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<document id="9D7BADBEF259E68BF4BDCD584FBEE506" ID-CLB-Dataset="3366" ID-DOI="10.5852/ejt.2021.778.1575" ID-GBIF-Dataset="e9b6381f-da1b-47c3-b232-596c9c9af970" ID-ISSN="2118-9773" ID-Zenodo-Dep="5718409" ID-ZooBank="33DA5677-6AC0-4A27-A9B8-763FCCE4C416" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1637588133144" checkinUser="felipe" docAuthor="Żyła, Dagmara &amp; Koszela, Katarzyna" docDate="2021" docId="11718260FF96060EFDA7FEEBFC31C43C" docLanguage="en" docName="ejt-778_138-147.pdf" docOrigin="European Journal of Taxonomy 778" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Megastilicus iowaensis Żyła &amp; Koszela 2021, sp. nov." docType="treatment" docUuid="9603D57D-FAA2-4DAD-B68F-C91647D98CD0" docUuidSource="ZooBank" docVersion="7" lastPageNumber="145" masterDocId="ED48FA18FF930609FFF9FFE3FF9AC56C" masterDocTitle="Redescription of the genus Megastilicus Casey (Staphylinidae, Paederinae) with the description of a new species" masterLastPageNumber="147" masterPageNumber="138" pageNumber="143" updateTime="1699469157285" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="3E4FA52AFD4BCAB692C9E98B3D82B61C">Redescription of the genus Megastilicus Casey (Staphylinidae, Paederinae) with the description of a new species</mods:title>
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<mods:name id="5C77A8D1326ED6B4F2E87AD3FEEE5073" type="personal">
<mods:role id="ED90AA000DFBE6350534935041B65B75">
<mods:roleTerm id="4E21BDEEF3E1B18323350108647404D4">Author</mods:roleTerm>
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<mods:namePart id="39047AB0849FBC46C04B6B300DB8979C">Żyła, Dagmara</mods:namePart>
<mods:nameIdentifier id="AE73AEB63CEFE7C74408D0165A20C1D6" type="LSID">0A7FEB0F-308C-4FDF-AD0D-8074BC69F0B3</mods:nameIdentifier>
<mods:affiliation id="B211E4A7E23167C16A5337CF083D001F">Department of Invertebrate Zoology and Parasitology, University of Gdańsk, 80 - 308, Gdańsk, Poland. &amp; Department of Ecology, Evolution and Organismal Biology, Iowa State University, Ames, IA 50011, USA. &amp; Museum and Institute of Zoology, Polish Academy of Sciences, Wilcza 64, 00 - 679, Warszawa, Poland.</mods:affiliation>
<mods:nameIdentifier id="B22B85A98CAC4A7C9DC6355E3263808A" type="email">zyladagmara@gmail.com</mods:nameIdentifier>
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<mods:namePart id="ED5B84FC5E868B76C233E21DA4A912C7">Koszela, Katarzyna</mods:namePart>
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<mods:affiliation id="90436D85D8534450375837BD63B3D1DC">Museum and Institute of Zoology, Polish Academy of Sciences, Wilcza 64, 00 - 679, Warszawa, Poland.</mods:affiliation>
<mods:nameIdentifier id="21A0C8D8BF2E1EB5E2E9CDC8B6D5496B" type="email">kat.koszela@gmail.com</mods:nameIdentifier>
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<mods:title id="E93415FBEC179822AE8FBFE58D959BCF">European Journal of Taxonomy</mods:title>
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<mods:part id="EBF56B57DF8C5B5E0E7346E82595A02A">
<mods:date id="0CB46DEB4A49A96FC4BAA57611D93879">2021</mods:date>
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<mods:number id="B71FA22BD0F8A8A96035528BB052E558">2021-11-22</mods:number>
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<treatment id="11718260FF96060EFDA7FEEBFC31C43C" ID-DOI="http://doi.org/10.5281/zenodo.5718527" ID-GBIF-Taxon="190503018" ID-Zenodo-Dep="5718527" LSID="urn:lsid:zoobank.org:act:9603D57D-FAA2-4DAD-B68F-C91647D98CD0" httpUri="http://treatment.plazi.org/id/11718260FF96060EFDA7FEEBFC31C43C" lastPageId="7" lastPageNumber="145" pageId="5" pageNumber="143">
<subSubSection id="D1C260FDFF96060CFDA7FEEBFC4CC448" box="[606,982,264,292]" pageId="5" pageNumber="143" type="nomenclature">
<paragraph id="99673376FF96060CFDA7FEEBFC4CC448" blockId="5.[355,1232,264,366]" box="[606,982,264,292]" pageId="5" pageNumber="143">
<heading id="C22F841AFF96060CFDA7FEEBFC4CC448" box="[606,982,264,292]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="143" reason="2">
<taxonomicName id="5ED848F5FF96060CFDA7FEEBFCEEC44F" ID-CoL="9GCP7" authority="Żyła &amp; Koszela, 2021" authorityName="Żyła &amp; Koszela" authorityYear="2021" box="[606,884,264,291]" class="Insecta" family="Staphylinidae" genus="Megastilicus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="143" phylum="Arthropoda" rank="species" species="iowaensis" status="sp. nov.">
<emphasis id="ABACEF64FF96060CFDA7FEEBFCEEC44F" bold="true" box="[606,884,264,291]" italics="true" pageId="5" pageNumber="143">Megastilicus iowaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B09F521FFF96060CFC82FEE9FC4CC448" box="[891,982,266,292]" pageId="5" pageNumber="143" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<uri id="ED493F74FF96060CFE9AFECDFB4AC425" box="[355,1232,302,329]" pageId="5" pageNumber="143">
urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
<paragraph id="99673376FF96060CFD0EFEB7FCA6C402" blockId="5.[355,1232,264,366]" box="[759,828,340,366]" pageId="5" pageNumber="143">
<figureCitation id="01E32FF3FF96060CFD0EFEB7FCA6C402" box="[759,828,340,366]" captionStart="Fig" captionStartId="6.[189,232,1928,1954]" captionTargetBox="[280,1309,265,1861]" captionTargetId="figure-11@6.[279,1309,265,1890]" captionTargetPageId="6" captionText="Fig. 2. Morphology of Megastilicus iowaensis sp. nov. A. Dorsal habitus of holotype (NMNH). B. Dorsal habitus of paratype (NMNH). C. Sternum VIII. D. Aedeagus in parameral view. E. Aedeagus in lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5718415" httpUri="https://zenodo.org/record/5718415/files/figure.png" pageId="5" pageNumber="143">Fig. 2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="D1C260FDFF96060CFF44FE74FB0AC742" pageId="5" pageNumber="143" type="diagnosis">
<paragraph id="99673376FF96060CFF44FE74FEACC4DD" blockId="5.[189,310,407,433]" box="[189,310,407,433]" pageId="5" pageNumber="143">
<heading id="C22F841AFF96060CFF44FE74FEACC4DD" bold="true" box="[189,310,407,433]" fontSize="11" level="3" pageId="5" pageNumber="143" reason="3">
<emphasis id="ABACEF64FF96060CFF44FE74FEACC4DD" bold="true" box="[189,310,407,433]" pageId="5" pageNumber="143">Diagnosis</emphasis>
</heading>
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Head as long as wide.Antennomere 4 more transverse compared to elongated in
<taxonomicName id="5ED848F5FF96060CFBB7FE2BFAF4C48E" authorityName="Casey" authorityYear="1889" box="[1102,1390,456,482]" class="Insecta" family="Staphylinidae" genus="Megastilicus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="143" phylum="Arthropoda" rank="species" species="formicarius">
<emphasis id="ABACEF64FF96060CFBB7FE2BFAF4C48E" box="[1102,1390,456,482]" italics="true" pageId="5" pageNumber="143">Megastilicus formicarius</emphasis>
</taxonomicName>
. Teeth on labrum pointed. Eyes larger than in
<taxonomicName id="5ED848F5FF96060CFD33FE0CFCE0C764" authorityName="Casey" authorityYear="1889" box="[714,890,494,520]" class="Insecta" family="Staphylinidae" genus="Megastilicus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="143" phylum="Arthropoda" rank="species" species="formicarius">
<emphasis id="ABACEF64FF96060CFD33FE0CFCE0C764" box="[714,890,494,520]" italics="true" pageId="5" pageNumber="143">M. formicarius</emphasis>
</taxonomicName>
, ca ⅕ of head length. Median emargination of posterior margin of sternum VIII shallower and narrower than in
<taxonomicName id="5ED848F5FF96060CFC20FDF7FB13C741" authorityName="Casey" authorityYear="1889" box="[985,1161,531,557]" class="Insecta" family="Staphylinidae" genus="Megastilicus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="143" phylum="Arthropoda" rank="species" species="formicarius">
<emphasis id="ABACEF64FF96060CFC20FDF7FB13C741" box="[985,1161,531,557]" italics="true" pageId="5" pageNumber="143">M. formicarius</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="D1C260FDFF96060CFF44FDB4FEB5C7AB" pageId="5" pageNumber="143" type="etymology">
<paragraph id="99673376FF96060CFF44FDB4FEDEC71D" blockId="5.[189,324,599,625]" box="[189,324,599,625]" pageId="5" pageNumber="143">
<heading id="C22F841AFF96060CFF44FDB4FEDEC71D" bold="true" box="[189,324,599,625]" fontSize="11" level="3" pageId="5" pageNumber="143" reason="3">
<emphasis id="ABACEF64FF96060CFF44FDB4FEDEC71D" bold="true" box="[189,324,599,625]" pageId="5" pageNumber="143">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="99673376FF96060CFF44FD6BFEB5C7AB" blockId="5.[189,1398,648,711]" pageId="5" pageNumber="143">
The species name ʻ
<taxonomicName id="5ED848F5FF96060CFE62FD6AFD95C7CE" authorityName="Żyła &amp; Koszela" authorityYear="2021" box="[411,527,649,674]" class="Insecta" family="Staphylinidae" genus="Megastilicus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="143" phylum="Arthropoda" rank="species" species="iowaensis">
<emphasis id="ABACEF64FF96060CFE62FD6AFD95C7CE" box="[411,527,649,674]" italics="true" pageId="5" pageNumber="143">iowaensis</emphasis>
</taxonomicName>
ʼ is derived from the name of the
<collectingCountry id="E1CF73E6FF96060CFC76FD6BFC2EC7CE" box="[911,948,648,674]" name="United States of America" pageId="5" pageNumber="143">US</collectingCountry>
state
<collectingRegion id="5B1CFD94FF96060CFC0EFD6BFBA8C7CE" box="[1015,1074,648,674]" country="United States of America" name="Iowa" pageId="5" pageNumber="143">Iowa</collectingRegion>
where this species has been collected.
</paragraph>
</subSubSection>
<subSubSection id="D1C260FDFF96060CFF44FD12FBA8C10C" pageId="5" pageNumber="143" type="materials_examined">
<paragraph id="99673376FF96060CFF44FD12FEF6C667" blockId="5.[189,364,753,779]" box="[189,364,753,779]" pageId="5" pageNumber="143">
<heading id="C22F841AFF96060CFF44FD12FEF6C667" bold="true" box="[189,364,753,779]" fontSize="11" level="3" pageId="5" pageNumber="143" reason="3">
<emphasis id="ABACEF64FF96060CFF44FD12FEF6C667" bold="true" box="[189,364,753,779]" pageId="5" pageNumber="143">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="99673376FF96060CFF13FCC1FE6DC6EB" blockId="5.[189,1398,802,903]" pageId="5" pageNumber="143">
<materialsCitation id="29B0392BFF96060CFF13FCC1FE6DC6EB" ID-GBIF-Occurrence="3410541317" collectingDate="2010-04" collectionCode="NMNH" country="United States of America" location="Iowa" pageId="5" pageNumber="143" specimenCount="1" specimenCount-male="1" stateProvince="Iowa" typeStatus="holotype">
<emphasis id="ABACEF64FF96060CFF13FCC1FEC6C650" bold="true" box="[234,348,802,828]" pageId="5" pageNumber="143">
<typeStatus id="46638DD4FF96060CFF13FCC1FEC6C650" box="[234,348,802,828]" pageId="5" pageNumber="143" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="E1CF73E6FF96060CFF44FCA4FF6DC60D" box="[189,247,839,865]" name="United States of America" pageId="5" pageNumber="143">USA</collectingCountry>
<specimenCount id="8FDEF8FFFF96060CFEEFFCABFEB6C60E" box="[278,300,840,866]" pageId="5" pageNumber="143" type="male"></specimenCount>
with ant [identified as
<taxonomicName id="5ED848F5FF96060CFDA9FCABFCD9C60D" authorityName="Say" authorityYear="1836" box="[592,835,839,865]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="143" phylum="Arthropoda" rank="species" species="subsericea">
<emphasis id="ABACEF64FF96060CFDA9FCABFCD9C60D" box="[592,835,839,865]" italics="true" pageId="5" pageNumber="143">Formica subsericea</emphasis>
</taxonomicName>
]; “
<collectingRegion id="5B1CFD94FF96060CFC92FCA4FC30C60D" box="[875,938,839,865]" country="United States of America" name="Iowa" pageId="5" pageNumber="143">Iowa</collectingRegion>
City Wickham
<date id="ED6615B6FF96060CFB89FCABFB22C60D" box="[1136,1208,839,866]" pageId="5" pageNumber="143" value="2010-04">
<collectingDate id="FD22EC5EFF96060CFB89FCABFB22C60D" box="[1136,1208,839,866]" pageId="5" pageNumber="143" value="2010-04">IV 10</collectingDate>
</date>
? / WICKHAM Collection 1933”;
<collectionCode id="FFC9ABB3FF96060CFE6FFC8EFE6EC6EB" box="[406,500,877,903]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="143" type="Museum">NMNH</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="99673376FF96060CFF13FC53FBA8C10C" blockId="5.[189,1399,944,1121]" pageId="5" pageNumber="143">
<materialsCitation id="29B0392BFF96060CFF13FC53FB25C69C" ID-GBIF-Occurrence="3410541311" collectingDate="2010-04" collectionCode="NMNH" country="United States of America" county="City" location="Iowa" municipality="Iowa" pageId="5" pageNumber="143" specimenCount="1" specimenCount-female="1" stateProvince="Iowa" typeStatus="paratype">
<emphasis id="ABACEF64FF96060CFF13FC53FEFDC6A6" bold="true" box="[234,359,944,970]" pageId="5" pageNumber="143">
<typeStatus id="46638DD4FF96060CFF13FC53FEFDC6A6" box="[234,359,944,970]" pageId="5" pageNumber="143" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="E1CF73E6FF96060CFF44FC35FF6CC69C" box="[189,246,982,1008]" name="United States of America" pageId="5" pageNumber="143">USA</collectingCountry>
<specimenCount id="8FDEF8FFFF96060CFEE8FC35FEA5C69C" box="[273,319,982,1008]" pageId="5" pageNumber="143" type="female">1 ♀</specimenCount>
; “
<collectingMunicipality id="7903A90CFF96060CFEA1FC35FE0FC69C" box="[344,405,982,1008]" pageId="5" pageNumber="143">Iowa</collectingMunicipality>
City
<collectingRegion id="5B1CFD94FF96060CFE21FC35FD89C69C" box="[472,531,982,1008]" country="United States of America" name="Iowa" pageId="5" pageNumber="143">Iowa</collectingRegion>
<date id="ED6615B6FF96060CFDE2FC35FDC5C69C" box="[539,607,982,1008]" pageId="5" pageNumber="143" value="2010-04">
<collectingDate id="FD22EC5EFF96060CFDE2FC35FDC5C69C" box="[539,607,982,1008]" pageId="5" pageNumber="143" value="2010-04">IV 10</collectingDate>
</date>
Wickham / WICKHAM Collection 1933”;
<collectionCode id="FFC9ABB3FF96060CFB9CFC35FB25C69C" box="[1125,1215,982,1008]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="143" type="Museum">NMNH</collectionCode>
</materialsCitation>
<materialsCitation id="29B0392BFF96060CFB20FC35FBBDC17A" ID-GBIF-Occurrence="3410541318" collectingDate="2013-04" collectionCode="NMNH" country="United States of America" location="Iowa" municipality="Iowa" pageId="5" pageNumber="143" specimenCount="1" specimenCount-female="1" stateProvince="Iowa" typeStatus="paratype">
<specimenCount id="8FDEF8FFFF96060CFB20FC35FA9CC69C" box="[1241,1286,982,1008]" pageId="5" pageNumber="143" type="female">1 ♀</specimenCount>
with ant; “
<collectingMunicipality id="7903A90CFF96060CFF3EFC18FE9FC179" box="[199,261,1019,1045]" pageId="5" pageNumber="143">Iowa</collectingMunicipality>
City Wickham
<date id="ED6615B6FF96060CFE31FC18FD8AC179" box="[456,528,1019,1045]" pageId="5" pageNumber="143" value="2013-04">
<collectingDate id="FD22EC5EFF96060CFE31FC18FD8AC179" box="[456,528,1019,1045]" pageId="5" pageNumber="143" value="2013-04">IV-13</collectingDate>
</date>
-?8 / WICKHAM Collection 1933”;
<collectionCode id="FFC9ABB3FF96060CFC35FC1FFBBDC17A" box="[972,1063,1020,1046]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="143" type="Museum">NMNH</collectionCode>
</materialsCitation>
<materialsCitation id="29B0392BFF96060CFBBDFC18FBB4C10C" ID-GBIF-Occurrence="3410541304" collectingDate="1968-04-29" collectionCode="MCZ" country="United States of America" county="City Wickham" location="Wickham" municipality="Iowa City" pageId="5" pageNumber="143" specimenCode="MCZ ENT 00744565" specimenCount="1" specimenCount-female="1" stateProvince="Iowa" typeStatus="paratype">
<specimenCount id="8FDEF8FFFF96060CFBBDFC18FBE8C17A" box="[1092,1138,1019,1046]" pageId="5" pageNumber="143" type="female">1 ♀</specimenCount>
with ant; “
<collectingMunicipality id="7903A90CFF96060CFB02FC18FAECC179" box="[1275,1398,1019,1045]" pageId="5" pageNumber="143">Iowa City</collectingMunicipality>
<location id="9C0765ADFF96060CFF44FBC3FEB6C156" LSID="urn:lsid:plazi:treatment:11718260FF96060EFDA7FEEBFC31C43C:9C0765ADFF96060CFF44FBC3FEB6C156" box="[189,300,1056,1082]" country="United States of America" county="City Wickham" municipality="Iowa City" name="Wickham" pageId="5" pageNumber="143" stateProvince="Iowa">Wickham</location>
<date id="ED6615B6FF96060CFECBFBC3FE07C156" box="[306,413,1056,1082]" pageId="5" pageNumber="143" value="1968-04-29">
<collectingDate id="FD22EC5EFF96060CFECBFBC3FE07C156" box="[306,413,1056,1082]" pageId="5" pageNumber="143" value="1968-04-29">IV-29-68</collectingDate>
</date>
? /
<taxonomicName id="5ED848F5FF96060CFE42FBC3FC88C157" authority="Csy." authorityName="Casey" authorityYear="1889" box="[443,786,1056,1083]" class="Insecta" family="Staphylinidae" genus="Megastilicus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="143" phylum="Arthropoda" rank="species" species="formicarius">Megastilicus formicarius Csy.</taxonomicName>
/ Host ant:
<taxonomicName id="5ED848F5FF96060CFC68FBC3FBF5C156" authorityName="Say" authorityYear="1836" box="[913,1135,1056,1082]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="143" phylum="Arthropoda" rank="species" species="subsericea">Formica subsericea</taxonomicName>
/ 9650 / Percy Gardner Bolster Colln /
<taxonomicName id="5ED848F5FF96060CFE82FBA4FDE0C10C" authority="Casey" authorityName="Casey" authorityYear="1889" box="[379,634,1094,1121]" class="Insecta" family="Staphylinidae" genus="Megastilicus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="143" phylum="Arthropoda" rank="species" species="formicarius">M. formicarius Casey</taxonomicName>
9650 /
<specimenCode id="C97E9B0DFF96060CFD2AFBA5FC4FC10D" box="[723,981,1094,1121]" pageId="5" pageNumber="143">MCZ ENT 00744565</specimenCode>
”;
<collectionCode id="FFC9ABB3FF96060CFC12FBA5FBB4C10C" box="[1003,1070,1094,1120]" collectionName="USA, Massachusetts, Cambridge, Harvard University, Museum of Comparative Zoology" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="5" pageNumber="143" type="Museum">MCZ</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D1C260FDFF96060EFF44FB6AFC31C43C" lastPageId="7" lastPageNumber="145" pageId="5" pageNumber="143" type="description">
<paragraph id="99673376FF96060CFF44FB6AFED4C1CF" blockId="5.[189,334,1161,1187]" box="[189,334,1161,1187]" pageId="5" pageNumber="143">
<heading id="C22F841AFF96060CFF44FB6AFED4C1CF" bold="true" box="[189,334,1161,1187]" fontSize="11" level="3" pageId="5" pageNumber="143" reason="3">
<emphasis id="ABACEF64FF96060CFF44FB6AFED4C1CF" bold="true" box="[189,334,1161,1187]" pageId="5" pageNumber="143">Description</emphasis>
</heading>
</paragraph>
<paragraph id="99673376FF96060CFF44FB58FE5AC196" blockId="5.[189,1398,1210,1274]" pageId="5" pageNumber="143">
BODY. Colour of specimens pale brown/sandy (
<figureCitation id="01E32FF3FF96060CFD15FB59FCF4C1B9" box="[748,878,1210,1237]" captionStart="Fig" captionStartId="6.[189,232,1928,1954]" captionTargetBox="[280,1309,265,1861]" captionTargetId="figure-11@6.[279,1309,265,1890]" captionTargetPageId="6" captionText="Fig. 2. Morphology of Megastilicus iowaensis sp. nov. A. Dorsal habitus of holotype (NMNH). B. Dorsal habitus of paratype (NMNH). C. Sternum VIII. D. Aedeagus in parameral view. E. Aedeagus in lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5718415" httpUri="https://zenodo.org/record/5718415/files/figure.png" pageId="5" pageNumber="143">Fig. 2AB</figureCitation>
). Head darker. Body length from tip of head to tip of abdomen 6.0.
</paragraph>
<paragraph id="99673376FF96060CFF44FAC7FC0CC372" blockId="5.[188,1400,1316,1567]" pageId="5" pageNumber="143">HEAD. Width 1.0, length 1.11.2. Antennomere 1 elongate, widest at tip; antennomeres 2 and 3 elongate; antennomeres 4 to 10 gradually becoming strongly transverse; antennomere 11 elongate, 1.5 × as long as antennomere 10. Antennomeres 1 and 2 sparsely pubescent, antennomeres 3 and 4 more densely pubescent, antennomeres 5 to 11 densely covered with tomentose pubescence. Emargination on anterior margin of labrum shallow. Maxillary palpomere 2 slender and short, with only few setae, maxillary palpomere 3 slightly expanded towards apex with denser setation, maxillary palpomere 4 glossy. Eyes about ⅕ of head length, shifted anteriorly, ⅓ of temple length.</paragraph>
<paragraph id="99673376FF96060CFF44F9ABFB17C30E" blockId="5.[189,1165,1608,1635]" box="[189,1165,1608,1635]" pageId="5" pageNumber="143">THORAX. Pronotum width 0.9, length 1.01.1. Elytra width 1.41.5, length 1.31.4.</paragraph>
<paragraph id="99673376FF96060CFF44F96FFD93C3A7" blockId="5.[188,1399,1676,1740]" pageId="5" pageNumber="143">LEGS. Two long setae on outer margin of metacoxa. Seta closer to abdomen shorter and thinner than seta more distant from abdomen.</paragraph>
<paragraph id="99673376FF96060CFF44F916FE3CC213" blockId="5.[189,1400,1781,1919]" pageId="5" pageNumber="143">
ABDOMEN. Tergum VIII with slightly pointed posterior margin. Male with sternum VIII with shallow and broad median emargination of posterior margin; depth of emargination of ca ⅐ of sternum length; sides of emargination rounded (
<figureCitation id="01E32FF3FF96060CFDCBF8A3FD15C236" box="[562,655,1856,1882]" captionStart="Fig" captionStartId="6.[189,232,1928,1954]" captionTargetBox="[280,1309,265,1861]" captionTargetId="figure-11@6.[279,1309,265,1890]" captionTargetPageId="6" captionText="Fig. 2. Morphology of Megastilicus iowaensis sp. nov. A. Dorsal habitus of holotype (NMNH). B. Dorsal habitus of paratype (NMNH). C. Sternum VIII. D. Aedeagus in parameral view. E. Aedeagus in lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5718415" httpUri="https://zenodo.org/record/5718415/files/figure.png" pageId="5" pageNumber="143">Fig. 2C</figureCitation>
); surface anterior to emargination without setae, while margins with dense setation.
</paragraph>
<paragraph id="99673376FF96060CFF44F84AFC1EC284" blockId="5.[189,1398,1961,2025]" pageId="5" pageNumber="143">
AEDEAGUS. With ventral process in parameral view widened in middle; lateral plates of internal sac welldeveloped and attached to sides of median lobe (
<figureCitation id="01E32FF3FF96060CFD01F82DFCEDC284" box="[760,887,1998,2024]" captionStart="Fig" captionStartId="6.[189,232,1928,1954]" captionTargetBox="[280,1309,265,1861]" captionTargetId="figure-11@6.[279,1309,265,1890]" captionTargetPageId="6" captionText="Fig. 2. Morphology of Megastilicus iowaensis sp. nov. A. Dorsal habitus of holotype (NMNH). B. Dorsal habitus of paratype (NMNH). C. Sternum VIII. D. Aedeagus in parameral view. E. Aedeagus in lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5718415" httpUri="https://zenodo.org/record/5718415/files/figure.png" pageId="5" pageNumber="143">Fig. 2DE</figureCitation>
).
</paragraph>
<caption id="CDA763FEFF95060FFF44F86BFE7CC285" ID-DOI="http://doi.org/10.5281/zenodo.5718415" ID-Zenodo-Dep="5718415" httpUri="https://zenodo.org/record/5718415/files/figure.png" pageId="6" pageNumber="144" startId="6.[189,232,1928,1954]" targetBox="[280,1309,265,1861]" targetPageId="6">
<paragraph id="99673376FF95060FFF44F86BFE7CC285" blockId="6.[189,1399,1928,2025]" pageId="6" pageNumber="144">
<emphasis id="ABACEF64FF95060FFF44F86BFE93C2CE" bold="true" box="[189,265,1928,1954]" pageId="6" pageNumber="144">Fig. 2.</emphasis>
Morphology of
<taxonomicName id="5ED848F5FF95060FFE3BF86BFD4AC2CE" authority="Żyła &amp; Koszela, 2021" authorityName="Żyła &amp; Koszela" authorityYear="2021" box="[450,720,1928,1954]" class="Insecta" family="Staphylinidae" genus="Megastilicus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="144" phylum="Arthropoda" rank="species" species="iowaensis" status="sp. nov.">
<emphasis id="ABACEF64FF95060FFE3BF86BFD4AC2CE" box="[450,720,1928,1954]" italics="true" pageId="6" pageNumber="144">Megastilicus iowaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B09F521FFF95060FFD2EF86AFCAAC2CF" box="[727,816,1929,1955]" pageId="6" pageNumber="144" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="ABACEF64FF95060FFCCDF86BFCD3C2CE" bold="true" box="[820,841,1928,1954]" pageId="6" pageNumber="144">A</emphasis>
. Dorsal habitus of holotype (NMNH).
<emphasis id="ABACEF64FF95060FFAF1F86BFA81C2CE" bold="true" box="[1288,1307,1928,1954]" pageId="6" pageNumber="144">B</emphasis>
. Dorsal habitus of paratype (NMNH).
<emphasis id="ABACEF64FF95060FFDDAF848FDA2C2A9" bold="true" box="[547,568,1963,1989]" pageId="6" pageNumber="144">C</emphasis>
. Sternum VIII.
<emphasis id="ABACEF64FF95060FFD0AF848FC92C2A9" bold="true" box="[755,776,1963,1989]" pageId="6" pageNumber="144">D</emphasis>
. Aedeagus in parameral view.
<emphasis id="ABACEF64FF95060FFB8AF848FB1CC2A9" bold="true" box="[1139,1158,1963,1989]" pageId="6" pageNumber="144">E</emphasis>
. Aedeagus in lateral view. Scale bars = 1 mm.
</paragraph>
</caption>
<paragraph id="99673376FF94060EFF44FEEBFED5C44E" blockId="7.[189,335,264,290]" box="[189,335,264,290]" pageId="7" pageNumber="145">
<heading id="C22F841AFF94060EFF44FEEBFED5C44E" bold="true" box="[189,335,264,290]" fontSize="11" level="3" pageId="7" pageNumber="145" reason="3">
<emphasis id="ABACEF64FF94060EFF44FEEBFED5C44E" bold="true" box="[189,335,264,290]" pageId="7" pageNumber="145">Occurrence</emphasis>
</heading>
</paragraph>
<paragraph id="99673376FF94060EFF44FED5FC31C43C" blockId="7.[189,939,310,336]" box="[189,939,310,336]" pageId="7" pageNumber="145">
The new species has so far only been recorded from
<collectingRegion id="5B1CFD94FF94060EFCDFFED5FC3CC43C" box="[806,934,310,336]" country="United States of America" name="Iowa" pageId="7" pageNumber="145">Iowa State</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>