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<mods:title id="4479B2D823754CFC25A62C5BCFB16E03">Revision of the Seashore-dwelling Subgenera Emplenota Casey and Triochara Bernhauer (Coleoptera: Staphylinidae: genus Aleochara) from Japan</mods:title>
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<mods:namePart id="7078C9B4A425985811449027C8F9E517">Yamamoto, Shûhei</mods:namePart>
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<treatment id="03DE87D80830FFEFFF28FE1129BC3F24" ID-DOI="http://doi.org/10.5281/zenodo.6175378" ID-GBIF-Taxon="119528577" ID-Zenodo-Dep="6175378" LSID="urn:lsid:plazi:treatment:03DE87D80830FFEFFF28FE1129BC3F24" httpUri="http://treatment.plazi.org/id/03DE87D80830FFEFFF28FE1129BC3F24" lastPageId="30" lastPageNumber="31" pageId="28" pageNumber="29">
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<paragraph id="8BC836CE0830FFEDFF28FE112A603910" blockId="28.[151,613,487,546]" box="[151,613,487,513]" pageId="28" pageNumber="29">
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Subgenus
<taxonomicName id="4C774D4D0830FFEDFEA8FE112A603910" authority="Bernhauer, 1901" authorityName="Bernhauer" authorityYear="1901" box="[279,613,487,513]" class="Insecta" family="Staphylinidae" genus="Triochara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B903EADC0830FFEDFEA8FE11298A3910" bold="true" box="[279,399,487,513]" italics="true" pageId="28" pageNumber="29">Triochara</emphasis>
Bernhauer, 1901
</taxonomicName>
</emphasis>
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</paragraph>
<paragraph id="8BC836CE0830FFEDFF28FDFC2A053933" blockId="28.[151,613,487,546]" box="[151,512,522,546]" pageId="28" pageNumber="29">
(
<figureCitation id="134C2A4B0830FFEDFF20FDFC29223933" box="[159,295,522,546]" captionStart-0="FIGURES 53 64" captionStart-1="FIGURES 65 70" captionStart-2="FIGURES 71 78" captionStart-3="FIGURES 79 85" captionStart-4="FIGURES 86 92" captionStartId-0="26.[151,264,1753,1776]" captionStartId-1="29.[151,264,1965,1988]" captionStartId-2="33.[151,264,1821,1844]" captionStartId-3="36.[151,264,1915,1938]" captionStartId-4="41.[151,264,1942,1965]" captionTargetBox-0="[151,1433,193,1728]" captionTargetBox-1="[179,1408,270,1932]" captionTargetBox-2="[152,1426,187,1820]" captionTargetBox-3="[178,1416,289,1854]" captionTargetBox-4="[153,1404,302,1941]" captionTargetId-0="figure@26.[151,1436,193,1732]" captionTargetId-1="figure@29.[162,1432,253,1954]" captionTargetId-2="figure@33.[151,1436,141,1822]" captionTargetId-3="figure@36.[151,1436,265,1894]" captionTargetId-4="figure@41.[151,1436,265,1951]" captionTargetPageId-0="26" captionTargetPageId-1="29" captionTargetPageId-2="33" captionTargetPageId-3="36" captionTargetPageId-4="41" captionText-0="FIGURES 53 64. Japanese species of Aleochara (Triochara). 53 - 56. Aleochara (Triochara) trisulcata; 57 - 60. A. (T.) zerchei; 61 - 64. A. (T.) nubis. (Habitus: 53, 57, 61; Pronotum: 54, 58, 62; Mesoventrite and metaventrite: 55, 59, 63; Inter coxal process of mesoventrite: 56, 60, 64)." captionText-1="FIGURES 65 70. Body parts of Aleochara (Triochara) trisulcata of male. 65. right antenna; 66. labium; 67. maxilla; 68. mentum; 69. labrum; 70. right foreleg in ventral view." captionText-2="FIGURES 71 78. Terminalia of Aleochara (Triochara) trisulcata. 71. tergite VIII of male; 72. tergite VIII of female; 73. sternite VIII of male; 74. sternite VIII of female; 75. male genitalia: median lobe of aedeagus in lateral view; 76. male genitalia: median lobe of aedeagus in ventral view; 77. apical lobe of paramerite, lateral view; 78. female genitalia: spermatheca." captionText-3="FIGURES 79 85. Terminalia of Aleochara (Triochara) zerchei. 79. tergite VIII of male; 80. tergite VIII of female; 81. sternite VIII of male; 82. sternite VIII of female; 83. male genitalia: median lobe of aedeagus in lateral view; 84. male genitalia: median lobe of aedeagus in ventral view; 85. female genitalia: spermatheca." captionText-4="FIGURES 86 92. Terminalia of Aleochara (Triochara) nubis. 86. tergite VIII of male; 87. tergite VIII of female; 88. sternite VIII of male; 89. sternite VIII of female; 90. male genitalia: median lobe of aedeagus in lateral view; 91. male genitalia: median lobe of aedeagus in ventral view; 92. female genitalia: spermatheca." httpUri-0="https://zenodo.org/record/282603/files/figure.png" httpUri-1="https://zenodo.org/record/282604/files/figure.png" httpUri-2="https://zenodo.org/record/282605/files/figure.png" httpUri-3="https://zenodo.org/record/282606/files/figure.png" httpUri-4="https://zenodo.org/record/282607/files/figure.png" pageId="28" pageNumber="29">Figs. 5392</figureCitation>
,
<figureCitation id="134C2A4B0830FFEDFE8DFDFC298F3933" box="[306,394,522,546]" captionStart="FIGURES 93 100" captionStartId="43.[151,264,1848,1871]" captionTargetBox="[201,1397,201,1819]" captionTargetId="figure@43.[186,1410,193,1833]" captionTargetPageId="43" captionText="FIGURES 93 100. Collecting sites of the subgenera Emplenota and Triochara of the genus Aleochara in Japan. 93. Kourakaigan (Matsue-shi, Shimane-ken; habitat of A. (E.) fucicola, A. (E.) segregata, and A. (E.) hayamai; arrow); 94. drifted seaweed and flotsams on beach (Koura-kaigan); 95. Mochirippu-numa (Hamanaka-chô, Hokkaidô; A. (E.) puetzi, arrow); 96. Ochiishi-misaki (Nemuro-shi, Hokkaidô; A. puetzi); 97. Kakanokuketo (Shimane-ken; habitat of A. fucicola, A. segregata, A. hayamai, and the type locality of A. (E.) yamato; A. fucicola, arrow); 98. Hamamatsu (Nemuro-shi, Hokkaidô; habitat of A. (T.) nubis, arrow); 99. drifted seaweed on beach (Hamamatsu); 100. mating behaviour of A. nubis at Hamamatsu." httpUri="https://zenodo.org/record/282608/files/figure.png" pageId="28" pageNumber="29">98100</figureCitation>
,
<figureCitation id="134C2A4B0830FFEDFE2BFDFC29F23933" box="[404,503,522,546]" captionStart="FIGURES 104 105" captionStartId="45.[151,264,1157,1180]" captionTargetBox="[184,1387,212,1100]" captionTargetId="figure@45.[151,1436,193,1107]" captionTargetPageId="45" captionText="FIGURES 104 105. Collecting localities of Aleochara (Triochara) spp. based on specimens examined. 104. A. (T.) trisulcata (Pointed by white circle); A. (T.) nubis (Pointed by white square); 105. A. (T.) zerchei (Pointed by white triangular)." httpUri="https://zenodo.org/record/282610/files/figure.png" pageId="28" pageNumber="29">104105</figureCitation>
)
</paragraph>
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<taxonomicName id="4C774D4D0830FFEDFF28FDA429F43979" authority="Bernhauer, 1901b: 373" authorityName="Bernhauer" authorityPageNumber="373" authorityYear="1901" box="[151,497,594,616]" class="Insecta" family="Staphylinidae" genus="Triochara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B903EADC0830FFEDFF28FDA428FE3979" box="[151,251,594,616]" italics="true" pageId="28" pageNumber="29">Triochara</emphasis>
<treatmentCitation id="0AD610DF0830FFEDFEBAFDA429F43979" author="Bernhauer" box="[261,497,594,616]" page="373" pageId="28" pageNumber="29" year="1901">
<bibRefCitation id="EFE64B3F0830FFEDFEBAFDA429BA3979" author="Bernhauer" box="[261,447,594,616]" pageId="28" pageNumber="29" refString="Bernhauer, M. (1901 b) Neue exotische Arten der Gattung Aleochara Gravh. Stettiner Entomologische Zeitung, 62, 366 - 373." type="journal article" year="1901" yearSuffix="b">Bernhauer, 1901b</bibRefCitation>
: 373
</treatmentCitation>
</taxonomicName>
(original description;
<typeStatus id="54CC886C0830FFEDFD65FDA52B033979" box="[730,774,595,616]" pageId="28" pageNumber="29">type</typeStatus>
species:
<taxonomicName id="4C774D4D0830FFEDFCD8FDA42CC03978" ID-CoL="BN7H" authority="Weise, 1877" authorityName="Weise" authorityYear="1877" box="[871,1221,594,617]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="trisulcata">
<emphasis id="B903EADC0830FFEDFCD8FDA42C3F3979" box="[871,1082,594,616]" italics="true" pageId="28" pageNumber="29">Aleochara trisulcata</emphasis>
<bibRefCitation id="EFE64B3F0830FFEDFBFCFDA42CC03978" author="Weise" box="[1091,1221,594,617]" pageId="28" pageNumber="29" refString="Weise, J. (1877) Japanische Staphilinidae [sic] und Pselaphidae. In: Beitrage zur Kaferfauna von Japan, meist auf R. Hiller's Sammlungen basiert. Deutsche Entomologische Zeitschrift, 21, 88 - 100." type="journal article" year="1877">Weise, 1877</bibRefCitation>
</taxonomicName>
);
<treatmentCitation id="0AD610DF0830FFEDFB65FDA428E83996" author="Eichelbaum" page="246" pageId="28" pageNumber="29" year="1909">
<bibRefCitation id="EFE64B3F0830FFEDFB65FDA42D9D3979" author="Eichelbaum" box="[1242,1432,594,616]" pageId="28" pageNumber="29" refString="Eichelbaum, F. (1909) Katalog der Staphyliniden-Gattungen nebst Angabe ihrer Literatur, Synonyme, Artenzahl, geographischen Verbreitung und ihrer bekannten Larvenzustande. Memoires de la Societe Entomologique de Belgique, 17, 71 - 280." type="journal article" year="1909">Eichelbaum, 1909</bibRefCitation>
: 246
</treatmentCitation>
(as subgenus; generic catalogue of world
<taxonomicName id="4C774D4D0830FFEDFD23FD872B293996" ID-CoL="8VR4S" box="[668,812,625,647]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="family">Staphylinidae</taxonomicName>
); Fenyes, 1920: 398 (as subgenus; key to world subgenera of
<taxonomicName id="4C774D4D0830FFEDFF78FD66293539B7" ID-CoL="8VRG3" box="[199,304,656,678]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B903EADC0830FFEDFF78FD66293539B7" box="[199,304,656,678]" italics="true" pageId="28" pageNumber="29">Aleochara</emphasis>
</taxonomicName>
), 414 (as subgenus; catalogue of world species; diagnosis; addition of 5 species to genus);
<treatmentCitation id="0AD610DF0830FFEDFB61FD6628E839D5" author="Scheerpeltz" page="447" pageId="28" pageNumber="29" year="1925">
<bibRefCitation id="EFE64B3F0830FFEDFB61FD662D9D39B7" author="Scheerpeltz" box="[1246,1432,656,678]" pageId="28" pageNumber="29" refString="Scheerpeltz, O. (1925). Staphylinidae [II]. In: Winkler, A. (Ed.), Catalogus Coleopterorum regionis palaearcticae, pars 4, Winkler &amp; Wagner, Wien, pp. 369 - 496." type="journal article" year="1925">Scheerpeltz, 1925</bibRefCitation>
: 447
</treatmentCitation>
(as subgenus; catalogue of Palaearctic species of
<taxonomicName id="4C774D4D0830FFEDFD4DFD582B7B39D5" ID-CoL="8VR4S" box="[754,894,686,708]" class="Insecta" family="Staphylinidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="family">Aleocharinae</taxonomicName>
);
<treatmentCitation id="0AD610DF0830FFEDFC2FFD582D0C39D5" author="Bernhauer" box="[912,1289,686,708]" page="795" pageId="28" pageNumber="29" year="1926">
<bibRefCitation id="EFE64B3F0830FFEDFC2FFD582CDC39D5" author="Bernhauer" box="[912,1241,686,708]" pageId="28" pageNumber="29" refString="Bernhauer, M. &amp; Scheerpeltz, O. (1926) Staphylinidae. VI. (Pars 82). In: Junk, W. &amp; Schenkling, S. (Eds.), Coleopterorum Catalogus. Vol. 5. Staphylinidae. W. Junk, Berlin, pp. 499 - 988." type="book chapter" year="1926">Bernhauer &amp; Scheerpeltz, 1926</bibRefCitation>
: 795
</treatmentCitation>
(as subgenus; catalogue of world species of
<taxonomicName id="4C774D4D0830FFEDFDB4FD3B2A9339F2" ID-CoL="8VR4S" box="[523,662,717,739]" class="Insecta" family="Staphylinidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="family">Aleocharinae</taxonomicName>
; exclusion of these 5 species and only one species,
<taxonomicName id="4C774D4D0830FFEDFB77FD382D4F39F2" ID-CoL="BN7H" box="[1224,1354,717,739]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="trisulcata">
<emphasis id="B903EADC0830FFEDFB77FD382D4F39F2" box="[1224,1354,717,739]" italics="true" pageId="28" pageNumber="29">A. trisulcata</emphasis>
</taxonomicName>
, left as subgenus);
<treatmentCitation id="0AD610DF0830FFEDFE81FD1D2A323810" author="Blackwelder" box="[318,567,747,769]" page="397" pageId="28" pageNumber="29" year="1952">
<bibRefCitation id="EFE64B3F0830FFEDFE81FD1D2A013810" author="Blackwelder" box="[318,516,747,769]" pageId="28" pageNumber="29" refString="Blackwelder, R. E. (1952) The generic names of the beetle family Staphylinidae, with an essay on genotypy. United States National Museum Bulletin, 200, I - IV + 1 - 483." type="journal article" year="1952">Blackwelder, 1952</bibRefCitation>
: 397
</treatmentCitation>
(as subgenus; catalogue entry; generic catalogue of world
<taxonomicName id="4C774D4D0830FFEDFB14FD1D2D3E3810" ID-CoL="8VR4S" box="[1195,1339,747,769]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="family">Staphylinidae</taxonomicName>
);
<treatmentCitation id="0AD610DF0830FFEDFAEFFD1D29283831" author="Assing" page="219" pageId="28" pageNumber="29" year="1995">
<bibRefCitation id="EFE64B3F0830FFEDFAEFFD1D28F83831" author="Assing" pageId="28" pageNumber="29" refString="Assing, V. (1995) The Palaearctic species of Emplenota Casey, Polystomota Casey, Triochara Bernhauer and Skenochara Bernhauer &amp; Scheerpeltz, with descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). Beitrage zur Entomologie, 45, 217 - 237." type="journal article" year="1995">Assing, 1995</bibRefCitation>
: 219
</treatmentCitation>
(as genus; diagnostic key to genera), 229 (notes on genus), 230 (key to all species of genus), 230 (descriptions of each species);
<bibRefCitation id="EFE64B3F0830FFEDFE36FCDC2A71382E" author="Maus" box="[393,628,809,831]" pageId="28" pageNumber="29" refString="Maus, C. &amp; Ashe, J. S. (1998 c) Aleochara (Triochara). Version 11 September 1998 (under construction). Available from http: // tolweb. org / Aleochara _ % 28 Triochara % 29 / 9905 / 1998.09.11 in The Tree of Life Web Project, http: // tolweb. org / (accessed 20 July 2012)." type="journal article" year="1998" yearSuffix="c">Maus &amp; Ashe, 1998c</bibRefCitation>
(online) (world checklist of subgenus; diagnosis; bionomics; phylogenetic relationships);
<treatmentCitation id="0AD610DF0830FFEDFEE1FCBE2A2D384F" author="Smetana" box="[350,552,840,862]" page="358" pageId="28" pageNumber="29" year="2004">
<bibRefCitation id="EFE64B3F0830FFEDFEE1FCBE29FD384C" author="Smetana" box="[350,504,840,861]" pageId="28" pageNumber="29" refString="Smetana, A. (2004) Aleocharinae. In: Lobl, I &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera, Vol. 2. Apollo Books, Stenstrup, Denmark, pp. 353 - 494." type="book" year="2004">Smetana, 2004</bibRefCitation>
: 358
</treatmentCitation>
(as subgenus; catalogue of Palaearctic species of
<taxonomicName id="4C774D4D0830FFEDFB9FFCBE2CA9384F" ID-CoL="8VR4S" box="[1056,1196,840,862]" class="Insecta" family="Staphylinidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="family">Aleocharinae</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C36D65450830FFEFFF28FC7C29983AB3" lastPageId="30" lastPageNumber="31" pageId="28" pageNumber="29" type="description">
<paragraph id="8BC836CE0830FFEDFF28FC7C2C803C7F" blockId="28.[151,1437,906,2011]" pageId="28" pageNumber="29">
<emphasis id="B903EADC0830FFEDFF28FC7C298C38B2" bold="true" box="[151,393,906,931]" pageId="28" pageNumber="29">Redescription. Body</emphasis>
(
<figureCitation id="134C2A4B0830FFEDFE26FC7C2A4838B3" box="[409,589,906,930]" captionStart="FIGURES 53 64" captionStartId="26.[151,264,1753,1776]" captionTargetBox="[151,1433,193,1728]" captionTargetId="figure@26.[151,1436,193,1732]" captionTargetPageId="26" captionText="FIGURES 53 64. Japanese species of Aleochara (Triochara). 53 - 56. Aleochara (Triochara) trisulcata; 57 - 60. A. (T.) zerchei; 61 - 64. A. (T.) nubis. (Habitus: 53, 57, 61; Pronotum: 54, 58, 62; Mesoventrite and metaventrite: 55, 59, 63; Inter coxal process of mesoventrite: 56, 60, 64)." httpUri="https://zenodo.org/record/282603/files/figure.png" pageId="28" pageNumber="29">Figs. 53, 57, 61</figureCitation>
): narrowly elongated extremely, and almost subparallel sided; whole length varying from 2.8 to 5.0 mm, normally around
<quantity id="4C8F9B2B0830FFEDFD0BFC582B3938D7" box="[692,828,942,966]" metricMagnitude="-3" metricUnit="m" metricValue="3.5999999999999996" metricValueMax="3.9" metricValueMin="3.3" pageId="28" pageNumber="29" unit="mm" value="3.5999999999999996" valueMax="3.9" valueMin="3.3">3.3-3.9 mm</quantity>
; dorsal surface shining to less shining, covered with coarse and distinct punctures. Surface of head and pronotum partly covered with coarse and short but thick setae.
<emphasis id="B903EADC0830FFEDFF28FC0028EE3F1E" bold="true" box="[151,235,1014,1039]" pageId="28" pageNumber="29">Colour</emphasis>
(
<figureCitation id="134C2A4B0830FFEDFF42FC0029BF3F1F" box="[253,442,1014,1038]" captionStart="FIGURES 53 64" captionStartId="26.[151,264,1753,1776]" captionTargetBox="[151,1433,193,1728]" captionTargetId="figure@26.[151,1436,193,1732]" captionTargetPageId="26" captionText="FIGURES 53 64. Japanese species of Aleochara (Triochara). 53 - 56. Aleochara (Triochara) trisulcata; 57 - 60. A. (T.) zerchei; 61 - 64. A. (T.) nubis. (Habitus: 53, 57, 61; Pronotum: 54, 58, 62; Mesoventrite and metaventrite: 55, 59, 63; Inter coxal process of mesoventrite: 56, 60, 64)." httpUri="https://zenodo.org/record/282603/files/figure.png" pageId="28" pageNumber="29">Figs. 53, 57, 61</figureCitation>
): uniformly blackish gray to black sometimes with somewhat lighter colour in elytra.
<emphasis id="B903EADC0830FFEDFF28FBED28D23F22" bold="true" box="[151,215,1051,1075]" pageId="28" pageNumber="29">Head</emphasis>
: rounded subquadrangular (HW/HL ≈1.18), moderately convex above, widest around base; dorsal surface of head with two longitudinal and subparallel deep furrows and connected by a transverse deep furrow or by a somewhat shallow impression. Eyes very small, not clearly protruding laterally.
<emphasis id="B903EADC0830FFEDFB93FB942C9A3F6A" bold="true" box="[1068,1183,1122,1147]" pageId="28" pageNumber="29">Antennae</emphasis>
(
<figureCitation id="134C2A4B0830FFEDFB0FFB942D033F6B" box="[1200,1286,1122,1146]" captionStart="FIGURES 65 70" captionStartId="29.[151,264,1965,1988]" captionTargetBox="[179,1408,270,1932]" captionTargetId="figure@29.[162,1432,253,1954]" captionTargetPageId="29" captionText="FIGURES 65 70. Body parts of Aleochara (Triochara) trisulcata of male. 65. right antenna; 66. labium; 67. maxilla; 68. mentum; 69. labrum; 70. right foreleg in ventral view." httpUri="https://zenodo.org/record/282604/files/figure.png" pageId="28" pageNumber="29">Fig. 65</figureCitation>
): moniliform (similar to filiform); robust, thick and extremely short, clearly shorter than combined length of head and pronotum.
<emphasis id="B903EADC0830FFEDFF28FB5D292A3FD2" bold="true" box="[151,303,1195,1219]" pageId="28" pageNumber="29">Mouth Parts</emphasis>
: mandibles asymmetric, left mandible with small tooth near apex. Clypeus truncate at apex. Labrum (
<figureCitation id="134C2A4B0830FFEDFF20FB3828F13FF7" box="[159,244,1230,1254]" captionStart="FIGURES 65 70" captionStartId="29.[151,264,1965,1988]" captionTargetBox="[179,1408,270,1932]" captionTargetId="figure@29.[162,1432,253,1954]" captionTargetPageId="29" captionText="FIGURES 65 70. Body parts of Aleochara (Triochara) trisulcata of male. 65. right antenna; 66. labium; 67. maxilla; 68. mentum; 69. labrum; 70. right foreleg in ventral view." httpUri="https://zenodo.org/record/282604/files/figure.png" pageId="28" pageNumber="29">Fig. 69</figureCitation>
) transverse, about 1.7 times wider than long; anterior margin slightly emarginated medially; basal half semi-transparent. Mentum (
<figureCitation id="134C2A4B0830FFEDFE72FB042A1B3E1B" box="[461,542,1266,1290]" captionStart="FIGURES 65 70" captionStartId="29.[151,264,1965,1988]" captionTargetBox="[179,1408,270,1932]" captionTargetId="figure@29.[162,1432,253,1954]" captionTargetPageId="29" captionText="FIGURES 65 70. Body parts of Aleochara (Triochara) trisulcata of male. 65. right antenna; 66. labium; 67. maxilla; 68. mentum; 69. labrum; 70. right foreleg in ventral view." httpUri="https://zenodo.org/record/282604/files/figure.png" pageId="28" pageNumber="29">Fig. 68</figureCitation>
) trapezoidal and strongly emarginated transversely along anterior margin, about 3.2 times as wide as long. Labial palpus (
<figureCitation id="134C2A4B0830FFEDFDCAFAE02ACD3E3F" box="[629,712,1302,1326]" captionStart="FIGURES 65 70" captionStartId="29.[151,264,1965,1988]" captionTargetBox="[179,1408,270,1932]" captionTargetId="figure@29.[162,1432,253,1954]" captionTargetPageId="29" captionText="FIGURES 65 70. Body parts of Aleochara (Triochara) trisulcata of male. 65. right antenna; 66. labium; 67. maxilla; 68. mentum; 69. labrum; 70. right foreleg in ventral view." httpUri="https://zenodo.org/record/282604/files/figure.png" pageId="28" pageNumber="29">Fig. 66</figureCitation>
) with segment I prominently thick and longer than segment II; segment III prominently narrower and shorter than II, with minute pseudosegment. Segments of maxillary palpus (
<figureCitation id="134C2A4B0830FFEDFF20FAA828F23E67" box="[159,247,1374,1398]" captionStart="FIGURES 65 70" captionStartId="29.[151,264,1965,1988]" captionTargetBox="[179,1408,270,1932]" captionTargetId="figure@29.[162,1432,253,1954]" captionTargetPageId="29" captionText="FIGURES 65 70. Body parts of Aleochara (Triochara) trisulcata of male. 65. right antenna; 66. labium; 67. maxilla; 68. mentum; 69. labrum; 70. right foreleg in ventral view." httpUri="https://zenodo.org/record/282604/files/figure.png" pageId="28" pageNumber="29">Fig. 67</figureCitation>
) robust, especially, segment II and III; pseudosegment prominent; segment II same length as III; pseudosegment prominent; lacinia with numerous hairs and with about 13 very thick spines pectinately.
<emphasis id="B903EADC0830FFEDFA83FA752D913E8A" bold="true" box="[1340,1428,1411,1435]" pageId="28" pageNumber="29">Thorax</emphasis>
: pronotum (
<figureCitation id="134C2A4B0830FFEDFEA9FA5029D43EAF" box="[278,465,1446,1470]" captionStart="FIGURES 53 64" captionStartId="26.[151,264,1753,1776]" captionTargetBox="[151,1433,193,1728]" captionTargetId="figure@26.[151,1436,193,1732]" captionTargetPageId="26" captionText="FIGURES 53 64. Japanese species of Aleochara (Triochara). 53 - 56. Aleochara (Triochara) trisulcata; 57 - 60. A. (T.) zerchei; 61 - 64. A. (T.) nubis. (Habitus: 53, 57, 61; Pronotum: 54, 58, 62; Mesoventrite and metaventrite: 55, 59, 63; Inter coxal process of mesoventrite: 56, 60, 64)." httpUri="https://zenodo.org/record/282603/files/figure.png" pageId="28" pageNumber="29">Figs. 54, 58, 62</figureCitation>
) largely octagonal to oblong (PW/PL ≈1.18) with prominent furrows (usually three longitudinal sulci) along midline; slightly longer than head length (PL/HL ≈1.29), slightly broader than head (PW/ HW ≈1.29). Hypomera visible in lateral view. Mesocoxae narrowly separated (
<figureCitation id="134C2A4B0830FFEDFBACFA182CCD3D17" box="[1043,1224,1518,1542]" captionStart="FIGURES 53 64" captionStartId="26.[151,264,1753,1776]" captionTargetBox="[151,1433,193,1728]" captionTargetId="figure@26.[151,1436,193,1732]" captionTargetPageId="26" captionText="FIGURES 53 64. Japanese species of Aleochara (Triochara). 53 - 56. Aleochara (Triochara) trisulcata; 57 - 60. A. (T.) zerchei; 61 - 64. A. (T.) nubis. (Habitus: 53, 57, 61; Pronotum: 54, 58, 62; Mesoventrite and metaventrite: 55, 59, 63; Inter coxal process of mesoventrite: 56, 60, 64)." httpUri="https://zenodo.org/record/282603/files/figure.png" pageId="28" pageNumber="29">Figs. 55, 59, 63</figureCitation>
). Mesoventrite not carinate at all (
<figureCitation id="134C2A4B0830FFEDFEFAF9E42A7A3D3B" box="[325,639,1554,1578]" captionStart="FIGURES 53 64" captionStartId="26.[151,264,1753,1776]" captionTargetBox="[151,1433,193,1728]" captionTargetId="figure@26.[151,1436,193,1732]" captionTargetPageId="26" captionText="FIGURES 53 64. Japanese species of Aleochara (Triochara). 53 - 56. Aleochara (Triochara) trisulcata; 57 - 60. A. (T.) zerchei; 61 - 64. A. (T.) nubis. (Habitus: 53, 57, 61; Pronotum: 54, 58, 62; Mesoventrite and metaventrite: 55, 59, 63; Inter coxal process of mesoventrite: 56, 60, 64)." httpUri="https://zenodo.org/record/282603/files/figure.png" pageId="28" pageNumber="29">Figs. 5556, 5960, 6364</figureCitation>
).
<emphasis id="B903EADC0830FFEDFD21F9E52AEF3D3A" bold="true" box="[670,746,1555,1579]" pageId="28" pageNumber="29">Elytra</emphasis>
: widened toward posterior margin, much broader than long (EW/EL ≈1.44), and slightly wider than pronotum (EW/PW ≈1.16); posterior margin of each elytra nearly truncate but gently rounded toward posterior margins; surface rugosely punctured both hexagonal microsculptures and prominent distinct punctures in the species (
<taxonomicName id="4C774D4D0830FFEDFD2CF9882AF73D87" box="[659,754,1662,1686]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="nubis">
<emphasis id="B903EADC0830FFEDFD2CF9882AF73D87" box="[659,754,1662,1686]" italics="true" pageId="28" pageNumber="29">A. nubis</emphasis>
</taxonomicName>
); entire surface densely covered with brown setae; anterior margin with a relatively long blackish bristle.
<emphasis id="B903EADC0830FFEDFD16F9552AE43DAA" bold="true" box="[681,737,1699,1723]" pageId="28" pageNumber="29">Legs</emphasis>
(e. g.,
<figureCitation id="134C2A4B0830FFEDFC8AF9542B893DAB" box="[821,908,1698,1722]" captionStart="FIGURES 65 70" captionStartId="29.[151,264,1965,1988]" captionTargetBox="[179,1408,270,1932]" captionTargetId="figure@29.[162,1432,253,1954]" captionTargetPageId="29" captionText="FIGURES 65 70. Body parts of Aleochara (Triochara) trisulcata of male. 65. right antenna; 66. labium; 67. maxilla; 68. mentum; 69. labrum; 70. right foreleg in ventral view." httpUri="https://zenodo.org/record/282604/files/figure.png" pageId="28" pageNumber="29">Fig. 70</figureCitation>
): short, thick, robust and with numerous thick spines on surface, especially on fore and mid tibia; tarsi quite short, hindtarsi extremely short (hind tarsal length/ mid tarsal length ≈1.25).
<emphasis id="B903EADC0830FFEDFE05F91C2A3F3C12" bold="true" box="[442,570,1770,1795]" pageId="28" pageNumber="29">
<collectingCountry id="F360765E0830FFEDFE05F91C29F23C12" box="[442,503,1770,1795]" name="India" pageId="28" pageNumber="29">Hind</collectingCountry>
wing
</emphasis>
: entire; veins weakly sclerotized and very obscure; posterior margin with a row of numerous white hairs.
<emphasis id="B903EADC0830FFEDFE50F8F82A613C36" bold="true" box="[495,612,1806,1831]" pageId="28" pageNumber="29">Abdomen</emphasis>
(
<figureCitation id="134C2A4B0830FFEDFDCAF8F82B283C37" box="[629,813,1806,1830]" captionStart="FIGURES 53 64" captionStartId="26.[151,264,1753,1776]" captionTargetBox="[151,1433,193,1728]" captionTargetId="figure@26.[151,1436,193,1732]" captionTargetPageId="26" captionText="FIGURES 53 64. Japanese species of Aleochara (Triochara). 53 - 56. Aleochara (Triochara) trisulcata; 57 - 60. A. (T.) zerchei; 61 - 64. A. (T.) nubis. (Habitus: 53, 57, 61; Pronotum: 54, 58, 62; Mesoventrite and metaventrite: 55, 59, 63; Inter coxal process of mesoventrite: 56, 60, 64)." httpUri="https://zenodo.org/record/282603/files/figure.png" pageId="28" pageNumber="29">Figs. 53, 57, 61</figureCitation>
): narrowly elongated and almost pararell sided; tergite III-VI impressed weakly and transversely at base. Posterior margin of tergite VIII (
<figureCitation id="134C2A4B0830FFEDFBE4F8C42CE33C5B" box="[1115,1254,1842,1866]" captionStart="FIGURES 71 78" captionStartId="33.[151,264,1821,1844]" captionTargetBox="[152,1426,187,1820]" captionTargetId="figure@33.[151,1436,141,1822]" captionTargetPageId="33" captionText="FIGURES 71 78. Terminalia of Aleochara (Triochara) trisulcata. 71. tergite VIII of male; 72. tergite VIII of female; 73. sternite VIII of male; 74. sternite VIII of female; 75. male genitalia: median lobe of aedeagus in lateral view; 76. male genitalia: median lobe of aedeagus in ventral view; 77. apical lobe of paramerite, lateral view; 78. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282605/files/figure.png" pageId="28" pageNumber="29">Figs. 7172</figureCitation>
), with a row of thick and short several sensory setae or a row of thin and long setae (
<figureCitation id="134C2A4B0830FFEDFC24F8A02C273C7F" box="[923,1058,1878,1902]" captionStart="FIGURES 79 85" captionStartId="36.[151,264,1915,1938]" captionTargetBox="[178,1416,289,1854]" captionTargetId="figure@36.[151,1436,265,1894]" captionTargetPageId="36" captionText="FIGURES 79 85. Terminalia of Aleochara (Triochara) zerchei. 79. tergite VIII of male; 80. tergite VIII of female; 81. sternite VIII of male; 82. sternite VIII of female; 83. male genitalia: median lobe of aedeagus in lateral view; 84. male genitalia: median lobe of aedeagus in ventral view; 85. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282606/files/figure.png" pageId="28" pageNumber="29">Figs. 7980</figureCitation>
,
<figureCitation id="134C2A4B0830FFEDFB91F8A02C7C3C7F" box="[1070,1145,1878,1902]" captionStart="FIGURES 86 92" captionStartId="41.[151,264,1942,1965]" captionTargetBox="[153,1404,302,1941]" captionTargetId="figure@41.[151,1436,265,1951]" captionTargetPageId="41" captionText="FIGURES 86 92. Terminalia of Aleochara (Triochara) nubis. 86. tergite VIII of male; 87. tergite VIII of female; 88. sternite VIII of male; 89. sternite VIII of female; 90. male genitalia: median lobe of aedeagus in lateral view; 91. male genitalia: median lobe of aedeagus in ventral view; 92. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282607/files/figure.png" pageId="28" pageNumber="29">8687</figureCitation>
).
</paragraph>
<paragraph id="8BC836CE0830FFECFF78F88D2D923BC5" blockId="28.[151,1437,906,2011]" lastBlockId="29.[151,1436,151,212]" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
[
<emphasis id="B903EADC0830FFEDFF6FF88D29093C82" bold="true" box="[208,268,1915,1939]" pageId="28" pageNumber="29">Male</emphasis>
]: posterior margin of sternite VIII (
<figureCitation id="134C2A4B0830FFEDFD29F88C2AF73C83" box="[662,754,1914,1938]" captionStart="FIGURES 71 78" captionStartId="33.[151,264,1821,1844]" captionTargetBox="[152,1426,187,1820]" captionTargetId="figure@33.[151,1436,141,1822]" captionTargetPageId="33" captionText="FIGURES 71 78. Terminalia of Aleochara (Triochara) trisulcata. 71. tergite VIII of male; 72. tergite VIII of female; 73. sternite VIII of male; 74. sternite VIII of female; 75. male genitalia: median lobe of aedeagus in lateral view; 76. male genitalia: median lobe of aedeagus in ventral view; 77. apical lobe of paramerite, lateral view; 78. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282605/files/figure.png" pageId="28" pageNumber="29">Figs. 73</figureCitation>
,
<figureCitation id="134C2A4B0830FFEDFD41F88C2B1E3C83" box="[766,795,1914,1938]" captionStart="FIGURES 79 85" captionStartId="36.[151,264,1915,1938]" captionTargetBox="[178,1416,289,1854]" captionTargetId="figure@36.[151,1436,265,1894]" captionTargetPageId="36" captionText="FIGURES 79 85. Terminalia of Aleochara (Triochara) zerchei. 79. tergite VIII of male; 80. tergite VIII of female; 81. sternite VIII of male; 82. sternite VIII of female; 83. male genitalia: median lobe of aedeagus in lateral view; 84. male genitalia: median lobe of aedeagus in ventral view; 85. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282606/files/figure.png" pageId="28" pageNumber="29">81</figureCitation>
,
<figureCitation id="134C2A4B0830FFEDFC98F88C2B463C83" box="[807,835,1914,1938]" captionStart="FIGURES 86 92" captionStartId="41.[151,264,1942,1965]" captionTargetBox="[153,1404,302,1941]" captionTargetId="figure@41.[151,1436,265,1951]" captionTargetPageId="41" captionText="FIGURES 86 92. Terminalia of Aleochara (Triochara) nubis. 86. tergite VIII of male; 87. tergite VIII of female; 88. sternite VIII of male; 89. sternite VIII of female; 90. male genitalia: median lobe of aedeagus in lateral view; 91. male genitalia: median lobe of aedeagus in ventral view; 92. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282607/files/figure.png" pageId="28" pageNumber="29">88</figureCitation>
) not pointing strongly, and its shape not greatly varies among species. Median lobe (
<figureCitation id="134C2A4B0830FFEDFE4CF8682A513CA7" box="[499,596,1950,1974]" captionStart="FIGURES 71 78" captionStartId="33.[151,264,1821,1844]" captionTargetBox="[152,1426,187,1820]" captionTargetId="figure@33.[151,1436,141,1822]" captionTargetPageId="33" captionText="FIGURES 71 78. Terminalia of Aleochara (Triochara) trisulcata. 71. tergite VIII of male; 72. tergite VIII of female; 73. sternite VIII of male; 74. sternite VIII of female; 75. male genitalia: median lobe of aedeagus in lateral view; 76. male genitalia: median lobe of aedeagus in ventral view; 77. apical lobe of paramerite, lateral view; 78. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282605/files/figure.png" pageId="28" pageNumber="29">Figs. 75</figureCitation>
,
<figureCitation id="134C2A4B0830FFEDFDDBF8682A873CA7" box="[612,642,1950,1974]" captionStart="FIGURES 79 85" captionStartId="36.[151,264,1915,1938]" captionTargetBox="[178,1416,289,1854]" captionTargetId="figure@36.[151,1436,265,1894]" captionTargetPageId="36" captionText="FIGURES 79 85. Terminalia of Aleochara (Triochara) zerchei. 79. tergite VIII of male; 80. tergite VIII of female; 81. sternite VIII of male; 82. sternite VIII of female; 83. male genitalia: median lobe of aedeagus in lateral view; 84. male genitalia: median lobe of aedeagus in ventral view; 85. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282606/files/figure.png" pageId="28" pageNumber="29">83</figureCitation>
,
<figureCitation id="134C2A4B0830FFEDFD2CF8682AB53CA7" box="[659,688,1950,1974]" captionStart="FIGURES 86 92" captionStartId="41.[151,264,1942,1965]" captionTargetBox="[153,1404,302,1941]" captionTargetId="figure@41.[151,1436,265,1951]" captionTargetPageId="41" captionText="FIGURES 86 92. Terminalia of Aleochara (Triochara) nubis. 86. tergite VIII of male; 87. tergite VIII of female; 88. sternite VIII of male; 89. sternite VIII of female; 90. male genitalia: median lobe of aedeagus in lateral view; 91. male genitalia: median lobe of aedeagus in ventral view; 92. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282607/files/figure.png" pageId="28" pageNumber="29">90</figureCitation>
) compactly elongated. Median lobe of aedeagus with a pair of subapico-ventral projections (
<figureCitation id="134C2A4B0830FFEDFE5AF8342A403CCB" box="[485,581,1986,2010]" captionStart="FIGURES 71 78" captionStartId="33.[151,264,1821,1844]" captionTargetBox="[152,1426,187,1820]" captionTargetId="figure@33.[151,1436,141,1822]" captionTargetPageId="33" captionText="FIGURES 71 78. Terminalia of Aleochara (Triochara) trisulcata. 71. tergite VIII of male; 72. tergite VIII of female; 73. sternite VIII of male; 74. sternite VIII of female; 75. male genitalia: median lobe of aedeagus in lateral view; 76. male genitalia: median lobe of aedeagus in ventral view; 77. apical lobe of paramerite, lateral view; 78. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282605/files/figure.png" pageId="28" pageNumber="29">Figs. 75</figureCitation>
,
<figureCitation id="134C2A4B0830FFEDFDEBF8342A773CCB" box="[596,626,1986,2010]" captionStart="FIGURES 79 85" captionStartId="36.[151,264,1915,1938]" captionTargetBox="[178,1416,289,1854]" captionTargetId="figure@36.[151,1436,265,1894]" captionTargetPageId="36" captionText="FIGURES 79 85. Terminalia of Aleochara (Triochara) zerchei. 79. tergite VIII of male; 80. tergite VIII of female; 81. sternite VIII of male; 82. sternite VIII of female; 83. male genitalia: median lobe of aedeagus in lateral view; 84. male genitalia: median lobe of aedeagus in ventral view; 85. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282606/files/figure.png" pageId="28" pageNumber="29">83</figureCitation>
,
<figureCitation id="134C2A4B0830FFEDFD3DF8342A9A3CCB" box="[642,671,1986,2010]" captionStart="FIGURES 86 92" captionStartId="41.[151,264,1942,1965]" captionTargetBox="[153,1404,302,1941]" captionTargetId="figure@41.[151,1436,265,1951]" captionTargetPageId="41" captionText="FIGURES 86 92. Terminalia of Aleochara (Triochara) nubis. 86. tergite VIII of male; 87. tergite VIII of female; 88. sternite VIII of male; 89. sternite VIII of female; 90. male genitalia: median lobe of aedeagus in lateral view; 91. male genitalia: median lobe of aedeagus in ventral view; 92. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282607/files/figure.png" pageId="28" pageNumber="29">90</figureCitation>
) as well as
<taxonomicName id="4C774D4D0830FFEDFC8EF8352BAD3CCA" box="[817,936,1987,2011]" class="Insecta" family="Staphylinidae" genus="Emplenota" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B903EADC0830FFEDFC8EF8352BAD3CCA" box="[817,936,1987,2011]" italics="true" pageId="28" pageNumber="29">Emplenota</emphasis>
</taxonomicName>
species; shape of these projections varying greatly among species. Median lobe of aedeagus (
<figureCitation id="134C2A4B0831FFECFD57FF612B743BBE" box="[744,881,151,175]" captionStart="FIGURES 71 78" captionStartId="33.[151,264,1821,1844]" captionTargetBox="[152,1426,187,1820]" captionTargetId="figure@33.[151,1436,141,1822]" captionTargetPageId="33" captionText="FIGURES 71 78. Terminalia of Aleochara (Triochara) trisulcata. 71. tergite VIII of male; 72. tergite VIII of female; 73. sternite VIII of male; 74. sternite VIII of female; 75. male genitalia: median lobe of aedeagus in lateral view; 76. male genitalia: median lobe of aedeagus in ventral view; 77. apical lobe of paramerite, lateral view; 78. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282605/files/figure.png" pageId="29" pageNumber="30">Figs. 7576</figureCitation>
,
<figureCitation id="134C2A4B0831FFECFC3DFF612BCD3BBE" box="[898,968,151,175]" captionStart="FIGURES 79 85" captionStartId="36.[151,264,1915,1938]" captionTargetBox="[178,1416,289,1854]" captionTargetId="figure@36.[151,1436,265,1894]" captionTargetPageId="36" captionText="FIGURES 79 85. Terminalia of Aleochara (Triochara) zerchei. 79. tergite VIII of male; 80. tergite VIII of female; 81. sternite VIII of male; 82. sternite VIII of female; 83. male genitalia: median lobe of aedeagus in lateral view; 84. male genitalia: median lobe of aedeagus in ventral view; 85. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282606/files/figure.png" pageId="29" pageNumber="30">8384</figureCitation>
,
<figureCitation id="134C2A4B0831FFECFC66FF612C1A3BBE" box="[985,1055,151,175]" captionStart="FIGURES 86 92" captionStartId="41.[151,264,1942,1965]" captionTargetBox="[153,1404,302,1941]" captionTargetId="figure@41.[151,1436,265,1951]" captionTargetPageId="41" captionText="FIGURES 86 92. Terminalia of Aleochara (Triochara) nubis. 86. tergite VIII of male; 87. tergite VIII of female; 88. sternite VIII of male; 89. sternite VIII of female; 90. male genitalia: median lobe of aedeagus in lateral view; 91. male genitalia: median lobe of aedeagus in ventral view; 92. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282607/files/figure.png" pageId="29" pageNumber="30">9091</figureCitation>
) with rather short but prominent flagellum, as long as median lobe of aedeagus. Apical lobe of paramerite (
<figureCitation id="134C2A4B0831FFECFC06FF4A2C0C3BC5" box="[953,1033,188,212]" captionStart="FIGURES 71 78" captionStartId="33.[151,264,1821,1844]" captionTargetBox="[152,1426,187,1820]" captionTargetId="figure@33.[151,1436,141,1822]" captionTargetPageId="33" captionText="FIGURES 71 78. Terminalia of Aleochara (Triochara) trisulcata. 71. tergite VIII of male; 72. tergite VIII of female; 73. sternite VIII of male; 74. sternite VIII of female; 75. male genitalia: median lobe of aedeagus in lateral view; 76. male genitalia: median lobe of aedeagus in ventral view; 77. apical lobe of paramerite, lateral view; 78. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282605/files/figure.png" pageId="29" pageNumber="30">Fig. 77</figureCitation>
) narrowly elongated with four setae.
</paragraph>
<caption id="DF0866460831FFECFF28F85B2ABF3CF7" httpUri="https://zenodo.org/record/282604/files/figure.png" pageId="29" pageNumber="30" targetBox="[179,1408,270,1932]" targetPageId="29">
<paragraph id="8BC836CE0831FFECFF28F85B2ABF3CF7" blockId="29.[151,1437,1965,2022]" pageId="29" pageNumber="30">
<emphasis id="B903EADC0831FFECFF28F85B29523CD3" bold="true" box="[151,343,1965,1988]" pageId="29" pageNumber="30">FIGURES 6570.</emphasis>
Body parts of
<taxonomicName id="4C774D4D0831FFECFE46F85B2B483CD2" box="[505,845,1965,1987]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="trisulcata" subGenus="Triochara">
<emphasis id="B903EADC0831FFECFE46F85B2A673CD2" box="[505,610,1965,1987]" italics="true" pageId="29" pageNumber="30">Aleochara</emphasis>
(
<emphasis id="B903EADC0831FFECFDCBF85B2ADD3CD2" box="[628,728,1965,1987]" italics="true" pageId="29" pageNumber="30">Triochara</emphasis>
)
<emphasis id="B903EADC0831FFECFD53F85B2B483CD2" box="[748,845,1965,1987]" italics="true" pageId="29" pageNumber="30">trisulcata</emphasis>
</taxonomicName>
of male. 65. right antenna; 66. labium; 67. maxilla; 68. mentum; 69. labrum; 70. right foreleg in ventral view.
</paragraph>
</caption>
<paragraph id="8BC836CE0832FFEFFF78FF6E29983AB3" blockId="30.[151,1436,151,1077]" pageId="30" pageNumber="31">
[
<emphasis id="B903EADC0832FFEFFF6FFF6E29223BA1" bold="true" box="[208,295,152,176]" pageId="30" pageNumber="31">Female</emphasis>
]: tergite VIII (
<figureCitation id="134C2A4B0832FFEFFE6BFF612A303BBE" box="[468,565,151,175]" captionStart="FIGURES 71 78" captionStartId="33.[151,264,1821,1844]" captionTargetBox="[152,1426,187,1820]" captionTargetId="figure@33.[151,1436,141,1822]" captionTargetPageId="33" captionText="FIGURES 71 78. Terminalia of Aleochara (Triochara) trisulcata. 71. tergite VIII of male; 72. tergite VIII of female; 73. sternite VIII of male; 74. sternite VIII of female; 75. male genitalia: median lobe of aedeagus in lateral view; 76. male genitalia: median lobe of aedeagus in ventral view; 77. apical lobe of paramerite, lateral view; 78. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282605/files/figure.png" pageId="30" pageNumber="31">Figs. 72</figureCitation>
,
<figureCitation id="134C2A4B0832FFEFFDFCFF612A643BBE" box="[579,609,151,175]" captionStart="FIGURES 79 85" captionStartId="36.[151,264,1915,1938]" captionTargetBox="[178,1416,289,1854]" captionTargetId="figure@36.[151,1436,265,1894]" captionTargetPageId="36" captionText="FIGURES 79 85. Terminalia of Aleochara (Triochara) zerchei. 79. tergite VIII of male; 80. tergite VIII of female; 81. sternite VIII of male; 82. sternite VIII of female; 83. male genitalia: median lobe of aedeagus in lateral view; 84. male genitalia: median lobe of aedeagus in ventral view; 85. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282606/files/figure.png" pageId="30" pageNumber="31">80</figureCitation>
,
<figureCitation id="134C2A4B0832FFEFFDD0FF612A893BBE" box="[623,652,151,175]" captionStart="FIGURES 86 92" captionStartId="41.[151,264,1942,1965]" captionTargetBox="[153,1404,302,1941]" captionTargetId="figure@41.[151,1436,265,1951]" captionTargetPageId="41" captionText="FIGURES 86 92. Terminalia of Aleochara (Triochara) nubis. 86. tergite VIII of male; 87. tergite VIII of female; 88. sternite VIII of male; 89. sternite VIII of female; 90. male genitalia: median lobe of aedeagus in lateral view; 91. male genitalia: median lobe of aedeagus in ventral view; 92. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282607/files/figure.png" pageId="30" pageNumber="31">87</figureCitation>
) similar to that of male. Sternite VIII (
<figureCitation id="134C2A4B0832FFEFFBEDFF612CB73BBE" box="[1106,1202,151,175]" captionStart="FIGURES 71 78" captionStartId="33.[151,264,1821,1844]" captionTargetBox="[152,1426,187,1820]" captionTargetId="figure@33.[151,1436,141,1822]" captionTargetPageId="33" captionText="FIGURES 71 78. Terminalia of Aleochara (Triochara) trisulcata. 71. tergite VIII of male; 72. tergite VIII of female; 73. sternite VIII of male; 74. sternite VIII of female; 75. male genitalia: median lobe of aedeagus in lateral view; 76. male genitalia: median lobe of aedeagus in ventral view; 77. apical lobe of paramerite, lateral view; 78. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282605/files/figure.png" pageId="30" pageNumber="31">Figs. 74</figureCitation>
,
<figureCitation id="134C2A4B0832FFEFFB7EFF612CE53BBE" box="[1217,1248,151,175]" captionStart="FIGURES 79 85" captionStartId="36.[151,264,1915,1938]" captionTargetBox="[178,1416,289,1854]" captionTargetId="figure@36.[151,1436,265,1894]" captionTargetPageId="36" captionText="FIGURES 79 85. Terminalia of Aleochara (Triochara) zerchei. 79. tergite VIII of male; 80. tergite VIII of female; 81. sternite VIII of male; 82. sternite VIII of female; 83. male genitalia: median lobe of aedeagus in lateral view; 84. male genitalia: median lobe of aedeagus in ventral view; 85. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282606/files/figure.png" pageId="30" pageNumber="31">82</figureCitation>
,
<figureCitation id="134C2A4B0832FFEFFB52FF612D0F3BBE" box="[1261,1290,151,175]" captionStart="FIGURES 86 92" captionStartId="41.[151,264,1942,1965]" captionTargetBox="[153,1404,302,1941]" captionTargetId="figure@41.[151,1436,265,1951]" captionTargetPageId="41" captionText="FIGURES 86 92. Terminalia of Aleochara (Triochara) nubis. 86. tergite VIII of male; 87. tergite VIII of female; 88. sternite VIII of male; 89. sternite VIII of female; 90. male genitalia: median lobe of aedeagus in lateral view; 91. male genitalia: median lobe of aedeagus in ventral view; 92. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282607/files/figure.png" pageId="30" pageNumber="31">86</figureCitation>
) having less pointed apex than male, but sometimes very similar to that of male. Spermatheca (
<figureCitation id="134C2A4B0832FFEFFBFAFF4C2CA13BC3" box="[1093,1188,186,210]" captionStart="FIGURES 71 78" captionStartId="33.[151,264,1821,1844]" captionTargetBox="[152,1426,187,1820]" captionTargetId="figure@33.[151,1436,141,1822]" captionTargetPageId="33" captionText="FIGURES 71 78. Terminalia of Aleochara (Triochara) trisulcata. 71. tergite VIII of male; 72. tergite VIII of female; 73. sternite VIII of male; 74. sternite VIII of female; 75. male genitalia: median lobe of aedeagus in lateral view; 76. male genitalia: median lobe of aedeagus in ventral view; 77. apical lobe of paramerite, lateral view; 78. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282605/files/figure.png" pageId="30" pageNumber="31">Figs. 78</figureCitation>
,
<figureCitation id="134C2A4B0832FFEFFB0EFF4C2CCA3BC3" box="[1201,1231,186,210]" captionStart="FIGURES 79 85" captionStartId="36.[151,264,1915,1938]" captionTargetBox="[178,1416,289,1854]" captionTargetId="figure@36.[151,1436,265,1894]" captionTargetPageId="36" captionText="FIGURES 79 85. Terminalia of Aleochara (Triochara) zerchei. 79. tergite VIII of male; 80. tergite VIII of female; 81. sternite VIII of male; 82. sternite VIII of female; 83. male genitalia: median lobe of aedeagus in lateral view; 84. male genitalia: median lobe of aedeagus in ventral view; 85. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282606/files/figure.png" pageId="30" pageNumber="31">85</figureCitation>
,
<figureCitation id="134C2A4B0832FFEFFB63FF4C2CFC3BC3" box="[1244,1273,186,210]" captionStart="FIGURES 86 92" captionStartId="41.[151,264,1942,1965]" captionTargetBox="[153,1404,302,1941]" captionTargetId="figure@41.[151,1436,265,1951]" captionTargetPageId="41" captionText="FIGURES 86 92. Terminalia of Aleochara (Triochara) nubis. 86. tergite VIII of male; 87. tergite VIII of female; 88. sternite VIII of male; 89. sternite VIII of female; 90. male genitalia: median lobe of aedeagus in lateral view; 91. male genitalia: median lobe of aedeagus in ventral view; 92. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282607/files/figure.png" pageId="30" pageNumber="31">92</figureCitation>
) with shallow apical invagination of spermatheca (ai); head (sh) large, moderately swollen, less or longer than twice length of apical portion of stem (sa); spermathecal neck (sn) very short and united with (sh); without a distinct collar in spermatheca (different from
<taxonomicName id="4C774D4D0832FFEFFE63FED52A503A2A" box="[476,597,291,315]" class="Insecta" family="Staphylinidae" genus="Emplenota" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B903EADC0832FFEFFE63FED52A503A2A" box="[476,597,291,315]" italics="true" pageId="30" pageNumber="31">Emplenota</emphasis>
</taxonomicName>
:
<figureCitation id="134C2A4B0832FFEFFDD9FED42ABE3A2B" box="[614,699,290,314]" captionStart="FIGURES 14 22" captionStartId="9.[151,264,1909,1932]" captionTargetBox="[153,1426,203,1908]" captionTargetId="figure@9.[151,1436,203,1924]" captionTargetPageId="9" captionText="FIGURES 14 22. Terminalia of Aleochara (Emplenota) fucicola. 14. tergite VIII of male; 15. tergite VIII of female; 16. sternite VIII of male; 17. sternite VIII of female; 18. male genitalia: median lobe of aedeagus in lateral view; 19. male genitalia: median lobe of aedeagus in ventral view; 20. apical lobe of median lobe in ventral aspect; 21. apical lobe of paramerite, lateral view; 22. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282598/files/figure.png" pageId="30" pageNumber="31">Fig. 22</figureCitation>
); basal portion of stem (sb) narrow and distorted, and with short and erect sclerotized portion of spermathecal stem (ss); membraneous portion of spermathecal duct (sm) moderate in length; inner wall of (sh) and (sa) moderately striate; each part of spermatheca except for (sm) entirely and moderately sclerotized.
</paragraph>
</subSubSection>
<subSubSection id="C36D65450832FFEFFF78FE5B2B813853" pageId="30" pageNumber="31" type="diagnosis">
<paragraph id="8BC836CE0832FFEFFF78FE5B2B813853" blockId="30.[151,1436,151,1077]" pageId="30" pageNumber="31">
<emphasis id="B903EADC0832FFEFFF78FE5B29443AD7" bold="true" box="[199,321,429,454]" pageId="30" pageNumber="31">Diagnosis.</emphasis>
The distinct subgenus
<taxonomicName id="4C774D4D0832FFEFFDF6FE5B2B333AD4" authority="Bernhauer" authorityName="Bernhauer" box="[585,822,429,453]" class="Insecta" family="Staphylinidae" genus="Triochara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B903EADC0832FFEFFDF6FE5B2ABC3AD4" box="[585,697,429,453]" italics="true" pageId="30" pageNumber="31">Triochara</emphasis>
Bernhauer
</taxonomicName>
is somewhat similar to the members of the subfamily
<taxonomicName id="4C774D4D0832FFEFFF28FE3929C03AF6" box="[151,453,463,487]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="subFamily" subFamily="Oxytelinae">Oxytelinae (Staphylinidae)</taxonomicName>
rather than the members of
<taxonomicName id="4C774D4D0832FFEFFD42FE392B973AF6" box="[765,914,463,487]" class="Insecta" family="Staphylinidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="family">Aleocharinae</taxonomicName>
by having furrows on head and pronotum, and by heavily sclerotized, subparallel-sided body. This subgenus is therefore easily discriminated from other littoral staphylinids by following character states: antennae (e. g.,
<figureCitation id="134C2A4B0832FFEFFC9AFDE32B70393C" box="[805,885,533,557]" captionStart="FIGURES 65 70" captionStartId="29.[151,264,1965,1988]" captionTargetBox="[179,1408,270,1932]" captionTargetId="figure@29.[162,1432,253,1954]" captionTargetPageId="29" captionText="FIGURES 65 70. Body parts of Aleochara (Triochara) trisulcata of male. 65. right antenna; 66. labium; 67. maxilla; 68. mentum; 69. labrum; 70. right foreleg in ventral view." httpUri="https://zenodo.org/record/282604/files/figure.png" pageId="30" pageNumber="31">Fig. 65</figureCitation>
) very short, almost moniliform, and quite robust; dorsal surface of head (
<figureCitation id="134C2A4B0832FFEFFE1AFDC12A5E395E" box="[421,603,567,591]" captionStart="FIGURES 53 64" captionStartId="26.[151,264,1753,1776]" captionTargetBox="[151,1433,193,1728]" captionTargetId="figure@26.[151,1436,193,1732]" captionTargetPageId="26" captionText="FIGURES 53 64. Japanese species of Aleochara (Triochara). 53 - 56. Aleochara (Triochara) trisulcata; 57 - 60. A. (T.) zerchei; 61 - 64. A. (T.) nubis. (Habitus: 53, 57, 61; Pronotum: 54, 58, 62; Mesoventrite and metaventrite: 55, 59, 63; Inter coxal process of mesoventrite: 56, 60, 64)." httpUri="https://zenodo.org/record/282603/files/figure.png" pageId="30" pageNumber="31">Figs. 53, 57, 61</figureCitation>
) with two deep furrows along midline; pronotum (
<figureCitation id="134C2A4B0832FFEFFB1CFDC12D5C395E" box="[1187,1369,567,591]" captionStart="FIGURES 53 64" captionStartId="26.[151,264,1753,1776]" captionTargetBox="[151,1433,193,1728]" captionTargetId="figure@26.[151,1436,193,1732]" captionTargetPageId="26" captionText="FIGURES 53 64. Japanese species of Aleochara (Triochara). 53 - 56. Aleochara (Triochara) trisulcata; 57 - 60. A. (T.) zerchei; 61 - 64. A. (T.) nubis. (Habitus: 53, 57, 61; Pronotum: 54, 58, 62; Mesoventrite and metaventrite: 55, 59, 63; Inter coxal process of mesoventrite: 56, 60, 64)." httpUri="https://zenodo.org/record/282603/files/figure.png" pageId="30" pageNumber="31">Figs. 54, 58, 62</figureCitation>
) with longitudinal sulci and three-dimensional pattern on dorsal surface; tarsal fomula: 5-5-5; mesoventrite without carina at all (
<figureCitation id="134C2A4B0832FFEFFE95FD8B2A5B3984" box="[298,606,637,661]" captionStart="FIGURES 53 64" captionStartId="26.[151,264,1753,1776]" captionTargetBox="[151,1433,193,1728]" captionTargetId="figure@26.[151,1436,193,1732]" captionTargetPageId="26" captionText="FIGURES 53 64. Japanese species of Aleochara (Triochara). 53 - 56. Aleochara (Triochara) trisulcata; 57 - 60. A. (T.) zerchei; 61 - 64. A. (T.) nubis. (Habitus: 53, 57, 61; Pronotum: 54, 58, 62; Mesoventrite and metaventrite: 55, 59, 63; Inter coxal process of mesoventrite: 56, 60, 64)." httpUri="https://zenodo.org/record/282603/files/figure.png" pageId="30" pageNumber="31">Figs. 5556, 5960, 6364</figureCitation>
); legs (e. g.,
<figureCitation id="134C2A4B0832FFEFFD51FD8B2B453984" box="[750,832,637,661]" captionStart="FIGURES 65 70" captionStartId="29.[151,264,1965,1988]" captionTargetBox="[179,1408,270,1932]" captionTargetId="figure@29.[162,1432,253,1954]" captionTargetPageId="29" captionText="FIGURES 65 70. Body parts of Aleochara (Triochara) trisulcata of male. 65. right antenna; 66. labium; 67. maxilla; 68. mentum; 69. labrum; 70. right foreleg in ventral view." httpUri="https://zenodo.org/record/282604/files/figure.png" pageId="30" pageNumber="31">Fig. 70</figureCitation>
) short but thick, covered with numerous thick spines, especially on tibia. [
<emphasis id="B903EADC0832FFEFFE2DFD5629CB39A9" bold="true" box="[402,462,672,696]" pageId="30" pageNumber="31">Male</emphasis>
]: sternite VIII (
<figureCitation id="134C2A4B0832FFEFFD28FD692AF839A6" box="[663,765,671,695]" captionStart="FIGURES 71 78" captionStartId="33.[151,264,1821,1844]" captionTargetBox="[152,1426,187,1820]" captionTargetId="figure@33.[151,1436,141,1822]" captionTargetPageId="33" captionText="FIGURES 71 78. Terminalia of Aleochara (Triochara) trisulcata. 71. tergite VIII of male; 72. tergite VIII of female; 73. sternite VIII of male; 74. sternite VIII of female; 75. male genitalia: median lobe of aedeagus in lateral view; 76. male genitalia: median lobe of aedeagus in ventral view; 77. apical lobe of paramerite, lateral view; 78. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282605/files/figure.png" pageId="30" pageNumber="31">Figs. 73</figureCitation>
,
<figureCitation id="134C2A4B0832FFEFFCAEFD692B2A39A6" box="[785,815,671,695]" captionStart="FIGURES 79 85" captionStartId="36.[151,264,1915,1938]" captionTargetBox="[178,1416,289,1854]" captionTargetId="figure@36.[151,1436,265,1894]" captionTargetPageId="36" captionText="FIGURES 79 85. Terminalia of Aleochara (Triochara) zerchei. 79. tergite VIII of male; 80. tergite VIII of female; 81. sternite VIII of male; 82. sternite VIII of female; 83. male genitalia: median lobe of aedeagus in lateral view; 84. male genitalia: median lobe of aedeagus in ventral view; 85. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282606/files/figure.png" pageId="30" pageNumber="31">81</figureCitation>
,
<figureCitation id="134C2A4B0832FFEFFCFCFD692B6539A6" box="[835,864,671,695]" captionStart="FIGURES 86 92" captionStartId="41.[151,264,1942,1965]" captionTargetBox="[153,1404,302,1941]" captionTargetId="figure@41.[151,1436,265,1951]" captionTargetPageId="41" captionText="FIGURES 86 92. Terminalia of Aleochara (Triochara) nubis. 86. tergite VIII of male; 87. tergite VIII of female; 88. sternite VIII of male; 89. sternite VIII of female; 90. male genitalia: median lobe of aedeagus in lateral view; 91. male genitalia: median lobe of aedeagus in ventral view; 92. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282607/files/figure.png" pageId="30" pageNumber="31">88</figureCitation>
) similar shape that of female, and not greatly discriminated from female (
<figureCitation id="134C2A4B0832FFEFFE63FD342A3B39CB" box="[476,574,706,730]" captionStart="FIGURES 71 78" captionStartId="33.[151,264,1821,1844]" captionTargetBox="[152,1426,187,1820]" captionTargetId="figure@33.[151,1436,141,1822]" captionTargetPageId="33" captionText="FIGURES 71 78. Terminalia of Aleochara (Triochara) trisulcata. 71. tergite VIII of male; 72. tergite VIII of female; 73. sternite VIII of male; 74. sternite VIII of female; 75. male genitalia: median lobe of aedeagus in lateral view; 76. male genitalia: median lobe of aedeagus in ventral view; 77. apical lobe of paramerite, lateral view; 78. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282605/files/figure.png" pageId="30" pageNumber="31">Figs. 74</figureCitation>
,
<figureCitation id="134C2A4B0832FFEFFDF1FD342A6939CB" box="[590,620,706,730]" captionStart="FIGURES 79 85" captionStartId="36.[151,264,1915,1938]" captionTargetBox="[178,1416,289,1854]" captionTargetId="figure@36.[151,1436,265,1894]" captionTargetPageId="36" captionText="FIGURES 79 85. Terminalia of Aleochara (Triochara) zerchei. 79. tergite VIII of male; 80. tergite VIII of female; 81. sternite VIII of male; 82. sternite VIII of female; 83. male genitalia: median lobe of aedeagus in lateral view; 84. male genitalia: median lobe of aedeagus in ventral view; 85. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282606/files/figure.png" pageId="30" pageNumber="31">82</figureCitation>
,
<figureCitation id="134C2A4B0832FFEFFDC3FD342A9E39CB" box="[636,667,706,730]" captionStart="FIGURES 86 92" captionStartId="41.[151,264,1942,1965]" captionTargetBox="[153,1404,302,1941]" captionTargetId="figure@41.[151,1436,265,1951]" captionTargetPageId="41" captionText="FIGURES 86 92. Terminalia of Aleochara (Triochara) nubis. 86. tergite VIII of male; 87. tergite VIII of female; 88. sternite VIII of male; 89. sternite VIII of female; 90. male genitalia: median lobe of aedeagus in lateral view; 91. male genitalia: median lobe of aedeagus in ventral view; 92. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282607/files/figure.png" pageId="30" pageNumber="31">89</figureCitation>
;
<taxonomicName id="4C774D4D0832FFEFFD14FD352B2139CA" box="[683,804,707,731]" class="Insecta" family="Staphylinidae" genus="Emplenota" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B903EADC0832FFEFFD14FD352B2139CA" box="[683,804,707,731]" italics="true" pageId="30" pageNumber="31">Emplenota</emphasis>
</taxonomicName>
species differes between both sexes); median lobe of aedeagus (
<figureCitation id="134C2A4B0832FFEFFEABFD1329A739EC" box="[276,418,741,765]" captionStart="FIGURES 71 78" captionStartId="33.[151,264,1821,1844]" captionTargetBox="[152,1426,187,1820]" captionTargetId="figure@33.[151,1436,141,1822]" captionTargetPageId="33" captionText="FIGURES 71 78. Terminalia of Aleochara (Triochara) trisulcata. 71. tergite VIII of male; 72. tergite VIII of female; 73. sternite VIII of male; 74. sternite VIII of female; 75. male genitalia: median lobe of aedeagus in lateral view; 76. male genitalia: median lobe of aedeagus in ventral view; 77. apical lobe of paramerite, lateral view; 78. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282605/files/figure.png" pageId="30" pageNumber="31">Figs. 7576</figureCitation>
,
<figureCitation id="134C2A4B0832FFEFFE0BFD1329F839EC" box="[436,509,741,765]" captionStart="FIGURES 79 85" captionStartId="36.[151,264,1915,1938]" captionTargetBox="[178,1416,289,1854]" captionTargetId="figure@36.[151,1436,265,1894]" captionTargetPageId="36" captionText="FIGURES 79 85. Terminalia of Aleochara (Triochara) zerchei. 79. tergite VIII of male; 80. tergite VIII of female; 81. sternite VIII of male; 82. sternite VIII of female; 83. male genitalia: median lobe of aedeagus in lateral view; 84. male genitalia: median lobe of aedeagus in ventral view; 85. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282606/files/figure.png" pageId="30" pageNumber="31">8384</figureCitation>
,
<figureCitation id="134C2A4B0832FFEFFDB0FD132A5339EC" box="[527,598,741,765]" captionStart="FIGURES 86 92" captionStartId="41.[151,264,1942,1965]" captionTargetBox="[153,1404,302,1941]" captionTargetId="figure@41.[151,1436,265,1951]" captionTargetPageId="41" captionText="FIGURES 86 92. Terminalia of Aleochara (Triochara) nubis. 86. tergite VIII of male; 87. tergite VIII of female; 88. sternite VIII of male; 89. sternite VIII of female; 90. male genitalia: median lobe of aedeagus in lateral view; 91. male genitalia: median lobe of aedeagus in ventral view; 92. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282607/files/figure.png" pageId="30" pageNumber="31">9091</figureCitation>
) with short but projecting flagellum, and subapico-ventral projections varying in shape among species. [
<emphasis id="B903EADC0832FFEFFDAAFCFE2A693831" bold="true" box="[533,620,776,800]" pageId="30" pageNumber="31">Female</emphasis>
]: spermatheca (
<figureCitation id="134C2A4B0832FFEFFC9EFCF12B7A380E" box="[801,895,775,799]" captionStart="FIGURES 71 78" captionStartId="33.[151,264,1821,1844]" captionTargetBox="[152,1426,187,1820]" captionTargetId="figure@33.[151,1436,141,1822]" captionTargetPageId="33" captionText="FIGURES 71 78. Terminalia of Aleochara (Triochara) trisulcata. 71. tergite VIII of male; 72. tergite VIII of female; 73. sternite VIII of male; 74. sternite VIII of female; 75. male genitalia: median lobe of aedeagus in lateral view; 76. male genitalia: median lobe of aedeagus in ventral view; 77. apical lobe of paramerite, lateral view; 78. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282605/files/figure.png" pageId="30" pageNumber="31">Figs. 78</figureCitation>
,
<figureCitation id="134C2A4B0832FFEFFC33FCF12BAF380E" box="[908,938,775,799]" captionStart="FIGURES 79 85" captionStartId="36.[151,264,1915,1938]" captionTargetBox="[178,1416,289,1854]" captionTargetId="figure@36.[151,1436,265,1894]" captionTargetPageId="36" captionText="FIGURES 79 85. Terminalia of Aleochara (Triochara) zerchei. 79. tergite VIII of male; 80. tergite VIII of female; 81. sternite VIII of male; 82. sternite VIII of female; 83. male genitalia: median lobe of aedeagus in lateral view; 84. male genitalia: median lobe of aedeagus in ventral view; 85. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282606/files/figure.png" pageId="30" pageNumber="31">85</figureCitation>
,
<figureCitation id="134C2A4B0832FFEFFC09FCF12BD6380E" box="[950,979,775,799]" captionStart="FIGURES 86 92" captionStartId="41.[151,264,1942,1965]" captionTargetBox="[153,1404,302,1941]" captionTargetId="figure@41.[151,1436,265,1951]" captionTargetPageId="41" captionText="FIGURES 86 92. Terminalia of Aleochara (Triochara) nubis. 86. tergite VIII of male; 87. tergite VIII of female; 88. sternite VIII of male; 89. sternite VIII of female; 90. male genitalia: median lobe of aedeagus in lateral view; 91. male genitalia: median lobe of aedeagus in ventral view; 92. female genitalia: spermatheca." httpUri="https://zenodo.org/record/282607/files/figure.png" pageId="30" pageNumber="31">92</figureCitation>
) simple; head (sh) large; neck (sn) short and fused with (sh); lacking distinct collar, but with erect duct (ss).
</paragraph>
</subSubSection>
<subSubSection id="C36D65450832FFEFFF78FCBB29BC3F24" pageId="30" pageNumber="31" type="discussion">
<paragraph id="8BC836CE0832FFEFFF78FCBB292E38DC" blockId="30.[151,1436,151,1077]" pageId="30" pageNumber="31">
<emphasis id="B903EADC0832FFEFFF78FCBB294A3877" bold="true" box="[199,335,845,870]" pageId="30" pageNumber="31">Comments.</emphasis>
All of the three known species of the subgenus are similar to each other in habitus, but pronotal patterns, the median lobe of the aedeagus, and tergite VIII of both sexes can be easily differentiated. However, the shape of sternite VIII, which usually act as important character state, do not provide precise identification method in
<taxonomicName id="4C774D4D0832FFEFFF0BFC43292738DC" box="[180,290,949,973]" class="Insecta" family="Staphylinidae" genus="Triochara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B903EADC0832FFEFFF0BFC43292738DC" box="[180,290,949,973]" italics="true" pageId="30" pageNumber="31">Triochara</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BC836CE0832FFEFFF78FC2129BC3F24" blockId="30.[151,1436,151,1077]" pageId="30" pageNumber="31">
The life history of
<taxonomicName id="4C774D4D0832FFEFFE28FC2E2A0238E1" box="[407,519,984,1008]" class="Insecta" family="Staphylinidae" genus="Triochara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B903EADC0832FFEFFE28FC2E2A0238E1" box="[407,519,984,1008]" italics="true" pageId="30" pageNumber="31">Triochara</emphasis>
</taxonomicName>
is thought to be similar to that of
<taxonomicName id="4C774D4D0832FFEFFC3FFC2E2BFC38E1" box="[896,1017,984,1008]" class="Insecta" family="Staphylinidae" genus="Emplenota" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B903EADC0832FFEFFC3FFC2E2BFC38E1" box="[896,1017,984,1008]" italics="true" pageId="30" pageNumber="31">Emplenota</emphasis>
</taxonomicName>
and
<taxonomicName id="4C774D4D0832FFEFFB90FC2E2D6F38FE" authority="Assing, 1995" authorityName="Assing" authorityYear="1995" box="[1071,1386,983,1008]" class="Insecta" family="Staphylinidae" genus="Polystomota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B903EADC0832FFEFFB90FC2E2CBE38E1" box="[1071,1211,984,1008]" italics="true" pageId="30" pageNumber="31">Polystomota</emphasis>
(
<bibRefCitation id="EFE64B3F0832FFEFFB76FC212D6738FE" author="Assing" box="[1225,1378,983,1007]" pageId="30" pageNumber="31" refString="Assing, V. (1995) The Palaearctic species of Emplenota Casey, Polystomota Casey, Triochara Bernhauer and Skenochara Bernhauer &amp; Scheerpeltz, with descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). Beitrage zur Entomologie, 45, 217 - 237." type="journal article" year="1995">Assing, 1995</bibRefCitation>
)
</taxonomicName>
. For information on the dipteran host of
<taxonomicName id="4C774D4D0832FFEFFD99FC0D2B773F03" authority="Weise, 1877" authorityName="Weise" authorityYear="1877" box="[550,882,1018,1043]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="trisulcata" subGenus="Triochara">
<emphasis id="B903EADC0832FFEFFD99FC0D2A323F02" box="[550,567,1019,1043]" italics="true" pageId="30" pageNumber="31">A</emphasis>
. (
<emphasis id="B903EADC0832FFEFFDF1FC0A2A593F02" box="[590,604,1020,1043]" italics="true" pageId="30" pageNumber="31">T</emphasis>
.)
<emphasis id="B903EADC0832FFEFFDCCFC0D2ADA3F02" box="[627,735,1019,1043]" italics="true" pageId="30" pageNumber="31">trisulcata</emphasis>
<bibRefCitation id="EFE64B3F0832FFEFFD59FC0C2B773F03" author="Weise" box="[742,882,1018,1042]" pageId="30" pageNumber="31" refString="Weise, J. (1877) Japanische Staphilinidae [sic] und Pselaphidae. In: Beitrage zur Kaferfauna von Japan, meist auf R. Hiller's Sammlungen basiert. Deutsche Entomologische Zeitschrift, 21, 88 - 100." type="journal article" year="1877">Weise, 1877</bibRefCitation>
</taxonomicName>
, see
<bibRefCitation id="EFE64B3F0832FFEFFC16FC0C2C673F03" author="Yamazaki" box="[937,1122,1018,1042]" pageId="30" pageNumber="31" refString="Yamazaki, K. (2008) Aleochara fucicola and A. trisulcata (Coleoptera, Staphylinidae) as parasitoids of a kelp fly, Fucellia apicalis (Diptera, Anthomyiidae). Elytra, 36 (1), 151 - 152." type="journal article" year="2008">Yamazaki (2008</bibRefCitation>
,
<bibRefCitation id="EFE64B3F0832FFEFFBD1FC0C2CAE3F03" author="Yamazaki" box="[1134,1195,1018,1042]" pageId="30" pageNumber="31" refString="Yamazaki, K. (2012) Seasonal changes in seaweed deposition, seaweed fly abundance, and parasitism at the pupal stage along sandy beaches in central Japan. Entomological Science, 15, 28 - 32." type="journal article" year="2012">2012</bibRefCitation>
). A larval description has never been published.
</paragraph>
</subSubSection>
</treatment>
</document>