMF-41982:
<typeStatus id="54F38827FF98FF8BFDC1FD3071E372A7" box="[576,698,675,701]" pageId="13" pageNumber="14" type="lectotype">Lectotype</typeStatus>
v.
<taxonomicName id="4C484D06FF98FF8BFD60FD3770BA72A7" authority="Mlldff." authorityName="Mlldff." box="[737,995,675,701]" class="Gastropoda" family="Enidae" genus="Buliminus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="eurystoma">
<emphasis id="B93CEA97FF98FF8BFD60FD3771AA72A7" box="[737,755,676,701]" italics="true" pageId="13" pageNumber="14">B</emphasis>
.
<emphasis id="B93CEA97FF98FF8BFC85FD3770DA72A7" box="[772,899,676,701]" italics="true" pageId="13" pageNumber="14">eurystoma</emphasis>
Mlldff.
</taxonomicName>
Liang-dshia-pa, Kansu,
<collectingCountry id="F35F7615FF98FF8BFA92FD30763972A7" box="[1299,1376,675,701]" name="China" pageId="13" pageNumber="14">China</collectingCountry>
. Slg.
<collectingCountry id="F35F7615FF98FF8BFF16FD5973F272FE" box="[151,171,714,740]" name="Iceland" pageId="13" pageNumber="14">O</collectingCountry>
.v.Moellendorff. SMF-41983:
<typeStatus id="54F38827FF98FF8BFD91FD5971DB72FE" box="[528,642,714,740]" pageId="13" pageNumber="14" type="paratype">Paratypes</typeStatus>
v.
<taxonomicName id="4C484D06FF98FF8BFD25FD5970C272FE" authority="Mlldff." authorityName="Mlldff." box="[676,923,714,740]" class="Gastropoda" family="Enidae" genus="Buliminus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="eurystoma">
<emphasis id="B93CEA97FF98FF8BFD25FD5971EF72F9" box="[676,694,714,739]" italics="true" pageId="13" pageNumber="14">B</emphasis>
.
<emphasis id="B93CEA97FF98FF8BFD44FD59706672F9" box="[709,831,714,739]" italics="true" pageId="13" pageNumber="14">eurystoma</emphasis>
Mlldff.
</taxonomicName>
2 fmss. Liang-dshia-pa, Kansu,
<collectingCountry id="F35F7615FF98FF8BFA98FD59763A72FE" box="[1305,1379,714,740]" name="China" pageId="13" pageNumber="14">China</collectingCountry>
. Slg.
<collectingCountry id="F35F7615FF98FF8BFF16FD6373F27310" box="[151,171,752,778]" name="Iceland" pageId="13" pageNumber="14">O</collectingCountry>
.v.Moellendorff (measurement of
<typeStatus id="54F38827FF98FF8BFDB7FD6371217310" box="[566,632,752,778]" pageId="13" pageNumber="14">types</typeStatus>
: ewh 1.5001.5631.625, wh 6.8757.1297.375, h 21.122.223.8, diam. maj. 12.913.413.9, aph 9.910.711.2, apw 8.58.89.0, h/diam. maj. 1.581.661.72, h/aph 1.972.062.13). SMF-41984: One juvenile. Wen-hsien, Kansu,
<collectingCountry id="F35F7615FF98FF8BFC0CFCAF708E734C" box="[909,983,828,854]" name="China" pageId="13" pageNumber="14">China</collectingCountry>
. Slg.
<collectingCountry id="F35F7615FF98FF8BFB96FCAF7772734C" box="[1047,1067,828,854]" name="Iceland" pageId="13" pageNumber="14">O</collectingCountry>
.v.Moellendorff.
</paragraph>
</subSubSection>
<subSubSection id="C352650EFF98FF8BFF44FCF177627366" box="[197,1083,866,892]" pageId="13" pageNumber="14" type="distribution">
<paragraph id="8BF73685FF98FF8BFF44FCF177627366" blockId="13.[151,1437,446,2038]" box="[197,1083,866,892]" pageId="13" pageNumber="14">
<emphasis id="B93CEA97FF98FF8BFF44FCF1723C7366" bold="true" box="[197,357,866,892]" pageId="13" pageNumber="14">Distribution.</emphasis>
Gansu (
<typeStatus id="54F38827FF98FF8BFE46FCF172A27366" box="[455,507,866,892]" pageId="13" pageNumber="14">type</typeStatus>
locality, Liang-dshia-pa, Lianghekou,
<taxonomicName id="4C484D06FF98FF8BFC40FCF177777366" box="[961,1070,866,892]" class="Gastropoda" family="Enidae" genus="Pupinidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="30" phylum="Mollusca" rank="species" species="wenxian" status="sp. nov.">Wenxian</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C352650EFF98FF8BFF44FC1B710E73B9" box="[197,599,904,931]" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="8BF73685FF98FF8BFF44FC1B710E73B9" blockId="13.[151,1437,446,2038]" box="[197,599,904,931]" pageId="13" pageNumber="14">
<emphasis id="B93CEA97FF98FF8BFF44FC1B723373B8" bold="true" box="[197,362,904,930]" pageId="13" pageNumber="14">
<typeStatus id="54F38827FF98FF8BFF44FC1B725A73B8" box="[197,259,904,930]" pageId="13" pageNumber="14">Type</typeStatus>
locality.
</emphasis>
Schy-pa (S Gansu).
</paragraph>
</subSubSection>
<subSubSection id="C352650EFF98FF89FF44FC3D70E37169" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14" type="diagnosis">
<paragraph id="8BF73685FF98FF8BFF44FC3D70C97421" blockId="13.[151,1437,446,2038]" pageId="13" pageNumber="14">
<emphasis id="B93CEA97FF98FF8BFF44FC3D721D73D2" bold="true" box="[197,324,942,968]" pageId="13" pageNumber="14">Diagnosis.</emphasis>
Shell bandless; striate. Aperture faintly with or without angular tubercle. Most swollen part at body whorl. With 5.7507.375 whorls; height 15.923.8, diam. maj. 10.413.9. Penis without penial caecum; internally without fine longitudinal pilaster. Papilla developed by fused proximal pilasters forming V-shaped structure present. Epiphallus straight. Epiphallic caecum present.
</paragraph>
<paragraph id="8BF73685FF98FF8BFF44FBDB73817648" blockId="13.[151,1437,446,2038]" pageId="13" pageNumber="14">
Re-description of shell. Pointed-ovate; apex protruded. Most swollen part at body whorl. Shell dextral; thin; solid; opaque; glossy; speckled or not; not spirally grooved;with 5.7506.2506.625 whorls. Whorls flattened; not shouldered. Growth lines usually not very clear. Embryonic shell smooth; polished; with 1.5001.5511.750 whorls. Postnuclear whorls smooth. Suture normal, without narrow defined zone on beneath whorl. Last whorl slightly gradually ascending towards aperture; slightly angular at periphery (in
<typeStatus id="54F38827FF98FF8BFF16FA95725A753A" box="[151,259,1286,1312]" pageId="13" pageNumber="14" type="lectotype">lectotype</typeStatus>
indistinct); without smoothed spiral peripheral depression nor rugate region. Shell 15.919.521.5 high; 10.411.312.0 in diam. maj. Height/width ratio 1.441.732.06. Aperture in a plane; completely attached to body whorl; truncate-ovate; not adnate; oblique; not armed; faintly with angular tubercle, or without angular tubercle; 8.29.610.2 high; 6.77.48.0 wide. Ratio of shell height to aperture height 1.772.032.28. Peristome narrowly reflexed with distinct cuff; expanded; sharp. Parietal callus indistinct. Columellar margin reflexed. Columella vertical. Outer edge of columellar lip vertical. Umbilicus very narrowly open. Apical whorls normally coloured. Shell multicoloured; in darker or lighter brown. With narrow or thick white stripes below protoconch. Aperture white; not brightly banded; spirally without colour band.
</paragraph>
<paragraph id="8BF73685FF98FF8BFF44F9CD76CC77EC" blockId="13.[151,1437,446,2038]" pageId="13" pageNumber="14">Re-description of genitalia. Vas deferens slightly swollen proximally; entering epiphallus apically with distinct demarcation. Epiphallus more or less short; slightly narrowed towards proximal end; normally straight; externally smooth. Epiphallic caecum present; blunt apically; located near vas deferens entrance. Flagellum short; tubular; proximally normal; with tip pointed. Penis with terminal entrance of epiphallus; clavate; without penial caecum; distally enlarged; thin-walled; internally without fine pilasters, with with 23 circular ridges and 1 V-shaped pilaster. V-shaped pilaster with free proximal end approaching at penial retractor insertion, distally developing a small nipple-like papilla. Penis without process. Penial verge absent. Penial appendix moderately long; branched off from proximal portion of penis; divided into sections including A-1+A-2, A-3 and A-4+A-5. A-1 and A-2 fused, interiorly smooth. A-2 and A-3 not fused. Boundary between A-4 and A-5 indistinct. A-5 long; almost straight. Penial retractor biramous; attaching to penis in middle part and to A-1+ A-2; with penial retractor arms arising from diaphragm closed to each other.</paragraph>
<caption id="DF37660DFF9BFF88FF16F8B077B77765" httpUri="https://zenodo.org/record/186651/files/figure.png" pageId="14" pageNumber="15" targetBox="[212,1373,178,1801]" targetPageId="14">
<paragraph id="8BF73685FF9BFF88FF16F8B077B77765" blockId="14.[151,1437,1827,1920]" pageId="14" pageNumber="15">
<emphasis id="B93CEA97FF9BFF88FF16F8B0727B7721" bold="true" box="[151,290,1827,1851]" pageId="14" pageNumber="15">FIGURE 5.</emphasis>
AB,
<taxonomicName id="4C484D06FF9BFF88FEEFF8B077497720" authority="Moellendorff, 1901" authorityName="Moellendorff" authorityYear="1901" box="[366,1040,1827,1850]" class="Gastropoda" family="Enidae" genus="Pupinidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="15" phylum="Mollusca" rank="subSpecies" species="obrutschewi" subSpecies="obrutschewi">
<emphasis id="B93CEA97FF9BFF88FEEFF8B072B37720" box="[366,490,1827,1850]" italics="true" pageId="14" pageNumber="15">Pupinidius</emphasis>
<emphasis id="B93CEA97FF9BFF88FE70F8B070487720" box="[497,785,1827,1850]" italics="true" pageId="14" pageNumber="15">obrutschewi obrutschewi</emphasis>
(Moellendorff, 1901)
</taxonomicName>
, 2 paratypes of SMF-41983. CD,
<taxonomicName id="4C484D06FF9BFF88FF16F8D571A57747" authority="Moellendorff, 1901" authorityName="Moellendorff" authorityYear="1901" box="[151,764,1862,1885]" class="Gastropoda" family="Enidae" genus="Pupinidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="15" phylum="Mollusca" rank="subSpecies" species="obrutschewi" subSpecies="contractus">
<emphasis id="B93CEA97FF9BFF88FF16F8D572527747" box="[151,267,1862,1885]" italics="true" pageId="14" pageNumber="15">Pupinidius</emphasis>
<emphasis id="B93CEA97FF9BFF88FE93F8D571577747" box="[274,526,1862,1885]" italics="true" pageId="14" pageNumber="15">obrutschewi contractus</emphasis>
(Moellendorff, 1901)
</taxonomicName>
. C, HBUMM-04282-specimen 11; D, Lectotype, SMF-41981. EF,
<taxonomicName id="4C484D06FF9BFF88FF4FF8FA71C37765" authority="Moellendorff, 1901" authorityName="Moellendorff" authorityYear="1901" box="[206,666,1896,1920]" class="Gastropoda" family="Enidae" genus="Pupinidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="porrectus">
<emphasis id="B93CEA97FF9BFF88FF4FF8FA7218779A" box="[206,321,1897,1920]" italics="true" pageId="14" pageNumber="15">Pupinidius</emphasis>
<emphasis id="B93CEA97FF9BFF88FEC8F8FA72F7779A" box="[329,430,1897,1920]" italics="true" pageId="14" pageNumber="15">porrectus</emphasis>
(Moellendorff, 1901)
</taxonomicName>
. E, Lectotype, SMF-41998; F, NHM-1902.5.13.20 Reg.
</paragraph>
</caption>
<paragraph id="8BF73685FF9BFF89FF44F83D705F713C" blockId="14.[151,1436,1966,2030]" lastBlockId="15.[151,1436,152,522]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">Additional retractor other than penial or appendicular absent. Muscular band connecting vagina and epiphallus absent. Atrium short; with weak retractor. Free oviduct longer than vagina. Vagina short; not swollen; straight; not lined with loose, spongy tissue; unpigmented. Bursa copulatrix duct moderately long; proximally straight. Bursa copulatrix with stalk; without apical ligament; normal in size; with short neck; well defined. Diverticle normally present; longer than reservoir; unexpanded. Bursa copulatrix and diverticle distinguishable; forked more distally from their base.</paragraph>
<paragraph id="8BF73685FF9AFF89FF44FEA170E37169" blockId="15.[151,1436,152,522]" pageId="15" pageNumber="16">Measurement of genitalia. P: 3; Ep: 7.8; Fl: 1.4; VD: 15.0; Va: 2.3; FO: 4.3; BCD: 5.9; BC: 1.1; D: 4.5; A- 1+A-2: 3.8; A3: 1.8; A-4+A-5: 12 (in HBUMM-05672-specimen 1).</paragraph>
</subSubSection>
<subSubSection id="C352650EFF9AFF89FF16FEEC73A37210" pageId="15" pageNumber="16" type="discussion">
<paragraph id="8BF73685FF9AFF89FF16FEEC73A37210" blockId="15.[151,1436,152,522]" pageId="15" pageNumber="16">
<emphasis id="B93CEA97FF9AFF89FF16FEEC72487183" bold="true" box="[151,273,383,409]" pageId="15" pageNumber="16">Remarks.</emphasis>
The new collected specimens are more or less smaller than the
<typeStatus id="54F38827FF9AFF89FB90FE13771D7180" box="[1041,1092,384,410]" pageId="15" pageNumber="16">type</typeStatus>
specimens in shell size. The population (HBUMM-05678), habited in the opposite bank of Bailongjiang River, differs slightly from the individuals represented by HBUMM-05672 by larger shell size, darker colouration, and more dense striation on shell.
</paragraph>
</subSubSection>
</treatment>
</document>