treatments-xml/data/03/F5/FA/03F5FA39BC67FFC1B1D2FD6C3520F84A.xml

<document ID-DOI="10.5252/a2015n2a2" ID-ISSN="1639-4798" ID-Zenodo-Dep="5208903" approvalRequired="50" approvalRequired_for_taxonomicNames="1" approvalRequired_for_textStreams="44" approvalRequired_for_treatments="5" checkinTime="1629150308770" checkinUser="carolina" docAuthor="Daly, Douglas C., Raharimampionona, Jeannie &amp; Federman, Sarah" docDate="2015" docId="03F5FA39BC67FFC1B1D2FD6C3520F84A" docLanguage="en" docName="Adansonia.37.2.277-345.pdf" docOrigin="Adansonia 37 (2)" docSource="https://www.mendeley.com/catalogue/2522e05a-0ab0-3e53-96aa-d45147a2ca6c/" docStyle="DocumentStyle:E7A860AFEA63D44BCC37955D07FCAFC7.2:Adansonia.2015-2017.journal_article" docStyleId="E7A860AFEA63D44BCC37955D07FCAFC7" docStyleName="Adansonia.2015-2017.journal_article" docStyleVersion="2" docTitle="Canarium subsidarium Daly, Raharim. &amp; Federman 2015, sp. nov." docType="treatment" docVersion="2" lastPageNumber="340" masterDocId="FFCC8241BC58FFFEB157FF99310AFFA1" masterDocTitle="A revision of Canarium L. (Burseraceae) in Madagascar" masterLastPageNumber="345" masterPageNumber="277" pageId="63" pageNumber="340" updateTime="1629150974147" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title>A revision of Canarium L. (Burseraceae) in Madagascar</mods:title>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Daly, Douglas C.</mods:namePart>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Raharimampionona, Jeannie</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
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<mods:namePart>Federman, Sarah</mods:namePart>
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<mods:title>Adansonia</mods:title>
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<mods:date>2015</mods:date>
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<mods:number>2015-12-31</mods:number>
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<mods:number>37</mods:number>
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<treatment LSID="urn:lsid:plazi:treatment:03F5FA39BC67FFC1B1D2FD6C3520F84A" httpUri="http://treatment.plazi.org/id/03F5FA39BC67FFC1B1D2FD6C3520F84A" lastPageNumber="340" pageId="63" pageNumber="340">
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<taxonomicName authority="Daly, Raharim. &amp; Federman" authorityName="Daly, Raharim. &amp; Federman" authorityYear="2015" box="[133,684,757,784]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="63" pageNumber="340" phylum="Tracheophyta" rank="species" species="subsidarium" status="sp. nov.">
<emphasis bold="true" box="[133,377,757,784]" italics="true" pageId="63" pageNumber="340">Canarium subsidarium</emphasis>
Daly, Raharim. &amp; Federman
</taxonomicName>
, 
<taxonomicNameLabel box="[693,775,758,784]" pageId="63" pageNumber="340" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection pageId="63" pageNumber="340" type="description">
<paragraph blockId="63.[133,775,757,815]" box="[391,516,789,815]" pageId="63" pageNumber="340">
(
<figureCitation box="[398,467,789,815]" captionStart="FIG" captionStartId="13.[132,143,1067,1085]" captionTargetBox="[140,667,763,917]" captionTargetPageId="13" captionText="FIG. 4. — Distributions of Canarium angulatum Daly, Raharim. &amp; Federman, sp. nov., C. ferrugineum Daly, Raharim. &amp; Federman, sp. nov., C. globosum Daly,Raharim.&amp; Federman,sp.nov., C. nitidifolium Daly,Raharim.&amp; Federman, sp. nov., and C. subsidarium Daly, Raharim. &amp; Federman, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5208909" httpUri="https://zenodo.org/record/5208909/files/figure.png" pageId="63" pageNumber="340">Figs 4</figureCitation>
; 
<figureCitation box="[479,508,789,815]" captionStart="FIG" captionStartId="62.[132,143,1917,1935]" captionTargetBox="[206,1388,215,1869]" captionTargetId="figure-19@62.[195,1392,209,1872]" captionTargetPageId="62" captionText="FIG. 29. — Canarium subsidarium Daly, Raharim. &amp; Federman, sp. nov.: A, flowering branchlet with detail of leaflet margin (A’) and petiole base with stipular scar (A’’); B, detail of leaf rachis, petiolule and leaflet base; C, staminate inflorescence and apical view of stipular scars (C’); D, portion of immature inflorescence; E, staminate bud; F, longisection of staminate flower; G, stamens and ovariodisk; H, abaxial, adaxial and lateral views of stamen; I, leaf minus base of petiole; J, portion of infructescence; K, pyrene; A, B, Malcomber 2596 (NY); C-H, Capuron (SF) 11513 (NY); I, J, Rakotomalala &amp; Narison 106 (MO); K, from Malcomber et al. 920 (NY). Scale bars: A, C, I, J, K, 2 cm; B, D, 5 mm; E, F, G, 2 mm; H, 1 mm; A’, A’’, C’, not to scale." figureDoi="http://doi.org/10.5281/zenodo.5208961" httpUri="https://zenodo.org/record/5208961/files/figure.png" pageId="63" pageNumber="340">29</figureCitation>
)
</paragraph>
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<emphasis italics="true" pageId="63" pageNumber="340">
Small trees, leaves 3-4(5)-jugate, stipular scars 
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long, fruits ovoid, green, glabrous, becoming finely lenticellate; distinguished from
</emphasis>
<taxonomicName authority="Daly, Raharim. &amp; Federman" authorityName="Daly, Raharim. &amp; Federman" authorityYear="2015" box="[132,560,899,923]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="63" pageNumber="340" phylum="Tracheophyta" rank="species" species="betamponae" status="sp. nov.">
C. betamponae 
<emphasis box="[286,560,899,923]" italics="true" pageId="63" pageNumber="340">Daly, Raharim. &amp; Federman</emphasis>
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, 
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,
</emphasis>
<taxonomicName authority="Daly, Raharim. &amp; Federman" authorityName="Daly, Raharim. &amp; Federman" authorityYear="2015" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="63" pageNumber="340" phylum="Tracheophyta" rank="species" species="galokense" status="sp. nov.">
C. galokense 
<emphasis box="[131,397,926,950]" italics="true" pageId="63" pageNumber="340">Daly, Raharim. &amp; Federman</emphasis>
</taxonomicName>
<emphasis box="[396,472,926,950]" italics="true" pageId="63" pageNumber="340">
, 
<taxonomicNameLabel box="[404,472,926,949]" pageId="63" pageNumber="340" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName authority="Daly, Raharim. &amp; Federman" authorityName="Daly, Raharim. &amp; Federman" authorityYear="2015" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="63" pageNumber="340" phylum="Tracheophyta" rank="species" species="globosum" status="sp.nov.">
C. globosum 
<emphasis italics="true" pageId="63" pageNumber="340">Daly, Raharim. &amp; Federman</emphasis>
</taxonomicName>
<emphasis box="[222,295,952,976]" italics="true" pageId="63" pageNumber="340">
, 
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</emphasis>
, 
<emphasis box="[305,339,952,976]" italics="true" pageId="63" pageNumber="340">and</emphasis>
<taxonomicName authority="Daly, Raharim.&amp; Federman" authorityName="Daly, Raharim. &amp; Federman" authorityYear="2015" box="[343,770,952,976]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="63" pageNumber="340" phylum="Tracheophyta" rank="species" species="manongarivum" status="sp. nov.">
C. manongarivum 
<emphasis box="[514,770,952,976]" italics="true" pageId="63" pageNumber="340">Daly, Raharim.&amp; Federman</emphasis>
</taxonomicName>
<emphasis italics="true" pageId="63" pageNumber="340">
, 
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by the petiole and rachis with dense to scattered hairs to 
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long (vs scattered to sparse hairs to 
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long in the others) and occurrence in montane forests 
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elevation (vs maximum recorded 
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), further distinguished from
</emphasis>
<taxonomicName authority="Daly &amp; Raharimampionona &amp; Federman, 2015" authorityName="Daly &amp; Raharimampionona &amp; Federman" authorityYear="2015" box="[542,666,1058,1082]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="63" pageNumber="340" phylum="Tracheophyta" rank="species" species="galokense" status="sp. nov.">C. galokense</taxonomicName>
, 
<emphasis italics="true" pageId="63" pageNumber="340">
<taxonomicNameLabel box="[678,749,1059,1082]" pageId="63" pageNumber="340" rank="species">sp. nov.</taxonomicNameLabel>
by the fruit ovoid (vs broadly oblong or broadly ovoid) with the surface coarsely (vs finely) lenticellate.
</emphasis>
</paragraph>
<paragraph blockId="63.[130,777,846,1696]" pageId="63" pageNumber="340">
<materialsCitation collectingDate="1994-01-10" collectionCode="MO, TAN" collectorName="Speciale de Manongarivo, E &amp; L. Rakotomalala &amp; Narison" country="Madagascar" elevation="850" latitude="-14.051389" location="Bekolosy" longLatPrecision="21" longitude="48.28528" municipality="Reserve Speciale de Manongarivo" pageId="63" pageNumber="340" specimenCount="1" stateProvince="Antsiranana">
TYPUS. — 
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<collectingCountry box="[236,356,1158,1182]" name="Madagascar" pageId="63" pageNumber="340">Madagascar</collectingCountry>
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. 
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, 
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Réserve 
<collectorName pageId="63" pageNumber="340">Spéciale de Manongarivo</collectorName>
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, E d’Ankaramibe; 
<location LSID="urn:lsid:plazi:treatment:03F5FA39BC67FFC1B1D2FD6C3520F84A:8E831DF4BC67FFC1B0C7FB3830E5FB18" box="[400,495,1185,1209]" country="Madagascar" latitude="-14.051389" longLatPrecision="21" longitude="48.28528" municipality="Reserve Speciale de Manongarivo" name="Bekolosy" pageId="63" pageNumber="340" stateProvince="Antsiranana">Bekolosy</location>
, 
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<elevation box="[508,577,1185,1209]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="63" pageNumber="340" unit="m" value="850.0">850 m</elevation>
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elev., 
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, 
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, 
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<collectingDate box="[265,367,1211,1235]" pageId="63" pageNumber="340" value="1994-01-10">10.I.1994</collectingDate>
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, 
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<collectorName box="[380,542,1211,1235]" pageId="63" pageNumber="340">L. Rakotomalala</collectorName>
&amp; 
<collectorName box="[576,653,1212,1235]" pageId="63" pageNumber="340">Narison</collectorName>
106
</emphasis>
(holo-, 
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!; iso-, 
<collectionCode box="[245,301,1238,1262]" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13842" lsid="urn:lsid:biocol.org:col:13842" name="Parc de Tsimbazaza" pageId="63" pageNumber="340" type="Herbarium">TAN</collectionCode>
)
</materialsCitation>
.
</paragraph>
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<materialsCitation collectingDate="1954-11" collectionCode="NY" collectorName="de l'Antsatrotro &amp; Capuron" country="Madagascar" elevation="1100" location="Massif de l'Antsatrotro" municipality="Diana" pageId="63" pageNumber="340" specimenCount="1" stateProvince="Antsiranana" typeStatus="paratype">
<typeStatus box="[132,219,1286,1309]" pageId="63" pageNumber="340">PARATYPI</typeStatus>
. — 
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<collectingCountry box="[260,377,1285,1309]" name="Madagascar" pageId="63" pageNumber="340">Madagascar</collectingCountry>
.
</emphasis>
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, 
<collectingMunicipality box="[510,571,1285,1309]" pageId="63" pageNumber="340">Diana</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:03F5FA39BC67FFC1B1D2FD6C3520F84A:8E831DF4BC67FFC1B313FA9C31AAFA96" country="Madagascar" municipality="Diana" name="Massif de l'Antsatrotro" pageId="63" pageNumber="340" stateProvince="Antsiranana">
Massif 
<collectorName pageId="63" pageNumber="340">de l’Antsatrotro</collectorName>
</location>
(Manongarivo), 
<emphasis box="[323,338,1313,1336]" italics="true" pageId="63" pageNumber="340">c.</emphasis>
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<elevation box="[344,421,1312,1336]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="63" pageNumber="340" unit="m" value="1100.0">1100 m</elevation>
</quantity>
, 
<date box="[430,514,1312,1336]" pageId="63" pageNumber="340" value="1954-11">
<collectingDate box="[430,514,1312,1336]" pageId="63" pageNumber="340" value="1954-11">XI.1954</collectingDate>
</date>
, 
<emphasis italics="true" pageId="63" pageNumber="340">
<collectorName box="[522,605,1312,1335]" pageId="63" pageNumber="340">Capuron</collectorName>
(Service Forestier) 11513
</emphasis>
(
<collectionCode box="[210,249,1338,1362]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="63" pageNumber="340" type="Herbarium">NY</collectionCode>
)
</materialsCitation>
; 
<materialsCitation collectingDate="1991-09-29" collectionCode="MO, NY" collectorName="S. Malcomber &amp; S. Razafimandimbison" country="Madagascar" elevation="1475" latitude="-14.1" location="Reserve Speciale Manongarivo" longLatPrecision="1289" longitude="48.35" pageId="63" pageNumber="340" specimenCount="1" stateProvince="Antsiranana" typeStatus="paratype">
<location LSID="urn:lsid:plazi:treatment:03F5FA39BC67FFC1B1D2FD6C3520F84A:8E831DF4BC67FFC1B05BFAA33333FAF3" box="[268,569,1338,1362]" country="Madagascar" latitude="-14.1" longLatPrecision="1289" longitude="48.35" name="Reserve Speciale Manongarivo" pageId="63" pageNumber="340" stateProvince="Antsiranana">Réserve Spéciale Manongarivo</location>
, 
<locationDeviation box="[580,705,1338,1363]" location="Beraty" pageId="63" pageNumber="340">SE of Beraty</locationDeviation>
, 
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.475" metricValueMax="1.6" metricValueMin="1.35" pageId="63" pageNumber="340" unit="m" value="1475.0" valueMax="1600.0" valueMin="1350.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.475" metricValueMax="1.6" metricValueMin="1.35" pageId="63" pageNumber="340" unit="m" value="1475.0" valueMax="1600.0" valueMin="1350.0">1350- 1600 m</elevation>
</quantity>
, 
<geoCoordinate box="[222,301,1365,1389]" degrees="14" direction="south" minutes="06" orientation="latitude" pageId="63" pageNumber="340" precision="925" value="-14.1">14°06’S</geoCoordinate>
, 
<geoCoordinate box="[312,394,1365,1389]" degrees="48" direction="east" minutes="21" orientation="longitude" pageId="63" pageNumber="340" precision="925" value="48.35">48°21’E</geoCoordinate>
, 
<date box="[404,520,1365,1389]" pageId="63" pageNumber="340" value="1991-09-29">
<collectingDate box="[404,520,1365,1389]" pageId="63" pageNumber="340" value="1991-09-29">29.IX.1991</collectingDate>
</date>
, 
<emphasis italics="true" pageId="63" pageNumber="340">
<collectorName box="[531,662,1365,1389]" pageId="63" pageNumber="340">S. Malcomber</collectorName>
&amp; 
<collectorName pageId="63" pageNumber="340">S. Razafimandimbison</collectorName>
920
</emphasis>
(
<collectionCode box="[334,380,1391,1415]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="63" pageNumber="340" type="Herbarium">MO</collectionCode>
, 
<collectionCode box="[392,430,1391,1415]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="63" pageNumber="340" type="Herbarium">NY</collectionCode>
)
</materialsCitation>
; 
<materialsCitation collectingDate="1993-12-04" collectionCode="MO, NY, TAN" collectorName="S. Malcomber &amp; L. Rakotomalala" country="Madagascar" elevation="1100" latitude="-14.033334" location="Ankaramibe" longLatPrecision="1289" longitude="48.3" pageId="63" pageNumber="340" specimenCount="1" stateProvince="Antsiranana" typeStatus="paratype">
E of 
<location LSID="urn:lsid:plazi:treatment:03F5FA39BC67FFC1B1D2FD6C3520F84A:8E831DF4BC67FFC1B0A5FAF63366FA26" box="[498,620,1391,1415]" country="Madagascar" latitude="-14.033334" longLatPrecision="1289" longitude="48.3" name="Ankaramibe" pageId="63" pageNumber="340" stateProvince="Antsiranana">Ankaramibe</location>
, 
<quantity box="[632,771,1391,1415]" metricMagnitude="3" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="63" pageNumber="340" unit="m" value="1100.0" valueMax="1200.0" valueMin="1000.0">
<elevation box="[632,771,1391,1415]" metricMagnitude="3" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="63" pageNumber="340" unit="m" value="1100.0" valueMax="1200.0" valueMin="1000.0">1000-1200 m</elevation>
</quantity>
, 
<geoCoordinate box="[134,212,1418,1442]" degrees="14" direction="south" minutes="02" orientation="latitude" pageId="63" pageNumber="340" precision="925" value="-14.033334">14°02’S</geoCoordinate>
, 
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, 
<date box="[317,441,1418,1442]" pageId="63" pageNumber="340" value="1993-12-04">
<collectingDate box="[317,441,1418,1442]" pageId="63" pageNumber="340" value="1993-12-04">04.XII.1993</collectingDate>
</date>
, 
<emphasis italics="true" pageId="63" pageNumber="340">
<collectorName box="[453,584,1418,1442]" pageId="63" pageNumber="340">S. Malcomber</collectorName>
&amp; 
<collectorName box="[617,775,1418,1442]" pageId="63" pageNumber="340">L. Rakotomalala</collectorName>
2596
</emphasis>
(
<collectionCode box="[197,243,1444,1468]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="63" pageNumber="340" type="Herbarium">MO</collectionCode>
, 
<collectionCode box="[253,288,1445,1468]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="63" pageNumber="340" type="Herbarium">NY</collectionCode>
, P, 
<collectionCode box="[319,374,1444,1468]" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13842" lsid="urn:lsid:biocol.org:col:13842" name="Parc de Tsimbazaza" pageId="63" pageNumber="340" type="Herbarium">TAN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="63" pageNumber="340" type="distribution">
<paragraph blockId="63.[130,777,846,1696]" pageId="63" pageNumber="340">
DISTRIBUTION AND ECOLOGY. — 
<taxonomicName authority="Daly &amp; Raharimampionona &amp; Federman, 2015" authorityName="Daly &amp; Raharimampionona &amp; Federman" authorityYear="2015" box="[446,658,1492,1516]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="63" pageNumber="340" phylum="Tracheophyta" rank="species" species="subsidarium" status="sp. nov.">
<emphasis box="[446,658,1492,1516]" italics="true" pageId="63" pageNumber="340">Canarium subsidarium</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel box="[670,741,1492,1515]" pageId="63" pageNumber="340" rank="species">sp. nov.</taxonomicNameLabel>
appears to be restricted to Manongarivo in N 
<collectingCountry box="[543,660,1518,1542]" name="Madagascar" pageId="63" pageNumber="340">Madagascar</collectingCountry>
. It is usually found in montane moss forest on ridgetops, between 
<quantity box="[639,773,1545,1569]" metricMagnitude="3" metricUnit="m" metricValue="1.35" metricValueMax="1.6" metricValueMin="1.1" pageId="63" pageNumber="340" unit="m" value="1350.0" valueMax="1600.0" valueMin="1100.0">1100-1600 m</quantity>
(collected once at 
<emphasis box="[300,315,1572,1595]" italics="true" pageId="63" pageNumber="340">c.</emphasis>
<quantity box="[320,386,1571,1595]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="63" pageNumber="340" unit="m" value="850.0">850 m</quantity>
). Flowering Nov.-Dec., fruiting Jan.-Sep. ETYMOLOGY. — The specific epithet, meaning “in reserve,” derives from the fact that the species is known only from the area of the Réserve Spéciale Manongarivo.
</paragraph>
</subSubSection>
<subSubSection pageId="63" pageNumber="340" type="description">
<paragraph blockId="63.[131,777,1746,2028]" box="[132,277,1746,1772]" pageId="63" pageNumber="340">DESCRIPTION</paragraph>
<paragraph blockId="63.[131,777,1746,2028]" lastBlockId="63.[811,1458,215,1676]" pageId="63" pageNumber="340">
Trees, reproductive height 
<quantity box="[429,514,1778,1805]" metricMagnitude="0" metricUnit="m" metricValue="8.5" metricValueMax="12.0" metricValueMin="5.0" pageId="63" pageNumber="340" unit="m" value="8.5" valueMax="12.0" valueMin="5.0">5-12 m</quantity>
, to 
<quantity box="[555,625,1778,1804]" metricMagnitude="-1" metricUnit="m" metricValue="4.0" pageId="63" pageNumber="340" unit="cm" value="40.0">40 cm</quantity>
diam. Leaves 
<quantity box="[132,244,1809,1836]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="63" pageNumber="340" unit="cm" value="20.0" valueMax="25.0" valueMin="15.0">15-25 cm</quantity>
long, 3-4(5)-jugate; petiole 
<quantity box="[579,707,1809,1836]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" metricValueMax="4.5" metricValueMin="2.5" pageId="63" pageNumber="340" unit="cm" value="3.5" valueMax="4.5" valueMin="2.5">2.5-4.5 cm</quantity>
long, petiole and rachis with dense to sparse capitate glands and bristles (fine erect hairs to 
<quantity box="[438,547,1873,1900]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="63" pageNumber="340" unit="mm" value="0.05">0.05 mm</quantity>
), also with dense to scattered fine erect hairs to 
<quantity box="[436,526,1905,1932]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="63" pageNumber="340" unit="mm" value="0.5">0.5 mm</quantity>
; stipules 
<quantity box="[627,716,1905,1932]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="9.0" metricValueMin="4.0" pageId="63" pageNumber="340" unit="mm" value="6.5" valueMax="9.0" valueMin="4.0">4-9 mm</quantity>
from petiole insertion, caducous, the scar 
<emphasis box="[525,542,1938,1963]" italics="true" pageId="63" pageNumber="340">c.</emphasis>
<quantity box="[549,636,1937,1964]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="63" pageNumber="340" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2-3 mm</quantity>
long, raised; basal petiolules 
<quantity box="[304,393,1969,1996]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="63" pageNumber="340" unit="mm" value="4.5" valueMax="6.0" valueMin="3.0">3-6 mm</quantity>
long, other laterals 
<quantity box="[610,712,1969,1996]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="11.0" metricValueMin="6.0" pageId="63" pageNumber="340" unit="mm" value="8.5" valueMax="11.0" valueMin="6.0">6-11 mm</quantity>
long, terminal one 
<quantity box="[280,397,2001,2028]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.5" metricValueMin="1.0" pageId="63" pageNumber="340" unit="mm" value="17.5" valueMax="25.0" valueMin="10.0">10-25 mm</quantity>
long, pulvinuli distal and moder- ately conspicuous; basal leaflets 3-6.5 
<emphasis box="[1236,1252,216,241]" italics="true" pageId="63" pageNumber="340">×</emphasis>
<quantity box="[1260,1386,215,242]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="63" pageNumber="340" unit="cm" value="2.0" valueMax="2.5" valueMin="1.5">1.5-2.5 cm</quantity>
, ovate to elliptic, other laterals 4.4-9.8 
<emphasis box="[1189,1205,248,273]" italics="true" pageId="63" pageNumber="340">×</emphasis>
<quantity box="[1217,1346,247,274]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" metricValueMax="3.2" metricValueMin="2.4" pageId="63" pageNumber="340" unit="cm" value="2.8" valueMax="3.2" valueMin="2.4">2.4-3.2 cm</quantity>
, oblong(- lanceolate) to elliptic, terminal one 5-8.7 
<emphasis box="[1285,1301,280,305]" italics="true" pageId="63" pageNumber="340">×</emphasis>
<quantity box="[1310,1416,279,306]" metricMagnitude="-2" metricUnit="m" metricValue="3.65" metricValueMax="4.3" metricValueMin="3.0" pageId="63" pageNumber="340" unit="cm" value="3.65" valueMax="4.3" valueMin="3.0">3-4.3 cm</quantity>
, elliptic to oblanceolate; leaflet apex gradually and broadly short-acuminate, the acumen 
<quantity box="[1147,1239,343,369]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="6.0" metricValueMin="2.0" pageId="63" pageNumber="340" unit="mm" value="4.0" valueMax="6.0" valueMin="2.0">2-6 mm</quantity>
, apex tip glandular; base truncate to rounded or obtuse, basally symmetrical or less often slightly oblique; margin flat; leaflets somewhat coriaceous, drying dark grayish-brown; secondary vein framework brochidodromous but looping near margin, secondaries 7-11 per side, spreading, discolorous, spacing usually decreasing toward apex and base, angle acute and increasing toward base; tertiary venation random-reticulate to alternate-percurrent; on abaxial side the midvein acutely prominent, secondary veins broadly prominulous, remainder narrowly prominent to prominulous, midvein with scattered ascending hairs to 
<quantity box="[1084,1186,693,720]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="63" pageNumber="340" unit="mm" value="0.15">0.15 mm</quantity>
and rest of surface with a few capitate glands and scattered thick appressed hairs to 
<quantity box="[812,911,757,784]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="63" pageNumber="340" unit="mm" value="0.05">0.05 mm</quantity>
long (sparse near base), the surface also appearing papillate; on adaxial side the midvein narrowly prominulous and sunk in a groove, rest of veins narrowly prominulous, surface glabrous. Inflorescences 
<quantity box="[1168,1283,853,880]" metricMagnitude="-1" metricUnit="m" metricValue="1.675" metricValueMax="2.5" metricValueMin="0.85" pageId="63" pageNumber="340" unit="cm" value="16.75" valueMax="25.0" valueMin="8.5">8.5-25 cm</quantity>
long, axes with sparse to dense bristles and capitate glands and with dense to sparse flexuous hairs to 
<quantity box="[1094,1182,917,943]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="63" pageNumber="340" unit="mm" value="0.3">0.3 mm</quantity>
; staminate inflorescences with secondary axes to 
<quantity box="[1076,1155,948,975]" metricMagnitude="-2" metricUnit="m" metricValue="9.5" pageId="63" pageNumber="340" unit="cm" value="9.5">9.5 cm</quantity>
long; bracts on secondary axes 
<quantity box="[865,975,981,1007]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" metricValueMax="3.0" metricValueMin="2.8" pageId="63" pageNumber="340" unit="mm" value="2.9" valueMax="3.0" valueMin="2.8">2.8-3 mm</quantity>
long, ovate and acuminate with constricted base, longer than subtended buds, bracteoles 
<quantity box="[1314,1449,1013,1039]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.2" metricValueMin="1.7" pageId="63" pageNumber="340" unit="mm" value="1.95" valueMax="2.2" valueMin="1.7">1.7-2.2 mm</quantity>
, subulate, shorter than buds, bracts and bracteoles usually semi-clasping; pedicel 1.2-3.0 
<emphasis box="[1163,1179,1077,1102]" italics="true" pageId="63" pageNumber="340">×</emphasis>
<quantity box="[1191,1330,1076,1103]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="63" pageNumber="340" unit="mm" value="0.65" valueMax="0.8" valueMin="0.5">0.5-0.8 mm</quantity>
. Staminate buds (flowers unknown) 
<quantity box="[1105,1222,1108,1135]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" metricValueMax="5.2" metricValueMin="5.0" pageId="63" pageNumber="340" unit="mm" value="5.1" valueMax="5.2" valueMin="5.0">5-5.2 mm</quantity>
long; calyx 3-4.5 
<emphasis box="[1439,1455,1109,1134]" italics="true" pageId="63" pageNumber="340">×</emphasis>
<quantity box="[813,945,1140,1167]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.9" metricValueMin="2.2" pageId="63" pageNumber="340" unit="mm" value="2.55" valueMax="2.9" valueMin="2.2">2.2-2.9 mm</quantity>
overall, much taller than the ovariodisk, lobes 
<quantity box="[812,940,1172,1198]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" metricValueMax="1.6" metricValueMin="0.6" pageId="63" pageNumber="340" unit="mm" value="1.1" valueMax="1.6" valueMin="0.6">0.6-1.6 mm</quantity>
long, deltate, abaxial surface with dense capitate glands and dense to sparse flexuous ascending to appressed golden hairs to 
<quantity box="[992,1082,1236,1262]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="63" pageNumber="340" unit="mm" value="0.2">0.2 mm</quantity>
long, adaxial surface with dense retrorse-appressed pale golden hairs to 
<quantity box="[1240,1325,1268,1294]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="63" pageNumber="340" unit="mm" value="0.2">0.2 mm</quantity>
long; petals cream, with flexuous ascending to appressed golden hairs to 
<quantity box="[843,934,1331,1358]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="63" pageNumber="340" unit="mm" value="0.3">0.3 mm</quantity>
; stamens inserted at two different levels toward base of ovariodisk, the antesepalous stamens 
<quantity box="[1320,1454,1363,1390]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" metricValueMax="3.3" metricValueMin="3.1" pageId="63" pageNumber="340" unit="mm" value="3.2" valueMax="3.3" valueMin="3.1">3.1-3.3 mm</quantity>
long, the antepetalous stamens 
<quantity box="[1168,1302,1395,1422]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.1" metricValueMin="2.9" pageId="63" pageNumber="340" unit="mm" value="3.0" valueMax="3.1" valueMin="2.9">2.9-3.1 mm</quantity>
long, anthers 
<emphasis box="[814,831,1428,1453]" italics="true" pageId="63" pageNumber="340">c.</emphasis>
<quantity box="[838,972,1427,1454]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="63" pageNumber="340" unit="mm" value="1.5" valueMax="1.6" valueMin="1.4">1.4-1.6 mm</quantity>
long, oblong to ovate in dorsiventral view, oblong-lanceolate in lateral view; ovariodisk 1.3-1.6 
<emphasis box="[1388,1404,1460,1485]" italics="true" pageId="63" pageNumber="340">×</emphasis>
<quantity metricMagnitude="-3" metricUnit="m" metricValue="1.05" metricValueMax="1.5" metricValueMin="0.6" pageId="63" pageNumber="340" unit="mm" value="1.05" valueMax="1.5" valueMin="0.6">0.6- 1.5 mm</quantity>
, essentially obovoid to columnar, apex truncate and craggy to bluntly and abruptly short-attenuate. Fruiting pedicel 3-5(6.5) 
<emphasis box="[1001,1017,1555,1580]" italics="true" pageId="63" pageNumber="340">×</emphasis>
<quantity box="[1026,1165,1554,1581]" metricMagnitude="-3" metricUnit="m" metricValue="3.45" metricValueMax="3.5" metricValueMin="3.4" pageId="63" pageNumber="340" unit="mm" value="3.45" valueMax="3.5" valueMin="3.4">3.4-3.5 mm</quantity>
; fruiting calyx patent, the lobes 
<quantity box="[881,972,1587,1613]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="63" pageNumber="340" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2-3 mm</quantity>
long. Fruits 3.6-3.8 
<emphasis box="[1216,1232,1587,1612]" italics="true" pageId="63" pageNumber="340">×</emphasis>
<quantity box="[1244,1371,1587,1613]" metricMagnitude="-2" metricUnit="m" metricValue="2.55" metricValueMax="2.8" metricValueMin="2.3" pageId="63" pageNumber="340" unit="cm" value="2.55" valueMax="2.8" valueMin="2.3">2.3-2.8 cm</quantity>
, ovoid, green, glabrous, densely micro-pustulate and becoming finely lenticellate.
</paragraph>
<paragraph blockId="63.[811,1457,1715,2027]" box="[813,887,1715,1741]" pageId="63" pageNumber="340">NOTES</paragraph>
<paragraph blockId="63.[811,1457,1715,2027]" pageId="63" pageNumber="340">
<taxonomicName authority="Daly &amp; Raharimampionona &amp; Federman, 2015" authorityName="Daly &amp; Raharimampionona &amp; Federman" authorityYear="2015" box="[813,1056,1746,1772]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="63" pageNumber="340" phylum="Tracheophyta" rank="species" species="subsidarium" status="sp. nov.">
<emphasis box="[813,1056,1746,1772]" italics="true" pageId="63" pageNumber="340">Canarium subsidarium</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel box="[1070,1153,1746,1772]" pageId="63" pageNumber="340" rank="species">sp. nov.</taxonomicNameLabel>
is part of a complex of species including 
<taxonomicName authority="Daly &amp; Raharimampionona &amp; Federman, 2015" authorityName="Daly &amp; Raharimampionona &amp; Federman" authorityYear="2015" box="[966,1121,1778,1804]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="63" pageNumber="340" phylum="Tracheophyta" rank="species" species="betamponae" status="sp. nov.">
<emphasis box="[966,1121,1778,1804]" italics="true" pageId="63" pageNumber="340">C. betamponae</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel box="[1134,1218,1778,1804]" pageId="63" pageNumber="340" rank="species">sp. nov.</taxonomicNameLabel>
, 
<taxonomicName authority="Daly &amp; Raharimampionona &amp; Federman, 2015" authorityName="Daly &amp; Raharimampionona &amp; Federman" authorityYear="2015" box="[1227,1354,1778,1804]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="63" pageNumber="340" phylum="Tracheophyta" rank="species" species="galokense" status="sp. nov.">
<emphasis box="[1227,1354,1778,1804]" italics="true" pageId="63" pageNumber="340">C. galokense</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel box="[1367,1449,1778,1804]" pageId="63" pageNumber="340" rank="species">sp. nov.</taxonomicNameLabel>
, 
<emphasis box="[813,943,1810,1836]" italics="true" pageId="63" pageNumber="340">
C, 
<taxonomicName authority="Daly &amp; Raharimampionona &amp; Federman, 2015" authorityName="Daly &amp; Raharimampionona &amp; Federman" authorityYear="2015" box="[847,943,1810,1836]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="63" pageNumber="340" phylum="Tracheophyta" rank="species" species="globosum" status="sp. nov.">globosum</taxonomicName>
</emphasis>
, 
<taxonomicNameLabel box="[959,1044,1810,1836]" pageId="63" pageNumber="340" rank="species">sp. nov.</taxonomicNameLabel>
, and 
<taxonomicName authority="Daly &amp; Raharimampionona &amp; Federman, 2015" authorityName="Daly &amp; Raharimampionona &amp; Federman" authorityYear="2015" box="[1108,1302,1810,1836]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="63" pageNumber="340" phylum="Tracheophyta" rank="species" species="manongarivum" status="sp. nov.">
<emphasis box="[1108,1302,1810,1836]" italics="true" pageId="63" pageNumber="340">C. manongarivum</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel box="[1319,1404,1810,1836]" pageId="63" pageNumber="340" rank="species">sp. nov.</taxonomicNameLabel>
that all have 3-5-jugate leaves with short stipular scars (except 
<taxonomicName authority="Daly &amp; Raharimampionona &amp; Federman, 2015" authorityName="Daly &amp; Raharimampionona &amp; Federman" authorityYear="2015" box="[815,1003,1874,1900]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="63" pageNumber="340" phylum="Tracheophyta" rank="species" species="manongarivum" status="sp. nov.">
<emphasis box="[815,1003,1874,1900]" italics="true" pageId="63" pageNumber="340">C. manongarivum</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel box="[1017,1102,1874,1900]" pageId="63" pageNumber="340" rank="species">sp. nov.</taxonomicNameLabel>
), small and often oblong leaflets, and relatively small fruits. They are compared in 
<tableCitation box="[1331,1410,1905,1931]" captionStart="TABLE" captionStartId="18.[392,402,219,237]" captionText="TABLE 1. — Comparison of the Canarium betamponae, sp. nov. complex (salient features in bold)." pageId="63" pageNumber="340">Table 1</tableCitation>
un- der 
<taxonomicName authority="Daly &amp; Raharimampionona &amp; Federman, 2015" authorityName="Daly &amp; Raharimampionona &amp; Federman" authorityYear="2015" box="[852,1006,1937,1964]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="63" pageNumber="340" phylum="Tracheophyta" rank="species" species="betamponae" status="sp. nov.">
<emphasis box="[852,1006,1937,1964]" italics="true" pageId="63" pageNumber="340">C. betamponae</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel box="[1018,1098,1938,1964]" pageId="63" pageNumber="340" rank="species">sp. nov.</taxonomicNameLabel>
<taxonomicName authority="Daly &amp; Raharimampionona &amp; Federman, 2015" authorityName="Daly &amp; Raharimampionona &amp; Federman" authorityYear="2015" box="[1101,1340,1937,1964]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="63" pageNumber="340" phylum="Tracheophyta" rank="species" species="subsidarium" status="sp. nov.">
<emphasis box="[1101,1340,1937,1964]" italics="true" pageId="63" pageNumber="340">Canarium subsidarium</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel box="[1354,1435,1938,1964]" pageId="63" pageNumber="340" rank="species">sp. nov.</taxonomicNameLabel>
is also contrasted with 
<taxonomicName authority="Daly &amp; Raharimampionona &amp; Federman, 2015" authorityName="Daly &amp; Raharimampionona &amp; Federman" authorityYear="2015" box="[1024,1187,1970,1996]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="63" pageNumber="340" phylum="Tracheophyta" rank="species" species="moramangae" status="sp. nov.">
<emphasis box="[1024,1187,1970,1996]" italics="true" pageId="63" pageNumber="340">C. moramangae</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel box="[1198,1278,1970,1996]" pageId="63" pageNumber="340" rank="species">sp. nov.</taxonomicNameLabel>
in the discussion under the latter species.
</paragraph>
</subSubSection>
</treatment>
</document>