171 lines
15 KiB
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171 lines
15 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.164.2201" ID-GBIF-Dataset="18e175c4-93e1-4cba-a7fd-3b05815a9af4" ID-PMC="PMC3272619" ID-Pensoft-Pub="1313-2970-164-1" ID-PubMed="22328849" ModsDocAuthor="" ModsDocDate="2012" ModsDocID="1313-2970-164-1" ModsDocOrigin="ZooKeys 164" ModsDocTitle="Five new species of the genera Heerz Marsh, Lissopsius Marsh and Ondigus Braet, Barbalho and van Achterberg (Braconidae, Doryctinae) from the Chamela-Cuixmala biosphere reserve in Jalisco, Mexico" checkinTime="1451249389041" checkinUser="pensoft" docAuthor="Zaldivar-Riveron, Alejandro, Martinez, Juan Jose, Ceccarelli, Fadia Sara & Shaw, Scott R." docDate="2012" docId="7AAFE2F722B40A128BED5FF9CCDF14DF" docLanguage="en" docName="ZooKeys 164: 1-23" docOrigin="ZooKeys 164" docSource="http://dx.doi.org/10.3897/zookeys.164.2201" docTitle="Heerz Marsh" docType="treatment" docVersion="3" lastPageNumber="6" masterDocId="4B553355FFC5FFA9AC7BFFE1FD18FFD2" masterDocTitle="Five new species of the genera Heerz Marsh, Lissopsius Marsh and Ondigus Braet, Barbalho and van Achterberg (Braconidae, Doryctinae) from the Chamela-Cuixmala biosphere reserve in Jalisco, Mexico" masterLastPageNumber="23" masterPageNumber="1" pageNumber="4" updateTime="1668153157884" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Five new species of the genera Heerz Marsh, Lissopsius Marsh and Ondigus Braet, Barbalho and van Achterberg (Braconidae, Doryctinae) from the Chamela-Cuixmala biosphere reserve in Jalisco, Mexico</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zaldivar-Riveron, Alejandro</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Martinez, Juan Jose</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ceccarelli, Fadia Sara</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Shaw, Scott R.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2012</mods:date>
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<mods:detail type="volume">
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<mods:number>164</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>23</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.164.2201</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.164.2201</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-164-1</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152033019" LSID="urn:lsid:plazi:treatment:7AAFE2F722B40A128BED5FF9CCDF14DF" httpUri="http://treatment.plazi.org/id/7AAFE2F722B40A128BED5FF9CCDF14DF" lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4">
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<subSubSection pageId="3" pageNumber="4" type="nomenclature">
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<paragraph pageId="3" pageNumber="4">
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<taxonomicName LSID="http://species-id.net/wiki/Heerz" authority="Marsh" class="Insecta" family="Braconidae" genus="Heerz" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heerz" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">Heerz Marsh</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="4" type="reference_group">
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<paragraph pageId="3" pageNumber="4">
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<taxonomicName class="Insecta" family="Braconidae" genus="Heerz" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heerz" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">Heerz</taxonomicName>
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Marsh, 1993: 17;
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<bibRefCitation author="Marsh, PM" journalOrPublisher="Memoirs of the American Entomological Institute" pageId="16" pageNumber="17" pagination="1 - 319" title="The Doryctinae of Costa Rica (excluding the genus Heterospilus)." volume="70" year="2002">Marsh 2002</bibRefCitation>
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: 102.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="4" type="type species">
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<paragraph pageId="3" pageNumber="4">Type species.</paragraph>
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<paragraph pageId="3" pageNumber="4">
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<taxonomicName class="Insecta" family="Braconidae" genus="Heerz" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heerz lukenatcha" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="lukenatcha">Heerz lukenatcha</taxonomicName>
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Marsh
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="4" type="diagnosis">
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<paragraph pageId="3" pageNumber="4">Diagnosis.</paragraph>
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<paragraph pageId="3" pageNumber="4">
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<taxonomicName class="Insecta" family="Braconidae" genus="Heerz" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heerz" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">Heerz</taxonomicName>
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distinguishes from other doryctine genera by the following combination of features: (1) frons excavated (Fig. 2D), (2) propodeum with a longitudinal median carina followed by a pentagonal areola (Figs 2C, 3D), (3) second metasomal tergite entirely or partially sculptured, contrasting with third one, which is smooth, polished and with a transverse furrow (Figs 2C, 3D), (4) vein r-m of fore wing present (Fig. 2F), (5) vein M+CU of hind about as long as vein 1M, (6) vein cu-a of hind wing straight or slightly curved apically towards wing apex, (7) male hind wing without pterostigma, and (8) hind coxa with a distinct basoventral tooth. Species of
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<taxonomicName class="Insecta" family="Braconidae" genus="Heerz" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heerz" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">Heerz</taxonomicName>
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are very similar in habitus to those of
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<taxonomicName class="Insecta" family="Braconidae" genus="Lissopsius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lissopsius" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">Lissopsius</taxonomicName>
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, all having a body mostly smooth and shiny, propodeum with a longitudinal median carina followed by a pentagonal areola, and vein M+CU of hind wing slightly shorter to larger than vein 1M. However,
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<taxonomicName class="Insecta" family="Braconidae" genus="Heerz" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heerz" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">Heerz</taxonomicName>
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differs from
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<taxonomicName class="Insecta" family="Braconidae" genus="Lissopsius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lissopsius" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">Lissopsius</taxonomicName>
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by having the vein r-m of fore wing present (Fig. 2F) (absent in
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<taxonomicName class="Insecta" family="Braconidae" genus="Lissopsius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lissopsius" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">Lissopsius</taxonomicName>
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), hind coxa with a basoventral tooth (absent in
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<taxonomicName class="Insecta" family="Braconidae" genus="Lissopsius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lissopsius" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">Lissopsius</taxonomicName>
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), and ovipositor distinctly sclerotised apically (uniformly slcerotised in
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<taxonomicName class="Insecta" family="Braconidae" genus="Lissopsius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lissopsius" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">Lissopsius</taxonomicName>
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).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4" type="description">
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<paragraph pageId="3" pageNumber="4">Description.</paragraph>
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<paragraph lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4">
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Small to moderate size, 2.5-7.0 mm; eyes large, moderately to distinctly emarginated opposite antennal sockets; frons concave; occipital carina present,
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<pageBreakToken pageId="4" pageNumber="5" start="start">meeting</pageBreakToken>
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hypostomal carina before mandible; labrum distinctly concave; hypoclypeal depression small and round; clypeus short; malar suture absent; maxillary palpi 5-segmented, labial palpi 4-segmented; head and mesosoma smooth or weakly sculptured; mesoscutum declivous anteriorly; prepectal carinae present; precoxal sulcus smooth; surface of propodeum smooth on anterior half, slightly rugose on posterior half, with a median longitudinal carina followed by a pentagonal areola; metapleural flange present; fore tibia with a row of spines along anterior edge; hind coxa with a distinct basoventral tooth; vein m-cu of fore wing antefurcal to vein 2RS, thus (RS+M)b present;
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<pageBreakToken pageId="5" pageNumber="6" start="start">vein</pageBreakToken>
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1cu-a postfurcal to vein 1M; vein r-m of fore wing present; second submarginal cell distinctly short; first subdiscal cell of fore wing open at apex; vein M+CU of hind wing slightly shorter to larger than vein 1M; males without pterostigma on hind wing; basal sternal plate (acrosternite) of first metasomal tergite short, 0.2-0.3 times the length of tergum; first and second metasomal tergites scupltured; third metasomal tergite smooth with a transverse furrow; remaining metasomal tergites smooth; ovipositor strongly sclerotised apically; nodes reduced, only one or absent.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="6" type="distribution">
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<paragraph pageId="5" pageNumber="6">Distribution.</paragraph>
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<paragraph pageId="5" pageNumber="6">Brazil, Costa Rica and Mexico.</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="6" type="remarks">
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<paragraph pageId="5" pageNumber="6">Remarks.</paragraph>
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<paragraph pageId="5" pageNumber="6">
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The two new species of
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<taxonomicName class="Insecta" family="Braconidae" genus="Heerz" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heerz" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">Heerz</taxonomicName>
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described below considerably modify the previous concept of the genus. The two previously described species,
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<taxonomicName class="Insecta" family="Braconidae" genus="Heerz" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heerz lukenatcha" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="lukenatcha">Heerz lukenatcha</taxonomicName>
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and
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<taxonomicName class="Insecta" family="Braconidae" genus="Heerz" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heerz tooya" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="tooya">Heerz tooya</taxonomicName>
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Marsh, are characterised by their smooth mesosoma, dusky wings and relatively large body size. The Mexican species, on the other hand, have a coriaceous mesoscutum, uniformly hyaline wings and are considerably smaller, especially
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<taxonomicName class="Insecta" family="Braconidae" genus="Heerz" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heerz ecmahla" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="ecmahla">Heerz ecmahla</taxonomicName>
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. Moreover, frons excavation is more conspicuous in the two new species compared to
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<taxonomicName class="Insecta" family="Braconidae" genus="Heerz" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heerz lukenatcha" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="lukenatcha">Heerz lukenatcha</taxonomicName>
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and
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<taxonomicName class="Insecta" family="Braconidae" genus="Heerz" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heerz tooya" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="tooya">Heerz tooya</taxonomicName>
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. Despite these morphological differences, our comparisons with type material and our DNA sequence data (see below) led us to include the new species within
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<taxonomicName class="Insecta" family="Braconidae" genus="Heerz" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heerz" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">Heerz</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="6" type="key to described species of heerz (modified from marsh 1993)">
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<paragraph pageId="5" pageNumber="6">
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Key to described species of
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<taxonomicName class="Insecta" family="Braconidae" genus="Heerz" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heerz" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">Heerz</taxonomicName>
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(modified from
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<bibRefCitation author="Marsh, PM" journalOrPublisher="Contributions of the American Entomological Institute" pageId="16" pageNumber="17" pagination="1 - 58" volume="28" year="1993">Marsh 1993</bibRefCitation>
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)
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</paragraph>
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<paragraph pageId="5" pageNumber="6">
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<table pageId="5" pageNumber="6">
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<tr pageId="5" pageNumber="6">
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<td colspan="1" pageId="5" pageNumber="6" rowspan="1">Fig. 2B</td>
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</tr>
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<tr pageId="5" pageNumber="6">
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<td colspan="1" pageId="5" pageNumber="6" rowspan="1">Fig. 2A3AFig. 3C</td>
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</tr>
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<tr pageId="5" pageNumber="6">
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<td colspan="1" pageId="5" pageNumber="6" rowspan="1">
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<taxonomicName class="Insecta" family="Braconidae" genus="Heerz" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heerz lukentacha" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="lukentacha">Heerz lukentacha</taxonomicName>
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</td>
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</tr>
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<tr pageId="5" pageNumber="6">
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<td colspan="1" pageId="5" pageNumber="6" rowspan="1">
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<taxonomicName class="Insecta" family="Braconidae" genus="Heerz" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heerz tooya" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="tooya">Heerz tooya</taxonomicName>
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</td>
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</tr>
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<tr pageId="5" pageNumber="6">
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<td colspan="1" pageId="5" pageNumber="6" rowspan="1">Fig. 3BFig. 3DFig. 3A</td>
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<td colspan="1" pageId="5" pageNumber="6" rowspan="1">
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<taxonomicName class="Insecta" family="Braconidae" genus="Heerz" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heerz macrophthalma" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="macrophthalma">Heerz macrophthalma</taxonomicName>
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</td>
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</tr>
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<tr pageId="5" pageNumber="6">
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<td colspan="1" pageId="5" pageNumber="6" rowspan="1">Fig. 2DFig. 2CFig. 2A</td>
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<td colspan="1" pageId="5" pageNumber="6" rowspan="1">
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<taxonomicName class="Insecta" family="Braconidae" genus="Heerz" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heerz ecmahla" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="ecmahla">Heerz ecmahla</taxonomicName>
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</td>
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</tr>
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</table>
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |