treatments-xml/data/03/F5/87/03F58791D3210C17C9CF9A8EFE9FFB4B.xml

<document id="511BFE51DDFD56109AE181DE75475F0B" ID-DOI="10.5281/zenodo.191909" ID-GBIF-Dataset="b5235354-ef06-4a49-b9d1-5498e97c3639" ID-ISSN="1175-5326" ID-Zenodo-Dep="191909" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461137864308" checkinUser="plazi" docAuthor="Ng, Peter K. L., Rahayu, Dwi Listyo &amp; Naser, Murtada D." docDate="2009" docId="03F58791D3210C17C9CF9A8EFE9FFB4B" docLanguage="en" docName="zt02312p026.pdf" docOrigin="Zootaxa 2312" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Opusia Ng, Rahayu &amp; Naser, 2009, new genus" docType="treatment" docVersion="9" lastPageNumber="6" masterDocId="FFCCFFE9D3200C12C9589874FFC1FFD4" masterDocTitle="The Camptandriidae of Iraq, with description of a new genus and notes on Leptochryseus Al-Khayat &amp; Jones, 1996 (Crustacea: Decapoda: Brachyura)" masterLastPageNumber="26" masterPageNumber="1" pageNumber="2" updateTime="1698598725934" updateUser="plazi">
<mods:mods id="5CF6DEF679F4243C878F4801E96EBAE5" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="04452049276FEBDC47688D306D761E1C">
<mods:title id="717E847BACAC092DD525431FE49FCE1F">The Camptandriidae of Iraq, with description of a new genus and notes on Leptochryseus Al-Khayat &amp; Jones, 1996 (Crustacea: Decapoda: Brachyura)</mods:title>
</mods:titleInfo>
<mods:name id="5F55306A9898EC73563545A3CCA797C6" type="personal">
<mods:role id="81967CAEB84EBF07E62646FFA7FB272A">
<mods:roleTerm id="4D4D03B1ADAA1DA956691DCA2D44CAEF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6E245565BFD72689E4F68511AAEB1693">Ng, Peter K. L.</mods:namePart>
</mods:name>
<mods:name id="4A276B816424B5B5F3A5F5D38678F79C" type="personal">
<mods:role id="3051B365CA8C8EE2793B5F0A20D3B9AE">
<mods:roleTerm id="318D4508EEB15FC83C7E27C3B1D925DB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2F1BA88F4B42FA6EE8D11514DEC461C8">Rahayu, Dwi Listyo</mods:namePart>
</mods:name>
<mods:name id="5C795E795310E334B32B63DDE85F736C" type="personal">
<mods:role id="4E4B387340335278956CA4A74EFF568C">
<mods:roleTerm id="B2D1733E9F1B16101D5EAE25B3B6612F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FBF26C4E386853086BEE49018F4D4CB6">Naser, Murtada D.</mods:namePart>
</mods:name>
<mods:typeOfResource id="95F4BFF9A1DA0460485B0E1510DE9956">text</mods:typeOfResource>
<mods:relatedItem id="C14B9762CBBF37A81A784E42172F2A0B" type="host">
<mods:titleInfo id="C939EF7955AE74C3363BC165C124E75F">
<mods:title id="1C14ACBB527772342228283482DDEF8A">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="BD69618D688E33A769680302066B5ACA">
<mods:date id="A564C754E960F4B3198CFDAB601AE2F1">2009</mods:date>
<mods:detail id="E4240CB78F59CDFC434DB72747CDCD9B" type="volume">
<mods:number id="CAE9459B0F8ED72AB4CE199119DF5077">2312</mods:number>
</mods:detail>
<mods:extent id="F0417BCB913EA46DAB338D965B8B479B" unit="page">
<mods:start id="9FA219646AB26277688CF2975FD47DD5">1</mods:start>
<mods:end id="281CA86D38308021725F6AFD7357F760">26</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="89082D387EF5C934BC6ABA83A6A5A78A">journal article</mods:classification>
<mods:identifier id="2EE1B374937A11FA2A532A75E7EDBA14" type="DOI">10.5281/zenodo.191909</mods:identifier>
<mods:identifier id="CDB1000A67761798A259A6289E25216D" type="GBIF-Dataset">b5235354-ef06-4a49-b9d1-5498e97c3639</mods:identifier>
<mods:identifier id="E9BFBC30395F05596255082D61006B09" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="4A5FD84DDC56235F678B5F00BA48F379" type="Zenodo-Dep">191909</mods:identifier>
</mods:mods>
<treatment id="03F58791D3210C17C9CF9A8EFE9FFB4B" ID-DOI="http://doi.org/10.5281/zenodo.6226783" ID-GBIF-Taxon="119627624" ID-Zenodo-Dep="6226783" LSID="urn:lsid:plazi:treatment:03F58791D3210C17C9CF9A8EFE9FFB4B" httpUri="http://treatment.plazi.org/id/03F58791D3210C17C9CF9A8EFE9FFB4B" lastPageId="5" lastPageNumber="6" pageId="1" pageNumber="2">
<subSubSection id="C346650CD3210C13C9CF9A8EFEB7FCC0" box="[151,374,762,788]" pageId="1" pageNumber="2" type="nomenclature">
<paragraph id="8BE33687D3210C13C9CF9A8EFEB7FCC0" blockId="1.[151,975,727,823]" box="[151,374,762,788]" pageId="1" pageNumber="2">
<heading id="D0AB81EBD3210C13C9CF9A8EFEB7FCC0" bold="true" box="[151,374,762,788]" fontSize="11" level="1" pageId="1" pageNumber="2" reason="1">
<emphasis id="B928EA95D3210C13C9CF9A8EFEB7FCC0" bold="true" box="[151,374,762,788]" pageId="1" pageNumber="2">
<taxonomicName id="4C5C4D04D3210C13C9CF9A8EFF2CFCC0" ID-CoL="7PR43" box="[151,237,762,788]" class="Malacostraca" family="Camptandriidae" genus="Opusia" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B928EA95D3210C13C9CF9A8EFF2CFCC0" bold="true" box="[151,237,762,788]" italics="true" pageId="1" pageNumber="2">Opusia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21B57EED3210C13C9AC9A8EFEB7FCC0" box="[244,374,762,788]" pageId="1" pageNumber="2" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C346650CD3210C13C9CF9B69FC0EFCE3" box="[151,975,797,823]" pageId="1" pageNumber="2" type="materials_examined">
<paragraph id="8BE33687D3210C13C9CF9B69FC0EFCE3" blockId="1.[151,975,727,823]" box="[151,975,797,823]" pageId="1" pageNumber="2">
<emphasis id="B928EA95D3210C13C9CF9B69FEF8FCE3" bold="true" box="[151,313,797,823]" pageId="1" pageNumber="2">
<typeStatus id="54E78825D3210C13C9CF9B69FF15FCE3" box="[151,212,797,823]" pageId="1" pageNumber="2">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C5C4D04D3210C13C8189B69FD75FCE3" authority="Alcock, 1900" authorityName="Alcock" authorityYear="1900" box="[320,692,797,823]" class="Malacostraca" family="Camptandriidae" genus="Tylodiplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="B928EA95D3210C13C8189B69FDCDFCE2" box="[320,524,797,822]" italics="true" pageId="1" pageNumber="2">Tylodiplax indica</emphasis>
<bibRefCitation id="EFCD4B76D3210C13CB4C9B69FD75FCE3" author="Alcock" box="[532,692,797,823]" pageId="1" pageNumber="2" refString="Alcock, A. (1900) Materials for a carcinological fauna of India. No. 6. The Brachyura Catometopa or Grapsoidea. Journal of the Asiatic Society of Bengal, Calcutta, 69 (3), 279 - 486." type="journal article" year="1900">Alcock, 1900</bibRefCitation>
</taxonomicName>
, by present designation.
</paragraph>
</subSubSection>
<subSubSection id="C346650CD3210C13C9CF9B1EFEBDF964" pageId="1" pageNumber="2" type="diagnosis">
<paragraph id="8BE33687D3210C13C9CF9B1EFEBDF964" blockId="1.[151,1437,874,2022]" pageId="1" pageNumber="2">
<emphasis id="B928EA95D3210C13C9CF9B1EFED8FC50" bold="true" box="[151,281,874,900]" pageId="1" pageNumber="2">Diagnosis.</emphasis>
Carapace transversely subovate, broader than long, weakly convex, regions well demarcated by grooves; anterolateral margin not separated from posterolateral margin by clearly demarcated angle; pterygostomial region swollen, produced beyond anterolateral margin, visible dorsally; dorsolateral margin not demarcated; posterolateral margin swollen, rounded; posterior margin weakly sinuous, thickened, with submarginal ridge (
<figureCitation id="13672A02D3210C13C8D99C71FE10FBCB" box="[385,465,1029,1055]" captionStart="FIGURE 1" captionStartId="2.[151,259,1488,1512]" captionTargetBox="[151,1434,335,1470]" captionTargetId="figure@2.[151,1436,335,1470]" captionTargetPageId="2" captionText="FIGURE 1. Overall views. A, Opusia indicum (Alcock, 1900), male (6.6 x 11.3 mm) (ZRC 2009.832), Iraq; B, Manningis arabicum (Jones &amp; Clayton, 1983), male (3.5 x 5.3 mm) (ZRC 2009.1097), Iraq." httpUri="https://zenodo.org/record/191910/files/figure.png" pageId="1" pageNumber="2">Figs. 1</figureCitation>
A, 2A). Front straight, entire from dorsal view, slightly thickened, lateral angles rounded, junction between base, supraorbital margin rounded; frontomedial projection broadly triangular, ending in truncate tip (
<figureCitation id="13672A02D3210C13C8F79C26FDC1FBB8" box="[431,512,1106,1132]" captionStart="FIGURE 1" captionStartId="2.[151,259,1488,1512]" captionTargetBox="[151,1434,335,1470]" captionTargetId="figure@2.[151,1436,335,1470]" captionTargetPageId="2" captionText="FIGURE 1. Overall views. A, Opusia indicum (Alcock, 1900), male (6.6 x 11.3 mm) (ZRC 2009.832), Iraq; B, Manningis arabicum (Jones &amp; Clayton, 1983), male (3.5 x 5.3 mm) (ZRC 2009.1097), Iraq." httpUri="https://zenodo.org/record/191910/files/figure.png" pageId="1" pageNumber="2">Figs. 1</figureCitation>
A, 2A). Eyestalks long, slender, with long, plumose setae (
<figureCitation id="13672A02D3210C13CD8F9C26FAE0FBB8" box="[1239,1313,1106,1132]" captionStart="FIGURE 2" captionStartId="3.[151,256,1918,1942]" captionTargetBox="[185,1383,306,1917]" captionTargetId="figure@3.[180,1404,301,1980]" captionTargetPageId="3" captionText="FIGURE 2. Opusia indicum (Alcock, 1900), male (6. 9 x 10.1 mm) (USNM 191107), Pakistan. A, carapace; B, outer view of left third maxilliped; C, male abdomen; D, frontal view of carapace showing orbit, eye and epistome; E, anterior part of thoracic sternum; F, outer view of right chela; G, left G 1; H, distal part of left G 1. Scales: A, C – G = 1.0 mm; B = 0.5 mm; H = 0.2 mm." httpUri="https://zenodo.org/record/191911/files/figure.png" pageId="1" pageNumber="2">Fig. 2</figureCitation>
A). Orbits sparsely setose, about as wide as front, external orbital margin distinct, triangular; supraorbital margin sinuous; inner two-thirds of infraorbital margin moderately cristate, inner infraorbital tooth distinctly demarcated from rest of infraorbital margin by broadly triangular notch (
<figureCitation id="13672A02D3210C13CABE9CB2FBEDFB34" box="[998,1068,1222,1248]" captionStart="FIGURE 2" captionStartId="3.[151,256,1918,1942]" captionTargetBox="[185,1383,306,1917]" captionTargetId="figure@3.[180,1404,301,1980]" captionTargetPageId="3" captionText="FIGURE 2. Opusia indicum (Alcock, 1900), male (6. 9 x 10.1 mm) (USNM 191107), Pakistan. A, carapace; B, outer view of left third maxilliped; C, male abdomen; D, frontal view of carapace showing orbit, eye and epistome; E, anterior part of thoracic sternum; F, outer view of right chela; G, left G 1; H, distal part of left G 1. Scales: A, C – G = 1.0 mm; B = 0.5 mm; H = 0.2 mm." httpUri="https://zenodo.org/record/191911/files/figure.png" pageId="1" pageNumber="2">Fig. 2</figureCitation>
A); suborbital ridge moderately developed, not visible dorsally; transverse pterygostomial sulcus present, open. Epistome with anterior half separated from posterior half by ridge, anterior half sunken slightly; posteromedial tooth large, long, acute, posterior margin on either side lobate under basal segment of antenna (
<figureCitation id="13672A02D3210C13CA829D4FFBE0FA81" box="[986,1057,1339,1365]" captionStart="FIGURE 2" captionStartId="3.[151,256,1918,1942]" captionTargetBox="[185,1383,306,1917]" captionTargetId="figure@3.[180,1404,301,1980]" captionTargetPageId="3" captionText="FIGURE 2. Opusia indicum (Alcock, 1900), male (6. 9 x 10.1 mm) (USNM 191107), Pakistan. A, carapace; B, outer view of left third maxilliped; C, male abdomen; D, frontal view of carapace showing orbit, eye and epistome; E, anterior part of thoracic sternum; F, outer view of right chela; G, left G 1; H, distal part of left G 1. Scales: A, C – G = 1.0 mm; B = 0.5 mm; H = 0.2 mm." httpUri="https://zenodo.org/record/191911/files/figure.png" pageId="1" pageNumber="2">Fig. 2</figureCitation>
D). Ischium of third maxillipeds longer than merus, with inner distal angle slightly produced; outer distal angle of merus produced, lobate, inner angle excavated for insertion of palp (
<figureCitation id="13672A02D3210C13CBE99DFCFD3DFA76" box="[689,764,1416,1442]" captionStart="FIGURE 2" captionStartId="3.[151,256,1918,1942]" captionTargetBox="[185,1383,306,1917]" captionTargetId="figure@3.[180,1404,301,1980]" captionTargetPageId="3" captionText="FIGURE 2. Opusia indicum (Alcock, 1900), male (6. 9 x 10.1 mm) (USNM 191107), Pakistan. A, carapace; B, outer view of left third maxilliped; C, male abdomen; D, frontal view of carapace showing orbit, eye and epistome; E, anterior part of thoracic sternum; F, outer view of right chela; G, left G 1; H, distal part of left G 1. Scales: A, C – G = 1.0 mm; B = 0.5 mm; H = 0.2 mm." httpUri="https://zenodo.org/record/191911/files/figure.png" pageId="1" pageNumber="2">Fig. 2</figureCitation>
B). Outer margins of chelipeds with long setae, fingers narrow, about twice palm length, setose, tips blade-like, sharp (
<figureCitation id="13672A02D3210C13CADC9DDAFC0DFA1C" box="[900,972,1454,1480]" captionStart="FIGURE 2" captionStartId="3.[151,256,1918,1942]" captionTargetBox="[185,1383,306,1917]" captionTargetId="figure@3.[180,1404,301,1980]" captionTargetPageId="3" captionText="FIGURE 2. Opusia indicum (Alcock, 1900), male (6. 9 x 10.1 mm) (USNM 191107), Pakistan. A, carapace; B, outer view of left third maxilliped; C, male abdomen; D, frontal view of carapace showing orbit, eye and epistome; E, anterior part of thoracic sternum; F, outer view of right chela; G, left G 1; H, distal part of left G 1. Scales: A, C – G = 1.0 mm; B = 0.5 mm; H = 0.2 mm." httpUri="https://zenodo.org/record/191911/files/figure.png" pageId="1" pageNumber="2">Fig. 2</figureCitation>
F); chelipeds in male similar to female, but slightly stouter. Ambulatory legs robust, setose, P3 broadest, P5 with dactylus directed upwards (
<figureCitation id="13672A02D3210C13CC689DA1FAB0FA3B" box="[1328,1393,1493,1519]" captionStart="FIGURE 1" captionStartId="2.[151,259,1488,1512]" captionTargetBox="[151,1434,335,1470]" captionTargetId="figure@2.[151,1436,335,1470]" captionTargetPageId="2" captionText="FIGURE 1. Overall views. A, Opusia indicum (Alcock, 1900), male (6.6 x 11.3 mm) (ZRC 2009.832), Iraq; B, Manningis arabicum (Jones &amp; Clayton, 1983), male (3.5 x 5.3 mm) (ZRC 2009.1097), Iraq." httpUri="https://zenodo.org/record/191910/files/figure.png" pageId="1" pageNumber="2">Fig. 1</figureCitation>
A); segments relatively less setose in females. Anterior part of male thoracic sternum rounded (
<figureCitation id="13672A02D3210C13CDB39D88FAF5F9C2" box="[1259,1332,1532,1558]" captionStart="FIGURE 2" captionStartId="3.[151,256,1918,1942]" captionTargetBox="[185,1383,306,1917]" captionTargetId="figure@3.[180,1404,301,1980]" captionTargetPageId="3" captionText="FIGURE 2. Opusia indicum (Alcock, 1900), male (6. 9 x 10.1 mm) (USNM 191107), Pakistan. A, carapace; B, outer view of left third maxilliped; C, male abdomen; D, frontal view of carapace showing orbit, eye and epistome; E, anterior part of thoracic sternum; F, outer view of right chela; G, left G 1; H, distal part of left G 1. Scales: A, C – G = 1.0 mm; B = 0.5 mm; H = 0.2 mm." httpUri="https://zenodo.org/record/191911/files/figure.png" pageId="1" pageNumber="2">Fig. 2</figureCitation>
E). Male abdomen with all sutures distinct, complete, somite 1 slightly wider than somite 2, not reaching P5 coxae; somite 5 slightly constricted close to proximal end; telson about as long as broad (
<figureCitation id="13672A02D3210C13CD389E3DFB66F9B7" box="[1120,1191,1609,1635]" captionStart="FIGURE 2" captionStartId="3.[151,256,1918,1942]" captionTargetBox="[185,1383,306,1917]" captionTargetId="figure@3.[180,1404,301,1980]" captionTargetPageId="3" captionText="FIGURE 2. Opusia indicum (Alcock, 1900), male (6. 9 x 10.1 mm) (USNM 191107), Pakistan. A, carapace; B, outer view of left third maxilliped; C, male abdomen; D, frontal view of carapace showing orbit, eye and epistome; E, anterior part of thoracic sternum; F, outer view of right chela; G, left G 1; H, distal part of left G 1. Scales: A, C – G = 1.0 mm; B = 0.5 mm; H = 0.2 mm." httpUri="https://zenodo.org/record/191911/files/figure.png" pageId="1" pageNumber="2">Fig. 2</figureCitation>
C). Female abdomen with all somites free, telson broadly triangular. G1 gently bent, apex conical with numerous strong subapical spines (
<figureCitation id="13672A02D3210C13C9A89EE2FEF4F964" box="[240,309,1686,1712]" captionStart="FIGURE 2" captionStartId="3.[151,256,1918,1942]" captionTargetBox="[185,1383,306,1917]" captionTargetId="figure@3.[180,1404,301,1980]" captionTargetPageId="3" captionText="FIGURE 2. Opusia indicum (Alcock, 1900), male (6. 9 x 10.1 mm) (USNM 191107), Pakistan. A, carapace; B, outer view of left third maxilliped; C, male abdomen; D, frontal view of carapace showing orbit, eye and epistome; E, anterior part of thoracic sternum; F, outer view of right chela; G, left G 1; H, distal part of left G 1. Scales: A, C – G = 1.0 mm; B = 0.5 mm; H = 0.2 mm." httpUri="https://zenodo.org/record/191911/files/figure.png" pageId="1" pageNumber="2">Fig. 2</figureCitation>
G, H).
</paragraph>
</subSubSection>
<subSubSection id="C346650CD3210C13C99E9EC9FB0DF92A" pageId="1" pageNumber="2" type="etymology">
<paragraph id="8BE33687D3210C13C99E9EC9FB0DF92A" blockId="1.[151,1437,874,2022]" pageId="1" pageNumber="2">
<emphasis id="B928EA95D3210C13C99E9EC9FE92F903" bold="true" box="[198,339,1725,1751]" pageId="1" pageNumber="2">Etymology.</emphasis>
The genus name is arbitrarily derived from the Latin 
<emphasis id="B928EA95D3210C13CD489EC9FBA4F902" box="[1040,1125,1725,1750]" italics="true" pageId="1" pageNumber="2">opimus</emphasis>
meaning &quot;well-fed, fat&quot;, alluding to the rounded and swollen appearance of the 
<typeStatus id="54E78825D3210C13CA409E90FC8AF92A" box="[792,843,1764,1790]" pageId="1" pageNumber="2">type</typeStatus>
species. The gender is feminine.
</paragraph>
</subSubSection>
<subSubSection id="C346650CD3210C11C99E9F7EFEB8FF2B" lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2" type="discussion">
<paragraph id="8BE33687D3210C10C99E9F7EFA54FEF2" blockId="1.[151,1437,874,2022]" lastBlockId="2.[151,1437,152,294]" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">
<emphasis id="B928EA95D3210C13C99E9F7EFE81F8F0" bold="true" box="[198,320,1802,1828]" pageId="1" pageNumber="2">Remarks.</emphasis>
The monotypic 
<taxonomicName id="4C5C4D04D3210C13CB509F7FFD9CF8F0" box="[520,605,1803,1828]" class="Malacostraca" family="Camptandriidae" genus="Opusia" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B928EA95D3210C13CB509F7FFD9CF8F0" box="[520,605,1803,1828]" italics="true" pageId="1" pageNumber="2">Opusia</emphasis>
</taxonomicName>
<emphasis id="B928EA95D3210C13CB3F9F7EFD2AF8F0" bold="true" box="[615,747,1802,1828]" pageId="1" pageNumber="2">
<taxonomicNameLabel id="A21B57EED3210C13CB3F9F7EFD2AF8F0" box="[615,747,1802,1828]" pageId="1" pageNumber="2" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
is only superficially similar to 
<taxonomicName id="4C5C4D04D3210C13CD319F7FFA5DF8F1" authority="De Man, 1895" authorityName="De Man" authorityYear="1895" box="[1129,1436,1803,1829]" class="Malacostraca" family="Camptandriidae" genus="Tylodiplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B928EA95D3210C13CD319F7FFB25F8F0" box="[1129,1252,1803,1828]" italics="true" pageId="1" pageNumber="2">Tylodiplax</emphasis>
De 
<bibRefCitation id="EFCD4B76D3210C13CC429F7FFA5DF8F1" author="Man" box="[1306,1436,1803,1829]" pageId="1" pageNumber="2" refString="Man, J. G. De (1895) Bericht uber die von Herrn Schiffscapitan Storm zur Atjeh, an den westlichen Kusten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8, 485 - 609, pls. 12 - 14." type="journal article" year="1895">Man, 1895</bibRefCitation>
</taxonomicName>
(
<typeStatus id="54E78825D3210C13C9C79F45FF14F89F" box="[159,213,1841,1867]" pageId="1" pageNumber="2">type</typeStatus>
species 
<taxonomicName id="4C5C4D04D3210C13C8659F45FCEEF89F" authority="De Man, 1895" authorityName="De Man" authorityYear="1895" box="[317,815,1841,1867]" class="Malacostraca" family="Camptandriidae" genus="Tylodiplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="tetratylophora">
<emphasis id="B928EA95D3210C13C8659F45FDB3F89E" box="[317,626,1841,1866]" italics="true" pageId="1" pageNumber="2">Tylodiplax tetratylophora</emphasis>
De 
<bibRefCitation id="EFCD4B76D3210C13CBFE9F45FCEEF89F" author="Man" box="[678,815,1841,1867]" pageId="1" pageNumber="2" refString="Man, J. G. De (1895) Bericht uber die von Herrn Schiffscapitan Storm zur Atjeh, an den westlichen Kusten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8, 485 - 609, pls. 12 - 14." type="journal article" year="1895">Man, 1895</bibRefCitation>
</taxonomicName>
). Members of these two genera do not show strong sexual dimorphism of the chelipeds, have a straight front, possess long, plumose setae on their eyestalks, the posterior margin of the epistome is lobed on either side, the dactylus of the P5 is upturned and the carapace lateral margins diverge posteriorly. 
<taxonomicName id="4C5C4D04D3210C13CB609FD1FD4CF86A" box="[568,653,1957,1982]" class="Malacostraca" family="Camptandriidae" genus="Opusia" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B928EA95D3210C13CB609FD1FD4CF86A" box="[568,653,1957,1982]" italics="true" pageId="1" pageNumber="2">Opusia</emphasis>
</taxonomicName>
<emphasis id="B928EA95D3210C13CBCC9FD1FCD7F86B" bold="true" box="[660,790,1957,1983]" pageId="1" pageNumber="2">
<taxonomicNameLabel id="A21B57EED3210C13CBCC9FD1FCD7F86B" box="[660,790,1957,1983]" pageId="1" pageNumber="2" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
can easily be distinguished from 
<taxonomicName id="4C5C4D04D3210C13CDF99FD1FADDF86A" box="[1185,1308,1957,1982]" class="Malacostraca" family="Camptandriidae" genus="Tylodiplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B928EA95D3210C13CDF99FD1FADDF86A" box="[1185,1308,1957,1982]" italics="true" pageId="1" pageNumber="2">Tylodiplax</emphasis>
</taxonomicName>
by several key characters. Most obvious is the differently shaped carapace. The carapace in 
<taxonomicName id="4C5C4D04D3210C13CD1D9FB8FB5BF831" box="[1093,1178,1996,2021]" class="Malacostraca" family="Camptandriidae" genus="Opusia" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B928EA95D3210C13CD1D9FB8FB5BF831" box="[1093,1178,1996,2021]" italics="true" pageId="1" pageNumber="2">Opusia</emphasis>
</taxonomicName>
<emphasis id="B928EA95D3210C13CDF99FBFFAE3F831" bold="true" box="[1185,1314,1995,2021]" pageId="1" pageNumber="2">
<taxonomicNameLabel id="A21B57EED3210C13CDF99FBFFAE3F831" box="[1185,1314,1995,2021]" pageId="1" pageNumber="2" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
is broadly ovate with no prominent angles on the anterolateral margins with the exception of the external orbital tooth, and its dorsal surface has no distinct ridges or tubercles (
<figureCitation id="13672A02D3220C10CA0598CAFC71FF0C" box="[861,944,190,216]" captionStart="FIGURE 1" captionStartId="2.[151,259,1488,1512]" captionTargetBox="[151,1434,335,1470]" captionTargetId="figure@2.[151,1436,335,1470]" captionTargetPageId="2" captionText="FIGURE 1. Overall views. A, Opusia indicum (Alcock, 1900), male (6.6 x 11.3 mm) (ZRC 2009.832), Iraq; B, Manningis arabicum (Jones &amp; Clayton, 1983), male (3.5 x 5.3 mm) (ZRC 2009.1097), Iraq." httpUri="https://zenodo.org/record/191910/files/figure.png" pageId="2" pageNumber="3">Figs. 1</figureCitation>
A, 2A). The anterolateral margin of the carapace of 
<taxonomicName id="4C5C4D04D3220C10C8719891FE66FF2A" box="[297,423,229,254]" class="Malacostraca" family="Camptandriidae" genus="Tylodiplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B928EA95D3220C10C8719891FE66FF2A" box="[297,423,229,254]" italics="true" pageId="2" pageNumber="3">Tylodiplax</emphasis>
</taxonomicName>
, however, bears a distinct angle with the posterolateral margins straighter (
<figureCitation id="13672A02D3220C10CC6E9891FA56FF2B" box="[1334,1431,229,255]" captionStart="FIGURE 3. A – D" captionStartId="4.[151,262,1554,1578]" captionTargetBox="[225,1341,213,1553]" captionTargetId="figure@4.[223,1363,194,1599]" captionTargetPageId="4" captionText="FIGURE 3. A – D, Tylodiplax tetratylophora De Man, 1895, male (6.8 x 10.2 mm) (ZRC), Malaysia; E – H, Paratylodiplax blephariskios (Stebbing, 1924), male (5.4 x 8.8 mm) (USNM 137247), South Africa. Scales: A, B, D – F, H = 1.0 mm; C, G = 0.5 mm." httpUri="https://zenodo.org/record/191912/files/figure.png" pageId="2" pageNumber="3">Fig. 3A</figureCitation>
; 
<bibRefCitation id="EFCD4B76D3220C10C9CF9978FE0EFEF2" author="Manning" box="[151,463,268,294]" pageId="2" pageNumber="3" refString="Manning, R. B. &amp; Holthuis, L. B. (1981) West African Brachyuran Crabs (Crustacea: Decapoda). Smithsonian Contributions to Zoology, 306, 1 - 379." type="journal article" year="1981">Manning &amp; Holthuis 1981</bibRefCitation>
: fig. 58a, c), and the anterolateral margin is not as swollen as in 
<taxonomicName id="4C5C4D04D3220C10CDEB9978FAC9FEF1" box="[1203,1288,268,293]" class="Malacostraca" family="Camptandriidae" genus="Opusia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B928EA95D3220C10CDEB9978FAC9FEF1" box="[1203,1288,268,293]" italics="true" pageId="2" pageNumber="3">Opusia</emphasis>
</taxonomicName>
<emphasis id="B928EA95D3220C10CC55997FFA4FFEF1" bold="true" box="[1293,1422,267,293]" pageId="2" pageNumber="3">
<taxonomicNameLabel id="A21B57EED3220C10CC55997FFA4FFEF1" box="[1293,1422,267,293]" pageId="2" pageNumber="3" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
.
</paragraph>
<caption id="DF23660FD3220C10C9CF9DA4FBACF9DE" httpUri="https://zenodo.org/record/191910/files/figure.png" pageId="2" pageNumber="3" targetBox="[151,1434,335,1470]" targetPageId="2">
<paragraph id="8BE33687D3220C10C9CF9DA4FBACF9DE" blockId="2.[151,1437,1488,1546]" pageId="2" pageNumber="3">
<emphasis id="B928EA95D3220C10C9CF9DA4FEE3FA3C" bold="true" box="[151,290,1488,1512]" pageId="2" pageNumber="3">FIGURE 1.</emphasis>
Overall views. A, 
<taxonomicName id="4C5C4D04D3220C10C8A49DA5FCABFA33" authority="Alcock, 1900" authorityName="Alcock" authorityYear="1900" box="[508,874,1488,1512]" class="Malacostraca" family="Camptandriidae" genus="Opusia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="indicum">
<emphasis id="B928EA95D3220C10C8A49DA5FD73FA3C" box="[508,690,1489,1512]" italics="true" pageId="2" pageNumber="3">Opusia indicum</emphasis>
(Alcock, 1900)
</taxonomicName>
, male (6.6 x 11.3 mm) (ZRC 2009.832), Iraq; B, 
<taxonomicName id="4C5C4D04D3220C10C9CF9D87FD45F9DE" authority="Jones &amp; Clayton, 1983" authorityName="Jones &amp; Clayton" authorityYear="1983" box="[151,644,1523,1546]" class="Malacostraca" family="Camptandriidae" genus="Manningis" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="arabicum">
<emphasis id="B928EA95D3220C10C9CF9D87FEB4F9DE" box="[151,373,1523,1546]" italics="true" pageId="2" pageNumber="3">Manningis arabicum</emphasis>
(Jones &amp; Clayton, 1983)
</taxonomicName>
, male (3.5 x 5.3 mm) (ZRC 2009.1097), Iraq.
</paragraph>
</caption>
<paragraph id="8BE33687D3220C11C9CF9E22FEB8FF2B" blockId="2.[151,1437,1622,2034]" lastBlockId="3.[151,1436,151,255]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
In addition, the dorsal carapace surface of 
<taxonomicName id="4C5C4D04D3220C10CBD79E22FCCBF9BB" box="[655,778,1622,1647]" class="Malacostraca" family="Camptandriidae" genus="Tylodiplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B928EA95D3220C10CBD79E22FCCBF9BB" box="[655,778,1622,1647]" italics="true" pageId="2" pageNumber="3">Tylodiplax</emphasis>
</taxonomicName>
bears several prominent ridges and protuberances (
<figureCitation id="13672A02D3220C10CC349E22FF7FF942" captionStart="FIGURE 3. A – D" captionStartId="4.[151,262,1554,1578]" captionTargetBox="[225,1341,213,1553]" captionTargetId="figure@4.[223,1363,194,1599]" captionTargetPageId="4" captionText="FIGURE 3. A – D, Tylodiplax tetratylophora De Man, 1895, male (6.8 x 10.2 mm) (ZRC), Malaysia; E – H, Paratylodiplax blephariskios (Stebbing, 1924), male (5.4 x 8.8 mm) (USNM 137247), South Africa. Scales: A, B, D – F, H = 1.0 mm; C, G = 0.5 mm." httpUri="https://zenodo.org/record/191912/files/figure.png" pageId="2" pageNumber="3">Fig. 3A</figureCitation>
; 
<bibRefCitation id="EFCD4B76D3220C10C9939E08FDC8F942" author="Manning" box="[203,521,1660,1686]" pageId="2" pageNumber="3" refString="Manning, R. B. &amp; Holthuis, L. B. (1981) West African Brachyuran Crabs (Crustacea: Decapoda). Smithsonian Contributions to Zoology, 306, 1 - 379." type="journal article" year="1981">Manning &amp; Holthuis 1981</bibRefCitation>
: 217, fig. 58a, c). The structures of the inner infraorbital tooth and suborbital ridge are also markedly different. In 
<taxonomicName id="4C5C4D04D3220C10CB159ED7FD62F968" box="[589,675,1699,1724]" class="Malacostraca" family="Camptandriidae" genus="Opusia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B928EA95D3220C10CB159ED7FD62F968" box="[589,675,1699,1724]" italics="true" pageId="2" pageNumber="3">Opusia</emphasis>
</taxonomicName>
<emphasis id="B928EA95D3220C10CBF39ED7FCEFF969" bold="true" box="[683,814,1699,1725]" pageId="2" pageNumber="3">
<taxonomicNameLabel id="A21B57EED3220C10CBF39ED7FCEFF969" box="[683,814,1699,1725]" pageId="2" pageNumber="3" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
, the inner infraorbital tooth is distinctly demarcated from the rest of the infraorbital margin by a triangular notch and the suborbital ridge is not distinctly visible dorsally (
<figureCitation id="13672A02D3220C10C8509E84FE91F8DE" box="[264,336,1776,1802]" captionStart="FIGURE 2" captionStartId="3.[151,256,1918,1942]" captionTargetBox="[185,1383,306,1917]" captionTargetId="figure@3.[180,1404,301,1980]" captionTargetPageId="3" captionText="FIGURE 2. Opusia indicum (Alcock, 1900), male (6. 9 x 10.1 mm) (USNM 191107), Pakistan. A, carapace; B, outer view of left third maxilliped; C, male abdomen; D, frontal view of carapace showing orbit, eye and epistome; E, anterior part of thoracic sternum; F, outer view of right chela; G, left G 1; H, distal part of left G 1. Scales: A, C – G = 1.0 mm; B = 0.5 mm; H = 0.2 mm." httpUri="https://zenodo.org/record/191911/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
D). In 
<taxonomicName id="4C5C4D04D3220C10C8C79E85FDDDF8DE" box="[415,540,1777,1802]" class="Malacostraca" family="Camptandriidae" genus="Tylodiplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B928EA95D3220C10C8C79E85FDDDF8DE" box="[415,540,1777,1802]" italics="true" pageId="2" pageNumber="3">Tylodiplax</emphasis>
</taxonomicName>
, however, the inner infraorbital tooth is weak and its margin is contiguous with the rest of the infraorbital margin with the suborbital ridge prominent, produced and clearly visible dorsally (
<figureCitation id="13672A02D3220C10C85D9F4AFE45F88C" box="[261,388,1854,1880]" captionStart="FIGURE 3. A – D" captionStartId="4.[151,262,1554,1578]" captionTargetBox="[225,1341,213,1553]" captionTargetId="figure@4.[223,1363,194,1599]" captionTargetPageId="4" captionText="FIGURE 3. A – D, Tylodiplax tetratylophora De Man, 1895, male (6.8 x 10.2 mm) (ZRC), Malaysia; E – H, Paratylodiplax blephariskios (Stebbing, 1924), male (5.4 x 8.8 mm) (USNM 137247), South Africa. Scales: A, B, D – F, H = 1.0 mm; C, G = 0.5 mm." httpUri="https://zenodo.org/record/191912/files/figure.png" pageId="2" pageNumber="3">Fig. 3A, B</figureCitation>
). The anterior part of the male thoracic sternum (first two sternites) in 
<taxonomicName id="4C5C4D04D3220C10CDE79F4AFAD5F883" box="[1215,1300,1854,1879]" class="Malacostraca" family="Camptandriidae" genus="Opusia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B928EA95D3220C10CDE79F4AFAD5F883" box="[1215,1300,1854,1879]" italics="true" pageId="2" pageNumber="3">Opusia</emphasis>
</taxonomicName>
<emphasis id="B928EA95D3220C10CC439F49FA5DF883" bold="true" box="[1307,1436,1853,1879]" pageId="2" pageNumber="3">
<taxonomicNameLabel id="A21B57EED3220C10CC439F49FA5DF883" box="[1307,1436,1853,1879]" pageId="2" pageNumber="3" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
is rounded whereas in 
<taxonomicName id="4C5C4D04D3220C10C8FB9F11FDDEF8AA" box="[419,543,1893,1918]" class="Malacostraca" family="Camptandriidae" genus="Tylodiplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B928EA95D3220C10C8FB9F11FDDEF8AA" box="[419,543,1893,1918]" italics="true" pageId="2" pageNumber="3">Tylodiplax</emphasis>
</taxonomicName>
, it forms a distinct triangle (
<figureCitation id="13672A02D3220C10CA2C9F10FC7AF8AA" box="[884,955,1892,1918]" captionStart="FIGURE 3. A – D" captionStartId="4.[151,262,1554,1578]" captionTargetBox="[225,1341,213,1553]" captionTargetId="figure@4.[223,1363,194,1599]" captionTargetPageId="4" captionText="FIGURE 3. A – D, Tylodiplax tetratylophora De Man, 1895, male (6.8 x 10.2 mm) (ZRC), Malaysia; E – H, Paratylodiplax blephariskios (Stebbing, 1924), male (5.4 x 8.8 mm) (USNM 137247), South Africa. Scales: A, B, D – F, H = 1.0 mm; C, G = 0.5 mm." httpUri="https://zenodo.org/record/191912/files/figure.png" pageId="2" pageNumber="3">Fig. 3</figureCitation>
E). In 
<taxonomicName id="4C5C4D04D3220C10CD5F9F11FB9DF8AA" box="[1031,1116,1893,1918]" class="Malacostraca" family="Camptandriidae" genus="Opusia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B928EA95D3220C10CD5F9F11FB9DF8AA" box="[1031,1116,1893,1918]" italics="true" pageId="2" pageNumber="3">Opusia</emphasis>
</taxonomicName>
<emphasis id="B928EA95D3220C10CD3C9F10FB26F8AA" bold="true" box="[1124,1255,1892,1918]" pageId="2" pageNumber="3">
<taxonomicNameLabel id="A21B57EED3220C10CD3C9F10FB26F8AA" box="[1124,1255,1892,1918]" pageId="2" pageNumber="3" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
, all the sutures between the male abdominal somites are complete and visible, although somites 3–5 are functionally fused, with somite 5 constricted at the proximal end (
<figureCitation id="13672A02D3220C10CB909FC6FCD0F818" box="[712,785,1970,1996]" captionStart="FIGURE 2" captionStartId="3.[151,256,1918,1942]" captionTargetBox="[185,1383,306,1917]" captionTargetId="figure@3.[180,1404,301,1980]" captionTargetPageId="3" captionText="FIGURE 2. Opusia indicum (Alcock, 1900), male (6. 9 x 10.1 mm) (USNM 191107), Pakistan. A, carapace; B, outer view of left third maxilliped; C, male abdomen; D, frontal view of carapace showing orbit, eye and epistome; E, anterior part of thoracic sternum; F, outer view of right chela; G, left G 1; H, distal part of left G 1. Scales: A, C – G = 1.0 mm; B = 0.5 mm; H = 0.2 mm." httpUri="https://zenodo.org/record/191911/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
C), while in 
<taxonomicName id="4C5C4D04D3220C10CAFF9FC6FBE5F81F" box="[935,1060,1970,1995]" class="Malacostraca" family="Camptandriidae" genus="Tylodiplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B928EA95D3220C10CAFF9FC6FBE5F81F" box="[935,1060,1970,1995]" italics="true" pageId="2" pageNumber="3">Tylodiplax</emphasis>
</taxonomicName>
somites 2–4 are fused with the sutures shallow and/or incomplete with somite 5 not constricted, with only a slight constriction close to the proximal end of somite 6 (
<figureCitation id="13672A02D3230C11C89598ECFDEBFF66" box="[461,554,152,178]" captionStart="FIGURE 3. A – D" captionStartId="4.[151,262,1554,1578]" captionTargetBox="[225,1341,213,1553]" captionTargetId="figure@4.[223,1363,194,1599]" captionTargetPageId="4" captionText="FIGURE 3. A – D, Tylodiplax tetratylophora De Man, 1895, male (6.8 x 10.2 mm) (ZRC), Malaysia; E – H, Paratylodiplax blephariskios (Stebbing, 1924), male (5.4 x 8.8 mm) (USNM 137247), South Africa. Scales: A, B, D – F, H = 1.0 mm; C, G = 0.5 mm." httpUri="https://zenodo.org/record/191912/files/figure.png" pageId="3" pageNumber="4">Fig. 3C</figureCitation>
). The structure of the G 
<quantity id="4CA49B62D3230C11CA1D98ECFCAFFF66" box="[837,878,152,178]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="3" pageNumber="4" unit="in" value="1.0">1 in</quantity>
<taxonomicName id="4C5C4D04D3230C11CA2D98ECFC0BFF65" box="[885,970,152,177]" class="Malacostraca" family="Camptandriidae" genus="Opusia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B928EA95D3230C11CA2D98ECFC0BFF65" box="[885,970,152,177]" italics="true" pageId="3" pageNumber="4">Opusia</emphasis>
</taxonomicName>
<emphasis id="B928EA95D3230C11CA8998E3FB92FF65" bold="true" box="[977,1107,151,177]" pageId="3" pageNumber="4">
<taxonomicNameLabel id="A21B57EED3230C11CA8998E3FB92FF65" box="[977,1107,151,177]" pageId="3" pageNumber="4" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
ends in a conical apex (
<figureCitation id="13672A02D3230C11CC3498ECFF64FF0C" captionStart="FIGURE 2" captionStartId="3.[151,256,1918,1942]" captionTargetBox="[185,1383,306,1917]" captionTargetId="figure@3.[180,1404,301,1980]" captionTargetPageId="3" captionText="FIGURE 2. Opusia indicum (Alcock, 1900), male (6. 9 x 10.1 mm) (USNM 191107), Pakistan. A, carapace; B, outer view of left third maxilliped; C, male abdomen; D, frontal view of carapace showing orbit, eye and epistome; E, anterior part of thoracic sternum; F, outer view of right chela; G, left G 1; H, distal part of left G 1. Scales: A, C – G = 1.0 mm; B = 0.5 mm; H = 0.2 mm." httpUri="https://zenodo.org/record/191911/files/figure.png" pageId="3" pageNumber="4">Fig. 2</figureCitation>
G, H), while in 
<taxonomicName id="4C5C4D04D3230C11C83A98CBFE21FF0C" box="[354,480,191,216]" class="Malacostraca" family="Camptandriidae" genus="Tylodiplax" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B928EA95D3230C11C83A98CBFE21FF0C" box="[354,480,191,216]" italics="true" pageId="3" pageNumber="4">Tylodiplax</emphasis>
</taxonomicName>
the distal part of the G1 broadens before reaching the apex, forming a bulbous structure (
<figureCitation id="13672A02D3230C11C8559891FEADFF2B" box="[269,364,229,255]" captionStart="FIGURE 3. A – D" captionStartId="4.[151,262,1554,1578]" captionTargetBox="[225,1341,213,1553]" captionTargetId="figure@4.[223,1363,194,1599]" captionTargetPageId="4" captionText="FIGURE 3. A – D, Tylodiplax tetratylophora De Man, 1895, male (6.8 x 10.2 mm) (ZRC), Malaysia; E – H, Paratylodiplax blephariskios (Stebbing, 1924), male (5.4 x 8.8 mm) (USNM 137247), South Africa. Scales: A, B, D – F, H = 1.0 mm; C, G = 0.5 mm." httpUri="https://zenodo.org/record/191912/files/figure.png" pageId="3" pageNumber="4">Fig. 3D</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF23660FD3230C11C9CF9F0AFEBEF821" httpUri="https://zenodo.org/record/191911/files/figure.png" pageId="3" pageNumber="4" targetBox="[185,1383,306,1917]" targetPageId="3">
<paragraph id="8BE33687D3230C11C9CF9F0AFEBEF821" blockId="3.[151,1436,1918,2037]" pageId="3" pageNumber="4">
<emphasis id="B928EA95D3230C11C9CF9F0AFEDAF842" bold="true" box="[151,283,1918,1942]" pageId="3" pageNumber="4">FIGURE 2.</emphasis>
<taxonomicName id="4C5C4D04D3230C11C87B9F0AFDBFF841" authority="Alcock, 1900" authorityName="Alcock" authorityYear="1900" box="[291,638,1918,1941]" class="Malacostraca" family="Camptandriidae" genus="Opusia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="indicum">
<emphasis id="B928EA95D3230C11C87B9F0AFE0EF841" box="[291,463,1918,1941]" italics="true" pageId="3" pageNumber="4">Opusia indicum</emphasis>
(Alcock, 1900)
</taxonomicName>
, male (6. 9 x 10.1 mm) (USNM 191107), Pakistan. A, carapace; B, outer view of left third maxilliped; C, male abdomen; D, frontal view of carapace showing orbit, eye and epistome; E, anterior part of thoracic sternum; F, outer view of right chela; G, left G1; H, distal part of left G1. Scales: A, C–G = 1.0 mm; B = 0.5 mm; H = 0.2 mm.
</paragraph>
</caption>
<caption id="DF23660FD3240C16C9CF9E66FE0FF9BB" httpUri="https://zenodo.org/record/191912/files/figure.png" pageId="4" pageNumber="5" targetBox="[225,1341,213,1553]" targetPageId="4">
<paragraph id="8BE33687D3240C16C9CF9E66FE0FF9BB" blockId="4.[151,1436,1554,1647]" pageId="4" pageNumber="5">
<emphasis id="B928EA95D3240C16C9CF9E66FEE7F9FE" bold="true" box="[151,294,1554,1578]" pageId="4" pageNumber="5">FIGURE 3.</emphasis>
A–D, 
<taxonomicName id="4C5C4D04D3240C16C8219E66FCA7F9FD" authority="De Man, 1895" authorityName="De Man" authorityYear="1895" box="[377,870,1554,1577]" class="Malacostraca" family="Camptandriidae" genus="Tylodiplax" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="tetratylophora">
<emphasis id="B928EA95D3240C16C8219E66FD6FF9FD" box="[377,686,1554,1577]" italics="true" pageId="4" pageNumber="5">Tylodiplax tetratylophora</emphasis>
De Man, 1895
</taxonomicName>
, male (6.8 x 10.2 mm) (ZRC), Malaysia; E–H, 
<taxonomicName id="4C5C4D04D3240C16C9CF9E41FD4AF998" authority="Stebbing, 1924" authorityName="Stebbing" authorityYear="1924" box="[151,651,1589,1612]" class="Malacostraca" family="Camptandriidae" genus="Paratylodiplax" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="blephariskios">
<emphasis id="B928EA95D3240C16C9CF9E41FE0CF998" box="[151,461,1589,1612]" italics="true" pageId="4" pageNumber="5">Paratylodiplax blephariskios</emphasis>
(Stebbing, 1924)
</taxonomicName>
, male (5.4 x 8.8 mm) (USNM 137247), South Africa. Scales: A, B, D–F, H = 1.0 mm; C, G = 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C346650CD3240C17C99E9EB0FEC2FC97" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="reference_group">
<paragraph id="8BE33687D3240C17C99E9EB0FEDCFF2B" blockId="4.[151,1436,1731,2028]" lastBlockId="5.[151,1437,152,1183]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
<taxonomicName id="4C5C4D04D3240C16C99E9EB0FEDAF909" ID-CoL="7PR43" box="[198,283,1732,1757]" class="Malacostraca" family="Camptandriidae" genus="Opusia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B928EA95D3240C16C99E9EB0FEDAF909" box="[198,283,1732,1757]" italics="true" pageId="4" pageNumber="5">Opusia</emphasis>
</taxonomicName>
<emphasis id="B928EA95D3240C16C8729EB7FE75F909" bold="true" box="[298,436,1731,1757]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="A21B57EED3240C16C8729EB7FE75F909" box="[298,436,1731,1757]" pageId="4" pageNumber="5" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
cannot be included in 
<taxonomicName id="4C5C4D04D3240C16CBBC9EB0FB84F90A" ID-CoL="7PSDK" authority="Serene, 1974" authorityName="Serene" authorityYear="1974" box="[740,1093,1732,1758]" class="Malacostraca" family="Camptandriidae" genus="Paratylodiplax" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B928EA95D3240C16CBBC9EB0FC55F909" box="[740,916,1732,1757]" italics="true" pageId="4" pageNumber="5">Paratylodiplax</emphasis>
<bibRefCitation id="EFCD4B76D3240C16CAFC9EB0FB84F90A" author="Serene" box="[932,1093,1732,1758]" pageId="4" pageNumber="5" refString="Serene, R. (1974) Note on the genera and species of the Camptandriinae Stimpson, 1858 (Decapoda, Brachyura: Ocypodidae). Treubia, 28 (3), 59 - 117." type="journal article" year="1974">Serène, 1974</bibRefCitation>
</taxonomicName>
(
<typeStatus id="54E78825D3240C16CD049EB0FB51F90A" box="[1116,1168,1732,1758]" pageId="4" pageNumber="5">type</typeStatus>
species 
<taxonomicName id="4C5C4D04D3240C16CC5B9EB0FE2FF8D0" ID-CoL="VXMP" authority="Stebbing, 1924" authorityName="Stebbing" authorityYear="1924" class="Malacostraca" family="Camptandriidae" genus="Cleistostoma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="blephariskios">
<emphasis id="B928EA95D3240C16CC5B9EB0FEF4F8D0" italics="true" pageId="4" pageNumber="5">Cleistostoma blephariskios</emphasis>
<bibRefCitation id="EFCD4B76D3240C16C8649E9EFE2FF8D0" author="Stebbing" box="[316,494,1770,1796]" pageId="4" pageNumber="5" refString="Stebbing, T. R. R. (1924) South African Crustacea (Part XII of South African Crustacea, for Marine Investigations in South Africa). Annals of the South African Museum, 19, 235 - 248, pls. 1 - 7 (pls. 116 - 122 of Crustacea)." type="journal article" year="1924">Stebbing, 1924</bibRefCitation>
</taxonomicName>
) as there are also key differences in terms of the structure of the orbital structure (supraorbital margin sinuous in 
<taxonomicName id="4C5C4D04D3240C16CB4E9F65FDADF8FE" ID-CoL="7PR43" box="[534,620,1809,1834]" class="Malacostraca" family="Camptandriidae" genus="Opusia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B928EA95D3240C16CB4E9F65FDADF8FE" box="[534,620,1809,1834]" italics="true" pageId="4" pageNumber="5">Opusia</emphasis>
</taxonomicName>
<emphasis id="B928EA95D3240C16CB2D9F65FD38F8FF" bold="true" box="[629,761,1809,1835]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="A21B57EED3240C16CB2D9F65FD38F8FF" box="[629,761,1809,1835]" pageId="4" pageNumber="5" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
, 
<figureCitation id="13672A02D3240C16CA509F65FC8EF8FF" box="[776,847,1809,1835]" captionStart="FIGURE 2" captionStartId="3.[151,256,1918,1942]" captionTargetBox="[185,1383,306,1917]" captionTargetId="figure@3.[180,1404,301,1980]" captionTargetPageId="3" captionText="FIGURE 2. Opusia indicum (Alcock, 1900), male (6. 9 x 10.1 mm) (USNM 191107), Pakistan. A, carapace; B, outer view of left third maxilliped; C, male abdomen; D, frontal view of carapace showing orbit, eye and epistome; E, anterior part of thoracic sternum; F, outer view of right chela; G, left G 1; H, distal part of left G 1. Scales: A, C – G = 1.0 mm; B = 0.5 mm; H = 0.2 mm." httpUri="https://zenodo.org/record/191911/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
A, against almost straight in 
<taxonomicName id="4C5C4D04D3240C16CDF29F65FA9DF8FE" ID-CoL="7PSDK" box="[1194,1372,1809,1834]" class="Malacostraca" family="Camptandriidae" genus="Paratylodiplax" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B928EA95D3240C16CDF29F65FA9DF8FE" box="[1194,1372,1809,1834]" italics="true" pageId="4" pageNumber="5">Paratylodiplax</emphasis>
</taxonomicName>
, 
<figureCitation id="13672A02D3240C16CC359F65FF64F886" captionStart="FIGURE 3. A – D" captionStartId="4.[151,262,1554,1578]" captionTargetBox="[225,1341,213,1553]" captionTargetId="figure@4.[223,1363,194,1599]" captionTargetPageId="4" captionText="FIGURE 3. A – D, Tylodiplax tetratylophora De Man, 1895, male (6.8 x 10.2 mm) (ZRC), Malaysia; E – H, Paratylodiplax blephariskios (Stebbing, 1924), male (5.4 x 8.8 mm) (USNM 137247), South Africa. Scales: A, B, D – F, H = 1.0 mm; C, G = 0.5 mm." httpUri="https://zenodo.org/record/191912/files/figure.png" pageId="4" pageNumber="5">Fig. 3</figureCitation>
E), epistome (posteromedial tooth triangular with lateral margins gently sinuous in 
<taxonomicName id="4C5C4D04D3240C16CD279F4CFB15F885" ID-CoL="7PR43" box="[1151,1236,1848,1873]" class="Malacostraca" family="Camptandriidae" genus="Opusia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B928EA95D3240C16CD279F4CFB15F885" box="[1151,1236,1848,1873]" italics="true" pageId="4" pageNumber="5">Opusia</emphasis>
</taxonomicName>
<emphasis id="B928EA95D3240C16CD849F4CFA9FF886" bold="true" box="[1244,1374,1848,1874]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="A21B57EED3240C16CD849F4CFA9FF886" box="[1244,1374,1848,1874]" pageId="4" pageNumber="5" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
, 
<figureCitation id="13672A02D3240C16CC359F4CFF66F8AC" captionStart="FIGURE 2" captionStartId="3.[151,256,1918,1942]" captionTargetBox="[185,1383,306,1917]" captionTargetId="figure@3.[180,1404,301,1980]" captionTargetPageId="3" captionText="FIGURE 2. Opusia indicum (Alcock, 1900), male (6. 9 x 10.1 mm) (USNM 191107), Pakistan. A, carapace; B, outer view of left third maxilliped; C, male abdomen; D, frontal view of carapace showing orbit, eye and epistome; E, anterior part of thoracic sternum; F, outer view of right chela; G, left G 1; H, distal part of left G 1. Scales: A, C – G = 1.0 mm; B = 0.5 mm; H = 0.2 mm." httpUri="https://zenodo.org/record/191911/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
D, against posteromedial tooth acutely triangular with lateral margins strongly sinuous in 
<taxonomicName id="4C5C4D04D3240C16CDB99F2BFA55F8AC" ID-CoL="7PSDK" box="[1249,1428,1887,1912]" class="Malacostraca" family="Camptandriidae" genus="Paratylodiplax" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B928EA95D3240C16CDB99F2BFA55F8AC" box="[1249,1428,1887,1912]" italics="true" pageId="4" pageNumber="5">Paratylodiplax</emphasis>
</taxonomicName>
, 
<figureCitation id="13672A02D3240C16C9CF9FF1FF1DF84B" box="[151,220,1925,1951]" captionStart="FIGURE 3. A – D" captionStartId="4.[151,262,1554,1578]" captionTargetBox="[225,1341,213,1553]" captionTargetId="figure@4.[223,1363,194,1599]" captionTargetPageId="4" captionText="FIGURE 3. A – D, Tylodiplax tetratylophora De Man, 1895, male (6.8 x 10.2 mm) (ZRC), Malaysia; E – H, Paratylodiplax blephariskios (Stebbing, 1924), male (5.4 x 8.8 mm) (USNM 137247), South Africa. Scales: A, B, D – F, H = 1.0 mm; C, G = 0.5 mm." httpUri="https://zenodo.org/record/191912/files/figure.png" pageId="4" pageNumber="5">Fig. 3</figureCitation>
F), male abdominal segmentation (sutures between all somites visible in 
<taxonomicName id="4C5C4D04D3240C16CD119FF1FB5EF84A" ID-CoL="7PR43" box="[1097,1183,1925,1950]" class="Malacostraca" family="Camptandriidae" genus="Opusia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B928EA95D3240C16CD119FF1FB5EF84A" box="[1097,1183,1925,1950]" italics="true" pageId="4" pageNumber="5">Opusia</emphasis>
</taxonomicName>
<emphasis id="B928EA95D3240C16CDFF9FF1FAEAF84B" bold="true" box="[1191,1323,1925,1951]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="A21B57EED3240C16CDFF9FF1FAEAF84B" box="[1191,1323,1925,1951]" pageId="4" pageNumber="5" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
, 
<figureCitation id="13672A02D3240C16CC639FF1FA43F84B" box="[1339,1410,1925,1951]" captionStart="FIGURE 2" captionStartId="3.[151,256,1918,1942]" captionTargetBox="[185,1383,306,1917]" captionTargetId="figure@3.[180,1404,301,1980]" captionTargetPageId="3" captionText="FIGURE 2. Opusia indicum (Alcock, 1900), male (6. 9 x 10.1 mm) (USNM 191107), Pakistan. A, carapace; B, outer view of left third maxilliped; C, male abdomen; D, frontal view of carapace showing orbit, eye and epistome; E, anterior part of thoracic sternum; F, outer view of right chela; G, left G 1; H, distal part of left G 1. Scales: A, C – G = 1.0 mm; B = 0.5 mm; H = 0.2 mm." httpUri="https://zenodo.org/record/191911/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
C, against somites 3–5 completely fused without trace of sutures in 
<taxonomicName id="4C5C4D04D3240C16CAF79FD8FBA5F811" ID-CoL="7PSDK" box="[943,1124,1964,1989]" class="Malacostraca" family="Camptandriidae" genus="Paratylodiplax" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B928EA95D3240C16CAF79FD8FBA5F811" box="[943,1124,1964,1989]" italics="true" pageId="4" pageNumber="5">Paratylodiplax</emphasis>
</taxonomicName>
, 
<figureCitation id="13672A02D3240C16CD2C9FD8FB7AF812" box="[1140,1211,1964,1990]" captionStart="FIGURE 3. A – D" captionStartId="4.[151,262,1554,1578]" captionTargetBox="[225,1341,213,1553]" captionTargetId="figure@4.[223,1363,194,1599]" captionTargetPageId="4" captionText="FIGURE 3. A – D, Tylodiplax tetratylophora De Man, 1895, male (6.8 x 10.2 mm) (ZRC), Malaysia; E – H, Paratylodiplax blephariskios (Stebbing, 1924), male (5.4 x 8.8 mm) (USNM 137247), South Africa. Scales: A, B, D – F, H = 1.0 mm; C, G = 0.5 mm." httpUri="https://zenodo.org/record/191912/files/figure.png" pageId="4" pageNumber="5">Fig. 3</figureCitation>
G), and G1 (bends gradually with several large subterminal spines covered with lateral spinules in 
<taxonomicName id="4C5C4D04D3240C16CD159FA7FB63F838" ID-CoL="7PR43" box="[1101,1186,2003,2028]" class="Malacostraca" family="Camptandriidae" genus="Opusia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B928EA95D3240C16CD159FA7FB63F838" box="[1101,1186,2003,2028]" italics="true" pageId="4" pageNumber="5">Opusia</emphasis>
</taxonomicName>
<emphasis id="B928EA95D3240C16CDF29FA6FAECF838" bold="true" box="[1194,1325,2002,2028]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="A21B57EED3240C16CDF29FA6FAECF838" box="[1194,1325,2002,2028]" pageId="4" pageNumber="5" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
, 
<figureCitation id="13672A02D3240C16CC659FA6FA43F838" box="[1341,1410,2002,2028]" captionStart="FIGURE 2" captionStartId="3.[151,256,1918,1942]" captionTargetBox="[185,1383,306,1917]" captionTargetId="figure@3.[180,1404,301,1980]" captionTargetPageId="3" captionText="FIGURE 2. Opusia indicum (Alcock, 1900), male (6. 9 x 10.1 mm) (USNM 191107), Pakistan. A, carapace; B, outer view of left third maxilliped; C, male abdomen; D, frontal view of carapace showing orbit, eye and epistome; E, anterior part of thoracic sternum; F, outer view of right chela; G, left G 1; H, distal part of left G 1. Scales: A, C – G = 1.0 mm; B = 0.5 mm; H = 0.2 mm." httpUri="https://zenodo.org/record/191911/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
G, H, against bending sharply with subdistal part only with one large subterminal spine in 
<taxonomicName id="4C5C4D04D3250C17CDF498ECFA9FFF65" ID-CoL="7PSDK" box="[1196,1374,152,177]" class="Malacostraca" family="Camptandriidae" genus="Paratylodiplax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B928EA95D3250C17CDF498ECFA9FFF65" box="[1196,1374,152,177]" italics="true" pageId="5" pageNumber="6">Paratylodiplax</emphasis>
</taxonomicName>
, 
<figureCitation id="13672A02D3250C17CC3598ECFF64FF0C" captionStart="FIGURE 3. A – D" captionStartId="4.[151,262,1554,1578]" captionTargetBox="[225,1341,213,1553]" captionTargetId="figure@4.[223,1363,194,1599]" captionTargetPageId="4" captionText="FIGURE 3. A – D, Tylodiplax tetratylophora De Man, 1895, male (6.8 x 10.2 mm) (ZRC), Malaysia; E – H, Paratylodiplax blephariskios (Stebbing, 1924), male (5.4 x 8.8 mm) (USNM 137247), South Africa. Scales: A, B, D – F, H = 1.0 mm; C, G = 0.5 mm." httpUri="https://zenodo.org/record/191912/files/figure.png" pageId="5" pageNumber="6">Fig. 3</figureCitation>
H), as well as absence of sexual dimorphism in the chelipeds (
<figureCitation id="13672A02D3250C17CAD798CAFC15FF0C" box="[911,980,190,216]" captionStart="FIGURE 3. A – D" captionStartId="4.[151,262,1554,1578]" captionTargetBox="[225,1341,213,1553]" captionTargetId="figure@4.[223,1363,194,1599]" captionTargetPageId="4" captionText="FIGURE 3. A – D, Tylodiplax tetratylophora De Man, 1895, male (6.8 x 10.2 mm) (ZRC), Malaysia; E – H, Paratylodiplax blephariskios (Stebbing, 1924), male (5.4 x 8.8 mm) (USNM 137247), South Africa. Scales: A, B, D – F, H = 1.0 mm; C, G = 0.5 mm." httpUri="https://zenodo.org/record/191912/files/figure.png" pageId="5" pageNumber="6">Fig. 3</figureCitation>
E–H; 
<bibRefCitation id="EFCD4B76D3250C17CD4498CAFA9DFF0C" author="Manning" box="[1052,1372,190,216]" pageId="5" pageNumber="6" refString="Manning, R. B. &amp; Holthuis, L. B. (1981) West African Brachyuran Crabs (Crustacea: Decapoda). Smithsonian Contributions to Zoology, 306, 1 - 379." type="journal article" year="1981">Manning &amp; Holthuis 1981</bibRefCitation>
: 209, fig. 54a–e).
</paragraph>
<paragraph id="8BE33687D3250C17C99E9978FEC2FC97" blockId="5.[151,1437,152,1183]" pageId="5" pageNumber="6">
<bibRefCitation id="EFCD4B76D3250C17C99E9978FDDDFEF2" author="Manning" box="[198,540,268,294]" pageId="5" pageNumber="6" refString="Manning, R. B. &amp; Holthuis, L. B. (1981) West African Brachyuran Crabs (Crustacea: Decapoda). Smithsonian Contributions to Zoology, 306, 1 - 379." type="journal article" year="1981">Manning &amp; Holthuis (1981)</bibRefCitation>
were unable to resolve the generic position of 
<taxonomicName id="4C5C4D04D3250C17CD3A9978FAF3FEF1" ID-CoL="7CYJX" box="[1122,1330,268,293]" class="Malacostraca" family="Camptandriidae" genus="Tylodiplax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="B928EA95D3250C17CD3A9978FAF3FEF1" box="[1122,1330,268,293]" italics="true" pageId="5" pageNumber="6">Tylodiplax indica</emphasis>
</taxonomicName>
, so they provisionally placed the species in 
<taxonomicName id="4C5C4D04D3250C17CB6F9946FD64FE9F" ID-CoL="7PZGS" box="[567,677,306,331]" class="Malacostraca" family="Camptandriidae" genus="Serenella" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B928EA95D3250C17CB6F9946FD64FE9F" box="[567,677,306,331]" italics="true" pageId="5" pageNumber="6">Serenella</emphasis>
</taxonomicName>
. Examination of the present material, however, indicates that 
<taxonomicName id="4C5C4D04D3250C17CCDE9946FF27FEA6" ID-CoL="7CYJX" class="Malacostraca" family="Camptandriidae" genus="Tylodiplax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="B928EA95D3250C17CCDE9946FF27FEA6" italics="true" pageId="5" pageNumber="6">T. indica</emphasis>
</taxonomicName>
does not belong to 
<taxonomicName id="4C5C4D04D3250C17C8BB992DFD98FEA6" ID-CoL="7PZGS" box="[483,601,345,370]" class="Malacostraca" family="Camptandriidae" genus="Serenella" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B928EA95D3250C17C8BB992DFD98FEA6" box="[483,601,345,370]" italics="true" pageId="5" pageNumber="6">Serenella</emphasis>
</taxonomicName>
because of important differences in several characters. The inner infraorbital tooth of the frontal region of 
<taxonomicName id="4C5C4D04D3250C17CBCB99F4FD3FFE4D" ID-CoL="7CYJX" box="[659,766,384,409]" class="Malacostraca" family="Camptandriidae" genus="Tylodiplax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="B928EA95D3250C17CBCB99F4FD3FFE4D" box="[659,766,384,409]" italics="true" pageId="5" pageNumber="6">T. indica</emphasis>
</taxonomicName>
, is clearly demarcated from the rest of the infraorbital margin, and the frontomedial margin is triangular, ending in a truncate apex (
<figureCitation id="13672A02D3250C17CD7899D2FBA7FE14" box="[1056,1126,422,448]" captionStart="FIGURE 2" captionStartId="3.[151,256,1918,1942]" captionTargetBox="[185,1383,306,1917]" captionTargetId="figure@3.[180,1404,301,1980]" captionTargetPageId="3" captionText="FIGURE 2. Opusia indicum (Alcock, 1900), male (6. 9 x 10.1 mm) (USNM 191107), Pakistan. A, carapace; B, outer view of left third maxilliped; C, male abdomen; D, frontal view of carapace showing orbit, eye and epistome; E, anterior part of thoracic sternum; F, outer view of right chela; G, left G 1; H, distal part of left G 1. Scales: A, C – G = 1.0 mm; B = 0.5 mm; H = 0.2 mm." httpUri="https://zenodo.org/record/191911/files/figure.png" pageId="5" pageNumber="6">Fig. 2</figureCitation>
A, D); but in 
<taxonomicName id="4C5C4D04D3250C17CC5B99D3FAB0FE14" ID-CoL="7PZGS" box="[1283,1393,423,448]" class="Malacostraca" family="Camptandriidae" genus="Serenella" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B928EA95D3250C17CC5B99D3FAB0FE14" box="[1283,1393,423,448]" italics="true" pageId="5" pageNumber="6">Serenella</emphasis>
</taxonomicName>
the inner infraorbital tooth is not sharply demarcated, and the frontomedial margin is bilobed (
<bibRefCitation id="EFCD4B76D3250C17CC5399B9FE72FDDA" author="Manning" pageId="5" pageNumber="6" refString="Manning, R. B. &amp; Holthuis, L. B. (1981) West African Brachyuran Crabs (Crustacea: Decapoda). Smithsonian Contributions to Zoology, 306, 1 - 379." type="journal article" year="1981" yearSuffix="a">Manning &amp; Holthuis 1981: fig. 55a</bibRefCitation>
). The front of 
<taxonomicName id="4C5C4D04D3250C17CB3A9980FD09FDD9" ID-CoL="7CYJX" box="[610,712,500,525]" class="Malacostraca" family="Camptandriidae" genus="Tylodiplax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="B928EA95D3250C17CB3A9980FD09FDD9" box="[610,712,500,525]" italics="true" pageId="5" pageNumber="6">T. indica</emphasis>
</taxonomicName>
is straight and entire from dorsal view and the anterolateral angles are rounded (
<figureCitation id="13672A02D3250C17C8D29A6EFE12FDE0" box="[394,467,538,564]" captionStart="FIGURE 2" captionStartId="3.[151,256,1918,1942]" captionTargetBox="[185,1383,306,1917]" captionTargetId="figure@3.[180,1404,301,1980]" captionTargetPageId="3" captionText="FIGURE 2. Opusia indicum (Alcock, 1900), male (6. 9 x 10.1 mm) (USNM 191107), Pakistan. A, carapace; B, outer view of left third maxilliped; C, male abdomen; D, frontal view of carapace showing orbit, eye and epistome; E, anterior part of thoracic sternum; F, outer view of right chela; G, left G 1; H, distal part of left G 1. Scales: A, C – G = 1.0 mm; B = 0.5 mm; H = 0.2 mm." httpUri="https://zenodo.org/record/191911/files/figure.png" pageId="5" pageNumber="6">Fig. 2</figureCitation>
A), whereas for 
<taxonomicName id="4C5C4D04D3250C17CBCD9A6EFCC5FDE7" ID-CoL="7PZGS" box="[661,772,538,563]" class="Malacostraca" family="Camptandriidae" genus="Serenella" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B928EA95D3250C17CBCD9A6EFCC5FDE7" box="[661,772,538,563]" italics="true" pageId="5" pageNumber="6">Serenella</emphasis>
</taxonomicName>
the front is distinctly bilobed from dorsal view and the anterolateral angles are pointed (
<bibRefCitation id="EFCD4B76D3250C17CB4C9A35FC79FD8F" author="Manning" box="[532,952,577,603]" pageId="5" pageNumber="6" refString="Manning, R. B. &amp; Holthuis, L. B. (1981) West African Brachyuran Crabs (Crustacea: Decapoda). Smithsonian Contributions to Zoology, 306, 1 - 379." type="journal article" year="1981" yearSuffix="a">Manning &amp; Holthuis 1981: fig. 55a</bibRefCitation>
). There is no obvious sexual dimorphism of the chelae in 
<taxonomicName id="4C5C4D04D3250C17C8029A1CFE02FD55" ID-CoL="7CYJX" box="[346,451,616,641]" class="Malacostraca" family="Camptandriidae" genus="Tylodiplax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="B928EA95D3250C17C8029A1CFE02FD55" box="[346,451,616,641]" italics="true" pageId="5" pageNumber="6">T. indica</emphasis>
</taxonomicName>
(
<figureCitation id="13672A02D3250C17C88D9A1CFDDFFD56" box="[469,542,616,642]" captionStart="FIGURE 2" captionStartId="3.[151,256,1918,1942]" captionTargetBox="[185,1383,306,1917]" captionTargetId="figure@3.[180,1404,301,1980]" captionTargetPageId="3" captionText="FIGURE 2. Opusia indicum (Alcock, 1900), male (6. 9 x 10.1 mm) (USNM 191107), Pakistan. A, carapace; B, outer view of left third maxilliped; C, male abdomen; D, frontal view of carapace showing orbit, eye and epistome; E, anterior part of thoracic sternum; F, outer view of right chela; G, left G 1; H, distal part of left G 1. Scales: A, C – G = 1.0 mm; B = 0.5 mm; H = 0.2 mm." httpUri="https://zenodo.org/record/191911/files/figure.png" pageId="5" pageNumber="6">Fig. 2</figureCitation>
F), but this is clearly present in 
<taxonomicName id="4C5C4D04D3250C17CAF89A1CFBD0FD55" ID-CoL="7PZGS" box="[928,1041,616,641]" class="Malacostraca" family="Camptandriidae" genus="Serenella" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B928EA95D3250C17CAF89A1CFBD0FD55" box="[928,1041,616,641]" italics="true" pageId="5" pageNumber="6">Serenella</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFCD4B76D3250C17CD799A1CFF04FD7C" author="Manning" pageId="5" pageNumber="6" refString="Manning, R. B. &amp; Holthuis, L. B. (1981) West African Brachyuran Crabs (Crustacea: Decapoda). Smithsonian Contributions to Zoology, 306, 1 - 379." type="journal article" year="1981" yearSuffix="a">Manning &amp; Holthuis 1981: fig. 55a</bibRefCitation>
). The supraorbital margins of 
<taxonomicName id="4C5C4D04D3250C17CB749AFBFD55FD7C" ID-CoL="7CYJX" box="[556,660,655,680]" class="Malacostraca" family="Camptandriidae" genus="Tylodiplax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="B928EA95D3250C17CB749AFBFD55FD7C" box="[556,660,655,680]" italics="true" pageId="5" pageNumber="6">T. indica</emphasis>
</taxonomicName>
are slightly sinuous but generally parallel to the posterior margin of the carapace (
<figureCitation id="13672A02D3250C17C8069AC1FE67FD1B" box="[350,422,693,719]" captionStart="FIGURE 2" captionStartId="3.[151,256,1918,1942]" captionTargetBox="[185,1383,306,1917]" captionTargetId="figure@3.[180,1404,301,1980]" captionTargetPageId="3" captionText="FIGURE 2. Opusia indicum (Alcock, 1900), male (6. 9 x 10.1 mm) (USNM 191107), Pakistan. A, carapace; B, outer view of left third maxilliped; C, male abdomen; D, frontal view of carapace showing orbit, eye and epistome; E, anterior part of thoracic sternum; F, outer view of right chela; G, left G 1; H, distal part of left G 1. Scales: A, C – G = 1.0 mm; B = 0.5 mm; H = 0.2 mm." httpUri="https://zenodo.org/record/191911/files/figure.png" pageId="5" pageNumber="6">Fig. 2</figureCitation>
A, D), but in 
<taxonomicName id="4C5C4D04D3250C17CB119AC1FD78FD1A" ID-CoL="7PZGS" box="[585,697,693,718]" class="Malacostraca" family="Camptandriidae" genus="Serenella" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B928EA95D3250C17CB119AC1FD78FD1A" box="[585,697,693,718]" italics="true" pageId="5" pageNumber="6">Serenella</emphasis>
</taxonomicName>
, the supraorbital margins are oblique relative to the posterior margin (
<bibRefCitation id="EFCD4B76D3250C17C9A09AA8FD5AFD22" author="Manning" box="[248,667,732,758]" pageId="5" pageNumber="6" refString="Manning, R. B. &amp; Holthuis, L. B. (1981) West African Brachyuran Crabs (Crustacea: Decapoda). Smithsonian Contributions to Zoology, 306, 1 - 379." type="journal article" year="1981" yearSuffix="a">Manning &amp; Holthuis 1981: fig. 55a</bibRefCitation>
). The junction between the recurved and straighter portions of the G 
<quantity id="4CA49B62D3250C17C9E89B76FF1BFCC8" box="[176,218,770,796]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="5" pageNumber="6" unit="in" value="1.0">1 in</quantity>
<taxonomicName id="4C5C4D04D3250C17C9BA9B76FE89FCCF" ID-CoL="7CYJX" box="[226,328,770,795]" class="Malacostraca" family="Camptandriidae" genus="Tylodiplax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="B928EA95D3250C17C9BA9B76FE89FCCF" box="[226,328,770,795]" italics="true" pageId="5" pageNumber="6">T. indica</emphasis>
</taxonomicName>
, is rounded and gradual (
<figureCitation id="13672A02D3250C17CB2F9B76FD7EFCC8" box="[631,703,770,796]" captionStart="FIGURE 2" captionStartId="3.[151,256,1918,1942]" captionTargetBox="[185,1383,306,1917]" captionTargetId="figure@3.[180,1404,301,1980]" captionTargetPageId="3" captionText="FIGURE 2. Opusia indicum (Alcock, 1900), male (6. 9 x 10.1 mm) (USNM 191107), Pakistan. A, carapace; B, outer view of left third maxilliped; C, male abdomen; D, frontal view of carapace showing orbit, eye and epistome; E, anterior part of thoracic sternum; F, outer view of right chela; G, left G 1; H, distal part of left G 1. Scales: A, C – G = 1.0 mm; B = 0.5 mm; H = 0.2 mm." httpUri="https://zenodo.org/record/191911/files/figure.png" pageId="5" pageNumber="6">Fig. 2</figureCitation>
G), while in 
<taxonomicName id="4C5C4D04D3250C17CA0C9B76FC03FCCF" ID-CoL="7PZGS" box="[852,962,770,795]" class="Malacostraca" family="Camptandriidae" genus="Serenella" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B928EA95D3250C17CA0C9B76FC03FCCF" box="[852,962,770,795]" italics="true" pageId="5" pageNumber="6">Serenella</emphasis>
</taxonomicName>
it is abrupt (
<bibRefCitation id="EFCD4B76D3250C17CD019B76FF37FC97" author="Manning" pageId="5" pageNumber="6" refString="Manning, R. B. &amp; Holthuis, L. B. (1981) West African Brachyuran Crabs (Crustacea: Decapoda). Smithsonian Contributions to Zoology, 306, 1 - 379." type="journal article" year="1981" yearSuffix="b">Manning &amp; Holthuis 1981: fig. 55b</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C346650CD3250C17C99E9B3BFE9FFB4B" pageId="5" pageNumber="6" type="materials_examined">
<paragraph id="8BE33687D3250C17C99E9B3BFE9FFB4B" blockId="5.[151,1437,152,1183]" pageId="5" pageNumber="6">
<emphasis id="B928EA95D3250C17C99E9B3BFE23FCBD" bold="true" box="[198,482,847,873]" pageId="5" pageNumber="6">Comparative material.</emphasis>
<taxonomicName id="4C5C4D04D3250C17C8B19B24FC02FCBE" authority="De Man, 1895" authorityName="De Man" authorityYear="1895" box="[489,963,848,874]" class="Malacostraca" family="Camptandriidae" genus="Tylodiplax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="tetratylophora">
<emphasis id="B928EA95D3250C17C8B19B24FCD5FCBD" box="[489,788,848,873]" italics="true" pageId="5" pageNumber="6">Tylodiplax tetratylophora</emphasis>
De 
<bibRefCitation id="EFCD4B76D3250C17CA1C9B24FC02FCBE" author="Man" box="[836,963,848,874]" pageId="5" pageNumber="6" refString="Man, J. G. De (1895) Bericht uber die von Herrn Schiffscapitan Storm zur Atjeh, an den westlichen Kusten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8, 485 - 609, pls. 12 - 14." type="journal article" year="1895">Man, 1895</bibRefCitation>
</taxonomicName>
– 
<emphasis id="B928EA95D3250C17CA879B3BFBBCFCBD" bold="true" box="[991,1149,847,873]" pageId="5" pageNumber="6">
<collectingCountry id="F34B7617D3250C17CA879B3BFBBCFCBD" box="[991,1149,847,873]" name="Thailand" pageId="5" pageNumber="6">THAILAND</collectingCountry>
</emphasis>
: 
<specimenCount id="9D5AFD0ED3250C17CDD49B24FB1BFCBE" box="[1164,1242,848,874]" pageId="5" pageNumber="6" type="male">1 male</specimenCount>
(6.9 x 11.0 mm) (
<collectionCode id="ED4DAE42D3250C17C9C79B02FF33FC44" box="[159,242,886,912]" httpUri="http://grbio.org/cool/c3cw-qjai" name="Phuket Marine Biological Centre" pageId="5" pageNumber="6">PMBC</collectionCode>
2227), Phang Nga mangrove forest, leg. S. Lundoer, 
<date id="FFE21047D3250C17CA3A9B02FBD6FC44" box="[866,1047,886,912]" pageId="5" pageNumber="6" value="1972-01-03">3 January 1972</date>
; 
<specimenCount id="9D5AFD0ED3250C17CD7B9B02FBBDFC44" box="[1059,1148,886,912]" pageId="5" pageNumber="6" type="male">4 males</specimenCount>
(
<quantity id="4CA49B62D3250C17CDD49B02FADBFC44" box="[1164,1306,886,912]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" metricValueMax="7.7" metricValueMin="5.9" pageId="5" pageNumber="6" unit="mm" value="6.8" valueMax="7.7" valueMin="5.9">5.9–7.7 mm</quantity>
x 
<quantity id="4CA49B62D3250C17CC6E9B02FF06FC63" metricMagnitude="-2" metricUnit="m" metricValue="1.005" metricValueMax="1.17" metricValueMin="0.84" pageId="5" pageNumber="6" unit="mm" value="10.05" valueMax="11.7" valueMin="8.4">8.4–11.7 mm</quantity>
) (
<collectionCode id="ED4DAE42D3250C17C9869BE9FEF0FC63" box="[222,305,925,951]" httpUri="http://grbio.org/cool/c3cw-qjai" name="Phuket Marine Biological Centre" pageId="5" pageNumber="6">PMBC</collectionCode>
4384), mangrove shore, Ao Nam Bor, Phuket; 
<specimenCount id="9D5AFD0ED3250C17CA069BE9FC78FC63" box="[862,953,925,951]" pageId="5" pageNumber="6" type="male">7 males</specimenCount>
(5.0– 
<quantity id="4CA49B62D3250C17CAA59BE9FB99FC63" box="[1021,1112,925,951]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="5" pageNumber="6" unit="mm" value="7.2">7.2 mm</quantity>
x 
<quantity id="4CA49B62D3250C17CD2E9BE9FAD7FC63" box="[1142,1302,925,951]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" metricValueMax="10.2" metricValueMin="8.2" pageId="5" pageNumber="6" unit="mm" value="9.2" valueMax="10.2" valueMin="8.2">8.2–10.2 mm</quantity>
), 
<specimenCount id="9D5AFD0ED3250C17CC739BE9FA5DFC63" box="[1323,1436,925,951]" pageId="5" pageNumber="6" type="female">3 females</specimenCount>
(
<quantity id="4CA49B62D3250C17C9C79BB0FEF0FC0A" box="[159,305,964,990]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" metricValueMax="7.2" metricValueMin="6.4" pageId="5" pageNumber="6" unit="mm" value="6.8" valueMax="7.2" valueMin="6.4">6.4–7.2 mm</quantity>
x 11.0– 
<quantity id="4CA49B62D3250C17C8CC9BB0FE3EFC0A" box="[404,511,964,990]" metricMagnitude="-2" metricUnit="m" metricValue="1.17" pageId="5" pageNumber="6" unit="mm" value="11.7">11.7 mm</quantity>
) (
<collectionCode id="ED4DAE42D3250C17CB409BB0FDACFC0A" box="[536,621,964,990]" httpUri="http://grbio.org/cool/c3cw-qjai" name="Phuket Marine Biological Centre" pageId="5" pageNumber="6">PMBC</collectionCode>
4390), mangrove shore, Ao Nam Bor, Phuket; 
<specimenCount id="9D5AFD0ED3250C17CDF39BB0FADEFC0A" box="[1195,1311,964,990]" pageId="5" pageNumber="6" type="female">2 females</specimenCount>
(5.1 x 
<quantity id="4CA49B62D3250C17CC2F9BB0FF08FBD0" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="5" pageNumber="6" unit="mm" value="7.5">7.5 mm</quantity>
, 6.6 x 
<quantity id="4CA49B62D3250C17C8439B9EFE4DFBD0" box="[283,396,1002,1028]" metricMagnitude="-2" metricUnit="m" metricValue="1.03" pageId="5" pageNumber="6" unit="mm" value="10.3">10.3 mm</quantity>
), 
<specimenCount id="9D5AFD0ED3250C17C8F99B9EFDDAFBD0" box="[417,539,1002,1028]" pageId="5" pageNumber="6" type="juvenile">1 juvenile</specimenCount>
(3.7 x 
<quantity id="4CA49B62D3250C17CB2B9B9EFD10FBD0" box="[627,721,1002,1028]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="5" pageNumber="6" unit="mm" value="5.8">5.8 mm</quantity>
) (
<collectionCode id="ED4DAE42D3250C17CBB29B9EFCFEFBD0" box="[746,831,1002,1028]" httpUri="http://grbio.org/cool/c3cw-qjai" name="Phuket Marine Biological Centre" pageId="5" pageNumber="6">PMBC</collectionCode>
4394), mudflat, Ko Yao Yai, Phang Nga, Phuket; 
<emphasis id="B928EA95D3250C17C9CF9C64FDC5FBFE" bold="true" box="[151,516,1040,1066]" pageId="5" pageNumber="6">
PENINSULAR 
<collectingCountry id="F34B7617D3250C17C83B9C64FDC5FBFE" box="[355,516,1040,1066]" name="Malaysia" pageId="5" pageNumber="6">MALAYSIA</collectingCountry>
</emphasis>
: 
<specimenCount id="9D5AFD0ED3250C17CB4D9C65FD47FBFF" box="[533,646,1041,1067]" pageId="5" pageNumber="6" type="male">10 males</specimenCount>
, 
<specimenCount id="9D5AFD0ED3250C17CBCB9C65FCD8FBFF" box="[659,793,1041,1067]" pageId="5" pageNumber="6" type="female">20 females</specimenCount>
(
<collectionCode id="ED4DAE42D3250C17CA729C65FCAAFBFF" box="[810,875,1041,1067]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="5" pageNumber="6">ZRC</collectionCode>
), mudflat, Matang mangroves, Perak, coll. A. Ramli, F. Mohamad, C. G. S. Tan, 
<date id="FFE21047D3250C17CB6F9C4CFD1AFB86" box="[567,731,1080,1106]" pageId="5" pageNumber="6" value="1995-06-10">10 June 1995</date>
. 
<taxonomicName id="4C5C4D04D3250C17CBB09C4CFAD5FB86" authority="Stebbing, 1924" authorityName="Stebbing" authorityYear="1924" box="[744,1300,1080,1106]" class="Malacostraca" family="Camptandriidae" genus="Paratylodiplax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="blephariskios">
<emphasis id="B928EA95D3250C17CBB09C4CFB85FB85" box="[744,1092,1080,1105]" italics="true" pageId="5" pageNumber="6">Paratylodiplax blephariskios</emphasis>
(
<bibRefCitation id="EFCD4B76D3250C17CD0C9C4CFACDFB86" author="Stebbing" box="[1108,1292,1080,1106]" pageId="5" pageNumber="6" refString="Stebbing, T. R. R. (1924) South African Crustacea (Part XII of South African Crustacea, for Marine Investigations in South Africa). Annals of the South African Museum, 19, 235 - 248, pls. 1 - 7 (pls. 116 - 122 of Crustacea)." type="journal article" year="1924">Stebbing, 1924</bibRefCitation>
)
</taxonomicName>
– 
<emphasis id="B928EA95D3250C17CC6C9C43FECFFBAC" bold="true" pageId="5" pageNumber="6">
<collectingCountry id="F34B7617D3250C17CC6C9C43FECFFBAC" name="South Africa" pageId="5" pageNumber="6">SOUTH AFRICA</collectingCountry>
</emphasis>
: 
<specimenCount id="9D5AFD0ED3250C17C8479C2AFEB2FBAC" box="[287,371,1118,1144]" pageId="5" pageNumber="6" type="male">1 male</specimenCount>
(5.4 x 
<quantity id="4CA49B62D3250C17C8979C2AFDF2FBAC" box="[463,563,1118,1144]" metricMagnitude="-3" metricUnit="m" metricValue="8.8" pageId="5" pageNumber="6" unit="mm" value="8.8">8.8 mm</quantity>
), 
<specimenCount id="9D5AFD0ED3250C17CB129C2AFD77FBAC" box="[586,694,1118,1144]" pageId="5" pageNumber="6" type="female">1 female</specimenCount>
(4.7 x 7.0 mm) (
<collectionCode id="ED4DAE42D3250C17CAD49C2AFC27FBAC" LSID="urn:lsid:biocol.org:col:34871" box="[908,998,1118,1144]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="6">USNM</collectionCode>
137247), Cape Town, coll. J. Day, 
<date id="FFE21047D3250C17C9CF9CF1FE98FB4B" box="[151,345,1157,1183]" pageId="5" pageNumber="6" value="1957-09">September 1957</date>
.
</paragraph>
</subSubSection>
</treatment>
</document>