346 lines
38 KiB
XML
346 lines
38 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1077.67081" ID-GBIF-Dataset="66b827c0-65ac-44a6-90e0-b78cd9ef81c2" ID-PMC="PMC8692307" ID-Pensoft-Pub="1313-2970-1077-1" ID-Pensoft-UUID="734088641608531C8E2CC69397B000ED" ID-PubMed="35027858" ID-ZooBank="C28AD65A76F242CFBED7DFB3702CABCE" ModsDocID="1313-2970-1077-1" checkinTime="1639570129388" checkinUser="pensoft" docAuthor="De Mattia, Willy, Reier, Susanne & Haring, Elisabeth" docDate="2021" docId="3B4D1DDD1672500C89696130F7AE9AEB" docLanguage="en" docName="ZooKeys 1077: 1-175" docOrigin="ZooKeys 1077" docPubDate="2021-12-14" docSource="http://dx.doi.org/10.3897/zookeys.1077.67081" docTitle="Siciliaria tiberii subsp. alcamoensis Brandt 1961, comb. nov." docType="treatment" docVersion="4" id="734088641608531C8E2CC69397B000ED" lastPageNumber="1" masterDocId="734088641608531C8E2CC69397B000ED" masterDocTitle="Morphological investigation of genital organs and first insights into the phylogeny of the genus Siciliaria Vest, 1867 as a basis for a taxonomic revision (Mollusca, Gastropoda, Clausiliidae)" masterLastPageNumber="175" masterPageNumber="1" pageNumber="1" updateTime="1668151235153" updateUser="ExternalLinkService">
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<mods:titleInfo>
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<mods:title>Morphological investigation of genital organs and first insights into the phylogeny of the genus Siciliaria Vest, 1867 as a basis for a taxonomic revision (Mollusca, Gastropoda, Clausiliidae)</mods:title>
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>De Mattia, Willy</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-0056-467X</mods:nameIdentifier>
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<mods:affiliation>Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
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<mods:nameIdentifier type="email">willy.demattia@icgeb.org</mods:nameIdentifier>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Reier, Susanne</mods:namePart>
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<mods:affiliation>Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Haring, Elisabeth</mods:namePart>
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<mods:affiliation>Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-12-14</mods:number>
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<mods:detail type="volume">
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<mods:number>1077</mods:number>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>175</mods:end>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1077.67081</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1077.67081</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1077-1</mods:identifier>
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<mods:identifier type="ZooBank">C28AD65A76F242CFBED7DFB3702CABCE</mods:identifier>
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<mods:identifier type="Pensoft-UUID">734088641608531C8E2CC69397B000ED</mods:identifier>
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<treatment ID-GBIF-Taxon="191645972" LSID="urn:lsid:plazi:treatment:3B4D1DDD1672500C89696130F7AE9AEB" httpUri="http://treatment.plazi.org/id/3B4D1DDD1672500C89696130F7AE9AEB" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="3B4D1DDD-1672-500C-8969-6130F7AE9AEB" authority="Brandt, 1961" authorityName="Brandt" authorityYear="1961" baseAuthorityName="De Mattia & Reier & Haring" baseAuthorityYear="1961" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria tiberii subsp. alcamoensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="tiberii" status="comb. nov." subSpecies="alcamoensis">Siciliaria tiberii alcamoensis Brandt, 1961</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">comb. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Wagner's (1913, 1925) and Brandt's (1961) Siciliaria genital drawings A Siciliaria septemplicata, Wagner, 1913: plate 572, fig. 14 B Siciliaria grohmanniana grohmanniana, Wagner, 1925: plate 1, fig. 8 C Siciliaria calcarae calcarae, 1925: plate 3, fig. 25 D Siciliaria ferrox, Brandt, 1961: fig. 1 E overall Siciliaria / Sicania and Sicilian Mauritanica distribution F Siciliaria / Sicania and Sicilian Mauritanica distribution of taxa." figureDoi="10.3897/zookeys.1077.67081.figure1" httpUri="https://binary.pensoft.net/fig/622793" pageId="0" pageNumber="1">Figs 1.F, 25.3-25.5, 26.6-26.9, 29.4, 29.5</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="reference_group">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="De Mattia & Reier & Haring" authorityYear="2021" baseAuthorityName="De Mattia & Reier & Haring" baseAuthorityYear="2021" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria (Siciliaria) alcamoensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="alcamoensis" status="comb. nov." subGenus="Siciliaria">Siciliaria (Siciliaria) alcamoensis</taxonomicName>
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<bibRefCitation author="Brandt, R" journalOrPublisher="Bullettino della Societa Malacologica Italiana" pageId="0" pageNumber="1" refId="B10" refString="Brandt, R, 1961. Diagnosen neuer Clausiliiden. Archiv fuer Molluskenkunde 90(1/3): 1-20." title="Diagnosen neuer Clausiliiden. Archiv fuer Molluskenkunde 90 (1 / 3): 1 - 20." year="1961">Brandt 1961</bibRefCitation>
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: 9.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="De Mattia & Reier & Haring" authorityYear="2021" baseAuthorityName="De Mattia & Reier & Haring" baseAuthorityYear="1961" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria (Siciliaria) septemplicata subsp. alcamoensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="septemplicata" status="comb. nov." subGenus="Siciliaria" subSpecies="alcamoensis">Siciliaria (Siciliaria) septemplicata alcamoensis</taxonomicName>
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-
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<bibRefCitation author="Manganelli, G" journalOrPublisher="Biodiversity Journal" pageId="0" pageNumber="1" refId="B43" refString="Manganelli, G, Bodon, M, Favilli, L, Giusti, F, 1995. Fascicolo 16, Gastropoda Pulmonta. In: Minelli A, Ruffo S, La Posta S (Eds) Checklist delle specie della fauna italiana 16: 1-60. Calderini, Bologna." title="Fascicolo 16, Gastropoda Pulmonta. In: Minelli A, Ruffo S, La Posta S (Eds) Checklist delle specie della fauna italiana 16: 1 - 60. Calderini, Bologna." year="1995">Manganelli et al. 1995</bibRefCitation>
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: 25.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="De Mattia & Reier & Haring" authorityYear="2021" baseAuthorityName="De Mattia & Reier & Haring" baseAuthorityYear="2021" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria septemplicata subsp. hemmeni" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="septemplicata" status="comb. nov." subSpecies="hemmeni">Siciliaria septemplicata hemmeni</taxonomicName>
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<bibRefCitation DOI="https://doi.org/10.1127/arch.moll/133/2004/185" author="Beckmann, KH" journalOrPublisher="Basteria" pageId="0" pageNumber="1" refId="B4" refString="Beckmann, KH, 2004. Zur Verbreitung der endemischen nordwestsizilianischen Clausiliidae der Untergattung Charpentieria (Siciliaria) mit Beschreibung von zwei neuen Unterarten (Gastropoda: Stylommatophora: Clausiliidae). Archiv fuer Molluskenkunde 133(1/2): 185-191. https://doi.org/10.1127/arch.moll/133/2004/185" title="Zur Verbreitung der endemischen nordwestsizilianischen Clausiliidae der Untergattung Charpentieria (Siciliaria) mit Beschreibung von zwei neuen Unterarten (Gastropoda: Stylommatophora: Clausiliidae). Archiv fuer Molluskenkunde 133 (1 / 2): 185 - 191." url="https://doi.org/10.1127/arch.moll/133/2004/185" year="2004">Beckmann 2004</bibRefCitation>
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: 190.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="De Mattia & Reier & Haring" authorityYear="2021" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria (Siciliaria) septemplicata subsp. alcamoensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="septemplicata" status="comb. nov." subGenus="Siciliaria" subSpecies="alcamoensis">Charpentieria (Siciliaria) septemplicata alcamoensis</taxonomicName>
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-
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<bibRefCitation author="Nordsieck, H" journalOrPublisher="Mitteilungen der deutschen malakozoologischen Gesellschaft" pageId="0" pageNumber="1" refId="B65" refString="Nordsieck, H, 2007. Worldwide Door Snails (Clausiliidae), recent and fossil. Conchbooks Hackenheim, 214 pp." title="Worldwide Door Snails (Clausiliidae), recent and fossil. Conchbooks Hackenheim, 214 pp." year="2007">Nordsieck 2007</bibRefCitation>
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: 54.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="De Mattia & Reier & Haring" authorityYear="2021" baseAuthorityName="De Mattia & Reier & Haring" baseAuthorityYear="1961" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria septemplicata subsp. alcamoensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="septemplicata" status="comb. nov." subSpecies="alcamoensis">Siciliaria septemplicata alcamoensis</taxonomicName>
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-
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<bibRefCitation author="Nordsieck, H" journalOrPublisher="Mitteilungen der deutschen malakozoologischen Gesellschaft" pageId="0" pageNumber="1" publicationUrl="http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" refId="B69" refString="Nordsieck, H, 2013b. Revisory remarks on the species of Siciliaria Vest from N. W. Sicily. http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" title="Revisory remarks on the species of Siciliaria Vest from N. W. Sicily." url="http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" year="2013 b">Nordsieck 2013b</bibRefCitation>
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: 10.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="examined specimens">
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<paragraph pageId="0" pageNumber="1">Examined specimens.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Italy, Sicily, Cinisi, Piano Margi, 670 m asl,
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<geoCoordinate degrees="38" direction="north" minutes="08" orientation="latitude" precision="1" seconds="58.56" value="38.1496">38°08'58.56"N</geoCoordinate>
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,
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<geoCoordinate degrees="13" direction="east" minutes="09" orientation="longitude" precision="1" seconds="25.83" value="13.157175">13°09'25.83"E</geoCoordinate>
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, [Lab ID 37_1, COI: MW758928, ITS2: MW757102, MW757101], W. De Mattia and J. Macor leg., 14.iv.2017. 2 dissected spm.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="shell">
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<paragraph pageId="0" pageNumber="1">Shell</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">
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(Figs
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<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Siciliaria tiberii tiberii (A. Schmidt, 1868), Capo Rama, Terrasini 26.1 shell 26.2 clausiliar plate double side 26.3 shell 26.4 detail of the aperture 26.5 clausiliar plate double side. Siciliaria tiberii alcamoensis Brandt, 1961 comb. nov., Piano Margi, Carini 26.6 shell 26.7 clausiliar plate double side 26.8 detail of the aperture 26.9 shell." figureDoi="10.3897/zookeys.1077.67081.figure26" httpUri="https://binary.pensoft.net/fig/622818" pageId="0" pageNumber="1">26</figureCitation>
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.6-26.9, 29.4-29.5).
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</emphasis>
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Shell decollate; whorls striated to rib-striated; dorsal keel very prominent; inferior lamella high; anterior upper palatal plica present, mostly separated from upper palatal plica, sometimes a second or also third upper palatal plica present; anterior lower palatal plica strong; palatal edge of clausilium plate distally not receding, palatal edge distally more or less strongly bent upwards.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Measurements</paragraph>
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<paragraph pageId="0" pageNumber="1">
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(n = 18, decollate): shell height 21.5
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<normalizedToken originalValue="±">+/-</normalizedToken>
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1.3, whorl width 5.2
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.3, aperture height 4.6
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.2, aperture width 3.4
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.4.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="external morphology of the genital organs">
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<paragraph pageId="0" pageNumber="1">External morphology of the genital organs</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">
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(Fig.
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<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Siciliaria tiberii tiberii (A. Schmidt, 1868), Capo Rama, Terrasini 25.1 whole distal genital organs 25.2 internal distal part of genital organs. Siciliaria tiberii alcamoensis Brandt, 1961 comb. nov., Piano Margi, Carini 25.3 whole distal genital organs 25.4 internal distal part of genital organs 25.5 detail of the pseudopapilla." figureDoi="10.3897/zookeys.1077.67081.figure25" httpUri="https://binary.pensoft.net/fig/622817" pageId="0" pageNumber="1">25</figureCitation>
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.3).
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</emphasis>
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The FO is longer than the V (FO/V range 1.4-1.6). The VD is thin along its whole course. The FDBC is as long as the BC+SDBC (DBC/BC+SDBC = 1.0). The BC+SDBC is spindle-like to cylindrical and longer than the V (BC+SDBC/V range 1.4-1.5), with no distinction between the SDBC and the BC. The apex is pointed. The D is longer than the V (D/V range 2.5-2.6) and longer that the BC+SDBC (D/BC+SDBC range 1.6-1.8), thinner than the BC+SDBC and with a pointed apex. The V is long, cylindrical and small in diameter. The A is large. The PC is much longer than the V (P+E/V range 2.4-2.6). The PR is long and thin. The ET is clearly visible. There is a clear distinction between P and E, there is a visible ER, and a proximal narrowing. The E is longer than the P (E/P range 2.0-2.1), gradually shrinking and turning into the VD.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="internal morphology of the genital organs">
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<paragraph pageId="0" pageNumber="1">Internal morphology of the genital organs</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">
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(Figs
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<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Siciliaria tiberii tiberii (A. Schmidt, 1868), Capo Rama, Terrasini 25.1 whole distal genital organs 25.2 internal distal part of genital organs. Siciliaria tiberii alcamoensis Brandt, 1961 comb. nov., Piano Margi, Carini 25.3 whole distal genital organs 25.4 internal distal part of genital organs 25.5 detail of the pseudopapilla." figureDoi="10.3897/zookeys.1077.67081.figure25" httpUri="https://binary.pensoft.net/fig/622817" pageId="0" pageNumber="1">25</figureCitation>
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.4-25.5).
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</emphasis>
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The A shows a set of irregular fleshy folds. The P presents two or three main large longitudinal pleats. These pleats are heavily transversely segmented, getting a comb-like shape. The fine structure of the wall is smooth.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">The PP is large, elongated, and heavily wrinkled. The P-E transition presents one ER with both the PP and ELP originating from the ER. The epiphallar formula is: 1ER(PP+ELP). The E shows a set of three or four smooth longitudinal pleats. The V is almost completely smooth. The wall shows a fine granulation.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="ecology">
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<paragraph pageId="0" pageNumber="1">Ecology.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Brandt" authorityYear="1961" baseAuthorityName="De Mattia & Reier & Haring" baseAuthorityYear="1961" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria tiberii subsp. alcamoensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="tiberii" subSpecies="alcamoensis">
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<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria tiberii alcamoensis</emphasis>
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</taxonomicName>
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comb. nov. inhabits scattered isolated limestone boulders, and is hiding in rock cracks and crevices. After careful field research in a wide area around Piano Margi, the taxon revealed to be remarkably rare and scattered.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Brandt" authorityYear="1961" baseAuthorityName="De Mattia & Reier & Haring" baseAuthorityYear="1961" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria tiberii subsp. alcamoensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="tiberii" subSpecies="alcamoensis">
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<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria tiberii alcamoensis</emphasis>
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</taxonomicName>
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comb. nov. is known only from a small area of less than 1 km2 in the surrounding of the type locality: Piano Margi south of Cinisi on the Montagna Longa. Further field research is needed to define its actual distribution range.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="remarks">
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<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<bibRefCitation author="Brandt, R" journalOrPublisher="Bullettino della Societa Malacologica Italiana" pageId="0" pageNumber="1" refId="B10" refString="Brandt, R, 1961. Diagnosen neuer Clausiliiden. Archiv fuer Molluskenkunde 90(1/3): 1-20." title="Diagnosen neuer Clausiliiden. Archiv fuer Molluskenkunde 90 (1 / 3): 1 - 20." year="1961">Brandt (1961</bibRefCitation>
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: 9) described
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<taxonomicName authorityName="De Mattia & Reier & Haring" authorityYear="2021" baseAuthorityName="De Mattia & Reier & Haring" baseAuthorityYear="2021" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria (Siciliaria) alcamoensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="alcamoensis" subGenus="Siciliaria">Siciliaria (Siciliaria) alcamoensis</taxonomicName>
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based on material from the Senckenberg Museum (SMF163949), provided by Monterosato and labelled as
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<taxonomicName authorityName="Retowski" authorityYear="1889" class="Gastropoda" family="Clausiliidae" genus="Clausilia" kingdom="Animalia" lsidName="Clausilia difficilis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="difficilis">
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<emphasis italics="true" pageId="0" pageNumber="1">Clausilia difficilis</emphasis>
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</taxonomicName>
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from "Alcamo in NW-Sizilien, ohne
|
||
<normalizedToken originalValue="nähere">naehere</normalizedToken>
|
||
Fundortangabe".
|
||
<bibRefCitation author="Brandt, R" journalOrPublisher="Bullettino della Societa Malacologica Italiana" pageId="0" pageNumber="1" refId="B10" refString="Brandt, R, 1961. Diagnosen neuer Clausiliiden. Archiv fuer Molluskenkunde 90(1/3): 1-20." title="Diagnosen neuer Clausiliiden. Archiv fuer Molluskenkunde 90 (1 / 3): 1 - 20." year="1961">Brandt (1961</bibRefCitation>
|
||
: 9) introduced the name
|
||
<taxonomicName authorityName="De Mattia & Reier & Haring" authorityYear="2021" baseAuthorityName="De Mattia & Reier & Haring" baseAuthorityYear="2021" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria alcamoensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="alcamoensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria alcamoensis</emphasis>
|
||
</taxonomicName>
|
||
as
|
||
<taxonomicName authorityName="Retowski" authorityYear="1889" class="Gastropoda" family="Clausiliidae" genus="Clausilia" kingdom="Animalia" lsidName="Clausilia difficilis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="difficilis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Clausilia difficilis</emphasis>
|
||
</taxonomicName>
|
||
sensu Monterosato [non
|
||
<taxonomicName authorityName="Retowski" authorityYear="1889" class="Gastropoda" family="Clausiliidae" genus="Clausilia" kingdom="Animalia" lsidName="Clausilia difficilis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="difficilis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Clausilia difficilis</emphasis>
|
||
</taxonomicName>
|
||
Retowski, 1889, currently
|
||
<taxonomicName baseAuthorityName="Retowski" baseAuthorityYear="1889" class="Gastropoda" family="Clausiliidae" genus="Roseniella" kingdom="Animalia" lsidName="Roseniella difficilis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="difficilis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Roseniella difficilis</emphasis>
|
||
</taxonomicName>
|
||
(Retowski, 1889)] is a nomen nudum.
|
||
<bibRefCitation author="Brandt, R" journalOrPublisher="Bullettino della Societa Malacologica Italiana" pageId="0" pageNumber="1" refId="B10" refString="Brandt, R, 1961. Diagnosen neuer Clausiliiden. Archiv fuer Molluskenkunde 90(1/3): 1-20." title="Diagnosen neuer Clausiliiden. Archiv fuer Molluskenkunde 90 (1 / 3): 1 - 20." year="1961">Brandt (1961</bibRefCitation>
|
||
: 10) stated that he was not able to find this species again during field collecting nearby Alcamo, but he also lists in the type material: "Paratypoide Coll. Brandt Cl. 2365/2 leg. Belluci". He did not provide the collecting site of the paratype material so its origin is unknown, but it is conceivable that their shell morphology matches with what he described as
|
||
<taxonomicName authorityName="De Mattia & Reier & Haring" authorityYear="2021" baseAuthorityName="De Mattia & Reier & Haring" baseAuthorityYear="2021" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria (Siciliaria) alcamoensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="alcamoensis" subGenus="Siciliaria">Siciliaria (Siciliaria) alcamoensis</taxonomicName>
|
||
indeed. Undoubtedly, the locality indication provided by Monterosato on the original SMF label is wrong. Despite intensive field research (WDM) all over the Alcamo area and Bosco
|
||
<normalizedToken originalValue="d’Alcamo">d'Alcamo</normalizedToken>
|
||
/Monte Bonifato, no specimens/population was found that matches
|
||
<normalizedToken originalValue="Brandt’s">Brandt's</normalizedToken>
|
||
description of
|
||
<taxonomicName authorityName="De Mattia & Reier & Haring" authorityYear="2021" baseAuthorityName="De Mattia & Reier & Haring" baseAuthorityYear="2021" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria alcamoensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="alcamoensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria alcamoensis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation DOI="https://doi.org/10.1127/arch.moll/133/2004/185" author="Beckmann, KH" journalOrPublisher="Basteria" pageId="0" pageNumber="1" refId="B4" refString="Beckmann, KH, 2004. Zur Verbreitung der endemischen nordwestsizilianischen Clausiliidae der Untergattung Charpentieria (Siciliaria) mit Beschreibung von zwei neuen Unterarten (Gastropoda: Stylommatophora: Clausiliidae). Archiv fuer Molluskenkunde 133(1/2): 185-191. https://doi.org/10.1127/arch.moll/133/2004/185" title="Zur Verbreitung der endemischen nordwestsizilianischen Clausiliidae der Untergattung Charpentieria (Siciliaria) mit Beschreibung von zwei neuen Unterarten (Gastropoda: Stylommatophora: Clausiliidae). Archiv fuer Molluskenkunde 133 (1 / 2): 185 - 191." url="https://doi.org/10.1127/arch.moll/133/2004/185" year="2004">Beckmann (2004</bibRefCitation>
|
||
: 190) described
|
||
<taxonomicName authorityName="De Mattia & Reier & Haring" authorityYear="2021" baseAuthorityName="De Mattia & Reier & Haring" baseAuthorityYear="2021" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria septemplicata subsp. hemmeni" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="septemplicata" subSpecies="hemmeni">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria septemplicata hemmeni</emphasis>
|
||
</taxonomicName>
|
||
from Piano Margi (Cinisi, Palermo). Its description and the pictures (
|
||
<bibRefCitation DOI="https://doi.org/10.1127/arch.moll/133/2004/185" author="Beckmann, KH" journalOrPublisher="Basteria" pageId="0" pageNumber="1" refId="B4" refString="Beckmann, KH, 2004. Zur Verbreitung der endemischen nordwestsizilianischen Clausiliidae der Untergattung Charpentieria (Siciliaria) mit Beschreibung von zwei neuen Unterarten (Gastropoda: Stylommatophora: Clausiliidae). Archiv fuer Molluskenkunde 133(1/2): 185-191. https://doi.org/10.1127/arch.moll/133/2004/185" title="Zur Verbreitung der endemischen nordwestsizilianischen Clausiliidae der Untergattung Charpentieria (Siciliaria) mit Beschreibung von zwei neuen Unterarten (Gastropoda: Stylommatophora: Clausiliidae). Archiv fuer Molluskenkunde 133 (1 / 2): 185 - 191." url="https://doi.org/10.1127/arch.moll/133/2004/185" year="2004">Beckmann 2004</bibRefCitation>
|
||
: 189 and 190) match with
|
||
<normalizedToken originalValue="Brandt’s">Brandt's</normalizedToken>
|
||
<taxonomicName authorityName="De Mattia & Reier & Haring" authorityYear="2021" baseAuthorityName="De Mattia & Reier & Haring" baseAuthorityYear="2021" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria (Siciliaria) alcamoensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="alcamoensis" subGenus="Siciliaria">Siciliaria (Siciliaria) alcamoensis</taxonomicName>
|
||
, as stated by
|
||
<bibRefCitation author="Nordsieck, H" journalOrPublisher="Mitteilungen der deutschen malakozoologischen Gesellschaft" pageId="0" pageNumber="1" publicationUrl="http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" refId="B69" refString="Nordsieck, H, 2013b. Revisory remarks on the species of Siciliaria Vest from N. W. Sicily. http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" title="Revisory remarks on the species of Siciliaria Vest from N. W. Sicily." url="http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" year="2013 b">Nordsieck (2013b</bibRefCitation>
|
||
: 7) and therefore, it is reasonable to assume that the Monterosato material (SMF163949) was collected from the same area of
|
||
<normalizedToken originalValue="Beckmann’s">Beckmann's</normalizedToken>
|
||
species.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<taxonomicName authorityName="Brandt" authorityYear="1961" baseAuthorityName="De Mattia & Reier & Haring" baseAuthorityYear="1961" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria tiberii subsp. alcamoensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="tiberii" subSpecies="alcamoensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria tiberii alcamoensis</emphasis>
|
||
</taxonomicName>
|
||
comb. nov. (Figs
|
||
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Siciliaria tiberii tiberii (A. Schmidt, 1868), Capo Rama, Terrasini 26.1 shell 26.2 clausiliar plate double side 26.3 shell 26.4 detail of the aperture 26.5 clausiliar plate double side. Siciliaria tiberii alcamoensis Brandt, 1961 comb. nov., Piano Margi, Carini 26.6 shell 26.7 clausiliar plate double side 26.8 detail of the aperture 26.9 shell." figureDoi="10.3897/zookeys.1077.67081.figure26" httpUri="https://binary.pensoft.net/fig/622818" pageId="0" pageNumber="1">26</figureCitation>
|
||
.6-26.9) was treated as a subspecies of
|
||
<taxonomicName lsidName="S. septemplicata" pageId="0" pageNumber="1" rank="species" species="septemplicata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S. septemplicata</emphasis>
|
||
</taxonomicName>
|
||
(MolluscaBase 2021). This is due to the fact that the typical form of
|
||
<taxonomicName lsidName="S. septemplicata" pageId="0" pageNumber="1" rank="species" species="septemplicata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S. septemplicata</emphasis>
|
||
</taxonomicName>
|
||
(e.g., Monte Gallo populations, Figs
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Siciliaria septemplicata (Philippi, 1836), Monte Gallo, Sferracavallo 10.1 shell 10.2 detail of the aperture 10.3 clausiliar plate 10.4 shell 10.5 detail of the aperture 10.6 detail of the columellar side of last whorl 10.7 clausiliar plate double side." figureDoi="10.3897/zookeys.1077.67081.figure10" httpUri="https://binary.pensoft.net/fig/622802" pageId="0" pageNumber="1">10</figureCitation>
|
||
.1-10.7), is quite similar regarding the morphology of the clausiliar apparatus, (
|
||
<bibRefCitation author="Nordsieck, H" journalOrPublisher="Mitteilungen der deutschen malakozoologischen Gesellschaft" pageId="0" pageNumber="1" publicationUrl="http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" refId="B69" refString="Nordsieck, H, 2013b. Revisory remarks on the species of Siciliaria Vest from N. W. Sicily. http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" title="Revisory remarks on the species of Siciliaria Vest from N. W. Sicily." url="http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" year="2013 b">Nordsieck 2013b</bibRefCitation>
|
||
), despite they greatly differ in the shell surface, sutures ornamentation, colouration, thickness and dimensions. The genital morphology (Figs
|
||
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Siciliaria tiberii tiberii (A. Schmidt, 1868), Capo Rama, Terrasini 25.1 whole distal genital organs 25.2 internal distal part of genital organs. Siciliaria tiberii alcamoensis Brandt, 1961 comb. nov., Piano Margi, Carini 25.3 whole distal genital organs 25.4 internal distal part of genital organs 25.5 detail of the pseudopapilla." figureDoi="10.3897/zookeys.1077.67081.figure25" httpUri="https://binary.pensoft.net/fig/622817" pageId="0" pageNumber="1">25</figureCitation>
|
||
.3-25.4) of
|
||
<taxonomicName lsidName="S. tiberii subsp. alcamoensis" pageId="0" pageNumber="1" rank="subSpecies" species="tiberii" subSpecies="alcamoensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S. tiberii alcamoensis</emphasis>
|
||
</taxonomicName>
|
||
gretaly differs from that of
|
||
<taxonomicName lsidName="S. s. subsp. septemplicata" pageId="0" pageNumber="1" rank="subSpecies" species="s." subSpecies="septemplicata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S. s. septemplicata</emphasis>
|
||
</taxonomicName>
|
||
(Figs
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Siciliaria septemplicata (Philippi, 1836), Monte Gallo, Sferracavallo 9.1 whole distal genital organs 9.2 internal distal part of genital organs 9.3 whole distal genital organs 9.4 internal distal part of genital organs." figureDoi="10.3897/zookeys.1077.67081.figure9" httpUri="https://binary.pensoft.net/fig/622801" pageId="0" pageNumber="1">9</figureCitation>
|
||
.1-9.2). In the COI and concatenated trees,
|
||
<taxonomicName lsidName="S. t. subsp. alcamoensis" pageId="0" pageNumber="1" rank="subSpecies" species="t." subSpecies="alcamoensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S. t. alcamoensis</emphasis>
|
||
</taxonomicName>
|
||
is is well embedded within the
|
||
<taxonomicName authorityName=", s. l." baseAuthorityName="A. Schmidt" baseAuthorityYear="1868" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria tiberii" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tiberii">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria tiberii</emphasis>
|
||
</taxonomicName>
|
||
clade. Since it is found well within the distribution range of
|
||
<taxonomicName lsidName="S. tiberii" pageId="0" pageNumber="1" rank="species" species="tiberii">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S. tiberii</emphasis>
|
||
</taxonomicName>
|
||
s. l., introgression due to hybridisation appears as an unlikely explanation. It might be better explained by incomplete lineage sorting. Possible interspecies hybridisation should be tested on a larger sample size with additional nuclear markers. There is no argument for considering
|
||
<taxonomicName lsidName="S. s. subsp. alcamoensis" pageId="0" pageNumber="1" rank="subSpecies" species="s." subSpecies="alcamoensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S. s. alcamoensis</emphasis>
|
||
</taxonomicName>
|
||
as a subspecies of
|
||
<taxonomicName lsidName="S. septemplicata" pageId="0" pageNumber="1" rank="species" species="septemplicata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S. septemplicata</emphasis>
|
||
</taxonomicName>
|
||
, we here consider it a subspecies
|
||
<taxonomicName lsidName="S. tiberii" pageId="0" pageNumber="1" rank="species" species="tiberii">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S. tiberii</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1077.67081.figure25" httpUri="https://binary.pensoft.net/fig/622817" pageId="0" pageNumber="1" start="Figure 25" startId="F25">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">Figure 25.</emphasis>
|
||
<taxonomicName authority="tiberii" authorityName="tiberii (A. Schmidt" authorityYear="1868" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria tiberii subsp. tiberii" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="tiberii" subSpecies="tiberii">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria tiberii tiberii</emphasis>
|
||
</taxonomicName>
|
||
(A. Schmidt, 1868), Capo Rama, Terrasini
|
||
<emphasis bold="true" pageId="0" pageNumber="1">25.1</emphasis>
|
||
whole distal genital organs
|
||
<emphasis bold="true" pageId="0" pageNumber="1">25.2</emphasis>
|
||
internal distal part of genital organs.
|
||
<taxonomicName authorityName="Brandt" authorityYear="1961" baseAuthorityName="De Mattia & Reier & Haring" baseAuthorityYear="1961" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria tiberii subsp. alcamoensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="tiberii" subSpecies="alcamoensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria tiberii alcamoensis</emphasis>
|
||
</taxonomicName>
|
||
Brandt, 1961 comb. nov., Piano Margi, Carini
|
||
<emphasis bold="true" pageId="0" pageNumber="1">25.3</emphasis>
|
||
whole distal genital organs
|
||
<emphasis bold="true" pageId="0" pageNumber="1">25.4</emphasis>
|
||
internal distal part of genital organs
|
||
<emphasis bold="true" pageId="0" pageNumber="1">25.5</emphasis>
|
||
detail of the pseudopapilla.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.1077.67081.figure26" httpUri="https://binary.pensoft.net/fig/622818" pageId="0" pageNumber="1" start="Figure 26" startId="F26">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">Figure 26.</emphasis>
|
||
<taxonomicName authority="tiberii" authorityName="tiberii (A. Schmidt" authorityYear="1868" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria tiberii subsp. tiberii" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="tiberii" subSpecies="tiberii">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria tiberii tiberii</emphasis>
|
||
</taxonomicName>
|
||
(A. Schmidt, 1868), Capo Rama, Terrasini
|
||
<emphasis bold="true" pageId="0" pageNumber="1">26.1</emphasis>
|
||
shell
|
||
<emphasis bold="true" pageId="0" pageNumber="1">26.2</emphasis>
|
||
clausiliar plate double side
|
||
<emphasis bold="true" pageId="0" pageNumber="1">26.3</emphasis>
|
||
shell
|
||
<emphasis bold="true" pageId="0" pageNumber="1">26.4</emphasis>
|
||
detail of the aperture
|
||
<emphasis bold="true" pageId="0" pageNumber="1">26.5</emphasis>
|
||
clausiliar plate double side.
|
||
<taxonomicName authorityName="Brandt" authorityYear="1961" baseAuthorityName="De Mattia & Reier & Haring" baseAuthorityYear="1961" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria tiberii subsp. alcamoensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="tiberii" subSpecies="alcamoensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria tiberii alcamoensis</emphasis>
|
||
</taxonomicName>
|
||
Brandt, 1961 comb. nov., Piano Margi, Carini
|
||
<emphasis bold="true" pageId="0" pageNumber="1">26.6</emphasis>
|
||
shell
|
||
<emphasis bold="true" pageId="0" pageNumber="1">26.7</emphasis>
|
||
clausiliar plate double side
|
||
<emphasis bold="true" pageId="0" pageNumber="1">26.8</emphasis>
|
||
detail of the aperture
|
||
<emphasis bold="true" pageId="0" pageNumber="1">26.9</emphasis>
|
||
shell.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |