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<document id="0D94054A70C57FD4E48BD79D11340571" ID-DOI="10.1080/00222933.2020.1781947" ID-ISSN="1464-5262" ID-Zenodo-Dep="5020903" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624477374180" checkinUser="carolina" docAuthor="Valois, Marcely C., Vaz-de-Mello, Fernando Z. &amp; Silva, Fernando A. B." docDate="2020" docId="C4290E04F419FFF911E0019A5A653C57" docLanguage="en" docName="JNATHIST.54.17-18.1081-1114.pdf" docOrigin="Journal of Natural History 54 (17 - 18)" docSource="http://dx.doi.org/10.1080/00222933.2020.1781947" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Scatonomus fasciculatus , Erichson 1835" docType="treatment" docVersion="4" lastPageNumber="1097" masterDocId="3810767CF408FFEB106003535B043E72" masterDocTitle="A taxonomic revision of the Neotropical genus Scatonomus Erichson, 1835 (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="1114" masterPageNumber="1081" pageNumber="1097" updateTime="1698973370024" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="CA92C6C6DB4FB579F8FCC18F90B55A24">
<mods:title id="502B5449BF50B54719572C8BE31F5345">A taxonomic revision of the Neotropical genus Scatonomus Erichson, 1835 (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:namePart id="202FBBC50B121CB67DB3F561CDACCF97">Valois, Marcely C.</mods:namePart>
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<mods:namePart id="A550F467F967896E20132BDD4C72A325">Vaz-de-Mello, Fernando Z.</mods:namePart>
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<mods:namePart id="FB27B5171359E2A55DC83411EF6284F9">Silva, Fernando A. B.</mods:namePart>
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<mods:title id="56EFDCC34DB63F490F90D91EBF0F9D8D">Journal of Natural History</mods:title>
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<mods:date id="B7ED39B4C9990BE7E34E3941C638FD99">2020</mods:date>
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<mods:number id="8C9513A4FE2F9E1715448099296E6DFC">2020-10-14</mods:number>
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<mods:number id="66C3661AE1191345FA536CA367CE226A">54</mods:number>
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<mods:number id="E0BA357B656F0EABFE7DB6EDD38A3836">17 - 18</mods:number>
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<mods:identifier id="6ED57A510A116D118F01D29CAA67F678" type="DOI">10.1080/00222933.2020.1781947</mods:identifier>
<mods:identifier id="31E1A75C4176C55FE7BB7D3FBBFA7E14" type="ISSN">1464-5262</mods:identifier>
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<treatment id="C4290E04F419FFF911E0019A5A653C57" ID-DOI="http://doi.org/10.5281/zenodo.5020998" ID-Zenodo-Dep="5020998" LSID="urn:lsid:plazi:treatment:C4290E04F419FFF911E0019A5A653C57" httpUri="http://treatment.plazi.org/id/C4290E04F419FFF911E0019A5A653C57" lastPageId="18" lastPageNumber="1097" pageId="17" pageNumber="1097">
<subSubSection id="049AEC99F419FFFA11E0019A58A03C96" box="[384,932,713,740]" pageId="17" pageNumber="1097" type="nomenclature">
<paragraph id="4C3FBF12F419FFFA11E0019A58A03C96" blockId="17.[384,932,713,740]" box="[384,932,713,740]" pageId="17" pageNumber="1097">
<heading id="1777087EF419FFFA11E0019A58A03C96" box="[384,932,713,740]" centered="true" fontSize="11" level="2" pageId="17" pageNumber="1097" reason="5">
<treatmentCitationGroup id="6C90983CF419FFFA11E0019A58A03C96" box="[384,932,713,740]" pageId="17" pageNumber="1097">
<treatmentCitation id="CD219903F419FFFA11E0019A58A43C96" author="Erichson WS" box="[384,928,713,740]" page="258" pageId="17" pageNumber="1097" year="1835">
<taxonomicName id="8B80C491F419FFFA11E0019A58A43C96" ID-CoL="4V3Z8" authority="Erichson, 1835: 258" authorityName=", Erichson" authorityPageNumber="258" authorityYear="1835" box="[384,928,713,740]" class="Insecta" family="Scarabaeidae" genus="Scatonomus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="1097" phylum="Arthropoda" rank="species" species="fasciculatus">
<emphasis id="7EF46300F419FFFA11E0019A59A23C96" box="[384,678,713,740]" italics="true" pageId="17" pageNumber="1097">Scatonomus fasciculatus</emphasis>
<bibRefCitation id="2811C2E3F419FFFA12CF019A58A43C96" author="Erichson WS" box="[687,928,713,740]" pageId="17" pageNumber="1097" pagination="256 - 270" refId="ref19426" refString="Erichson WS. 1835. Neue sudamerikanische Kafergattungen aus der Familie der Blatterhorner. Arch Naturgesch. 1: 256 - 270." type="journal article" year="1835">Erichson, 1835: 258</bibRefCitation>
</taxonomicName>
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.
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</heading>
</paragraph>
</subSubSection>
<subSubSection id="049AEC99F419FFFA10C000725F5A3A5F" pageId="17" pageNumber="1097" type="diagnosis">
<paragraph id="4C3FBF12F419FFFA10C000725A103D4B" blockId="17.[160,1156,801,1069]" box="[160,276,801,825]" pageId="17" pageNumber="1097">
<emphasis id="7EF46300F419FFFA10C000725A103D4B" bold="true" box="[160,276,801,825]" italics="true" pageId="17" pageNumber="1097">Diagnosis</emphasis>
</paragraph>
<paragraph id="4C3FBF12F419FFFA10C000175F5A3A5F" blockId="17.[160,1156,801,1069]" pageId="17" pageNumber="1097">
<taxonomicName id="8B80C491F419FFFA10C000175AAD3D2F" authorityName=", Erichson" authorityYear="1835" box="[160,425,836,861]" class="Insecta" family="Scarabaeidae" genus="Scatonomus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="1097" phylum="Arthropoda" rank="species" species="fasciculatus">
<emphasis id="7EF46300F419FFFA10C000175AAD3D2F" box="[160,425,836,861]" italics="true" pageId="17" pageNumber="1097">Scatonomus fasciculatus</emphasis>
</taxonomicName>
and
<taxonomicName id="8B80C491F419FFFA11BF0017595C3D2F" authorityName="Erichson" authorityYear="1835" box="[479,600,836,861]" class="Insecta" family="Scarabaeidae" genus="Scatonomus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="1097" phylum="Arthropoda" rank="species" species="barbatus">
<emphasis id="7EF46300F419FFFA11BF0017595C3D2F" box="[479,600,836,861]" italics="true" pageId="17" pageNumber="1097">S. barbatus</emphasis>
</taxonomicName>
are closely related. Both species have body longer than
<quantity id="8B7812F7F419FFFA10BF00345A2C3DF2" box="[223,296,871,896]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="17" pageNumber="1097" unit="mm" value="9.0">9 mm</quantity>
; dorsal surface shiny, almost smooth; genal edge not distinctly angulate laterally (
<figureCitation id="D4BBA397F419FFFA116B00D95A6F3DD1" box="[267,363,906,931]" captionStart="Figure 3" captionStartId="7.[160,225,1294,1317]" captionTargetBox="[171,1141,146,1237]" captionTargetId="figure-89@7.[163,1153,142,1262]" captionTargetPageId="7" captionText="Figure 3. Head, dorsal view. (a) Scatonomus barbatus stat. rev.; (b) S. canhedoae sp. nov. (holotype); (c) S. chalybaeus; (d) S. fasciculatus; (e) S. foresti sp. nov. (holotype); (f) S. insignis; (g) S. janssensi; (h) S. mitzae sp. nov. (holotype); (i) S. paulosawayai (holotype); (j, k) S. thalassinus; (l) S. viridis." figureDoi="http://doi.org/10.5281/zenodo.5020909" httpUri="https://zenodo.org/record/5020909/files/figure.png" pageId="17" pageNumber="1097">Figure 3</figureCitation>
(a, d)); and clypeal edge bidentate, with deep lateral emarginations filled with a tuft of long setae (
<figureCitation id="D4BBA397F419FFFA119000FF594E3DB7" box="[496,586,940,965]" captionStart="Figure 3" captionStartId="7.[160,225,1294,1317]" captionTargetBox="[171,1141,146,1237]" captionTargetId="figure-89@7.[163,1153,142,1262]" captionTargetPageId="7" captionText="Figure 3. Head, dorsal view. (a) Scatonomus barbatus stat. rev.; (b) S. canhedoae sp. nov. (holotype); (c) S. chalybaeus; (d) S. fasciculatus; (e) S. foresti sp. nov. (holotype); (f) S. insignis; (g) S. janssensi; (h) S. mitzae sp. nov. (holotype); (i) S. paulosawayai (holotype); (j, k) S. thalassinus; (l) S. viridis." figureDoi="http://doi.org/10.5281/zenodo.5020909" httpUri="https://zenodo.org/record/5020909/files/figure.png" pageId="17" pageNumber="1097">Figure 3</figureCitation>
(a, d)). However,
<taxonomicName id="8B80C491F419FFFA136100FF58933DB7" authorityName=", Erichson" authorityYear="1835" box="[769,919,940,965]" class="Insecta" family="Scarabaeidae" genus="Scatonomus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="1097" phylum="Arthropoda" rank="species" species="fasciculatus">
<emphasis id="7EF46300F419FFFA136100FF58933DB7" box="[769,919,940,965]" italics="true" pageId="17" pageNumber="1097">S. fasciculatus</emphasis>
</taxonomicName>
can be distinguished by the following combination of characters: clypeal median teeth elongated, approximately as long as the lateral tuft of setae (
<figureCitation id="D4BBA397F419FFFA12F600A159F23A79" box="[662,758,1010,1035]" captionStart="Figure 3" captionStartId="7.[160,225,1294,1317]" captionTargetBox="[171,1141,146,1237]" captionTargetId="figure-89@7.[163,1153,142,1262]" captionTargetPageId="7" captionText="Figure 3. Head, dorsal view. (a) Scatonomus barbatus stat. rev.; (b) S. canhedoae sp. nov. (holotype); (c) S. chalybaeus; (d) S. fasciculatus; (e) S. foresti sp. nov. (holotype); (f) S. insignis; (g) S. janssensi; (h) S. mitzae sp. nov. (holotype); (i) S. paulosawayai (holotype); (j, k) S. thalassinus; (l) S. viridis." figureDoi="http://doi.org/10.5281/zenodo.5020909" httpUri="https://zenodo.org/record/5020909/files/figure.png" pageId="17" pageNumber="1097">Figure 3</figureCitation>
(d)); anterior edge of gena slightly straight (
<figureCitation id="D4BBA397F419FFFA116507475A663A5F" box="[261,354,1044,1069]" captionStart="Figure 3" captionStartId="7.[160,225,1294,1317]" captionTargetBox="[171,1141,146,1237]" captionTargetId="figure-89@7.[163,1153,142,1262]" captionTargetPageId="7" captionText="Figure 3. Head, dorsal view. (a) Scatonomus barbatus stat. rev.; (b) S. canhedoae sp. nov. (holotype); (c) S. chalybaeus; (d) S. fasciculatus; (e) S. foresti sp. nov. (holotype); (f) S. insignis; (g) S. janssensi; (h) S. mitzae sp. nov. (holotype); (i) S. paulosawayai (holotype); (j, k) S. thalassinus; (l) S. viridis." figureDoi="http://doi.org/10.5281/zenodo.5020909" httpUri="https://zenodo.org/record/5020909/files/figure.png" pageId="17" pageNumber="1097">Figure 3</figureCitation>
(d), arrow); and dorsum with blue or purple reflections (
<figureCitation id="D4BBA397F419FFFA13B607475F2A3A5F" box="[982,1070,1044,1069]" captionStart="Figure 1" captionStartId="5.[160,225,1145,1168]" captionTargetBox="[194,1153,143,1113]" captionTargetId="figure-187@5.[163,1153,142,1113]" captionTargetPageId="5" captionText="Figure 1. Dorsal habitus. (a) S Scatonomus barbatus stat. rev.; (b) S. canhedoae sp. nov.; (holotype); (c) S. chalybaeus; (d) S. fasciculatus; (e) S. foresti sp. nov. (holotype); (f) S. insignis; (g) S. janssensi; (h) S. mitzae sp. nov. (holotype); (i) S. paulosawayai (holotype); (j, k) S. thalassinus; (l) S. viridis." figureDoi="http://doi.org/10.5281/zenodo.5020905" httpUri="https://zenodo.org/record/5020905/files/figure.png" pageId="17" pageNumber="1097">Figure 1</figureCitation>
(d)).
</paragraph>
</subSubSection>
<subSubSection id="049AEC99F419FFF910C007095A653C57" lastPageId="18" lastPageNumber="1098" pageId="17" pageNumber="1097" type="description">
<paragraph id="4C3FBF12F419FFFA10C007095A443A00" blockId="17.[160,1156,1114,1728]" box="[160,320,1114,1138]" pageId="17" pageNumber="1097">
<emphasis id="7EF46300F419FFFA10C007095A443A00" bold="true" box="[160,320,1114,1138]" italics="true" pageId="17" pageNumber="1097">Redescription</emphasis>
</paragraph>
<paragraph id="4C3FBF12F419FFF910C0072E5A653C57" blockId="17.[160,1156,1114,1728]" lastBlockId="18.[160,1156,142,549]" lastPageId="18" lastPageNumber="1098" pageId="17" pageNumber="1097">
<emphasis id="7EF46300F419FFFA10C0072E5BDE3AE4" bold="true" box="[160,218,1149,1174]" pageId="17" pageNumber="1097">Male</emphasis>
: length:
<quantity id="8B7812F7F419FFFA115A072E5AA23AE4" box="[314,422,1149,1174]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.3" metricValueMin="0.9" pageId="17" pageNumber="1097" unit="mm" value="11.0" valueMax="13.0" valueMin="9.0">9–13 mm</quantity>
. Dorsum with black, blue or purple reflections (
<figureCitation id="D4BBA397F419FFFA13D3072E5F0E3AE4" box="[947,1034,1149,1174]" captionStart="Figure 1" captionStartId="5.[160,225,1145,1168]" captionTargetBox="[194,1153,143,1113]" captionTargetId="figure-187@5.[163,1153,142,1113]" captionTargetPageId="5" captionText="Figure 1. Dorsal habitus. (a) S Scatonomus barbatus stat. rev.; (b) S. canhedoae sp. nov.; (holotype); (c) S. chalybaeus; (d) S. fasciculatus; (e) S. foresti sp. nov. (holotype); (f) S. insignis; (g) S. janssensi; (h) S. mitzae sp. nov. (holotype); (i) S. paulosawayai (holotype); (j, k) S. thalassinus; (l) S. viridis." figureDoi="http://doi.org/10.5281/zenodo.5020905" httpUri="https://zenodo.org/record/5020905/files/figure.png" pageId="17" pageNumber="1097">Figure 1</figureCitation>
(d)).
<emphasis id="7EF46300F419FFFA145F072E5F793AE4" bold="true" box="[1087,1149,1149,1174]" pageId="17" pageNumber="1097">Head</emphasis>
: densely punctate, slightly wrinkled medially (
<figureCitation id="D4BBA397F419FFFA12C607CC58013ACA" box="[678,773,1183,1208]" captionStart="Figure 3" captionStartId="7.[160,225,1294,1317]" captionTargetBox="[171,1141,146,1237]" captionTargetId="figure-89@7.[163,1153,142,1262]" captionTargetPageId="7" captionText="Figure 3. Head, dorsal view. (a) Scatonomus barbatus stat. rev.; (b) S. canhedoae sp. nov. (holotype); (c) S. chalybaeus; (d) S. fasciculatus; (e) S. foresti sp. nov. (holotype); (f) S. insignis; (g) S. janssensi; (h) S. mitzae sp. nov. (holotype); (i) S. paulosawayai (holotype); (j, k) S. thalassinus; (l) S. viridis." figureDoi="http://doi.org/10.5281/zenodo.5020909" httpUri="https://zenodo.org/record/5020909/files/figure.png" pageId="17" pageNumber="1097">Figure 3</figureCitation>
(d)). Anterior and posterior edges narrowly margined. Clypeo-genal and clypeo-frontal sutures carinate, arched. Clypeal median teeth upturned, separated by U-shaped emargination (
<figureCitation id="D4BBA397F419FFFA13EE07B658F53A8C" box="[910,1009,1253,1278]" captionStart="Figure 3" captionStartId="7.[160,225,1294,1317]" captionTargetBox="[171,1141,146,1237]" captionTargetId="figure-89@7.[163,1153,142,1262]" captionTargetPageId="7" captionText="Figure 3. Head, dorsal view. (a) Scatonomus barbatus stat. rev.; (b) S. canhedoae sp. nov. (holotype); (c) S. chalybaeus; (d) S. fasciculatus; (e) S. foresti sp. nov. (holotype); (f) S. insignis; (g) S. janssensi; (h) S. mitzae sp. nov. (holotype); (i) S. paulosawayai (holotype); (j, k) S. thalassinus; (l) S. viridis." figureDoi="http://doi.org/10.5281/zenodo.5020909" httpUri="https://zenodo.org/record/5020909/files/figure.png" pageId="17" pageNumber="1097">Figure 3</figureCitation>
(d)). Anterior portion of clypeus strongly concave medially, surface less punctate than remaining portions of head. Clypeal and genal edges with rows of long erect setae. Antennal club globose, densely coated with long setae.
<emphasis id="7EF46300F419FFFA1216061E59CE3B14" bold="true" box="[630,714,1357,1382]" pageId="17" pageNumber="1097">Thorax</emphasis>
: pronotum shiny, puncticulate, slightly wrinkled. Anteromedian portion of pronotal disc longitudinally wrinkled, and with welldefined microgranulation. Mid-longitudinal sulcus feebly impressed on pronotal disc. Transverse carina of hypomera almost complete, not reaching lateral edge. Anterior portion of hypomera with dense long setae; lateral edge with row of coarse setose punctures, setae short. Surface of mesepimeron filled with dense and fine striae. Surface of mesanepisternum with well-defined microgranulation. Anterior and lateral lobes of metaventrite with well-defined microgranulation, lacking shallow punctures. Disc flat, densely puncticulate.
<emphasis id="7EF46300F419FFFA11F605315AD83809" bold="true" box="[406,476,1634,1659]" pageId="17" pageNumber="1097">Elytra</emphasis>
: striae thin, shallow, indistinctly punctate; width of striae on disc 7 times narrower than the third interstria. Seventh and eighth striae effaced or absent at elytral base. Ninth stria present, weakly impressed at elytral base. Tenth stria inconspicuous. Interstriae plane, shiny, densely punctate, slightly wrinkled.
<emphasis id="7EF46300F41AFFF9138803DD5BFB3EB8" bold="true" pageId="18" pageNumber="1098">Legs [ventral surface]</emphasis>
: profemora densely punctate, surface between punctures smooth. Mesofemora and metafemor sparsely punctate. Outer edge of protibia with 3 distinct teeth separated by deep U-shaped emarginations. Outer edge of mesotibiae and metatibiae with row of short setae and small teeth.
<emphasis id="7EF46300F41AFFF911BC024A59553F40" bold="true" box="[476,593,281,306]" pageId="18" pageNumber="1098">Abdomen</emphasis>
: posterior half of abdominal ventrites II–VI convex at transverse midline. Ventrite V slightly depressed medially, as long as ventrite VI.
<emphasis id="7EF46300F41AFFF910C0020D5A163F05" bold="true" box="[160,274,350,375]" pageId="18" pageNumber="1098">Pygidium</emphasis>
: 1/3 wider than long. Surface shiny, with punctures spaced by 2 times their diameter.
<emphasis id="7EF46300F41AFFF9117002D25AC33FE8" bold="true" box="[272,455,385,410]" pageId="18" pageNumber="1098">Genital capsule</emphasis>
: paramera apex divided into apical and lateral lobes by deep Lshaped emargination. Apical lobe rounded; lateral lobe slightly acute (
<figureCitation id="D4BBA397F41AFFF9139802F75F5F3FCF" box="[1016,1115,420,445]" captionStart="Figure 5" captionStartId="9.[160,225,1191,1214]" captionTargetBox="[224,1152,142,1131]" captionTargetId="figure-205@9.[163,1153,142,1159]" captionTargetPageId="9" captionText="Figure 5. Aedeagus, detail of right paramera and endophallic sclerites. (a) Scatonomus barbatus stat. rev.; (b) S. chalybaeus; (c) S. fasciculatus; (d) S. foresti sp. nov.; (e) S. insignis; (f) S. janssensi; (g) S. mitzae sp. nov.; (h) S. thalassinus; (i) S. viridis; (j) endophallic sclerites of S. fasciculatus." figureDoi="http://doi.org/10.5281/zenodo.5020913" httpUri="https://zenodo.org/record/5020913/files/figure.png" pageId="18" pageNumber="1098">Figure 5</figureCitation>
(c)).
<emphasis id="7EF46300F41AFFF910C002955BF33FAD" bold="true" box="[160,247,454,479]" pageId="18" pageNumber="1098">Female</emphasis>
: differs from male by the abdominal ventrite VI twice longer than V. Disc of pygidium with punctures spaced by 2 times their diameter; apical edge of pygidium slightly angulate.
</paragraph>
</subSubSection>
</treatment>
</document>
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