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<document id="9E69BA164BDE1BDCB821030640087645" ID-DOI="10.1080/00222933.2020.1781947" ID-ISSN="1464-5262" ID-Zenodo-Dep="5020903" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624477374180" checkinUser="carolina" docAuthor="Valois, Marcely C., Vaz-de-Mello, Fernando Z. &amp; Silva, Fernando A. B." docDate="2020" docId="C4290E04F412FFF0110E03A55A013E9F" docLanguage="en" docName="JNATHIST.54.17-18.1081-1114.pdf" docOrigin="Journal of Natural History 54 (17 - 18)" docSource="http://dx.doi.org/10.1080/00222933.2020.1781947" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Scatonomus thalassinus Waterhouse 1891" docType="treatment" docVersion="3" lastPageNumber="1106" masterDocId="3810767CF408FFEB106003535B043E72" masterDocTitle="A taxonomic revision of the Neotropical genus Scatonomus Erichson, 1835 (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="1114" masterPageNumber="1081" pageNumber="1106" updateTime="1698973370024" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="72A4BAA7938A36B1995FD6AC4BFC4A88" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="66F3996CBC40FD00A9791DAE095F30F0">
<mods:title id="4BF881AE459C024B418F12334A263BC9">A taxonomic revision of the Neotropical genus Scatonomus Erichson, 1835 (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:name id="BF05F5C51AA2934EF74FD0E40CFB9D5C" type="personal">
<mods:role id="ED5F9C031C9B15ADD5A4FDA97FDAF519">
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<mods:namePart id="B46AFF7C0F38C7C7B11CCFC7D4729218">Valois, Marcely C.</mods:namePart>
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<mods:namePart id="8D57A307A949A53806783C637882F30C">Vaz-de-Mello, Fernando Z.</mods:namePart>
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<mods:namePart id="F0DEE5F41B143D14A2F584B579A7EADD">Silva, Fernando A. B.</mods:namePart>
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<mods:typeOfResource id="FCB48BD11C5D35ADD03B5EF32E22172C">text</mods:typeOfResource>
<mods:relatedItem id="C21DC93F2B252E560DEAEB74EABB944A" type="host">
<mods:titleInfo id="7EB1CD2EBEA86F3EB06DED166F5EF150">
<mods:title id="C164C424D14C447149A23938F16453DB">Journal of Natural History</mods:title>
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<mods:part id="51746249F0A85DD0E208B17073E372C1">
<mods:date id="EBC83CAB804A3945905E413DFB8574C9">2020</mods:date>
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<mods:number id="B1DFBF2EE86A07572230C7CE25698DE8">2020-10-14</mods:number>
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<mods:detail id="D0825A0AB72D183163AEAB66E2F93C9A" type="volume">
<mods:number id="A8FDCB32DAB773DDAF62C8AE25BC9235">54</mods:number>
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<mods:detail id="471B41D7701702DD1ED7A659B66C72DB" type="issue">
<mods:number id="3B1ECAE6AC0A7E1478DE6E6A628D56E2">17 - 18</mods:number>
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<mods:start id="6A852645B4EA8CDA8B81BB8B9C77F7E9">1081</mods:start>
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<mods:url id="1B8681440E816AFE7D96A32CB37FA410">http://dx.doi.org/10.1080/00222933.2020.1781947</mods:url>
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<mods:classification id="AB548135D2EBA7184417AA7B93CD99CC">journal article</mods:classification>
<mods:identifier id="D7B526DC0DB18C9E82D93D1936A53286" type="DOI">10.1080/00222933.2020.1781947</mods:identifier>
<mods:identifier id="132908E2C2023C8032D90A8B142BDCD2" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="F0E57AB9FF576534779BB5D9299969C4" type="Zenodo-Dep">5020903</mods:identifier>
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<treatment id="C4290E04F412FFF0110E03A55A013E9F" LSID="urn:lsid:plazi:treatment:C4290E04F412FFF0110E03A55A013E9F" httpUri="http://treatment.plazi.org/id/C4290E04F412FFF0110E03A55A013E9F" lastPageId="27" lastPageNumber="1106" pageId="26" pageNumber="1106">
<subSubSection id="049AEC99F412FFF1110E03A558B23F63" box="[366,950,246,273]" pageId="26" pageNumber="1106" type="nomenclature">
<paragraph id="4C3FBF12F412FFF1110E03A558B23F63" blockId="26.[366,950,246,273]" box="[366,950,246,273]" pageId="26" pageNumber="1106">
<heading id="1777087EF412FFF1110E03A558B23F63" box="[366,950,246,273]" centered="true" fontSize="11" level="2" pageId="26" pageNumber="1106" reason="5">
<treatmentCitationGroup id="6C90983CF412FFF1110E03A558B23F63" box="[366,950,246,273]" pageId="26" pageNumber="1106">
<treatmentCitation id="CD219903F412FFF1110E03A558B63F63" author="Waterhouse CO" box="[366,946,246,273]" page="350" pageId="26" pageNumber="1106" year="1891">
<taxonomicName id="8B80C491F412FFF1110E03A558B63F63" ID-CoL="6XW8M" authority="Waterhouse, 1891: 350" authorityName="Waterhouse" authorityPageNumber="350" authorityYear="1891" box="[366,946,246,273]" class="Insecta" family="Scarabaeidae" genus="Scatonomus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="1106" phylum="Arthropoda" rank="species" species="thalassinus">
<emphasis id="7EF46300F412FFF1110E03A5598A3F63" box="[366,654,246,273]" italics="true" pageId="26" pageNumber="1106">Scatonomus thalassinus</emphasis>
<bibRefCitation id="2811C2E3F412FFF112F603A558B63F63" author="Waterhouse CO" box="[662,946,246,273]" pageId="26" pageNumber="1106" pagination="347 - 350" refId="ref20807" refString="Waterhouse CO. 1891. New Scarabaeidae in the British Museum: a forth contribution. Ann Mag Nat Hist. 7 (6): 347 - 350." type="journal article" year="1891">Waterhouse, 1891: 350</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="049AEC99F412FFF110C00205584B3CF7" pageId="26" pageNumber="1106" type="diagnosis">
<paragraph id="4C3FBF12F412FFF110C002055A103F1C" blockId="26.[160,1156,342,645]" box="[160,276,342,366]" pageId="26" pageNumber="1106">
<emphasis id="7EF46300F412FFF110C002055A103F1C" bold="true" box="[160,276,342,366]" italics="true" pageId="26" pageNumber="1106">Diagnosis</emphasis>
</paragraph>
<paragraph id="4C3FBF12F412FFF110C0022A584B3CF7" blockId="26.[160,1156,342,645]" pageId="26" pageNumber="1106">
<taxonomicName id="8B80C491F412FFF110C0022A5AA13FE0" authorityName="Waterhouse" authorityYear="1891" box="[160,421,377,402]" class="Insecta" family="Scarabaeidae" genus="Scatonomus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="1106" phylum="Arthropoda" rank="species" species="thalassinus">
<emphasis id="7EF46300F412FFF110C0022A5AA13FE0" box="[160,421,377,402]" italics="true" pageId="26" pageNumber="1106">Scatonomus thalassinus</emphasis>
</taxonomicName>
resembles
<taxonomicName id="8B80C491F412FFF11243022A59893FE0" authorityName="Harold" authorityYear="1867" box="[547,653,377,402]" class="Insecta" family="Scarabaeidae" genus="Scatonomus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="1106" phylum="Arthropoda" rank="species" species="insignis">
<emphasis id="7EF46300F412FFF11243022A59893FE0" box="[547,653,377,402]" italics="true" pageId="26" pageNumber="1106">S. insignis</emphasis>
</taxonomicName>
and
<taxonomicName id="8B80C491F412FFF112A4022A58213FE0" authority="Valois &amp; Vaz-de-Mello &amp; Silva, 2020" authorityName="Valois &amp; Vaz-de-Mello &amp; Silva" authorityYear="2020" box="[708,805,377,402]" class="Insecta" family="Scarabaeidae" genus="Scatonomus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="1106" phylum="Arthropoda" rank="species" species="mitzae" status="sp. nov.">
<emphasis id="7EF46300F412FFF112A4022A58213FE0" box="[708,805,377,402]" italics="true" pageId="26" pageNumber="1106">S. mitzae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="65C7DE7BF412FFF1134C022A58863FE0" box="[812,898,377,402]" pageId="26" pageNumber="1106" rank="species">sp. nov.</taxonomicNameLabel>
in having clypeal edge with 3 emarginations (
<figureCitation id="D4BBA397F412FFF111F902CF5AF03FC7" box="[409,500,412,437]" captionStart="Figure 3" captionStartId="7.[160,225,1294,1317]" captionTargetBox="[171,1141,146,1237]" captionTargetId="figure-89@7.[163,1153,142,1262]" captionTargetPageId="7" captionText="Figure 3. Head, dorsal view. (a) Scatonomus barbatus stat. rev.; (b) S. canhedoae sp. nov. (holotype); (c) S. chalybaeus; (d) S. fasciculatus; (e) S. foresti sp. nov. (holotype); (f) S. insignis; (g) S. janssensi; (h) S. mitzae sp. nov. (holotype); (i) S. paulosawayai (holotype); (j, k) S. thalassinus; (l) S. viridis." figureDoi="http://doi.org/10.5281/zenodo.5020909" httpUri="https://zenodo.org/record/5020909/files/figure.png" pageId="26" pageNumber="1106">Figure 3</figureCitation>
(f, h, j, k)) and dorsum with coarse punctures (
<figureCitation id="D4BBA397F412FFF1138902CF5F3B3FC7" box="[1001,1087,412,437]" captionStart="Figure 1" captionStartId="5.[160,225,1145,1168]" captionTargetBox="[194,1153,143,1113]" captionTargetId="figure-187@5.[163,1153,142,1113]" captionTargetPageId="5" captionText="Figure 1. Dorsal habitus. (a) S Scatonomus barbatus stat. rev.; (b) S. canhedoae sp. nov.; (holotype); (c) S. chalybaeus; (d) S. fasciculatus; (e) S. foresti sp. nov. (holotype); (f) S. insignis; (g) S. janssensi; (h) S. mitzae sp. nov. (holotype); (i) S. paulosawayai (holotype); (j, k) S. thalassinus; (l) S. viridis." figureDoi="http://doi.org/10.5281/zenodo.5020905" httpUri="https://zenodo.org/record/5020905/files/figure.png" pageId="26" pageNumber="1106">Figure 1</figureCitation>
(f, h, j, k)). However,
<taxonomicName id="8B80C491F412FFF1115C02EC5ACB3FAA" authorityName="Waterhouse" authorityYear="1891" box="[316,463,447,472]" class="Insecta" family="Scarabaeidae" genus="Scatonomus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="1106" phylum="Arthropoda" rank="species" species="thalassinus">
<emphasis id="7EF46300F412FFF1115C02EC5ACB3FAA" box="[316,463,447,472]" italics="true" pageId="26" pageNumber="1106">S. thalassinus</emphasis>
</taxonomicName>
is distinguished from these two species by the dorsum with green metallic colouration (
<figureCitation id="D4BBA397F412FFF111B202B2592E3F88" box="[466,554,481,506]" captionStart="Figure 1" captionStartId="5.[160,225,1145,1168]" captionTargetBox="[194,1153,143,1113]" captionTargetId="figure-187@5.[163,1153,142,1113]" captionTargetPageId="5" captionText="Figure 1. Dorsal habitus. (a) S Scatonomus barbatus stat. rev.; (b) S. canhedoae sp. nov.; (holotype); (c) S. chalybaeus; (d) S. fasciculatus; (e) S. foresti sp. nov. (holotype); (f) S. insignis; (g) S. janssensi; (h) S. mitzae sp. nov. (holotype); (i) S. paulosawayai (holotype); (j, k) S. thalassinus; (l) S. viridis." figureDoi="http://doi.org/10.5281/zenodo.5020905" httpUri="https://zenodo.org/record/5020905/files/figure.png" pageId="26" pageNumber="1106">Figure 1</figureCitation>
(j, k)); clypeal edge moderately margined with median teeth acute, separated by a deep V-shaped emargination (some specimens have clypeal edge widely margined with margination produced into 4 rounded teeth, separated by Ushaped emarginations; see taxonomic remarks) (
<figureCitation id="D4BBA397F412FFF112AA011A582D3C10" box="[714,809,585,610]" captionStart="Figure 3" captionStartId="7.[160,225,1294,1317]" captionTargetBox="[171,1141,146,1237]" captionTargetId="figure-89@7.[163,1153,142,1262]" captionTargetPageId="7" captionText="Figure 3. Head, dorsal view. (a) Scatonomus barbatus stat. rev.; (b) S. canhedoae sp. nov. (holotype); (c) S. chalybaeus; (d) S. fasciculatus; (e) S. foresti sp. nov. (holotype); (f) S. insignis; (g) S. janssensi; (h) S. mitzae sp. nov. (holotype); (i) S. paulosawayai (holotype); (j, k) S. thalassinus; (l) S. viridis." figureDoi="http://doi.org/10.5281/zenodo.5020909" httpUri="https://zenodo.org/record/5020909/files/figure.png" pageId="26" pageNumber="1106">Figure 3</figureCitation>
(j, k); abdominal ventrites II–IV usually with a row of wrinkles on posterior edge (
<figureCitation id="D4BBA397F412FFF112B0013F582B3CF7" box="[720,815,620,645]" captionStart="Figure 4" captionStartId="8.[160,225,904,927]" captionTargetBox="[166,1143,144,862]" captionTargetId="figure-495@8.[163,1153,142,873]" captionTargetPageId="8" captionText="Figure 4. Details of Scatonomus morphology. (a) S. mitzae sp. nov., microgranulated interstrial surface (white arrow indicating the elytral stria); (b) S. janssensi, detail of elytral striae; (c) S. thalassinus, surface of pronotal disc (wrinkles, square); (d) S. mitzae sp. nov., puncticulate pronotal surface (white arrow, a coase puncture; black arrow, a fine puncture); (e) S. janssensi, ocellate punctures on pronotum; (f) S. fasciculatus, striate surface on mesepimeron; (g) S. janssensi, broad and transverse carina at the posterior portion of clypeus (white arrow) and longitudinal carina (black arrow); (h) holotype of S. xanthopygus (synonym of S. chalybaeus), dry body fat partially covering the pygidial disc; (i, j) Abdominal ventrites of S. thalassinus." figureDoi="http://doi.org/10.5281/zenodo.5020911" httpUri="https://zenodo.org/record/5020911/files/figure.png" pageId="26" pageNumber="1106">Figure 4</figureCitation>
(i)).
</paragraph>
</subSubSection>
<subSubSection id="049AEC99F412FFF010C001E95A013E9F" lastPageId="27" lastPageNumber="1107" pageId="26" pageNumber="1106" type="description">
<paragraph id="4C3FBF12F412FFF110C001E95A443CA0" blockId="26.[160,1156,698,1728]" box="[160,320,698,722]" pageId="26" pageNumber="1106">
<emphasis id="7EF46300F412FFF110C001E95A443CA0" bold="true" box="[160,320,698,722]" italics="true" pageId="26" pageNumber="1106">Redescription</emphasis>
</paragraph>
<paragraph id="4C3FBF12F412FFF010C0018F5A013E9F" blockId="26.[160,1156,698,1728]" lastBlockId="27.[160,1155,142,237]" lastPageId="27" lastPageNumber="1107" pageId="26" pageNumber="1106">
<emphasis id="7EF46300F412FFF110C0018F5BDE3C87" bold="true" box="[160,218,732,757]" pageId="26" pageNumber="1106">Male</emphasis>
: Length:
<quantity id="8B7812F7F412FFF11120018F5ABD3C87" box="[320,441,732,757]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="26" pageNumber="1106" unit="mm" value="11.5" valueMax="13.0" valueMin="10.0">10–13 mm</quantity>
. Dorsum metallic green, usually with red or yellow reflections on head and pronotum (
<figureCitation id="D4BBA397F412FFF111F301AC5AE93D6A" box="[403,493,767,792]" captionStart="Figure 1" captionStartId="5.[160,225,1145,1168]" captionTargetBox="[194,1153,143,1113]" captionTargetId="figure-187@5.[163,1153,142,1113]" captionTargetPageId="5" captionText="Figure 1. Dorsal habitus. (a) S Scatonomus barbatus stat. rev.; (b) S. canhedoae sp. nov.; (holotype); (c) S. chalybaeus; (d) S. fasciculatus; (e) S. foresti sp. nov. (holotype); (f) S. insignis; (g) S. janssensi; (h) S. mitzae sp. nov. (holotype); (i) S. paulosawayai (holotype); (j, k) S. thalassinus; (l) S. viridis." figureDoi="http://doi.org/10.5281/zenodo.5020905" httpUri="https://zenodo.org/record/5020905/files/figure.png" pageId="26" pageNumber="1106">Figure 1</figureCitation>
(j, k)).
<emphasis id="7EF46300F412FFF1125301AC59753D6A" bold="true" box="[563,625,767,792]" pageId="26" pageNumber="1106">Head</emphasis>
: dorsal surface barely shiny, with dense coarse punctures, except on anterior portion of clypeus with sparse punctures. Fronto-clypeal junction smooth medially (
<figureCitation id="D4BBA397F412FFF111AE0016592F3D2C" box="[462,555,837,862]" captionStart="Figure 3" captionStartId="7.[160,225,1294,1317]" captionTargetBox="[171,1141,146,1237]" captionTargetId="figure-89@7.[163,1153,142,1262]" captionTargetPageId="7" captionText="Figure 3. Head, dorsal view. (a) Scatonomus barbatus stat. rev.; (b) S. canhedoae sp. nov. (holotype); (c) S. chalybaeus; (d) S. fasciculatus; (e) S. foresti sp. nov. (holotype); (f) S. insignis; (g) S. janssensi; (h) S. mitzae sp. nov. (holotype); (i) S. paulosawayai (holotype); (j, k) S. thalassinus; (l) S. viridis." figureDoi="http://doi.org/10.5281/zenodo.5020909" httpUri="https://zenodo.org/record/5020909/files/figure.png" pageId="26" pageNumber="1106">Figure 3</figureCitation>
(j, k)). Clypeal edge distinctly margined (
<figureCitation id="D4BBA397F412FFF1138C00165F4D3D2C" box="[1004,1097,837,862]" captionStart="Figure 3" captionStartId="7.[160,225,1294,1317]" captionTargetBox="[171,1141,146,1237]" captionTargetId="figure-89@7.[163,1153,142,1262]" captionTargetPageId="7" captionText="Figure 3. Head, dorsal view. (a) Scatonomus barbatus stat. rev.; (b) S. canhedoae sp. nov. (holotype); (c) S. chalybaeus; (d) S. fasciculatus; (e) S. foresti sp. nov. (holotype); (f) S. insignis; (g) S. janssensi; (h) S. mitzae sp. nov. (holotype); (i) S. paulosawayai (holotype); (j, k) S. thalassinus; (l) S. viridis." figureDoi="http://doi.org/10.5281/zenodo.5020909" httpUri="https://zenodo.org/record/5020909/files/figure.png" pageId="26" pageNumber="1106">Figure 3</figureCitation>
(j, k)). Clypeo-genal suture sulcate, slightly curved outward on posterior half. Ventral clypeal process transverse, produced into an arched carina. Anterior clypeal and genal edges with a row of ventral setae, visible in dorsal view.
<emphasis id="7EF46300F412FFF112D100FE58013DB4" bold="true" box="[689,773,941,966]" pageId="26" pageNumber="1106">Thorax</emphasis>
: pronotal disc coarsely punctate, lacking wrinkles; mid-longitudinal sulcus smooth, broader medially. Transverse carina of hypomera incomplete, not reaching lateral edge. Anterior portion of hypomera finely striate, with erect yellow setae; setae on lateral edge visible in dorsal view. Posterior portion of hypomera smooth. Lateral edge of hypomera with sparse setae. Surface of mesepimeron and mesanepisternum coarsely punctate, slightly microsculptured. Surface of disc and anterior lobe of metaventrite shallowly punctate; lateral lobes with ill-defined microgranulation and sparse coarse punctures.
<emphasis id="7EF46300F412FFF112A507CC580F3ACA" bold="true" box="[709,779,1183,1208]" pageId="26" pageNumber="1106">Elytra</emphasis>
: striae thin, ill-impressed, inconspicuously punctate; on disc at least 10 times narrower than third interstria. Lateral edges of striae finely carinate. Seventh and eighth striae absent on basal one-fifth of elytra. Eighth stria carinate. Ninth stria visible only at the posterior half of elytra. Tenth stria absent. Interstriae shiny, filled with dense shallow punctures; in some individuals interstriae barely shiny with well-defined microgranulation.
<emphasis id="7EF46300F412FFF11363061E5F063B14" bold="true" box="[771,1026,1357,1382]" pageId="26" pageNumber="1106">Legs [ventral surface]</emphasis>
: profemora puncticulate. Posterior edge of profemora with setose punctures near trochanter. Meso- and metafemora finely punctate. Outer edge of protibiae with 3 teeth, apical and median teeth well developed, separated by a deep U-shaped emargination; basal teeth small, separated from median tooth by shallow V-shaped emargination. Outer edge of meso- and metatibiae with row of small teeth on basal two-thirds; apical third carinate. Inner edge of meso- and metatibiae carinate, except by the apical third.
<emphasis id="7EF46300F412FFF113F3054E5F0C3844" bold="true" box="[915,1032,1565,1590]" pageId="26" pageNumber="1106">Abdomen</emphasis>
: ventrite V approximately 2 times shorter than ventrites II–IV and VI at medial portion. Ventrites V–VI flat medially. Ventrites II–IV wrinkled on posterior edge.
<emphasis id="7EF46300F412FFF1137E053158943809" bold="true" box="[798,912,1634,1659]" pageId="26" pageNumber="1106">Pygidium</emphasis>
: surface barely shiny, with sparse punctures and well-defined microgranulation.
<emphasis id="7EF46300F412FFF1132B05D65F0C38EC" bold="true" box="[843,1032,1669,1694]" pageId="26" pageNumber="1106">Genital capsule</emphasis>
: paramera apex divided into apical and lateral lobes by shallow V-shaped emargination. Apical lobe rounded. Lateral lobe widely rounded (
<figureCitation id="D4BBA397F413FFF0120A03DD59CE3ED5" box="[618,714,142,167]" captionStart="Figure 5" captionStartId="9.[160,225,1191,1214]" captionTargetBox="[224,1152,142,1131]" captionTargetId="figure-205@9.[163,1153,142,1159]" captionTargetPageId="9" captionText="Figure 5. Aedeagus, detail of right paramera and endophallic sclerites. (a) Scatonomus barbatus stat. rev.; (b) S. chalybaeus; (c) S. fasciculatus; (d) S. foresti sp. nov.; (e) S. insignis; (f) S. janssensi; (g) S. mitzae sp. nov.; (h) S. thalassinus; (i) S. viridis; (j) endophallic sclerites of S. fasciculatus." figureDoi="http://doi.org/10.5281/zenodo.5020913" httpUri="https://zenodo.org/record/5020913/files/figure.png" pageId="27" pageNumber="1107">Figure 5</figureCitation>
(h)).
<emphasis id="7EF46300F413FFF0129F03DD58523ED5" bold="true" box="[767,854,142,167]" pageId="27" pageNumber="1107">Female</emphasis>
: differs from the male by abdominal ventrites II–IV only finely punctate on posterior half, and ventrites V–VI convex medially.
</paragraph>
</subSubSection>
</treatment>
</document>
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