246 lines
38 KiB
XML
246 lines
38 KiB
XML
<document id="645D78022F97A019F688305CA185B84D" ID-CLB-Dataset="27402" ID-DOI="10.11646/zootaxa.4577.1.12" ID-GBIF-Dataset="67e2ba86-b010-49d8-ba6e-a4845470e725" ID-ISSN="1175-5326" ID-Zenodo-Dep="2628556" ID-ZooBank="4ED3987D-586E-4429-9241-D760944947F5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554387032861" checkinUser="plazi" docAuthor="Kovalev, Alexey V., Kirejtshuk, Alexander G. & Shapovalov, Andrey M." docDate="2019" docId="03F287C2FFEBFF82AB8CFE7AFDE31F01" docLanguage="en" docName="zootaxa.4577.1.12.pdf" docOrigin="Zootaxa 4577 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Drilorhinus Kovalev & Kirejtshuk & Shapovalov 2019, gen. nov." docType="treatment" docVersion="8" lastPageNumber="189" masterDocId="FFCBFFBAFFEAFF80AB1BFF88FFBB1A0C" masterDocTitle="Drilorhinus, a new genus of the family Drilidae Lacordaire, 1857 (Coleoptera: Elateroidea) from Iran" masterLastPageNumber="194" masterPageNumber="187" pageNumber="188" updateTime="1698718418615" updateUser="ExternalLinkService">
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<mods:title id="C227E5BF8834D9FFEB717FF476C43D46">Drilorhinus, a new genus of the family Drilidae Lacordaire, 1857 (Coleoptera: Elateroidea) from Iran</mods:title>
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<mods:name id="D97D0FDEDA4575D2975285B869EDDBE1" type="personal">
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<mods:namePart id="640EDB511C14280DC68B7DB9B26F6DA0">Kovalev, Alexey V.</mods:namePart>
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<mods:namePart id="B4ED0DA48BE599BDB4E417C38A8DA3F5">Kirejtshuk, Alexander G.</mods:namePart>
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<mods:name id="91833B2D03A4E75735AEAB19528ACD6B" type="personal">
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<mods:roleTerm id="9F1628C2706E797513727A3A6281B64F">Author</mods:roleTerm>
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<mods:namePart id="B55D0668EB6AB19548208611E702391E">Shapovalov, Andrey M.</mods:namePart>
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<mods:title id="8E23B734F2F112208584ADD2F0A6CE45">Zootaxa</mods:title>
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<mods:part id="8024C5691782F2D4477DB0F4A23D49BD">
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<mods:date id="980BDB8740B825D6700D6D6BFEB42DA4">2019</mods:date>
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<treatment id="03F287C2FFEBFF82AB8CFE7AFDE31F01" ID-DOI="http://doi.org/10.5281/zenodo.5923877" ID-GBIF-Taxon="154553448" ID-Zenodo-Dep="5923877" LSID="urn:lsid:plazi:treatment:03F287C2FFEBFF82AB8CFE7AFDE31F01" httpUri="http://treatment.plazi.org/id/03F287C2FFEBFF82AB8CFE7AFDE31F01" lastPageId="2" lastPageNumber="189" pageId="1" pageNumber="188">
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<subSubSection id="C341655FFFEBFF81AB8CFE7AFE201800" box="[151,411,498,524]" pageId="1" pageNumber="188" type="nomenclature">
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<paragraph id="8BE436D4FFEBFF81AB8CFE7AFE201800" blockId="1.[151,411,498,524]" box="[151,411,498,524]" pageId="1" pageNumber="188">
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<heading id="D0AC81B8FFEBFF81AB8CFE7AFE201800" bold="true" box="[151,411,498,524]" fontSize="11" level="1" pageId="1" pageNumber="188" reason="1">
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<emphasis id="B92FEAC6FFEBFF81AB8CFE7AFE201800" bold="true" box="[151,411,498,524]" pageId="1" pageNumber="188">
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<taxonomicName id="4C5B4D57FFEBFF81AB8CFE7AFE9D1800" authority="Kovalev & Kirejtshuk & Shapovalov, 2019" authorityName="Kovalev & Kirejtshuk & Shapovalov" authorityYear="2019" box="[151,294,498,524]" class="Insecta" family="Drilidae" genus="Drilorhinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="188" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B92FEAC6FFEBFF81AB8CFE7AFE9D1800" bold="true" box="[151,294,498,524]" italics="true" pageId="1" pageNumber="188">Drilorhinus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21C57BDFFEBFF81AA36FE7AFE201800" box="[301,411,498,524]" pageId="1" pageNumber="188" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C341655FFFEBFF81AB8CFDB1FDE71841" box="[151,604,568,590]" pageId="1" pageNumber="188" type="materials_examined">
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<paragraph id="8BE436D4FFEBFF81AB8CFDB1FDE71841" blockId="1.[151,604,568,590]" box="[151,604,568,590]" pageId="1" pageNumber="188">
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<typeStatus id="54E08876FFEBFF81AB8CFDB1FF721842" box="[151,201,569,590]" pageId="1" pageNumber="188">Type</typeStatus>
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species:
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<taxonomicName id="4C5B4D57FFEBFF81AA3DFDB0FDBD1842" authority="Kovalev & Kirejtshuk & Shapovalov, 2019" authorityName="Kovalev & Kirejtshuk & Shapovalov" authorityYear="2019" box="[294,518,568,590]" class="Insecta" family="Drilidae" genus="Drilorhinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="188" phylum="Arthropoda" rank="species" species="klimenkoi" status="sp. nov.">
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<emphasis id="B92FEAC6FFEBFF81AA3DFDB0FDBD1842" box="[294,518,568,590]" italics="true" pageId="1" pageNumber="188">Drilorhinus klimenkoi</emphasis>
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</taxonomicName>
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<emphasis id="B92FEAC6FFEBFF81A917FDB0FDE71841" bold="true" box="[524,604,568,589]" pageId="1" pageNumber="188">
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<taxonomicNameLabel id="A21C57BDFFEBFF81A917FDB0FDE71841" box="[524,604,568,589]" pageId="1" pageNumber="188" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C341655FFFEBFF81AB8CFDF2FDDD18D7" pageId="1" pageNumber="188" type="etymology">
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<paragraph id="8BE436D4FFEBFF81AB8CFDF2FE0518BB" blockId="1.[151,1437,634,2027]" pageId="1" pageNumber="188">
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<emphasis id="B92FEAC6FFEBFF81AB8CFDF2FEA5189F" bold="true" box="[151,286,634,659]" pageId="1" pageNumber="188">Etymology.</emphasis>
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The name of the new genus is formed from the generic name “
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<taxonomicName id="4C5B4D57FFEBFF81A8E9FDF3FB81189F" authorityName="A. Olivier" authorityYear="1790" box="[1010,1082,635,659]" class="Insecta" family="Elateridae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="188" phylum="Arthropoda" rank="genus">
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<emphasis id="B92FEAC6FFEBFF81A8E9FDF3FB81189F" box="[1010,1082,635,659]" italics="true" pageId="1" pageNumber="188">Drilus</emphasis>
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</taxonomicName>
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” and Greek “ῥίς, ῥινός” (nose, snout). Gender masculine.
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</paragraph>
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<paragraph id="8BE436D4FFEBFF81ABDCFD4AFDDD18D7" blockId="1.[151,1437,634,2027]" box="[199,614,706,731]" pageId="1" pageNumber="188">
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<emphasis id="B92FEAC6FFEBFF81ABDCFD4AFEDC18D7" bold="true" box="[199,359,706,731]" pageId="1" pageNumber="188">Composition.</emphasis>
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The
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<typeStatus id="54E08876FFEBFF81AABBFD4BFE6B18D7" box="[416,464,707,731]" pageId="1" pageNumber="188">type</typeStatus>
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species only.
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</paragraph>
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</subSubSection>
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<subSubSection id="C341655FFFEBFF81ABDCFD6EFD961C8F" pageId="1" pageNumber="188" type="diagnosis">
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<paragraph id="8BE436D4FFEBFF81ABDCFD6EFED519F7" blockId="1.[151,1437,634,2027]" pageId="1" pageNumber="188">
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<emphasis id="B92FEAC6FFEBFF81ABDCFD6EFEFA18F3" bold="true" box="[199,321,742,767]" pageId="1" pageNumber="188">Diagnosis.</emphasis>
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Head somewhat wider than anterior margin of pronotum. Frons between antennal insertions strongly protruding forward and forming long subtriangular projection, covering mouthparts from above; antennal insertions located at base of sides of this projection. Antennae pectinate. Labrum small and slightly transverse, freely attached and fully exposed. Mandibles comparatively short and stout, moderately curved, with angularly curved outer margin. Pronotum slightly transverse, subtrapezoid, with lateral ridge distinct only in posterior half. Elytra shortened and somewhat divergent, covered with rough rugose-granulate sculpture. Prosternum with prosternal process short and widely rounded at apex. Mesoventrite V-shaped, with anterior margin deeply and widely emarginate.
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</paragraph>
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<paragraph id="8BE436D4FFEBFF81ABDCFB8EFD961C8F" blockId="1.[151,1437,634,2027]" pageId="1" pageNumber="188">
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<emphasis id="B92FEAC6FFEBFF81ABDCFB8EFED91E13" bold="true" box="[199,354,1030,1055]" pageId="1" pageNumber="188">Comparison.</emphasis>
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A strong projection of the anterior part of the frons between antennal insertions distinguishes the new genus from all known genera of
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<taxonomicName id="4C5B4D57FFEBFF81A924FBA2FD201E4F" authorityName="Lacordaire" authorityYear="1857" box="[575,667,1066,1091]" class="Insecta" family="Drilidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="188" phylum="Arthropoda" rank="family">Drilidae</taxonomicName>
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(
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<taxonomicName id="4C5B4D57FFEBFF81A9B7FBA3FC021E4F" authority="A. Olivier, 1790" authorityName="A. Olivier" authorityYear="1790" box="[684,953,1066,1091]" class="Insecta" family="Elateridae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="188" phylum="Arthropoda" rank="genus">
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<emphasis id="B92FEAC6FFEBFF81A9B7FBA3FD4F1E4F" box="[684,756,1067,1091]" italics="true" pageId="1" pageNumber="188">Drilus</emphasis>
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A. Olivier, 1790
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</taxonomicName>
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;
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<emphasis id="B92FEAC6FFEBFF81A8DFFBA3FBCA1E4F" box="[964,1137,1067,1091]" italics="true" pageId="1" pageNumber="188">Flabelloselasia</emphasis>
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<bibRefCitation id="EFCA4B25FFEBFF81AF62FBA2FA2C1E4F" author="Kundrata, R. & Bocak, L." box="[1145,1431,1066,1091]" pageId="1" pageNumber="188" pagination="441 - 492" refId="ref4148" refString="Kundrata, R. & Bocak, L. (2017) Taxonomic review of Drilini (Elateridae: Agrypninae) in Cameroon reveals high morphological diversity, including the discovery of five new genera. Insect systematics & evolution, 48 (5), 441 - 492. https: // doi. org / 10.1163 / 1876312 X- 48022161" type="journal article" year="2017">Kundrata et Bocak, 2017</bibRefCitation>
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||
;
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<emphasis id="B92FEAC6FFEBFF81AB8CFBC6FE9B1E6A" box="[151,288,1102,1126]" italics="true" pageId="1" pageNumber="188">Kupeselasia</emphasis>
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<bibRefCitation id="EFCA4B25FFEBFF81AA3CFBC6FDFF1E6B" author="Kundrata, R. & Bocak, L." box="[295,580,1102,1127]" pageId="1" pageNumber="188" pagination="441 - 492" refId="ref4148" refString="Kundrata, R. & Bocak, L. (2017) Taxonomic review of Drilini (Elateridae: Agrypninae) in Cameroon reveals high morphological diversity, including the discovery of five new genera. Insect systematics & evolution, 48 (5), 441 - 492. https: // doi. org / 10.1163 / 1876312 X- 48022161" type="journal article" year="2017">Kundrata et Bocak, 2017</bibRefCitation>
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;
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<emphasis id="B92FEAC6FFEBFF81A94BFBC6FD1F1E6A" box="[592,676,1102,1126]" italics="true" pageId="1" pageNumber="188">Lolosia</emphasis>
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<bibRefCitation id="EFCA4B25FFEBFF81A9B7FBC6FC721E6B" author="Kundrata, R. & Bocak, L." box="[684,969,1102,1127]" pageId="1" pageNumber="188" pagination="441 - 492" refId="ref4148" refString="Kundrata, R. & Bocak, L. (2017) Taxonomic review of Drilini (Elateridae: Agrypninae) in Cameroon reveals high morphological diversity, including the discovery of five new genera. Insect systematics & evolution, 48 (5), 441 - 492. https: // doi. org / 10.1163 / 1876312 X- 48022161" type="journal article" year="2017">Kundrata et Bocak, 2017</bibRefCitation>
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;
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<taxonomicName id="4C5B4D57FFEBFF81A8CEFBC6FB441E6B" authority="Bassi, 1834" authorityName="Bassi" authorityYear="1834" box="[981,1279,1102,1127]" class="Insecta" family="Drilidae" genus="Malacogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="188" phylum="Arthropoda" rank="genus">
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<emphasis id="B92FEAC6FFEBFF81A8CEFBC6FBCA1E6A" box="[981,1137,1102,1126]" italics="true" pageId="1" pageNumber="188">Malacogaster</emphasis>
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Bassi, 1834
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</taxonomicName>
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;
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<emphasis id="B92FEAC6FFEBFF81AE10FBC6FA271E6A" box="[1291,1436,1102,1126]" italics="true" pageId="1" pageNumber="188">Microselasia</emphasis>
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<bibRefCitation id="EFCA4B25FFEBFF81AB8CFBFAFE7B1E87" author="Kundrata, R. & Bocak, L." box="[151,448,1138,1163]" pageId="1" pageNumber="188" pagination="441 - 492" refId="ref4148" refString="Kundrata, R. & Bocak, L. (2017) Taxonomic review of Drilini (Elateridae: Agrypninae) in Cameroon reveals high morphological diversity, including the discovery of five new genera. Insect systematics & evolution, 48 (5), 441 - 492. https: // doi. org / 10.1163 / 1876312 X- 48022161" type="journal article" year="2017">Kundrata et Bocak, 2017</bibRefCitation>
|
||
;
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||
<taxonomicName id="4C5B4D57FFEBFF81AACBFBFBFCCB1E87" authority="Laporte de Castelnau, 1836" authorityName="Laporte de Castelnau" authorityYear="1836" box="[464,880,1138,1163]" class="Insecta" family="Drilidae" genus="Selasia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="188" phylum="Arthropoda" rank="genus">
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<emphasis id="B92FEAC6FFEBFF81AACBFBFBFD9A1E87" box="[464,545,1139,1163]" italics="true" pageId="1" pageNumber="188">Selasia</emphasis>
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Laporte de Castelnau, 1836
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</taxonomicName>
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, and
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<emphasis id="B92FEAC6FFEBFF81A8AFFBFBFBE11E87" box="[948,1114,1139,1163]" italics="true" pageId="1" pageNumber="188">Wittmerselasia</emphasis>
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<bibRefCitation id="EFCA4B25FFEBFF81AF7DFBFAFA2A1E87" author="Kundrata, R. & Bocak, L." box="[1126,1425,1138,1163]" pageId="1" pageNumber="188" pagination="441 - 492" refId="ref4148" refString="Kundrata, R. & Bocak, L. (2017) Taxonomic review of Drilini (Elateridae: Agrypninae) in Cameroon reveals high morphological diversity, including the discovery of five new genera. Insect systematics & evolution, 48 (5), 441 - 492. https: // doi. org / 10.1163 / 1876312 X- 48022161" type="journal article" year="2017">Kundrata et Bocak, 2017</bibRefCitation>
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).
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<taxonomicName id="4C5B4D57FFEBFF81AB8CFB1EFEA01EA2" authority="Kovalev & Kirejtshuk & Shapovalov, 2019" authorityName="Kovalev & Kirejtshuk & Shapovalov" authorityYear="2019" box="[151,283,1174,1198]" class="Insecta" family="Drilidae" genus="Drilorhinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="188" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B92FEAC6FFEBFF81AB8CFB1EFEA01EA2" box="[151,283,1174,1198]" italics="true" pageId="1" pageNumber="188">Drilorhinus</emphasis>
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</taxonomicName>
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<emphasis id="B92FEAC6FFEBFF81AA3DFB1EFE281EA3" bold="true" box="[294,403,1174,1199]" pageId="1" pageNumber="188">
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<taxonomicNameLabel id="A21C57BDFFEBFF81AA3DFB1EFE281EA3" box="[294,403,1174,1199]" pageId="1" pageNumber="188" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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is most similar to
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<taxonomicName id="4C5B4D57FFEBFF81A963FB1EFCAF1EA2" authorityName="Bassi" authorityYear="1834" box="[632,788,1174,1198]" class="Insecta" family="Drilidae" genus="Malacogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="188" phylum="Arthropoda" rank="genus">
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<emphasis id="B92FEAC6FFEBFF81A963FB1EFCAF1EA2" box="[632,788,1174,1198]" italics="true" pageId="1" pageNumber="188">Malacogaster</emphasis>
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</taxonomicName>
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in sharing the subtrapezoidal pronotum with shortened lateral ridge, shortened elytra (reaching only the base of abdominal segment 6), and V-shaped mesoventrite with deeply emarginate anterior margin, but the latter genus differs from
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<taxonomicName id="4C5B4D57FFEBFF81A882FB56FBA61EFA" authority="Kovalev & Kirejtshuk & Shapovalov, 2019" authorityName="Kovalev & Kirejtshuk & Shapovalov" authorityYear="2019" box="[921,1053,1246,1270]" class="Insecta" family="Drilidae" genus="Drilorhinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="188" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B92FEAC6FFEBFF81A882FB56FBA61EFA" box="[921,1053,1246,1270]" italics="true" pageId="1" pageNumber="188">Drilorhinus</emphasis>
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</taxonomicName>
|
||
<emphasis id="B92FEAC6FFEBFF81AF3EFB56FB341EFB" bold="true" box="[1061,1167,1246,1271]" pageId="1" pageNumber="188">
|
||
<taxonomicNameLabel id="A21C57BDFFEBFF81AF3EFB56FB341EFB" box="[1061,1167,1246,1271]" pageId="1" pageNumber="188" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in the serrate antennae, very transverse labrum, widely separated falcate mandibles, subparallel-sided pronotum, and shape of the parameres.
|
||
<taxonomicName id="4C5B4D57FFEBFF81AA0CFAAEFEE51F32" authorityName="A. Olivier" authorityYear="1790" box="[279,350,1318,1342]" class="Insecta" family="Elateridae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="188" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92FEAC6FFEBFF81AA0CFAAEFEE51F32" box="[279,350,1318,1342]" italics="true" pageId="1" pageNumber="188">Drilus</emphasis>
|
||
</taxonomicName>
|
||
resembles the new genus in the shape of the mesoventrite, however, in contrast to the new genus,
|
||
<taxonomicName id="4C5B4D57FFEBFF81AB8CFAC3FF641F6F" authorityName="A. Olivier" authorityYear="1790" box="[151,223,1355,1379]" class="Insecta" family="Elateridae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="188" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92FEAC6FFEBFF81AB8CFAC3FF641F6F" box="[151,223,1355,1379]" italics="true" pageId="1" pageNumber="188">Drilus</emphasis>
|
||
</taxonomicName>
|
||
demonstrates the convex lateral pronotal margins with almost complete lateral ridges and not shortened elytra.
|
||
<taxonomicName id="4C5B4D57FFEBFF81ABFFFAE6FE8E1F8A" authorityName="Laporte de Castelnau" authorityYear="1836" box="[228,309,1390,1414]" class="Insecta" family="Drilidae" genus="Selasia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="188" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92FEAC6FFEBFF81ABFFFAE6FE8E1F8A" box="[228,309,1390,1414]" italics="true" pageId="1" pageNumber="188">Selasia</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C5B4D57FFEBFF81AAD3FAE6FDF71F8A" authority="Kovalev & Kirejtshuk & Shapovalov, 2019" authorityName="Kovalev & Kirejtshuk & Shapovalov" authorityYear="2019" box="[456,588,1390,1414]" class="Insecta" family="Drilidae" genus="Drilorhinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="188" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B92FEAC6FFEBFF81AAD3FAE6FDF71F8A" box="[456,588,1390,1414]" italics="true" pageId="1" pageNumber="188">Drilorhinus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B92FEAC6FFEBFF81A948FAE6FD071F8B" bold="true" box="[595,700,1390,1415]" pageId="1" pageNumber="188">
|
||
<taxonomicNameLabel id="A21C57BDFFEBFF81A948FAE6FD071F8B" box="[595,700,1390,1415]" pageId="1" pageNumber="188" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in the concave anterior margin of the mesoventrite, elytra that are not shortened, and flabellate antennae. In addition to the above-mentioned diagnostic characters of
|
||
<taxonomicName id="4C5B4D57FFEBFF81AE03FA1BFA271FA7" authority="Kovalev & Kirejtshuk & Shapovalov, 2019" authorityName="Kovalev & Kirejtshuk & Shapovalov" authorityYear="2019" box="[1304,1436,1427,1451]" class="Insecta" family="Drilidae" genus="Drilorhinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="188" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B92FEAC6FFEBFF81AE03FA1BFA271FA7" box="[1304,1436,1427,1451]" italics="true" pageId="1" pageNumber="188">Drilorhinus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B92FEAC6FFEBFF81AB8CFA3EFEB81FC3" bold="true" box="[151,259,1462,1487]" pageId="1" pageNumber="188">
|
||
<taxonomicNameLabel id="A21C57BDFFEBFF81AB8CFA3EFEB81FC3" box="[151,259,1462,1487]" pageId="1" pageNumber="188" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, the anterior part of the head of the new genus differs from the Afrotropical members of this family (
|
||
<emphasis id="B92FEAC6FFEBFF81ABBBFA53FEF61FFF" box="[160,333,1499,1523]" italics="true" pageId="1" pageNumber="188">Flabelloselasia</emphasis>
|
||
<bibRefCitation id="EFCA4B25FFEBFF81AA43FA52FD3A1FFF" author="Kundrata, R. & Bocak, L." box="[344,641,1498,1523]" pageId="1" pageNumber="188" pagination="441 - 492" refId="ref4148" refString="Kundrata, R. & Bocak, L. (2017) Taxonomic review of Drilini (Elateridae: Agrypninae) in Cameroon reveals high morphological diversity, including the discovery of five new genera. Insect systematics & evolution, 48 (5), 441 - 492. https: // doi. org / 10.1163 / 1876312 X- 48022161" type="journal article" year="2017">Kundrata et Bocak, 2017</bibRefCitation>
|
||
;
|
||
<emphasis id="B92FEAC6FFEBFF81A98BFA53FCA21FFF" box="[656,793,1499,1523]" italics="true" pageId="1" pageNumber="188">Kupeselasia</emphasis>
|
||
<bibRefCitation id="EFCA4B25FFEBFF81A83FFA52FBF01FFF" author="Kundrata, R. & Bocak, L." box="[804,1099,1498,1523]" pageId="1" pageNumber="188" pagination="441 - 492" refId="ref4148" refString="Kundrata, R. & Bocak, L. (2017) Taxonomic review of Drilini (Elateridae: Agrypninae) in Cameroon reveals high morphological diversity, including the discovery of five new genera. Insect systematics & evolution, 48 (5), 441 - 492. https: // doi. org / 10.1163 / 1876312 X- 48022161" type="journal article" year="2017">Kundrata et Bocak, 2017</bibRefCitation>
|
||
;
|
||
<emphasis id="B92FEAC6FFEBFF81AF40FA53FB141FFF" box="[1115,1199,1499,1523]" italics="true" pageId="1" pageNumber="188">Lolosia</emphasis>
|
||
<bibRefCitation id="EFCA4B25FFEBFF81AFA0FA52FF681C1B" author="Kundrata, R. & Bocak, L." pageId="1" pageNumber="188" pagination="441 - 492" refId="ref4148" refString="Kundrata, R. & Bocak, L. (2017) Taxonomic review of Drilini (Elateridae: Agrypninae) in Cameroon reveals high morphological diversity, including the discovery of five new genera. Insect systematics & evolution, 48 (5), 441 - 492. https: // doi. org / 10.1163 / 1876312 X- 48022161" type="journal article" year="2017">Kundrata et Bocak, 2017</bibRefCitation>
|
||
;
|
||
<emphasis id="B92FEAC6FFEBFF81ABC4FA76FECB1C1A" box="[223,368,1534,1558]" italics="true" pageId="1" pageNumber="188">Microselasia</emphasis>
|
||
<bibRefCitation id="EFCA4B25FFEBFF81AA62FA76FD2D1C1B" author="Kundrata, R. & Bocak, L." box="[377,662,1534,1559]" pageId="1" pageNumber="188" pagination="441 - 492" refId="ref4148" refString="Kundrata, R. & Bocak, L. (2017) Taxonomic review of Drilini (Elateridae: Agrypninae) in Cameroon reveals high morphological diversity, including the discovery of five new genera. Insect systematics & evolution, 48 (5), 441 - 492. https: // doi. org / 10.1163 / 1876312 X- 48022161" type="journal article" year="2017">Kundrata et Bocak, 2017</bibRefCitation>
|
||
and
|
||
<emphasis id="B92FEAC6FFEBFF81A9CBFA76FCCD1C1A" box="[720,886,1534,1558]" italics="true" pageId="1" pageNumber="188">Wittmerselasia</emphasis>
|
||
<bibRefCitation id="EFCA4B25FFEBFF81A864FA76FB241C1B" author="Kundrata, R. & Bocak, L." box="[895,1183,1534,1559]" pageId="1" pageNumber="188" pagination="441 - 492" refId="ref4148" refString="Kundrata, R. & Bocak, L. (2017) Taxonomic review of Drilini (Elateridae: Agrypninae) in Cameroon reveals high morphological diversity, including the discovery of five new genera. Insect systematics & evolution, 48 (5), 441 - 492. https: // doi. org / 10.1163 / 1876312 X- 48022161" type="journal article" year="2017">Kundrata et Bocak, 2017</bibRefCitation>
|
||
, known to the authors after
|
||
<bibRefCitation id="EFCA4B25FFEBFF81ABC1F9AAFDAB1C37" author="Kundrata, R. & Bocak, L." box="[218,528,1570,1595]" pageId="1" pageNumber="188" pagination="441 - 492" refId="ref4148" refString="Kundrata, R. & Bocak, L. (2017) Taxonomic review of Drilini (Elateridae: Agrypninae) in Cameroon reveals high morphological diversity, including the discovery of five new genera. Insect systematics & evolution, 48 (5), 441 - 492. https: // doi. org / 10.1163 / 1876312 X- 48022161" type="journal article" year="2017">Kundrata & Bocak, 2017</bibRefCitation>
|
||
) in the shortened elytra, and also from
|
||
<emphasis id="B92FEAC6FFEBFF81AF14F9ABFAD81C37" box="[1039,1379,1571,1595]" italics="true" pageId="1" pageNumber="188">Flabelloselasia, Microselasia</emphasis>
|
||
and
|
||
<emphasis id="B92FEAC6FFEBFF81AB8CF9CEFE861C52" box="[151,317,1606,1630]" italics="true" pageId="1" pageNumber="188">Wittmerselasia</emphasis>
|
||
in the pectinate antennae (not flabellate), and from
|
||
<emphasis id="B92FEAC6FFEBFF81A860F9CEFBBF1C52" box="[891,1028,1606,1630]" italics="true" pageId="1" pageNumber="188">Kupeselasia</emphasis>
|
||
and
|
||
<emphasis id="B92FEAC6FFEBFF81AF22F9CEFB361C52" box="[1081,1165,1606,1630]" italics="true" pageId="1" pageNumber="188">Lolosia</emphasis>
|
||
in the much larger body and pectinate antennae (not serrate).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C341655FFFEBFF82ABDCF906FDE31F01" lastPageId="2" lastPageNumber="189" pageId="1" pageNumber="188" type="description">
|
||
<paragraph id="8BE436D4FFEBFF82ABDCF906FE471B10" blockId="1.[151,1437,634,2027]" lastBlockId="2.[151,1437,151,1293]" lastPageId="2" lastPageNumber="189" pageId="1" pageNumber="188">
|
||
<emphasis id="B92FEAC6FFEBFF81ABDCF906FEE21CAB" bold="true" box="[199,345,1678,1703]" pageId="1" pageNumber="188">Description.</emphasis>
|
||
Head slightly hypognathous, slightly wider than anterior margin of pronotum, dorsally convex, frons between antennal insertions strongly protruding forwards and forming long subtriangular projection, antennal insertions located at base of sides of projection, mouthparts completely concealed observing head from above; upper plane of this projection nearly horizontal and flattened, lateral planes strongly converging toward cariniform emarginate anterior edge, frontal plane strongly inclined ventroposteriorly toward emarginate anterior margin of epicranium (
|
||
<figureCitation id="13602A51FFEBFF81AA33F8CAFEC81D57" box="[296,371,1858,1883]" captionStart="FIGURE 1" captionStartId="3.[151,250,2000,2022]" captionTargetBox="[264,1323,233,1957]" captionTargetId="figure@3.[151,1436,218,1957]" captionTargetPageId="3" captionText="FIGURE 1. Holotype of Drilorhinus klimenkoi gen. et sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2628558/files/figure.png" pageId="1" pageNumber="188">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="13602A51FFEBFF81AA9EF8CBFE0E1D57" box="[389,437,1859,1883]" captionStart="FIGURES 2–8" captionStartId="4.[151,264,1937,1960]" captionTargetBox="[156,1425,717,1889]" captionTargetId="figure@4.[151,1436,717,1911]" captionTargetPageId="4" captionText="FIGURES 2–8. Drilorhinus klimenkoi gen. et sp. nov., holotype, details. 2: head, frontal. 3: antenna; 4: maxillary palpus; 5: labial palpus; 6: prosternum; 7: head and prothorax, lateral; 8: scutellum. Scale bars = 1 mm: a, for Figs. 2, 3, 6–8; b, for Figs. 4, 5." httpUri="https://zenodo.org/record/2628560/files/figure.png" pageId="1" pageNumber="188">2, 7</figureCitation>
|
||
). Eyes medium-sized, hemispherically prominent and rather coarsely facetted; frontal distance between eyes 1.85 times as great as eye diameter. Labrum small and slightly transverse, sclerotized, free and completely exposed, with anterior margin slightly concave (
|
||
<figureCitation id="13602A51FFEBFF81A8B7F802FC411DAF" box="[940,1018,1930,1955]" captionStart="FIGURES 2–8" captionStartId="4.[151,264,1937,1960]" captionTargetBox="[156,1425,717,1889]" captionTargetId="figure@4.[151,1436,717,1911]" captionTargetPageId="4" captionText="FIGURES 2–8. Drilorhinus klimenkoi gen. et sp. nov., holotype, details. 2: head, frontal. 3: antenna; 4: maxillary palpus; 5: labial palpus; 6: prosternum; 7: head and prothorax, lateral; 8: scutellum. Scale bars = 1 mm: a, for Figs. 2, 3, 6–8; b, for Figs. 4, 5." httpUri="https://zenodo.org/record/2628560/files/figure.png" pageId="1" pageNumber="188">Fig. 2</figureCitation>
|
||
). Mandibles moderately separated, comparatively short and robust, moderately curved, with short shining apical region and sharp subapical tooth, apex acute, outer margin angularly curved (
|
||
<figureCitation id="13602A51FFEBFF81A995F85AFD6E1DE7" box="[654,725,2002,2027]" captionStart="FIGURES 2–8" captionStartId="4.[151,264,1937,1960]" captionTargetBox="[156,1425,717,1889]" captionTargetId="figure@4.[151,1436,717,1911]" captionTargetPageId="4" captionText="FIGURES 2–8. Drilorhinus klimenkoi gen. et sp. nov., holotype, details. 2: head, frontal. 3: antenna; 4: maxillary palpus; 5: labial palpus; 6: prosternum; 7: head and prothorax, lateral; 8: scutellum. Scale bars = 1 mm: a, for Figs. 2, 3, 6–8; b, for Figs. 4, 5." httpUri="https://zenodo.org/record/2628560/files/figure.png" pageId="1" pageNumber="188">Fig. 2</figureCitation>
|
||
). Maxillary palpi short and slender, tetramerous, palpomere 1 short and wide; palpomere 2 about 1.5 times as long as wide; palpomere 3 short, about as long as wide; apical palpomere elongate, about twice as long as palpomere 3, obliquely truncate at apex (
|
||
<figureCitation id="13602A51FFE8FF82AF41FF34FB1A1AD8" box="[1114,1185,188,213]" captionStart="FIGURES 2–8" captionStartId="4.[151,264,1937,1960]" captionTargetBox="[156,1425,717,1889]" captionTargetId="figure@4.[151,1436,717,1911]" captionTargetPageId="4" captionText="FIGURES 2–8. Drilorhinus klimenkoi gen. et sp. nov., holotype, details. 2: head, frontal. 3: antenna; 4: maxillary palpus; 5: labial palpus; 6: prosternum; 7: head and prothorax, lateral; 8: scutellum. Scale bars = 1 mm: a, for Figs. 2, 3, 6–8; b, for Figs. 4, 5." httpUri="https://zenodo.org/record/2628560/files/figure.png" pageId="2" pageNumber="189">Fig. 4</figureCitation>
|
||
). Labium short; labial palpi trimerous, with apical palpomere subacute (
|
||
<figureCitation id="13602A51FFE8FF82A9EAFF57FC861AF4" box="[753,829,223,248]" captionStart="FIGURES 2–8" captionStartId="4.[151,264,1937,1960]" captionTargetBox="[156,1425,717,1889]" captionTargetId="figure@4.[151,1436,717,1911]" captionTargetPageId="4" captionText="FIGURES 2–8. Drilorhinus klimenkoi gen. et sp. nov., holotype, details. 2: head, frontal. 3: antenna; 4: maxillary palpus; 5: labial palpus; 6: prosternum; 7: head and prothorax, lateral; 8: scutellum. Scale bars = 1 mm: a, for Figs. 2, 3, 6–8; b, for Figs. 4, 5." httpUri="https://zenodo.org/record/2628560/files/figure.png" pageId="2" pageNumber="189">Fig. 5</figureCitation>
|
||
). Antennae 11-segmented, pectinate, covered with moderately dense setae (
|
||
<figureCitation id="13602A51FFE8FF82AAB1FE8CFE551B10" box="[426,494,260,285]" captionStart="FIGURES 2–8" captionStartId="4.[151,264,1937,1960]" captionTargetBox="[156,1425,717,1889]" captionTargetId="figure@4.[151,1436,717,1911]" captionTargetPageId="4" captionText="FIGURES 2–8. Drilorhinus klimenkoi gen. et sp. nov., holotype, details. 2: head, frontal. 3: antenna; 4: maxillary palpus; 5: labial palpus; 6: prosternum; 7: head and prothorax, lateral; 8: scutellum. Scale bars = 1 mm: a, for Figs. 2, 3, 6–8; b, for Figs. 4, 5." httpUri="https://zenodo.org/record/2628560/files/figure.png" pageId="2" pageNumber="189">Fig. 3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE436D4FFE8FF82ABDCFEA0FF491814" blockId="2.[151,1437,151,1293]" pageId="2" pageNumber="189">
|
||
Pronotum slightly transverse, subtrapezoidal, widest at base; anterior margin slightly concave medially; sides subrectilinear: slightly diverging posteriorly in anterior half and scarcely convex before posterior angles; posterior margin slightly bisinuate and nearly straight medially. Anterior angles obtuse; posterior angles subacute and narrowly rounded at apex. Lateral carina separating pronotum from prohypomeron distinct in posterior half and obliterated anteriorly (
|
||
<figureCitation id="13602A51FFE8FF82AA88FE3FFE631BDC" box="[403,472,439,464]" captionStart="FIGURES 2–8" captionStartId="4.[151,264,1937,1960]" captionTargetBox="[156,1425,717,1889]" captionTargetId="figure@4.[151,1436,717,1911]" captionTargetPageId="4" captionText="FIGURES 2–8. Drilorhinus klimenkoi gen. et sp. nov., holotype, details. 2: head, frontal. 3: antenna; 4: maxillary palpus; 5: labial palpus; 6: prosternum; 7: head and prothorax, lateral; 8: scutellum. Scale bars = 1 mm: a, for Figs. 2, 3, 6–8; b, for Figs. 4, 5." httpUri="https://zenodo.org/record/2628560/files/figure.png" pageId="2" pageNumber="189">Fig. 7</figureCitation>
|
||
). Prohypomeron carinate sublaterally; prosternal suture short. Prosternum transverse; chin-piece short, with anterior margin widely emarginate; prosternal process very short and widely rounded at apex (
|
||
<figureCitation id="13602A51FFE8FF82AB84FE77FF5F1814" box="[159,228,511,536]" captionStart="FIGURES 2–8" captionStartId="4.[151,264,1937,1960]" captionTargetBox="[156,1425,717,1889]" captionTargetId="figure@4.[151,1436,717,1911]" captionTargetPageId="4" captionText="FIGURES 2–8. Drilorhinus klimenkoi gen. et sp. nov., holotype, details. 2: head, frontal. 3: antenna; 4: maxillary palpus; 5: labial palpus; 6: prosternum; 7: head and prothorax, lateral; 8: scutellum. Scale bars = 1 mm: a, for Figs. 2, 3, 6–8; b, for Figs. 4, 5." httpUri="https://zenodo.org/record/2628560/files/figure.png" pageId="2" pageNumber="189">Fig. 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE436D4FFE8FF82ABDCFDACFC2918C1" blockId="2.[151,1437,151,1293]" pageId="2" pageNumber="189">Scutellum elongate, flattened, subtriangular, with concave sides and blunt apex. Elytra elongate, each gradually narrowing posteriorly, reaching only the base of abdominal segment 6, divergent, adjacent to each other only at its basal third, elytral apices separately rounded; elytral striae indistinct; each elytron somewhat raised along suture, with weakly expressed arcuate costa running from humeri towards apices and short oblique weak keel in scutellar region. Integument uniformly rugose-granulate and dull.</paragraph>
|
||
<paragraph id="8BE436D4FFE8FF82ABDCFD5FFEF91934" blockId="2.[151,1437,151,1293]" pageId="2" pageNumber="189">Mesoventrite widely V-shaped, with frontal margin widely and deeply emarginate; mesoventral cavity shallow, with weakly defined walls. Mesocoxal cavity open to both mesepimeron and mesepisternum. Metaventrite large, subtrapezoidal.</paragraph>
|
||
<paragraph id="8BE436D4FFE8FF82ABDCFCCDFDEF19E1" blockId="2.[151,1437,151,1293]" pageId="2" pageNumber="189">Legs slightly narrow and compressed; coxae robust; trochanters slender, elongate, slightly widened apically, obliquely attached to femora. Tarsi moderately long; tarsomeres 1 and 2 subequal in length; tarsomere 3 slightly shorter than preceding ones, about 1.5 times longer than tarsomere 4; tarsomere 4 shortest, extended ventrally; apical tarsomere slender and elongate, about 2.5 times as long as tarsomere 4; claws simple, slender, slightly curved, each with one long seta at base.</paragraph>
|
||
<paragraph id="8BE436D4FFE8FF82ABDCFC7FFB211E5B" blockId="2.[151,1437,151,1293]" pageId="2" pageNumber="189">Abdomen soft and slender; penultimate ventrite subtruncate at apex. Tergite 9 shallowly concave at apex; tergite 10 subquadrangular and widely rounded at apex. Sternite 9 about 1.8 times as long as wide, slightly notched at base and with widely rounded apical margin, its apex finely punctate and sparsely setose.</paragraph>
|
||
<paragraph id="8BE436D4FFE8FF82ABDCFBECFAE11EE4" blockId="2.[151,1437,151,1293]" pageId="2" pageNumber="189">
|
||
Male genitalia trilobate (
|
||
<figureCitation id="13602A51FFE8FF82AAF3FBECFDDB1E70" box="[488,608,1124,1149]" captionStart="FIGURES 9–11" captionStartId="6.[151,264,1221,1244]" captionTargetBox="[222,1393,210,1193]" captionTargetId="figure@6.[151,1436,193,1200]" captionTargetPageId="6" captionText="FIGURES 9–11. Drilorhinus klimenkoi gen. et sp. nov., holotype, details. 9, 10: aedeagus dorsal; 11: subapical hook of penis trunk, lateral. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/2628562/files/figure.png" pageId="2" pageNumber="189">Figs 9–11</figureCitation>
|
||
); phallobase robust, slightly longer than wide, basally narrowed; penis (median lobe) robust, longer than parameres and slightly longer than phallobase, moderately curved in lateral view, basally with pair of short divergent apophyses, dorsally with long robust subapical hook (
|
||
<figureCitation id="13602A51FFE8FF82AFA2FB24FAB41EC8" box="[1209,1295,1196,1221]" captionStart="FIGURES 9–11" captionStartId="6.[151,264,1221,1244]" captionTargetBox="[222,1393,210,1193]" captionTargetId="figure@6.[151,1436,193,1200]" captionTargetPageId="6" captionText="FIGURES 9–11. Drilorhinus klimenkoi gen. et sp. nov., holotype, details. 9, 10: aedeagus dorsal; 11: subapical hook of penis trunk, lateral. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/2628562/files/figure.png" pageId="2" pageNumber="189">Fig. 11</figureCitation>
|
||
); parameres robust, elongate and narrowing apically, about twice as long as wide at base, with rounded and setose apices.
|
||
</paragraph>
|
||
<paragraph id="8BE436D4FFE8FF82AB8CFB7CFDE31F01" blockId="2.[151,1437,151,1293]" box="[151,600,1268,1293]" pageId="2" pageNumber="189">Females and immature stages unknown.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |