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<document id="7F72C516536669B13171A1E66DD456E8" ID-DOI="10.1080/00222933.2020.1781947" ID-ISSN="1464-5262" ID-Zenodo-Dep="5020903" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624477374180" checkinUser="carolina" docAuthor="Valois, Marcely C., Vaz-de-Mello, Fernando Z. &amp; Silva, Fernando A. B." docDate="2020" docId="C4290E04F403FFE711F10555593D3B52" docLanguage="en" docName="JNATHIST.54.17-18.1081-1114.pdf" docOrigin="Journal of Natural History 54 (17 - 18)" docSource="http://dx.doi.org/10.1080/00222933.2020.1781947" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Scatonomus barbatus Erichson 1835" docType="treatment" docVersion="4" lastPageNumber="1092" masterDocId="3810767CF408FFEB106003535B043E72" masterDocTitle="A taxonomic revision of the Neotropical genus Scatonomus Erichson, 1835 (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="1114" masterPageNumber="1081" pageNumber="1091" updateTime="1698973370024" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="F14A9D8FC3EC643D73FDDBA002B91932" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="2F6B3B1D30CA883DF23CD9C73E3AA818">
<mods:title id="0FE782FF4ADD5633B95E224FDE3AEE89">A taxonomic revision of the Neotropical genus Scatonomus Erichson, 1835 (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:namePart id="D6D9DBCE5CEAD1C9765827D180283A22">Valois, Marcely C.</mods:namePart>
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<mods:namePart id="3F68BFB2742F7DDC5D66F296BE3FC3B3">Vaz-de-Mello, Fernando Z.</mods:namePart>
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<mods:namePart id="24647ED6BED6BB72C7EB2B3151636EF1">Silva, Fernando A. B.</mods:namePart>
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<mods:title id="745C9DA35AA64E470E5178FE5474FB79">Journal of Natural History</mods:title>
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<mods:date id="74359978C118C341FF019A358FFA80AF">2020</mods:date>
<mods:detail id="D84EE9C2178F28D1FEE66B7D6185B0B2" type="pubDate">
<mods:number id="A2A2F205161FF0DDBFB4F185BD088802">2020-10-14</mods:number>
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<mods:number id="401B922A2468C671BCDF9D55E6FE7374">54</mods:number>
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<mods:detail id="B58509A192C8FD036053A2F247D8FE94" type="issue">
<mods:number id="655B89A7C695598CF68AE0DAB925968E">17 - 18</mods:number>
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<mods:url id="4B377C9E3ADCAE82D2D4D9C15ACDF9DF">http://dx.doi.org/10.1080/00222933.2020.1781947</mods:url>
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<mods:classification id="4F90A991670D15B44BC7A82882C17403">journal article</mods:classification>
<mods:identifier id="8CCD7A3EE4328EB50C51E2C284392506" type="DOI">10.1080/00222933.2020.1781947</mods:identifier>
<mods:identifier id="5DFCB4C29582773ADE31864BB3A5D263" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="73D74A16CCFF8E7DE04A8F3492F25465" type="Zenodo-Dep">5020903</mods:identifier>
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<treatment id="C4290E04F403FFE711F10555593D3B52" ID-DOI="http://doi.org/10.5281/zenodo.5020988" ID-Zenodo-Dep="5020988" LSID="urn:lsid:plazi:treatment:C4290E04F403FFE711F10555593D3B52" httpUri="http://treatment.plazi.org/id/C4290E04F403FFE711F10555593D3B52" lastPageId="12" lastPageNumber="1092" pageId="11" pageNumber="1091">
<subSubSection id="049AEC99F403FFE011F1055558973853" box="[401,915,1542,1569]" pageId="11" pageNumber="1091" type="nomenclature">
<paragraph id="4C3FBF12F403FFE011F1055558973853" blockId="11.[401,915,1542,1569]" box="[401,915,1542,1569]" pageId="11" pageNumber="1091">
<heading id="1777087EF403FFE011F1055558973853" box="[401,915,1542,1569]" centered="true" fontSize="11" level="2" pageId="11" pageNumber="1091" reason="5">
<taxonomicName id="8B80C491F403FFE011F1055558553853" ID-CoL="7YYB4" authority="Erichson, 1835" authorityName="Erichson" authorityYear="1835" box="[401,849,1542,1569]" class="Insecta" family="Scarabaeidae" genus="Scatonomus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="1091" phylum="Arthropoda" rank="species" species="barbatus">
<emphasis id="7EF46300F403FFE011F1055559923853" box="[401,662,1542,1569]" italics="true" pageId="11" pageNumber="1091">Scatonomus barbatus</emphasis>
<bibRefCitation id="2811C2E3F403FFE012FE055558553853" author="Erichson WS" box="[670,849,1542,1569]" pageId="11" pageNumber="1091" pagination="256 - 270" refId="ref19426" refString="Erichson WS. 1835. Neue sudamerikanische Kafergattungen aus der Familie der Blatterhorner. Arch Naturgesch. 1: 256 - 270." type="journal article" year="1835">Erichson, 1835</bibRefCitation>
</taxonomicName>
, 258.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="049AEC99F403FFE710C0053158B23F40" lastPageId="12" lastPageNumber="1092" pageId="11" pageNumber="1091" type="diagnosis">
<paragraph id="4C3FBF12F403FFE010C005315A103808" blockId="11.[160,1155,1634,1728]" box="[160,276,1634,1658]" pageId="11" pageNumber="1091">
<emphasis id="7EF46300F403FFE010C005315A103808" bold="true" box="[160,276,1634,1658]" italics="true" pageId="11" pageNumber="1091">Diagnosis</emphasis>
</paragraph>
<paragraph id="4C3FBF12F403FFE710C005D658B23F40" blockId="11.[160,1155,1634,1728]" lastBlockId="12.[160,1156,142,306]" lastPageId="12" lastPageNumber="1092" pageId="11" pageNumber="1091">
<taxonomicName id="8B80C491F403FFE010C005D65A9538EC" authorityName="Erichson" authorityYear="1835" box="[160,401,1669,1694]" class="Insecta" family="Scarabaeidae" genus="Scatonomus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="1091" phylum="Arthropoda" rank="species" species="barbatus">
<emphasis id="7EF46300F403FFE010C005D65A9538EC" box="[160,401,1669,1694]" italics="true" pageId="11" pageNumber="1091">Scatonomus barbatus</emphasis>
</taxonomicName>
and
<taxonomicName id="8B80C491F403FFE011B005D6596F38EC" authorityName=", Erichson" authorityYear="1835" box="[464,619,1669,1694]" class="Insecta" family="Scarabaeidae" genus="Scatonomus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="1091" phylum="Arthropoda" rank="species" species="fasciculatus">
<emphasis id="7EF46300F403FFE011B005D6596F38EC" box="[464,619,1669,1694]" italics="true" pageId="11" pageNumber="1091">S. fasciculatus</emphasis>
</taxonomicName>
are closely related species. Both species have body longer than
<quantity id="8B7812F7F403FFE0110F05F45AB138B2" box="[367,437,1703,1728]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="11" pageNumber="1091" unit="mm" value="9.0">9 mm</quantity>
; dorsal surface shiny, almost smooth; genal edge not distinctly angulate laterally (
<figureCitation id="D4BBA397F404FFE7110E03DD5ACD3ED5" box="[366,457,142,167]" captionStart="Figure 3" captionStartId="7.[160,225,1294,1317]" captionTargetBox="[171,1141,146,1237]" captionTargetId="figure-89@7.[163,1153,142,1262]" captionTargetPageId="7" captionText="Figure 3. Head, dorsal view. (a) Scatonomus barbatus stat. rev.; (b) S. canhedoae sp. nov. (holotype); (c) S. chalybaeus; (d) S. fasciculatus; (e) S. foresti sp. nov. (holotype); (f) S. insignis; (g) S. janssensi; (h) S. mitzae sp. nov. (holotype); (i) S. paulosawayai (holotype); (j, k) S. thalassinus; (l) S. viridis." figureDoi="http://doi.org/10.5281/zenodo.5020909" httpUri="https://zenodo.org/record/5020909/files/figure.png" pageId="12" pageNumber="1092">Figure 3</figureCitation>
(a, d)); and clypeal edge bidentate, with deep lateral emarginations filled with a tuft of long setae (
<figureCitation id="D4BBA397F404FFE7120B03E259C93EB8" box="[619,717,177,202]" captionStart="Figure 3" captionStartId="7.[160,225,1294,1317]" captionTargetBox="[171,1141,146,1237]" captionTargetId="figure-89@7.[163,1153,142,1262]" captionTargetPageId="7" captionText="Figure 3. Head, dorsal view. (a) Scatonomus barbatus stat. rev.; (b) S. canhedoae sp. nov. (holotype); (c) S. chalybaeus; (d) S. fasciculatus; (e) S. foresti sp. nov. (holotype); (f) S. insignis; (g) S. janssensi; (h) S. mitzae sp. nov. (holotype); (i) S. paulosawayai (holotype); (j, k) S. thalassinus; (l) S. viridis." figureDoi="http://doi.org/10.5281/zenodo.5020909" httpUri="https://zenodo.org/record/5020909/files/figure.png" pageId="12" pageNumber="1092">Figure 3</figureCitation>
(a, d)). However,
<taxonomicName id="8B80C491F404FFE713FC03E25F193EB8" authorityName="Erichson" authorityYear="1835" box="[924,1053,177,202]" class="Insecta" family="Scarabaeidae" genus="Scatonomus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="1092" phylum="Arthropoda" rank="species" species="barbatus">
<emphasis id="7EF46300F404FFE713FC03E25F193EB8" box="[924,1053,177,202]" italics="true" pageId="12" pageNumber="1092">S. barbatus</emphasis>
</taxonomicName>
may be distinguished from
<taxonomicName id="8B80C491F404FFE711E2038759193E9F" authorityName=", Erichson" authorityYear="1835" box="[386,541,212,237]" class="Insecta" family="Scarabaeidae" genus="Scatonomus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="1092" phylum="Arthropoda" rank="species" species="fasciculatus">
<emphasis id="7EF46300F404FFE711E2038759193E9F" box="[386,541,212,237]" italics="true" pageId="12" pageNumber="1092">S. fasciculatus</emphasis>
</taxonomicName>
by the following combination of characters: median clypeal teeth shorter than the lateral tuft of setae (
<figureCitation id="D4BBA397F404FFE7129303A558563F7D" box="[755,850,246,271]" captionStart="Figure 3" captionStartId="7.[160,225,1294,1317]" captionTargetBox="[171,1141,146,1237]" captionTargetId="figure-89@7.[163,1153,142,1262]" captionTargetPageId="7" captionText="Figure 3. Head, dorsal view. (a) Scatonomus barbatus stat. rev.; (b) S. canhedoae sp. nov. (holotype); (c) S. chalybaeus; (d) S. fasciculatus; (e) S. foresti sp. nov. (holotype); (f) S. insignis; (g) S. janssensi; (h) S. mitzae sp. nov. (holotype); (i) S. paulosawayai (holotype); (j, k) S. thalassinus; (l) S. viridis." figureDoi="http://doi.org/10.5281/zenodo.5020909" httpUri="https://zenodo.org/record/5020909/files/figure.png" pageId="12" pageNumber="1092">Figure 3</figureCitation>
(a)); anterior edge of gena rounded (
<figureCitation id="D4BBA397F404FFE7116F024A5A683F40" box="[271,364,281,306]" captionStart="Figure 3" captionStartId="7.[160,225,1294,1317]" captionTargetBox="[171,1141,146,1237]" captionTargetId="figure-89@7.[163,1153,142,1262]" captionTargetPageId="7" captionText="Figure 3. Head, dorsal view. (a) Scatonomus barbatus stat. rev.; (b) S. canhedoae sp. nov. (holotype); (c) S. chalybaeus; (d) S. fasciculatus; (e) S. foresti sp. nov. (holotype); (f) S. insignis; (g) S. janssensi; (h) S. mitzae sp. nov. (holotype); (i) S. paulosawayai (holotype); (j, k) S. thalassinus; (l) S. viridis." figureDoi="http://doi.org/10.5281/zenodo.5020909" httpUri="https://zenodo.org/record/5020909/files/figure.png" pageId="12" pageNumber="1092">Figure 3</figureCitation>
(a)); and dorsum with green reflections (
<figureCitation id="D4BBA397F404FFE71351024A588C3F40" box="[817,904,281,306]" captionStart="Figure 1" captionStartId="5.[160,225,1145,1168]" captionTargetBox="[194,1153,143,1113]" captionTargetId="figure-187@5.[163,1153,142,1113]" captionTargetPageId="5" captionText="Figure 1. Dorsal habitus. (a) S Scatonomus barbatus stat. rev.; (b) S. canhedoae sp. nov.; (holotype); (c) S. chalybaeus; (d) S. fasciculatus; (e) S. foresti sp. nov. (holotype); (f) S. insignis; (g) S. janssensi; (h) S. mitzae sp. nov. (holotype); (i) S. paulosawayai (holotype); (j, k) S. thalassinus; (l) S. viridis." figureDoi="http://doi.org/10.5281/zenodo.5020905" httpUri="https://zenodo.org/record/5020905/files/figure.png" pageId="12" pageNumber="1092">Figure 1</figureCitation>
(a)).
</paragraph>
</subSubSection>
<subSubSection id="049AEC99F404FFE710C0020D593D3B52" pageId="12" pageNumber="1092" type="description">
<paragraph id="4C3FBF12F404FFE710C0020D5A443F04" blockId="12.[160,1156,350,1312]" box="[160,320,350,374]" pageId="12" pageNumber="1092">
<emphasis id="7EF46300F404FFE710C0020D5A443F04" bold="true" box="[160,320,350,374]" italics="true" pageId="12" pageNumber="1092">Redescription</emphasis>
</paragraph>
<paragraph id="4C3FBF12F404FFE710C002D2593D3B52" blockId="12.[160,1156,350,1312]" pageId="12" pageNumber="1092">
<emphasis id="7EF46300F404FFE710C002D25BDE3FE8" bold="true" box="[160,218,385,410]" pageId="12" pageNumber="1092">Male</emphasis>
: length:
<quantity id="8B7812F7F404FFE7112D02D25AC53FE8" box="[333,449,385,410]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="12" pageNumber="1092" unit="mm" value="10.0" valueMax="11.0" valueMin="9.0">9–11 mm</quantity>
. Dorsum black with green reflections (
<figureCitation id="D4BBA397F404FFE713C202D25F063FE8" box="[930,1026,385,410]" captionStart="Figure 1" captionStartId="5.[160,225,1145,1168]" captionTargetBox="[194,1153,143,1113]" captionTargetId="figure-187@5.[163,1153,142,1113]" captionTargetPageId="5" captionText="Figure 1. Dorsal habitus. (a) S Scatonomus barbatus stat. rev.; (b) S. canhedoae sp. nov.; (holotype); (c) S. chalybaeus; (d) S. fasciculatus; (e) S. foresti sp. nov. (holotype); (f) S. insignis; (g) S. janssensi; (h) S. mitzae sp. nov. (holotype); (i) S. paulosawayai (holotype); (j, k) S. thalassinus; (l) S. viridis." figureDoi="http://doi.org/10.5281/zenodo.5020905" httpUri="https://zenodo.org/record/5020905/files/figure.png" pageId="12" pageNumber="1092">Figure 1</figureCitation>
(a)).
<emphasis id="7EF46300F404FFE7145F02D25F7A3FE8" bold="true" box="[1087,1150,385,410]" pageId="12" pageNumber="1092">Head</emphasis>
: densely punctate, slightly wrinkled medially (
<figureCitation id="D4BBA397F404FFE712C802F758023FCF" box="[680,774,420,445]" captionStart="Figure 3" captionStartId="7.[160,225,1294,1317]" captionTargetBox="[171,1141,146,1237]" captionTargetId="figure-89@7.[163,1153,142,1262]" captionTargetPageId="7" captionText="Figure 3. Head, dorsal view. (a) Scatonomus barbatus stat. rev.; (b) S. canhedoae sp. nov. (holotype); (c) S. chalybaeus; (d) S. fasciculatus; (e) S. foresti sp. nov. (holotype); (f) S. insignis; (g) S. janssensi; (h) S. mitzae sp. nov. (holotype); (i) S. paulosawayai (holotype); (j, k) S. thalassinus; (l) S. viridis." figureDoi="http://doi.org/10.5281/zenodo.5020909" httpUri="https://zenodo.org/record/5020909/files/figure.png" pageId="12" pageNumber="1092">Figure 3</figureCitation>
(a)). Anterior and posterior edges narrowly margined. Clypeo-genal and fronto-clypeal sutures arched, moderately impressed, not carinate. Clypeal median teeth upturned, separated by a deep U-shaped emargination (
<figureCitation id="D4BBA397F404FFE71129015F5AAC3C57" box="[329,424,524,549]" captionStart="Figure 3" captionStartId="7.[160,225,1294,1317]" captionTargetBox="[171,1141,146,1237]" captionTargetId="figure-89@7.[163,1153,142,1262]" captionTargetPageId="7" captionText="Figure 3. Head, dorsal view. (a) Scatonomus barbatus stat. rev.; (b) S. canhedoae sp. nov. (holotype); (c) S. chalybaeus; (d) S. fasciculatus; (e) S. foresti sp. nov. (holotype); (f) S. insignis; (g) S. janssensi; (h) S. mitzae sp. nov. (holotype); (i) S. paulosawayai (holotype); (j, k) S. thalassinus; (l) S. viridis." figureDoi="http://doi.org/10.5281/zenodo.5020909" httpUri="https://zenodo.org/record/5020909/files/figure.png" pageId="12" pageNumber="1092">Figure 3</figureCitation>
(a)). Anterior portion of clypeus strongly concave medially, surface of concavity less punctate than remaining portions of head. Edges of clypeus and gena densely coated with rows of long erect setae. Antennal club globose, densely coated with long setae.
<emphasis id="7EF46300F404FFE711DE012759163CFF" bold="true" box="[446,530,628,653]" pageId="12" pageNumber="1092">Thorax</emphasis>
: pronotum shiny, sparsely punctate, mid-longitudinal sulcus feebly impressed on disc. Lateral edge of hypomera with a row of indistinct setose punctures, setae short. Transverse carina of hypomera almost complete, not reaching lateral edge. Surface of mesepimeron and mesanepisternum with green reflections, finely striate. Anterior and lateral lobes of metaventrite with inconspicuous microgranulation; lateral lobes also with shallow punctures. Metaventral disc densely puncticulate.
<emphasis id="7EF46300F404FFE7115E00175A803D2F" bold="true" box="[318,388,836,861]" pageId="12" pageNumber="1092">Elytra</emphasis>
: striae thin, shallow, indistinctly punctate; striae on disc 7 times narrower than third interstria. Seventh stria not reaching the basal and apical portions of elytra. Eighth stria weakly impressed, visible at the apical half. Ninth stria weakly impressed near elytral base. Interstriae flat, surface shiny and sparsely punctate.
<emphasis id="7EF46300F404FFE7142C00FF5A6F3D9A" bold="true" pageId="12" pageNumber="1092">Legs [ventral surface]</emphasis>
: profemora densely punctate, surface between punctures smooth. Mesofemur and metafemur sparsely punctate. External edge of protibia with 3 distinct teeth separated by U-shaped emarginations. Outer edge of mesotibia and metatibia with a row of sparse setae and small teeth throughout their length.
<emphasis id="7EF46300F404FFE713C807645F1A3A22" bold="true" box="[936,1054,1079,1104]" pageId="12" pageNumber="1092">Abdomen</emphasis>
: ventrite VI 1.5 times as long as ventrite V.
<emphasis id="7EF46300F404FFE7122F070A59C63A00" bold="true" box="[591,706,1113,1138]" pageId="12" pageNumber="1092">Pygidium</emphasis>
: 1/3 wider than long. Disc punctate, punctures spaced by 3 times their diameter.
<emphasis id="7EF46300F404FFE712CE072F586D3AE7" bold="true" box="[686,873,1148,1173]" pageId="12" pageNumber="1092">Genital capsule</emphasis>
: paramera apex divided into apical and lateral lobes by a deep L-shaped emargination. Apical and lateral lobes slightly acute (
<figureCitation id="D4BBA397F404FFE7112507925AA53AA8" box="[325,417,1217,1242]" captionStart="Figure 5" captionStartId="9.[160,225,1191,1214]" captionTargetBox="[224,1152,142,1131]" captionTargetId="figure-205@9.[163,1153,142,1159]" captionTargetPageId="9" captionText="Figure 5. Aedeagus, detail of right paramera and endophallic sclerites. (a) Scatonomus barbatus stat. rev.; (b) S. chalybaeus; (c) S. fasciculatus; (d) S. foresti sp. nov.; (e) S. insignis; (f) S. janssensi; (g) S. mitzae sp. nov.; (h) S. thalassinus; (i) S. viridis; (j) endophallic sclerites of S. fasciculatus." figureDoi="http://doi.org/10.5281/zenodo.5020913" httpUri="https://zenodo.org/record/5020913/files/figure.png" pageId="12" pageNumber="1092">Figure 5</figureCitation>
(a)).
<emphasis id="7EF46300F404FFE711B20792592D3AA8" bold="true" box="[466,553,1217,1242]" pageId="12" pageNumber="1092">Female</emphasis>
: differs from male by the abdominal ventrite VI twice as long as V. Disc of pygidium with punctures spaced by 2 times their diameter; apical edge of pygidium slightly angulate.
</paragraph>
</subSubSection>
</treatment>
</document>
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