135 lines
19 KiB
XML
135 lines
19 KiB
XML
<document id="34C7903FFB46E7F87313DD7E38E79D1F" ID-DOI="10.1080/00222933.2020.1781947" ID-ISSN="1464-5262" ID-Zenodo-Dep="5020903" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624477374180" checkinUser="carolina" docAuthor="Valois, Marcely C., Vaz-de-Mello, Fernando Z. & Silva, Fernando A. B." docDate="2020" docId="C4290E04F41CFFFE11C507155F2D3A78" docLanguage="en" docName="JNATHIST.54.17-18.1081-1114.pdf" docOrigin="Journal of Natural History 54 (17 - 18)" docSource="http://dx.doi.org/10.1080/00222933.2020.1781947" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Scatonomus marii Martinez 1949" docType="treatment" docVersion="4" lastPageNumber="1100" masterDocId="3810767CF408FFEB106003535B043E72" masterDocTitle="A taxonomic revision of the Neotropical genus Scatonomus Erichson, 1835 (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="1114" masterPageNumber="1081" pageNumber="1100" updateTime="1698973370024" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="C07F10927FE3862F6D4EE6D6F16C0F76">A taxonomic revision of the Neotropical genus Scatonomus Erichson, 1835 (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:name id="85246479921C535C6107FCE4B615A073" type="personal">
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<mods:namePart id="B47FA378864EDEDE1B951E662DDE449D">Valois, Marcely C.</mods:namePart>
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<mods:namePart id="3E741925E00AB5D7B789837E704F6EB4">Vaz-de-Mello, Fernando Z.</mods:namePart>
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<mods:name id="4210D0AF77EE9191281B2BDF8D91081B" type="personal">
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<mods:roleTerm id="C3B1CC92B9E1CC783068609185B501B9">Author</mods:roleTerm>
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<mods:namePart id="2BBE100F0E23A7D8D149A54661A910A0">Silva, Fernando A. B.</mods:namePart>
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<mods:typeOfResource id="D0387CE4B457646315C60D4AE8CA9B25">text</mods:typeOfResource>
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<mods:title id="5F76B4C81F3B46CA3EA1CA7EB6F66C10">Journal of Natural History</mods:title>
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<mods:part id="CF95A6C20B39E293F565996BF4A3C6CF">
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<mods:date id="08C6E9D431EF4C59C16F0BB5197105B3">2020</mods:date>
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<mods:number id="DCF33AC1661C3556043EF45E77988438">2020-10-14</mods:number>
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<mods:detail id="ADAE0C8EF1584AB455A05CAA04D6E366" type="volume">
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<mods:number id="C16F80FA45C9DBE3FF97E7F606EE9FFD">54</mods:number>
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<mods:detail id="BA23BA230AB3737B07813610341E279B" type="issue">
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<mods:number id="DF59DE0F8559736E1BD0FA7DE887617B">17 - 18</mods:number>
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<mods:start id="CDE9423FE39030C52E4B99B1D46AEDF5">1081</mods:start>
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<mods:url id="35757970D7058C6C3568BA7988731DE1">http://dx.doi.org/10.1080/00222933.2020.1781947</mods:url>
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<mods:classification id="9C9900A49430C4AD84C34FFB94352FEB">journal article</mods:classification>
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<mods:identifier id="9F42D315FF1D8943718F46457748BD75" type="DOI">10.1080/00222933.2020.1781947</mods:identifier>
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<mods:identifier id="5E9F61AFFEAAD310A0B47FE7295F0412" type="ISSN">1464-5262</mods:identifier>
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<treatment id="C4290E04F41CFFFE11C507155F2D3A78" ID-DOI="http://doi.org/10.5281/zenodo.5021002" ID-Zenodo-Dep="5021002" LSID="urn:lsid:plazi:treatment:C4290E04F41CFFFE11C507155F2D3A78" httpUri="http://treatment.plazi.org/id/C4290E04F41CFFFE11C507155F2D3A78" lastPageId="21" lastPageNumber="1100" pageId="20" pageNumber="1100">
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<subSubSection id="049AEC99F41CFFFF11C50715587B3A13" box="[421,895,1094,1121]" pageId="20" pageNumber="1100" type="nomenclature">
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<paragraph id="4C3FBF12F41CFFFF11C50715587B3A13" blockId="20.[421,895,1025,1121]" box="[421,895,1094,1121]" pageId="20" pageNumber="1100">
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<heading id="1777087EF41CFFFF11C50715587B3A13" box="[421,895,1094,1121]" centered="true" fontSize="11" level="2" pageId="20" pageNumber="1100" reason="5">
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<treatmentCitationGroup id="6C90983CF41CFFFF11C50715587B3A13" box="[421,895,1094,1121]" pageId="20" pageNumber="1100">
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<treatmentCitation id="CD219903F41CFFFF11C50715587F3A13" author="Martinez A" box="[421,891,1094,1121]" page="185" pageId="20" pageNumber="1100" year="1949">
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<taxonomicName id="8B80C491F41CFFFF11C50715587F3A13" ID-CoL="QPD8" authority=": Martinez 1949: 185" authorityName="Martinez" authorityPageNumber="185" authorityYear="1949" box="[421,891,1094,1121]" class="Insecta" family="Scarabaeidae" genus="Scatonomus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="1100" phylum="Arthropoda" rank="species" species="marii">
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<emphasis id="7EF46300F41CFFFF11C50715597A3A13" box="[421,638,1094,1121]" italics="true" pageId="20" pageNumber="1100">Scatonomus marii</emphasis>
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:
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<bibRefCitation id="2811C2E3F41CFFFF12EC0715587F3A13" author="Martinez A" box="[652,891,1094,1121]" pageId="20" pageNumber="1100" pagination="175 - 193" refId="ref20027" refString="Martinez A. 1949. Insectos nuevos o poco conocidos VII. Rev Soc Entomol Argent. 14: 175 - 193." type="journal article" year="1949">Martínez 1949: 185</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="049AEC99F41CFFFF10C007CC59323B82" pageId="20" pageNumber="1100" type="diagnosis">
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<paragraph id="4C3FBF12F41CFFFF10C007CC5A103AC5" blockId="20.[160,1156,1183,1520]" box="[160,276,1183,1207]" pageId="20" pageNumber="1100">
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<emphasis id="7EF46300F41CFFFF10C007CC5A103AC5" bold="true" box="[160,276,1183,1207]" italics="true" pageId="20" pageNumber="1100">Diagnosis</emphasis>
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</paragraph>
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<paragraph id="4C3FBF12F41CFFFF10C0079159323B82" blockId="20.[160,1156,1183,1520]" pageId="20" pageNumber="1100">
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<taxonomicName id="8B80C491F41CFFFF10C007915A853AA9" authorityName="Harold" authorityYear="1867" box="[160,385,1218,1243]" class="Insecta" family="Scarabaeidae" genus="Scatonomus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="1100" phylum="Arthropoda" rank="species" species="insignis">
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<emphasis id="7EF46300F41CFFFF10C007915A853AA9" box="[160,385,1218,1243]" italics="true" pageId="20" pageNumber="1100">Scatonomus insignis</emphasis>
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</taxonomicName>
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is similar to
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<taxonomicName id="8B80C491F41CFFFF126C0791596C3AA9" authority="Valois & Vaz-de-Mello & Silva, 2020" authorityName="Valois & Vaz-de-Mello & Silva" authorityYear="2020" box="[524,616,1218,1243]" class="Insecta" family="Scarabaeidae" genus="Scatonomus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="1100" phylum="Arthropoda" rank="species" species="foresti" status="sp. nov.">
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<emphasis id="7EF46300F41CFFFF126C0791596C3AA9" box="[524,616,1218,1243]" italics="true" pageId="20" pageNumber="1100">S. foresti</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="65C7DE7BF41CFFFF120E079159C13AA9" box="[622,709,1218,1243]" pageId="20" pageNumber="1100" rank="species">sp. nov.</taxonomicNameLabel>
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in having the following combination of characters: body length>
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<quantity id="8B7812F7F41CFFFF11B007B7592C3A8F" box="[464,552,1252,1277]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="20" pageNumber="1100" unit="mm" value="13.0">13 mm</quantity>
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; clypeal edge with two median teeth bounded by lateral emarginations (
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<figureCitation id="D4BBA397F41CFFFF11C7065459033B52" box="[423,519,1287,1312]" captionStart="Figure 3" captionStartId="7.[160,225,1294,1317]" captionTargetBox="[171,1141,146,1237]" captionTargetId="figure-89@7.[163,1153,142,1262]" captionTargetPageId="7" captionText="Figure 3. Head, dorsal view. (a) Scatonomus barbatus stat. rev.; (b) S. canhedoae sp. nov. (holotype); (c) S. chalybaeus; (d) S. fasciculatus; (e) S. foresti sp. nov. (holotype); (f) S. insignis; (g) S. janssensi; (h) S. mitzae sp. nov. (holotype); (i) S. paulosawayai (holotype); (j, k) S. thalassinus; (l) S. viridis." figureDoi="http://doi.org/10.5281/zenodo.5020909" httpUri="https://zenodo.org/record/5020909/files/figure.png" pageId="20" pageNumber="1100">Figure 3</figureCitation>
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(e, f); and dorsum coarsely punctate with dark blue to purple reflections (
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<figureCitation id="D4BBA397F41CFFFF11E206795ADB3B31" box="[386,479,1322,1347]" captionStart="Figure 1" captionStartId="5.[160,225,1145,1168]" captionTargetBox="[194,1153,143,1113]" captionTargetId="figure-187@5.[163,1153,142,1113]" captionTargetPageId="5" captionText="Figure 1. Dorsal habitus. (a) S Scatonomus barbatus stat. rev.; (b) S. canhedoae sp. nov.; (holotype); (c) S. chalybaeus; (d) S. fasciculatus; (e) S. foresti sp. nov. (holotype); (f) S. insignis; (g) S. janssensi; (h) S. mitzae sp. nov. (holotype); (i) S. paulosawayai (holotype); (j, k) S. thalassinus; (l) S. viridis." figureDoi="http://doi.org/10.5281/zenodo.5020905" httpUri="https://zenodo.org/record/5020905/files/figure.png" pageId="20" pageNumber="1100">Figure 1</figureCitation>
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(e, f)). However,
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<taxonomicName id="8B80C491F41CFFFF12CD0679581A3B31" authorityName="Harold" authorityYear="1867" box="[685,798,1322,1347]" class="Insecta" family="Scarabaeidae" genus="Scatonomus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="1100" phylum="Arthropoda" rank="species" species="insignis">
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<emphasis id="7EF46300F41CFFFF12CD0679581A3B31" box="[685,798,1322,1347]" italics="true" pageId="20" pageNumber="1100">S. insignis</emphasis>
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</taxonomicName>
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may be distinguished by the clypeal edge widely margined; clypeal median teeth rounded, separated by a V-shaped emargination (
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<figureCitation id="D4BBA397F41CFFFF112B063C5AA93BFA" box="[331,429,1391,1416]" captionStart="Figure 3" captionStartId="7.[160,225,1294,1317]" captionTargetBox="[171,1141,146,1237]" captionTargetId="figure-89@7.[163,1153,142,1262]" captionTargetPageId="7" captionText="Figure 3. Head, dorsal view. (a) Scatonomus barbatus stat. rev.; (b) S. canhedoae sp. nov. (holotype); (c) S. chalybaeus; (d) S. fasciculatus; (e) S. foresti sp. nov. (holotype); (f) S. insignis; (g) S. janssensi; (h) S. mitzae sp. nov. (holotype); (i) S. paulosawayai (holotype); (j, k) S. thalassinus; (l) S. viridis." figureDoi="http://doi.org/10.5281/zenodo.5020909" httpUri="https://zenodo.org/record/5020909/files/figure.png" pageId="20" pageNumber="1100">Figure 3</figureCitation>
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(f)); antennal club densely coated with yellow setae, antennomeres elongate, slightly pointed at apex; clypeal teeth prolonged beyond the remainder clypeal margin (
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<figureCitation id="D4BBA397F41CFFFF113C06E75AB93BBF" box="[348,445,1460,1485]" captionStart="Figure 3" captionStartId="7.[160,225,1294,1317]" captionTargetBox="[171,1141,146,1237]" captionTargetId="figure-89@7.[163,1153,142,1262]" captionTargetPageId="7" captionText="Figure 3. Head, dorsal view. (a) Scatonomus barbatus stat. rev.; (b) S. canhedoae sp. nov. (holotype); (c) S. chalybaeus; (d) S. fasciculatus; (e) S. foresti sp. nov. (holotype); (f) S. insignis; (g) S. janssensi; (h) S. mitzae sp. nov. (holotype); (i) S. paulosawayai (holotype); (j, k) S. thalassinus; (l) S. viridis." figureDoi="http://doi.org/10.5281/zenodo.5020909" httpUri="https://zenodo.org/record/5020909/files/figure.png" pageId="20" pageNumber="1100">Figure 3</figureCitation>
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(f)); pronotal disc densely puncticulate, slightly wrinkled; and elytral interstriae densely punctate.
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</paragraph>
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</subSubSection>
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<subSubSection id="049AEC99F41CFFFE10C0054F5F2D3A78" lastPageId="21" lastPageNumber="1101" pageId="20" pageNumber="1100" type="description">
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<paragraph id="4C3FBF12F41CFFFF10C0054F5A443846" blockId="20.[160,1156,1564,1728]" box="[160,320,1564,1588]" pageId="20" pageNumber="1100">
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<emphasis id="7EF46300F41CFFFF10C0054F5A443846" bold="true" box="[160,320,1564,1588]" italics="true" pageId="20" pageNumber="1100">Redescription</emphasis>
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</paragraph>
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<paragraph id="4C3FBF12F41CFFFE10C0056C5F2D3A78" blockId="20.[160,1156,1564,1728]" lastBlockId="21.[160,1157,142,1034]" lastPageId="21" lastPageNumber="1101" pageId="20" pageNumber="1100">
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<emphasis id="7EF46300F41CFFFF10C0056C5BD8382A" bold="true" box="[160,220,1599,1624]" pageId="20" pageNumber="1100">Male</emphasis>
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: Length:
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<quantity id="8B7812F7F41CFFFF1129056C5ADE382A" box="[329,474,1599,1624]" metricMagnitude="-2" metricUnit="m" metricValue="1.375" metricValueMax="1.45" metricValueMin="1.3" pageId="20" pageNumber="1100" unit="mm" value="13.75" valueMax="14.5" valueMin="13.0">13–14.5 mm</quantity>
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. Dorsum black, with dark blue or purple reflections (
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<figureCitation id="D4BBA397F41CFFFF145A056C5BAF3809" captionStart="Figure 1" captionStartId="5.[160,225,1145,1168]" captionTargetBox="[194,1153,143,1113]" captionTargetId="figure-187@5.[163,1153,142,1113]" captionTargetPageId="5" captionText="Figure 1. Dorsal habitus. (a) S Scatonomus barbatus stat. rev.; (b) S. canhedoae sp. nov.; (holotype); (c) S. chalybaeus; (d) S. fasciculatus; (e) S. foresti sp. nov. (holotype); (f) S. insignis; (g) S. janssensi; (h) S. mitzae sp. nov. (holotype); (i) S. paulosawayai (holotype); (j, k) S. thalassinus; (l) S. viridis." figureDoi="http://doi.org/10.5281/zenodo.5020905" httpUri="https://zenodo.org/record/5020905/files/figure.png" pageId="20" pageNumber="1100">Figure 1</figureCitation>
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(f)).
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<emphasis id="7EF46300F41CFFFF108005315A243809" bold="true" box="[224,288,1634,1659]" pageId="20" pageNumber="1100">Head</emphasis>
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: surface wrinkled and densely punctate; punctures less impressed near anterior edge (
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<figureCitation id="D4BBA397F41CFFFF112E05D65AAA38EC" box="[334,430,1669,1694]" captionStart="Figure 3" captionStartId="7.[160,225,1294,1317]" captionTargetBox="[171,1141,146,1237]" captionTargetId="figure-89@7.[163,1153,142,1262]" captionTargetPageId="7" captionText="Figure 3. Head, dorsal view. (a) Scatonomus barbatus stat. rev.; (b) S. canhedoae sp. nov. (holotype); (c) S. chalybaeus; (d) S. fasciculatus; (e) S. foresti sp. nov. (holotype); (f) S. insignis; (g) S. janssensi; (h) S. mitzae sp. nov. (holotype); (i) S. paulosawayai (holotype); (j, k) S. thalassinus; (l) S. viridis." figureDoi="http://doi.org/10.5281/zenodo.5020909" httpUri="https://zenodo.org/record/5020909/files/figure.png" pageId="20" pageNumber="1100">Figure 3</figureCitation>
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(f)). Clypeo-genal suture straight, finely sulcate. Fronto-clypeal suture inconspicuous between eyes. Anterior clypeal edge with a row of dorsal orange setae. Clypeus transversely elevated posteriorly. Ventral surface of clypeus densely setose; setae near the anterior edge visible in dorsal view. Ventral clypeal process produced into a transverse and arched carina.
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<emphasis id="7EF46300F41DFFFE12A7038758193E9F" bold="true" box="[711,797,212,237]" pageId="21" pageNumber="1101">Thorax</emphasis>
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: pronotum with an anteromedian knob and a pair of shallow impressions positioned more posteriorly to the knob. Pronotal disc densely puncticulate; mid-longitudinal sulcus present on posterior half. Transverse carina of hypomera incomplete, not reaching the lateral edge. Posterior portion of hypomera finely striate, bearing sparse setae along lateral edge. Surface of mesepimeron and mesanepisternum with green reflections, finely striate and shallowly punctate. Anterior lobe of metaventrite smooth. Lateral lobes of metaventrite finely striate and sparsely punctate. Metaventral disc slightly convex, punctate, with shallow mid-longitudinal sulcus posteriorly.
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<emphasis id="7EF46300F41DFFFE12D502BA59F93C70" bold="true" box="[693,765,489,514]" pageId="21" pageNumber="1101">Elytra</emphasis>
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: striae thin, shallow, indistinctly punctate; width of striae on disc at least 10 times narrower than third interstria. Lateral edge of striae slightly carinate. Seventh and eighth striae absent at the basal one-fifth of elytra. Ninth stria carinate on its apical half and narrowly sulcate on remaining parts.
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<emphasis id="7EF46300F41DFFFE10C001275AAA3CFF" bold="true" box="[160,430,628,653]" pageId="21" pageNumber="1101">Legs [ventral surface]</emphasis>
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: profemora puncticulate; posterior portion with setose punctures, setae long. Outer edge of protibiae with 2 distinct teeth separated by deep Ushaped emargination; basal tooth inconspicuous. Outer edge of mesotibiae and metatibiae with a distinct row of small teeth. Inner edge of mesotibiae and metatibiae carinate, eventually with few setae on apical third.
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<emphasis id="7EF46300F41DFFFE136601AD587B3D65" bold="true" box="[774,895,766,791]" pageId="21" pageNumber="1101">Abdomen</emphasis>
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: ventrites II–IV with a row of coarse punctures along posterior portion. Ventrite V slightly shorter than VI at medial portion.
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<emphasis id="7EF46300F41DFFFE113800175ACA3D2F" bold="true" box="[344,462,836,861]" pageId="21" pageNumber="1101">Pygidium</emphasis>
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: disc sparsely puncticulate.
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<emphasis id="7EF46300F41DFFFE136E001758C83D2F" bold="true" box="[782,972,836,861]" pageId="21" pageNumber="1101">Genital capsule</emphasis>
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: paramera apex divided into apical and lateral lobes by shallow V-shaped emargination. Apical lobe rounded. Lateral lobe truncate, approximately as wide as long; vertical portion of the edge slightly curved inward medially (
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<figureCitation id="D4BBA397F41DFFFE121C00FF59E63DB7" box="[636,738,940,965]" captionStart="Figure 5" captionStartId="9.[160,225,1191,1214]" captionTargetBox="[224,1152,142,1131]" captionTargetId="figure-205@9.[163,1153,142,1159]" captionTargetPageId="9" captionText="Figure 5. Aedeagus, detail of right paramera and endophallic sclerites. (a) Scatonomus barbatus stat. rev.; (b) S. chalybaeus; (c) S. fasciculatus; (d) S. foresti sp. nov.; (e) S. insignis; (f) S. janssensi; (g) S. mitzae sp. nov.; (h) S. thalassinus; (i) S. viridis; (j) endophallic sclerites of S. fasciculatus." figureDoi="http://doi.org/10.5281/zenodo.5020913" httpUri="https://zenodo.org/record/5020913/files/figure.png" pageId="21" pageNumber="1101">Figure 5</figureCitation>
|
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(e)).
|
||
<emphasis id="7EF46300F41DFFFE137B00FF58703DB7" bold="true" box="[795,884,940,965]" pageId="21" pageNumber="1101">Female</emphasis>
|
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: differs from male by having the ventral surface of ventrites II–IV with microgranulation on posterior portion, ventrites V–VI sparsely punctate. Disc of pygidium densely puncticulate.
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||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |