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<document id="00C81ECA226AAFE05BBA524E6F2403CC" ID-DOI="10.1080/00222933.2020.1781947" ID-ISSN="1464-5262" ID-Zenodo-Dep="5020903" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624477374180" checkinUser="carolina" docAuthor="Valois, Marcely C., Vaz-de-Mello, Fernando Z. &amp; Silva, Fernando A. B." docDate="2020" docId="C4290E04F414FFF611C303B759933EB8" docLanguage="en" docName="JNATHIST.54.17-18.1081-1114.pdf" docOrigin="Journal of Natural History 54 (17 - 18)" docSource="http://dx.doi.org/10.1080/00222933.2020.1781947" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Scatonomus viridis Erichson 1835" docType="treatment" docVersion="4" lastPageNumber="1108" masterDocId="3810767CF408FFEB106003535B043E72" masterDocTitle="A taxonomic revision of the Neotropical genus Scatonomus Erichson, 1835 (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="1114" masterPageNumber="1081" pageNumber="1108" updateTime="1698973370024" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="2FF664CAE833E27934EAD16AF1771C19">A taxonomic revision of the Neotropical genus Scatonomus Erichson, 1835 (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:namePart id="6C6A08DD35097462C7960E4AE0F72BD9">Valois, Marcely C.</mods:namePart>
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<mods:namePart id="B9D32DB6EE3EC436F02BAD84D09EBDAB">Vaz-de-Mello, Fernando Z.</mods:namePart>
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<mods:namePart id="5CFE580154B382BBA3542E45A3797C8F">Silva, Fernando A. B.</mods:namePart>
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<mods:typeOfResource id="D9818F114A2C4E30143C25BFCCF28D38">text</mods:typeOfResource>
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<mods:title id="131958EA52934013BEB5C9D42610EDA8">Journal of Natural History</mods:title>
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<mods:date id="F4F1DFA1044194A392105FAB9AE5687B">2020</mods:date>
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<mods:number id="B93F9FDC96F2884E5F1BDF73099E83E9">2020-10-14</mods:number>
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<mods:number id="F3616AC899E5CA4F60D22971C889A0FF">54</mods:number>
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<mods:detail id="E3301DFF11A4F26CEF56FE8DFB73BC32" type="issue">
<mods:number id="80FD954B8662B0FD939FF944044B74CA">17 - 18</mods:number>
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<mods:identifier id="0A3D453F2390FDAC78A3578F2D83DCBD" type="DOI">10.1080/00222933.2020.1781947</mods:identifier>
<mods:identifier id="F80269FFFAD991E66F0384A3F49F4C16" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="435F5C8BD8BF26DAD46D2EBA90166CE0" type="Zenodo-Dep">5020903</mods:identifier>
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<treatment id="C4290E04F414FFF611C303B759933EB8" ID-DOI="http://doi.org/10.5281/zenodo.5021008" ID-Zenodo-Dep="5021008" LSID="urn:lsid:plazi:treatment:C4290E04F414FFF611C303B759933EB8" httpUri="http://treatment.plazi.org/id/C4290E04F414FFF611C303B759933EB8" lastPageId="29" lastPageNumber="1108" pageId="28" pageNumber="1108">
<subSubSection id="049AEC99F414FFF711C303B758853E8D" box="[419,897,228,255]" pageId="28" pageNumber="1108" type="nomenclature">
<paragraph id="4C3FBF12F414FFF711C303B758853E8D" blockId="28.[419,897,228,255]" box="[419,897,228,255]" pageId="28" pageNumber="1108">
<heading id="1777087EF414FFF711C303B758853E8D" box="[419,897,228,255]" centered="true" fontSize="11" level="2" pageId="28" pageNumber="1108" reason="5">
<treatmentCitationGroup id="6C90983CF414FFF711C303B758853E8D" box="[419,897,228,255]" pageId="28" pageNumber="1108">
<treatmentCitation id="CD219903F414FFF711C303B758793E8D" author="Erichson WS" box="[419,893,228,255]" page="257" pageId="28" pageNumber="1108" year="1835">
<taxonomicName id="8B80C491F414FFF711C303B758793E8D" ID-CoL="4V3ZF" authority="Erichson, 1835: 257" authorityName="Erichson" authorityPageNumber="257" authorityYear="1835" box="[419,893,228,255]" class="Insecta" family="Scarabaeidae" genus="Scatonomus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="1108" phylum="Arthropoda" rank="species" species="viridis">
<emphasis id="7EF46300F414FFF711C303B759873E8D" box="[419,643,228,255]" italics="true" pageId="28" pageNumber="1108">Scatonomus viridis</emphasis>
<bibRefCitation id="2811C2E3F414FFF712EC03B758793E8D" author="Erichson WS" box="[652,893,228,255]" pageId="28" pageNumber="1108" pagination="256 - 270" refId="ref19426" refString="Erichson WS. 1835. Neue sudamerikanische Kafergattungen aus der Familie der Blatterhorner. Arch Naturgesch. 1: 256 - 270." type="journal article" year="1835">Erichson, 1835: 257</bibRefCitation>
</taxonomicName>
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.
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</heading>
</paragraph>
</subSubSection>
<subSubSection id="049AEC99F414FFF710C0026F58053CFF" pageId="28" pageNumber="1108" type="diagnosis">
<paragraph id="4C3FBF12F414FFF710C0026F5A103F26" blockId="28.[160,1156,316,653]" box="[160,276,316,340]" pageId="28" pageNumber="1108">
<emphasis id="7EF46300F414FFF710C0026F5A103F26" bold="true" box="[160,276,316,340]" italics="true" pageId="28" pageNumber="1108">Diagnosis</emphasis>
</paragraph>
<paragraph id="4C3FBF12F414FFF710C0020C58053CFF" blockId="28.[160,1156,316,653]" pageId="28" pageNumber="1108">
<taxonomicName id="8B80C491F414FFF710C0020C5AB73F0A" authority="viridis" authorityName="viridis Erichson" authorityYear="1835" box="[160,435,351,376]" class="Insecta" family="Scarabaeidae" genus="Scatonomus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="1108" phylum="Arthropoda" rank="subSpecies" species="viridis" subSpecies="viridis">
<emphasis id="7EF46300F414FFF710C0020C5AB73F0A" box="[160,435,351,376]" italics="true" pageId="28" pageNumber="1108">Scatonomus viridis viridis</emphasis>
</taxonomicName>
,
<taxonomicName id="8B80C491F414FFF711A1020C593F3F0A" authorityName="Pereira" authorityYear="1954" box="[449,571,351,376]" class="Insecta" family="Scarabaeidae" genus="Scatonomus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="1108" phylum="Arthropoda" rank="species" species="janssensi">
<emphasis id="7EF46300F414FFF711A1020C593F3F0A" box="[449,571,351,376]" italics="true" pageId="28" pageNumber="1108">S. janssensi</emphasis>
</taxonomicName>
and
<taxonomicName id="8B80C491F414FFF71216020C59DD3F0A" authority="Valois &amp; Vaz-de-Mello &amp; Silva, 2020" authorityName="Valois &amp; Vaz-de-Mello &amp; Silva" authorityYear="2020" box="[630,729,351,376]" class="Insecta" family="Scarabaeidae" genus="Scatonomus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="1108" phylum="Arthropoda" rank="species" species="mitzae" status="sp. nov.">
<emphasis id="7EF46300F414FFF71216020C59DD3F0A" box="[630,729,351,376]" italics="true" pageId="28" pageNumber="1108">S. mitzae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="65C7DE7BF414FFF71281020C583C3F0A" box="[737,824,351,376]" pageId="28" pageNumber="1108" rank="species">sp. nov.</taxonomicNameLabel>
are distinguished from other species of the genus by having a single central emargination on clypeal edge, without teeth or lateral emarginations (
<figureCitation id="D4BBA397F414FFF7119D02F7595F3FCF" box="[509,603,420,445]" captionStart="Figure 3" captionStartId="7.[160,225,1294,1317]" captionTargetBox="[171,1141,146,1237]" captionTargetId="figure-89@7.[163,1153,142,1262]" captionTargetPageId="7" captionText="Figure 3. Head, dorsal view. (a) Scatonomus barbatus stat. rev.; (b) S. canhedoae sp. nov. (holotype); (c) S. chalybaeus; (d) S. fasciculatus; (e) S. foresti sp. nov. (holotype); (f) S. insignis; (g) S. janssensi; (h) S. mitzae sp. nov. (holotype); (i) S. paulosawayai (holotype); (j, k) S. thalassinus; (l) S. viridis." figureDoi="http://doi.org/10.5281/zenodo.5020909" httpUri="https://zenodo.org/record/5020909/files/figure.png" pageId="28" pageNumber="1108">Figure 3</figureCitation>
(g, h, l)).
<taxonomicName id="8B80C491F414FFF712DB02F758C93FCF" authority="viridis" authorityName="viridis Erichson" authorityYear="1835" box="[699,973,420,445]" class="Insecta" family="Scarabaeidae" genus="Scatonomus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="1108" phylum="Arthropoda" rank="subSpecies" species="viridis" subSpecies="viridis">
<emphasis id="7EF46300F414FFF712DB02F758C93FCF" box="[699,973,420,445]" italics="true" pageId="28" pageNumber="1108">Scatonomus viridis viridis</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="8B80C491F414FFF710BB02945A563F92" authorityName="Pereira" authorityYear="1954" box="[219,338,455,480]" class="Insecta" family="Scarabaeidae" genus="Scatonomus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="1108" phylum="Arthropoda" rank="species" species="janssensi">
<emphasis id="7EF46300F414FFF710BB02945A563F92" box="[219,338,455,480]" italics="true" pageId="28" pageNumber="1108">S. janssensi</emphasis>
</taxonomicName>
by the clypeal edge rounded, projected forward, neither widely margined nor strongly upturned (
<figureCitation id="D4BBA397F414FFF711CD02B9590A3C71" box="[429,526,490,515]" captionStart="Figure 3" captionStartId="7.[160,225,1294,1317]" captionTargetBox="[171,1141,146,1237]" captionTargetId="figure-89@7.[163,1153,142,1262]" captionTargetPageId="7" captionText="Figure 3. Head, dorsal view. (a) Scatonomus barbatus stat. rev.; (b) S. canhedoae sp. nov. (holotype); (c) S. chalybaeus; (d) S. fasciculatus; (e) S. foresti sp. nov. (holotype); (f) S. insignis; (g) S. janssensi; (h) S. mitzae sp. nov. (holotype); (i) S. paulosawayai (holotype); (j, k) S. thalassinus; (l) S. viridis." figureDoi="http://doi.org/10.5281/zenodo.5020909" httpUri="https://zenodo.org/record/5020909/files/figure.png" pageId="28" pageNumber="1108">Figure 3</figureCitation>
(l)); and dorsal surface of clypeus lacking anteromedian longitudinal carina. In comparison with
<taxonomicName id="8B80C491F414FFF7123C015F59B93C57" authority="Valois &amp; Vaz-de-Mello &amp; Silva, 2020" authorityName="Valois &amp; Vaz-de-Mello &amp; Silva" authorityYear="2020" box="[604,701,524,549]" class="Insecta" family="Scarabaeidae" genus="Scatonomus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="1108" phylum="Arthropoda" rank="species" species="mitzae" status="sp. nov.">
<emphasis id="7EF46300F414FFF7123C015F59B93C57" box="[604,701,524,549]" italics="true" pageId="28" pageNumber="1108">S. mitzae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="65C7DE7BF414FFF712A4015F58183C57" box="[708,796,524,549]" pageId="28" pageNumber="1108" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="8B80C491F414FFF71345015F58C63C57" authority="viridis" authorityName="viridis Erichson" authorityYear="1835" box="[805,962,524,549]" class="Insecta" family="Scarabaeidae" genus="Scatonomus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="1108" phylum="Arthropoda" rank="subSpecies" species="viridis" subSpecies="viridis">
<emphasis id="7EF46300F414FFF71345015F58C63C57" box="[805,962,524,549]" italics="true" pageId="28" pageNumber="1108">S. viridis viridis</emphasis>
</taxonomicName>
differs by having a shallow central clypeal emargination (
<figureCitation id="D4BBA397F414FFF71206017C59C23C3A" box="[614,710,559,584]" captionStart="Figure 3" captionStartId="7.[160,225,1294,1317]" captionTargetBox="[171,1141,146,1237]" captionTargetId="figure-89@7.[163,1153,142,1262]" captionTargetPageId="7" captionText="Figure 3. Head, dorsal view. (a) Scatonomus barbatus stat. rev.; (b) S. canhedoae sp. nov. (holotype); (c) S. chalybaeus; (d) S. fasciculatus; (e) S. foresti sp. nov. (holotype); (f) S. insignis; (g) S. janssensi; (h) S. mitzae sp. nov. (holotype); (i) S. paulosawayai (holotype); (j, k) S. thalassinus; (l) S. viridis." figureDoi="http://doi.org/10.5281/zenodo.5020909" httpUri="https://zenodo.org/record/5020909/files/figure.png" pageId="28" pageNumber="1108">Figure 3</figureCitation>
(l)); antenal club coated with very short setae; elytral striae slightly narrowed, lateral margins distinctly carinate, width of striae on disc at least 7 times narrower than the third interstria.
</paragraph>
</subSubSection>
<subSubSection id="049AEC99F414FFF610C001E959933EB8" lastPageId="29" lastPageNumber="1109" pageId="28" pageNumber="1108" type="description">
<paragraph id="4C3FBF12F414FFF710C001E95A443CA0" blockId="28.[160,1156,698,1728]" box="[160,320,698,722]" pageId="28" pageNumber="1108">
<emphasis id="7EF46300F414FFF710C001E95A443CA0" bold="true" box="[160,320,698,722]" italics="true" pageId="28" pageNumber="1108">Redescription</emphasis>
</paragraph>
<paragraph id="4C3FBF12F414FFF610C0018F59933EB8" blockId="28.[160,1156,698,1728]" lastBlockId="29.[160,1155,142,202]" lastPageId="29" lastPageNumber="1109" pageId="28" pageNumber="1108">
Male: length:
<quantity id="8B7812F7F414FFF71159018F5AB03C87" box="[313,436,732,757]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="28" pageNumber="1108" unit="mm" value="11.5" valueMax="12.0" valueMin="11.0">
11
<emphasis id="7EF46300F414FFF71134018F5A653C87" bold="true" box="[340,353,732,757]" pageId="28" pageNumber="1108">–</emphasis>
12 mm
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. Dorsum black, with greenish or reddish reflections (
<figureCitation id="D4BBA397F414FFF71464018F5F5B3C87" box="[1028,1119,732,757]" captionStart="Figure 1" captionStartId="5.[160,225,1145,1168]" captionTargetBox="[194,1153,143,1113]" captionTargetId="figure-187@5.[163,1153,142,1113]" captionTargetPageId="5" captionText="Figure 1. Dorsal habitus. (a) S Scatonomus barbatus stat. rev.; (b) S. canhedoae sp. nov.; (holotype); (c) S. chalybaeus; (d) S. fasciculatus; (e) S. foresti sp. nov. (holotype); (f) S. insignis; (g) S. janssensi; (h) S. mitzae sp. nov. (holotype); (i) S. paulosawayai (holotype); (j, k) S. thalassinus; (l) S. viridis." figureDoi="http://doi.org/10.5281/zenodo.5020905" httpUri="https://zenodo.org/record/5020905/files/figure.png" pageId="28" pageNumber="1108">Figure 1</figureCitation>
(l)).
<emphasis id="7EF46300F414FFF710C001AC5BDA3D6A" bold="true" box="[160,222,767,792]" pageId="28" pageNumber="1108">Head</emphasis>
: densely punctate, wrinkled (
<figureCitation id="D4BBA397F414FFF7124401AC59863D6A" box="[548,642,767,792]" captionStart="Figure 3" captionStartId="7.[160,225,1294,1317]" captionTargetBox="[171,1141,146,1237]" captionTargetId="figure-89@7.[163,1153,142,1262]" captionTargetPageId="7" captionText="Figure 3. Head, dorsal view. (a) Scatonomus barbatus stat. rev.; (b) S. canhedoae sp. nov. (holotype); (c) S. chalybaeus; (d) S. fasciculatus; (e) S. foresti sp. nov. (holotype); (f) S. insignis; (g) S. janssensi; (h) S. mitzae sp. nov. (holotype); (i) S. paulosawayai (holotype); (j, k) S. thalassinus; (l) S. viridis." figureDoi="http://doi.org/10.5281/zenodo.5020909" httpUri="https://zenodo.org/record/5020909/files/figure.png" pageId="28" pageNumber="1108">Figure 3</figureCitation>
(l)). Anterior edge narrowly margined (
<figureCitation id="D4BBA397F414FFF7144901AC5F873D6A" box="[1065,1155,767,792]" captionStart="Figure 3" captionStartId="7.[160,225,1294,1317]" captionTargetBox="[171,1141,146,1237]" captionTargetId="figure-89@7.[163,1153,142,1262]" captionTargetPageId="7" captionText="Figure 3. Head, dorsal view. (a) Scatonomus barbatus stat. rev.; (b) S. canhedoae sp. nov. (holotype); (c) S. chalybaeus; (d) S. fasciculatus; (e) S. foresti sp. nov. (holotype); (f) S. insignis; (g) S. janssensi; (h) S. mitzae sp. nov. (holotype); (i) S. paulosawayai (holotype); (j, k) S. thalassinus; (l) S. viridis." figureDoi="http://doi.org/10.5281/zenodo.5020909" httpUri="https://zenodo.org/record/5020909/files/figure.png" pageId="28" pageNumber="1108">Figure 3</figureCitation>
(l)). Clypeo-genal suture sulcate, almost straight. Fronto-clypeal suture weakly defined between eyes. Clypeus with a transverse elevation at posterior portion. Two shallow impressions on interocular space. Anterior clypeal and genal edges with a row of sparse ventral setae. Ventral surface of clypeus densely punctate, wrinkled and glabrous.
<emphasis id="7EF46300F414FFF7144A00D95F7A3DD1" bold="true" box="[1066,1150,906,931]" pageId="28" pageNumber="1108">Thorax</emphasis>
: surface of pronotal disc wrinkled, densely puncticulate. Lateral portions of pronotal disc with ocellate punctures. Anteromedian portion of pronotum with a knob and a pair of shallow impressions positioned posteriorly to the knob. Transversal carina of hypomera complete, reaching the lateral edge. Surface of mesepimeron and mesanepisternum with green or dark blue reflections, finely striate. Mesepimeron with sparse and short setae. Anterior lobe of metaventrite with well-defined microgranulation. Metasternal disc flat, punctate, with a distinct mid- longitudinal sulcus. Lateral lobes of metaventrite with welldefined microgranulation and shallow punctures.
<emphasis id="7EF46300F414FFF712B407CC581E3ACA" bold="true" box="[724,794,1183,1208]" pageId="28" pageNumber="1108">Elytra</emphasis>
: striae shallow, feebly punctate. Seventh stria absent at the basal one-seventh of elytra. Eighth stria carinate, interrupted on basal half, absent near the anterior portion of elytra. Ninth stria carinate, present only at the apical fifth of elytra, punctate on its apical half. Tenth stria carinate, present at medial portion of elytra. Interstriae with punctate and microgranulated surface.
<emphasis id="7EF46300F414FFF7142C06795A623B14" bold="true" pageId="28" pageNumber="1108">Legs [ventral surface]</emphasis>
: profemora densely puncticulate, surface between punctures slightly wrinkled. Mesofemur and metafemur sparsely punctate. Outer edge of protibia with 3 indistinct teeth separated by feeble U-shaped emarginations. Outer edge of mesotibia and metatibia with a row of coarse teeth. Outer carina of mesotibia and metatibia with weakly defined teeth, glabrous. Inner edge of metatibia carinate, except by apical onethird that bears few setae.
<emphasis id="7EF46300F414FFF711A906A9593A3861" bold="true" box="[457,574,1530,1555]" pageId="28" pageNumber="1108">Abdomen</emphasis>
: ventrites sparsely punctate, ventrites I–V with welldefined microgranulation on anterior half. Posterior half of ventrites II–IV with a row of coarse, transversely elongated punctures. Ventrite V slightly shorter than ventrites II–IV and 2 times shorter than ventrite VI.
<emphasis id="7EF46300F414FFF71225053159B33809" bold="true" box="[581,695,1634,1659]" pageId="28" pageNumber="1108">Pygidium</emphasis>
: shiny, slightly wrinkled, densely puncticulate.
<emphasis id="7EF46300F414FFF7109005D65AAC38EC" bold="true" box="[240,424,1669,1694]" pageId="28" pageNumber="1108">Genital capsule</emphasis>
: paramera apex divided into apical and lateral lobes by a deep Vshaped emargination. Apical lobe slightly acute. Lateral lobe truncate, slightly wider than long (
<figureCitation id="D4BBA397F415FFF6108103DD5A393ED5" box="[225,317,142,167]" captionStart="Figure 5" captionStartId="9.[160,225,1191,1214]" captionTargetBox="[224,1152,142,1131]" captionTargetId="figure-205@9.[163,1153,142,1159]" captionTargetPageId="9" captionText="Figure 5. Aedeagus, detail of right paramera and endophallic sclerites. (a) Scatonomus barbatus stat. rev.; (b) S. chalybaeus; (c) S. fasciculatus; (d) S. foresti sp. nov.; (e) S. insignis; (f) S. janssensi; (g) S. mitzae sp. nov.; (h) S. thalassinus; (i) S. viridis; (j) endophallic sclerites of S. fasciculatus." figureDoi="http://doi.org/10.5281/zenodo.5020913" httpUri="https://zenodo.org/record/5020913/files/figure.png" pageId="29" pageNumber="1109">Figure 5</figureCitation>
(j)).
<emphasis id="7EF46300F415FFF6110403DD5ABF3ED5" bold="true" box="[356,443,142,167]" pageId="29" pageNumber="1109">Female</emphasis>
: differs from male by posterior half of ventrites II–IV lacking row of coarse, transversally elongated punctures.
</paragraph>
</subSubSection>
</treatment>
</document>
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