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<document id="59B01849382A29B7C2BD9512A5F8F2E4" ID-DOI="10.5281/zenodo.4689879" ID-GBIF-Dataset="cccd251f-0aff-456c-aecc-5eccc5824fe8" ID-ISSN="1638-9387" ID-Zenodo-Dep="4689879" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="juliana" IM.taxonomicNames_approvedBy="admin" IM.treatments_approvedBy="admin" checkinTime="1618437896095" checkinUser="felipe" docAuthor="Biseswar, Ramlall" docDate="2006" docId="03F53669FFE3FFF9FCA1BC0C2212C4B0" docLanguage="en" docName="Zoosystema.28.4.853-864.pdf" docOrigin="Zoosystema 28 (4)" docStyle="DocumentStyle:2A9C6F9EA5F90D98170D9E0D9CA29375.4:Zoosystema.2000-2008.journal_article.type1" docStyleId="2A9C6F9EA5F90D98170D9E0D9CA29375" docStyleName="Zoosystema.2000-2008.journal_article.type1" docStyleVersion="4" docTitle="Bengalus longiductus Biseswar 2006, n. sp." docType="treatment" docVersion="9" lastPageNumber="860" masterDocId="FFCC4E11FFE6FFFEFFBCB83C205BC137" masterDocTitle="Additions to the deep-sea echiuran (Echiura) fauna of the North-East Atlantic" masterLastPageNumber="864" masterPageNumber="853" pageNumber="858" updateTime="1720370326658" updateUser="admin" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="1BAC0FCE32CB767D285A77D2F53569A0" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DCBA6D3F1E438CA8F5F70C405CE52CA2">
<mods:title id="38357B4416F0FE157DD3B90E2928A2F2">Additions to the deep-sea echiuran (Echiura) fauna of the North-East Atlantic</mods:title>
</mods:titleInfo>
<mods:name id="589749436002CF159ACFBF7862DF3AB7" type="personal">
<mods:role id="7EF7F98066F0B3BD6D670E202CB65361">
<mods:roleTerm id="421D8609BE9E7844F5F19D9347A6DB15">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="47FBCF2981AC407762BA44567B1BABA4">Biseswar, Ramlall</mods:namePart>
<mods:affiliation id="FDB38E804E95ECAC7DA8FCBF53E2398D">School of Biological and Environmental Sciences, University of KwaZulu-Natal, Westville Campus, Private Bag x 54001, Durban 4000 (Republic of South Africa) moodleygk @ ukzn. ac. za</mods:affiliation>
<mods:nameIdentifier id="7B745A16FB7E9CE5A261EB020C8735CA" type="email">moodleygk@ukzn.ac.za</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="52D73AEE32EEFCEA0B2D0E1F78E2596C">text</mods:typeOfResource>
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<mods:titleInfo id="137B3657E26269A828FAAAE2F5955E37">
<mods:title id="8099ECA46B1E72BB207FA288DFFE08AB">Zoosystema</mods:title>
</mods:titleInfo>
<mods:part id="9439C6265B895D5AEB3BE3EB65B5C496">
<mods:date id="50868270448E87320379944C843DA6EF">2006</mods:date>
<mods:detail id="D2550D17ECF11CE7FF4FFCF13022DC52" type="volume">
<mods:number id="5AAE85F2CDC8A30A5414E8E359E97A77">28</mods:number>
</mods:detail>
<mods:detail id="6495226C76287037DF5DBC2D8976148C" type="issue">
<mods:number id="68FBADC9BC6C6F5ADB065583B2FF79AE">4</mods:number>
</mods:detail>
<mods:extent id="93F0965D46C3839D56FE37857FCDEB05" unit="page">
<mods:start id="375F6612BFEBF18430C82B0B84628546">853</mods:start>
<mods:end id="390E3CD2004D432EC5FD883EC4BC8F96">864</mods:end>
</mods:extent>
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<mods:classification id="1299639D8ED437D74C37985176D2E01F">journal article</mods:classification>
<mods:identifier id="3B9E5762203670F6CE270E312A838CC6" type="DOI">10.5281/zenodo.4689879</mods:identifier>
<mods:identifier id="FF5CBAF02699B99F4AD3687C5D92ACD3" type="GBIF-Dataset">cccd251f-0aff-456c-aecc-5eccc5824fe8</mods:identifier>
<mods:identifier id="D07C778F16AEF713592F2B88096B0688" type="ISSN">1638-9387</mods:identifier>
<mods:identifier id="26BBF40AC2C60B8BFF904661E7C521E3" type="Zenodo-Dep">4689879</mods:identifier>
</mods:mods>
<treatment id="03F53669FFE3FFF9FCA1BC0C2212C4B0" ID-DOI="http://doi.org/10.5281/zenodo.4893235" ID-GBIF-Taxon="182484731" ID-Zenodo-Dep="4893235" LSID="urn:lsid:plazi:treatment:03F53669FFE3FFF9FCA1BC0C2212C4B0" httpUri="http://treatment.plazi.org/id/03F53669FFE3FFF9FCA1BC0C2212C4B0" lastPageId="7" lastPageNumber="860" pageId="5" pageNumber="858">
<subSubSection id="C346D4F4FFE3FFFBFCA1BC0C2419C57D" box="[797,1090,1072,1098]" pageId="5" pageNumber="858" type="nomenclature">
<paragraph id="8BE3877FFFE3FFFBFCA1BC0C2419C57D" blockId="5.[797,1090,1072,1129]" box="[797,1090,1072,1098]" pageId="5" pageNumber="858">
<heading id="D0AB3013FFE3FFFBFCA1BC0C2419C57D" box="[797,1090,1072,1098]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="858" reason="2">
<taxonomicName id="4C5CFCFCFFE3FFFBFCA1BC0C23A4C57D" ID-CoL="LH7K" authority="Biseswar, 2006" authorityName="Biseswar" authorityYear="2006" box="[797,1023,1072,1098]" class="Polychaeta" family="Bonelliidae" genus="Bengalus" kingdom="Animalia" order="Echiuroidea" pageId="5" pageNumber="858" phylum="Annelida" rank="species" species="longiductus" status="sp. nov.">
<emphasis id="B9285B6DFFE3FFFBFCA1BC0C23A4C57D" bold="true" box="[797,1023,1072,1098]" italics="true" pageId="5" pageNumber="858">Bengalus longiductus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21BE616FFE3FFFBFBBBBC0C2419C57D" box="[1031,1090,1072,1098]" pageId="5" pageNumber="858" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C346D4F4FFE3FFFBFCCCBC7323B5C55E" box="[880,1006,1103,1129]" pageId="5" pageNumber="858" type="description">
<paragraph id="8BE3877FFFE3FFFBFCCCBC7323B5C55E" blockId="5.[797,1090,1072,1129]" box="[880,1006,1103,1129]" pageId="5" pageNumber="858">
(
<figureCitation id="13679BFAFFE3FFFBFCC4BC7323EDC55E" box="[888,950,1103,1129]" captionStart="FIG" captionStartId="6.[113,124,1589,1606]" captionTargetBox="[167,1200,222,1556]" captionTargetId="figure-14@6.[116,1212,131,1560]" captionTargetPageId="6" captionText="FIG. 2. — A-D, Eubonellia noratlanticum DattaGupta, 1981; A, ventral view of anterior part of female; B, gonoduct; C, anterior end of trunk cavity showing the blood system; D, anal vesicles; E-G, Bengalus longiductus n. gen., n. sp.; E, ventral view of anterior part of female; F, gonoduct; G, anal vesicle. Abbreviations: av, anal vesicle; cc, cloacal chamber; dv, dorsal vessel; gd, gonoduct; gp, gonopore; gs, gonostome; int, intestine; nc, nerve cord; niv, neurointestinal vessel; re, rectum; vv, ventral vessel. Scale bars: A, C, F, 3 mm; B, 1.5 mm; D, G, 1 mm; E, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4689883" httpUri="https://zenodo.org/record/4689883/files/figure.png" pageId="5" pageNumber="858">Fig. 2</figureCitation>
E-G)
</paragraph>
</subSubSection>
<subSubSection id="C346D4F4FFE3FFFBFD19BCB824F1C432" pageId="5" pageNumber="858" type="materials_examined">
<paragraph id="8BE3877FFFE3FFFBFD19BCB824FEC5E7" blockId="5.[675,1210,1156,1354]" pageId="5" pageNumber="858">
<materialsCitation id="3B348D22FFE3FFFBFD19BCB824FEC5E7" ID-GBIF-Occurrence="3125549302" collectingDate="1997-04-01" collectionCode="TYPE, MATERIAL" elevation="4844" latitude="48.8865" longLatPrecision="11" longitude="-16.599667" pageId="5" pageNumber="858" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<collectionCode id="ED4D1FBAFFE3FFFBFD19BCB8236CC5AD" box="[677,823,1156,1179]" pageId="5" pageNumber="858">TYPE MATERIAL</collectionCode>
. —
<typeStatus id="54E739DDFFE3FFFBFCDEBCB82399C5AC" box="[866,962,1156,1179]" pageId="5" pageNumber="858" type="holotype">Holotype</typeStatus>
: Porcupine Abyssal Plain, BENGAL 2, Discovery 226, stn 13078#11,
<geoCoordinate id="EE68E1B8FFE3FFFBFBFDBCA224EEC582" box="[1089,1205,1182,1205]" degrees="48" direction="north" minutes="53.19" orientation="latitude" pageId="5" pageNumber="858" precision="9" value="48.8865">48°53.19’N</geoCoordinate>
,
<geoCoordinate id="EE68E1B8FFE3FFFBFD1FBC852341C5E7" box="[675,794,1209,1232]" degrees="16" direction="west" minutes="35.98" orientation="longitude" pageId="5" pageNumber="858" precision="9" value="-16.599667">16°35.98’W</geoCoordinate>
,
<quantity id="4CA42A9AFFE3FFFBFC99BC85232FC5E7" box="[805,884,1209,1232]" metricMagnitude="3" metricUnit="m" metricValue="4.843999999999999" pageId="5" pageNumber="858" unit="m" value="4844.0">
<elevation id="0071604CFFE3FFFBFC99BC85232FC5E7" box="[805,884,1209,1232]" metricMagnitude="3" metricUnit="m" metricValue="4.843999999999999" pageId="5" pageNumber="858" unit="m" value="4844.0">4844 m</elevation>
</quantity>
,
<date id="FFE2A1BFFFE3FFFBFCC2BC8523B8C5E7" box="[894,995,1209,1232]" pageId="5" pageNumber="858" value="1997-04-01">
<collectingDate id="EFA65857FFE3FFFBFCC2BC8523B8C5E7" box="[894,995,1209,1232]" pageId="5" pageNumber="858" value="1997-04-01">1.IV.1997</collectingDate>
</date>
, sexually mature
<specimenCount id="9D5A4CF6FFE3FFFBFB31BC8624C5C5E6" box="[1165,1182,1210,1233]" pageId="5" pageNumber="858" type="female">♀</specimenCount>
.
</materialsCitation>
</paragraph>
<paragraph id="8BE3877FFFE3FFFBFD19BCEF24F1C432" blockId="5.[675,1210,1156,1354]" pageId="5" pageNumber="858">
<typeStatus id="54E739DDFFE3FFFBFD19BCEF22A7C5DD" box="[677,764,1235,1258]" pageId="5" pageNumber="858" type="paratype">Paratype</typeStatus>
: BENGAL 2, Discovery 226, stn 13078#06,
<geoCoordinate id="EE68E1B8FFE3FFFBFD19BCD22341C432" box="[677,794,1262,1285]" degrees="48" direction="north" minutes="44.55" orientation="latitude" pageId="5" pageNumber="858" precision="9" value="48.7425">48°44.55’N</geoCoordinate>
,
<geoCoordinate id="EE68E1B8FFE3FFFBFC99BCD223C7C432" box="[805,924,1262,1285]" degrees="16" direction="west" minutes="32.77" orientation="longitude" pageId="5" pageNumber="858" precision="9" value="-16.546167">16°32.77’W</geoCoordinate>
,
<quantity id="4CA42A9AFFE3FFFBFC1BBCD223ADC432" box="[935,1014,1262,1285]" metricMagnitude="3" metricUnit="m" metricValue="4.838" pageId="5" pageNumber="858" unit="m" value="4838.0">4838 m</quantity>
,
<date id="FFE2A1BFFFE3FFFBFBBCBCD2242EC432" box="[1024,1141,1262,1285]" pageId="5" pageNumber="858" value="1997-03-30">30.III.1997</date>
,
<specimenCount id="9D5A4CF6FFE3FFFBFBC3BCD224F8C432" box="[1151,1187,1262,1285]" pageId="5" pageNumber="858" type="female">1 ♀</specimenCount>
.
</paragraph>
</subSubSection>
<subSubSection id="C346D4F4FFE3FFFBFD19BD2523E5C47D" pageId="5" pageNumber="858" type="etymology">
<paragraph id="8BE3877FFFE3FFFBFD19BD2523E5C47D" blockId="5.[675,1210,1156,1354]" pageId="5" pageNumber="858">ETYMOLOGY. — The species is named on account of the presence of a long gonoduct.</paragraph>
</subSubSection>
<subSubSection id="C346D4F4FFE3FFF9FD19BD512279C2DC" lastPageId="7" lastPageNumber="860" pageId="5" pageNumber="858" type="description">
<paragraph id="8BE3877FFFE3FFFBFD19BD51236EC4B2" blockId="5.[675,1212,1389,1700]" box="[677,821,1389,1415]" pageId="5" pageNumber="858">DESCRIPTION</paragraph>
<paragraph id="8BE3877FFFE3FFF9FD19BDB021E0C086" blockId="5.[675,1212,1389,1700]" lastBlockId="7.[111,648,216,1003]" lastPageId="7" lastPageNumber="860" pageId="5" pageNumber="858">
Colour of proboscis of preserved specimen is cream, trunk is light pink. Only basal stump of proboscis is attached to trunk, distal part is missing. Junction of proboscis and trunk is stout. Lateral margins of proboscis curl inwards proximally but do not fuse to form funnel around mouth (
<figureCitation id="13679BFAFFE3FFFBFBBEBE172403C772" box="[1026,1112,1579,1605]" captionStart="FIG" captionStartId="6.[113,124,1589,1606]" captionTargetBox="[167,1200,222,1556]" captionTargetId="figure-14@6.[116,1212,131,1560]" captionTargetPageId="6" captionText="FIG. 2. — A-D, Eubonellia noratlanticum DattaGupta, 1981; A, ventral view of anterior part of female; B, gonoduct; C, anterior end of trunk cavity showing the blood system; D, anal vesicles; E-G, Bengalus longiductus n. gen., n. sp.; E, ventral view of anterior part of female; F, gonoduct; G, anal vesicle. Abbreviations: av, anal vesicle; cc, cloacal chamber; dv, dorsal vessel; gd, gonoduct; gp, gonopore; gs, gonostome; int, intestine; nc, nerve cord; niv, neurointestinal vessel; re, rectum; vv, ventral vessel. Scale bars: A, C, F, 3 mm; B, 1.5 mm; D, G, 1 mm; E, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4689883" httpUri="https://zenodo.org/record/4689883/files/figure.png" pageId="5" pageNumber="858">Fig. 2E</figureCitation>
). Mouth is an oval aperture located on round muscular projection. Trunk is cylindrical. In
<typeStatus id="54E739DDFFE3FFFBFBA6BE562422C7B3" box="[1050,1145,1642,1668]" pageId="5" pageNumber="858" type="holotype">holotype</typeStatus>
trunk measures
<quantity id="4CA42A9AFFE3FFFBFCB9BEB62314C793" box="[773,847,1674,1700]" metricMagnitude="-2" metricUnit="m" metricValue="4.1" pageId="5" pageNumber="858" unit="mm" value="41.0">41 mm</quantity>
in length and
<quantity id="4CA42A9AFFE3FFFBFC63BEB62447C793" box="[991,1052,1674,1700]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="858" unit="mm" value="8.0">8 mm</quantity>
across broadest part. Corresponding measurements of
<typeStatus id="54E739DDFFE1FFF9FDBBB8E42239C1C5" box="[519,610,216,242]" pageId="7" pageNumber="860" type="paratype">paratype</typeStatus>
are 60 and
<quantity id="4CA42A9AFFE1FFF9FF7AB8C42156C025" box="[198,269,248,274]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="7" pageNumber="860" unit="mm" value="7.0">7 mm</quantity>
. Integument is thick and opaque. Papillae are microscopic, more closely aggregated at extremities of trunk. Genital pore is conspicuous transverse slit located on round eminence of body wall about
<quantity id="4CA42A9AFFE1FFF9FF5BB94B217CC0A6" box="[231,295,375,401]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="860" unit="mm" value="6.0">6 mm</quantity>
away from anterior end of trunk (
<figureCitation id="13679BFAFFE1FFF9FFC5B9AB2097C086" box="[121,204,407,433]" captionStart="FIG" captionStartId="6.[113,124,1589,1606]" captionTargetBox="[167,1200,222,1556]" captionTargetId="figure-14@6.[116,1212,131,1560]" captionTargetPageId="6" captionText="FIG. 2. — A-D, Eubonellia noratlanticum DattaGupta, 1981; A, ventral view of anterior part of female; B, gonoduct; C, anterior end of trunk cavity showing the blood system; D, anal vesicles; E-G, Bengalus longiductus n. gen., n. sp.; E, ventral view of anterior part of female; F, gonoduct; G, anal vesicle. Abbreviations: av, anal vesicle; cc, cloacal chamber; dv, dorsal vessel; gd, gonoduct; gp, gonopore; gs, gonostome; int, intestine; nc, nerve cord; niv, neurointestinal vessel; re, rectum; vv, ventral vessel. Scale bars: A, C, F, 3 mm; B, 1.5 mm; D, G, 1 mm; E, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4689883" httpUri="https://zenodo.org/record/4689883/files/figure.png" pageId="7" pageNumber="860">Fig. 2E</figureCitation>
). Ventral setae absent.
</paragraph>
<caption id="DF23D7F7FFE0FFF8FFCDBE0921B7C796" ID-DOI="http://doi.org/10.5281/zenodo.4689883" ID-Zenodo-Dep="4689883" httpUri="https://zenodo.org/record/4689883/files/figure.png" pageId="6" pageNumber="859" startId="6.[113,124,1589,1606]" targetBox="[167,1200,222,1556]" targetPageId="6">
<paragraph id="8BE3877FFFE0FFF8FFCDBE0921B7C796" blockId="6.[113,1211,1589,1697]" pageId="6" pageNumber="859">
FIG. 2. —
<emphasis id="B9285B6DFFE0FFF8FF79BE0920BCC771" bold="true" box="[197,231,1589,1606]" pageId="6" pageNumber="859">A -D</emphasis>
,
<taxonomicName id="4C5CFCFCFFE0FFF8FF4EBE092205C771" authority="DattaGupta, 1981" authorityName="DattaGupta" authorityYear="1981" box="[242,606,1589,1606]" class="Polychaeta" family="Bonelliidae" genus="Eubonellia" kingdom="Animalia" order="Echiuroidea" pageId="6" pageNumber="859" phylum="Annelida" rank="species" species="noratlanticum">
<emphasis id="B9285B6DFFE0FFF8FF4EBE0921E4C771" box="[242,447,1589,1606]" italics="true" pageId="6" pageNumber="859">Eubonellia noratlanticum</emphasis>
<bibRefCitation id="EFCDFA8EFFE0FFF8FE7ABE092205C771" author="DATTAGUPTA A. K." box="[454,606,1589,1606]" pageId="6" pageNumber="859" pagination="353 - 378" refId="ref6182" refString="DATTAGUPTA A. K. 1981. - Atlantic echiurans. Part 1. Report on twenty-two species of deep-sea echiurans of the North and the South Atlantic Oceans. Bulletin du Museum national d'Histoire naturelle Paris, 4 e ser., sect. A, 3 (2): 353 - 378." type="journal article" year="1981">DattaGupta, 1981</bibRefCitation>
</taxonomicName>
;
<emphasis id="B9285B6DFFE0FFF8FDDBBE09222FC771" bold="true" box="[615,628,1589,1606]" pageId="6" pageNumber="859">A</emphasis>
, ventral view of anterior part of female;
<emphasis id="B9285B6DFFE0FFF8FC7BBE09238FC771" bold="true" box="[967,980,1589,1606]" pageId="6" pageNumber="859">B</emphasis>
, gonoduct;
<emphasis id="B9285B6DFFE0FFF8FB87BE092412C771" bold="true" box="[1083,1097,1589,1606]" pageId="6" pageNumber="859">C</emphasis>
, anterior end of trunk cavity showing the blood system;
<emphasis id="B9285B6DFFE0FFF8FE63BE7021B6C76A" bold="true" box="[479,493,1612,1629]" pageId="6" pageNumber="859">D</emphasis>
, anal vesicles;
<emphasis id="B9285B6DFFE0FFF8FDCDBE7022C8C76A" bold="true" box="[625,659,1612,1629]" pageId="6" pageNumber="859">E -G</emphasis>
,
<taxonomicName id="4C5CFCFCFFE0FFF8FD23BE70230EC76A" authority="Biseswar, 2006" authorityName="Biseswar" authorityYear="2006" box="[671,853,1612,1629]" class="Polychaeta" family="Bonelliidae" genus="Bengalus" kingdom="Animalia" order="Echiuroidea" pageId="6" pageNumber="859" phylum="Annelida" rank="species" species="longiductus" status="n. gen., n. sp.">
<emphasis id="B9285B6DFFE0FFF8FD23BE70230EC76A" box="[671,853,1612,1629]" italics="true" pageId="6" pageNumber="859">Bengalus longiductus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21BE616FFE0FFF8FCE0BE70238DC76A" box="[860,982,1612,1629]" pageId="6" pageNumber="859" rank="species">n. gen., n. sp.</taxonomicNameLabel>
;
<emphasis id="B9285B6DFFE0FFF8FC62BE7023B1C76A" bold="true" box="[990,1002,1612,1629]" pageId="6" pageNumber="859">E</emphasis>
, ventral view of anterior part of female;
<emphasis id="B9285B6DFFE0FFF8FF4EBE5F20A6C743" bold="true" box="[242,253,1635,1652]" pageId="6" pageNumber="859">F</emphasis>
, gonoduct;
<emphasis id="B9285B6DFFE0FFF8FEDFBE5F212AC743" bold="true" box="[355,369,1635,1652]" pageId="6" pageNumber="859">G</emphasis>
, anal vesicle. Abbreviations:
<emphasis id="B9285B6DFFE0FFF8FDDBBE5F2220C743" bold="true" box="[615,635,1635,1652]" pageId="6" pageNumber="859">av</emphasis>
, anal vesicle;
<emphasis id="B9285B6DFFE0FFF8FD4FBE5F2353C743" bold="true" box="[755,776,1635,1652]" pageId="6" pageNumber="859">cc</emphasis>
, cloacal chamber;
<emphasis id="B9285B6DFFE0FFF8FC15BE5F23E5C743" bold="true" box="[937,958,1635,1652]" pageId="6" pageNumber="859">dv</emphasis>
, dorsal vessel;
<emphasis id="B9285B6DFFE0FFF8FBFEBE5F2402C743" bold="true" box="[1090,1113,1635,1652]" pageId="6" pageNumber="859">gd</emphasis>
, gonoduct;
<emphasis id="B9285B6DFFE0FFF8FFCDBE4520D3C7BD" bold="true" box="[113,136,1657,1674]" pageId="6" pageNumber="859">gp</emphasis>
, gonopore;
<emphasis id="B9285B6DFFE0FFF8FF51BE452159C7BD" bold="true" box="[237,258,1657,1674]" pageId="6" pageNumber="859">gs</emphasis>
, gonostome;
<emphasis id="B9285B6DFFE0FFF8FECABE4521D7C7BD" bold="true" box="[374,396,1657,1674]" pageId="6" pageNumber="859">int</emphasis>
, intestine;
<emphasis id="B9285B6DFFE0FFF8FE5BBE4521A6C7BD" bold="true" box="[487,509,1657,1674]" pageId="6" pageNumber="859">nc</emphasis>
, nerve cord;
<emphasis id="B9285B6DFFE0FFF8FDD6BE4522DFC7BD" bold="true" box="[618,644,1657,1674]" pageId="6" pageNumber="859">niv</emphasis>
, neurointestinal vessel;
<emphasis id="B9285B6DFFE0FFF8FCF0BE452305C7BD" bold="true" box="[844,862,1657,1674]" pageId="6" pageNumber="859">re</emphasis>
, rectum;
<emphasis id="B9285B6DFFE0FFF8FC10BE4523E4C7BD" bold="true" box="[940,959,1657,1674]" pageId="6" pageNumber="859">vv</emphasis>
, ventral vessel. Scale bars: A, C, F, 3 mm; B, 1.5 mm; D, G, 1 mm; E, 2 mm.
</paragraph>
</caption>
<paragraph id="8BE3877FFFE1FFF9FF34B98A2279C2DC" blockId="7.[111,648,216,1003]" pageId="7" pageNumber="860">
Gonoduct single, elongated, tubular, about threequarters length of trunk, on right side of nerve cord (
<figureCitation id="13679BFAFFE1FFF9FFCBB9CA209FC327" box="[119,196,502,528]" captionStart="FIG" captionStartId="6.[113,124,1589,1606]" captionTargetBox="[167,1200,222,1556]" captionTargetId="figure-14@6.[116,1212,131,1560]" captionTargetPageId="6" captionText="FIG. 2. — A-D, Eubonellia noratlanticum DattaGupta, 1981; A, ventral view of anterior part of female; B, gonoduct; C, anterior end of trunk cavity showing the blood system; D, anal vesicles; E-G, Bengalus longiductus n. gen., n. sp.; E, ventral view of anterior part of female; F, gonoduct; G, anal vesicle. Abbreviations: av, anal vesicle; cc, cloacal chamber; dv, dorsal vessel; gd, gonoduct; gp, gonopore; gs, gonostome; int, intestine; nc, nerve cord; niv, neurointestinal vessel; re, rectum; vv, ventral vessel. Scale bars: A, C, F, 3 mm; B, 1.5 mm; D, G, 1 mm; E, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4689883" httpUri="https://zenodo.org/record/4689883/files/figure.png" pageId="7" pageNumber="860">Fig. 2F</figureCitation>
). Gonoduct of
<typeStatus id="54E739DDFFE1FFF9FEE3B9CA21E0C327" box="[351,443,502,528]" pageId="7" pageNumber="860" type="holotype">holotype</typeStatus>
contains few round eggs. Gonostome stalked, located on one side near distal end of gonoduct, with minute lobes around margin (
<figureCitation id="13679BFAFFE1FFF9FF74BA69214EC358" box="[200,277,597,623]" captionStart="FIG" captionStartId="6.[113,124,1589,1606]" captionTargetBox="[167,1200,222,1556]" captionTargetId="figure-14@6.[116,1212,131,1560]" captionTargetPageId="6" captionText="FIG. 2. — A-D, Eubonellia noratlanticum DattaGupta, 1981; A, ventral view of anterior part of female; B, gonoduct; C, anterior end of trunk cavity showing the blood system; D, anal vesicles; E-G, Bengalus longiductus n. gen., n. sp.; E, ventral view of anterior part of female; F, gonoduct; G, anal vesicle. Abbreviations: av, anal vesicle; cc, cloacal chamber; dv, dorsal vessel; gd, gonoduct; gp, gonopore; gs, gonostome; int, intestine; nc, nerve cord; niv, neurointestinal vessel; re, rectum; vv, ventral vessel. Scale bars: A, C, F, 3 mm; B, 1.5 mm; D, G, 1 mm; E, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4689883" httpUri="https://zenodo.org/record/4689883/files/figure.png" pageId="7" pageNumber="860">Fig. 2F</figureCitation>
). Except for foregut, rest of intestine and blood system damaged in
<typeStatus id="54E739DDFFE1FFF9FE03BA48227AC3B9" box="[447,545,628,654]" pageId="7" pageNumber="860" type="holotype">holotype</typeStatus>
. In
<typeStatus id="54E739DDFFE1FFF9FDF1BA4820F9C399" pageId="7" pageNumber="860" type="paratype">paratype</typeStatus>
, foregut is straight tube. Pharynx is somewhat dilated, clearly demarcated from oesophagus. Intestine filled with fine sand not moulded into faecal pellets. Cloacal chamber is thick and bulbous. In
<typeStatus id="54E739DDFFE1FFF9FFCEBB2F2097C21A" box="[114,204,787,813]" pageId="7" pageNumber="860" type="paratype">paratype</typeStatus>
dorsal vessel is long, passing anteriorly and entering proboscis. Neurointestinal vessel arises as single vessel and splits into two vessels which open separately into ventral vessel. Anal vesicles small, tubular, slightly distended distally, opening into cloacal bulb (
<figureCitation id="13679BFAFFE1FFF9FF69BB8D2177C2FC" box="[213,300,945,971]" captionStart="FIG" captionStartId="6.[113,124,1589,1606]" captionTargetBox="[167,1200,222,1556]" captionTargetId="figure-14@6.[116,1212,131,1560]" captionTargetPageId="6" captionText="FIG. 2. — A-D, Eubonellia noratlanticum DattaGupta, 1981; A, ventral view of anterior part of female; B, gonoduct; C, anterior end of trunk cavity showing the blood system; D, anal vesicles; E-G, Bengalus longiductus n. gen., n. sp.; E, ventral view of anterior part of female; F, gonoduct; G, anal vesicle. Abbreviations: av, anal vesicle; cc, cloacal chamber; dv, dorsal vessel; gd, gonoduct; gp, gonopore; gs, gonostome; int, intestine; nc, nerve cord; niv, neurointestinal vessel; re, rectum; vv, ventral vessel. Scale bars: A, C, F, 3 mm; B, 1.5 mm; D, G, 1 mm; E, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4689883" httpUri="https://zenodo.org/record/4689883/files/figure.png" pageId="7" pageNumber="860">Fig. 2G</figureCitation>
). Distal ends of vesicles fastened to body wall by thin mesenteric strands.
</paragraph>
</subSubSection>
<subSubSection id="C346D4F4FFE1FFF9FFCDBC2C2212C4B0" pageId="7" pageNumber="860" type="discussion">
<paragraph id="8BE3877FFFE1FFF9FFCDBC2C208EC51F" blockId="7.[111,648,1040,1415]" box="[113,213,1040,1066]" pageId="7" pageNumber="860">REMARKS</paragraph>
<paragraph id="8BE3877FFFE1FFF9FFCDBC0C2212C4B0" blockId="7.[111,648,1040,1415]" pageId="7" pageNumber="860">
The new genus
<taxonomicName id="4C5CFCFCFFE1FFF9FEB7BC0C213AC57D" authority="Biseswar, 2006" authorityName="Biseswar" authorityYear="2006" box="[267,353,1072,1098]" class="Polychaeta" family="Bonelliidae" genus="Bengalus" kingdom="Animalia" order="Echiuroidea" pageId="7" pageNumber="860" phylum="Annelida" rank="genus" status="gen. nov.">
<emphasis id="B9285B6DFFE1FFF9FEB7BC0C213AC57D" box="[267,353,1072,1098]" italics="true" pageId="7" pageNumber="860">Bengalus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21BE616FFE1FFF9FEDABC0C21F0C57D" box="[358,427,1072,1098]" pageId="7" pageNumber="860" rank="genus">n. gen.</taxonomicNameLabel>
is nearest to the genus
<taxonomicName id="4C5CFCFCFFE1FFF9FFD3BC6C208DC55D" authorityName="DattaGupta" authorityYear="1981" box="[111,214,1104,1130]" class="Polychaeta" family="Bonelliidae" genus="Charcotus" kingdom="Animalia" order="Echiuroidea" pageId="7" pageNumber="860" phylum="Annelida" rank="genus">
<emphasis id="B9285B6DFFE1FFF9FFD3BC6C208DC55D" box="[111,214,1104,1130]" italics="true" pageId="7" pageNumber="860">Charcotus</emphasis>
</taxonomicName>
in possessing a single gonoduct, tubular anal vesicles and in lacking ventral setae. In both genera, the anal vesicles open into a cloacal bulb. In the genus
<taxonomicName id="4C5CFCFCFFE1FFF9FF61BC932163C5FE" authority="Biseswar, 2006" authorityName="Biseswar" authorityYear="2006" box="[221,312,1199,1225]" class="Polychaeta" family="Bonelliidae" genus="Bengalus" kingdom="Animalia" order="Echiuroidea" pageId="7" pageNumber="860" phylum="Annelida" rank="genus" status="gen. nov.">
<emphasis id="B9285B6DFFE1FFF9FF61BC932163C5FE" box="[221,312,1199,1225]" italics="true" pageId="7" pageNumber="860">Bengalus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21BE616FFE1FFF9FE82BC9321D2C5FE" box="[318,393,1199,1225]" pageId="7" pageNumber="860" rank="genus">n. gen.</taxonomicNameLabel>
, however, the gonoduct is long and tubular and the gonostome is located on one side near the distal end of the gonoduct. Furthermore, in the genus
<taxonomicName id="4C5CFCFCFFE1FFF9FE2DBD3221A3C41F" authorityName="DattaGupta" authorityYear="1981" box="[401,504,1294,1320]" class="Polychaeta" family="Bonelliidae" genus="Charcotus" kingdom="Animalia" order="Echiuroidea" pageId="7" pageNumber="860" phylum="Annelida" rank="genus">
<emphasis id="B9285B6DFFE1FFF9FE2DBD3221A3C41F" box="[401,504,1294,1320]" italics="true" pageId="7" pageNumber="860">Charcotus</emphasis>
</taxonomicName>
the mouth is located in an oral cup formed by the inflection of the lateral margins of the proboscis proximally and the body wall is covered with thick papillae.
</paragraph>
</subSubSection>
</treatment>
</document>
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