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<mods:title id="9C81BD653C28DB57A393DB5530391F2A">Revision of the Mexican ladybird genus Vedalia Mulsant (Coleoptera: Coccinellidae)</mods:title>
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<mods:namePart id="1D5D6ECCD4959D854F11419B450A4B0B">Czerwiński, Tomasz</mods:namePart>
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<mods:namePart id="406332F67EA49ACB48CAA50957C9F881">Szawaryn, Karol</mods:namePart>
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<paragraph id="8BF6E802FFCABD6AFF1BF98D9B4770EE" blockId="1.[151,513,1662,1689]" box="[151,513,1662,1689]" pageId="1" pageNumber="154">
<heading id="D0BE5F6EFFCABD6AFF1BF98D9B4770EE" bold="true" box="[151,513,1662,1689]" fontSize="11" level="1" pageId="1" pageNumber="154" reason="1">
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Genus
<taxonomicName id="4C499381FFCABD6AFF62F98D9B4770EE" authority="Mulsant, 1850" authorityName="Mulsant" authorityYear="1850" box="[238,513,1662,1689]" class="Insecta" family="Coccinellidae" genus="Vedalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="154" phylum="Arthropoda" rank="genus">
<emphasis id="B93D3410FFCABD6AFF62F98D980E70EF" bold="true" box="[238,328,1662,1689]" italics="true" pageId="1" pageNumber="154">Vedalia</emphasis>
<bibRefCitation id="EFD895F3FFCABD6AFEC3F98C9B4770EE" author="Mulsant, E." box="[335,513,1662,1689]" pageId="1" pageNumber="154" pagination="1 - 1104" refId="ref6251" refString="Mulsant, E. (1850) Species des Coleopteres trimeres securipalpes. Annales des Sciences Physiques et Naturelles, d'Agriculture et d'Industrie, Lyon, Series 2, 2, 1 - 1104. https: // doi. org / 10.5962 / bhl. title. 8953" type="journal article" year="1850">Mulsant, 1850</bibRefCitation>
</taxonomicName>
</emphasis>
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<paragraph id="8BF6E802FFCABD6AFF1BF93098F670AD" blockId="1.[151,1094,1731,1786]" box="[151,432,1731,1755]" pageId="1" pageNumber="154">
<treatmentCitationGroup id="AB59CF2CFFCABD6AFF1BF93098F670AD" box="[151,432,1731,1755]" pageId="1" pageNumber="154">
<treatmentCitation id="0AE8CE13FFCABD6AFF1BF93098EA70AD" author="Mulsant, E." box="[151,428,1731,1755]" page="905" pageId="1" pageNumber="154" year="1850">
<taxonomicName id="4C499381FFCABD6AFF1BF93098EA70AD" authority="Mulsant 1850: 905" authorityName="Mulsant" authorityPageNumber="905" authorityYear="1850" box="[151,428,1731,1755]" class="Insecta" family="Coccinellidae" genus="Vedalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="154" phylum="Arthropoda" rank="genus">
<emphasis id="B93D3410FFCABD6AFF1BF93099A570AD" box="[151,227,1731,1755]" italics="true" pageId="1" pageNumber="154">Vedalia</emphasis>
<bibRefCitation id="EFD895F3FFCABD6AFF66F93098EA70AD" author="Mulsant, E." box="[234,428,1731,1755]" pageId="1" pageNumber="154" pagination="1 - 1104" refId="ref6251" refString="Mulsant, E. (1850) Species des Coleopteres trimeres securipalpes. Annales des Sciences Physiques et Naturelles, d'Agriculture et d'Industrie, Lyon, Series 2, 2, 1 - 1104. https: // doi. org / 10.5962 / bhl. title. 8953" type="journal article" year="1850">Mulsant 1850: 905</bibRefCitation>
</taxonomicName>
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.
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</paragraph>
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<paragraph id="8BF6E802FFCABD6AFF1BF9119D00708C" blockId="1.[151,1094,1731,1786]" box="[151,1094,1762,1786]" pageId="1" pageNumber="154">
<typeStatus id="54F256A0FFCABD6AFF1BF911998F708C" box="[151,201,1762,1786]" pageId="1" pageNumber="154">Type</typeStatus>
species:
<taxonomicName id="4C499381FFCABD6AFEAAF9119B2A708C" authority="Mulsant, 1850" authorityName="Mulsant. Since" authorityYear="1850" box="[294,620,1762,1786]" class="Insecta" family="Coccinellidae" genus="Vedalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="154" phylum="Arthropoda" rank="species" species="sieboldii">
<emphasis id="B93D3410FFCABD6AFEAAF9119896708C" box="[294,464,1762,1786]" italics="true" pageId="1" pageNumber="154">Vedalia sieboldii</emphasis>
<bibRefCitation id="EFD895F3FFCABD6AFE5AF9119B2A708C" author="Mulsant, E." box="[470,620,1762,1786]" pageId="1" pageNumber="154" pagination="1 - 1104" refId="ref6251" refString="Mulsant, E. (1850) Species des Coleopteres trimeres securipalpes. Annales des Sciences Physiques et Naturelles, d'Agriculture et d'Industrie, Lyon, Series 2, 2, 1 - 1104. https: // doi. org / 10.5962 / bhl. title. 8953" type="journal article" year="1850">Mulsant, 1850</bibRefCitation>
</taxonomicName>
; subsequent designation by
<bibRefCitation id="EFD895F3FFCABD6AFC04F9119D04708C" author="Crotch, G. R." box="[904,1090,1762,1786]" pageId="1" pageNumber="154" refId="ref5678" refString="Crotch, G. R. (1874) A revision of the coleopterous family Coccinellidae. E. W. Janson, London, xv + 311 pp. https: // doi. org / 10.5962 / bhl. title. 8975" type="book" year="1874">Crotch, 1874: 281</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C353BB89FFCABD68FF1BF8DB99BF7737" lastPageId="3" lastPageNumber="156" pageId="1" pageNumber="154" type="diagnosis">
<paragraph id="8BF6E802FFCABD6AFF1BF8DB9C2471A4" blockId="1.[151,1437,1832,2003]" pageId="1" pageNumber="154">
<emphasis id="B93D3410FFCABD6AFF1BF8DB98547134" bold="true" box="[151,274,1832,1858]" pageId="1" pageNumber="154">Diagnosis.</emphasis>
Based on the following set of characters
<taxonomicName id="4C499381FFCABD6AFD58F8DB9A6E7134" authorityName="Mulsant" authorityYear="1850" box="[724,808,1832,1858]" class="Insecta" family="Coccinellidae" genus="Vedalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="154" phylum="Arthropoda" rank="genus">
<emphasis id="B93D3410FFCABD6AFD58F8DB9A6E7134" box="[724,808,1832,1858]" italics="true" pageId="1" pageNumber="154">Vedalia</emphasis>
</taxonomicName>
belongs to the tribe
<taxonomicName id="4C499381FFCABD6AFB8AF8DB9DC77134" authorityName="Mulsant" authorityYear="1850" box="[1030,1153,1832,1858]" class="Insecta" family="Coccinellidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="154" phylum="Arthropoda" rank="tribe" tribe="Chnoodini">Chnoodini</taxonomicName>
: antenna short, composed of 11 antennomeres (
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other
<taxonomicName id="4C499381FFCABD6AFDB4F8BF9BF67110" authorityName="Mulsant" authorityYear="1850" box="[568,688,1868,1894]" class="Insecta" family="Coccinellidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="161" phylum="Arthropoda" rank="tribe" tribe="Chnoodini">Chnoodini</taxonomicName>
) (
<figureCitation id="1372F487FFCABD6AFD44F8BF9A5D7110" box="[712,795,1868,1894]" captionStart="FIGURE 3" captionStartId="4.[151,250,1895,1919]" captionTargetBox="[190,1397,237,1871]" captionTargetId="figure-62@4.[190,1397,237,1871]" captionTargetPageId="4" captionText="FIGURE 3. Vedalia sieboldii Mulsant, 1850; SEM illustrations: (a) antenna; (b) mouthparts; (c) pronotum; (d) head, prothorax and mesothorax, ventral; (e) scutellar shield; (f) meso- and metathorax, ventral; (g) tibial spurs on hind leg; (h) protarsal claw, male; (i) hind tarsal claw, female." figureDoi="http://doi.org/10.5281/zenodo.10611487" httpUri="https://zenodo.org/record/10611487/files/figure.png" pageId="1" pageNumber="154">Fig. 3a</figureCitation>
), scape greatly expanded, swollen (
<figureCitation id="1372F487FFCABD6AFB3DF8BF9C427110" box="[1201,1284,1868,1894]" captionStart="FIGURE 3" captionStartId="4.[151,250,1895,1919]" captionTargetBox="[190,1397,237,1871]" captionTargetId="figure-62@4.[190,1397,237,1871]" captionTargetPageId="4" captionText="FIGURE 3. Vedalia sieboldii Mulsant, 1850; SEM illustrations: (a) antenna; (b) mouthparts; (c) pronotum; (d) head, prothorax and mesothorax, ventral; (e) scutellar shield; (f) meso- and metathorax, ventral; (g) tibial spurs on hind leg; (h) protarsal claw, male; (i) hind tarsal claw, female." figureDoi="http://doi.org/10.5281/zenodo.10611487" httpUri="https://zenodo.org/record/10611487/files/figure.png" pageId="1" pageNumber="154">Fig. 3a</figureCitation>
); distinct eye canthus present (
<figureCitation id="1372F487FFCABD6AFED1F88398F571FC" box="[349,435,1904,1930]" captionStart="FIGURE 1" captionStartId="2.[151,250,1514,1538]" captionTargetBox="[158,1430,214,1489]" captionTargetId="figure-34@2.[158,1430,179,1489]" captionTargetPageId="2" captionText="FIGURE 1. Vedalia sieboldii Mulsant, 1850: habitus: (a) dorsal view; (b) lateral view; (c) frontal view; (d) head; (e) living specimen on cactus Cylindropuntia tunicata (Zumpango); (f) living specimen (Monte Albán); Photos: (e) Zahir Santillán, (f) John Kemner (modified)." figureDoi="http://doi.org/10.5281/zenodo.10611483" httpUri="https://zenodo.org/record/10611483/files/figure.png" pageId="1" pageNumber="154">Fig. 1d</figureCitation>
); terminal maxillary palpomere broad, securiform (
<figureCitation id="1372F487FFCABD6AFB9CF8839D2E71FC" box="[1040,1128,1904,1930]" captionStart="FIGURE 3" captionStartId="4.[151,250,1895,1919]" captionTargetBox="[190,1397,237,1871]" captionTargetId="figure-62@4.[190,1397,237,1871]" captionTargetPageId="4" captionText="FIGURE 3. Vedalia sieboldii Mulsant, 1850; SEM illustrations: (a) antenna; (b) mouthparts; (c) pronotum; (d) head, prothorax and mesothorax, ventral; (e) scutellar shield; (f) meso- and metathorax, ventral; (g) tibial spurs on hind leg; (h) protarsal claw, male; (i) hind tarsal claw, female." figureDoi="http://doi.org/10.5281/zenodo.10611487" httpUri="https://zenodo.org/record/10611487/files/figure.png" pageId="1" pageNumber="154">Fig. 3d</figureCitation>
); labial palp composed of three palpomeres (
<figureCitation id="1372F487FFCABD6AFEE5F86798FB71D8" box="[361,445,1940,1966]" captionStart="FIGURE 3" captionStartId="4.[151,250,1895,1919]" captionTargetBox="[190,1397,237,1871]" captionTargetId="figure-62@4.[190,1397,237,1871]" captionTargetPageId="4" captionText="FIGURE 3. Vedalia sieboldii Mulsant, 1850; SEM illustrations: (a) antenna; (b) mouthparts; (c) pronotum; (d) head, prothorax and mesothorax, ventral; (e) scutellar shield; (f) meso- and metathorax, ventral; (g) tibial spurs on hind leg; (h) protarsal claw, male; (i) hind tarsal claw, female." figureDoi="http://doi.org/10.5281/zenodo.10611487" httpUri="https://zenodo.org/record/10611487/files/figure.png" pageId="1" pageNumber="154">Fig. 3b</figureCitation>
); femora and tibiae broadened (
<figureCitation id="1372F487FFCABD6AFCAAF8679A3D71D8" box="[806,891,1940,1966]" captionStart="FIGURE 2" captionStartId="3.[151,250,1457,1481]" captionTargetBox="[190,1397,453,1434]" captionTargetId="figure-129@3.[190,1397,453,1434]" captionTargetPageId="3" captionText="FIGURE 2. Vedalia sieboldii Mulsant, 1850; SEM illustrations: (a) habitus, dorsal; (b) hind leg; (c) habitus, ventral; (d) abdomen." figureDoi="http://doi.org/10.5281/zenodo.10611485" httpUri="https://zenodo.org/record/10611485/files/figure.png" pageId="1" pageNumber="154">Fig. 2b</figureCitation>
); tibiae with two apical spurs (
<figureCitation id="1372F487FFCABD6AFB55F8679C6D71D8" box="[1241,1323,1940,1966]" captionStart="FIGURE 3" captionStartId="4.[151,250,1895,1919]" captionTargetBox="[190,1397,237,1871]" captionTargetId="figure-62@4.[190,1397,237,1871]" captionTargetPageId="4" captionText="FIGURE 3. Vedalia sieboldii Mulsant, 1850; SEM illustrations: (a) antenna; (b) mouthparts; (c) pronotum; (d) head, prothorax and mesothorax, ventral; (e) scutellar shield; (f) meso- and metathorax, ventral; (g) tibial spurs on hind leg; (h) protarsal claw, male; (i) hind tarsal claw, female." figureDoi="http://doi.org/10.5281/zenodo.10611487" httpUri="https://zenodo.org/record/10611487/files/figure.png" pageId="1" pageNumber="154">
Fig.
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</figureCitation>
) (in other
<taxonomicName id="4C499381FFCABD6AFF1BF84B984B71A4" authorityName="Mulsant" authorityYear="1850" box="[151,269,1976,2002]" class="Insecta" family="Coccinellidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="161" phylum="Arthropoda" rank="tribe" tribe="Chnoodini">Chnoodini</taxonomicName>
mostly absent or single, rarely double spurs present in some species of
<taxonomicName id="4C499381FFCABD6AFBBCF84B9DED71A4" box="[1072,1195,1976,2002]" class="Insecta" family="Coccinellidae" genus="Exoplectra" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="154" phylum="Arthropoda" rank="genus">
<emphasis id="B93D3410FFCABD6AFBBCF84B9DED71A4" box="[1072,1195,1976,2002]" italics="true" pageId="1" pageNumber="154">Exoplectra</emphasis>
</taxonomicName>
and
<taxonomicName id="4C499381FFCABD6AFB6DF84B9C1771A4" authorityName="Chevrolat" authorityYear="1837" box="[1249,1361,1976,2002]" class="Insecta" family="Coccinellidae" genus="Chnoodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="154" phylum="Arthropoda" rank="genus">
<emphasis id="B93D3410FFCABD6AFB6DF84B9C1771A4" box="[1249,1361,1976,2002]" italics="true" pageId="1" pageNumber="154">Chnoodes</emphasis>
</taxonomicName>
).
</paragraph>
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<paragraph id="8BF6E802FFC9BD69FF1BFA1998DF703D" blockId="2.[151,1436,1514,1611]" pageId="2" pageNumber="155">
<emphasis id="B93D3410FFC9BD69FF1BFA1998537074" bold="true" box="[151,277,1514,1538]" pageId="2" pageNumber="155">FIGURE 1.</emphasis>
<taxonomicName id="4C499381FFC9BD69FE91FA189B2C7074" authority="Mulsant, 1850" authorityName="Mulsant. Since" authorityYear="1850" box="[285,618,1514,1539]" class="Insecta" family="Coccinellidae" genus="Vedalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="155" phylum="Arthropoda" rank="species" species="sieboldii">
<emphasis id="B93D3410FFC9BD69FE91FA18988C7075" box="[285,458,1515,1539]" italics="true" pageId="2" pageNumber="155">Vedalia sieboldii</emphasis>
<bibRefCitation id="EFD895F3FFC9BD69FE5FFA189B2C7074" author="Mulsant, E." box="[467,618,1514,1539]" pageId="2" pageNumber="155" pagination="1 - 1104" refId="ref6251" refString="Mulsant, E. (1850) Species des Coleopteres trimeres securipalpes. Annales des Sciences Physiques et Naturelles, d'Agriculture et d'Industrie, Lyon, Series 2, 2, 1 - 1104. https: // doi. org / 10.5962 / bhl. title. 8953" type="journal article" year="1850">Mulsant, 1850</bibRefCitation>
</taxonomicName>
: habitus: (
<emphasis id="B93D3410FFC9BD69FD5AFA189BA57075" bold="true" box="[726,739,1515,1539]" pageId="2" pageNumber="155">a</emphasis>
) dorsal view; (
<emphasis id="B93D3410FFC9BD69FC0EFA189AD67075" bold="true" box="[898,912,1515,1539]" pageId="2" pageNumber="155">b</emphasis>
) lateral view; (
<emphasis id="B93D3410FFC9BD69FBBCFA189D7D7075" bold="true" box="[1072,1083,1515,1539]" pageId="2" pageNumber="155">c</emphasis>
) frontal view; (
<emphasis id="B93D3410FFC9BD69FB6CFA189DA87075" bold="true" box="[1248,1262,1515,1539]" pageId="2" pageNumber="155">d</emphasis>
) head; (
<emphasis id="B93D3410FFC9BD69FAC9FA189C167075" bold="true" box="[1349,1360,1515,1539]" pageId="2" pageNumber="155">e</emphasis>
) living specimen on cactus
<taxonomicName id="4C499381FFC9BD69FEEBF9FC9BA07051" authority="(Zumpango)" baseAuthorityName="Zumpango" box="[359,742,1551,1575]" class="Magnoliopsida" family="Cactaceae" genus="Cylindropuntia" kingdom="Plantae" order="Caryophyllales" pageId="2" pageNumber="155" phylum="Tracheophyta" rank="species" species="tunicata">
<emphasis id="B93D3410FFC9BD69FEEBF9FC9B1A7050" box="[359,604,1551,1575]" italics="true" pageId="2" pageNumber="155">Cylindropuntia tunicata</emphasis>
(Zumpango)
</taxonomicName>
; (
<emphasis id="B93D3410FFC9BD69FD76F9FD9A447050" bold="true" box="[762,770,1550,1574]" pageId="2" pageNumber="155">f</emphasis>
) living specimen (Monte Albán); Photos: (
<emphasis id="B93D3410FFC9BD69FB4DF9FC9D8A7051" bold="true" box="[1217,1228,1551,1575]" pageId="2" pageNumber="155">e</emphasis>
) Zahir Santillán, (
<emphasis id="B93D3410FFC9BD69FA07F9FD9CD57050" bold="true" box="[1419,1427,1550,1574]" pageId="2" pageNumber="155">f</emphasis>
) John Kemner (modified).
</paragraph>
</caption>
<paragraph id="8BF6E802FFC9BD68FF4BF96699BF7737" blockId="2.[151,1437,1685,2035]" lastBlockId="3.[151,1437,151,429]" lastPageId="3" lastPageNumber="156" pageId="2" pageNumber="155">
<taxonomicName id="4C499381FFC9BD69FF4BF966985D70D9" authorityName="Mulsant" authorityYear="1850" box="[199,283,1685,1711]" class="Insecta" family="Coccinellidae" genus="Vedalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="B93D3410FFC9BD69FF4BF966985D70D9" box="[199,283,1685,1711]" italics="true" pageId="2" pageNumber="155">Vedalia</emphasis>
</taxonomicName>
can be distinguished from all other genera of
<taxonomicName id="4C499381FFC9BD69FCBEF9669AEE70D9" authorityName="Mulsant" authorityYear="1850" box="[818,936,1685,1711]" class="Insecta" family="Coccinellidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="161" phylum="Arthropoda" rank="tribe" tribe="Chnoodini">Chnoodini</taxonomicName>
by the shape of abdominal postcoxal lines: incomplete, recurved, not reaching hind edge of ventrite 1, while in genera
<taxonomicName id="4C499381FFC9BD69FC78F94A9DA770A5" authority="Crotch" authorityName="Crotch" authorityYear="1874" box="[1012,1249,1721,1747]" class="Insecta" family="Coccinellidae" genus="Neorhizobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="B93D3410FFC9BD69FC78F94A9DCF70A5" box="[1012,1161,1721,1747]" italics="true" pageId="2" pageNumber="155">Neorhizobius</emphasis>
Crotch
</taxonomicName>
,
<taxonomicName id="4C499381FFC9BD69FB61F9499CDA70A5" authority="Mulsant" authorityName="Mulsant" authorityYear="1850" box="[1261,1436,1721,1747]" class="Insecta" family="Coccinellidae" genus="Dioria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="B93D3410FFC9BD69FB61F9499C7E70A5" box="[1261,1336,1722,1747]" italics="true" pageId="2" pageNumber="155">Dioria</emphasis>
Mulsant
</taxonomicName>
and
<taxonomicName id="4C499381FFC9BD69FF45F92E98C77081" authority="Mulsant" authorityName="Mulsant" authorityYear="1850" box="[201,385,1757,1783]" class="Insecta" family="Coccinellidae" genus="Sidonis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="B93D3410FFC9BD69FF45F92E985D7081" box="[201,283,1757,1783]" italics="true" pageId="2" pageNumber="155">Sidonis</emphasis>
Mulsant
</taxonomicName>
they are complete; in
<taxonomicName id="4C499381FFC9BD69FD0BF92E9A377081" authority="Gonzalez" authorityName="Gonzalez" box="[647,881,1757,1783]" class="Insecta" family="Coccinellidae" genus="Gordonita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="B93D3410FFC9BD69FD0BF92E9BBD7081" box="[647,763,1757,1783]" italics="true" pageId="2" pageNumber="155">Gordonita</emphasis>
González
</taxonomicName>
,
<taxonomicName id="4C499381FFC9BD69FCF2F92E9D207081" authority="Chevrolat" authorityName="Chevrolat" authorityYear="1837" box="[894,1126,1757,1783]" class="Insecta" family="Coccinellidae" genus="Chnoodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="B93D3410FFC9BD69FCF2F92E9AA87081" box="[894,1006,1757,1783]" italics="true" pageId="2" pageNumber="155">Chnoodes</emphasis>
Chevrolat
</taxonomicName>
and
<taxonomicName id="4C499381FFC9BD69FB2EF92E9C5B7081" box="[1186,1309,1757,1783]" class="Insecta" family="Coccinellidae" genus="Exoplectra" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="B93D3410FFC9BD69FB2EF92E9C5B7081" box="[1186,1309,1757,1783]" italics="true" pageId="2" pageNumber="155">Exoplectra</emphasis>
</taxonomicName>
they reach hind edge of ventrite 1; in
<taxonomicName id="4C499381FFC9BD69FE35F8F29BCF716D" authority="Mulsant" authorityName="Mulsant" authorityYear="1850" box="[441,649,1793,1819]" class="Insecta" family="Coccinellidae" genus="Coeliaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="B93D3410FFC9BD69FE35F8F29B62716D" box="[441,548,1793,1819]" italics="true" pageId="2" pageNumber="155">Coeliaria</emphasis>
Mulsant
</taxonomicName>
,
<taxonomicName id="4C499381FFC9BD69FD1FF8F29A15716D" authority="Mulsant" authorityName="Mulsant" authorityYear="1850" box="[659,851,1793,1819]" class="Insecta" family="Coccinellidae" genus="Dapolia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="B93D3410FFC9BD69FD1FF8F29BA9716D" box="[659,751,1793,1819]" italics="true" pageId="2" pageNumber="155">Dapolia</emphasis>
Mulsant
</taxonomicName>
,
<taxonomicName id="4C499381FFC9BD69FCD1F8F19D74716D" authority="Gonzalez" authorityName="Gonzalez" box="[861,1074,1793,1819]" class="Insecta" family="Coccinellidae" genus="Incurvus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="B93D3410FFC9BD69FCD1F8F19AF9716D" box="[861,959,1794,1819]" italics="true" pageId="2" pageNumber="155">Incurvus</emphasis>
González
</taxonomicName>
,
<taxonomicName id="4C499381FFC9BD69FBB7F8F29D91716D" authority="Mulsant" authorityName="Mulsant" authorityYear="1850" box="[1083,1239,1793,1819]" class="Insecta" family="Coccinellidae" genus="Siola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="B93D3410FFC9BD69FBB7F8F29D33716D" box="[1083,1141,1793,1819]" italics="true" pageId="2" pageNumber="155">Siola</emphasis>
Mulsant
</taxonomicName>
and some species of
<taxonomicName id="4C499381FFC9BD69FF39F8D698637149" authorityName="Chevrolat" authorityYear="1837" box="[181,293,1829,1855]" class="Insecta" family="Coccinellidae" genus="Chnoodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="B93D3410FFC9BD69FF39F8D698637149" box="[181,293,1829,1855]" italics="true" pageId="2" pageNumber="155">Chnoodes</emphasis>
</taxonomicName>
and
<taxonomicName id="4C499381FFC9BD69FED6F8D698937149" box="[346,469,1829,1855]" class="Insecta" family="Coccinellidae" genus="Exoplectra" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="B93D3410FFC9BD69FED6F8D698937149" box="[346,469,1829,1855]" italics="true" pageId="2" pageNumber="155">Exoplectra</emphasis>
</taxonomicName>
an additional oblique line is present.
<taxonomicName id="4C499381FFC9BD69FCF8F8D69A8E7149" authorityName="Mulsant" authorityYear="1850" box="[884,968,1829,1855]" class="Insecta" family="Coccinellidae" genus="Vedalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="B93D3410FFC9BD69FCF8F8D69A8E7149" box="[884,968,1829,1855]" italics="true" pageId="2" pageNumber="155">Vedalia</emphasis>
</taxonomicName>
in general appearance resembles the most
<taxonomicName id="4C499381FFC9BD69FF1BF8BA98417115" authorityName="Chevrolat" authorityYear="1837" box="[151,263,1865,1891]" class="Insecta" family="Coccinellidae" genus="Chnoodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="B93D3410FFC9BD69FF1BF8BA98417115" box="[151,263,1865,1891]" italics="true" pageId="2" pageNumber="155">Chnoodes</emphasis>
</taxonomicName>
,
<taxonomicName id="4C499381FFC9BD69FE9BF8BA98D47115" box="[279,402,1865,1891]" class="Insecta" family="Coccinellidae" genus="Exoplectra" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="B93D3410FFC9BD69FE9BF8BA98D47115" box="[279,402,1865,1891]" italics="true" pageId="2" pageNumber="155">Exoplectra</emphasis>
</taxonomicName>
and
<taxonomicName id="4C499381FFC9BD69FE40F8BA9B587115" authorityName="Mulsant" authorityYear="1850" box="[460,542,1865,1891]" class="Insecta" family="Coccinellidae" genus="Sidonis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="B93D3410FFC9BD69FE40F8BA9B587115" box="[460,542,1865,1891]" italics="true" pageId="2" pageNumber="155">Sidonis</emphasis>
</taxonomicName>
. However, it differs from
<taxonomicName id="4C499381FFC9BD69FCC0F8BA9AFA7115" authorityName="Chevrolat" authorityYear="1837" box="[844,956,1865,1891]" class="Insecta" family="Coccinellidae" genus="Chnoodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="B93D3410FFC9BD69FCC0F8BA9AFA7115" box="[844,956,1865,1891]" italics="true" pageId="2" pageNumber="155">Chnoodes</emphasis>
</taxonomicName>
, not only by shape of postcoxal lines, but also by having larger body size, antennae composed with 11 antennomeres with 3-segmented club (most species of
<taxonomicName id="4C499381FFC9BD69FF34F862986E71DD" authorityName="Chevrolat" authorityYear="1837" box="[184,296,1937,1963]" class="Insecta" family="Coccinellidae" genus="Chnoodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="B93D3410FFC9BD69FF34F862986E71DD" box="[184,296,1937,1963]" italics="true" pageId="2" pageNumber="155">Chnoodes</emphasis>
</taxonomicName>
have antennae composed with 10 antennomeres with 4-segmented club) and double tibial spurs (in
<taxonomicName id="4C499381FFC9BD69FF1BF846984171B9" authorityName="Chevrolat" authorityYear="1837" box="[151,263,1973,1999]" class="Insecta" family="Coccinellidae" genus="Chnoodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="B93D3410FFC9BD69FF1BF846984171B9" box="[151,263,1973,1999]" italics="true" pageId="2" pageNumber="155">Chnoodes</emphasis>
</taxonomicName>
often with single spur or without spurs). From
<taxonomicName id="4C499381FFC9BD69FCA7F8469AE071B9" box="[811,934,1973,1999]" class="Insecta" family="Coccinellidae" genus="Exoplectra" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="B93D3410FFC9BD69FCA7F8469AE071B9" box="[811,934,1973,1999]" italics="true" pageId="2" pageNumber="155">Exoplectra</emphasis>
</taxonomicName>
it differs by having simple tibiae, epipleura without foveae for reception of femora, and sexual dimorphism in shape of tarsal claws (angulate tibiae, foveae on epipleurae present and non-dimorphic tarsal claws in
<taxonomicName id="4C499381FFC8BD68FD64FF649A2576C7" box="[744,867,151,177]" class="Insecta" family="Coccinellidae" genus="Exoplectra" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="156" phylum="Arthropoda" rank="genus">
<emphasis id="B93D3410FFC8BD68FD64FF649A2576C7" box="[744,867,151,177]" italics="true" pageId="3" pageNumber="156">Exoplectra</emphasis>
</taxonomicName>
). From
<taxonomicName id="4C499381FFC8BD68FC30FF649D4876C7" authorityName="Mulsant" authorityYear="1850" box="[956,1038,151,177]" class="Insecta" family="Coccinellidae" genus="Sidonis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="156" phylum="Arthropoda" rank="genus">
<emphasis id="B93D3410FFC8BD68FC30FF649D4876C7" box="[956,1038,151,177]" italics="true" pageId="3" pageNumber="156">Sidonis</emphasis>
</taxonomicName>
it can be easily separated by having oval body shape, antennae with 11 antennomeres, incomplete postcoxal lines, short and weak grooves on prosternal process, short and adjacent pubescence on dorsum (
<taxonomicName id="4C499381FFC8BD68FD6BFF2C9A7F768F" authorityName="Mulsant" authorityYear="1850" box="[743,825,223,249]" class="Insecta" family="Coccinellidae" genus="Sidonis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="156" phylum="Arthropoda" rank="genus">
<emphasis id="B93D3410FFC8BD68FD6BFF2C9A7F768F" box="[743,825,223,249]" italics="true" pageId="3" pageNumber="156">Sidonis</emphasis>
</taxonomicName>
has cordiform body, antennae with 10 antennomeres, complete postcoxal lines, complete carinae on prosternal process, long and erect pubescence on dorsal body surface).
</paragraph>
</subSubSection>
<subSubSection id="C353BB89FFC8BD6DFF4BFEB999B0759B" lastPageId="6" lastPageNumber="159" pageId="3" pageNumber="156" type="description">
<paragraph id="8BF6E802FFC8BD68FF4BFEB99DB777DB" blockId="3.[151,1437,151,429]" pageId="3" pageNumber="156">
<emphasis id="B93D3410FFC8BD68FF4BFEB998337712" bold="true" box="[199,373,330,356]" pageId="3" pageNumber="156">Redescription.</emphasis>
TL: 5.0
<quantity id="4CB145E7FFC8BD68FE53FEB99B717713" box="[479,567,330,357]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="3" pageNumber="156" unit="mm" value="6.5">6.5 mm</quantity>
; EW: 4.05.0 mm. Body oval (
<figureCitation id="1372F487FFC8BD68FC18FEB89D667713" box="[916,1056,331,357]" captionStart="FIGURE 1" captionStartId="2.[151,250,1514,1538]" captionTargetBox="[158,1430,214,1489]" captionTargetId="figure-34@2.[158,1430,179,1489]" captionTargetPageId="2" captionText="FIGURE 1. Vedalia sieboldii Mulsant, 1850: habitus: (a) dorsal view; (b) lateral view; (c) frontal view; (d) head; (e) living specimen on cactus Cylindropuntia tunicata (Zumpango); (f) living specimen (Monte Albán); Photos: (e) Zahir Santillán, (f) John Kemner (modified)." figureDoi="http://doi.org/10.5281/zenodo.10611483" httpUri="https://zenodo.org/record/10611483/files/figure.png" pageId="3" pageNumber="156">Figs 1a, d, e</figureCitation>
;
<figureCitation id="1372F487FFC8BD68FBA1FEB89D0C7713" box="[1069,1098,331,357]" captionStart="FIGURE 2" captionStartId="3.[151,250,1457,1481]" captionTargetBox="[190,1397,453,1434]" captionTargetId="figure-129@3.[190,1397,453,1434]" captionTargetPageId="3" captionText="FIGURE 2. Vedalia sieboldii Mulsant, 1850; SEM illustrations: (a) habitus, dorsal; (b) hind leg; (c) habitus, ventral; (d) abdomen." figureDoi="http://doi.org/10.5281/zenodo.10611485" httpUri="https://zenodo.org/record/10611485/files/figure.png" pageId="3" pageNumber="156">2a</figureCitation>
), moderately convex, winged. Dorsum with single size punctures, punctures on pronotum coarser than on elytra. Pubescence moderately long and uniform, directed anteriorly on pronotum, radially on scutellar shield and posterolaterally on elytra.
</paragraph>
<caption id="DF36B88AFFC8BD68FF1BFA4299BD739B" ID-DOI="http://doi.org/10.5281/zenodo.10611485" ID-Zenodo-Dep="10611485" httpUri="https://zenodo.org/record/10611485/files/figure.png" pageId="3" pageNumber="156" startId="3.[151,250,1457,1481]" targetBox="[190,1397,453,1434]" targetPageId="3" targetType="figure">
<paragraph id="8BF6E802FFC8BD68FF1BFA4299BD739B" blockId="3.[151,1436,1457,1517]" pageId="3" pageNumber="156">
<emphasis id="B93D3410FFC8BD68FF1BFA42985073BF" bold="true" box="[151,278,1457,1481]" pageId="3" pageNumber="156">FIGURE 2.</emphasis>
<taxonomicName id="4C499381FFC8BD68FEACFA429B3573BF" authority="Mulsant, 1850" authorityName="Mulsant. Since" authorityYear="1850" box="[288,627,1457,1481]" class="Insecta" family="Coccinellidae" genus="Vedalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="156" phylum="Arthropoda" rank="species" species="sieboldii">
<emphasis id="B93D3410FFC8BD68FEACFA42989673BF" box="[288,464,1457,1481]" italics="true" pageId="3" pageNumber="156">Vedalia sieboldii</emphasis>
<bibRefCitation id="EFD895F3FFC8BD68FE56FA429B3573BF" author="Mulsant, E." box="[474,627,1457,1481]" pageId="3" pageNumber="156" pagination="1 - 1104" refId="ref6251" refString="Mulsant, E. (1850) Species des Coleopteres trimeres securipalpes. Annales des Sciences Physiques et Naturelles, d'Agriculture et d'Industrie, Lyon, Series 2, 2, 1 - 1104. https: // doi. org / 10.5962 / bhl. title. 8953" type="journal article" year="1850">Mulsant, 1850</bibRefCitation>
</taxonomicName>
; SEM illustrations: (
<emphasis id="B93D3410FFC8BD68FCDCFA429A1B73BF" bold="true" box="[848,861,1457,1481]" pageId="3" pageNumber="156">a</emphasis>
) habitus, dorsal; (
<emphasis id="B93D3410FFC8BD68FBACFA429D6873BF" bold="true" box="[1056,1070,1457,1481]" pageId="3" pageNumber="156">b</emphasis>
) hind leg; (
<emphasis id="B93D3410FFC8BD68FB23FA429DFC73BF" bold="true" box="[1199,1210,1457,1481]" pageId="3" pageNumber="156">c</emphasis>
) habitus, ventral; (
<emphasis id="B93D3410FFC8BD68FA0AFA429CD273BF" bold="true" box="[1414,1428,1457,1481]" pageId="3" pageNumber="156">d</emphasis>
) abdomen.
</paragraph>
</caption>
<paragraph id="8BF6E802FFC8BD6FFF4BF9C49A2A76A3" blockId="3.[151,1436,1591,2014]" lastBlockId="4.[151,1436,151,213]" lastPageId="4" lastPageNumber="157" pageId="3" pageNumber="156">
<emphasis id="B93D3410FFC8BD68FF4BF9C498417027" bold="true" box="[199,263,1591,1617]" pageId="3" pageNumber="156">Head</emphasis>
partially covered by pronotum (
<figureCitation id="1372F487FFC8BD68FD0BF9C49BA67027" box="[647,736,1591,1617]" captionStart="FIGURE 1" captionStartId="2.[151,250,1514,1538]" captionTargetBox="[158,1430,214,1489]" captionTargetId="figure-34@2.[158,1430,179,1489]" captionTargetPageId="2" captionText="FIGURE 1. Vedalia sieboldii Mulsant, 1850: habitus: (a) dorsal view; (b) lateral view; (c) frontal view; (d) head; (e) living specimen on cactus Cylindropuntia tunicata (Zumpango); (f) living specimen (Monte Albán); Photos: (e) Zahir Santillán, (f) John Kemner (modified)." figureDoi="http://doi.org/10.5281/zenodo.10611483" httpUri="https://zenodo.org/record/10611483/files/figure.png" pageId="3" pageNumber="156">Figs 1c</figureCitation>
,
<figureCitation id="1372F487FFC8BD68FD7DF9C49A487027" box="[753,782,1591,1617]" captionStart="FIGURE 3" captionStartId="4.[151,250,1895,1919]" captionTargetBox="[190,1397,237,1871]" captionTargetId="figure-62@4.[190,1397,237,1871]" captionTargetPageId="4" captionText="FIGURE 3. Vedalia sieboldii Mulsant, 1850; SEM illustrations: (a) antenna; (b) mouthparts; (c) pronotum; (d) head, prothorax and mesothorax, ventral; (e) scutellar shield; (f) meso- and metathorax, ventral; (g) tibial spurs on hind leg; (h) protarsal claw, male; (i) hind tarsal claw, female." figureDoi="http://doi.org/10.5281/zenodo.10611487" httpUri="https://zenodo.org/record/10611487/files/figure.png" pageId="3" pageNumber="156">3c</figureCitation>
); ventral antennal grooves deep, distinctly circular bent towards outer margin of an eye (
<figureCitation id="1372F487FFC8BD68FD8CF9A89B157003" box="[512,595,1627,1653]" captionStart="FIGURE 3" captionStartId="4.[151,250,1895,1919]" captionTargetBox="[190,1397,237,1871]" captionTargetId="figure-62@4.[190,1397,237,1871]" captionTargetPageId="4" captionText="FIGURE 3. Vedalia sieboldii Mulsant, 1850; SEM illustrations: (a) antenna; (b) mouthparts; (c) pronotum; (d) head, prothorax and mesothorax, ventral; (e) scutellar shield; (f) meso- and metathorax, ventral; (g) tibial spurs on hind leg; (h) protarsal claw, male; (i) hind tarsal claw, female." figureDoi="http://doi.org/10.5281/zenodo.10611487" httpUri="https://zenodo.org/record/10611487/files/figure.png" pageId="3" pageNumber="156">Fig. 3d</figureCitation>
). Eyes dorsally with inner orbits sub-parallel, coarsely facetted; interfacetal setae present, sparse; eye canthus large, reaching half width of an eye (
<figureCitation id="1372F487FFC8BD68FC4BF98C9D5C70EF" box="[967,1050,1663,1689]" captionStart="FIGURE 1" captionStartId="2.[151,250,1514,1538]" captionTargetBox="[158,1430,214,1489]" captionTargetId="figure-34@2.[158,1430,179,1489]" captionTargetPageId="2" captionText="FIGURE 1. Vedalia sieboldii Mulsant, 1850: habitus: (a) dorsal view; (b) lateral view; (c) frontal view; (d) head; (e) living specimen on cactus Cylindropuntia tunicata (Zumpango); (f) living specimen (Monte Albán); Photos: (e) Zahir Santillán, (f) John Kemner (modified)." figureDoi="http://doi.org/10.5281/zenodo.10611483" httpUri="https://zenodo.org/record/10611483/files/figure.png" pageId="3" pageNumber="156">Fig. 1d</figureCitation>
); interocular distance about 0.6 × head width across eyes. Antenna (
<figureCitation id="1372F487FFC8BD68FD82F9509B1B70CB" box="[526,605,1699,1725]" captionStart="FIGURE 3" captionStartId="4.[151,250,1895,1919]" captionTargetBox="[190,1397,237,1871]" captionTargetId="figure-62@4.[190,1397,237,1871]" captionTargetPageId="4" captionText="FIGURE 3. Vedalia sieboldii Mulsant, 1850; SEM illustrations: (a) antenna; (b) mouthparts; (c) pronotum; (d) head, prothorax and mesothorax, ventral; (e) scutellar shield; (f) meso- and metathorax, ventral; (g) tibial spurs on hind leg; (h) protarsal claw, male; (i) hind tarsal claw, female." figureDoi="http://doi.org/10.5281/zenodo.10611487" httpUri="https://zenodo.org/record/10611487/files/figure.png" pageId="3" pageNumber="156">Fig. 3a</figureCitation>
) 0.6 × as long as head capsule width, composed of 11 antennomeres; scape enlarged, distinctly roundly projected anteriorly; pedicel narrower than scape, barrel-shaped, about the same length as width; terminal antennomere enlarged. Antennal insertions partially visible from above. Anterior clypeal margin emarginate with anterolateral lobes roundly projected (
<figureCitation id="1372F487FFC8BD68FD78F8FC9A02715F" box="[756,836,1807,1833]" captionStart="FIGURE 1" captionStartId="2.[151,250,1514,1538]" captionTargetBox="[158,1430,214,1489]" captionTargetId="figure-34@2.[158,1430,179,1489]" captionTargetPageId="2" captionText="FIGURE 1. Vedalia sieboldii Mulsant, 1850: habitus: (a) dorsal view; (b) lateral view; (c) frontal view; (d) head; (e) living specimen on cactus Cylindropuntia tunicata (Zumpango); (f) living specimen (Monte Albán); Photos: (e) Zahir Santillán, (f) John Kemner (modified)." figureDoi="http://doi.org/10.5281/zenodo.10611483" httpUri="https://zenodo.org/record/10611483/files/figure.png" pageId="3" pageNumber="156">Fig. 1d</figureCitation>
). Labrum truncate at apex. Mandible bidentate at apex (
<figureCitation id="1372F487FFC8BD68FF13F8C099B4713B" box="[159,242,1843,1869]" captionStart="FIGURE 3" captionStartId="4.[151,250,1895,1919]" captionTargetBox="[190,1397,237,1871]" captionTargetId="figure-62@4.[190,1397,237,1871]" captionTargetPageId="4" captionText="FIGURE 3. Vedalia sieboldii Mulsant, 1850; SEM illustrations: (a) antenna; (b) mouthparts; (c) pronotum; (d) head, prothorax and mesothorax, ventral; (e) scutellar shield; (f) meso- and metathorax, ventral; (g) tibial spurs on hind leg; (h) protarsal claw, male; (i) hind tarsal claw, female." figureDoi="http://doi.org/10.5281/zenodo.10611487" httpUri="https://zenodo.org/record/10611487/files/figure.png" pageId="3" pageNumber="156">Fig. 3b</figureCitation>
). Maxilla (
<figureCitation id="1372F487FFC8BD68FEE6F8C098FC713B" box="[362,442,1843,1869]" captionStart="FIGURE 3" captionStartId="4.[151,250,1895,1919]" captionTargetBox="[190,1397,237,1871]" captionTargetId="figure-62@4.[190,1397,237,1871]" captionTargetPageId="4" captionText="FIGURE 3. Vedalia sieboldii Mulsant, 1850; SEM illustrations: (a) antenna; (b) mouthparts; (c) pronotum; (d) head, prothorax and mesothorax, ventral; (e) scutellar shield; (f) meso- and metathorax, ventral; (g) tibial spurs on hind leg; (h) protarsal claw, male; (i) hind tarsal claw, female." figureDoi="http://doi.org/10.5281/zenodo.10611487" httpUri="https://zenodo.org/record/10611487/files/figure.png" pageId="3" pageNumber="156">Fig. 3b</figureCitation>
) with cardo transverse; maxillary stipes deeply foveate to accommodate maxillary palp in repose, basistipes with distinct ridge; lacinia elongate, covered with moderately long setae; galea narrow, C-shaped, densely setose apically; maxillary palp composed of 4 palpomeres, palpomere 2 subtriangular, elongate, longer than palpomere 3; terminal palpomere distinctly securiform. Submentum short, transverse; mentum sub-quadrate with sides sub-parallel, anterior edge emarginate with large rounded lobes (
<figureCitation id="1372F487FFC8BD68FC36F8309D4971AB" box="[954,1039,1987,2013]" captionStart="FIGURE 3" captionStartId="4.[151,250,1895,1919]" captionTargetBox="[190,1397,237,1871]" captionTargetId="figure-62@4.[190,1397,237,1871]" captionTargetPageId="4" captionText="FIGURE 3. Vedalia sieboldii Mulsant, 1850; SEM illustrations: (a) antenna; (b) mouthparts; (c) pronotum; (d) head, prothorax and mesothorax, ventral; (e) scutellar shield; (f) meso- and metathorax, ventral; (g) tibial spurs on hind leg; (h) protarsal claw, male; (i) hind tarsal claw, female." figureDoi="http://doi.org/10.5281/zenodo.10611487" httpUri="https://zenodo.org/record/10611487/files/figure.png" pageId="3" pageNumber="156">Fig. 3b</figureCitation>
); prementum transverse, anteriorly truncate; labial palps separated by distance about equal to width of palpiger; palp composed of 3 palpomeres, apical palpomere sub-parallel, as long and as broad as penultimate one.
</paragraph>
<caption id="DF36B88AFFCFBD6FFF1BF89498AB71B1" ID-DOI="http://doi.org/10.5281/zenodo.10611487" ID-Zenodo-Dep="10611487" httpUri="https://zenodo.org/record/10611487/files/figure.png" pageId="4" pageNumber="157" startId="4.[151,250,1895,1919]" targetBox="[190,1397,237,1871]" targetPageId="4" targetType="figure">
<paragraph id="8BF6E802FFCFBD6FFF1BF89498AB71B1" blockId="4.[151,1436,1895,1991]" pageId="4" pageNumber="157">
<emphasis id="B93D3410FFCFBD6FFF1BF89498547109" bold="true" box="[151,274,1895,1919]" pageId="4" pageNumber="157">FIGURE 3.</emphasis>
<taxonomicName id="4C499381FFCFBD6FFE94F8949B197109" authority="Mulsant, 1850" authorityName="Mulsant. Since" authorityYear="1850" box="[280,607,1895,1919]" class="Insecta" family="Coccinellidae" genus="Vedalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="157" phylum="Arthropoda" rank="species" species="sieboldii">
<emphasis id="B93D3410FFCFBD6FFE94F89498827109" box="[280,452,1895,1919]" italics="true" pageId="4" pageNumber="157">Vedalia sieboldii</emphasis>
<bibRefCitation id="EFD895F3FFCFBD6FFE46F8949B197109" author="Mulsant, E." box="[458,607,1895,1919]" pageId="4" pageNumber="157" pagination="1 - 1104" refId="ref6251" refString="Mulsant, E. (1850) Species des Coleopteres trimeres securipalpes. Annales des Sciences Physiques et Naturelles, d'Agriculture et d'Industrie, Lyon, Series 2, 2, 1 - 1104. https: // doi. org / 10.5962 / bhl. title. 8953" type="journal article" year="1850">Mulsant, 1850</bibRefCitation>
</taxonomicName>
; SEM illustrations: (
<emphasis id="B93D3410FFCFBD6FFCA3F8949A7A7109" bold="true" box="[815,828,1895,1919]" pageId="4" pageNumber="157">a</emphasis>
) antenna; (
<emphasis id="B93D3410FFCFBD6FFC21F8949AFD7109" bold="true" box="[941,955,1895,1919]" pageId="4" pageNumber="157">b</emphasis>
) mouthparts; (
<emphasis id="B93D3410FFCFBD6FFBDDF8949D1A7109" bold="true" box="[1105,1116,1895,1919]" pageId="4" pageNumber="157">c</emphasis>
) pronotum; (
<emphasis id="B93D3410FFCFBD6FFB6EF8949DB67109" bold="true" box="[1250,1264,1895,1919]" pageId="4" pageNumber="157">d</emphasis>
) head, prothorax and mesothorax, ventral; (
<emphasis id="B93D3410FFCFBD6FFE2EF87898EB71D5" bold="true" box="[418,429,1931,1955]" pageId="4" pageNumber="157">e</emphasis>
) scutellar shield; (
<emphasis id="B93D3410FFCFBD6FFDE7F8789B3571D5" bold="true" box="[619,627,1931,1955]" pageId="4" pageNumber="157">f</emphasis>
) meso- and metathorax, ventral; (
<emphasis id="B93D3410FFCFBD6FFC42F8789A9D71D5" bold="true" box="[974,987,1931,1955]" pageId="4" pageNumber="157">g</emphasis>
) tibial spurs on hind leg; (
<emphasis id="B93D3410FFCFBD6FFB66F8789DBE71D5" bold="true" box="[1258,1272,1931,1955]" pageId="4" pageNumber="157">h</emphasis>
) protarsal claw, male; (
<emphasis id="B93D3410FFCFBD6FFF52F85C99A371B1" bold="true" box="[222,229,1967,1991]" pageId="4" pageNumber="157">i</emphasis>
) hind tarsal claw, female.
</paragraph>
</caption>
<caption id="DF36B88AFFCEBD6EFF1BF89098017191" ID-DOI="http://doi.org/10.5281/zenodo.10611489" ID-Zenodo-Dep="10611489" httpUri="https://zenodo.org/record/10611489/files/figure.png" pageId="5" pageNumber="158" startId="5.[151,250,1891,1915]" targetBox="[221,1366,181,1866]" targetPageId="5" targetType="figure">
<paragraph id="8BF6E802FFCEBD6EFF1BF89098017191" blockId="5.[151,1436,1891,2023]" pageId="5" pageNumber="158">
<emphasis id="B93D3410FFCEBD6EFF1BF8909850710D" bold="true" box="[151,278,1891,1915]" pageId="5" pageNumber="158">FIGURE 4.</emphasis>
<taxonomicName id="4C499381FFCEBD6EFE93F8909B36710D" authority="Mulsant, 1850" authorityName="Mulsant. Since" authorityYear="1850" box="[287,624,1891,1915]" class="Insecta" family="Coccinellidae" genus="Vedalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="158" phylum="Arthropoda" rank="species" species="sieboldii">
<emphasis id="B93D3410FFCEBD6EFE93F8909888710D" box="[287,462,1891,1915]" italics="true" pageId="5" pageNumber="158">Vedalia sieboldii</emphasis>
<bibRefCitation id="EFD895F3FFCEBD6EFE5BF8909B36710D" author="Mulsant, E." box="[471,624,1891,1915]" pageId="5" pageNumber="158" pagination="1 - 1104" refId="ref6251" refString="Mulsant, E. (1850) Species des Coleopteres trimeres securipalpes. Annales des Sciences Physiques et Naturelles, d'Agriculture et d'Industrie, Lyon, Series 2, 2, 1 - 1104. https: // doi. org / 10.5962 / bhl. title. 8953" type="journal article" year="1850">Mulsant, 1850</bibRefCitation>
</taxonomicName>
; abdomen and genitalia details: (
<emphasis id="B93D3410FFCEBD6EFC47F8909A9E710D" bold="true" box="[971,984,1891,1915]" pageId="5" pageNumber="158">a</emphasis>
) abdomen, male; (
<emphasis id="B93D3410FFCEBD6EFB13F8909DEB710D" bold="true" box="[1183,1197,1891,1915]" pageId="5" pageNumber="158">b</emphasis>
) abdomen, female; (
<emphasis id="B93D3410FFCEBD6EFA05F8909CD2710D" bold="true" box="[1417,1428,1891,1915]" pageId="5" pageNumber="158">c</emphasis>
) tergite VIII, male; (
<emphasis id="B93D3410FFCEBD6EFEECF874982871E9" bold="true" box="[352,366,1927,1951]" pageId="5" pageNumber="158">d</emphasis>
) ventrite 6, male; (
<emphasis id="B93D3410FFCEBD6EFDB8F8749B7971E9" bold="true" box="[564,575,1927,1951]" pageId="5" pageNumber="158">e</emphasis>
) tergite VIII, female; (
<emphasis id="B93D3410FFCEBD6EFCA7F8749A7571E9" bold="true" box="[811,819,1927,1951]" pageId="5" pageNumber="158">f</emphasis>
) ventrite 6, female; (
<emphasis id="B93D3410FFCEBD6EFB80F8749D5F71E9" bold="true" box="[1036,1049,1927,1951]" pageId="5" pageNumber="158">g</emphasis>
) segments IX and X, male; (
<emphasis id="B93D3410FFCEBD6EFACFF8749C1771E9" bold="true" box="[1347,1361,1927,1951]" pageId="5" pageNumber="158">h</emphasis>
) penis, lateral; (
<emphasis id="B93D3410FFCEBD6EFF60F85899B571B5" bold="true" box="[236,243,1963,1987]" pageId="5" pageNumber="158">i</emphasis>
) penis tip, lateral; (
<emphasis id="B93D3410FFCEBD6EFE35F858988771B5" bold="true" box="[441,449,1963,1987]" pageId="5" pageNumber="158">j</emphasis>
) penis tip, inner; (
<emphasis id="B93D3410FFCEBD6EFDF0F8589BCC71B5" bold="true" box="[636,650,1963,1987]" pageId="5" pageNumber="158">k</emphasis>
) tegmen, inner; (
<emphasis id="B93D3410FFCEBD6EFCB6F8589A0771B5" bold="true" box="[826,833,1963,1987]" pageId="5" pageNumber="158">l</emphasis>
) tegmen, lateral; (
<emphasis id="B93D3410FFCEBD6EFC77F8589D5671B5" bold="true" box="[1019,1040,1963,1987]" pageId="5" pageNumber="158">m</emphasis>
) female genitalia; (
<emphasis id="B93D3410FFCEBD6EFB5AF8589DA271B5" bold="true" box="[1238,1252,1963,1987]" pageId="5" pageNumber="158">n</emphasis>
) spermatheca; (
<emphasis id="B93D3410FFCEBD6EFA0BF8589CD271B5" bold="true" box="[1415,1428,1963,1987]" pageId="5" pageNumber="158">o</emphasis>
) apices of coxites.
</paragraph>
</caption>
<paragraph id="8BF6E802FFCDBD6DFF4BFF649C6077FF" blockId="6.[151,1437,151,1041]" pageId="6" pageNumber="159">
<emphasis id="B93D3410FFCDBD6DFF4BFF64980F76C7" bold="true" box="[199,329,151,177]" pageId="6" pageNumber="159">Prothorax.</emphasis>
Pronotum (
<figureCitation id="1372F487FFCDBD6DFE57FF649B7676C7" box="[475,560,151,177]" captionStart="FIGURE 3" captionStartId="4.[151,250,1895,1919]" captionTargetBox="[190,1397,237,1871]" captionTargetId="figure-62@4.[190,1397,237,1871]" captionTargetPageId="4" captionText="FIGURE 3. Vedalia sieboldii Mulsant, 1850; SEM illustrations: (a) antenna; (b) mouthparts; (c) pronotum; (d) head, prothorax and mesothorax, ventral; (e) scutellar shield; (f) meso- and metathorax, ventral; (g) tibial spurs on hind leg; (h) protarsal claw, male; (i) hind tarsal claw, female." figureDoi="http://doi.org/10.5281/zenodo.10611487" httpUri="https://zenodo.org/record/10611487/files/figure.png" pageId="6" pageNumber="159">Fig. 3c</figureCitation>
) with anterior corners rounded, slightly produced anteriorly, not swollen with regular border; anterior and posterior margin without border; lateral margin with entire border. Prothoracic hypomeron smooth (
<figureCitation id="1372F487FFCDBD6DFE0CFF2C9895768F" box="[384,467,223,249]" captionStart="FIGURE 3" captionStartId="4.[151,250,1895,1919]" captionTargetBox="[190,1397,237,1871]" captionTargetId="figure-62@4.[190,1397,237,1871]" captionTargetPageId="4" captionText="FIGURE 3. Vedalia sieboldii Mulsant, 1850; SEM illustrations: (a) antenna; (b) mouthparts; (c) pronotum; (d) head, prothorax and mesothorax, ventral; (e) scutellar shield; (f) meso- and metathorax, ventral; (g) tibial spurs on hind leg; (h) protarsal claw, male; (i) hind tarsal claw, female." figureDoi="http://doi.org/10.5281/zenodo.10611487" httpUri="https://zenodo.org/record/10611487/files/figure.png" pageId="6" pageNumber="159">Fig. 3d</figureCitation>
), densely covered with hairs. Prosternum in front of coxa about 0.3 × as long as coxal longitudinal diameter at the same position; anterior pronotal margin straight, with visible bordering line; notosternal suture visible (
<figureCitation id="1372F487FFCDBD6DFEB1FED498D77737" box="[317,401,295,321]" captionStart="FIGURE 3" captionStartId="4.[151,250,1895,1919]" captionTargetBox="[190,1397,237,1871]" captionTargetId="figure-62@4.[190,1397,237,1871]" captionTargetPageId="4" captionText="FIGURE 3. Vedalia sieboldii Mulsant, 1850; SEM illustrations: (a) antenna; (b) mouthparts; (c) pronotum; (d) head, prothorax and mesothorax, ventral; (e) scutellar shield; (f) meso- and metathorax, ventral; (g) tibial spurs on hind leg; (h) protarsal claw, male; (i) hind tarsal claw, female." figureDoi="http://doi.org/10.5281/zenodo.10611487" httpUri="https://zenodo.org/record/10611487/files/figure.png" pageId="6" pageNumber="159">Fig. 3b</figureCitation>
); procoxal cavity distinctly transverse with lateral slit, with complete bordering line close to the cavity; prosternal process narrow, about 0.2 × as wide as corresponding coxal width, with sides parallel and apex rounded, its surface without carinae, with a pair of shallow, longitudinal grooves in apical part (
<figureCitation id="1372F487FFCDBD6DFB49FE9C9C5F77FF" box="[1221,1305,367,393]" captionStart="FIGURE 3" captionStartId="4.[151,250,1895,1919]" captionTargetBox="[190,1397,237,1871]" captionTargetId="figure-62@4.[190,1397,237,1871]" captionTargetPageId="4" captionText="FIGURE 3. Vedalia sieboldii Mulsant, 1850; SEM illustrations: (a) antenna; (b) mouthparts; (c) pronotum; (d) head, prothorax and mesothorax, ventral; (e) scutellar shield; (f) meso- and metathorax, ventral; (g) tibial spurs on hind leg; (h) protarsal claw, male; (i) hind tarsal claw, female." figureDoi="http://doi.org/10.5281/zenodo.10611487" httpUri="https://zenodo.org/record/10611487/files/figure.png" pageId="6" pageNumber="159">Fig. 3d</figureCitation>
).
</paragraph>
<paragraph id="8BF6E802FFCDBD6DFF4BFE609ACE74DF" blockId="6.[151,1437,151,1041]" pageId="6" pageNumber="159">
<emphasis id="B93D3410FFCDBD6DFF4BFE60981877DB" bold="true" box="[199,350,403,429]" pageId="6" pageNumber="159">Pterothorax.</emphasis>
Mesoventrite (
<figureCitation id="1372F487FFCDBD6DFD88FE609B1277DB" box="[516,596,403,429]" captionStart="FIGURE 3" captionStartId="4.[151,250,1895,1919]" captionTargetBox="[190,1397,237,1871]" captionTargetId="figure-62@4.[190,1397,237,1871]" captionTargetPageId="4" captionText="FIGURE 3. Vedalia sieboldii Mulsant, 1850; SEM illustrations: (a) antenna; (b) mouthparts; (c) pronotum; (d) head, prothorax and mesothorax, ventral; (e) scutellar shield; (f) meso- and metathorax, ventral; (g) tibial spurs on hind leg; (h) protarsal claw, male; (i) hind tarsal claw, female." figureDoi="http://doi.org/10.5281/zenodo.10611487" httpUri="https://zenodo.org/record/10611487/files/figure.png" pageId="6" pageNumber="159">Fig. 3d</figureCitation>
) transverse, its anterior edge straight with complete raised border; its surface with shallow depression in middle. Mesoventral process (
<figureCitation id="1372F487FFCDBD6DFC92FE449A2977A7" box="[798,879,439,465]" captionStart="FIGURE 3" captionStartId="4.[151,250,1895,1919]" captionTargetBox="[190,1397,237,1871]" captionTargetId="figure-62@4.[190,1397,237,1871]" captionTargetPageId="4" captionText="FIGURE 3. Vedalia sieboldii Mulsant, 1850; SEM illustrations: (a) antenna; (b) mouthparts; (c) pronotum; (d) head, prothorax and mesothorax, ventral; (e) scutellar shield; (f) meso- and metathorax, ventral; (g) tibial spurs on hind leg; (h) protarsal claw, male; (i) hind tarsal claw, female." figureDoi="http://doi.org/10.5281/zenodo.10611487" httpUri="https://zenodo.org/record/10611487/files/figure.png" pageId="6" pageNumber="159">Fig. 3d</figureCitation>
) in middle about 0.5 × as broad as corresponding coxal diameter; meso-metaventral articulation with suture visible; junction straight. Scutellar shield triangular, large (
<figureCitation id="1372F487FFCDBD6DFF13FE0C99A9746F" box="[159,239,511,537]" captionStart="FIGURE 3" captionStartId="4.[151,250,1895,1919]" captionTargetBox="[190,1397,237,1871]" captionTargetId="figure-62@4.[190,1397,237,1871]" captionTargetPageId="4" captionText="FIGURE 3. Vedalia sieboldii Mulsant, 1850; SEM illustrations: (a) antenna; (b) mouthparts; (c) pronotum; (d) head, prothorax and mesothorax, ventral; (e) scutellar shield; (f) meso- and metathorax, ventral; (g) tibial spurs on hind leg; (h) protarsal claw, male; (i) hind tarsal claw, female." figureDoi="http://doi.org/10.5281/zenodo.10611487" httpUri="https://zenodo.org/record/10611487/files/figure.png" pageId="6" pageNumber="159">Fig. 3e</figureCitation>
), its surface punctate and densely setose. Elytra with sides rounded; lateral margins narrow but visible from above throughout; humeral calli present; surface with single size punctures; elytral epipleuron wide, about 2.0 × as wide as corresponding metepisternum, apically incomplete, not foveate, inner margin with border line reaching level of ventrite 2 (
<figureCitation id="1372F487FFCDBD6DFEF1FD98989574F3" box="[381,467,619,645]" captionStart="FIGURE 2" captionStartId="3.[151,250,1457,1481]" captionTargetBox="[190,1397,453,1434]" captionTargetId="figure-129@3.[190,1397,453,1434]" captionTargetPageId="3" captionText="FIGURE 2. Vedalia sieboldii Mulsant, 1850; SEM illustrations: (a) habitus, dorsal; (b) hind leg; (c) habitus, ventral; (d) abdomen." figureDoi="http://doi.org/10.5281/zenodo.10611485" httpUri="https://zenodo.org/record/10611485/files/figure.png" pageId="6" pageNumber="159">Fig. 2c</figureCitation>
). Metaventrite with discrimen visible; metaventral postcoxal lines joined at middle forming arcuate line, laterally slightly recurved, complete (
<figureCitation id="1372F487FFCDBD6DFCA0FD7C9A3D74DF" box="[812,891,655,681]" captionStart="FIGURE 3" captionStartId="4.[151,250,1895,1919]" captionTargetBox="[190,1397,237,1871]" captionTargetId="figure-62@4.[190,1397,237,1871]" captionTargetPageId="4" captionText="FIGURE 3. Vedalia sieboldii Mulsant, 1850; SEM illustrations: (a) antenna; (b) mouthparts; (c) pronotum; (d) head, prothorax and mesothorax, ventral; (e) scutellar shield; (f) meso- and metathorax, ventral; (g) tibial spurs on hind leg; (h) protarsal claw, male; (i) hind tarsal claw, female." figureDoi="http://doi.org/10.5281/zenodo.10611487" httpUri="https://zenodo.org/record/10611487/files/figure.png" pageId="6" pageNumber="159">Fig. 3f</figureCitation>
).
</paragraph>
<paragraph id="8BF6E802FFCDBD6DFF4BFD409B70754F" blockId="6.[151,1437,151,1041]" pageId="6" pageNumber="159">
<emphasis id="B93D3410FFCDBD6DFF4BFD40984074BB" bold="true" box="[199,262,691,717]" pageId="6" pageNumber="159">Legs.</emphasis>
Trochanters roundly produced (
<figureCitation id="1372F487FFCDBD6DFDF9FD409B8974BB" box="[629,719,691,717]" captionStart="FIGURE 2" captionStartId="3.[151,250,1457,1481]" captionTargetBox="[190,1397,453,1434]" captionTargetId="figure-129@3.[190,1397,453,1434]" captionTargetPageId="3" captionText="FIGURE 2. Vedalia sieboldii Mulsant, 1850; SEM illustrations: (a) habitus, dorsal; (b) hind leg; (c) habitus, ventral; (d) abdomen." figureDoi="http://doi.org/10.5281/zenodo.10611485" httpUri="https://zenodo.org/record/10611485/files/figure.png" pageId="6" pageNumber="159">Figs 2b</figureCitation>
;
<figureCitation id="1372F487FFCDBD6DFD51FD409BB174BB" box="[733,759,691,717]" captionStart="FIGURE 3" captionStartId="4.[151,250,1895,1919]" captionTargetBox="[190,1397,237,1871]" captionTargetId="figure-62@4.[190,1397,237,1871]" captionTargetPageId="4" captionText="FIGURE 3. Vedalia sieboldii Mulsant, 1850; SEM illustrations: (a) antenna; (b) mouthparts; (c) pronotum; (d) head, prothorax and mesothorax, ventral; (e) scutellar shield; (f) meso- and metathorax, ventral; (g) tibial spurs on hind leg; (h) protarsal claw, male; (i) hind tarsal claw, female." figureDoi="http://doi.org/10.5281/zenodo.10611487" httpUri="https://zenodo.org/record/10611487/files/figure.png" pageId="6" pageNumber="159">3f</figureCitation>
); femora stout, excavated to receive tibiae; tibiae distinctly broadened, with outer margin smooth (
<figureCitation id="1372F487FFCDBD6DFDDDFD249BE07487" box="[593,678,727,753]" captionStart="FIGURE 2" captionStartId="3.[151,250,1457,1481]" captionTargetBox="[190,1397,453,1434]" captionTargetId="figure-129@3.[190,1397,453,1434]" captionTargetPageId="3" captionText="FIGURE 2. Vedalia sieboldii Mulsant, 1850; SEM illustrations: (a) habitus, dorsal; (b) hind leg; (c) habitus, ventral; (d) abdomen." figureDoi="http://doi.org/10.5281/zenodo.10611485" httpUri="https://zenodo.org/record/10611485/files/figure.png" pageId="6" pageNumber="159">Fig. 2b</figureCitation>
); tibial apices of mid and hind leg with double spurs in both sexes (
<figureCitation id="1372F487FFCDBD6DFF13FD0899B57563" box="[159,243,763,789]" captionStart="FIGURE 3" captionStartId="4.[151,250,1895,1919]" captionTargetBox="[190,1397,237,1871]" captionTargetId="figure-62@4.[190,1397,237,1871]" captionTargetPageId="4" captionText="FIGURE 3. Vedalia sieboldii Mulsant, 1850; SEM illustrations: (a) antenna; (b) mouthparts; (c) pronotum; (d) head, prothorax and mesothorax, ventral; (e) scutellar shield; (f) meso- and metathorax, ventral; (g) tibial spurs on hind leg; (h) protarsal claw, male; (i) hind tarsal claw, female." figureDoi="http://doi.org/10.5281/zenodo.10611487" httpUri="https://zenodo.org/record/10611487/files/figure.png" pageId="6" pageNumber="159">
Fig.
<quantity id="4CB145E7FFCDBD6DFF59FD0899B57563" box="[213,243,763,789]" metricMagnitude="-3" metricUnit="kg" metricValue="3.0" pageId="6" pageNumber="159" unit="g" value="3.0">3g</quantity>
</figureCitation>
). Tarsi with four tarsomeres, pro- and mid-tarsal claws in males cleft (
<figureCitation id="1372F487FFCDBD6DFB9EFD089D207563" box="[1042,1126,763,789]" captionStart="FIGURE 3" captionStartId="4.[151,250,1895,1919]" captionTargetBox="[190,1397,237,1871]" captionTargetId="figure-62@4.[190,1397,237,1871]" captionTargetPageId="4" captionText="FIGURE 3. Vedalia sieboldii Mulsant, 1850; SEM illustrations: (a) antenna; (b) mouthparts; (c) pronotum; (d) head, prothorax and mesothorax, ventral; (e) scutellar shield; (f) meso- and metathorax, ventral; (g) tibial spurs on hind leg; (h) protarsal claw, male; (i) hind tarsal claw, female." figureDoi="http://doi.org/10.5281/zenodo.10611487" httpUri="https://zenodo.org/record/10611487/files/figure.png" pageId="6" pageNumber="159">Fig. 3h</figureCitation>
), hind tarsal claws bifid; in female all tarsal claws bifid (
<figureCitation id="1372F487FFCDBD6DFE50FCEC9B6E754F" box="[476,552,799,825]" captionStart="FIGURE 3" captionStartId="4.[151,250,1895,1919]" captionTargetBox="[190,1397,237,1871]" captionTargetId="figure-62@4.[190,1397,237,1871]" captionTargetPageId="4" captionText="FIGURE 3. Vedalia sieboldii Mulsant, 1850; SEM illustrations: (a) antenna; (b) mouthparts; (c) pronotum; (d) head, prothorax and mesothorax, ventral; (e) scutellar shield; (f) meso- and metathorax, ventral; (g) tibial spurs on hind leg; (h) protarsal claw, male; (i) hind tarsal claw, female." figureDoi="http://doi.org/10.5281/zenodo.10611487" httpUri="https://zenodo.org/record/10611487/files/figure.png" pageId="6" pageNumber="159">Fig. 3i</figureCitation>
).
</paragraph>
<paragraph id="8BF6E802FFCDBD6DFF4BFCB199B0759B" blockId="6.[151,1437,151,1041]" pageId="6" pageNumber="159">
<emphasis id="B93D3410FFCDBD6DFF4BFCB1987A752A" bold="true" box="[199,316,834,860]" pageId="6" pageNumber="159">Abdomen</emphasis>
(
<figureCitation id="1372F487FFCDBD6DFEC3FCB098ED752B" box="[335,427,835,861]" captionStart="FIGURE 2" captionStartId="3.[151,250,1457,1481]" captionTargetBox="[190,1397,453,1434]" captionTargetId="figure-129@3.[190,1397,453,1434]" captionTargetPageId="3" captionText="FIGURE 2. Vedalia sieboldii Mulsant, 1850; SEM illustrations: (a) habitus, dorsal; (b) hind leg; (c) habitus, ventral; (d) abdomen." figureDoi="http://doi.org/10.5281/zenodo.10611485" httpUri="https://zenodo.org/record/10611485/files/figure.png" pageId="6" pageNumber="159">Figs 2d</figureCitation>
;
<figureCitation id="1372F487FFCDBD6DFE36FCB098BE752B" box="[442,504,835,861]" captionStart="FIGURE 4" captionStartId="5.[151,250,1891,1915]" captionTargetBox="[221,1366,181,1866]" captionTargetId="figure-17@5.[221,1366,181,1866]" captionTargetPageId="5" captionText="FIGURE 4. Vedalia sieboldii Mulsant, 1850; abdomen and genitalia details: (a) abdomen, male; (b) abdomen, female; (c) tergite VIII, male; (d) ventrite 6, male; (e) tergite VIII, female; (f) ventrite 6, female; (g) segments IX and X, male; (h) penis, lateral; (i) penis tip, lateral; (j) penis tip, inner; (k) tegmen, inner; (l) tegmen, lateral; (m) female genitalia; (n) spermatheca; (o) apices of coxites." figureDoi="http://doi.org/10.5281/zenodo.10611489" httpUri="https://zenodo.org/record/10611489/files/figure.png" pageId="6" pageNumber="159">4a, b</figureCitation>
). Abdomen with six ventrites in both sexes (
<figureCitation id="1372F487FFCDBD6DFB86FCB09D11752B" box="[1034,1111,835,861]" captionStart="FIGURE 2" captionStartId="3.[151,250,1457,1481]" captionTargetBox="[190,1397,453,1434]" captionTargetId="figure-129@3.[190,1397,453,1434]" captionTargetPageId="3" captionText="FIGURE 2. Vedalia sieboldii Mulsant, 1850; SEM illustrations: (a) habitus, dorsal; (b) hind leg; (c) habitus, ventral; (d) abdomen." figureDoi="http://doi.org/10.5281/zenodo.10611485" httpUri="https://zenodo.org/record/10611485/files/figure.png" pageId="6" pageNumber="159">Fig.2d</figureCitation>
); abdominal postcoxal lines separate medially, incomplete laterally, posteriorly not reaching hind edge of ventrite 1, recurved; hind margin of ventrite 5 straight in male (
<figureCitation id="1372F487FFCDBD6DFE43FC789B6575D3" box="[463,547,907,933]" captionStart="FIGURE 4" captionStartId="5.[151,250,1891,1915]" captionTargetBox="[221,1366,181,1866]" captionTargetId="figure-17@5.[221,1366,181,1866]" captionTargetPageId="5" captionText="FIGURE 4. Vedalia sieboldii Mulsant, 1850; abdomen and genitalia details: (a) abdomen, male; (b) abdomen, female; (c) tergite VIII, male; (d) ventrite 6, male; (e) tergite VIII, female; (f) ventrite 6, female; (g) segments IX and X, male; (h) penis, lateral; (i) penis tip, lateral; (j) penis tip, inner; (k) tegmen, inner; (l) tegmen, lateral; (m) female genitalia; (n) spermatheca; (o) apices of coxites." figureDoi="http://doi.org/10.5281/zenodo.10611489" httpUri="https://zenodo.org/record/10611489/files/figure.png" pageId="6" pageNumber="159">Fig. 4a</figureCitation>
), slightly emarginate in female (
<figureCitation id="1372F487FFCDBD6DFC1AFC789AAA75D3" box="[918,1004,907,933]" captionStart="FIGURE 4" captionStartId="5.[151,250,1891,1915]" captionTargetBox="[221,1366,181,1866]" captionTargetId="figure-17@5.[221,1366,181,1866]" captionTargetPageId="5" captionText="FIGURE 4. Vedalia sieboldii Mulsant, 1850; abdomen and genitalia details: (a) abdomen, male; (b) abdomen, female; (c) tergite VIII, male; (d) ventrite 6, male; (e) tergite VIII, female; (f) ventrite 6, female; (g) segments IX and X, male; (h) penis, lateral; (i) penis tip, lateral; (j) penis tip, inner; (k) tegmen, inner; (l) tegmen, lateral; (m) female genitalia; (n) spermatheca; (o) apices of coxites." figureDoi="http://doi.org/10.5281/zenodo.10611489" httpUri="https://zenodo.org/record/10611489/files/figure.png" pageId="6" pageNumber="159">Fig. 4b</figureCitation>
); in male ventrite 6 (
<figureCitation id="1372F487FFCDBD6DFB52FC789C7775D3" box="[1246,1329,907,933]" captionStart="FIGURE 4" captionStartId="5.[151,250,1891,1915]" captionTargetBox="[221,1366,181,1866]" captionTargetId="figure-17@5.[221,1366,181,1866]" captionTargetPageId="5" captionText="FIGURE 4. Vedalia sieboldii Mulsant, 1850; abdomen and genitalia details: (a) abdomen, male; (b) abdomen, female; (c) tergite VIII, male; (d) ventrite 6, male; (e) tergite VIII, female; (f) ventrite 6, female; (g) segments IX and X, male; (h) penis, lateral; (i) penis tip, lateral; (j) penis tip, inner; (k) tegmen, inner; (l) tegmen, lateral; (m) female genitalia; (n) spermatheca; (o) apices of coxites." figureDoi="http://doi.org/10.5281/zenodo.10611489" httpUri="https://zenodo.org/record/10611489/files/figure.png" pageId="6" pageNumber="159">Fig. 4d</figureCitation>
) truncate apically, tergite VIII (
<figureCitation id="1372F487FFCDBD6DFE1CFC5C98A575BF" box="[400,483,943,969]" captionStart="FIGURE 4" captionStartId="5.[151,250,1891,1915]" captionTargetBox="[221,1366,181,1866]" captionTargetId="figure-17@5.[221,1366,181,1866]" captionTargetPageId="5" captionText="FIGURE 4. Vedalia sieboldii Mulsant, 1850; abdomen and genitalia details: (a) abdomen, male; (b) abdomen, female; (c) tergite VIII, male; (d) ventrite 6, male; (e) tergite VIII, female; (f) ventrite 6, female; (g) segments IX and X, male; (h) penis, lateral; (i) penis tip, lateral; (j) penis tip, inner; (k) tegmen, inner; (l) tegmen, lateral; (m) female genitalia; (n) spermatheca; (o) apices of coxites." figureDoi="http://doi.org/10.5281/zenodo.10611489" httpUri="https://zenodo.org/record/10611489/files/figure.png" pageId="6" pageNumber="159">Fig. 4c</figureCitation>
) rounded apically; in female ventrite 6 (
<figureCitation id="1372F487FFCDBD6DFC3BFC5C9D4175BF" box="[951,1031,943,969]" captionStart="FIGURE 4" captionStartId="5.[151,250,1891,1915]" captionTargetBox="[221,1366,181,1866]" captionTargetId="figure-17@5.[221,1366,181,1866]" captionTargetPageId="5" captionText="FIGURE 4. Vedalia sieboldii Mulsant, 1850; abdomen and genitalia details: (a) abdomen, male; (b) abdomen, female; (c) tergite VIII, male; (d) ventrite 6, male; (e) tergite VIII, female; (f) ventrite 6, female; (g) segments IX and X, male; (h) penis, lateral; (i) penis tip, lateral; (j) penis tip, inner; (k) tegmen, inner; (l) tegmen, lateral; (m) female genitalia; (n) spermatheca; (o) apices of coxites." figureDoi="http://doi.org/10.5281/zenodo.10611489" httpUri="https://zenodo.org/record/10611489/files/figure.png" pageId="6" pageNumber="159">Fig. 4f</figureCitation>
) and tergite VIII (
<figureCitation id="1372F487FFCDBD6DFB50FC5C9C6975BF" box="[1244,1327,943,969]" captionStart="FIGURE 4" captionStartId="5.[151,250,1891,1915]" captionTargetBox="[221,1366,181,1866]" captionTargetId="figure-17@5.[221,1366,181,1866]" captionTargetPageId="5" captionText="FIGURE 4. Vedalia sieboldii Mulsant, 1850; abdomen and genitalia details: (a) abdomen, male; (b) abdomen, female; (c) tergite VIII, male; (d) ventrite 6, male; (e) tergite VIII, female; (f) ventrite 6, female; (g) segments IX and X, male; (h) penis, lateral; (i) penis tip, lateral; (j) penis tip, inner; (k) tegmen, inner; (l) tegmen, lateral; (m) female genitalia; (n) spermatheca; (o) apices of coxites." figureDoi="http://doi.org/10.5281/zenodo.10611489" httpUri="https://zenodo.org/record/10611489/files/figure.png" pageId="6" pageNumber="159">Fig. 4e</figureCitation>
) rounded apically.
</paragraph>
</subSubSection>
<subSubSection id="C353BB89FFCDBD6DFF4BFC0598837267" box="[199,453,1014,1041]" pageId="6" pageNumber="159" type="distribution">
<paragraph id="8BF6E802FFCDBD6DFF4BFC0598837267" blockId="6.[151,1437,151,1041]" box="[199,453,1014,1041]" pageId="6" pageNumber="159">
<emphasis id="B93D3410FFCDBD6DFF4BFC0598277266" bold="true" box="[199,353,1014,1040]" pageId="6" pageNumber="159">Distribution.</emphasis>
<collectingCountry id="F35EA892FFCDBD6DFEEBFC0498867267" box="[359,448,1015,1041]" name="Mexico" pageId="6" pageNumber="159">Mexico</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>