174 lines
20 KiB
XML
174 lines
20 KiB
XML
<document id="974C5251D652E370410F4013E116DE0B" ID-DOI="10.5281/zenodo.5183220" ID-ISSN="0253-116X" ID-Zenodo-Dep="5183220" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628561451650" checkinUser="felipe" docAuthor="Guilbert, Eric & Heiss, Ernst" docDate="2016" docId="03F587E7FFBA2D75F69CFB67FE96FC95" docLanguage="en" docName="LinzerbiolBeitr.48.2.1081-1090.pdf" docOrigin="Linzer biologische Beiträge 48 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Phatnoma madagascariensis Guilbert & Heiss 2016, nov.sp." docType="treatment" docVersion="3" lastPageNumber="1084" masterDocId="FFCCFF9FFFBB2D76F62EFFA7FFCBFF86" masterDocTitle="First fossil records of Tingidae from Madagascan Copal with description of two new species (Hemiptera, Heteroptera)" masterLastPageNumber="1090" masterPageNumber="1081" pageNumber="1082" updateTime="1699046987057" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:mods id="F15DB798077E378D5800AD189152ACD7" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="8A078D6BBB8E3BDB37E8593E5D91EF87">
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<mods:title id="E6A305DCAAB348919F78E8B23497B4D7">First fossil records of Tingidae from Madagascan Copal with description of two new species (Hemiptera, Heteroptera)</mods:title>
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<mods:name id="FA0FB25C9C9283478065DFEED1330F82" type="personal">
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<mods:namePart id="A4C359D1CE13A9534C4D777026CCD425">Guilbert, Eric</mods:namePart>
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<mods:name id="301E205532A661FBBD255A1A7B1A055B" type="personal">
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<mods:roleTerm id="03A759399867F875910D500A399E00D7">Author</mods:roleTerm>
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<mods:namePart id="E83CDCC0B94ACBCE9221A8D6AB4FE9B3">Heiss, Ernst</mods:namePart>
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<mods:typeOfResource id="4DE0670114E6E49C57303CF558A8444D">text</mods:typeOfResource>
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<mods:titleInfo id="749CC5B80E66D8A018FA957224B47240">
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<mods:title id="A2F15066A75FB9A69331A791368E8AA0">Linzer biologische Beiträge</mods:title>
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</mods:titleInfo>
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<mods:part id="71DD868331750515FE069DA45C41949A">
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<mods:date id="1911261A48B5706A0D53C021AA461C66">2016</mods:date>
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<mods:detail id="2354FDFC247A3556AF76E74557C152A8" type="pubDate">
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<mods:number id="4965603016DCAF53DB4B690DBABE85E1">2016-12-19</mods:number>
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<mods:detail id="BF1A01B5859D09AA7FBDAD1A4D1CDD1A" type="volume">
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<mods:number id="CA580963283D0A921B26425F655A5788">48</mods:number>
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<mods:detail id="2295C5D67006482D5BE02CC5BBCA1907" type="issue">
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<mods:number id="E701C0B9002C1C792EBA031CA9DC57AE">2</mods:number>
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<mods:start id="E4B9F33C0D7C7773A0771A9567BF98E8">1081</mods:start>
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<mods:classification id="1F65385440F33536162B7A937CA13DFC">journal article</mods:classification>
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<mods:identifier id="79F87FB55829DA404F03AF968BE66A3E" type="DOI">10.5281/zenodo.5183220</mods:identifier>
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<mods:identifier id="1178C1D15884FC3BB6B18B6A90BD5450" type="ISSN">0253-116X</mods:identifier>
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<mods:identifier id="9E0D9C6B1048326561247A528548154A" type="Zenodo-Dep">5183220</mods:identifier>
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<treatment id="03F587E7FFBA2D75F69CFB67FE96FC95" LSID="urn:lsid:plazi:treatment:03F587E7FFBA2D75F69CFB67FE96FC95" httpUri="http://treatment.plazi.org/id/03F587E7FFBA2D75F69CFB67FE96FC95" lastPageId="3" lastPageNumber="1084" pageId="1" pageNumber="1082">
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<subSubSection id="C346657AFFBA2D77F69CFB67FD58FA92" pageId="1" pageNumber="1082" type="nomenclature">
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<paragraph id="8BE336F1FFBA2D77F69CFB67FD69FB50" blockId="1.[178,674,1216,1238]" box="[178,674,1216,1238]" pageId="1" pageNumber="1082">
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<heading id="D0AB819DFFBA2D77F69CFB67FD69FB50" box="[178,674,1216,1238]" fontSize="8" level="2" pageId="1" pageNumber="1082" reason="2">
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<emphasis id="B928EAE3FFBA2D77F69CFB67FD69FB50" bold="true" box="[178,674,1216,1238]" pageId="1" pageNumber="1082">
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<taxonomicName id="4C5C4D72FFBA2D77F69CFB67FE14FB50" authority="Guilbert & Heiss, 2016" authorityName="Guilbert & Heiss" authorityYear="2016" box="[178,479,1216,1238]" class="Insecta" family="Tingidae" genus="Phatnoma" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="1082" phylum="Arthropoda" rank="species" species="madagascariensis" status="sp. nov.">
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<emphasis id="B928EAE3FFBA2D77F69CFB67FE14FB50" bold="true" box="[178,479,1216,1238]" italics="true" pageId="1" pageNumber="1082">Phatnoma madagascariensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21B5798FFBA2D77F7CBFB66FDFAFB50" box="[485,561,1217,1238]" pageId="1" pageNumber="1082" rank="species">nov.sp.</taxonomicNameLabel>
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(
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<figureCitation id="13672A74FFBA2D77F410FB67FD52FB50" box="[574,665,1216,1238]" captionStart="Figs 1-6" captionStartId="2.[178,218,1301,1323]" captionTargetBox="[179,1083,212,1292]" captionTargetId="figure-180@2.[179,1083,212,1292]" captionTargetPageId="2" captionText="Figs 1-6: Tingidae in Madagascan Copal: (1) Phatnoma madadascariensis nov.sp., holotype male, dorsal view; (2) ditto, ventral view; (3) Cysteochila impressa, male dorsal view; (4) Cysteochila copal nov.sp., holotype female, dorsal view; (5) ditto, ventral view; (6) Cysteochila impressa, male, ventral view. Scales 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5183222" httpUri="https://zenodo.org/record/5183222/files/figure.png" pageId="1" pageNumber="1082">Figs 1, 2</figureCitation>
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)
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BE336F1FFBA2D77F69CFB40FD58FA92" blockId="1.[178,1086,1254,1300]" pageId="1" pageNumber="1082">
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M a t e r i a l e x a m i n e d:
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<typeStatus id="54E78853FFBA2D77F7DAFB41FD8DFB7A" box="[500,582,1254,1276]" pageId="1" pageNumber="1082" type="holotype">Holotype</typeStatus>
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male in a honey-coloured block of Copal, antennae and legs bent ventrally, dorsal and ventral side visible.
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</paragraph>
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</subSubSection>
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<subSubSection id="C346657AFFBA2D75F69CFA84FE96FC95" lastPageId="3" lastPageNumber="1084" pageId="1" pageNumber="1082" type="description">
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<paragraph id="8BE336F1FFBA2D77F69CFA84FE19FAF5" blockId="1.[178,1085,1315,1395]" pageId="1" pageNumber="1082">
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D e s c r i p t i o n: Body length, 3.00 mm; width,
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<quantity id="4CA49B14FFBA2D77F4CBFA84FC83FABE" box="[741,840,1315,1337]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" pageId="1" pageNumber="1082" unit="mm" value="1.55">1.55 mm</quantity>
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; colouration uniformly yellowish, with some secondary transversal veinlets darker (brownish) on the costal area, antennal segment IV brown.
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</paragraph>
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<paragraph id="8BE336F1FFBA2D77F69CFA24FDBAF9CE" blockId="1.[178,1085,1411,1609]" pageId="1" pageNumber="1082">
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H e a d: Oblong, armed with seven spiny tubercles, an occipital pair, a frontal pair, a jugal pair and a median clypeal tubercle; the occipital tubercles the longest, the clypeal ones longer than the frontal and the jugal pairs; antenniferous lobes short and spiny; sulcus straight and narrow; bucculae long, extending in front of clypeus, apex straight and not contiguous; rostrum long, reaching the fourth visible abdominal segment; antennae long and slender, length of antennal segments: I and II much shorter than III and IV but not measurable; III 1.0 mm; IV
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<quantity id="4CA49B14FFBA2D77F432F994FDA6F9CE" box="[540,621,1587,1608]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="1082" unit="mm" value="0.2">0.2 mm</quantity>
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.
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</paragraph>
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<caption id="DF236679FFB92D74F69CFAB2FE62FAE9" ID-DOI="http://doi.org/10.5281/zenodo.5183222" ID-Zenodo-Dep="5183222" httpUri="https://zenodo.org/record/5183222/files/figure.png" pageId="2" pageNumber="1083" startId="2.[178,218,1301,1323]" targetBox="[179,1083,212,1292]" targetPageId="2">
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<paragraph id="8BE336F1FFB92D74F69CFAB2FE62FAE9" blockId="2.[178,1086,1301,1392]" pageId="2" pageNumber="1083">
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<emphasis id="B928EAE3FFB92D74F69CFAB2FECBFAAC" bold="true" box="[178,256,1301,1323]" pageId="2" pageNumber="1083">Figs 1-6</emphasis>
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:
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<taxonomicName id="4C5C4D72FFB92D74F722FAB1FE96FAAA" box="[268,349,1302,1324]" class="Insecta" family="Tingidae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="1083" phylum="Arthropoda" rank="family">Tingidae</taxonomicName>
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in Madagascan Copal: (
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<emphasis id="B928EAE3FFB92D74F411FAB2FD81FAAC" bold="true" box="[575,586,1301,1322]" pageId="2" pageNumber="1083">1</emphasis>
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)
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<taxonomicName id="4C5C4D72FFB92D74F477FAB1FCAAFAAD" authority="Guilbert & Heiss, 2016" authorityName="Guilbert & Heiss" authorityYear="2016" box="[601,865,1302,1323]" class="Insecta" family="Tingidae" genus="Phatnoma" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="1083" phylum="Arthropoda" rank="species" species="madadascariensis" status="sp. nov.">
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<emphasis id="B928EAE3FFB92D74F477FAB1FCAAFAAD" box="[601,865,1302,1323]" italics="true" pageId="2" pageNumber="1083">Phatnoma madadascariensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21B5798FFB92D74F549FAB1FC60FAAD" box="[871,939,1302,1323]" pageId="2" pageNumber="1083" rank="species">nov.sp.</taxonomicNameLabel>
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, holotype male, dorsal view; (
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<emphasis id="B928EAE3FFB92D74F71AFA8BFEF4FAC7" bold="true" box="[308,319,1324,1345]" pageId="2" pageNumber="1083">2</emphasis>
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) ditto, ventral view; (
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<emphasis id="B928EAE3FFB92D74F43FFA8BFDD7FAC7" bold="true" box="[529,540,1324,1345]" pageId="2" pageNumber="1083">3</emphasis>
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)
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<taxonomicName id="4C5C4D72FFB92D74F402FA8BFD39FAC4" authorityName="HORVATH" authorityYear="1910" box="[556,754,1324,1346]" class="Insecta" family="Tingidae" genus="Cysteochila" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="1083" phylum="Arthropoda" rank="species" species="impressa">
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<emphasis id="B928EAE3FFB92D74F402FA8BFD39FAC4" box="[556,754,1324,1346]" italics="true" pageId="2" pageNumber="1083">Cysteochila impressa</emphasis>
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</taxonomicName>
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, male dorsal view; (
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<emphasis id="B928EAE3FFB92D74F599FA8BFC09FAC7" bold="true" box="[951,962,1324,1345]" pageId="2" pageNumber="1083">4</emphasis>
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)
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<taxonomicName id="4C5C4D72FFB92D74F5FCFA8BFF2FFADE" authority="Guilbert & Heiss, 2016" authorityName="Guilbert & Heiss" authorityYear="2016" class="Insecta" family="Tingidae" genus="Cysteochila" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="1083" phylum="Arthropoda" rank="species" species="copal" status="sp. nov.">
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<emphasis id="B928EAE3FFB92D74F5FCFA8BFF2FFADE" italics="true" pageId="2" pageNumber="1083">Cysteochila copal</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21B5798FFB92D74F6C4FAE4FEE5FADE" box="[234,302,1347,1368]" pageId="2" pageNumber="1083" rank="species">nov.sp.</taxonomicNameLabel>
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, holotype female, dorsal view; (
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<emphasis id="B928EAE3FFB92D74F47CFAE4FD96FADE" bold="true" box="[594,605,1347,1368]" pageId="2" pageNumber="1083">5</emphasis>
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) ditto, ventral view; (
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<emphasis id="B928EAE3FFB92D74F50AFAE4FCE4FADE" bold="true" box="[804,815,1347,1368]" pageId="2" pageNumber="1083">6</emphasis>
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)
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<taxonomicName id="4C5C4D72FFB92D74F513FAE4FBCBFADF" authorityName="HORVATH" authorityYear="1910" box="[829,1024,1347,1369]" class="Insecta" family="Tingidae" genus="Cysteochila" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="1083" phylum="Arthropoda" rank="species" species="impressa">
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<emphasis id="B928EAE3FFB92D74F513FAE4FBCBFADF" box="[829,1024,1347,1369]" italics="true" pageId="2" pageNumber="1083">Cysteochila impressa</emphasis>
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</taxonomicName>
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, male, ventral view. Scales 1 mm.
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</paragraph>
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</caption>
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<paragraph id="8BE336F1FFB92D74F69CFAD9FE31F9C2" blockId="2.[178,1086,1405,1604]" pageId="2" pageNumber="1083">
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P r o n o t u m Wider than long, length
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<quantity id="4CA49B14FFB92D74F444FADAFD00FA15" box="[618,715,1405,1427]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" pageId="2" pageNumber="1083" unit="mm" value="0.58">0.58 mm</quantity>
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; width
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<quantity id="4CA49B14FFB92D74F532FAD9FCA4FA15" box="[796,879,1406,1427]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="2" pageNumber="1083" unit="mm" value="1.1">1.1 mm</quantity>
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; tricarinate, carinae with one row of areolae, the median carina slightly higher than the lateral ones; paranota wide and expanded anterolaterally, widening in front, with five areolae at its widest part, with two anterolateral angles projecting like a spiniform process and directed forward; the posterior pronotal process reduced to a row of areolae; rostral groove wider on meso- and metasternum than bucculae, their lateral laminae straight, with one row of areolae. Scutellum reduced to a tubercle.
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</paragraph>
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<paragraph id="8BE336F1FFB82D75F69CFF7DFBD1FE49" blockId="3.[178,1085,217,464]" pageId="3" pageNumber="1084">
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H e m e l y t r a: Length
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<quantity id="4CA49B14FFB82D75F7E4FF7DFDE7FF69" box="[458,556,218,239]" metricMagnitude="-3" metricUnit="m" metricValue="1.92" pageId="3" pageNumber="1084" unit="mm" value="1.92">1.92 mm</quantity>
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; width
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<quantity id="4CA49B14FFB82D75F4AEFF7EFD29FF69" box="[640,738,217,239]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" pageId="3" pageNumber="1084" unit="mm" value="1.55">1.55 mm</quantity>
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; lateral margin rounded, surface divided into the usual areas, all areolae round and of the same size; R+M and Cu veins raised with one row of areolae, these are larger than those of the hemelytral areas; costal area almost of the same width along its whole length, with four to five small round areolae, six at the base and about four at the apex; subcostal area wider than costal area with six areolae wide at its widest part, with three or four secondary transversal veins, dividing the area into four to five subareas; discoidal area longer than half the length of hemeytra with five areolae at its widest part, with at least two secondary transversal veins dividing the area; sutural area enlarging backward with nine areolae at the apex.
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</paragraph>
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<paragraph id="8BE336F1FFB82D75F69CFE79FD38FE72" blockId="3.[178,755,478,500]" box="[178,755,478,500]" pageId="3" pageNumber="1084">E t y m o l o g y: Named after the country of its origin.</paragraph>
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<paragraph id="8BE336F1FFB82D75F69CFDA5FE96FC95" blockId="3.[178,1086,514,788]" pageId="3" pageNumber="1084">
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D i s c u s s i o n: The general habitus of his new species is very similar to
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<taxonomicName id="4C5C4D72FFB82D75F5FBFDA5FE7BFDB5" authority="SCHOUTEDEN 1957" authorityName="SCHOUTEDEN" authorityYear="1957" class="Insecta" family="Tingidae" genus="Phatnoma" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="1084" phylum="Arthropoda" rank="species" species="hova">
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<emphasis id="B928EAE3FFB82D75F5FBFDA5FF28FDB5" italics="true" pageId="3" pageNumber="1084">Phatnoma hova</emphasis>
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<bibRefCitation id="EFCD4B00FFB82D75F6C5FDB9FE7BFDB5" author="SCHOUTEDEN H" box="[235,432,541,563]" pageId="3" pageNumber="1084" pagination="162 - 168" refId="ref4593" refString="SCHOUTEDEN H. (1957): Tingides de Madagascar. - Revue de Zoologie et Botanique Africaine 55: 162 - 168." type="journal article" year="1957">SCHOUTEDEN 1957</bibRefCitation>
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</taxonomicName>
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, which is the only known species of
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<taxonomicName id="4C5C4D72FFB82D75F51EFDB9FC53FDB5" authorityName="Fieber" authorityYear="1844" box="[816,920,542,563]" class="Insecta" family="Tingidae" genus="Phatnoma" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="1084" phylum="Arthropoda" rank="genus">
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<emphasis id="B928EAE3FFB82D75F51EFDB9FC53FDB5" box="[816,920,542,563]" italics="true" pageId="3" pageNumber="1084">Phatnoma</emphasis>
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</taxonomicName>
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from
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<collectingCountry id="F34B7661FFB82D75F5F7FDB9FF1FFDC9" name="Madagascar" pageId="3" pageNumber="1084">Madagascar</collectingCountry>
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. However, it shows a different combination of areolae on the hemelytra and paranota. There are four rows of areolae at the apex of the costal area and six at the base, while
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<taxonomicName id="4C5C4D72FFB82D75F209FDF1FF28FD01" authorityName="SCHOUTEDEN" authorityYear="1957" class="Insecta" family="Tingidae" genus="Phatnoma" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="1084" phylum="Arthropoda" rank="species" species="hova">
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<emphasis id="B928EAE3FFB82D75F209FDF1FF28FD01" italics="true" pageId="3" pageNumber="1084">P. hova</emphasis>
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</taxonomicName>
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has three rows of areolae at the apex and seven at the base. The subcostal area of
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<taxonomicName id="4C5C4D72FFB82D75F209FDD5FEA2FD25" authority="Guilbert & Heiss, 2016" authorityName="Guilbert & Heiss" authorityYear="2016" class="Insecta" family="Tingidae" genus="Phatnoma" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="1084" phylum="Arthropoda" rank="species" species="madagascariensis" status="sp. nov.">
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<emphasis id="B928EAE3FFB82D75F209FDD5FEA2FD25" italics="true" pageId="3" pageNumber="1084">P. madagascariensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21B5798FFB82D75F75AFD29FE75FD25" box="[372,446,654,675]" pageId="3" pageNumber="1084" rank="species">nov.sp.</taxonomicNameLabel>
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is six areolae wide, while there are seven in
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<taxonomicName id="4C5C4D72FFB82D75F59CFD29FBCFFD25" authorityName="SCHOUTEDEN" authorityYear="1957" box="[946,1028,654,675]" class="Insecta" family="Tingidae" genus="Phatnoma" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="1084" phylum="Arthropoda" rank="species" species="hova">
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<emphasis id="B928EAE3FFB82D75F59CFD29FBCFFD25" box="[946,1028,654,675]" italics="true" pageId="3" pageNumber="1084">P. hova</emphasis>
|
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</taxonomicName>
|
|
. The discoidal area is five areolae wide and shows six areolae in
|
|
<taxonomicName id="4C5C4D72FFB82D75F530FD0DFCA6FD39" authorityName="SCHOUTEDEN" authorityYear="1957" box="[798,877,682,703]" class="Insecta" family="Tingidae" genus="Phatnoma" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="1084" phylum="Arthropoda" rank="species" species="hova">
|
|
<emphasis id="B928EAE3FFB82D75F530FD0DFCA6FD39" box="[798,877,682,703]" italics="true" pageId="3" pageNumber="1084">P. hova</emphasis>
|
|
</taxonomicName>
|
|
. The width of paranota of this new species ranges from three to five areolae, while there are four areolae in
|
|
<taxonomicName id="4C5C4D72FFB82D75F69CFD45FEC9FD71" authorityName="SCHOUTEDEN" authorityYear="1957" box="[178,258,738,759]" class="Insecta" family="Tingidae" genus="Phatnoma" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="1084" phylum="Arthropoda" rank="species" species="hova">
|
|
<emphasis id="B928EAE3FFB82D75F69CFD45FEC9FD71" box="[178,258,738,759]" italics="true" pageId="3" pageNumber="1084">P. hova</emphasis>
|
|
</taxonomicName>
|
|
. In addition, the areolae of
|
|
<emphasis id="B928EAE3FFB82D75F405FD45FD29FD71" box="[555,738,738,759]" italics="true" pageId="3" pageNumber="1084">madagascariensis</emphasis>
|
|
<taxonomicNameLabel id="A21B5798FFB82D75F4C5FD45FCFEFD71" box="[747,821,738,759]" pageId="3" pageNumber="1084" rank="species">nov.sp.</taxonomicNameLabel>
|
|
are generally larger than those of
|
|
<taxonomicName id="4C5C4D72FFB82D75F727FD59FE9CFC95" authorityName="SCHOUTEDEN" authorityYear="1957" box="[265,343,766,787]" class="Insecta" family="Tingidae" genus="Phatnoma" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="1084" phylum="Arthropoda" rank="species" species="hova">
|
|
<emphasis id="B928EAE3FFB82D75F727FD59FE9CFC95" box="[265,343,766,787]" italics="true" pageId="3" pageNumber="1084">P. hova</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |