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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.101.1182" ID-GBIF-Dataset="27b97ed3-f3cd-4468-af5e-0c3701b28f46" ID-PMC="PMC3118703" ID-Pensoft-Pub="1313-2970-101-51" ID-PubMed="21747670" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-101-51" ModsDocOrigin="ZooKeys 101" ModsDocTitle="Alocanthedon, a new subgenus of Chalicodoma from Southeast Asia (Hymenoptera, Megachilidae)" checkinTime="1451250346471" checkinUser="pensoft" docAuthor="Engel, Michael S. &amp; Gonzalez, Victor H." docDate="2011" docId="9B7588E6A1C5B2380954F7E83A84EC3D" docLanguage="en" docName="ZooKeys 101: 51-80" docOrigin="ZooKeys 101" docSource="http://dx.doi.org/10.3897/zookeys.101.1182" docTitle="Chalicodoma ((Alocanthedon)) atratiforme, comb. n." docType="treatment" docVersion="3" lastPageNumber="67" masterDocId="FFE06A70FFD9FFB6D71F3A05AE130042" masterDocTitle="Alocanthedon, a new subgenus of Chalicodoma from Southeast Asia (Hymenoptera, Megachilidae)" masterLastPageNumber="80" masterPageNumber="51" pageNumber="66" updateTime="1668149769182" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Alocanthedon, a new subgenus of Chalicodoma from Southeast Asia (Hymenoptera, Megachilidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Engel, Michael S.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gonzalez, Victor H.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2011</mods:date>
<mods:detail type="volume">
<mods:number>101</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>51</mods:start>
<mods:end>80</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.101.1182</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.101.1182</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-101-51</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152029511" LSID="urn:lsid:plazi:treatment:9B7588E6A1C5B2380954F7E83A84EC3D" httpUri="http://treatment.plazi.org/id/9B7588E6A1C5B2380954F7E83A84EC3D" lastPageId="16" lastPageNumber="67" pageId="15" pageNumber="66">
<subSubSection pageId="15" pageNumber="66" type="nomenclature">
<paragraph pageId="15" pageNumber="66">
<taxonomicName LSID="http://species-id.net/wiki/Chalicodoma_(Alocanthedon)_atratiforme" authority="Meade-Waldo" class="Insecta" family="Megachilidae" genus="Chalicodoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chalicodoma ((Alocanthedon)) atratiforme" order="Hymenoptera" pageId="15" pageNumber="66" phylum="Arthropoda" rank="species" species="atratiforme" subGenus="(Alocanthedon)">Chalicodoma (Alocanthedon) atratiforme (Meade-Waldo)</taxonomicName>
<taxonomicNameLabel pageId="15" pageNumber="66">comb. n.</taxonomicNameLabel>
Figs 32
<normalizedToken originalValue="–3444–">-3444-</normalizedToken>
46
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="66" type="reference_group">
<paragraph pageId="15" pageNumber="66">
<taxonomicName class="Insecta" family="Megachilidae" genus="Megachile" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megachile atratiformis" order="Hymenoptera" pageId="15" pageNumber="66" phylum="Arthropoda" rank="species" species="atratiformis">Megachile atratiformis</taxonomicName>
<bibRefCitation author="Meade-Waldo, G" journalOrPublisher="Annals and Magazine of Natural History, Series 8," pageId="20" pageNumber="71" pagination="450 - 464" title="Notes on the Hymenoptera in the collection of the British Museum, with descriptions of new species." volume="14" year="1914">Meade-Waldo 1914</bibRefCitation>
: 456.
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="66" type="additional material">
<paragraph pageId="15" pageNumber="66">Additional material.</paragraph>
<paragraph pageId="15" pageNumber="66">
Malaysia (Peninsular): 3♀♀, Pahang, Batu Balai Estate, 18 March 1927, E. Seimund, Ex F.M.S. [Federated Malay States] Museum (NHML); 1♂, 1♀, Negri Sembilan, Gunong Angsi, 2000
<normalizedToken originalValue="–2790’">-2790'</normalizedToken>
[feet], April 1918, Ex F.M.S. [Federated Malay States] Museum (NHML).
</paragraph>
<paragraph pageId="15" pageNumber="66">
Thailand: 1♀, Uthai Thani Province [western Thailand], Huay Kha Khaeng Wildlife Sanctuary, 15
<normalizedToken originalValue="º36’">°36'</normalizedToken>
N, 99
<normalizedToken originalValue="º20’">°20'</normalizedToken>
E, 1.xi.1995 [1 November 1995], J. Gazhoul, captured at
<taxonomicName class="Magnoliopsida" family="Dipterocarpaceae" genus="Dipterocarpus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dipterocarpus obtusifolius" order="Malvales" pageId="15" pageNumber="66" phylum="Tracheophyta" rank="species" species="obtusifolius">Dipterocarpus obtusifolius</taxonomicName>
Teijsman &amp; Miquel (
<taxonomicName genus="Dipterocarpaceae" lsidName="Dipterocarpaceae" pageId="15" pageNumber="66" rank="genus">Dipterocarpaceae</taxonomicName>
), D.B. &amp; M.W. Baker Collection (SEMC); 1♀, 150 n.w. Bangkok, Huay Kha Khaeng, 5.xi.1995 [5 November 1995], J. Ghazoul (NHML).
</paragraph>
</subSubSection>
<subSubSection lastPageId="16" lastPageNumber="67" pageId="15" pageNumber="66" type="diagnosis">
<paragraph pageId="15" pageNumber="66">Diagnosisn.</paragraph>
<paragraph lastPageId="16" lastPageNumber="67" pageId="15" pageNumber="66">
The female of this species can be recognized by the clypeus lacking a medioapical tubercle, short mandibles (Fig. 45), labrum with an apical fringe of erect setae running along relatively straight apical margin, and the yellow wings (Fig. 44). The male can be recognized by the dense setal patch in forewing medial cell, tergum
<pageBreakToken pageId="16" pageNumber="67" start="start">VI</pageBreakToken>
with shallow medioapical concavity on preapical carina, protibia with outer, apical surface not depressed, and genitalia as in figures 32-34.
</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="67" type="remarks">
<paragraph pageId="16" pageNumber="67">Remarks.</paragraph>
<paragraph pageId="16" pageNumber="67">
It should be noted that
<normalizedToken originalValue="Meade-Waldo’s">Meade-Waldo's</normalizedToken>
(1914) material from &quot;Middle Tenasserim&quot; is not conspecific with his type (B.M. Type Hym.17a2037) for
<taxonomicName class="Insecta" family="Megachilidae" genus="Chalicodoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chalicodoma atratiforme" order="Hymenoptera" pageId="16" pageNumber="67" phylum="Arthropoda" rank="species" species="atratiforme">Chalicodoma atratiforme</taxonomicName>
and are actually specimens of
<taxonomicName class="Insecta" family="Megachilidae" genus="Chalicodoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chalicodoma odontophorum" order="Hymenoptera" pageId="16" pageNumber="67" phylum="Arthropoda" rank="species" species="odontophorum">Chalicodoma odontophorum</taxonomicName>
. Additionally, despite
<bibRefCitation author="Cockerell, TDA" journalOrPublisher="Proceedings of the Entomological Society of Washington" pageId="20" pageNumber="71" pagination="160 - 162" title="Bees collected by Dr. H. M. Smith on Turtle Island (Koh Tao), Gulf of Siam." volume="29" year="1927">
<normalizedToken originalValue="Cockerell’s">Cockerell's</normalizedToken>
(1927)
</bibRefCitation>
assignment of the subspecies
<taxonomicName class="Insecta" family="Megachilidae" genus="Chalicodoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chalicodoma atratiforme subsp. sininsulae" order="Hymenoptera" pageId="16" pageNumber="67" phylum="Arthropoda" rank="subSpecies" species="atratiforme" subSpecies="sininsulae">Chalicodoma atratiforme sininsulae</taxonomicName>
(Cockerell) (Type USNM 70455; ♀; Turtle Island [Koh Tao], Gulf of Siam; visum) to
<normalizedToken originalValue="Meade-Waldo’s">Meade-Waldo's</normalizedToken>
species, his female has nothing whatever to do with
<taxonomicName class="Insecta" family="Megachilidae" genus="Chalicodoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chalicodoma atratiforme" order="Hymenoptera" pageId="16" pageNumber="67" phylum="Arthropoda" rank="species" species="atratiforme">Chalicodoma atratiforme</taxonomicName>
and is instead a synonym of
<taxonomicName class="Insecta" family="Megachilidae" genus="Callomegachile" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Callomegachile (Callomegachile) fulvipenne" order="Hymenoptera" pageId="16" pageNumber="67" phylum="Arthropoda" rank="species" species="fulvipenne" subGenus="Callomegachile">Callomegachile (Callomegachile) fulvipenne</taxonomicName>
(
<bibRefCitation author="Smith, F" journalOrPublisher="Taylor and Francis, London" pageId="20" pageNumber="71" title="Descriptions of New Species of Hymenoptera in the Collection of the British Museum." year="1879">Smith 1879</bibRefCitation>
) (syn. n.).
</paragraph>
<caption pageId="16" pageNumber="67">
<paragraph pageId="16" pageNumber="67">
Figures 35-40. Photomicrographs of male of
<taxonomicName class="Insecta" family="Megachilidae" genus="Chalicodoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chalicodoma" order="Hymenoptera" pageId="16" pageNumber="67" phylum="Arthropoda" rank="genus">Chalicodoma</taxonomicName>
(
<taxonomicName class="Insecta" family="Megachilidae" genus="Alocanthedon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Alocanthedon" order="Hymenoptera" pageId="16" pageNumber="67" phylum="Arthropoda" rank="genus">Alocanthedon</taxonomicName>
) aterrimum (Smith) 35 Dorsal habitus 36 Facial aspect 37 Dorsal aspect of mesoscutum and mesoscutellum (small portion of first metasomal tergum also visible) 38 Protarsus, pro-pretarsus, and apex of protibia 39 Detail of forewing medial cell 40 Dorsal aspect of metasomal terga V, VI, and apical portion of IV.
</paragraph>
</caption>
<caption pageId="16" pageNumber="67">
<paragraph pageId="16" pageNumber="67">
Figures 41-43. Photomicrographs of female of
<taxonomicName class="Insecta" family="Megachilidae" genus="Chalicodoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chalicodoma" order="Hymenoptera" pageId="16" pageNumber="67" phylum="Arthropoda" rank="genus">Chalicodoma</taxonomicName>
(
<taxonomicName class="Insecta" family="Megachilidae" genus="Alocanthedon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Alocanthedon" order="Hymenoptera" pageId="16" pageNumber="67" phylum="Arthropoda" rank="genus">Alocanthedon</taxonomicName>
) aterrimum (Smith) 41 Dorsal habitus 42 Facial aspect 43 Dorsal aspect of mesoscutum and mesoscutellum.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="16" pageNumber="67" type="">
<paragraph pageId="16" pageNumber="67">Key to Species of Alocanthedon</paragraph>
<paragraph pageId="16" pageNumber="67">
Note: The female of
<taxonomicName class="Insecta" family="Megachilidae" genus="Chalicodoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chalicodoma apoicola" order="Hymenoptera" pageId="16" pageNumber="67" phylum="Arthropoda" rank="species" species="apoicola">Chalicodoma apoicola</taxonomicName>
is unknown.
</paragraph>
<paragraph pageId="16" pageNumber="67">
<table pageId="16" pageNumber="67">
<tr pageId="16" pageNumber="67">
<td colspan="1" pageId="16" pageNumber="67" rowspan="1">Figs. 3541Fig. 10Figs. 1040</td>
</tr>
<tr pageId="16" pageNumber="67">
<td colspan="1" pageId="16" pageNumber="67" rowspan="1">Figs. 17172344Figs. 622</td>
</tr>
<tr pageId="16" pageNumber="67">
<td colspan="1" pageId="16" pageNumber="67" rowspan="1">Fig. 36figure 38</td>
<td colspan="1" pageId="16" pageNumber="67" rowspan="1">
<taxonomicName class="Insecta" family="Megachilidae" genus="Chalicodoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chalicodoma aterrimum" order="Hymenoptera" pageId="16" pageNumber="67" phylum="Arthropoda" rank="species" species="aterrimum">Chalicodoma aterrimum</taxonomicName>
</td>
</tr>
<tr pageId="16" pageNumber="67">
<td colspan="1" pageId="16" pageNumber="67" rowspan="1">Fig. 12figure 14</td>
<td colspan="1" pageId="16" pageNumber="67" rowspan="1">
<taxonomicName class="Insecta" family="Megachilidae" genus="Chalicodoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chalicodoma apoicola" order="Hymenoptera" pageId="16" pageNumber="67" phylum="Arthropoda" rank="species" species="apoicola">Chalicodoma apoicola</taxonomicName>
</td>
</tr>
<tr pageId="16" pageNumber="67">
<td colspan="1" pageId="16" pageNumber="67" rowspan="1">Fig. 8Fig. 8Fig. 2figure 5</td>
<td colspan="1" pageId="16" pageNumber="67" rowspan="1">
<taxonomicName class="Insecta" family="Megachilidae" genus="Chalicodoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chalicodoma odontophorum" order="Hymenoptera" pageId="16" pageNumber="67" phylum="Arthropoda" rank="species" species="odontophorum">Chalicodoma odontophorum</taxonomicName>
</td>
</tr>
<tr pageId="16" pageNumber="67">
<td colspan="1" pageId="16" pageNumber="67" rowspan="1">Figs. 2545Fig. 18figure 5</td>
</tr>
<tr pageId="16" pageNumber="67">
<td colspan="1" pageId="16" pageNumber="67" rowspan="1">Fig. 25Figs. 17, 20figures 29-31</td>
<td colspan="1" pageId="16" pageNumber="67" rowspan="1">
<taxonomicName class="Insecta" family="Megachilidae" genus="Chalicodoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chalicodoma memecylonae" order="Hymenoptera" pageId="16" pageNumber="67" phylum="Arthropoda" rank="species" species="memecylonae">Chalicodoma memecylonae</taxonomicName>
</td>
</tr>
<tr pageId="16" pageNumber="67">
<td colspan="1" pageId="16" pageNumber="67" rowspan="1">Fig. 46figures 32-34</td>
<td colspan="1" pageId="16" pageNumber="67" rowspan="1">
<taxonomicName class="Insecta" family="Megachilidae" genus="Chalicodoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chalicodoma atratiforme" order="Hymenoptera" pageId="16" pageNumber="67" phylum="Arthropoda" rank="species" species="atratiforme">Chalicodoma atratiforme</taxonomicName>
</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</document>
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