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<document id="D3ECA2943E9179AF9F47E45E1DDA2E70" ID="10.11646/zootaxa.4147.1.4" ID-DOI="10.11646/zootaxa.4147.1.4" ID-GBIF-Dataset="3166e193-d052-4efa-953b-4e5ac1b6f568" ID-ISSN="1175-5326" ID-Zenodo-Dep="266251" ID-ZooBank="39ECCC38-BC63-40D2-9B04-F435841AA0D9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1470120521773" checkinUser="plazi" docAuthor="Daniel, Gimo M., Davis, Adrian Lv. &amp; Scholtz, Clarke H." docDate="2016" docId="03E087A1E556FFCA88BE997964B47855" docLanguage="en" docName="zootaxa.4147.1.4.pdf" docOrigin="Zootaxa 4147 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sisyphus umbraphilus Daniel &amp; Davis, sp. nov." docType="treatment" docVersion="7" lastPageNumber="69" masterDocId="FFD9FFD9E557FFC888299C0B66117C33" masterDocTitle="Three new Sisyphus species (Coleoptera: Scarabaeidae: Scarabaeinae) from shaded vegetation in southern Africa" masterLastPageNumber="74" masterPageNumber="67" pageNumber="68" updateTime="1698366689639" updateUser="plazi">
<mods:mods id="ACC031AC57E8261F7B831170F54C51D6" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0EBE49C90158FCA5EED64BC08637A08A">
<mods:title id="35E3E0D875D317E270F80F25CE816033">Three new Sisyphus species (Coleoptera: Scarabaeidae: Scarabaeinae) from shaded vegetation in southern Africa</mods:title>
</mods:titleInfo>
<mods:name id="44A1A1336D5B5C9975A1AE47311A0334" type="personal">
<mods:role id="852F49577EE1C0C85A998A44C988E200">
<mods:roleTerm id="30C020E0431EA101F570BD3C6AC85A4C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="988DC82A35DC13C5228D225755A23DF3">Daniel, Gimo M.</mods:namePart>
</mods:name>
<mods:name id="5200245ABA9BA0EC906DA6284DEA15CB" type="personal">
<mods:role id="3465EDB6B872C14EC31E926979455EAF">
<mods:roleTerm id="E2FB74C4D2022E820E07B5E7671E7889">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="27CEA5DBB413D0FD59502FAF9DEA10C5">Davis, Adrian Lv.</mods:namePart>
</mods:name>
<mods:name id="C25B2676D117CAFB1790C6CC8CFDF41F" type="personal">
<mods:role id="62A79E82811D44EE3B4CFBF0F351189A">
<mods:roleTerm id="EDE5F448F49BB63DE27AAE0081BF7EE0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="905AD115C5B6E5CC29CC7F0977248270">Scholtz, Clarke H.</mods:namePart>
</mods:name>
<mods:typeOfResource id="448431891D97CFAD300342C043C709A4">text</mods:typeOfResource>
<mods:relatedItem id="229061C9F34C6BCF2B2641639258D72C" type="host">
<mods:titleInfo id="5565B96ED0F077C6D53C6DD7079AC679">
<mods:title id="D408882F77768557288B54469993F9FE">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="CF73A1321D418251B51377B82EF9DE70">
<mods:date id="995FBAB34DAF560797193EB895221302">2016</mods:date>
<mods:detail id="6FC0FC1F031456CE257D12DB4114CB7D" type="volume">
<mods:number id="122FF588D89CA5068DE2BD93CFB55083">4147</mods:number>
</mods:detail>
<mods:detail id="AEBA783A7CF207C52A22D643202D2F9A" type="issue">
<mods:number id="44AA9ED3C464B8E7C3B489BCB5CFFE82">1</mods:number>
</mods:detail>
<mods:extent id="41EE52B34C8E4E935429223A0EDB81DA" unit="page">
<mods:start id="73930031573CDFA6DCA2DEC42D422F62">67</mods:start>
<mods:end id="AB9DF644A7A89A067C40F5476E6F96DD">74</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="DD53EECBC84769BD344D17A0E9856FAC">journal article</mods:classification>
<mods:identifier id="4E5EDC5A617ABF3308E8A97C82ED7D32" type="DOI">10.11646/zootaxa.4147.1.4</mods:identifier>
<mods:identifier id="BC35EC4835068E52007BFC4082DB2953" type="GBIF-Dataset">3166e193-d052-4efa-953b-4e5ac1b6f568</mods:identifier>
<mods:identifier id="7ADD4BC8F093B034B2137D47EFBA8D55" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="49654E6B89BFCF92D5EA6A857523CEB2" type="Zenodo-Dep">266251</mods:identifier>
<mods:identifier id="94AEE352E30D6970B746F28CB83E91C7" type="ZooBank">39ECCC38-BC63-40D2-9B04-F435841AA0D9</mods:identifier>
</mods:mods>
<treatment id="03E087A1E556FFCA88BE997964B47855" ID-DOI="http://doi.org/10.5281/zenodo.6087199" ID-GBIF-Taxon="123513970" ID-Zenodo-Dep="6087199" LSID="urn:lsid:plazi:treatment:03E087A1E556FFCA88BE997964B47855" httpUri="http://treatment.plazi.org/id/03E087A1E556FFCA88BE997964B47855" lastPageId="2" lastPageNumber="69" pageId="1" pageNumber="68">
<subSubSection id="C353653CE556FFC988BE997966E9799D" pageId="1" pageNumber="68" type="nomenclature">
<paragraph id="8BF636B7E556FFC988BE997964DE79BE" blockId="1.[151,719,1394,1454]" box="[151,719,1394,1421]" pageId="1" pageNumber="68">
<heading id="D0BE81DBE556FFC988BE997964DE79BE" bold="true" box="[151,719,1394,1421]" fontSize="11" level="1" pageId="1" pageNumber="68" reason="1">
<emphasis id="B93DEAA5E556FFC988BE997964DE79BE" bold="true" box="[151,719,1394,1421]" pageId="1" pageNumber="68">
<taxonomicName id="4C494D34E556FFC988BE9979647B79BE" ID-CoL="7ZRHK" authority="Daniel &amp; Davis" authorityName="Daniel &amp; Davis" box="[151,618,1394,1421]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="68" phylum="Arthropoda" rank="species" species="umbraphilus" status="sp. nov.">
<emphasis id="B93DEAA5E556FFC988BE997967B479BF" bold="true" box="[151,421,1394,1420]" italics="true" pageId="1" pageNumber="68">Sisyphus umbraphilus</emphasis>
Daniel &amp; Davis
</taxonomicName>
<taxonomicNameLabel id="A20E57DEE556FFC98A58997864DE79BE" box="[625,719,1395,1421]" pageId="1" pageNumber="68" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF636B7E556FFC988BE999E66E9799D" blockId="1.[151,719,1394,1454]" box="[151,248,1429,1454]" pageId="1" pageNumber="68">
(
<figureCitation id="13722A32E556FFC988B6999E66FC799D" box="[159,237,1429,1454]" captionStart="FIGURE 1. A. S" captionStartId="2.[151,250,1918,1940]" captionTargetBox="[345,1244,1186,1888]" captionTargetId="figure@2.[332,1255,1181,1893]" captionTargetPageId="2" captionText="FIGURE 1. A. S. umbraphilus sp. nov. habitus. Diagnostic characters: Bi. Lateral genal margins almost parallel. Bii. Setae absent from vertical medial line on pygidium. Biii. T ufts of setae on epipleurae. C. Aedeagus." httpUri="https://zenodo.org/record/266252/files/figure.png" pageId="1" pageNumber="68">Fig. 1a</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C353653CE556FFC988BE99D6673A7A51" pageId="1" pageNumber="68" type="diagnosis">
<paragraph id="8BF636B7E556FFC988BE99D6673A7A51" blockId="1.[151,1437,1501,2030]" pageId="1" pageNumber="68">
<emphasis id="B93DEAA5E556FFC988BE99D6670079C5" bold="true" box="[151,273,1501,1526]" pageId="1" pageNumber="68">Diagnosis.</emphasis>
This new species is similar to
<taxonomicName id="4C494D34E556FFC98AAA99D5651B79C6" box="[643,778,1501,1525]" class="Insecta" family="Scarabaeidae" genus="Sisyphus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="68" phylum="Arthropoda" rank="species" species="oralensis" status="sp. nov.">
<emphasis id="B93DEAA5E556FFC98AAA99D5651B79C6" box="[643,778,1501,1525]" italics="true" pageId="1" pageNumber="68">S. oralensis</emphasis>
</taxonomicName>
<emphasis id="B93DEAA5E556FFC98B3D99D6656079C5" bold="true" box="[788,881,1501,1526]" pageId="1" pageNumber="68">
<taxonomicNameLabel id="A20E57DEE556FFC98B3D99D6656079C5" box="[788,881,1501,1526]" pageId="1" pageNumber="68" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
but can be distinguished from the latter by the following characters: lateral edges of genae virtually parallel; second and third abdominal epipleurae with welldeveloped tufts of setae; pygidium with uniformly scattered setae, except for a distinct non-setose line medially (
<emphasis id="B93DEAA5E556FFC988899A42670A7A51" bold="true" box="[160,283,1609,1634]" pageId="1" pageNumber="68">
<figureCitation id="13722A32E556FFC988899A4266F17A51" box="[160,224,1609,1634]" captionStart="FIGURE 1. A. S" captionStartId="2.[151,250,1918,1940]" captionTargetBox="[345,1244,1186,1888]" captionTargetId="figure@2.[332,1255,1181,1893]" captionTargetPageId="2" captionText="FIGURE 1. A. S. umbraphilus sp. nov. habitus. Diagnostic characters: Bi. Lateral genal margins almost parallel. Bii. Setae absent from vertical medial line on pygidium. Biii. T ufts of setae on epipleurae. C. Aedeagus." httpUri="https://zenodo.org/record/266252/files/figure.png" pageId="1" pageNumber="68">Fig. 1</figureCitation>
bi-iii
</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="C353653CE556FFCA88EE9A6665387E9B" lastPageId="2" lastPageNumber="69" pageId="1" pageNumber="68" type="description">
<paragraph id="8BF636B7E556FFC988EE9A6666E57A9A" blockId="1.[151,1437,1501,2030]" pageId="1" pageNumber="68">
<emphasis id="B93DEAA5E556FFC988EE9A6667407AB5" bold="true" box="[199,337,1645,1670]" pageId="1" pageNumber="68">Description</emphasis>
.
<emphasis id="B93DEAA5E556FFC989499A6667CF7AB5" bold="true" box="[352,478,1645,1670]" pageId="1" pageNumber="68">Size: Male</emphasis>
: Length:
<quantity id="4CB19B52E556FFC98A649A6664C97AB6" box="[589,728,1645,1670]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" metricValueMax="7.1" metricValueMin="5.1" pageId="1" pageNumber="68" unit="mm" value="6.1" valueMax="7.1" valueMin="5.1">5.1–7.1 mm</quantity>
, width:
<quantity id="4CB19B52E556FFC98B1D9A6565AE7AB6" box="[820,959,1646,1670]" metricMagnitude="-3" metricUnit="m" metricValue="3.55" metricValueMax="4.2" metricValueMin="2.9" pageId="1" pageNumber="68" unit="mm" value="3.55" valueMax="4.2" valueMin="2.9">2.9–4.2 mm</quantity>
; Female: Length:
<quantity id="4CB19B52E556FFC98CA49A6663067AB6" box="[1165,1303,1645,1670]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" metricValueMax="7.4" metricValueMin="5.4" pageId="1" pageNumber="68" unit="mm" value="6.4" valueMax="7.4" valueMin="5.4">5.4–7.4 mm</quantity>
, width
<quantity id="4CB19B52E556FFC98D429A6566E17A9A" metricMagnitude="-3" metricUnit="m" metricValue="3.7" metricValueMax="4.2" metricValueMin="3.2" pageId="1" pageNumber="68" unit="mm" value="3.7" valueMax="4.2" valueMin="3.2">3.2– 4.2 mm</quantity>
.
</paragraph>
<paragraph id="8BF636B7E556FFC988BE9ABE67FB7AFD" blockId="1.[151,1437,1501,2030]" box="[151,490,1717,1742]" pageId="1" pageNumber="68">
<emphasis id="B93DEAA5E556FFC988BE9ABE66E57AFD" bold="true" box="[151,244,1717,1742]" pageId="1" pageNumber="68">Colour:</emphasis>
Dark brown to black.
</paragraph>
<paragraph id="8BF636B7E556FFC988EE9AD267417B09" blockId="1.[151,1437,1501,2030]" pageId="1" pageNumber="68">
<emphasis id="B93DEAA5E556FFC988EE9AD267017AC1" bold="true" box="[199,272,1753,1778]" pageId="1" pageNumber="68">Head:</emphasis>
Clypeus dark and punctate; anterior margin with two medial teeth and a convex lateral sinuosity between teeth and genal sutures; lateral margins of genae almost parallel, post-occipital suture not fully rounded; antennae brown.
</paragraph>
<paragraph id="8BF636B7E556FFC988EE9B4E672C7BF9" blockId="1.[151,1437,1501,2030]" pageId="1" pageNumber="68">
<emphasis id="B93DEAA5E556FFC988EE9B4E67587B6D" bold="true" box="[199,329,1861,1886]" pageId="1" pageNumber="68">Pronotum:</emphasis>
Convex, shiny black, densely punctate, lateral edges slightly compressed, maximum length shorter than maximum width, complete lateral prothoracic ridge between prothoracic disc and prothoracic episternum, disc showing an indistinct pattern of bare patches, some with sparse short setae, arranged within a matrix of dense, long setae (
<figureCitation id="13722A32E556FFC988F79BBA673F7BF9" box="[222,302,1969,1994]" captionStart="FIGURE 1. A. S" captionStartId="2.[151,250,1918,1940]" captionTargetBox="[345,1244,1186,1888]" captionTargetId="figure@2.[332,1255,1181,1893]" captionTargetPageId="2" captionText="FIGURE 1. A. S. umbraphilus sp. nov. habitus. Diagnostic characters: Bi. Lateral genal margins almost parallel. Bii. Setae absent from vertical medial line on pygidium. Biii. T ufts of setae on epipleurae. C. Aedeagus." httpUri="https://zenodo.org/record/266252/files/figure.png" pageId="1" pageNumber="68">Fig. 1a</figureCitation>
).
</paragraph>
<paragraph id="8BF636B7E556FFCA88EE9BDE673B7CE7" blockId="1.[151,1437,1501,2030]" lastBlockId="2.[151,1437,151,1126]" lastPageId="2" lastPageNumber="69" pageId="1" pageNumber="68">
<emphasis id="B93DEAA5E556FFC988EE9BDE670D7BDD" bold="true" box="[199,284,2005,2030]" pageId="1" pageNumber="68">Elytra:</emphasis>
Dark brown, shiny in some individuals, ovoid posteriorly, interstriae finely punctate bearing lightcoloured setae with recurved tips, epipleurae of the second and third abdominal segments each bearing a tuft of yellow setae.
</paragraph>
<paragraph id="8BF636B7E555FFCA88EE9CD463937CC4" blockId="2.[151,1437,151,1126]" box="[199,1410,223,248]" pageId="2" pageNumber="69">
<emphasis id="B93DEAA5E555FFCA88EE9CD467527CCB" bold="true" box="[199,323,223,248]" pageId="2" pageNumber="69">Pygidium:</emphasis>
Narrow with ocellate punctation and scattered setae but with a distinct medial line lacking setae.
</paragraph>
<paragraph id="8BF636B7E555FFCA88EE9D0F64377D57" blockId="2.[151,1437,151,1126]" pageId="2" pageNumber="69">
<emphasis id="B93DEAA5E555FFCA88EE9D0F67287D2E" bold="true" box="[199,313,260,285]" pageId="2" pageNumber="69">Sternites:</emphasis>
Setae arranged in rows laterally, except on first sternite where setae form a ventral row; visible meso-metasternal suture; meso- and metasternum black shiny, punctate with lateral setae; mesepimerum and metepisternum punctate and setose.
</paragraph>
<paragraph id="8BF636B7E555FFCA88EE9D64636A7E0E" blockId="2.[151,1437,151,1126]" pageId="2" pageNumber="69">
<emphasis id="B93DEAA5E555FFCA88EE9D6467197DBB" bold="true" box="[199,264,367,392]" pageId="2" pageNumber="69">Legs:</emphasis>
Profemur black, punctate, internally pubescent with a lateral row of well-developed setae in ventral view; meso- and metafemur densely punctate and setose; meso- and metatrochanter contiguous with femur, sharp tip of metatrochanter projecting away from femur; metacoxa punctate and shagreened ventro-anteriorly; protibia with three teeth and single terminal spine; mesotibia punctate and setose with two terminal spines; metatibia densely setose, serrated laterally with two spurs; pro- meso- and metatarsus five segmented with two claws, setose laterally; first tarsal segment of the meso- and meta-thoracic legs with a row of strong setae on the ventral edge.
</paragraph>
<paragraph id="8BF636B7E555FFCA88EE9E4C65387E9B" blockId="2.[151,1437,151,1126]" pageId="2" pageNumber="69">
<emphasis id="B93DEAA5E555FFCA88EE9E4C67557E53" bold="true" box="[199,324,583,608]" pageId="2" pageNumber="69">Aedeagus:</emphasis>
Phallobase almost straight, slight angular depression immediately distal from the phallobaseparamere suture; parameres symetrical, sligthly curved inward at the tip, separated from each other by fine membranous area visible dorsally on apical edge (
<emphasis id="B93DEAA5E555FFCA8AE09E8465087E9B" bold="true" box="[713,793,655,680]" pageId="2" pageNumber="69">
<figureCitation id="13722A32E555FFCA8AE09E84651A7E9B" box="[713,779,655,680]" captionStart="FIGURE 1. A. S" captionStartId="2.[151,250,1918,1940]" captionTargetBox="[345,1244,1186,1888]" captionTargetId="figure@2.[332,1255,1181,1893]" captionTargetPageId="2" captionText="FIGURE 1. A. S. umbraphilus sp. nov. habitus. Diagnostic characters: Bi. Lateral genal margins almost parallel. Bii. Setae absent from vertical medial line on pygidium. Biii. T ufts of setae on epipleurae. C. Aedeagus." httpUri="https://zenodo.org/record/266252/files/figure.png" pageId="2" pageNumber="69">Fig. 1</figureCitation>
c
</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="C353653CE555FFCA88EE9EBF631E7EFE" box="[199,1295,692,717]" pageId="2" pageNumber="69" type="etymology">
<paragraph id="8BF636B7E555FFCA88EE9EBF631E7EFE" blockId="2.[151,1437,151,1126]" box="[199,1295,692,717]" pageId="2" pageNumber="69">
<emphasis id="B93DEAA5E555FFCA88EE9EBF675F7EFE" bold="true" box="[199,334,692,717]" pageId="2" pageNumber="69">Etymology.</emphasis>
The species name reflects the shaded vegetation with which the species is associated.
</paragraph>
</subSubSection>
<subSubSection id="C353653CE555FFCA88EE9ED164B47855" pageId="2" pageNumber="69" type="materials_examined">
<paragraph id="8BF636B7E555FFCA88EE9ED167137FC9" blockId="2.[151,1437,151,1126]" pageId="2" pageNumber="69">
<materialsCitation id="3B213CEAE555FFCA88EE9ED164957F24" ID-GBIF-Occurrence="1290984953" collectingDate="1970-10-01" collectingDateMax="1970-10-07" collectingDateMin="1970-10-01" collectionCode="SANC" collectorName="Bornemissza &amp; Aschenborn" country="South Africa" location="Examined" pageId="2" pageNumber="69" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B93DEAA5E555FFCA88EE9ED167A27EC0" bold="true" box="[199,435,730,755]" pageId="2" pageNumber="69">
<location id="8E96606CE555FFCA88EE9ED167517EC0" LSID="urn:lsid:plazi:treatment:03E087A1E556FFCA88BE997964B47855:8E96606CE555FFCA88EE9ED167517EC0" box="[199,320,730,755]" country="South Africa" name="Examined" pageId="2" pageNumber="69">Examined</location>
material.
</emphasis>
<typeStatus id="54F28815E555FFCA89939ED164377EC0" box="[442,550,730,755]" pageId="2" pageNumber="69" type="holotype">Holotype</typeStatus>
: (
<specimenCount id="9D4FFD3EE555FFCA8A109ED0645E7EC0" box="[569,591,731,755]" pageId="2" pageNumber="69" type="male">♂</specimenCount>
<emphasis id="B93DEAA5E555FFCA8A7C9ED164B07EC0" bold="true" box="[597,673,730,755]" pageId="2" pageNumber="69">
<collectionCode id="ED58AE72E555FFCA8A7C9ED164B07EC0" box="[597,673,730,755]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="2" pageNumber="69">SANC</collectionCode>
</emphasis>
):
<emphasis id="B93DEAA5E555FFCA8A909ED1659E7EC0" bold="true" box="[697,911,730,755]" pageId="2" pageNumber="69">
<collectingCountry id="F35E7627E555FFCA8A909ED1659E7EC0" box="[697,911,730,755]" name="South Africa" pageId="2" pageNumber="69">SOUTH AFRICA</collectingCountry>
</emphasis>
, KZN, Umfolozi 28[◦] 15[’] S; 31[◦] 55[’] E,
<date id="FFF71077E555FFCA8DAD9ED067157F24" pageId="2" pageNumber="69" value="1970-10-01" valueMax="1970-10-07" valueMin="1970-10-01">
<collectingDate id="EFB3E99FE555FFCA8DAD9ED067157F24" pageId="2" pageNumber="69" value="1970-10-01" valueMax="1970-10-07" valueMin="1970-10-01">1- 7.x. 1970</collectingDate>
</date>
, leg:
<collectorName id="26BC5361E555FFCA896B9EF567C27F24" box="[322,467,766,791]" pageId="2" pageNumber="69">Bornemissza</collectorName>
&amp;
<collectorName id="26BC5361E555FFCA89D09EF564957F24" box="[505,644,766,791]" pageId="2" pageNumber="69">Aschenborn</collectorName>
</materialsCitation>
.
<materialsCitation id="3B213CEAE555FFCA8AA69F0B63227F24" ID-GBIF-Occurrence="1290984958" box="[655,1331,766,792]" collectingDate="1970-10-01" collectingDateMax="1970-10-07" collectingDateMin="1970-10-01" collectionCode="SANC" collectorName="Bornemissza &amp; Aschenborn" country="South Africa" location="Examined" pageId="2" pageNumber="69" specimenCount="12" specimenCount-female="9" specimenCount-male="3" typeStatus="paratype">
<typeStatus id="54F28815E555FFCA8AA69F0B65117F2B" box="[655,768,768,792]" pageId="2" pageNumber="69" type="paratype">Paratypes</typeStatus>
: (
<specimenCount id="9D4FFD3EE555FFCA8B3C9EF4652A7F24" box="[789,827,767,791]" pageId="2" pageNumber="69" type="male">3♂</specimenCount>
,
<specimenCount id="9D4FFD3EE555FFCA8B6E9EF4657A7F24" box="[839,875,767,791]" pageId="2" pageNumber="69" type="female">9♀</specimenCount>
<emphasis id="B93DEAA5E555FFCA8B5A9EF565D97F24" bold="true" box="[883,968,766,791]" pageId="2" pageNumber="69">
<collectionCode id="ED58AE72E555FFCA8B5A9EF565D07F24" box="[883,961,766,791]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="2" pageNumber="69">SANC</collectionCode>
)
</emphasis>
with the same data as holotype
</materialsCitation>
. (1♂, 2♀
<emphasis id="B93DEAA5E555FFCA88BE9F2E66F17F0D" bold="true" box="[151,224,805,830]" pageId="2" pageNumber="69">UPSA</emphasis>
):
<materialsCitation id="3B213CEAE555FFCA88D09F2E64627F56" ID-GBIF-Occurrence="1290984957" collectingDate="1999-01-13" collectionCode="SANC" collectorName="Chown &amp; McGeogh &amp; Davis." country="South Africa" location="Itala Game Reserve" pageId="2" pageNumber="69" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<collectingCountry id="F35E7627E555FFCA88D09F2E67C07F0D" box="[249,465,805,830]" name="South Africa" pageId="2" pageNumber="69">
<emphasis id="B93DEAA5E555FFCA88D09F2E67C07F0D" bold="true" box="[249,465,805,830]" pageId="2" pageNumber="69">SOUTH AFRICA</emphasis>
</collectingCountry>
, KZN,
<location id="8E96606CE555FFCA8A019F2E65167F0E" LSID="urn:lsid:plazi:treatment:03E087A1E556FFCA88BE997964B47855:8E96606CE555FFCA8A019F2E65167F0E" box="[552,775,805,830]" country="South Africa" name="Itala Game Reserve" pageId="2" pageNumber="69">Itala Game Reserve</location>
27[◦] 27[’] 53.6[”] S; 31[◦] 15[’] 36.1[”] E,
<date id="FFF71077E555FFCA8CD09F2E63747F0D" box="[1273,1381,805,830]" pageId="2" pageNumber="69" value="1999-01-13">
<collectingDate id="EFB3E99FE555FFCA8CD09F2E63747F0D" box="[1273,1381,805,830]" pageId="2" pageNumber="69" value="1999-01-13">13.i.1999</collectingDate>
</date>
, leg:
<collectorName id="26BC5361E555FFCA88BE9F4766FA7F56" box="[151,235,844,869]" pageId="2" pageNumber="69">Chown</collectorName>
,
<collectorName id="26BC5361E555FFCA88D09F4767787F56" box="[249,361,844,869]" pageId="2" pageNumber="69">McGeogh</collectorName>
&amp;
<collectorName id="26BC5361E555FFCA89BB9F4767CA7F56" box="[402,475,844,869]" pageId="2" pageNumber="69">Davis.</collectorName>
(
<specimenCount id="9D4FFD3EE555FFCA89C49F4664007F56" box="[493,529,845,869]" pageId="2" pageNumber="69" type="male">2♂</specimenCount>
<emphasis id="B93DEAA5E555FFCA8A329F4764767F56" bold="true" box="[539,615,844,869]" pageId="2" pageNumber="69">
<collectionCode id="ED58AE72E555FFCA8A329F4764767F56" box="[539,615,844,869]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="2" pageNumber="69">SANC</collectionCode>
</emphasis>
)
</materialsCitation>
:
<materialsCitation id="3B213CEAE555FFCA8AAB9F4765757FBA" ID-GBIF-Occurrence="1290984950" collectingDate="1978-10-15" collectionCode="SANC" collectorName="Bornemissza &amp; Aschenborn." country="South Africa" location="Jozini" pageId="2" pageNumber="69" specimenCount="21" specimenCount-female="14" specimenCount-male="7" typeStatus="paratype">
<emphasis id="B93DEAA5E555FFCA8AAB9F47654A7F56" bold="true" box="[642,859,844,869]" pageId="2" pageNumber="69">
<collectingCountry id="F35E7627E555FFCA8AAB9F47654A7F56" box="[642,859,844,869]" name="South Africa" pageId="2" pageNumber="69">SOUTH AFRICA</collectingCountry>
</emphasis>
, KZN,
<location id="8E96606CE555FFCA8B9F9F4765E87F56" LSID="urn:lsid:plazi:treatment:03E087A1E556FFCA88BE997964B47855:8E96606CE555FFCA8B9F9F4765E87F56" box="[950,1017,844,869]" country="South Africa" name="Jozini" pageId="2" pageNumber="69">Jozini</location>
27[◦] 26[’] 0[”] S; 32[◦] 4[’] 0[”] E,
<date id="FFF71077E555FFCA88BE9F7B67187FBA" box="[151,265,880,905]" pageId="2" pageNumber="69" value="1978-10-15">
<collectingDate id="EFB3E99FE555FFCA88BE9F7B67187FBA" box="[151,265,880,905]" pageId="2" pageNumber="69" value="1978-10-15">15.x.1978</collectingDate>
</date>
, leg:
<collectorName id="26BC5361E555FFCA89609F7B67CB7FBA" box="[329,474,880,905]" pageId="2" pageNumber="69">Bornemissza</collectorName>
&amp;
<collectorName id="26BC5361E555FFCA8A299F7B649E7FBA" box="[512,655,880,905]" pageId="2" pageNumber="69">Aschenborn.</collectorName>
(
<specimenCount id="9D4FFD3EE555FFCA8A899F7A64D47FBA" box="[672,709,881,905]" pageId="2" pageNumber="69" type="male">7♂</specimenCount>
,
<specimenCount id="9D4FFD3EE555FFCA8AFB9F7A65127FBA" box="[722,771,881,905]" pageId="2" pageNumber="69" type="female">14♀</specimenCount>
<emphasis id="B93DEAA5E555FFCA8B259F7B65497FBA" bold="true" box="[780,856,880,905]" pageId="2" pageNumber="69">
<collectionCode id="ED58AE72E555FFCA8B259F7B65497FBA" box="[780,856,880,905]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="2" pageNumber="69">SANC</collectionCode>
</emphasis>
)
</materialsCitation>
:
<materialsCitation id="3B213CEAE555FFCA8B589F7B62A17F83" ID-GBIF-Occurrence="1290984955" collectingDate="1983-12-12" collectingDateMax="1983-12-13" collectingDateMin="1983-12-12" collectionCode="SANC" collectorName="Davis." country="South Africa" location="Pretoria" pageId="2" pageNumber="69" specimenCount="1" specimenCount-male="1" stateProvince="Gauteng" typeStatus="paratype">
<emphasis id="B93DEAA5E555FFCA8B589F7B62587FBA" bold="true" box="[881,1097,880,905]" pageId="2" pageNumber="69">
<collectingCountry id="F35E7627E555FFCA8B589F7B62587FBA" box="[881,1097,880,905]" name="South Africa" pageId="2" pageNumber="69">SOUTH AFRICA</collectingCountry>
</emphasis>
,
<collectingRegion id="498DF855E555FFCA8C719F7A62AB7FBA" box="[1112,1210,881,905]" country="South Africa" name="Gauteng" pageId="2" pageNumber="69">Gauteng</collectingRegion>
,
<location id="8E96606CE555FFCA8CEF9F7B63337FBA" LSID="urn:lsid:plazi:treatment:03E087A1E556FFCA88BE997964B47855:8E96606CE555FFCA8CEF9F7B63337FBA" box="[1222,1314,880,905]" country="South Africa" name="Pretoria" pageId="2" pageNumber="69" stateProvince="Gauteng">Pretoria</location>
,
<locationDeviation id="9A442F95E555FFCA8D079F7B67417F9C" pageId="2" pageNumber="69">
<quantity id="4CB19B52E555FFCA8D079F7B637F7FBA" box="[1326,1390,880,905]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="69" unit="km" value="35.0">35km</quantity>
NE of Pretoria Farm
</locationDeviation>
25[◦] 42[’] 0[”] S; 28[◦] 13[’] 0[”] E,
<date id="FFF71077E555FFCA8ADA9F9C658B7F83" box="[755,922,919,944]" pageId="2" pageNumber="69" value="1983-12-12" valueMax="1983-12-13" valueMin="1983-12-12">
<collectingDate id="EFB3E99FE555FFCA8ADA9F9C658B7F83" box="[755,922,919,944]" pageId="2" pageNumber="69" value="1983-12-12" valueMax="1983-12-13" valueMin="1983-12-12">12-13.xii.1983</collectingDate>
</date>
, leg:
<collectorName id="26BC5361E555FFCA8BFE9F9C62317F83" box="[983,1056,919,944]" pageId="2" pageNumber="69">Davis.</collectorName>
(
<specimenCount id="9D4FFD3EE555FFCA8C189F9C62427F9C" box="[1073,1107,919,943]" pageId="2" pageNumber="69" type="male">1♂</specimenCount>
<emphasis id="B93DEAA5E555FFCA8C739F9C62A17F83" bold="true" box="[1114,1200,919,944]" pageId="2" pageNumber="69">
<collectionCode id="ED58AE72E555FFCA8C739F9C62B97F83" box="[1114,1192,919,944]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="2" pageNumber="69">SANC</collectionCode>
)
</emphasis>
</materialsCitation>
:
<materialsCitation id="3B213CEAE555FFCA8C969F9C67137FC9" ID-GBIF-Occurrence="1290984956" collectingDate="2010-11-11" collectorName="AJ Armstrong &amp; G Van Bassouw." country="South Africa" location="Shongweni Farm" municipality="Weke-Weke Stream" pageId="2" pageNumber="69" specimenCount="1" typeStatus="paratype">
<emphasis id="B93DEAA5E555FFCA8C969F9C63847F83" bold="true" box="[1215,1429,919,944]" pageId="2" pageNumber="69">
<collectingCountry id="F35E7627E555FFCA8C969F9C63847F83" box="[1215,1429,919,944]" name="South Africa" pageId="2" pageNumber="69">SOUTH AFRICA</collectingCountry>
</emphasis>
, KZN,
<collectingMunicipality id="6B92ACCDE555FFCA88C99FB667D37FE5" box="[224,450,957,982]" pageId="2" pageNumber="69">Weke-Weke Stream</collectingMunicipality>
,
<location id="8E96606CE555FFCA89E79FB664817FE5" LSID="urn:lsid:plazi:treatment:03E087A1E556FFCA88BE997964B47855:8E96606CE555FFCA89E79FB664817FE5" box="[462,656,957,982]" country="South Africa" municipality="Weke-Weke Stream" name="Shongweni Farm" pageId="2" pageNumber="69">Shongweni Farm</location>
29[◦] 48[’] S; 30[◦] 43[’] E,
<date id="FFF71077E555FFCA8BFA9FB6625D7FE5" box="[979,1100,957,982]" pageId="2" pageNumber="69" value="2010-11-11">
<collectingDate id="EFB3E99FE555FFCA8BFA9FB6625D7FE5" box="[979,1100,957,982]" pageId="2" pageNumber="69" value="2010-11-11">11.xi.2010</collectingDate>
</date>
, leg:
<collectorName id="26BC5361E555FFCA8CA59FB5633C7FE5" box="[1164,1325,958,982]" pageId="2" pageNumber="69">AJ Armstrong</collectorName>
&amp;
<collectorName id="26BC5361E555FFCA8D7D9FB567137FC9" pageId="2" pageNumber="69">G Van Bassouw.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8BF636B7E555FFCA88EE980E64B47855" blockId="2.[151,1437,151,1126]" pageId="2" pageNumber="69">
<emphasis id="B93DEAA5E555FFCA88EE980E673B782D" bold="true" box="[199,298,1029,1054]" pageId="2" pageNumber="69">Habitat.</emphasis>
This new species has been recorded in patches of dense woodland and thicket. On the bushveld near Pretoria, it was sampled only in thickets on deep sand and not in open woodland or grassland (
<bibRefCitation id="EFD84B46E555FFCA8CED9821635B7870" author="Davis" box="[1220,1354,1066,1091]" pageId="2" pageNumber="69" refString="Davis, A. L. V. (1996) Habitat associations in a South Africa, Summer rainfall, dung beetle community (Coleoptera: Scarabaeidae, Aphodiidae, Staphylinidae, Histeridae, Hydrophilidae), Pedologia, 40, 260 - 280." type="journal article" year="1996">Davis 1996</bibRefCitation>
). It has been recorded only from
<collectingCountry id="F35E7627E555FFCA899B984664507855" box="[434,577,1101,1126]" name="South Africa" pageId="2" pageNumber="69">South Africa</collectingCountry>
(
<figureCitation id="13722A32E555FFCA8A7B984664847855" box="[594,661,1101,1126]" captionStart="FIGURE 4" captionStartId="6.[151,250,1720,1742]" captionTargetBox="[408,1246,841,1677]" captionTargetId="figure@6.[340,1247,841,1698]" captionTargetPageId="6" captionText="FIGURE 4. Map of southern Africa showing the geographic distribution of the new species: S. umbraphilus, S. oralensis and S. neobornemisszanus." httpUri="https://zenodo.org/record/266255/files/figure.png" pageId="2" pageNumber="69">
<emphasis id="B93DEAA5E555FFCA8A7B984664847855" bold="true" box="[594,661,1101,1126]" pageId="2" pageNumber="69">Fig. 4</emphasis>
</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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