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<document id="2F70A1EABAA2064270565306624F352E" ID-CLB-Dataset="20167" ID-DOI="10.5852/ejt.2022.803.1691" ID-GBIF-Dataset="af09c8e1-b481-4de0-b9d0-83ba26bf9876" ID-ISSN="2118-9773" ID-Zenodo-Dep="6359066" ID-ZooBank="8B66C8AE-F00A-42F6-9641-26B0ECC49F78" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1647350252755" checkinUser="felipe" docAuthor="Enghoff, Henrik" docDate="2022" docId="03F15C39D63B0616FDC9FA6AC03EFD1D" docLanguage="en" docName="ejt-803.1-136.pdf" docOrigin="European Journal of Taxonomy 803" docSource="http://dx.doi.org/10.5852/ejt.2022.803.1691" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Calyptomastix xystopygoides Enghoff 2022, sp. nov." docType="treatment" docUuid="0F641CA1-EB6E-4EB4-82A4-0BE936074D2E" docUuidSource="ZooBank" docVersion="8" lastPageNumber="34" masterDocId="FFC82441D6250637FFF3FFE3C52CFFD8" masterDocTitle="Mountains of millipedes. The family Odontopygidae in the Eastern Arc Mountains of Tanzania (Diplopoda, Spirostreptida)" masterLastPageNumber="136" masterPageNumber="1" pageNumber="31" updateTime="1699318071475" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="1CB3D8FE73100B1E093746425C415192">Mountains of millipedes. The family Odontopygidae in the Eastern Arc Mountains of Tanzania (Diplopoda, Spirostreptida)</mods:title>
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<mods:namePart id="491353630687EEA7CF543F4D27E7CBED">Enghoff, Henrik</mods:namePart>
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<mods:affiliation id="1B35F2300F757FFE3ED12FE626B84DEC">Natural History Museum of Denmark, University of Copenhagen, Universitetsparken 15, DK- 2100 Copenhagen Ø, Denmark.</mods:affiliation>
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<treatment id="03F15C39D63B0616FDC9FA6AC03EFD1D" ID-DOI="http://doi.org/10.5281/zenodo.6373800" ID-GBIF-Taxon="194181648" ID-Zenodo-Dep="6373800" LSID="urn:lsid:zoobank.org:act:0F641CA1-EB6E-4EB4-82A4-0BE936074D2E" httpUri="http://treatment.plazi.org/id/03F15C39D63B0616FDC9FA6AC03EFD1D" lastPageId="33" lastPageNumber="34" pageId="30" pageNumber="31">
<subSubSection id="C342BEA4D63B0629FDC9FA6AC6D6FA7C" box="[570,1018,1417,1444]" pageId="30" pageNumber="31" type="nomenclature">
<paragraph id="8BE7ED2FD63B0629FDC9FA6AC6D6FA7C" blockId="30.[364,1223,1417,1520]" box="[570,1018,1417,1444]" pageId="30" pageNumber="31">
<heading id="D0AF5A43D63B0629FDC9FA6AC6D6FA7C" box="[570,1018,1417,1444]" centered="true" fontSize="11" level="2" pageId="30" pageNumber="31" reason="2">
<taxonomicName id="4C5896ACD63B0629FDC9FA6AC6B4FA7C" ID-CoL="8ZDSK" authority="Enghoff, 2022" authorityName="Enghoff" authorityYear="2022" box="[570,920,1417,1444]" class="Diplopoda" family="Odontopygidae" genus="Calyptomastix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="xystopygoides" status="sp. nov.">
<emphasis id="B92C313DD63B0629FDC9FA6AC6B4FA7C" bold="true" box="[570,920,1417,1444]" italics="true" pageId="30" pageNumber="31">Calyptomastix xystopygoides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21F8C46D63B0629FC6CFA69C6D6FA7C" box="[927,1018,1418,1444]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="C342BEA4D63B0629FE9FFA53C677FA28" pageId="30" pageNumber="31" type="description">
<paragraph id="8BE7ED2FD63B0629FE9FFA53C1EBFA12" blockId="30.[364,1223,1417,1520]" box="[364,1223,1456,1482]" pageId="30" pageNumber="31">
<uri id="FFC9E12DD63B0629FE9FFA53C1EBFA12" box="[364,1223,1456,1482]" pageId="30" pageNumber="31">
urn:lsid:zoobank.org:act:
<uuid id="FFFED7FAD63B0629FD7EFA53C1EBFA12" box="[653,1223,1456,1482]" pageId="30" pageNumber="31">0F641CA1-EB6E-4EB4-82A4-0BE936074D2E</uuid>
</uri>
</paragraph>
<paragraph id="8BE7ED2FD63B0629FD2BFA35C677FA28" blockId="30.[364,1223,1417,1520]" box="[728,859,1494,1520]" pageId="30" pageNumber="31">
<figureCitation id="1363F1AAD63B0629FD2BFA35C677FA28" box="[728,859,1494,1520]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="31.[189,232,1785,1811]" captionStartId-1="32.[189,232,1857,1883]" captionTargetBox-0="[189,1398,464,1754]" captionTargetBox-1="[189,1398,854,1826]" captionTargetId-0="figure-170@31.[189,1398,464,1754]" captionTargetId-1="figure-278@32.[189,1398,854,1826]" captionTargetPageId-0="31" captionTargetPageId-1="32" captionText-0="Fig. 19. Calyptomastix xystopygoides sp. nov., holotype, ♂ (NHMD 621718). AC. First pair of legs. A. Sublateral view. B. Anterior view. C. Ventral view. D. Midbody dorsal limbus (worn). E. Midbody ventrolateral limbus. FI. Left gonopod coxa. F. Anterior view. G. Anterior-lateral view. H. Posteriormesal view. I. lateral view. Abbreviations: APS =mesapical prefemoral setae; CU = cucullus; cus = cucullar spine; CXS =coxosternal setae; hh =hemisphaerical hump; LCS= lateral coxal spine; LPS =lateral prefemoral setae; MP=metaplica; ms =metazonital striae; PP =proplica, Scale bars: AC, FI =0.1 mm; DE=0.02 mm." captionText-1="Fig. 20. Calyptomastix xystopygoides sp. nov., holotype, ♂ (NHMD 621718), left gonopod telopodite. A. anterior-mesal view. B. Posterior view. C. Ventral view. D. Dorsal view. E. Detail of telomere and ridged solenomere. Abbreviations: BSS =basal solenomeral spine; PN =post-torsal narrowing; SLM =solenomere; TL =torsotope lobe; tml =telomeral spinelike lobe; TT=torsotope; PTS = post-torsal spine. Scale bars: AD =0.2 mm; E =0.05 mm." figureDoi-0="http://doi.org/10.5281/zenodo.6359127" figureDoi-1="http://doi.org/10.5281/zenodo.6359131" httpUri-0="https://zenodo.org/record/6359127/files/figure.png" httpUri-1="https://zenodo.org/record/6359131/files/figure.png" pageId="30" pageNumber="31">Figs 1920</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C342BEA4D63B0629FF4EF9F9C787F969" pageId="30" pageNumber="31" type="diagnosis">
<paragraph id="8BE7ED2FD63B0629FF4EF9F9C41AF9EC" blockId="30.[189,310,1562,1588]" box="[189,310,1562,1588]" pageId="30" pageNumber="31">
<heading id="D0AF5A43D63B0629FF4EF9F9C41AF9EC" bold="true" box="[189,310,1562,1588]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="B92C313DD63B0629FF4EF9F9C41AF9EC" bold="true" box="[189,310,1562,1588]" pageId="30" pageNumber="31">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BE7ED2FD63B0629FF4EF9A8C787F969" blockId="30.[188,1398,1611,1713]" pageId="30" pageNumber="31">
Differs from other species of
<taxonomicName id="4C5896ACD63B0629FDE3F9A8C795F9BD" authorityName="Hoffman &amp; Howell" authorityYear="2012" box="[528,697,1611,1637]" class="Diplopoda" family="Odontopygidae" genus="Calyptomastix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B92C313DD63B0629FDE3F9A8C795F9BD" box="[528,697,1611,1637]" italics="true" pageId="30" pageNumber="31">Calyptomastix</emphasis>
</taxonomicName>
by having the post-torsal spine straight and directed laterad, by having the telomere more slender and curved in a
<taxonomicName id="4C5896ACD63B0629FCD0F991C6B0F953" authorityName="Attems" authorityYear="1909" box="[803,924,1650,1675]" class="Diplopoda" family="Odontopygidae" genus="Xystopyge" kingdom="Animalia" order="Spirostreptida" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B92C313DD63B0629FCD0F991C6B0F953" box="[803,924,1650,1675]" italics="true" pageId="30" pageNumber="31">Xystopyge</emphasis>
</taxonomicName>
-like way, and having a spinelike lobe on the subdistal ectal surface of the telomere.
</paragraph>
</subSubSection>
<subSubSection id="C342BEA4D63B0629FF4EF93FC000F8FF" box="[189,1324,1756,1831]" pageId="30" pageNumber="31" type="etymology">
<paragraph id="8BE7ED2FD63B0629FF4EF93FC468F92E" blockId="30.[189,324,1756,1782]" box="[189,324,1756,1782]" pageId="30" pageNumber="31">
<heading id="D0AF5A43D63B0629FF4EF93FC468F92E" bold="true" box="[189,324,1756,1782]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="B92C313DD63B0629FF4EF93FC468F92E" bold="true" box="[189,324,1756,1782]" pageId="30" pageNumber="31">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BE7ED2FD63B0629FF4EF8EEC000F8FF" blockId="30.[189,1324,1805,1831]" box="[189,1324,1805,1831]" pageId="30" pageNumber="31">
Named after the superficial gonopodal similarity with species of the genus
<taxonomicName id="4C5896ACD63B0629FBDFF8EDC189F8FF" authorityName="Attems" authorityYear="1909" box="[1068,1189,1806,1831]" class="Diplopoda" family="Odontopygidae" genus="Xystopyge" kingdom="Animalia" order="Spirostreptida" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B92C313DD63B0629FBDFF8EDC189F8FF" box="[1068,1189,1806,1831]" italics="true" pageId="30" pageNumber="31">Xystopyge</emphasis>
</taxonomicName>
. Adjective.
</paragraph>
</subSubSection>
<subSubSection id="C342BEA4D63B0629FF4EF8B2C1ABF830" pageId="30" pageNumber="31" type="materials_examined">
<paragraph id="8BE7ED2FD63B0629FF4EF8B2C700F8B4" blockId="30.[189,556,1873,1900]" box="[189,556,1873,1900]" pageId="30" pageNumber="31">
<emphasis id="B92C313DD63B0629FF4EF8B2C486F8B3" bold="true" box="[189,426,1873,1899]" pageId="30" pageNumber="31">Material examined</emphasis>
(total
<specimenCount id="9D5E26A6D63B0629FE04F8B2C708F8B4" box="[503,548,1873,1900]" pageId="30" pageNumber="31" type="male">1 ♂</specimenCount>
)
</paragraph>
<paragraph id="8BE7ED2FD63B0629FF19F861C1ABF830" blockId="30.[189,1399,1922,2025]" pageId="30" pageNumber="31">
<materialsCitation id="3B30E772D63B0629FF19F861C1ABF830" ID-GBIF-Occurrence="3712026358" collectedFrom="forest" collectingDate="1995-10-27" collectingDateMax="1995-11-09" collectingDateMin="1995-10-27" collectionCode="NHMD" collectorName="C. E. Griswold &amp; N. Scharff &amp; D. Ubick" country="Tanzania" elevation="950" latitude="-5.095" location="Amani" longLatPrecision="926" longitude="38.633335" municipality="East Usambara Mts" pageId="30" pageNumber="31" specimenCode="NHMD 621718" specimenCount="1" specimenCount-male="1" stateProvince="Tanga Region" typeStatus="holotype">
<emphasis id="B92C313DD63B0629FF19F861C470F844" bold="true" box="[234,348,1922,1948]" pageId="30" pageNumber="31">
<typeStatus id="54E3538DD63B0629FF19F861C470F844" box="[234,348,1922,1948]" pageId="30" pageNumber="31" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F34FADBFD63B0629FF4EF84AC47EF81B" box="[189,338,1961,1987]" name="Tanzania" pageId="30" pageNumber="31">TANZANIA</collectingCountry>
<specimenCount id="9D5E26A6D63B0629FE99F84AC4AFF81B" box="[362,387,1961,1987]" pageId="30" pageNumber="31" type="male"></specimenCount>
;
<collectingRegion id="499C23CDD63B0629FE7CF84AC71AF81A" box="[399,566,1960,1987]" country="Tanzania" name="Tanga" pageId="30" pageNumber="31">Tanga Region</collectingRegion>
,
<collectingMunicipality id="6B837755D63B0629FDB1F84AC607F81B" box="[578,811,1960,1987]" pageId="30" pageNumber="31">East Usambara Mts</collectingMunicipality>
,
<location id="8E87BBF4D63B0629FCC5F84BC6A6F81A" LSID="urn:lsid:plazi:treatment:03F15C39D63B0616FDC9FA6AC03EFD1D:8E87BBF4D63B0629FCC5F84BC6A6F81A" box="[822,906,1960,1986]" country="Tanzania" latitude="-5.095" longLatPrecision="926" longitude="38.633335" municipality="East Usambara Mts" name="Amani" pageId="30" pageNumber="31" stateProvince="Tanga Region">Amani</location>
;
<geoCoordinate id="EE6C8BE8D63B0629FC65F844C6D9F81A" box="[918,1013,1959,1987]" degrees="5" direction="south" minutes="5.7" orientation="latitude" pageId="30" pageNumber="31" precision="92" value="-5.095">5°5.7 S</geoCoordinate>
,
<geoCoordinate id="EE6C8BE8D63B0629FBF1F844C140F81B" box="[1026,1132,1959,1987]" degrees="38" direction="east" minutes="38" orientation="longitude" pageId="30" pageNumber="31" precision="925" value="38.633335">38°38 E</geoCoordinate>
;
<elevation id="00750A1CD63B0629FB8BF84BC02FF81A" box="[1144,1283,1960,1987]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="30" pageNumber="31" unit="m" value="950.0">
<quantity id="4CA040CAD63B0629FB8BF84BC1EFF81B" box="[1144,1219,1960,1987]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="30" pageNumber="31" unit="m" value="950.0">950 m</quantity>
a.s.l.
</elevation>
;
<date id="FFE6CBEFD63B0629FAE2F84BC47CF831" pageId="30" pageNumber="31" value="1995-10-27" valueMax="1995-11-09" valueMin="1995-10-27">
<collectingDate id="EFA23207D63B0629FAE2F84BC47CF831" pageId="30" pageNumber="31" value="1995-10-27" valueMax="1995-11-09" valueMin="1995-10-27">27 Oct. 9 Nov. 1995</collectingDate>
</date>
;
<collectorName id="26AD88F9D63B0629FEAFF82DC724F830" box="[348,520,1998,2024]" pageId="30" pageNumber="31">C.E. Griswold</collectorName>
,
<collectorName id="26AD88F9D63B0629FDE7F82CC7A2F830" box="[532,654,1998,2025]" pageId="30" pageNumber="31">N. Scharff</collectorName>
and
<collectorName id="26AD88F9D63B0629FD35F82CC61EF830" box="[710,818,1998,2025]" pageId="30" pageNumber="31">D. Ubick</collectorName>
leg.;
<collectedFrom id="3844AED2D63B0629FC87F82DC697F830" box="[884,955,1998,2024]" pageId="30" pageNumber="31">forest</collectedFrom>
;
<specimenCode id="DBFE4554D63B0629FC35F82CC1AFF830" box="[966,1155,1998,2025]" collectionCode="NHMD" pageId="30" pageNumber="31">NHMD 621718</specimenCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C342BEA4D63A0616FF4EFEEBC03EFD1D" lastPageId="33" lastPageNumber="34" pageId="31" pageNumber="32" type="description">
<paragraph id="8BE7ED2FD63A0628FF4EFEEBC463FEFA" blockId="31.[189,335,264,290]" box="[189,335,264,290]" pageId="31" pageNumber="32">
<heading id="D0AF5A43D63A0628FF4EFEEBC463FEFA" bold="true" box="[189,335,264,290]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
<emphasis id="B92C313DD63A0628FF4EFEEBC463FEFA" bold="true" box="[189,335,264,290]" pageId="31" pageNumber="32">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BE7ED2FD63A0628FF4EFED6C1D7FE97" blockId="31.[189,1275,309,336]" box="[189,1275,309,336]" pageId="31" pageNumber="32">
SIZE. Length
<quantity id="4CA040CAD63A0628FEA4FED6C481FE88" box="[343,429,309,336]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="31" pageNumber="32" unit="mm" value="45.0">45 mm</quantity>
. Diameter
<quantity id="4CA040CAD63A0628FDDFFED6C7A5FE88" box="[556,649,309,336]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="31" pageNumber="32" unit="mm" value="2.5">2.5 mm</quantity>
. 63 podous rings, no apodous rings in front of telson.
</paragraph>
<paragraph id="8BE7ED2FD63A0628FF4EFE96C685FE68" blockId="31.[189,1398,373,433]" pageId="31" pageNumber="32">COLOUR. After 26 years in alcohol faded to light grey with irregular black blotches; posterior parts of metazonites amber; with faint traces of a pale mid-dorsal stripe.</paragraph>
<caption id="DF27BDA7D63A0628FF4EF91AC708F83F" ID-DOI="http://doi.org/10.5281/zenodo.6359127" ID-Zenodo-Dep="6359127" httpUri="https://zenodo.org/record/6359127/files/figure.png" pageId="31" pageNumber="32" startId="31.[189,232,1785,1811]" targetBox="[189,1398,464,1754]" targetPageId="31">
<paragraph id="8BE7ED2FD63A0628FF4EF91AC708F83F" blockId="31.[185,1399,1785,2023]" pageId="31" pageNumber="32">
<emphasis id="B92C313DD63A0628FF4EF91AC437F8CB" bold="true" box="[189,283,1785,1811]" pageId="31" pageNumber="32">Fig. 19.</emphasis>
<taxonomicName id="4C5896ACD63A0628FED6F919C752F8CC" authority="Enghoff, 2022" authorityName="Enghoff" authorityYear="2022" box="[293,638,1786,1812]" class="Diplopoda" family="Odontopygidae" genus="Calyptomastix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="xystopygoides" status="sp. nov.">
<emphasis id="B92C313DD63A0628FED6F919C752F8CC" box="[293,638,1786,1812]" italics="true" pageId="31" pageNumber="32">Calyptomastix xystopygoides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21F8C46D63A0628FD75F919C7C8F8CC" box="[646,740,1786,1812]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (NHMD 621718).
<emphasis id="B92C313DD63A0628FB91F91AC1B7F8CB" bold="true" box="[1122,1179,1785,1811]" pageId="31" pageNumber="32">AC</emphasis>
. First pair of legs. A. Sublateral view. B. Anterior view. C. Ventral view.
<emphasis id="B92C313DD63A0628FCB5F8FEC676F8EF" bold="true" box="[838,858,1821,1847]" pageId="31" pageNumber="32">D</emphasis>
. Midbody dorsal limbus (worn).
<emphasis id="B92C313DD63A0628FB1BF8FEC1D6F8EF" bold="true" box="[1256,1274,1821,1847]" pageId="31" pageNumber="32">E</emphasis>
. Midbody ventrolateral limbus.
<emphasis id="B92C313DD63A0628FE48F8A3C4CAF882" bold="true" box="[443,486,1856,1882]" pageId="31" pageNumber="32">FI</emphasis>
. Left gonopod coxa. F. Anterior view. G. Anterior-lateral view. H. Posteriormesal view. I. lateral view. Abbreviations:
<emphasis id="B92C313DD63A0628FCF4F887C616F8A6" box="[775,826,1892,1918]" italics="true" pageId="31" pageNumber="32">APS</emphasis>
=mesapical prefemoral setae;
<emphasis id="B92C313DD63A0628FB3BF887C1DDF8A6" box="[1224,1265,1892,1918]" italics="true" pageId="31" pageNumber="32">CU</emphasis>
= cucullus;
<emphasis id="B92C313DD63A0628FF4AF864C5CCF878" box="[185,224,1927,1952]" italics="true" pageId="31" pageNumber="32">cus</emphasis>
= cucullar spine;
<emphasis id="B92C313DD63A0628FE4BF864C4C0F879" box="[440,492,1927,1953]" italics="true" pageId="31" pageNumber="32">CXS</emphasis>
=coxosternal setae;
<emphasis id="B92C313DD63A0628FD1AF865C62AF878" box="[745,774,1926,1952]" italics="true" pageId="31" pageNumber="32">hh</emphasis>
=hemisphaerical hump; LCS= lateral coxal spine;
<emphasis id="B92C313DD63A0628FF4EF849C5C2F81C" box="[189,238,1962,1988]" italics="true" pageId="31" pageNumber="32">LPS</emphasis>
=lateral prefemoral setae;
<emphasis id="B92C313DD63A0628FDD9F849C779F81B" box="[554,597,1962,1987]" italics="true" pageId="31" pageNumber="32">MP</emphasis>
=metaplica;
<emphasis id="B92C313DD63A0628FD19F849C627F81B" box="[746,779,1962,1987]" italics="true" pageId="31" pageNumber="32">ms</emphasis>
=metazonital striae;
<emphasis id="B92C313DD63A0628FBF0F849C10BF81B" box="[1027,1063,1962,1987]" italics="true" pageId="31" pageNumber="32">PP</emphasis>
=proplica, Scale bars: AC, FI =0.1 mm; DE=0.02 mm.
</paragraph>
</caption>
<paragraph id="8BE7ED2FD6050617FF4EFEEAC621FEFB" blockId="32.[189,781,265,291]" box="[189,781,265,291]" pageId="32" pageNumber="33">SUPRALABRAL SETAE. Not countable due to damage.</paragraph>
<paragraph id="8BE7ED2FD6050617FF4EFEA9C1A7FEBC" blockId="32.[189,1163,329,356]" box="[189,1163,329,356]" pageId="32" pageNumber="33">MANDIBULAR STIPES. With large disto-ventral lobe, distal margin shallowly concave.</paragraph>
<paragraph id="8BE7ED2FD6050617FF4EFE69C443FE1D" blockId="32.[188,1398,393,453]" pageId="32" pageNumber="33">ANAL VALVES. Each with a very small dorsal spine, an even smaller ventral one and three setae on slightly raised margins.</paragraph>
<paragraph id="8BE7ED2FD6050617FF4EFE0FC059FD92" blockId="32.[187,1399,492,586]" pageId="32" pageNumber="33">
LIMBUS (
<figureCitation id="1363F1AAD6050617FED4FE0FC49AFDDE" box="[295,438,492,518]" captionStart="Fig" captionStartId="31.[189,232,1785,1811]" captionTargetBox="[189,1398,464,1754]" captionTargetId="figure-170@31.[189,1398,464,1754]" captionTargetPageId="31" captionText="Fig. 19. Calyptomastix xystopygoides sp. nov., holotype, ♂ (NHMD 621718). AC. First pair of legs. A. Sublateral view. B. Anterior view. C. Ventral view. D. Midbody dorsal limbus (worn). E. Midbody ventrolateral limbus. FI. Left gonopod coxa. F. Anterior view. G. Anterior-lateral view. H. Posteriormesal view. I. lateral view. Abbreviations: APS =mesapical prefemoral setae; CU = cucullus; cus = cucullar spine; CXS =coxosternal setae; hh =hemisphaerical hump; LCS= lateral coxal spine; LPS =lateral prefemoral setae; MP=metaplica; ms =metazonital striae; PP =proplica, Scale bars: AC, FI =0.1 mm; DE=0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.6359127" httpUri="https://zenodo.org/record/6359127/files/figure.png" pageId="32" pageNumber="33">Fig. 19DE</figureCitation>
). Dorsal and lateral limbus worn, appearing as with rounded-triangular lobes. Ventrolateral limbus with very slender, spinelike lobes. (There may be considerable differences in limbus structure between the dorsal, lateral and ventrolateral parts of a body ring, see
<bibRefCitation id="EFC990DED6050617FBA0FDD3C1D7FD92" author="Schmidt D." box="[1107,1275,560,586]" pageId="32" pageNumber="33" pagination="65 - 80" refId="ref49396" refString="Schmidt D. 1962. Uber die taxionomische Wertigkeit von Strukturen des Metazonit-Hinterrandes bei Diplopoden. Senckenbergiana Biologica 43 (1): 65 - 80." type="journal article" year="1962">Schmidt 1962</bibRefCitation>
: figs 12.)
</paragraph>
<paragraph id="8BE7ED2FD6050617FF4EFD93C714FD74" blockId="32.[188,1398,624,684]" pageId="32" pageNumber="33">LEGS. With small postfemoral and tibial pads on anterior postgonopodal legs; pads decreasing in size and absent from midbody backward.</paragraph>
<paragraph id="8BE7ED2FD6050617FF4EFD30C14AFCEA" blockId="32.[189,1398,722,818]" pageId="32" pageNumber="33">
FIRST PAIR OF LEGS (
<figureCitation id="1363F1AAD6050617FE6DFD31C702FD34" box="[414,558,722,749]" captionStart="Fig" captionStartId="31.[189,232,1785,1811]" captionTargetBox="[189,1398,464,1754]" captionTargetId="figure-170@31.[189,1398,464,1754]" captionTargetPageId="31" captionText="Fig. 19. Calyptomastix xystopygoides sp. nov., holotype, ♂ (NHMD 621718). AC. First pair of legs. A. Sublateral view. B. Anterior view. C. Ventral view. D. Midbody dorsal limbus (worn). E. Midbody ventrolateral limbus. FI. Left gonopod coxa. F. Anterior view. G. Anterior-lateral view. H. Posteriormesal view. I. lateral view. Abbreviations: APS =mesapical prefemoral setae; CU = cucullus; cus = cucullar spine; CXS =coxosternal setae; hh =hemisphaerical hump; LCS= lateral coxal spine; LPS =lateral prefemoral setae; MP=metaplica; ms =metazonital striae; PP =proplica, Scale bars: AC, FI =0.1 mm; DE=0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.6359127" httpUri="https://zenodo.org/record/6359127/files/figure.png" pageId="32" pageNumber="33">Fig. 19AC</figureCitation>
). Prefemoral lobes rounded-triangular in ventral view. Four coxosternal setae (
<emphasis id="B92C313DD6050617FEF9FD16C412FCD7" box="[266,318,757,783]" italics="true" pageId="32" pageNumber="33">CXS</emphasis>
) close to lateral margin of coxosternum, well separated from prefemoral lobes. Prefemora with one or two short mesapical setae (
<emphasis id="B92C313DD6050617FD7BFCFBC797FCEA" box="[648,699,792,818]" italics="true" pageId="32" pageNumber="33">APS</emphasis>
) and four to five lateral setae (
<emphasis id="B92C313DD6050617FBD0FCFBC178FCEA" box="[1059,1108,792,818]" italics="true" pageId="32" pageNumber="33">LPS</emphasis>
).
</paragraph>
<caption id="DF27BDA7D6050617FF4EF8A2C7CFF830" ID-DOI="http://doi.org/10.5281/zenodo.6359131" ID-Zenodo-Dep="6359131" httpUri="https://zenodo.org/record/6359131/files/figure.png" pageId="32" pageNumber="33" startId="32.[189,232,1857,1883]" targetBox="[189,1398,854,1826]" targetPageId="32">
<paragraph id="8BE7ED2FD6050617FF4EF8A2C7CFF830" blockId="32.[185,1399,1857,2024]" pageId="32" pageNumber="33">
<emphasis id="B92C313DD6050617FF4EF8A2C437F883" bold="true" box="[189,283,1857,1883]" pageId="32" pageNumber="33">Fig. 20.</emphasis>
<taxonomicName id="4C5896ACD6050617FED0F8A2C750F883" authority="Enghoff, 2022" authorityName="Enghoff" authorityYear="2022" box="[291,636,1857,1883]" class="Diplopoda" family="Odontopygidae" genus="Calyptomastix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="xystopygoides" status="sp. nov.">
<emphasis id="B92C313DD6050617FED0F8A2C750F883" box="[291,636,1857,1883]" italics="true" pageId="32" pageNumber="33">Calyptomastix xystopygoides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21F8C46D6050617FD77F8A2C7CDF883" box="[644,737,1857,1883]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (NHMD 621718), left gonopod telopodite.
<emphasis id="B92C313DD6050617FF4EF887C5FEF8A6" bold="true" box="[189,210,1892,1918]" pageId="32" pageNumber="33">A</emphasis>
. anterior-mesal view.
<emphasis id="B92C313DD6050617FE15F887C4D6F8A6" bold="true" box="[486,506,1892,1918]" pageId="32" pageNumber="33">B</emphasis>
. Posterior view.
<emphasis id="B92C313DD6050617FD3DF887C7CFF8A6" bold="true" box="[718,739,1892,1918]" pageId="32" pageNumber="33">C</emphasis>
. Ventral view.
<emphasis id="B92C313DD6050617FC51F887C69BF8A6" bold="true" box="[930,951,1892,1918]" pageId="32" pageNumber="33">D</emphasis>
. Dorsal view.
<emphasis id="B92C313DD6050617FB83F887C1A8F8A6" bold="true" box="[1136,1156,1892,1918]" pageId="32" pageNumber="33">E</emphasis>
. Detail of telomere and ridged solenomere. Abbreviations:
<emphasis id="B92C313DD6050617FD5BF86BC7F4F87A" box="[680,728,1928,1954]" italics="true" pageId="32" pageNumber="33">BSS</emphasis>
=basal solenomeral spine;
<emphasis id="B92C313DD6050617FBD9F86BC17CF879" box="[1066,1104,1928,1953]" italics="true" pageId="32" pageNumber="33">PN</emphasis>
=post-torsal narrowing;
<emphasis id="B92C313DD6050617FF4AF848C5DDF81D" box="[185,241,1963,1989]" italics="true" pageId="32" pageNumber="33">SLM</emphasis>
=solenomere;
<emphasis id="B92C313DD6050617FE53F848C4EDF81C" box="[416,449,1963,1988]" italics="true" pageId="32" pageNumber="33">TL</emphasis>
=torsotope lobe;
<emphasis id="B92C313DD6050617FD63F849C79AF81C" box="[656,694,1962,1988]" italics="true" pageId="32" pageNumber="33">tml</emphasis>
=telomeral spinelike lobe;
<emphasis id="B92C313DD6050617FC0FF848C131F81C" box="[1020,1053,1963,1988]" italics="true" pageId="32" pageNumber="33">TT</emphasis>
=torsotope;
<emphasis id="B92C313DD6050617FB5DF848C1F3F81D" box="[1198,1247,1963,1989]" italics="true" pageId="32" pageNumber="33">PTS</emphasis>
= post-torsal spine. Scale bars: AD =0.2 mm; E =0.05 mm.
</paragraph>
</caption>
<paragraph id="8BE7ED2FD6040616FF4EFEEAC77BFEFB" blockId="33.[189,599,265,291]" box="[189,599,265,291]" pageId="33" pageNumber="34">STERNUM 9. Lost during dissection.</paragraph>
<paragraph id="8BE7ED2FD6040616FF4EFEABC066FE1C" blockId="33.[188,1399,328,452]" pageId="33" pageNumber="34">
GONOPOD COXA (
<figureCitation id="1363F1AAD6040616FE88FEABC4D0FEBA" box="[379,508,328,354]" captionStart="Fig" captionStartId="31.[189,232,1785,1811]" captionTargetBox="[189,1398,464,1754]" captionTargetId="figure-170@31.[189,1398,464,1754]" captionTargetPageId="31" captionText="Fig. 19. Calyptomastix xystopygoides sp. nov., holotype, ♂ (NHMD 621718). AC. First pair of legs. A. Sublateral view. B. Anterior view. C. Ventral view. D. Midbody dorsal limbus (worn). E. Midbody ventrolateral limbus. FI. Left gonopod coxa. F. Anterior view. G. Anterior-lateral view. H. Posteriormesal view. I. lateral view. Abbreviations: APS =mesapical prefemoral setae; CU = cucullus; cus = cucullar spine; CXS =coxosternal setae; hh =hemisphaerical hump; LCS= lateral coxal spine; LPS =lateral prefemoral setae; MP=metaplica; ms =metazonital striae; PP =proplica, Scale bars: AC, FI =0.1 mm; DE=0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.6359127" httpUri="https://zenodo.org/record/6359127/files/figure.png" pageId="33" pageNumber="34">Fig. 19FI</figureCitation>
). Rather compact, with a large lateral spine (
<emphasis id="B92C313DD6040616FBF7FEABC11BFEBA" box="[1028,1079,328,354]" italics="true" pageId="33" pageNumber="34">LCS</emphasis>
) directed obliquely apicad. Distal part of proplica (
<emphasis id="B92C313DD6040616FE21FE8AC4DAFE5A" box="[466,502,361,386]" italics="true" pageId="33" pageNumber="34">PP</emphasis>
) overlaid by rounded-rectangular anterior lobe of metaplica (
<emphasis id="B92C313DD6040616FB38FE8AC1DAFE5A" box="[1227,1270,361,386]" italics="true" pageId="33" pageNumber="34">MP</emphasis>
), proplical lobe hence not visible. Cucullus (
<emphasis id="B92C313DD6040616FDBFFE6AC759FE7B" box="[588,629,393,419]" italics="true" pageId="33" pageNumber="34">CU</emphasis>
) very small, carrying a long, slender, overall straight but basally curved spine (
<emphasis id="B92C313DD6040616FE90FE49C4A6FE1B" box="[355,394,426,451]" italics="true" pageId="33" pageNumber="34">cus</emphasis>
) directed obliquely posteriad. Posterior surface with a hemisphaerical hump (
<emphasis id="B92C313DD6040616FAEFFE4AC015FE1B" box="[1308,1337,425,451]" italics="true" pageId="33" pageNumber="34">hh</emphasis>
).
</paragraph>
<paragraph id="8BE7ED2FD6040616FF4EFE0BC03EFD1D" blockId="33.[188,1400,488,710]" pageId="33" pageNumber="34">
GONOPOD TELOPODITE (
<figureCitation id="1363F1AAD6040616FE32FE0BC735FDDA" box="[449,537,488,515]" captionStart="Fig" captionStartId="32.[189,232,1857,1883]" captionTargetBox="[189,1398,854,1826]" captionTargetId="figure-278@32.[189,1398,854,1826]" captionTargetPageId="32" captionText="Fig. 20. Calyptomastix xystopygoides sp. nov., holotype, ♂ (NHMD 621718), left gonopod telopodite. A. anterior-mesal view. B. Posterior view. C. Ventral view. D. Dorsal view. E. Detail of telomere and ridged solenomere. Abbreviations: BSS =basal solenomeral spine; PN =post-torsal narrowing; SLM =solenomere; TL =torsotope lobe; tml =telomeral spinelike lobe; TT=torsotope; PTS = post-torsal spine. Scale bars: AD =0.2 mm; E =0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.6359131" httpUri="https://zenodo.org/record/6359131/files/figure.png" pageId="33" pageNumber="34">Fig. 20</figureCitation>
). Arculus 90°. Torsotope (
<emphasis id="B92C313DD6040616FCBCFE0AC65CFDDA" box="[847,880,489,514]" italics="true" pageId="33" pageNumber="34">
<collectingRegion id="499C23CDD6040616FCBCFE0AC65CFDDA" box="[847,880,489,514]" country="Mozambique" name="Tete" pageId="33" pageNumber="34">TT</collectingRegion>
</emphasis>
) extended, with semicircular torsotope lobe (
<emphasis id="B92C313DD6040616FF34FDEAC5C4FDFA" box="[199,232,521,546]" italics="true" pageId="33" pageNumber="34">TL</emphasis>
). Post-torsal narrowing (
<emphasis id="B92C313DD6040616FDF9FDEAC71CFDFA" box="[522,560,521,546]" italics="true" pageId="33" pageNumber="34">PN</emphasis>
) moderately pronounced. A very long, straight post-torsal spine directed laterad, i.e., towards arculus. Solenomere (SLM) with a stout, basal spine (
<emphasis id="B92C313DD6040616FBC2FDC9C148FD9C" box="[1073,1124,554,580]" italics="true" pageId="33" pageNumber="34">BBS</emphasis>
); apart from basalmost part slender, whiplike, almost completely hidden inside telomere, only a small, obliquely ridged part of the distal half visible (
<figureCitation id="1363F1AAD6040616FE38FD89C714FD5C" box="[459,568,618,644]" captionStart="Fig" captionStartId="32.[189,232,1857,1883]" captionTargetBox="[189,1398,854,1826]" captionTargetId="figure-278@32.[189,1398,854,1826]" captionTargetPageId="32" captionText="Fig. 20. Calyptomastix xystopygoides sp. nov., holotype, ♂ (NHMD 621718), left gonopod telopodite. A. anterior-mesal view. B. Posterior view. C. Ventral view. D. Dorsal view. E. Detail of telomere and ridged solenomere. Abbreviations: BSS =basal solenomeral spine; PN =post-torsal narrowing; SLM =solenomere; TL =torsotope lobe; tml =telomeral spinelike lobe; TT=torsotope; PTS = post-torsal spine. Scale bars: AD =0.2 mm; E =0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.6359131" httpUri="https://zenodo.org/record/6359131/files/figure.png" pageId="33" pageNumber="34">Fig. 20E</figureCitation>
), tip (not visible on
<figureCitation id="1363F1AAD6040616FCDDFD89C6A9FD5C" box="[814,901,618,644]" captionStart="Fig" captionStartId="32.[189,232,1857,1883]" captionTargetBox="[189,1398,854,1826]" captionTargetId="figure-278@32.[189,1398,854,1826]" captionTargetPageId="32" captionText="Fig. 20. Calyptomastix xystopygoides sp. nov., holotype, ♂ (NHMD 621718), left gonopod telopodite. A. anterior-mesal view. B. Posterior view. C. Ventral view. D. Dorsal view. E. Detail of telomere and ridged solenomere. Abbreviations: BSS =basal solenomeral spine; PN =post-torsal narrowing; SLM =solenomere; TL =torsotope lobe; tml =telomeral spinelike lobe; TT=torsotope; PTS = post-torsal spine. Scale bars: AD =0.2 mm; E =0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.6359131" httpUri="https://zenodo.org/record/6359131/files/figure.png" pageId="33" pageNumber="34">Fig. 20</figureCitation>
) simple, pointed. Telomere (
<emphasis id="B92C313DD6040616FB2CFD88C024FD5C" box="[1247,1288,619,644]" italics="true" pageId="33" pageNumber="34">TM</emphasis>
) slender, tightly rolled around solenomere and in addition forming a regular, almost 180° curve; distally expanded into sub-rectangular sheet, subdistall with a spinelike lobe (
<emphasis id="B92C313DD6040616FC84FD48C6B1FD1D" box="[887,925,683,709]" italics="true" pageId="33" pageNumber="34">tml</emphasis>
) originating from ectal surface.
</paragraph>
</subSubSection>
</treatment>
</document>