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<document id="84DE269A138A81FE46525B61AD4490BA" ID-CLB-Dataset="3366" ID-DOI="10.5852/ejt.2021.778.1575" ID-GBIF-Dataset="e9b6381f-da1b-47c3-b232-596c9c9af970" ID-ISSN="2118-9773" ID-Zenodo-Dep="5718409" ID-ZooBank="33DA5677-6AC0-4A27-A9B8-763FCCE4C416" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1637588133144" checkinUser="felipe" docAuthor="Żyła, Dagmara &amp; Koszela, Katarzyna" docDate="2021" docId="11718260FF92060AFDA0F84AFC5AC49F" docLanguage="en" docName="ejt-778_138-147.pdf" docOrigin="European Journal of Taxonomy 778" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Megastilicus Casey 1889" docType="treatment" docVersion="8" lastPageNumber="141" masterDocId="ED48FA18FF930609FFF9FFE3FF9AC56C" masterDocTitle="Redescription of the genus Megastilicus Casey (Staphylinidae, Paederinae) with the description of a new species" masterLastPageNumber="147" masterPageNumber="138" pageNumber="139" updateTime="1699469157285" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="BC4A1966DE12898AFEF6C712EA09C244">Redescription of the genus Megastilicus Casey (Staphylinidae, Paederinae) with the description of a new species</mods:title>
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<mods:name id="2FBC087216D3A34CCA2EEBE6CB1951EC" type="personal">
<mods:role id="7DFD137A8483C7CDBDFC5411FD8C3701">
<mods:roleTerm id="F26BEB3CD7B3A67C1B55A496762CF1CB">Author</mods:roleTerm>
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<mods:namePart id="627B8159865C8D56D7B5EAEFAC25ADA5">Żyła, Dagmara</mods:namePart>
<mods:nameIdentifier id="45CF1D3420B326C84A720B8747BFF0DF" type="LSID">0A7FEB0F-308C-4FDF-AD0D-8074BC69F0B3</mods:nameIdentifier>
<mods:affiliation id="183966895098527920CAC460AAB05109">Department of Invertebrate Zoology and Parasitology, University of Gdańsk, 80 - 308, Gdańsk, Poland. &amp; Department of Ecology, Evolution and Organismal Biology, Iowa State University, Ames, IA 50011, USA. &amp; Museum and Institute of Zoology, Polish Academy of Sciences, Wilcza 64, 00 - 679, Warszawa, Poland.</mods:affiliation>
<mods:nameIdentifier id="53C003E430C4F5BB48F369877E48E8DF" type="email">zyladagmara@gmail.com</mods:nameIdentifier>
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<mods:name id="174C543A78EBCB846EECBE6207DEEC70" type="personal">
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<mods:namePart id="F0D70F47402B9F317AAC0563F6D290A1">Koszela, Katarzyna</mods:namePart>
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<mods:affiliation id="6CE2EA1D4FCC6CB40445F2DD8F147A03">Museum and Institute of Zoology, Polish Academy of Sciences, Wilcza 64, 00 - 679, Warszawa, Poland.</mods:affiliation>
<mods:nameIdentifier id="FB86E7D6C7BA961193E5130FD718C14E" type="email">kat.koszela@gmail.com</mods:nameIdentifier>
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<mods:title id="C093C61E590110DAF155F928E3D18D38">European Journal of Taxonomy</mods:title>
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<mods:date id="B5E24E8C79488DCC1A92972E5500E08E">2021</mods:date>
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<mods:number id="505D9F36D0CBBF5170A12483253552BF">2021-11-22</mods:number>
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<mods:number id="4700C714946E8E2CB92B43F84D96711E">778</mods:number>
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<subSubSection id="D1C260FDFF920608FDA0F84AFC41C2AF" box="[601,987,1960,1987]" pageId="1" pageNumber="139" type="nomenclature">
<paragraph id="99673376FF920608FDA0F84AFC41C2AF" blockId="1.[601,987,1960,2024]" box="[601,987,1960,1987]" pageId="1" pageNumber="139">
<heading id="C22F841AFF920608FDA0F84AFC41C2AF" box="[601,987,1960,1987]" centered="true" fontSize="11" level="2" pageId="1" pageNumber="139" reason="2">
Genus
<taxonomicName id="5ED848F5FF920608FD52F84BFC41C2AF" ID-CoL="8VRTL" authority="Casey, 1889" authorityName="Casey" authorityYear="1889" box="[683,987,1960,1987]" class="Insecta" family="Staphylinidae" genus="Megastilicus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="139" phylum="Arthropoda" rank="genus">
<emphasis id="ABACEF64FF920608FD52F84BFCDEC2AF" bold="true" box="[683,836,1960,1987]" italics="true" pageId="1" pageNumber="139">Megastilicus</emphasis>
<bibRefCitation id="FD494E87FF920608FCB5F84AFC41C2AF" author="Casey T. L." box="[844,987,1961,1987]" pageId="1" pageNumber="139" pagination="182 - 184" refId="ref4337" refString="Casey T. L. 1889. Notes on the Paederini. Entomologica Americana 5: 182 - 184." type="journal article" year="1889">Casey, 1889</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="D1C260FDFF920608FD1EF82DFCD7C284" box="[743,845,1998,2024]" pageId="1" pageNumber="139" type="description">
<paragraph id="99673376FF920608FD1EF82DFCD7C284" blockId="1.[601,987,1960,2024]" box="[743,845,1998,2024]" pageId="1" pageNumber="139">
<figureCitation id="01E32FF3FF920608FD1EF82DFCD7C284" box="[743,845,1998,2024]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="4.[189,231,1928,1954]" captionStartId-1="6.[189,232,1928,1954]" captionTargetBox-0="[189,1398,975,1890]" captionTargetBox-1="[280,1309,265,1861]" captionTargetId-0="figure-168@4.[189,1398,975,1890]" captionTargetId-1="figure-11@6.[279,1309,265,1890]" captionTargetPageId-0="4" captionTargetPageId-1="6" captionText-0="Fig. 1. Morphology of Megastilicus formicarius Casey, 1889. A. Dorsal habitus of paralectotype (NMNH). B. Dorsal habitus of lectotype (NMNH). C. Sternum VIII. D. Aedeagus in parameral view. E. Aedeagus in lateral view. Scale bars = 1 mm." captionText-1="Fig. 2. Morphology of Megastilicus iowaensis sp. nov. A. Dorsal habitus of holotype (NMNH). B. Dorsal habitus of paratype (NMNH). C. Sternum VIII. D. Aedeagus in parameral view. E. Aedeagus in lateral view. Scale bars = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5718413" figureDoi-1="http://doi.org/10.5281/zenodo.5718415" httpUri-0="https://zenodo.org/record/5718413/files/figure.png" httpUri-1="https://zenodo.org/record/5718415/files/figure.png" pageId="1" pageNumber="139">Figs 1–2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="D1C260FDFF91060BFF44FEEAFE3DC77D" pageId="2" pageNumber="140" type="reference_group">
<paragraph id="99673376FF91060BFF44FEEAFCB7C448" blockId="2.[189,813,265,292]" box="[189,813,265,292]" pageId="2" pageNumber="140">
<treatmentCitationGroup id="B9C81458FF91060BFF44FEEAFCB7C448" box="[189,813,265,292]" pageId="2" pageNumber="140">
<treatmentCitation id="18791567FF91060BFF44FEEAFDBFC44F" author="Casey T. L." box="[189,549,265,291]" page="183" pageId="2" pageNumber="140" year="1889">
<taxonomicName id="5ED848F5FF91060BFF44FEEAFDBFC44F" ID-CoL="8VRTL" authority="Casey, 1889: 183" authorityName="Casey" authorityPageNumber="183" authorityYear="1889" box="[189,549,265,291]" class="Insecta" family="Staphylinidae" genus="Megastilicus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="140" phylum="Arthropoda" rank="genus">
<emphasis id="ABACEF64FF91060BFF44FEEAFEC9C44F" box="[189,339,265,291]" italics="true" pageId="2" pageNumber="140">Megastilicus</emphasis>
<bibRefCitation id="FD494E87FF91060BFEA3FEEAFDBFC44F" author="Casey T. L." box="[346,549,265,291]" pageId="2" pageNumber="140" pagination="182 - 184" refId="ref4337" refString="Casey T. L. 1889. Notes on the Paederini. Entomologica Americana 5: 182 - 184." type="journal article" year="1889">Casey, 1889: 183</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description).
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</paragraph>
<paragraph id="99673376FF91060BFF44FEA9FE3DC77D" blockId="2.[189,1399,330,529]" pageId="2" pageNumber="140">
<treatmentCitationGroup id="B9C81458FF91060BFF44FEA9FE3DC77D" pageId="2" pageNumber="140">
<taxonomicName id="5ED848F5FF91060BFF44FEA9FEC9C408" ID-CoL="8VRTL" authorityName="Casey" authorityYear="1889" box="[189,339,330,356]" class="Insecta" family="Staphylinidae" genus="Megastilicus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="140" phylum="Arthropoda" rank="genus">
<emphasis id="ABACEF64FF91060BFF44FEA9FEC9C408" box="[189,339,330,356]" italics="true" pageId="2" pageNumber="140">Megastilicus</emphasis>
</taxonomicName>
–
<treatmentCitation id="18791567FF91060BFE97FEA9FDABC408" author="Casey T. L." box="[366,561,330,357]" page="227" pageId="2" pageNumber="140" year="1905">
<bibRefCitation id="FD494E87FF91060BFE97FEA9FDABC408" author="Casey T. L." box="[366,561,330,357]" pageId="2" pageNumber="140" refId="ref4357" refString="Casey T. L. 1905. A revision of the American Paederini. Vol. 15, No. 2. Academy of Science of St. Louis, Saint Louis. https: // doi. org / 10.5962 / bhl. title. 9845" type="book" year="1905">Casey 1905: 227</bibRefCitation>
</treatmentCitation>
(description and key). —
<treatmentCitation id="18791567FF91060BFCA6FEA9FBB4C408" author="Wickham H. F." box="[863,1070,330,356]" page="3" pageId="2" pageNumber="140" year="1900">
<bibRefCitation id="FD494E87FF91060BFCA6FEA9FBB4C408" author="Wickham H. F." box="[863,1070,330,356]" pageId="2" pageNumber="140" pagination="3 - 5" refId="ref4885" refString="Wickham H. F. 1900. On Coleoptera found with ants. Fifth paper. Psyche 9: 3 - 5. https: // doi. org / 10.1155 / 1900 / 185328" type="journal article" year="1900">Wickham 1900: 3</bibRefCitation>
</treatmentCitation>
(note);
<treatmentCitation id="18791567FF91060BFB71FEA9FAE9C409" author="Wickham" box="[1160,1395,330,357]" page="15" pageId="2" pageNumber="140" year="1911">1911: 15 (checklist)</treatmentCitation>
. —
<treatmentCitation id="18791567FF91060BFEF6FE8EFDCDC4EA" author="Leng C. W. &amp; Wickham H. F." box="[271,599,364,391]" page="24" pageId="2" pageNumber="140" year="1920">
<bibRefCitation id="FD494E87FF91060BFEF6FE8EFDCDC4EA" author="Leng C. W. &amp; Wickham H. F." box="[271,599,364,391]" pageId="2" pageNumber="140" refId="ref4523" refString="Leng C. W. &amp; Wickham H. F. 1920. Catalogue of the Coleoptera of America, North of Mexico. JD Sherman Jr., Mount Vernon. https: // doi. org / 10.5962 / bhl. title. 8777" type="book" year="1920">Leng &amp; Wickham 1920: 24</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="18791567FF91060BFCE3FE8FFB8EC4EA" author="Holmquist A. M." box="[794,1044,364,390]" page="84" pageId="2" pageNumber="140" year="1928">
<bibRefCitation id="FD494E87FF91060BFCE3FE8FFB8EC4EA" author="Holmquist A. M." box="[794,1044,364,390]" pageId="2" pageNumber="140" pagination="70 - 87" refId="ref4438" refString="Holmquist A. M. 1928 a. Notes on the life-history and habits of the mound-building ant, Formica ulkei Emery. Ecology 9: 70 - 87. https: // doi. org / 10.2307 / 1929544" type="journal article" year="1928">Holmquist 1928a: 84</bibRefCitation>
</treatmentCitation>
(note);
<treatmentCitation id="18791567FF91060BFB8DFE8FFB60C4EB" author="Holmquist" box="[1140,1274,364,391]" page="335" pageId="2" pageNumber="140" year="1928">1928b: 335</treatmentCitation>
(note). —
<treatmentCitation id="18791567FF91060BFF13FE6CFE3AC4C5" author="Park O." box="[234,416,399,425]" page="199" pageId="2" pageNumber="140" year="1929">
<bibRefCitation id="FD494E87FF91060BFF13FE6CFE3AC4C5" author="Park O." box="[234,416,399,425]" pageId="2" pageNumber="140" pagination="195 - 215" refId="ref4751" refString="Park O. 1929. Ecological observations upon the myrmecocoles of Formica ulkei, especially Leptinus testaceus Mueller. Psyche 36: 195 - 215. https: // doi. org / 10.1155 / 1929 / 58409" type="journal article" year="1929">Park 1929: 199</bibRefCitation>
</treatmentCitation>
(record/note);
<treatmentCitation id="18791567FF91060BFDAEFE6CFCD2C4C6" author="Park" box="[599,840,399,426]" page="217" pageId="2" pageNumber="140" year="1935">1935: 217 (biology)</treatmentCitation>
. —
<treatmentCitation id="18791567FF91060BFC83FE6CFB0BC4C5" author="Blackwelder R. E." box="[890,1169,399,425]" page="100" pageId="2" pageNumber="140" year="1939">
<bibRefCitation id="FD494E87FF91060BFC83FE6CFB0BC4C5" author="Blackwelder R. E." box="[890,1169,399,425]" pageId="2" pageNumber="140" pagination="93 - 125" refId="ref4172" refString="Blackwelder R. E. 1939. A generic revision of the staphylinid beetles of the tribe Paederini. Proceedings of the United States National Museum 87: 93 - 125. https: // doi. org / 10.5479 / si. 00963801.87 - 3069.93" type="journal article" year="1939">Blackwelder 1939: 100</bibRefCitation>
</treatmentCitation>
, 119 (key and list). —
<treatmentCitation id="18791567FF91060BFEF4FE51FE52C4A0" author="Herman Jr L. H." box="[269,456,434,460]" page="3" pageId="2" pageNumber="140" year="1970">
<bibRefCitation id="FD494E87FF91060BFEF4FE51FE52C4A0" author="Herman Jr L. H." box="[269,456,434,460]" pageId="2" pageNumber="140" pagination="1 - 26" refId="ref4404" refString="Herman Jr L. H. 1970. The ecology, phylogeny, and taxonomy of Stilicolina (Coleoptera, Staphylinidae, Paederinae). American Museum Novitates 2412: 1 - 26." type="journal article" year="1970">Herman 1970: 3</bibRefCitation>
</treatmentCitation>
(note). —
<treatmentCitation id="18791567FF91060BFDBAFE51FC20C4A0" author="Moore I. &amp; Legner E. F." box="[579,954,434,460]" page="107" pageId="2" pageNumber="140" year="1979">
<bibRefCitation id="FD494E87FF91060BFDBAFE51FC20C4A0" author="Moore I. &amp; Legner E. F." box="[579,954,434,460]" pageId="2" pageNumber="140" refId="ref4629" refString="Moore I. &amp; Legner E. F. 1979. An Illustrated Guide to the Genera of the Staphylinidae of America North of Mexico Exclusive of the Aleocharinae (Coleoptera). University of California Division of Agricultural Sciences Priced Publication, Berkeley." type="book" year="1979">Moore &amp; Legner 1979: 107–108</bibRefCitation>
</treatmentCitation>
(description and drawing). —
<treatmentCitation id="18791567FF91060BFAE3FE51FD1DC483" author="Newton" page="327,387" pageId="2" pageNumber="140" year="2020">
Newton
<emphasis id="ABACEF64FF91060BFF13FE36FEBEC482" box="[234,292,468,494]" italics="true" pageId="2" pageNumber="140">et al.</emphasis>
2020: 327, 387 (key and note)
</treatmentCitation>
. —
<treatmentCitation id="18791567FF91060BFD4CFE36FC32C482" author="Bousquet et al" box="[693,936,468,495]" pageId="2" pageNumber="140" year="2013">
<bibRefCitation id="FD494E87FF91060BFD4CFE36FC32C482" author="Bousquet Y. &amp; Bouchard P. &amp; Davies A. E. &amp; Sikes D. S." box="[693,936,468,495]" pageId="2" pageNumber="140" pagination="1 - 44" refId="ref4286" refString="Bousquet Y., Bouchard P., Davies A. E. &amp; Sikes D. S. 2013. Checklist of beetles (Coleoptera) of Canada and Alaska. ZooKeys 360: 1 - 44. https: // doi. org / 10.3897 / zookeys. 360.4742" type="journal article" year="2013">
Bousquet
<emphasis id="ABACEF64FF91060BFCD1FE36FCF9C482" box="[808,867,468,494]" italics="true" pageId="2" pageNumber="140">et al.</emphasis>
2013
</bibRefCitation>
</treatmentCitation>
: DwC-A database (checklist). —
<treatmentCitation id="18791567FF91060BFAD3FE37FECBC77D" author="Parker J." page="78" pageId="2" pageNumber="140" year="2016">
<bibRefCitation id="FD494E87FF91060BFAD3FE37FECBC77D" author="Parker J." pageId="2" pageNumber="140" pagination="65 - 108" refId="ref4824" refString="Parker J. 2016. Myrmecophily in beetles (Coleoptera): evolutionary patterns and biological mechanisms. Myrmecological News 22: 65 - 108." type="journal article" year="2016">Parker 2016: 78</bibRefCitation>
</treatmentCitation>
(note).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="D1C260FDFF91060BFF44FDD4FD1BC713" pageId="2" pageNumber="140" type="type_taxon">
<paragraph id="99673376FF91060BFF44FDD4FEC2C73D" blockId="2.[189,344,567,593]" box="[189,344,567,593]" pageId="2" pageNumber="140">
<heading id="C22F841AFF91060BFF44FDD4FEC2C73D" bold="true" box="[189,344,567,593]" fontSize="11" level="3" pageId="2" pageNumber="140" reason="3">
<emphasis id="ABACEF64FF91060BFF44FDD4FEC2C73D" bold="true" box="[189,344,567,593]" pageId="2" pageNumber="140">
<typeStatus id="46638DD4FF91060BFF44FDD4FF60C73D" box="[189,250,567,593]" pageId="2" pageNumber="140">Type</typeStatus>
species
</emphasis>
</heading>
</paragraph>
<paragraph id="99673376FF91060BFF44FD86FD1BC713" blockId="2.[189,641,613,639]" box="[189,641,613,639]" pageId="2" pageNumber="140">
<taxonomicName id="5ED848F5FF91060BFF44FD86FDE7C713" ID-CoL="3YZH5" authority="Casey, 1889" authorityName="Casey" authorityYear="1889" box="[189,637,613,639]" class="Insecta" family="Staphylinidae" genus="Megastilicus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="140" phylum="Arthropoda" rank="species" species="formicarius">
<emphasis id="ABACEF64FF91060BFF44FD86FE79C713" box="[189,483,613,639]" italics="true" pageId="2" pageNumber="140">Megastilicus formicarius</emphasis>
<bibRefCitation id="FD494E87FF91060BFE13FD86FDE7C713" author="Casey T. L." box="[490,637,613,639]" pageId="2" pageNumber="140" pagination="182 - 184" refId="ref4337" refString="Casey T. L. 1889. Notes on the Paederini. Entomologica Americana 5: 182 - 184." type="journal article" year="1889">Casey, 1889</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="D1C260FDFF91060BFF44FD45FC09C67C" pageId="2" pageNumber="140" type="diagnosis">
<paragraph id="99673376FF91060BFF44FD45FEACC7AC" blockId="2.[189,310,678,704]" box="[189,310,678,704]" pageId="2" pageNumber="140">
<heading id="C22F841AFF91060BFF44FD45FEACC7AC" bold="true" box="[189,310,678,704]" fontSize="11" level="3" pageId="2" pageNumber="140" reason="3">
<emphasis id="ABACEF64FF91060BFF44FD45FEACC7AC" bold="true" box="[189,310,678,704]" pageId="2" pageNumber="140">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="99673376FF91060BFF44FD37FC09C67C" blockId="2.[189,1398,724,785]" pageId="2" pageNumber="140">Body robust, antennae short and stout. Body covered with short, black, stout setae. Dorsal surface of head, pronotum and elytra granulate. Head orbicular in shape.</paragraph>
</subSubSection>
<subSubSection id="D1C260FDFF91060AFF44FCD4FC5AC49F" lastPageId="3" lastPageNumber="141" pageId="2" pageNumber="140" type="description">
<paragraph id="99673376FF91060BFF44FCD4FEF1C63D" blockId="2.[189,363,823,849]" box="[189,363,823,849]" pageId="2" pageNumber="140">
<heading id="C22F841AFF91060BFF44FCD4FEF1C63D" bold="true" box="[189,363,823,849]" fontSize="11" level="3" pageId="2" pageNumber="140" reason="3">
<emphasis id="ABACEF64FF91060BFF44FCD4FEF1C63D" bold="true" box="[189,363,823,849]" pageId="2" pageNumber="140">Redescription</emphasis>
</heading>
</paragraph>
<paragraph id="99673376FF91060BFF44FC86FB8FC6A8" blockId="2.[189,1399,869,965]" pageId="2" pageNumber="140">BODY. Medium size beetle; body robust, somewhat flattened. Colour of specimens from pale brown/sandy to brown/reddish brown. Head, pronotum and elytra covered with short, black setae. Dorsal surface of body densely granulato-reticulate, abdomen smoother, and not granulate.</paragraph>
<paragraph id="99673376FF91060BFF44FC08FEDCC085" blockId="2.[188,1400,1003,1513]" pageId="2" pageNumber="140">HEAD. Orbicular, with rounded hind angles; widest in middle, tapering towards antennal insertions anteriorly and towards neck posteriorly; disc of head without punctation, with leather-like integument. Head wider and longer than pronotum. Antennae short, reaching only to anterior part of pronotum, compact, mostly transverse, with well-developed stems in between, pubescent. Labrum large, almost completely covering mandibles from above, notched, broadly emarginate in middle at apex, with two distinct lobes, each bearing long and pale macroseta and comb of setae near emargination; emargination bidentate; weakly sclerotised, connected with head by membrane. Mandibulae without prostheca, with two teeth on right and one on left mandibula. Maxillary palpomere 3 about twice as long as palpomere 2; palpomere 4 elongate and thin, around ⅓ of palpomere 3 length and ⅓ of palpomere 3 width. Labial palpomere 1 shorter and thinner than palpomere 2; palpomere 2 slightly elongate, expanded toward apex, twice as long as palpomere 1; palpomere 3 thin and long, longer than half of palpomere 2 and ¼ of palpomere 2 width. Mentum transverse. Submentum with pair of setae on sides. Ligula bilobed. Gular sutures fused. Eyes small, from ⅕ to ⅙ of head length. Temples long, more than half of head length. Shallow longitudinal depression on each side of head ventrally. Neck distinct and narrow, about ⅕ of head width.</paragraph>
<paragraph id="99673376FF91060BFF44F9F3FC2DC395" blockId="2.[189,1401,1551,1785]" pageId="2" pageNumber="140">THORAX. Pronotum longer than wide, anterior angles obtuse, widest in middle. Superior marginal line deflexed, becoming indistinct close to apical portion of prosternum, but reaching prosternum. Basisternum without macrosetae or microsculpture, but surface wrinkled. Furcasternum triangular, acute. Pronotal disc dull, with leather-like integument, without punctures. Elytra longer and wider than pronotum, without punctures, without epipleural ridge; humeral angles indistinct, rounded; surface dull with leather-like integument, with unorganised rows of small, black macrosetae. Anterior margin of scutellum rounded. Posterior (?) scutellar ridge present. Hind wings fully developed.</paragraph>
<paragraph id="99673376FF91060BFF44F8C3FED5C285" blockId="2.[187,1399,1823,2025]" pageId="2" pageNumber="140">LEGS. Long and slender, uniformly covered with glossy black macrosetae. All tarsi 5-segmented. Tarsomeres 1–4 of protarsus not inflated. Protibia with three fully developed protibial combs. One pair of empodial setae on each tarsus, equal to or slightly shorter than claws (never shorter than half of claw length). Meso and metatarsomere 1 longer than 2. Metatarsi almost as long as metatibiae, with tarsomeres 1–4 decreasing gradually in length, metarsomere 1 shorter than 2 and 3 combined, slightly longer than tarsomere 5.</paragraph>
<paragraph id="99673376FF90060AFF44FEEAFC97C406" blockId="3.[188,1398,265,363]" pageId="3" pageNumber="141">ABDOMEN. Sternite III with medial longitudinal keel. Abdomen widest at tergite V. Abdominal segments III to VII with pair of paratergites on each side; setae along length of abdomen. Tergite/sternite IX or X with dense setation. [Similar structure for female.]</paragraph>
<paragraph id="99673376FF90060AFF44FE71FC5AC49F" blockId="3.[189,1401,402,499]" pageId="3" pageNumber="141">
AEDEAGUS. With parameres reduced and fused to median lobe; ventral process slightly longer than uneverted internal sac; in parameral view with apex of ventral process acute, sides sinuate; dorsal plate rather large and relatively weakly sclerotised (
<figureCitation id="01E32FF3FF90060AFD22FE3AFCC7C49F" box="[731,861,473,499]" captionStart="Fig" captionStartId="4.[189,231,1928,1954]" captionTargetBox="[189,1398,975,1890]" captionTargetId="figure-168@4.[189,1398,975,1890]" captionTargetPageId="4" captionText="Fig. 1. Morphology of Megastilicus formicarius Casey, 1889. A. Dorsal habitus of paralectotype (NMNH). B. Dorsal habitus of lectotype (NMNH). C. Sternum VIII. D. Aedeagus in parameral view. E. Aedeagus in lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5718413" httpUri="https://zenodo.org/record/5718413/files/figure.png" pageId="3" pageNumber="141">Figs 1D–E</figureCitation>
,
<figureCitation id="01E32FF3FF90060AFC93FE3AFC29C49F" box="[874,947,473,499]" captionStart="Fig" captionStartId="6.[189,232,1928,1954]" captionTargetBox="[280,1309,265,1861]" captionTargetId="figure-11@6.[279,1309,265,1890]" captionTargetPageId="6" captionText="Fig. 2. Morphology of Megastilicus iowaensis sp. nov. A. Dorsal habitus of holotype (NMNH). B. Dorsal habitus of paratype (NMNH). C. Sternum VIII. D. Aedeagus in parameral view. E. Aedeagus in lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5718415" httpUri="https://zenodo.org/record/5718415/files/figure.png" pageId="3" pageNumber="141">2D–E</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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