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<document id="20EADBE4D573969167A9F21D5488F78C" ID-DOI="10.1080/00222933.2020.1781947" ID-ISSN="1464-5262" ID-Zenodo-Dep="5020903" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624477374180" checkinUser="carolina" docAuthor="Valois, Marcely C., Vaz-de-Mello, Fernando Z. &amp; Silva, Fernando A. B." docDate="2020" docId="C4290E04F411FFF2114B03A55A363B20" docLanguage="en" docName="JNATHIST.54.17-18.1081-1114.pdf" docOrigin="Journal of Natural History 54 (17 - 18)" docSource="http://dx.doi.org/10.1080/00222933.2020.1781947" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Scatonomus paulosawayai Vulcano and Pereira 1973" docType="treatment" docVersion="3" lastPageNumber="1105" masterDocId="3810767CF408FFEB106003535B043E72" masterDocTitle="A taxonomic revision of the Neotropical genus Scatonomus Erichson, 1835 (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="1114" masterPageNumber="1081" pageNumber="1105" updateTime="1698973370024" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="FD3342F84A6992893E9CC3AAB2BE129E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7B47F55BFE7A8393919BCAE17F4549DF">
<mods:title id="7209BCB4E71CD6A6A54F295433AAA1A3">A taxonomic revision of the Neotropical genus Scatonomus Erichson, 1835 (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:namePart id="CEAC42D2482351010F367088A8BB88A4">Valois, Marcely C.</mods:namePart>
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<mods:namePart id="0B611F0961B076A947C72D4411B907BD">Vaz-de-Mello, Fernando Z.</mods:namePart>
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<mods:name id="356592098DE312BC744D481D6BFB13CD" type="personal">
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<mods:namePart id="4263FE521912CAEADC70D844834314CC">Silva, Fernando A. B.</mods:namePart>
</mods:name>
<mods:typeOfResource id="646D8E92D20C41E571B5EC25D713A84F">text</mods:typeOfResource>
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<mods:title id="7B76EC3BB4511177D0EDEF7C2B4A9521">Journal of Natural History</mods:title>
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<mods:part id="8320162DD2FCD17D5B8308B9BA2EB5D1">
<mods:date id="28F06D17D82557BEA2A8E48038FE4DC2">2020</mods:date>
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<mods:number id="BAB61AEDCF0B6B4190B2FC4079863577">2020-10-14</mods:number>
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<mods:detail id="8B359AE6D61F6871266963D40E51D3B0" type="volume">
<mods:number id="96A9E3342853F85383EC3AA9DF3603F9">54</mods:number>
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<mods:detail id="70D87832C7AF5E8C9493A87883076A0B" type="issue">
<mods:number id="3C5DD42DC44A4443DD5752600CF522E1">17 - 18</mods:number>
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<mods:start id="A47033A438E8413BD45867D8DF6E98C1">1081</mods:start>
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<mods:url id="63E5A288549E7E063E4556E2839157ED">http://dx.doi.org/10.1080/00222933.2020.1781947</mods:url>
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<mods:classification id="3C381429297E1922C62D15A9FABCB3A7">journal article</mods:classification>
<mods:identifier id="7B2D429FE0FCE56AAAA08231BFE7DEBA" type="DOI">10.1080/00222933.2020.1781947</mods:identifier>
<mods:identifier id="B223488715CAE51453C5A83D5F9297A5" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="8296D9EDAC170699299F0C4268B39A85" type="Zenodo-Dep">5020903</mods:identifier>
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<subSubSection id="049AEC99F411FFF2114B03A558FD3F63" box="[299,1017,246,273]" pageId="25" pageNumber="1105" type="nomenclature">
<paragraph id="4C3FBF12F411FFF2114B03A558FD3F63" blockId="25.[299,1017,246,273]" box="[299,1017,246,273]" pageId="25" pageNumber="1105">
<heading id="1777087EF411FFF2114B03A558FD3F63" box="[299,1017,246,273]" centered="true" fontSize="11" level="2" pageId="25" pageNumber="1105" reason="5">
<treatmentCitationGroup id="6C90983CF411FFF2114B03A558FD3F63" box="[299,1017,246,273]" pageId="25" pageNumber="1105">
<treatmentCitation id="CD219903F411FFF2114B03A558F13F63" author="Vulcano M &amp; Pereira FS" box="[299,1013,246,273]" page="536" pageId="25" pageNumber="1105" year="1973">
<taxonomicName id="8B80C491F411FFF2114B03A558F13F63" ID-CoL="4V3ZC" authority="Vulcano and Pereira, 1973: 536" authorityName="Vulcano and Pereira" authorityPageNumber="536" authorityYear="1973" box="[299,1013,246,273]" class="Insecta" family="Scarabaeidae" genus="Scatonomus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="1105" phylum="Arthropoda" rank="species" species="paulosawayai">
<emphasis id="7EF46300F411FFF2114B03A5596F3F63" box="[299,619,246,273]" italics="true" pageId="25" pageNumber="1105">Scatonomus paulosawayai</emphasis>
<bibRefCitation id="2811C2E3F411FFF2121403A558F13F63" author="Vulcano M &amp; Pereira FS" box="[628,1013,246,273]" pageId="25" pageNumber="1105" pagination="535 - 551" refId="ref20522" refString="Vulcano M A., Pereira FS. 1973. Duas novas especies do genero Scatonomus Er. 1835 (Col. scarabaeidae). Bol Zool Biol. 30 (30): 535 - 551. doi: 10.11606 /. 2526 - 3366. bzbm. 1973.121359." type="journal article" year="1973">Vulcano and Pereira, 1973: 536</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="049AEC99F411FFF210C0020D5AFF3C35" pageId="25" pageNumber="1105" type="diagnosis">
<paragraph id="4C3FBF12F411FFF210C0020D5A103F04" blockId="25.[160,1156,350,583]" box="[160,276,350,374]" pageId="25" pageNumber="1105">
<emphasis id="7EF46300F411FFF210C0020D5A103F04" bold="true" box="[160,276,350,374]" italics="true" pageId="25" pageNumber="1105">Diagnosis</emphasis>
</paragraph>
<paragraph id="4C3FBF12F411FFF210C002D25AFF3C35" blockId="25.[160,1156,350,583]" pageId="25" pageNumber="1105">
<emphasis id="7EF46300F411FFF210C002D259233FE8" box="[160,551,385,410]" italics="true" pageId="25" pageNumber="1105">
<taxonomicName id="8B80C491F411FFF210C002D25A893FE8" authorityName="Erichson" authorityYear="1835" box="[160,397,385,410]" class="Insecta" family="Scarabaeidae" genus="Scatonomus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="1105" phylum="Arthropoda" rank="species" species="barbatus">Scatonomus barbatus</taxonomicName>
,
<taxonomicName id="8B80C491F411FFF211F802D259233FE8" authority="Valois &amp; Vaz-de-Mello &amp; Silva, 2020" authorityName="Valois &amp; Vaz-de-Mello &amp; Silva" authorityYear="2020" box="[408,551,385,410]" class="Insecta" family="Scarabaeidae" genus="Scatonomus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="1105" phylum="Arthropoda" rank="species" species="canhedoae" status="sp. nov.">S. canhedoae</taxonomicName>
</emphasis>
<taxonomicNameLabel id="65C7DE7BF411FFF2124D02D259813FE8" box="[557,645,385,410]" pageId="25" pageNumber="1105" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="8B80C491F411FFF212ED02D258273FE8" authorityName=", Erichson" authorityYear="1835" box="[653,803,385,410]" class="Insecta" family="Scarabaeidae" genus="Scatonomus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="1105" phylum="Arthropoda" rank="species" species="fasciculatus">
<emphasis id="7EF46300F411FFF212ED02D258273FE8" box="[653,803,385,410]" italics="true" pageId="25" pageNumber="1105">S. fasciculatus</emphasis>
</taxonomicName>
and
<taxonomicName id="8B80C491F411FFF2133A02D25F0C3FE8" authorityName="Vulcano and Pereira" authorityYear="1973" box="[858,1032,385,410]" class="Insecta" family="Scarabaeidae" genus="Scatonomus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="1105" phylum="Arthropoda" rank="species" species="paulosawayai">
<emphasis id="7EF46300F411FFF2133A02D25F0C3FE8" box="[858,1032,385,410]" italics="true" pageId="25" pageNumber="1105">S. paulosawayai</emphasis>
</taxonomicName>
have deep lateral emarginations filled with setae at the clypeal edge (
<figureCitation id="D4BBA397F411FFF2134E02F0588D3FCE" box="[814,905,419,444]" captionStart="Figure 3" captionStartId="7.[160,225,1294,1317]" captionTargetBox="[171,1141,146,1237]" captionTargetId="figure-89@7.[163,1153,142,1262]" captionTargetPageId="7" captionText="Figure 3. Head, dorsal view. (a) Scatonomus barbatus stat. rev.; (b) S. canhedoae sp. nov. (holotype); (c) S. chalybaeus; (d) S. fasciculatus; (e) S. foresti sp. nov. (holotype); (f) S. insignis; (g) S. janssensi; (h) S. mitzae sp. nov. (holotype); (i) S. paulosawayai (holotype); (j, k) S. thalassinus; (l) S. viridis." figureDoi="http://doi.org/10.5281/zenodo.5020909" httpUri="https://zenodo.org/record/5020909/files/figure.png" pageId="25" pageNumber="1105">Figure 3</figureCitation>
(a, b, d, i)). However,
<taxonomicName id="8B80C491F411FFF2141102F05A323FAD" authorityName="Vulcano and Pereira" authorityYear="1973" class="Insecta" family="Scarabaeidae" genus="Scatonomus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="1105" phylum="Arthropoda" rank="species" species="paulosawayai">
<emphasis id="7EF46300F411FFF2141102F05A323FAD" italics="true" pageId="25" pageNumber="1105">S. paulosawayai</emphasis>
</taxonomicName>
has the clypeal edge lacking median teeth and the median portion between the lateral emarginations with rectangular shape (
<figureCitation id="D4BBA397F411FFF212B102BA58343C70" box="[721,816,489,514]" captionStart="Figure 3" captionStartId="7.[160,225,1294,1317]" captionTargetBox="[171,1141,146,1237]" captionTargetId="figure-89@7.[163,1153,142,1262]" captionTargetPageId="7" captionText="Figure 3. Head, dorsal view. (a) Scatonomus barbatus stat. rev.; (b) S. canhedoae sp. nov. (holotype); (c) S. chalybaeus; (d) S. fasciculatus; (e) S. foresti sp. nov. (holotype); (f) S. insignis; (g) S. janssensi; (h) S. mitzae sp. nov. (holotype); (i) S. paulosawayai (holotype); (j, k) S. thalassinus; (l) S. viridis." figureDoi="http://doi.org/10.5281/zenodo.5020909" httpUri="https://zenodo.org/record/5020909/files/figure.png" pageId="25" pageNumber="1105">Figure 3</figureCitation>
(i)), dorsum with green or blue reflections (
<figureCitation id="D4BBA397F411FFF2114301585A793C56" box="[291,381,523,548]" captionStart="Figure 1" captionStartId="5.[160,225,1145,1168]" captionTargetBox="[194,1153,143,1113]" captionTargetId="figure-187@5.[163,1153,142,1113]" captionTargetPageId="5" captionText="Figure 1. Dorsal habitus. (a) S Scatonomus barbatus stat. rev.; (b) S. canhedoae sp. nov.; (holotype); (c) S. chalybaeus; (d) S. fasciculatus; (e) S. foresti sp. nov. (holotype); (f) S. insignis; (g) S. janssensi; (h) S. mitzae sp. nov. (holotype); (i) S. paulosawayai (holotype); (j, k) S. thalassinus; (l) S. viridis." figureDoi="http://doi.org/10.5281/zenodo.5020905" httpUri="https://zenodo.org/record/5020905/files/figure.png" pageId="25" pageNumber="1105">Figure 1</figureCitation>
(i)), head and pronotum wrinkled and elytral interstriae opaque, with well-defined microgranulation.
</paragraph>
</subSubSection>
<subSubSection id="049AEC99F411FFF210C001D05A363B20" pageId="25" pageNumber="1105" type="description">
<paragraph id="4C3FBF12F411FFF210C001D05A443CE9" blockId="25.[160,1156,643,1362]" box="[160,320,643,667]" pageId="25" pageNumber="1105">
<emphasis id="7EF46300F411FFF210C001D05A443CE9" bold="true" box="[160,320,643,667]" italics="true" pageId="25" pageNumber="1105">Redescription</emphasis>
</paragraph>
<paragraph id="4C3FBF12F411FFF210C001F55A363B20" blockId="25.[160,1156,643,1362]" pageId="25" pageNumber="1105">
<emphasis id="7EF46300F411FFF210C001F55BDE3CCD" bold="true" box="[160,218,678,703]" pageId="25" pageNumber="1105">Male</emphasis>
: unknown.
<emphasis id="7EF46300F411FFF2110201F55ABD3CCD" bold="true" box="[354,441,678,703]" pageId="25" pageNumber="1105">Female</emphasis>
: length:
<quantity id="8B7812F7F411FFF2124001F559703CCD" box="[544,628,678,703]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="25" pageNumber="1105" unit="mm" value="13.0">13 mm</quantity>
. Dorsum with black, green or blue reflections (
<figureCitation id="D4BBA397F411FFF210C8019A5A053C90" box="[168,257,713,738]" captionStart="Figure 1" captionStartId="5.[160,225,1145,1168]" captionTargetBox="[194,1153,143,1113]" captionTargetId="figure-187@5.[163,1153,142,1113]" captionTargetPageId="5" captionText="Figure 1. Dorsal habitus. (a) S Scatonomus barbatus stat. rev.; (b) S. canhedoae sp. nov.; (holotype); (c) S. chalybaeus; (d) S. fasciculatus; (e) S. foresti sp. nov. (holotype); (f) S. insignis; (g) S. janssensi; (h) S. mitzae sp. nov. (holotype); (i) S. paulosawayai (holotype); (j, k) S. thalassinus; (l) S. viridis." figureDoi="http://doi.org/10.5281/zenodo.5020905" httpUri="https://zenodo.org/record/5020905/files/figure.png" pageId="25" pageNumber="1105">Figure 1</figureCitation>
(i)).
<emphasis id="7EF46300F411FFF2114D019A5A6F3C90" bold="true" box="[301,363,713,738]" pageId="25" pageNumber="1105">Head</emphasis>
: surface wrinkled (
<figureCitation id="D4BBA397F411FFF2125D019A59983C90" box="[573,668,713,738]" captionStart="Figure 3" captionStartId="7.[160,225,1294,1317]" captionTargetBox="[171,1141,146,1237]" captionTargetId="figure-89@7.[163,1153,142,1262]" captionTargetPageId="7" captionText="Figure 3. Head, dorsal view. (a) Scatonomus barbatus stat. rev.; (b) S. canhedoae sp. nov. (holotype); (c) S. chalybaeus; (d) S. fasciculatus; (e) S. foresti sp. nov. (holotype); (f) S. insignis; (g) S. janssensi; (h) S. mitzae sp. nov. (holotype); (i) S. paulosawayai (holotype); (j, k) S. thalassinus; (l) S. viridis." figureDoi="http://doi.org/10.5281/zenodo.5020909" httpUri="https://zenodo.org/record/5020909/files/figure.png" pageId="25" pageNumber="1105">Figure 3</figureCitation>
(i)). Clypeo-genal and fronto-clypeal sutures wide, sulcate. Anterior clypeal and genal edges with rows of ventral, orange setae (
<figureCitation id="D4BBA397F411FFF2145C01B85BB53D55" captionStart="Figure 3" captionStartId="7.[160,225,1294,1317]" captionTargetBox="[171,1141,146,1237]" captionTargetId="figure-89@7.[163,1153,142,1262]" captionTargetPageId="7" captionText="Figure 3. Head, dorsal view. (a) Scatonomus barbatus stat. rev.; (b) S. canhedoae sp. nov. (holotype); (c) S. chalybaeus; (d) S. fasciculatus; (e) S. foresti sp. nov. (holotype); (f) S. insignis; (g) S. janssensi; (h) S. mitzae sp. nov. (holotype); (i) S. paulosawayai (holotype); (j, k) S. thalassinus; (l) S. viridis." figureDoi="http://doi.org/10.5281/zenodo.5020909" httpUri="https://zenodo.org/record/5020909/files/figure.png" pageId="25" pageNumber="1105">Figure 3</figureCitation>
(i)). Antennal club filled with short setae.
<emphasis id="7EF46300F411FFF2121B005D59CB3D55" bold="true" box="[635,719,782,807]" pageId="25" pageNumber="1105">Thorax</emphasis>
: pronotal surface wrinkled and puncticulate. Pronotal disc slightly elevated at the anteromedian portion. Mid-longitudinal sulcus on pronotal disc filled with fine punctures. Transverse carina of hypomera complete, reaching the lateral edge. Anterior and lateral edges of hypomera densely coated with setae. Surface of mesepimeron and mesanepisternum with green reflections, finely striate. Disc of metaventrite flat, punctate, with shallow longitudinal sulcus medially. Lateral lobes of metaventrite with well-defined microgranulation and sparse shallow punctures.
<emphasis id="7EF46300F411FFF2117D07525A673A68" bold="true" box="[285,355,1025,1050]" pageId="25" pageNumber="1105">Elytra</emphasis>
: striae thin, punctures shallowly impressed. Seventh stria almost reaching the elytral base. Eighth stria absent at anterior one-fifth. Ninth stria carinate, inconspicuous near basal portion of elytra. Interstriae punctate, with well-defined microgranulation.
<emphasis id="7EF46300F411FFF21111073A59733AF0" bold="true" box="[369,631,1129,1154]" pageId="25" pageNumber="1105">Legs [ventral surface]</emphasis>
: profemora puncticulate, posterior edge with setose punctures near trochanter. Outer edge of protibiae with two acute teeth, apical tooth longer than median and basal teeth. Meso- and metafemora finely punctate. Outer carina of mesotibiae and metatibiae with row of small teeth. Inner edge of mesotibiae and metatibiae carinate, except by its apical one-third.
<emphasis id="7EF46300F411FFF212AD07A058463B7E" bold="true" box="[717,834,1267,1292]" pageId="25" pageNumber="1105">Abdomen</emphasis>
: posterior half of ventrites II– IV slightly depressed. Ventrite V 3 times shorter than ventrite VI.
<emphasis id="7EF46300F411FFF213F206455F003B5D" bold="true" box="[914,1028,1302,1327]" pageId="25" pageNumber="1105">Pygidium</emphasis>
: shiny and puncticulate.
</paragraph>
</subSubSection>
</treatment>
</document>
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