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<mods:title id="3C9B5B474BAFD3CF4B135504E33C15C6">New species and new records of South and North American tiger beetles (Coleoptera: Cicindelidae)</mods:title>
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<paragraph id="8BDB8507C16E8613D9BD58C3275DF923" blockId="1.[812,1399,1750,2071]" box="[930,1280,1750,1774]" pageId="1" pageNumber="150">
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<taxonomicName id="4C64FE84C16E8613D9BD58C3275DF923" ID-CoL="6NWYB" authority="Sumlin, 1993" authorityName="Sumlin" authorityYear="1993" box="[930,1280,1750,1774]" class="Insecta" family="Carabidae" genus="Langea" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="150" phylum="Arthropoda" rank="species" species="mellicollis">
<emphasis id="B9105915C16E8613D9BD58C32635F923" box="[930,1128,1750,1774]" italics="true" pageId="1" pageNumber="150">Langea mellicollis</emphasis>
<bibRefCitation id="EFF5F8F6C16E8613DE6F58C2275DF923" author="Sumlin W. D." box="[1136,1280,1750,1774]" pageId="1" pageNumber="150" pagination="1 - 12" refId="ref5098" refString="Sumlin W. D. 1993. Studies on the Neotropical Cicindelidae III: A review of the genus Langea (Coleoptera) // Cicindela. Vol. 25. Nos 1 - 2. P. 1 - 12." type="book chapter" year="1993">Sumlin, 1993</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37ED68CC16E8613DE0058E026D9F8C1" box="[1055,1156,1780,1804]" pageId="1" pageNumber="150" type="description">
<paragraph id="8BDB8507C16E8613DE0058E026D9F8C1" blockId="1.[812,1399,1750,2071]" box="[1055,1156,1780,1804]" pageId="1" pageNumber="150">
<figureCitation id="135F9982C16E8613DE0058E026DDF8C1" box="[1055,1152,1780,1804]" captionStart="Figs 14" captionStartId="2.[226,263,2017,2036]" captionTargetBox="[189,1399,629,1995]" captionTargetId="figure-268@2.[189,1399,578,2001]" captionTargetPageId="2" captionText="Figs 14. Langea mellicollis, male: 1 — habitus, dorsal view; 2 — labrum; 3 — left fore-tarsus; 4 — aedeagus, left lateral view. Рис. 14. Langea mellicollis, самец: 1 — внеШний вид, сверху; 2 — верхнЯЯ губа; 3 — леваЯ переднЯЯ лапка; 4 — Эдеагус, слева." figureDoi="http://doi.org/10.5281/zenodo.13177538" httpUri="https://zenodo.org/record/13177538/files/figure.png" pageId="1" pageNumber="150">Figs 14</figureCitation>
.
</paragraph>
</subSubSection>
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<paragraph id="8BDB8507C16E8613D94D590D2656F891" blockId="1.[812,1399,1750,2071]" pageId="1" pageNumber="150">
<materialsCitation id="3B0C8F5AC16E8613D94D590D21E8F891" collectingDate="2022-10-30" collectionCode="MATERIAL" collectorName="A. Sokolov" country="Peru" elevation="1122" latitude="-11.19645" location="Rio Venado" longLatPrecision="1" longitude="-74.76947" municipality="Satipo Prov." pageId="1" pageNumber="150" specimenCount="1" specimenCount-male="1" stateProvince="Junin">
<collectionCode id="ED751DC2C16E8613D94D590D2199F8E1" box="[850,964,1817,1836]" pageId="1" pageNumber="150">MATERIAL</collectionCode>
.
<specimenCount id="9D624E8EC16E8613D9D2590D21B1F8E1" box="[973,1004,1814,1836]" count="1" pageId="1" pageNumber="150" type="male">1♂</specimenCount>
<collectingCountry id="F373C597C16E8613DE0E590D2666F8E1" box="[1041,1083,1817,1836]" name="Peru" pageId="1" pageNumber="150">Peru</collectingCountry>
,
<collectingRegion id="49A04BE5C16E8613DE5A590D2629F8E1" box="[1093,1140,1817,1836]" country="Peru" name="Junin" pageId="1" pageNumber="150">Junin</collectingRegion>
Reg.,
<collectingMunicipality id="6BBF1F7DC16E8613DEAE590D277CF8E1" box="[1201,1313,1817,1836]" pageId="1" pageNumber="150">Satipo Prov.</collectingMunicipality>
, near
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vill., h =
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<elevation id="00496234C16E8613D9A4592521A1F889" box="[955,1020,1841,1860]" metricMagnitude="3" metricUnit="m" metricValue="1.122" pageId="1" pageNumber="150" unit="m" value="1122.0">1122 m</elevation>
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,
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<geoCoordinate id="EE50E3C0C16E8613DE6D592526B4F889" box="[1138,1257,1841,1860]" degrees="74" direction="west" minutes="46.168" orientation="longitude" pageId="1" pageNumber="150" precision="1" value="-74.76947">74°46.168´W</geoCoordinate>
, 20
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<collectingDate id="EF9E5A2FC16E8613DF0F5925272EF889" box="[1296,1395,1841,1860]" pageId="1" pageNumber="150" value="2022-10-30">30.10.2022</collectingDate>
</date>
, leg.
<collectorName id="2691E0D1C16E8613D94D595D21E8F891" box="[850,949,1865,1884]" pageId="1" pageNumber="150">A. Sokolov</collectorName>
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(
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).
</paragraph>
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<collectionCode id="ED751DC2C16E8613D94D597521FEF8BB" box="[850,931,1889,1910]" pageId="1" pageNumber="150">NOTES</collectionCode>
. This species was described from a single female from Manu Biosphere Reserve, Pakitza Park Ranger Post in the Peruvian Department of
<collectingRegion id="49A04BE5C16E8613DE5C5982268BF861" box="[1091,1238,1942,1964]" country="Peru" name="Madre de Dios" pageId="1" pageNumber="150">Madre de Dios</collectingRegion>
[
<bibRefCitation id="EFF5F8F6C16E8613DEFD59822730F866" author="Sumlin W. D." box="[1250,1389,1942,1964]" pageId="1" pageNumber="150" pagination="1 - 12" refId="ref5098" refString="Sumlin W. D. 1993. Studies on the Neotropical Cicindelidae III: A review of the genus Langea (Coleoptera) // Cicindela. Vol. 25. Nos 1 - 2. P. 1 - 12." type="book chapter" year="1993">Sumlin, 1993</bibRefCitation>
]. Until now in
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<taxonomicName id="4C64FE84C16E8613D9FD59A5263FF80B" authorityName="Sumlin" authorityYear="1993" box="[994,1122,1969,1990]" class="Insecta" family="Carabidae" genus="Langea" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="150" phylum="Arthropoda" rank="species" species="mellicollis">
<emphasis id="B9105915C16E8613D9FD59A5263FF80B" box="[994,1122,1969,1990]" italics="true" pageId="1" pageNumber="150">L. mellicollis</emphasis>
</taxonomicName>
was known only from mentioned above locality [
<bibRefCitation id="EFF5F8F6C16E8613DE1859DF26CDF82D" author="Sumlin W. D." box="[1031,1168,1995,2017]" pageId="1" pageNumber="150" pagination="1 - 12" refId="ref5098" refString="Sumlin W. D. 1993. Studies on the Neotropical Cicindelidae III: A review of the genus Langea (Coleoptera) // Cicindela. Vol. 25. Nos 1 - 2. P. 1 - 12." type="book chapter" year="1993">Sumlin, 1993</bibRefCitation>
;
<bibRefCitation id="EFF5F8F6C16E8613DE8459D8272FF82D" author="Pearson D. L. &amp; Huber R. L." box="[1179,1394,1995,2017]" pageId="1" pageNumber="150" pagination="1 - 28" refId="ref4845" refString="Pearson D. L., Huber R. L. 1994. The tiger beetles of Pakitza Madre de Dios Peru: identification, natural history and a comparison to the Peruvian fauna (Coleoptera: Cicindelidae) // Cicindela. Vol. 27. Nos 1 - 2. P. 1 - 28." type="book chapter" year="1994">Pearson, Huber, 1994</bibRefCitation>
;
<bibRefCitation id="EFF5F8F6C16E8613D93359F2265FF836" author="Erwin T. L. &amp; Pearson D. L." box="[812,1026,2022,2044]" pageId="1" pageNumber="150" refId="ref4482" refString="Erwin T. L., Pearson D. L. 2008. A Treatise on the Western Hemisphere Caraboidea (Coleoptera) their classification, distribution, and ways of life. Vol. 2. Carabidae - Nebriiformes 2 - Cicindelitae. Sofia - Moscow: Pensoft. 365 pp. + 33 pl." type="book" year="2008">Erwin, Pearson, 2008</bibRefCitation>
]. The main measurements of the
<typeStatus id="54DF3BA5C16E8613DF5C59F22108F7DB" pageId="1" pageNumber="150" type="holotype">holotype</typeStatus>
(female) are — TL =
<quantity id="4C9C28E2C16E8613DE2A561526DEF7DB" box="[1077,1155,2049,2070]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="1" pageNumber="150" unit="mm" value="7.7">7.7 mm</quantity>
, EW =
<quantity id="4C9C28E2C16E8613DECF56142746F7DB" box="[1232,1307,2048,2070]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="1" pageNumber="150" unit="mm" value="2.6">2.6 mm</quantity>
[
<bibRefCitation id="EFF5F8F6C16E8610DF37561522ACFE9D" author="Sumlin W. D." lastPageId="2" lastPageNumber="151" pageId="1" pageNumber="150" pagination="1 - 12" refId="ref5098" refString="Sumlin W. D. 1993. Studies on the Neotropical Cicindelidae III: A review of the genus Langea (Coleoptera) // Cicindela. Vol. 25. Nos 1 - 2. P. 1 - 12." type="book chapter" year="1993">Sumlin, 1993</bibRefCitation>
], and in collected early female TL = 9.0 mm [
<bibRefCitation id="EFF5F8F6C16D8610D8DC5F2F22ADFEA6" author="Erwin T. L." pageId="2" pageNumber="151" pagination="1 - 85" refId="ref4441" refString="Erwin T. L. 1990. Natural history of the carabid beetles at the BIOLAT Biological Station, Rio Manu, Pakitza, Peru // Revista Peruana de Entomologica. Vol. 33. P. 1 - 85." type="journal article" year="1990">Erwin, 1990</bibRefCitation>
;
<bibRefCitation id="EFF5F8F6C16D8610DAE55F422390FEA6" author="Erwin T. L. &amp; Pearson D. L." box="[250,461,342,364]" pageId="2" pageNumber="151" refId="ref4482" refString="Erwin T. L., Pearson D. L. 2008. A Treatise on the Western Hemisphere Caraboidea (Coleoptera) their classification, distribution, and ways of life. Vol. 2. Carabidae - Nebriiformes 2 - Cicindelitae. Sofia - Moscow: Pensoft. 365 pp. + 33 pl." type="book" year="2008">Erwin, Pearson, 2008</bibRefCitation>
: Pl. 22]. In addition, this species was found in the Colombian Department of Amazonas (according to the map — in Tarapacá Municipality) without sex and number mentions, but with the range of TL = 7.0
<quantity id="4C9C28E2C16D8610D8A05FB2215BFE76" box="[703,774,422,444]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="2" pageNumber="151" unit="mm" value="8.5">8.5 mm</quantity>
[
<bibRefCitation id="EFF5F8F6C16D8610DADD5FD523FCFE1B" author="Vitolo A. L. &amp; Pearson D. L." box="[194,417,449,471]" pageId="2" pageNumber="151" pagination="167 - 174" refId="ref5306" refString="Vitolo A. L., Pearson D. L. 2003. Escarabajos tigre (Coleoptera: Cicindelidae) de Colombia // Biota Colombiana. Vol. 4. No. 2. P. 167 - 174." type="book chapter" year="2003">Vítolo, Pearson, 2003</bibRefCitation>
;
<bibRefCitation id="EFF5F8F6C16D8610DBB15FD5206FFE1B" author="Vitolo A. L." box="[430,562,449,471]" pageId="2" pageNumber="151" refId="ref5266" refString="Vitolo A. L. 2004. Guia para la identificacion de los escarabajos tigre (Coleoptera: Cicindelidae) de Colombia. Instituto de Investigacion de Recursos Biologicos Alexander von Humboldt: Bogota, Colombia. 198 pp." type="book" year="2004">Vítolo, 2004</bibRefCitation>
]. The information on morphology of the male is still absent [
<bibRefCitation id="EFF5F8F6C16D8610D8205FCF2095FE3D" author="Sumlin W. D." box="[575,712,475,497]" pageId="2" pageNumber="151" pagination="1 - 12" refId="ref5098" refString="Sumlin W. D. 1993. Studies on the Neotropical Cicindelidae III: A review of the genus Langea (Coleoptera) // Cicindela. Vol. 25. Nos 1 - 2. P. 1 - 12." type="book chapter" year="1993">Sumlin, 1993</bibRefCitation>
;
<bibRefCitation id="EFF5F8F6C16D8610D8CB5FC8233AFDC6" author="Pearson D. L. &amp; Huber R. L." pageId="2" pageNumber="151" pagination="1 - 28" refId="ref4845" refString="Pearson D. L., Huber R. L. 1994. The tiger beetles of Pakitza Madre de Dios Peru: identification, natural history and a comparison to the Peruvian fauna (Coleoptera: Cicindelidae) // Cicindela. Vol. 27. Nos 1 - 2. P. 1 - 28." type="book chapter" year="1994">Pearson, Huber, 1994</bibRefCitation>
;
<bibRefCitation id="EFF5F8F6C16D8610DB6E5FE2201AFDC6" author="Erwin T. L. &amp; Pearson D. L." box="[369,583,502,524]" pageId="2" pageNumber="151" refId="ref4482" refString="Erwin T. L., Pearson D. L. 2008. A Treatise on the Western Hemisphere Caraboidea (Coleoptera) their classification, distribution, and ways of life. Vol. 2. Carabidae - Nebriiformes 2 - Cicindelitae. Sofia - Moscow: Pensoft. 365 pp. + 33 pl." type="book" year="2008">Erwin, Pearson, 2008</bibRefCitation>
]. Thus, in the Peruvian Region of
<collectingRegion id="49A04BE5C16D8610DB465C0523D0FDEA" box="[345,397,529,551]" country="Peru" name="Junin" pageId="2" pageNumber="151">Junin</collectingRegion>
<taxonomicName id="4C64FE84C16D8610DB895C05204AFDEB" authorityName="Sumlin" authorityYear="1993" box="[406,535,529,550]" class="Insecta" family="Carabidae" genus="Langea" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="151" phylum="Arthropoda" rank="species" species="mellicollis">
<emphasis id="B9105915C16D8610DB895C05204AFDEB" box="[406,535,529,550]" italics="true" pageId="2" pageNumber="151">L. mellicollis</emphasis>
</taxonomicName>
is recorded for the first time. Moreover, the male of this species is also illustrated for the first time (
<figureCitation id="135F9982C16D8610D9AA5F422650FEA6" box="[949,1037,342,364]" captionStart="Figs 14" captionStartId="2.[226,263,2017,2036]" captionTargetBox="[189,1399,629,1995]" captionTargetId="figure-268@2.[189,1399,578,2001]" captionTargetPageId="2" captionText="Figs 14. Langea mellicollis, male: 1 — habitus, dorsal view; 2 — labrum; 3 — left fore-tarsus; 4 — aedeagus, left lateral view. Рис. 14. Langea mellicollis, самец: 1 — внеШний вид, сверху; 2 — верхнЯЯ губа; 3 — леваЯ переднЯЯ лапка; 4 — Эдеагус, слева." figureDoi="http://doi.org/10.5281/zenodo.13177538" httpUri="https://zenodo.org/record/13177538/files/figure.png" pageId="2" pageNumber="151">Figs 14</figureCitation>
). Its main measurements are (in mm) — TL = 7.9, HW = 2.4, OW = 1.5, LL = 0.8, LW = 1.0,
<collectionCode id="ED751DC2C16D8610DF585F65273EFE4B" box="[1351,1379,369,390]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="2" pageNumber="151" type="Museum">PL</collectionCode>
= 1.4,
<collectionCode id="ED751DC2C16D8610D9475F982121FE6C" box="[856,892,396,417]" country="Thailand" name="Paleontological Collections" pageId="2" pageNumber="151">PW</collectionCode>
= 1.5, EL = 5.0, EW = 2.6, AL = 2.5. Labrum pale with light brown lateral and basal margins, six teeth and relatively broad and flat apical portion, as well as six long marginal setae (
<figureCitation id="135F9982C16D8610D9D65FCF265BFE3D" box="[969,1030,475,497]" captionStart="Figs 14" captionStartId="2.[226,263,2017,2036]" captionTargetBox="[189,1399,629,1995]" captionTargetId="figure-268@2.[189,1399,578,2001]" captionTargetPageId="2" captionText="Figs 14. Langea mellicollis, male: 1 — habitus, dorsal view; 2 — labrum; 3 — left fore-tarsus; 4 — aedeagus, left lateral view. Рис. 14. Langea mellicollis, самец: 1 — внеШний вид, сверху; 2 — верхнЯЯ губа; 3 — леваЯ переднЯЯ лапка; 4 — Эдеагус, слева." figureDoi="http://doi.org/10.5281/zenodo.13177538" httpUri="https://zenodo.org/record/13177538/files/figure.png" pageId="2" pageNumber="151">Fig. 2</figureCitation>
). Three basal tarsomeres of fore legs are very wide and densely haired by long black setae (
<figureCitation id="135F9982C16D8610DF2B5FE22164FDEB" captionStart="Figs 14" captionStartId="2.[226,263,2017,2036]" captionTargetBox="[189,1399,629,1995]" captionTargetId="figure-268@2.[189,1399,578,2001]" captionTargetPageId="2" captionText="Figs 14. Langea mellicollis, male: 1 — habitus, dorsal view; 2 — labrum; 3 — left fore-tarsus; 4 — aedeagus, left lateral view. Рис. 14. Langea mellicollis, самец: 1 — внеШний вид, сверху; 2 — верхнЯЯ губа; 3 — леваЯ переднЯЯ лапка; 4 — Эдеагус, слева." figureDoi="http://doi.org/10.5281/zenodo.13177538" httpUri="https://zenodo.org/record/13177538/files/figure.png" pageId="2" pageNumber="151">Figs 1, 3</figureCitation>
). Aedeagus with very large, curved ventrally hook (
<figureCitation id="135F9982C16D8610DF305C052736FDEB" box="[1327,1387,529,551]" captionStart="Figs 14" captionStartId="2.[226,263,2017,2036]" captionTargetBox="[189,1399,629,1995]" captionTargetId="figure-268@2.[189,1399,578,2001]" captionTargetPageId="2" captionText="Figs 14. Langea mellicollis, male: 1 — habitus, dorsal view; 2 — labrum; 3 — left fore-tarsus; 4 — aedeagus, left lateral view. Рис. 14. Langea mellicollis, самец: 1 — внеШний вид, сверху; 2 — верхнЯЯ губа; 3 — леваЯ переднЯЯ лапка; 4 — Эдеагус, слева." figureDoi="http://doi.org/10.5281/zenodo.13177538" httpUri="https://zenodo.org/record/13177538/files/figure.png" pageId="2" pageNumber="151">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>