160 lines
19 KiB
XML
160 lines
19 KiB
XML
<document id="7FF3D07CE9BE8CD190F742AD9A1E0512" ID-CLB-Dataset="24567" ID-DOI="10.11646/zootaxa.4711.3.3" ID-GBIF-Dataset="21de8a76-6a04-45d4-8e81-5fb524718c58" ID-ISSN="1175-5326" ID-Zenodo-Dep="3586554" ID-ZooBank="85D81C2D-0B66-4C0D-B708-AAF1DAD6018B" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1576823765851" checkinUser="plazi" docAuthor="Hechinger, Ryan F." docDate="2019" docId="EF6AD37789478B3BFF39FBDFFEFEFF61" docLanguage="en" docName="zootaxa.4711.3.3.pdf" docOrigin="Zootaxa 4711 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Himasthla Dietz 1909" docType="treatment" docVersion="7" lastPageNumber="475" masterDocId="1353AB0F89488B2BFFAEFF89FFC5FF9A" masterDocTitle="Guide to the trematodes (Platyhelminthes) that infect the California horn snail (Cerithideopsis californica: Potamididae: Gastropoda) as first intermediate host" masterLastPageNumber="494" masterPageNumber="459" pageNumber="474" updateTime="1698775255321" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="30FD6086113C95C6B598B7D64F0BB214">Guide to the trematodes (Platyhelminthes) that infect the California horn snail (Cerithideopsis californica: Potamididae: Gastropoda) as first intermediate host</mods:title>
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<mods:namePart id="075A8DC2C69B5635BA306D56DB59F52D">Hechinger, Ryan F.</mods:namePart>
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<mods:title id="7D7991CB305025E66CECD0B3003F812C">Zootaxa</mods:title>
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<mods:date id="A6491DCCD488BE41F417D64A8527FE0C">2019</mods:date>
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<treatment id="EF6AD37789478B3BFF39FBDFFEFEFF61" ID-DOI="http://doi.org/10.5281/zenodo.5658144" ID-GBIF-Taxon="160797598" ID-Zenodo-Dep="5658144" LSID="urn:lsid:plazi:treatment:EF6AD37789478B3BFF39FBDFFEFEFF61" httpUri="http://treatment.plazi.org/id/EF6AD37789478B3BFF39FBDFFEFEFF61" lastPageId="16" lastPageNumber="475" pageId="15" pageNumber="474">
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<subSubSection id="2FD931EA89478B24FF39FBDFFE6FFB0E" pageId="15" pageNumber="474" type="nomenclature">
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<paragraph id="677C626189478B24FF39FBDFFE99FBEB" blockId="15.[151,426,1110,1172]" box="[151,348,1110,1137]" pageId="15" pageNumber="474">
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<heading id="3C34D50D89478B24FF39FBDFFE99FBEB" bold="true" box="[151,348,1110,1137]" fontSize="11" level="1" pageId="15" pageNumber="474" reason="1">
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<emphasis id="55B7BE7389478B24FF39FBDFFE99FBEB" bold="true" box="[151,348,1110,1137]" pageId="15" pageNumber="474">
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<taxonomicName id="A0C319E289478B24FF39FBDFFED3FBEB" authorityName="Dietz" authorityYear="1909" box="[151,278,1110,1137]" class="Trematoda" family="Echinostomatidae" genus="Himasthla" kingdom="Animalia" order="Plagiorchiida" pageId="15" pageNumber="474" phylum="Platyhelminthes" rank="genus">
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<emphasis id="55B7BE7389478B24FF39FBDFFED3FBEB" bold="true" box="[151,278,1110,1137]" italics="true" pageId="15" pageNumber="474">Himasthla</emphasis>
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</taxonomicName>
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sp. B
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="677C626189478B24FF39FBF3FE6FFB0E" blockId="15.[151,426,1110,1172]" box="[151,426,1146,1172]" pageId="15" pageNumber="474">
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(7. Hisb;
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<figureCitation id="FFF87EE489478B24FEAEFBF3FE89FB0E" box="[256,332,1146,1172]" captionStart="FIGURE 1" captionStartId="3.[151,250,1428,1453]" captionTargetBox="[151,1436,522,1400]" captionTargetId="figure@3.[151,1436,522,1400]" captionTargetPageId="3" captionText="FIGURE 1. General characteristics of the parthenitae and cercariae of the trematodes infecting Cerithideopsis californica as first intermediate host. Species numbers and codes follow Table 1 and species accounts. Cercariae are all to scale, with additional magnified views of six small species (indicated by dashed lines). Note the oral stylets (presented in right lateral view) for Pruc and Smmi. Parthenitae are not to scale. Scale bars consistently indicate 100 µm." figureDoi="http://doi.org/10.5281/zenodo.3586556" httpUri="https://zenodo.org/record/3586556/files/figure.png" pageId="15" pageNumber="474">Figs. 1</figureCitation>
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,
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<figureCitation id="FFF87EE489478B24FEF4FBF3FE67FB0E" box="[346,418,1146,1172]" captionStart="FIGURES 27–29" captionStartId="16.[151,264,1431,1456]" captionTargetBox="[188,1423,270,1404]" captionTargetId="figure@16.[164,1423,270,1404]" captionTargetPageId="16" captionText="FIGURES 27–29. Himasthla sp. B (Hisb). 27, Overview of a colony in a freshly deshelled, infected horn snail in sea water. The arrow indicates the colony, which is localized in the gonadal region and part of the basal visceral mass. Scale bar = 1 cm. Base photo credit: Todd Huspeni. 28, Reproductive redia, live, with developing cercariae, under coverslip pressure, with an adjacent soldier redia indicated by the black arrow. Scale bar = 100 µm. 29, Cercariae, live, under coverslip pressure. Scale bar = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.3586576" httpUri="https://zenodo.org/record/3586576/files/figure.png" pageId="15" pageNumber="474">27–29</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="2FD931EA89478B24FF39FB48FAF8F9BA" pageId="15" pageNumber="474" type="diagnosis">
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<paragraph id="677C626189478B24FF39FB48FBFBFABE" blockId="15.[151,1437,1217,2037]" pageId="15" pageNumber="474">
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<emphasis id="55B7BE7389478B24FF39FB48FED1FB46" bold="true" box="[151,276,1217,1244]" pageId="15" pageNumber="474">Diagnosis:</emphasis>
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<emphasis id="55B7BE7389478B24FEB3FB4BFE6CFB46" box="[285,425,1218,1244]" italics="true" pageId="15" pageNumber="474">Parthenitae.</emphasis>
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Colony comprised of active rediae, densely concentrated in snail gonad with dispersion into basal visceral mass regions. Rediae translucent orange to colorless; ~
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<quantity id="A03BCF8489478B24FC70FB6CFBBAFA9B" box="[990,1151,1253,1281]" metricMagnitude="0" metricUnit="m" metricValue="1.375" metricValueMax="2.0" metricValueMin="0.75" pageId="15" pageNumber="474" unit="mm" value="1375.0" valueMax="2000.0" valueMin="750.0">750–2000 µm</quantity>
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long, oblong to elongate (length:width up to ~5:1), with posterior appendages that are often not pronounced.
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</paragraph>
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<paragraph id="677C626189478B24FF69FAA6FCF9FA2E" blockId="15.[151,1437,1217,2037]" pageId="15" pageNumber="474">
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<emphasis id="55B7BE7389478B24FF69FAA6FEF6FAD3" box="[199,307,1327,1353]" italics="true" pageId="15" pageNumber="474">
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<taxonomicName id="A0C319E289478B24FF69FAA6FEEAFAD3" authorityName="O.F.Muller" authorityYear="1773" box="[199,303,1327,1353]" class="Trematoda" family="Colepidae" genus="Cercaria" kingdom="Animalia" order="Prostomatida" pageId="15" pageNumber="474" phylum="Platyhelminthes" rank="genus">Cercaria</taxonomicName>
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.
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</emphasis>
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Body opaque white; non-oculate; with oral and ventral sucker; with main excretory ducts forming a broad “v”; with basal portions of main excretory ducts being particularly inflated and filled with large, distinct granules; collar with 25 collar spines in a single row, and
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<quantity id="A03BCF8489478B24FCB4FAFFFC80FA0A" box="[794,837,1398,1424]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="15" pageNumber="474" unit="in" value="4.0">4 in</quantity>
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a second row (2 pairs of “corner spines”); body ~
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<quantity id="A03BCF8489478B24FADFFAFCFF78FA2F" metricMagnitude="-1" metricUnit="m" metricValue="5.5" pageId="15" pageNumber="474" unit="mm" value="550.0">550 µm</quantity>
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long, ~equal in length to tail; tail dorso-ventrally finned.
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</paragraph>
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<paragraph id="677C626189478B24FF69FA34FC16FA66" blockId="15.[151,1437,1217,2037]" pageId="15" pageNumber="474">
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<emphasis id="55B7BE7389478B24FF69FA34FE68FA42" bold="true" box="[199,429,1469,1496]" pageId="15" pageNumber="474">
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<taxonomicName id="A0C319E289478B24FF69FA34FEF5FA42" authorityName="O.F.Muller" authorityYear="1773" box="[199,304,1469,1496]" class="Trematoda" family="Colepidae" genus="Cercaria" kingdom="Animalia" order="Prostomatida" pageId="15" pageNumber="474" phylum="Platyhelminthes" rank="genus">Cercaria</taxonomicName>
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behavior:
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</emphasis>
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Fresh, emerged cercariae remain in water column, swim ~continuously, body ventrally flexed (~forming an overall spheroidal shape), lashing tail back and forth.
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</paragraph>
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<paragraph id="677C626189478B24FF69F98CFAF8F9BA" blockId="15.[151,1437,1217,2037]" box="[199,1341,1541,1569]" pageId="15" pageNumber="474">
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<emphasis id="55B7BE7389478B24FF69F98CFE47F9BA" bold="true" box="[199,386,1541,1568]" pageId="15" pageNumber="474">Similar species:</emphasis>
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Hisb is most readily distinguished from all the echinostomatoids by having a tail fin.
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</paragraph>
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</subSubSection>
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<subSubSection id="2FD931EA89478B3BFF69F9A3FEFEFF61" lastPageId="16" lastPageNumber="475" pageId="15" pageNumber="474" type="discussion">
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<paragraph id="677C626189478B24FF69F9A3FCC9F9F2" blockId="15.[151,1437,1217,2037]" pageId="15" pageNumber="474">
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<emphasis id="55B7BE7389478B24FF69F9A3FEF8F9DE" bold="true" box="[199,317,1578,1604]" pageId="15" pageNumber="474">Remarks:</emphasis>
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The rediae and cercariae of this species have not been thoroughly described. However,
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<bibRefCitation id="03521F9089478B24FAA3F9A3FED6F9F3" author="Hechinger, R. F. & Wood, A. C. & Kuris, A. M." pageId="15" pageNumber="474" pagination="656 - 665" refId="ref20780" refString="Hechinger, R. F., Wood, A. C. & Kuris, A. M. (2011 b) Social organization in a flatworm: trematode parasites form soldier and reproductive castes. Proceedings of the Royal Society: Biological sciences, 278, 656 - 665. https: // doi. org / 10.1098 / rspb. 2010.1753" type="journal article" year="2011">
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Hechinger
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<emphasis id="55B7BE7389478B24FA26F9A2FF68F9F2" italics="true" pageId="15" pageNumber="474">et al</emphasis>
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. (2011b)
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</bibRefCitation>
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present several aspects of redia morphology.
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</paragraph>
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<paragraph id="677C626189478B24FF69F9FBFA54F92A" blockId="15.[151,1437,1217,2037]" pageId="15" pageNumber="474">
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This species corresponds to the ‘fin-tailed echinostome” of
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<bibRefCitation id="03521F9089478B24FCC1F9FBFBCBF916" author="Martin, W. E." box="[879,1038,1649,1676]" pageId="15" pageNumber="474" pagination="203 - 210" refId="ref21967" refString="Martin, W. E. (1955) Seasonal infections of the snail, Cerithidea californica Haldeman, with larval trematodes. In: Essays in Natural Science in Honor of Captain Alan Hancock on the occasion of his birthday. University of Southern California Press, Los Angeles, California, pp. 203 - 210." type="book chapter" year="1955">Martin (1955)</bibRefCitation>
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and the “
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<taxonomicName id="A0C319E289478B24FBD1F9FBFA8DF916" authorityName="Dietz" authorityYear="1909" box="[1151,1352,1650,1676]" class="Trematoda" family="Echinostomatidae" genus="Echinoparyphium" kingdom="Animalia" order="Plagiorchiida" pageId="15" pageNumber="474" phylum="Platyhelminthes" rank="genus">
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<emphasis id="55B7BE7389478B24FBD1F9FBFA8DF916" box="[1151,1352,1650,1676]" italics="true" pageId="15" pageNumber="474">Echinoparyphium</emphasis>
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</taxonomicName>
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sp.” of
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<bibRefCitation id="03521F9089478B24FF39F91FFEF2F92A" author="Martin, W. E." box="[151,311,1686,1712]" pageId="15" pageNumber="474" pagination="39 - 43" refId="ref22195" refString="Martin, W. E. (1972) An annotated key to the cercariae that develop in the snail Cerithidea californica. Bulletin of the Southern California Academy of Sciences, 71, 39 - 43." type="journal article" year="1972">Martin (1972)</bibRefCitation>
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, but it has been considered to be a species of
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<taxonomicName id="A0C319E289478B24FC9BF91FFC6FF92A" authorityName="Dietz" authorityYear="1909" box="[821,938,1686,1712]" class="Trematoda" family="Echinostomatidae" genus="Himasthla" kingdom="Animalia" order="Plagiorchiida" pageId="15" pageNumber="474" phylum="Platyhelminthes" rank="genus">
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<emphasis id="55B7BE7389478B24FC9BF91FFC6FF92A" box="[821,938,1686,1712]" italics="true" pageId="15" pageNumber="474">Himasthla</emphasis>
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</taxonomicName>
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since Huspeni’s unpublished thesis (2000).
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</paragraph>
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<paragraph id="677C626189478B24FF39F933FA8DF963" blockId="15.[151,1437,1217,2037]" pageId="15" pageNumber="474">Based on observations of natural infections at various stages of development, it appears that the initial mother sporocyst frequently invades the snail heart lumen (first pointed out to me by TC Huspeni; pers. observations).</paragraph>
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<paragraph id="677C626189478B24FF39F88AFD56F8DA" blockId="15.[151,1437,1217,2037]" pageId="15" pageNumber="474">
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Mature, ripe colonies comprise ~24% the soft-tissue weight of an infected snail (summer-time estimate derived from information in [
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<bibRefCitation id="03521F9089478B24FE2DF8AFFDBAF8DA" author="Hechinger, R. F. & Lafferty, K. D. & Mancini III, F. T. & Warner, R. R. & Kuris, A. M." box="[387,639,1829,1857]" pageId="15" pageNumber="474" pagination="651 - 667" refId="ref20498" refString="Hechinger, R. F., Lafferty, K. D., Mancini III, F. T., Warner, R. R. & Kuris, A. M. (2009) How large is the hand in the puppet? Ecological and evolutionary factors affecting body mass of 15 trematode parasitic castrators in their snail host. Evolutionary Ecology, 23, 651 - 667. https: // doi. org / 10.1007 / s 10682 - 008 - 9262 - 4" type="journal article" year="2009">
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Hechinger
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||
<emphasis id="55B7BE7389478B24FDACF8AEFDF6F8DA" box="[514,563,1830,1856]" italics="true" pageId="15" pageNumber="474">et al</emphasis>
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. 2009
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</bibRefCitation>
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]).
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</paragraph>
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<paragraph id="677C626189478B24FF69F8C3FA81F8FE" blockId="15.[151,1437,1217,2037]" box="[199,1348,1866,1893]" pageId="15" pageNumber="474">
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Hisb infection causes (stolen) snail bodies to grow ~
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<date id="137D44A189478B24FCBCF8C3FCEAF8FE" box="[786,815,1866,1892]" pageId="15" pageNumber="474">2x</date>
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faster than uninfected snails (
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<bibRefCitation id="03521F9089478B24FBD3F8C3FAFDF8FE" author="Hechinger, R. F." box="[1149,1336,1866,1892]" pageId="15" pageNumber="474" pagination="1 - 14" refId="ref20237" refString="Hechinger, R. F. (2010) Mortality affects adaptive allocation to growth and reproduction: field evidence from a guild of body snatchers. BMC Evolutionary Biology, 10 (136), 1 - 14. https: // doi. org / 10.1186 / 1471 - 2148 - 10 - 136" type="journal article" year="2010">Hechinger 2010</bibRefCitation>
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).
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</paragraph>
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<paragraph id="677C626189478B24FF69F8E7FB44F837" blockId="15.[151,1437,1217,2037]" pageId="15" pageNumber="474">
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Hisb has a caste of soldier rediae (
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<bibRefCitation id="03521F9089478B24FDE8F8E7FC89F813" author="Hechinger, R. F. & Wood, A. C. & Kuris, A. M." box="[582,844,1902,1929]" pageId="15" pageNumber="474" pagination="656 - 665" refId="ref20780" refString="Hechinger, R. F., Wood, A. C. & Kuris, A. M. (2011 b) Social organization in a flatworm: trematode parasites form soldier and reproductive castes. Proceedings of the Royal Society: Biological sciences, 278, 656 - 665. https: // doi. org / 10.1098 / rspb. 2010.1753" type="journal article" year="2011">
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||
Hechinger
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||
<emphasis id="55B7BE7389478B24FD6DF8E6FD31F812" box="[707,756,1902,1928]" italics="true" pageId="15" pageNumber="474">et al</emphasis>
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||
. 2011b
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</bibRefCitation>
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||
). In fact, it is first trematode species for which it was documented that trematodes can have a reproductive division of labor and a soldier caste.
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</paragraph>
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<paragraph id="677C626189478B24FF69F83FFE42F86F" blockId="15.[151,1437,1217,2037]" pageId="15" pageNumber="474">
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<bibRefCitation id="03521F9089478B24FF69F83FFE40F84B" author="Nadakal, A. M." box="[199,389,1974,2001]" pageId="15" pageNumber="474" pagination="777 - 786" refId="ref22487" refString="Nadakal, A. M. (1960 b) Types and sources of pigments in certain species of larval trematodes. Journal of Parasitology, 46, 777 - 786. https: // doi. org / 10.2307 / 3275532" type="journal article" year="1960">Nadakal (1960b)</bibRefCitation>
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presents information on the pigments of the rediae and cercariae of this species (as his “fintailed echinostome”).
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</paragraph>
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<paragraph id="677C626189588B3BFF69FF10FEFEFF61" blockId="16.[151,1436,153,251]" pageId="16" pageNumber="475">
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As part of one of the first studies documenting the syncytial nature of trematode integuments,
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<bibRefCitation id="03521F9089588B3BFB42FF10FF27FF42" author="Bils, R. F. & Martin, W. E." pageId="16" pageNumber="475" pagination="78 - 88" refId="ref19555" refString="Bils, R. F. & Martin, W. E. (1966) Fine Structure and Development of the Trematode Integument. Transactions of the American Microscopical Society, 85, 78 - 88. https: // doi. org / 10.2307 / 3224777" type="journal article" year="1966">Bils and Martin (1966)</bibRefCitation>
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||
examined the fine structure and development of the tegument for the rediae and cercariae of this species (as
|
||
<taxonomicName id="A0C319E289588B3BFF39FF68FEC1FF61" authorityName="Navas" authorityYear="1910" box="[151,260,225,251]" class="Insecta" family="Mantispidae" genus="Molinella" higherTaxonomySource="CoL" kingdom="Animalia" order="Neuroptera" pageId="16" pageNumber="475" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="55B7BE7389588B3BFF39FF68FEC1FF61" box="[151,260,225,251]" italics="true" pageId="16" pageNumber="475">Molinella</emphasis>
|
||
</taxonomicName>
|
||
sp.).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |