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<document id="A04CFDB99E923335CA7DBBD3C24BC196" ID-CLB-Dataset="284910" ID-DOI="10.5852/ejt.2023.908.2337" ID-GBIF-Dataset="6156c195-5953-4af2-9999-a1a03fd0f87e" ID-ISSN="2118-9773" ID-Zenodo-Dep="10149298" ID-ZooBank="1728B5B0-A130-4370-9610-9DA9CEDF4717" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1700226465688" checkinUser="plazi" docAuthor="Antić, Dragan, Šević, Mirko, Barjadze, Shalva &amp; Makarov, Slobodan" docDate="2023" docId="03E187FCFFC36974FDB10E05FDB8F9EB" docLanguage="en" docName="EJT.2023.908.39-76.pdf" docOrigin="European Journal of Taxonomy 908" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2337/10171" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Acanthophorella devi Antic 2023, sp. nov." docType="treatment" docUuid="DDAE0A65-7AB2-42AF-914C-1CE779454840" docUuidSource="ZooBank" docVersion="6" lastPageNumber="56" masterDocId="FFD8FF84FFC96965FFF20A17FFD4FFE7" masterDocTitle="Review of the genus Acanthophorella Antić &amp; Makarov, 2016 (Diplopoda, Chordeumatida, Anthroleucosomatidae), with descriptions of three new species from the Caucasus" masterLastPageNumber="76" masterPageNumber="39" pageNumber="49" updateTime="1705069979865" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="B8E1312EAE9652CA0585AFE49FFC2116">Review of the genus Acanthophorella Antić &amp; Makarov, 2016 (Diplopoda, Chordeumatida, Anthroleucosomatidae), with descriptions of three new species from the Caucasus</mods:title>
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<mods:namePart id="63C92E0768A6C22F7F2AD6EE6A853683">Antić, Dragan</mods:namePart>
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<mods:affiliation id="775B8597B14A48374A930CE4F685159F">University of Belgrade – Faculty of Biology, Institute of Zoology, Studentski Trg 16, 11000 Belgrade, Serbia. &amp; Serbian Biospeleological Society, Trg Dositeja Obradovića 2, 21000 Novi Sad, Serbia.</mods:affiliation>
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<mods:namePart id="2BE4507005601CEA03B076E4FE2711AA">Šević, Mirko</mods:namePart>
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<mods:affiliation id="BDF46D7B8988159D7C070067F1E7A12B">University of Belgrade – Faculty of Biology, Institute of Zoology, Studentski Trg 16, 11000 Belgrade, Serbia. &amp; Serbian Biospeleological Society, Trg Dositeja Obradovića 2, 21000 Novi Sad, Serbia.</mods:affiliation>
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<mods:namePart id="7ED0E6A9B043F175A4693DDD75287646">Barjadze, Shalva</mods:namePart>
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<mods:affiliation id="11E49FEBB8CB1D3C1D0B564307FFB86B">Institute of Zoology, Ilia State University, Giorgi Tsereteli 3, 0162, Tbilisi, Georgia.</mods:affiliation>
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<mods:affiliation id="2A095DC688E26C5CFE07BC8036B4573E">University of Belgrade – Faculty of Biology, Institute of Zoology, Studentski Trg 16, 11000 Belgrade, Serbia. &amp; Serbian Biospeleological Society, Trg Dositeja Obradovića 2, 21000 Novi Sad, Serbia.</mods:affiliation>
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<taxonomicName id="4C484D69FFC3696FFDB10E05FC5BFBCB" ID-CoL="BW7HW" authority="Antic" authorityName="Antic" authorityYear="2023" box="[579,911,1042,1069]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="10" pageNumber="49" phylum="Arthropoda" rank="species" species="devi" status="sp. nov.">
<emphasis id="B93CEAF8FFC3696FFDB10E05FC93FBCA" bold="true" box="[579,839,1042,1069]" italics="true" pageId="10" pageNumber="49">Acanthophorella devi</emphasis>
Antić
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<paragraph id="8BF736EAFFC3696FFD710E4FFC64FB94" blockId="10.[362,1225,1042,1139]" box="[643,944,1112,1139]" pageId="10" pageNumber="49">
<figureCitation id="13732A6FFFC3696FFD710E4FFD29FB94" box="[643,765,1112,1139]" captionStart="Fig" captionStartId="1.[189,232,1857,1883]" captionTargetBox="[189,1398,420,1810]" captionTargetId="figure-57@1.[189,1398,420,1810]" captionTargetPageId="1" captionText="Fig. 1. Troglobiotic members of the genus Acanthophorella Antić &amp; Makarov, 2016 in situ. A–B. Acanthophorella barjadzei Antić &amp; Makarov, 2016, ♂ from Nikortsminda Sakinule Cave. C–D. Acanthophorella barjadzei Antić &amp; Makarov, 2016, ♂ from Muradi Cave. E–F. Acanthophorella devi Antić sp. nov., types, ♂♂. G–H. Acanthophorella valerii Antić sp. nov., holotype, ♂ (NHMW MY10366). Photos by Dragan Antić." figureDoi="http://doi.org/10.5281/zenodo.10149300" httpUri="https://zenodo.org/record/10149300/files/figure.png" pageId="10" pageNumber="49">Figs 1E–F</figureCitation>
, 
<figureCitation id="13732A6FFFC3696FFCFB0E4FFC91FB95" box="[777,837,1112,1138]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStartId-0="12.[189,232,1879,1905]" captionStartId-1="13.[189,232,1770,1796]" captionStartId-2="14.[189,232,1892,1918]" captionStartId-3="15.[189,232,1787,1813]" captionStartId-4="16.[189,232,1787,1813]" captionTargetBox-0="[189,1398,270,1829]" captionTargetBox-1="[189,1398,265,1700]" captionTargetBox-2="[229,1359,265,1842]" captionTargetBox-3="[282,1305,265,1742]" captionTargetBox-4="[293,1287,282,1718]" captionTargetId-0="figure-16@12.[189,1398,270,1829]" captionTargetId-1="figure-11@13.[189,1398,265,1700]" captionTargetId-2="figure-17@14.[229,1359,265,1842]" captionTargetId-3="figure-12@15.[282,1305,265,1742]" captionTargetId-4="figure-17@16.[287,1300,265,1743]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionTargetPageId-2="14" captionTargetPageId-3="15" captionTargetPageId-4="16" captionText-0="Fig. 6. Acanthophorella devi Antić sp. nov. A–B. Paratype, ♀ (IZB). C–E. Holotype, ♂ (NHMW MY10364). A. Habitus, lateral view. B. Head and collum, lateral view. C. Habitus, lateral view. D. Midbody rings, dorsal view. E. Midbody rings, lateral view. Scale bars: A, C–E = 1 mm; B = 0.5 mm." captionText-1="Fig. 7. Acanthophorella devi Antić sp. nov., paratype, ♂ (IZB). A. Head, anterior view. B. Labrum, anterior view. C. Right side of head showing field with ommatidia, organ of Tömösvary and group of papilliform outgrowths, lateral view. D. Right field with ommatidia, anterior view. E. Ring 16, dorsal view.F. Ring 16, dorsolateral view. G. Rings 14–16, posterior view. H. Left interior macrochaeta of ring 15, lateral view. Scale bars: A–B, E–G = 0.1 mm; C, H = 0.05 mm; D = 0.01 mm." captionText-2="Fig. 8. Acanthophorella devi Antić sp. nov. A. Paratype, ♀ (IZB), B–G. Paratype, ♂ (IZB). A. Legpair 2, anterior view. B. Leg-pair 2, anterior view. C. Right leg 4, anterior view, white arrows indicate distoventral pads. D. Leg-pair 7, posterior view. E. Leg-pair 10, anterior view. F. Anterior gonopods, anterior view. G. Posterior gonopods, posterior view. Scale bars = 0.2 mm." captionText-3="Fig. 9. Acanthophorella devi Antić sp. nov. A–F. Paratype, ♂ (IZB). G–I. Paratype, ♀ (IZB). A. Anterior gonopods, anterior view. B. Anterior gonopods, anterodistal view. C. Anterior gonopods, distal view. D. Anterior gonopods, lateral view. E. Anterior gonopods, lateral lamella and tuft, posterodistal view. F. Anterior gonopods, posterodistal view. G. Left vulva, lateral view. H. Vulvae, distal view. I. Left vulva, anterior view.Abbreviations: a = angiocoxite; ap = anterior process; b = bursa; ll = lateral lamella; mp = medial part; o = operculum; pl = posterior lobe; s = sternum; sa = synangiocoxal base; sl = sternal lamella; tf = angiocoxal tuft. Scale bars: A–D, F–I = 0.1 mm; E = 0.02 mm." captionText-4="Fig. 10. Acanthophorella devi Antić sp. nov. A–E. Paratype, ♂ (IZB). F–G. Paratype, ♀ (IZB). A. Anterior gonopods, anterior view. B. Anterior gonopods, posterior view. C. Anterior gonopods, lateral view. D. Posterior gonopods, posterior view. E. Left posterior gonopod, lateral view. F. Right vulva, distal view. G. Right vulva, lateral view. Abbreviations: a = angiocoxite; ap = anterior process; b = bursa; c = colpocoxite; ll = lateral lamella; mp = medial part; o = operculum; pl = posterior lobe; s = sternum; sa = synangiocoxal base; sl = sternal lamella; t = telopodite; tf = angiocoxal tuft. Drawings by DA (A–E) and MŠ (F–G). Scale bars = 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.10149312" figureDoi-1="http://doi.org/10.5281/zenodo.10149314" figureDoi-2="http://doi.org/10.5281/zenodo.10149316" figureDoi-3="http://doi.org/10.5281/zenodo.10149318" figureDoi-4="http://doi.org/10.5281/zenodo.10149320" httpUri-0="https://zenodo.org/record/10149312/files/figure.png" httpUri-1="https://zenodo.org/record/10149314/files/figure.png" httpUri-2="https://zenodo.org/record/10149316/files/figure.png" httpUri-3="https://zenodo.org/record/10149318/files/figure.png" httpUri-4="https://zenodo.org/record/10149320/files/figure.png" pageId="10" pageNumber="49">6–10</figureCitation>
, 
<figureCitation id="13732A6FFFC3696FFCA00E4EFC53FB94" box="[850,903,1113,1139]" captionStart="Fig" captionStartId="33.[189,232,1822,1848]" captionTargetBox="[202,1378,705,1770]" captionTargetId="figure-170@33.[189,1398,686,1780]" captionTargetPageId="33" captionText="Fig. 21. Anterior gonopods in the genera Acanthophorella Antić &amp; Makarov, 2016 and PseudoflagellophorellaAntić &amp; Makarov, 2016, anterior views. A. A. auritaAntić sp. nov. B. A. barjadzei Antić &amp; Makarov, 2016. C. A. chegemi Antić &amp; Makarov, 2016. D. A. devi Antić sp. nov. E. A. irystoni Antić &amp; Makarov, 2016. F. A. valerii Antić sp. nov. G. P. eskovi Antić &amp; Makarov, 2016. H. P. mirabilis Antić &amp; Makarov, 2016. I. P. papilioformis Antić &amp; Makarov, 2016. B–C, E, G–I after Antić &amp; Makarov (2016). Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10149345" httpUri="https://zenodo.org/record/10149345/files/figure.png" pageId="10" pageNumber="49">21D</figureCitation>
, 
<figureCitation id="13732A6FFFC3696FFC610E4EFC64FB94" box="[915,944,1113,1139]" captionStart="Fig" captionStartId="35.[189,232,1998,2024]" captionTargetBox="[189,1398,1153,1959]" captionTargetId="figure-355@35.[189,1398,1150,1959]" captionTargetPageId="35" captionText="Fig. 22. Distribution of the genus Acanthophorella Antić &amp; Makarov, 2016." figureDoi="http://doi.org/10.5281/zenodo.10149347" httpUri="https://zenodo.org/record/10149347/files/figure.png" pageId="10" pageNumber="49">22</figureCitation>
</paragraph>
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<heading id="D0BF8186FFC3696FFF4F0E8DFEE2FB53" bold="true" box="[189,310,1178,1204]" fontSize="11" level="3" pageId="10" pageNumber="49" reason="3">
<emphasis id="B93CEAF8FFC3696FFF4F0E8DFEE2FB53" bold="true" box="[189,310,1178,1204]" pageId="10" pageNumber="49">Diagnosis</emphasis>
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Distinguished from congeners and members of the genus 
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<emphasis id="B93CEAF8FFC3696FFC670EDFFB72FB05" box="[917,1190,1224,1250]" italics="true" pageId="10" pageNumber="49">Pseudoflagellophorella</emphasis>
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by the presence of a poorly developed or the complete absence of a protrusions on coxae 
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males (
<figureCitation id="13732A6FFFC3696FFB1C0EFBFA98FAE1" box="[1262,1356,1260,1286]" captionStart="Fig" captionStartId="14.[189,232,1892,1918]" captionTargetBox="[229,1359,265,1842]" captionTargetId="figure-17@14.[229,1359,265,1842]" captionTargetPageId="14" captionText="Fig. 8. Acanthophorella devi Antić sp. nov. A. Paratype, ♀ (IZB), B–G. Paratype, ♂ (IZB). A. Legpair 2, anterior view. B. Leg-pair 2, anterior view. C. Right leg 4, anterior view, white arrows indicate distoventral pads. D. Leg-pair 7, posterior view. E. Leg-pair 10, anterior view. F. Anterior gonopods, anterior view. G. Posterior gonopods, posterior view. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10149316" httpUri="https://zenodo.org/record/10149316/files/figure.png" pageId="10" pageNumber="49">Fig. 8E</figureCitation>
, vs presence of well-developed subtriangular or rounded protrusion or even a long process in congeners or members of 
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<emphasis id="B93CEAF8FFC3696FFEA10F25FDB6FAAB" box="[339,610,1330,1356]" italics="true" pageId="10" pageNumber="49">Pseudoflagellophorella</emphasis>
</taxonomicName>
) and, aside from 
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<emphasis id="B93CEAF8FFC3696FFCC50F24FC1CFAAB" box="[823,968,1330,1357]" italics="true" pageId="10" pageNumber="49">A. barjadzei</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C484D69FFC3696FFBF10F24FBA6FAAB" authority="Antić &amp; Šević &amp; Barjadze &amp; Makarov, 2023" authorityName="Antić &amp; Šević &amp; Barjadze &amp; Makarov" authorityYear="2023" box="[1027,1138,1330,1357]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="10" pageNumber="49" phylum="Arthropoda" rank="species" species="valerii" status="sp. nov.">
<emphasis id="B93CEAF8FFC3696FFBF10F24FBA6FAAB" box="[1027,1138,1330,1357]" italics="true" pageId="10" pageNumber="49">A. valerii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20F5783FFC3696FFB8B0F24FB03FAAA" box="[1145,1239,1331,1357]" pageId="10" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
, by a whitish and unpigmented body (vs pigmented body in others). From 
<taxonomicName id="4C484D69FFC3696FFC710F41FC25FA88" authorityName="Antić &amp; Šević &amp; Barjadze &amp; Makarov" authorityYear="2023" box="[899,1009,1365,1392]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="10" pageNumber="49" phylum="Arthropoda" rank="species" species="valerii">
<emphasis id="B93CEAF8FFC3696FFC710F41FC25FA88" box="[899,1009,1365,1392]" italics="true" pageId="10" pageNumber="49">A. valerii</emphasis>
</taxonomicName>
, it differs by the presence of pale brownish ommatidia (vs black in 
<taxonomicName id="4C484D69FFC3696FFDB70F6EFD66FA75" authorityName="Antić &amp; Šević &amp; Barjadze &amp; Makarov" authorityYear="2023" box="[581,690,1400,1427]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="10" pageNumber="49" phylum="Arthropoda" rank="species" species="valerii">
<emphasis id="B93CEAF8FFC3696FFDB70F6EFD66FA75" box="[581,690,1400,1427]" italics="true" pageId="10" pageNumber="49">A. valerii</emphasis>
</taxonomicName>
), the absence of a mesal tooth on the coxal processes of leg-pair 7 (vs presence in 
<taxonomicName id="4C484D69FFC3696FFE300F8BFDFBFA51" authorityName="Antić &amp; Šević &amp; Barjadze &amp; Makarov" authorityYear="2023" box="[450,559,1436,1462]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="10" pageNumber="49" phylum="Arthropoda" rank="species" species="valerii">
<emphasis id="B93CEAF8FFC3696FFE300F8BFDFBFA51" box="[450,559,1436,1462]" italics="true" pageId="10" pageNumber="49">A. valerii</emphasis>
</taxonomicName>
) and in the general shape of the anterior and posterior gonopods. From the most similar 
<taxonomicName id="4C484D69FFC3696FFE7A0FA8FDCDFA3E" authorityName="Antic &amp; Makarov" authorityYear="2016" box="[392,537,1471,1497]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="10" pageNumber="49" phylum="Arthropoda" rank="species" species="barjadzei">
<emphasis id="B93CEAF8FFC3696FFE7A0FA8FDCDFA3E" box="[392,537,1471,1497]" italics="true" pageId="10" pageNumber="49">A. barjadzei</emphasis>
</taxonomicName>
(in terms of anterior and posterior gonopods), 
<taxonomicName id="4C484D69FFC3696FFBBE0FA8FB4BFA3E" authority="Antić &amp; Šević &amp; Barjadze &amp; Makarov, 2023" authorityName="Antić &amp; Šević &amp; Barjadze &amp; Makarov" authorityYear="2023" box="[1100,1183,1471,1497]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="10" pageNumber="49" phylum="Arthropoda" rank="species" species="devi" status="sp. nov.">
<emphasis id="B93CEAF8FFC3696FFBBE0FA8FB4BFA3E" box="[1100,1183,1471,1497]" italics="true" pageId="10" pageNumber="49">A. devi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20F5783FFC3696FFB5A0FA8FAD0FA3E" box="[1192,1284,1471,1497]" pageId="10" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
differs in the absence of a mesal tooth on the coxal processes of leg-pair 7 (vs presence in 
<taxonomicName id="4C484D69FFC3696FFB890FF4FADEFA1B" authorityName="Antic &amp; Makarov" authorityYear="2016" box="[1147,1290,1506,1533]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="10" pageNumber="49" phylum="Arthropoda" rank="species" species="barjadzei">
<emphasis id="B93CEAF8FFC3696FFB890FF4FADEFA1B" box="[1147,1290,1506,1533]" italics="true" pageId="10" pageNumber="49">A. barjadzei</emphasis>
</taxonomicName>
), as well as in the general appearance: 
<taxonomicName id="4C484D69FFC3696FFDE80C11FDB8F9F8" authorityName="Antić &amp; Šević &amp; Barjadze &amp; Makarov" authorityYear="2023" box="[538,620,1541,1568]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="10" pageNumber="49" phylum="Arthropoda" rank="species" species="devi">
<emphasis id="B93CEAF8FFC3696FFDE80C11FDB8F9F8" box="[538,620,1541,1568]" italics="true" pageId="10" pageNumber="49">A. devi</emphasis>
</taxonomicName>
is smaller, 
<quantity id="4CB09B0FFFC3696FFD010C12FCA5F9C7" box="[755,881,1541,1568]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="10" pageNumber="49" unit="mm" value="12.0" valueMax="13.0" valueMin="11.0">11–13 mm</quantity>
long (vs 
<quantity id="4CB09B0FFFC3696FFC120C12FB8BF9C7" box="[992,1119,1541,1568]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" metricValueMax="2.6" metricValueMin="1.6" pageId="10" pageNumber="49" unit="mm" value="21.0" valueMax="26.0" valueMin="16.0">16–26 mm</quantity>
in 
<taxonomicName id="4C484D69FFC3696FFB770C11FAC0F9F8" authorityName="Antic &amp; Makarov" authorityYear="2016" box="[1157,1300,1541,1568]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="10" pageNumber="49" phylum="Arthropoda" rank="species" species="barjadzei">
<emphasis id="B93CEAF8FFC3696FFB770C11FAC0F9F8" box="[1157,1300,1541,1568]" italics="true" pageId="10" pageNumber="49">A. barjadzei</emphasis>
</taxonomicName>
), by the shape of paranota, and by the shorter and rodlike, rather bacilliform macrochaetae (vs longer trichoid macrochaetae in 
<taxonomicName id="4C484D69FFC3696FFE760C5BFDC7F981" authorityName="Antic &amp; Makarov" authorityYear="2016" box="[388,531,1612,1638]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="10" pageNumber="49" phylum="Arthropoda" rank="species" species="barjadzei">
<emphasis id="B93CEAF8FFC3696FFE760C5BFDC7F981" box="[388,531,1612,1638]" italics="true" pageId="10" pageNumber="49">A. barjadzei</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C3526561FFC3696FFF4F0C9AFAFEF91E" pageId="10" pageNumber="49" type="etymology">
<paragraph id="8BF736EAFFC3696FFF4F0C9AFE90F940" blockId="10.[189,324,1677,1703]" box="[189,324,1677,1703]" pageId="10" pageNumber="49">
<heading id="D0BF8186FFC3696FFF4F0C9AFE90F940" bold="true" box="[189,324,1677,1703]" fontSize="11" level="3" pageId="10" pageNumber="49" reason="3">
<emphasis id="B93CEAF8FFC3696FFF4F0C9AFE90F940" bold="true" box="[189,324,1677,1703]" pageId="10" pageNumber="49">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BF736EAFFC3696FFF4F0CABFAFEF91E" blockId="10.[188,1399,1723,1786]" pageId="10" pageNumber="49">
In Georgian mythology, Devi (‘ 
<emphasis id="B93CEAF8FFC3696FFDDE0CACFDA3F930" bold="true" box="[556,631,1723,1751]" pageId="10" pageNumber="49">დევი</emphasis>
’) is a many-headed malevolent giant who lives in the underworld. The 
<typeStatus id="54F38848FFC3696FFF030CC8FEF0F91E" box="[241,292,1759,1785]" pageId="10" pageNumber="49">type</typeStatus>
locality Devis Khvreli (‘ 
<emphasis id="B93CEAF8FFC3696FFDB90CC9FCF8F91D" bold="true" box="[587,812,1758,1786]" pageId="10" pageNumber="49">დევის ხვრელი</emphasis>
’) means ‘Devi’s hole’. Noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C3526561FFC3696EFF4F0D37FAC5FE29" lastPageId="11" lastPageNumber="50" pageId="10" pageNumber="49" type="materials_examined">
<paragraph id="8BF736EAFFC3696FFF4F0D37FE7EF8DD" blockId="10.[189,426,1824,1850]" box="[189,426,1824,1850]" pageId="10" pageNumber="49">
<heading id="D0BF8186FFC3696FFF4F0D37FE7EF8DD" bold="true" box="[189,426,1824,1850]" fontSize="11" level="3" pageId="10" pageNumber="49" reason="3">
<emphasis id="B93CEAF8FFC3696FFF4F0D37FE7EF8DD" bold="true" box="[189,426,1824,1850]" pageId="10" pageNumber="49">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BF736EAFFC3696FFF180D58FEE9F835" blockId="10.[189,1399,1871,2002]" pageId="10" pageNumber="49">
<materialsCitation id="3B203CB7FFC3696FFF180D58FEE9F835" collectingDate="2022-07-26" collectionCode="NHMW" collectorName="D. Antic &amp; E. Kiria &amp; L. Shavadze" country="Georgia" county="Chiatura Municipality" elevation="691" location="Zemo Imereti plateau karst massif" pageId="10" pageNumber="49" specimenCode="MY10364" specimenCount="1" specimenCount-male="1" stateProvince="Imereti Region" typeStatus="holotype">
<emphasis id="B93CEAF8FFC3696FFF180D58FE88F88E" bold="true" box="[234,348,1871,1897]" pageId="10" pageNumber="49">
<typeStatus id="54F38848FFC3696FFF180D58FE88F88E" box="[234,348,1871,1897]" pageId="10" pageNumber="49" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F35F767AFFC3696FFF4F0D65FEEBF86B" box="[189,319,1906,1932]" name="Georgia" pageId="10" pageNumber="49">GEORGIA</collectingCountry>
• 
<specimenCount id="9D4EFD63FFC3696FFE910D67FEAEF86B" box="[355,378,1904,1932]" count="1" pageId="10" pageNumber="49" type="male">♂</specimenCount>
; 
<collectingRegion id="498CF808FFC3696FFE7D0D65FD9CF86B" box="[399,584,1906,1932]" country="Georgia" name="Imereti" pageId="10" pageNumber="49">Imereti Region</collectingRegion>
, 
<collectingCounty id="62964E66FFC3696FFDAB0D65FCB0F86B" box="[601,868,1906,1932]" pageId="10" pageNumber="49">Chiatura Municipality</collectingCounty>
, 
<location id="8E976031FFC3696FFC870D64FAF4F86B" LSID="urn:lsid:plazi:treatment:03E187FCFFC36974FDB10E05FDB8F9EB:8E976031FFC3696FFC870D64FAF4F86B" box="[885,1312,1906,1933]" country="Georgia" county="Chiatura Municipality" name="Zemo Imereti plateau karst massif" pageId="10" pageNumber="49" stateProvince="Imereti Region">Zemo Imereti plateau karst massif</location>
, 
<location id="8E976031FFC3696FFAC20D65FEC4F848" LSID="urn:lsid:plazi:treatment:03E187FCFFC36974FDB10E05FDB8F9EB:8E976031FFC3696FFAC20D65FEC4F848" country="Georgia" county="Chiatura Municipality" municipality="Zemo" name="Rgani village" pageId="10" pageNumber="49" stateProvince="Imereti Region">Rgani village</location>
, 
<location id="8E976031FFC3696FFEE90D82FDD2F848" LSID="urn:lsid:plazi:treatment:03E187FCFFC36974FDB10E05FDB8F9EB:8E976031FFC3696FFEE90D82FDD2F848" box="[283,518,1941,1967]" country="Georgia" county="Chiatura Municipality" municipality="Zemo" name="Devis Khvreli Cave" pageId="10" pageNumber="49" stateProvince="Imereti Region">Devis Khvreli Cave</location>
; 
<elevation id="0065D1D9FFC3696FFDE30D82FD4DF848" box="[529,665,1941,1968]" metricMagnitude="2" metricUnit="m" metricValue="6.909999999999999" pageId="10" pageNumber="49" unit="m" value="691.0">
<quantity id="4CB09B0FFFC3696FFDE30D82FD8EF857" box="[529,602,1941,1968]" metricMagnitude="2" metricUnit="m" metricValue="6.909999999999999" pageId="10" pageNumber="49" unit="m" value="691.0">691 m</quantity>
a.s.l.
</elevation>
; 
<date id="FFF6102AFFC3696FFD540D82FCE3F848" box="[678,823,1941,1967]" pageId="10" pageNumber="49" value="2022-07-26">
<collectingDate id="EFB2E9C2FFC3696FFD540D82FCE3F848" box="[678,823,1941,1967]" pageId="10" pageNumber="49" value="2022-07-26">26 Jul. 2022</collectingDate>
</date>
; 
<collectorName id="26BD533CFFC3696FFCB10D81FC7CF848" box="[835,936,1941,1968]" pageId="10" pageNumber="49">D. Antić</collectorName>
, 
<collectorName id="26BD533CFFC3696FFC410D81FBDBF848" box="[947,1039,1941,1968]" pageId="10" pageNumber="49">E. Kiria</collectorName>
and 
<collectorName id="26BD533CFFC3696FFBB40D81FB02F848" box="[1094,1238,1941,1968]" pageId="10" pageNumber="49">L. Shavadze</collectorName>
leg.; 
<collectionCode id="ED59AE2FFFC3696FFAE50D81FAA3F857" box="[1303,1399,1942,1968]" collectionName="Austria, Wien, Naturhistorisches Museum Wien" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="10" pageNumber="49" type="Museum">NHMW</collectionCode>
<specimenCode id="DBEE9E91FFC3696FFF4F0DAFFEEDF835" box="[189,313,1976,2002]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="10" pageNumber="49" type="Museum">MY10364</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BF736EAFFC2696EFF180B1EFAB9FE8D" blockId="11.[189,1398,265,362]" pageId="11" pageNumber="50">
<materialsCitation id="3B203CB7FFC2696EFF180B1EFB4EFEA1" collectingDate="2022-07-26" collectionCode="IZB" collectorName="D. Antic &amp; E. Kiria &amp; L. Shavadze" country="Georgia" county="Chiatura Municipality" elevation="691" location="GEORGIA" pageId="11" pageNumber="50" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Imereti Region" typeStatus="paratype">
<emphasis id="B93CEAF8FFC2696EFF180B1EFEB3FEC4" bold="true" box="[234,359,265,291]" pageId="11" pageNumber="50">
<typeStatus id="54F38848FFC2696EFF180B1EFEB3FEC4" box="[234,359,265,291]" pageId="11" pageNumber="50" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="F35F767AFFC2696EFF4F0B3BFEEBFEA1" box="[189,319,300,326]" name="Georgia" pageId="11" pageNumber="50">GEORGIA</collectingCountry>
• 
<specimenCount id="9D4EFD63FFC2696EFEAC0B3BFE58FEA1" box="[350,396,298,326]" count="1" pageId="11" pageNumber="50" type="male">1 ♂</specimenCount>
, 
<specimenCount id="9D4EFD63FFC2696EFE6C0B3BFE19FEA1" box="[414,461,300,326]" count="1" pageId="11" pageNumber="50" type="female">1 ♀</specimenCount>
(used for SEM); same collection data as for holotype; 
<collectionCode id="ED59AE2FFFC2696EFB990B3BFB4EFEA1" box="[1131,1178,300,326]" collectionName="IZB" pageId="11" pageNumber="50">IZB</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B203CB7FFC2696EFB4B0B3BFDAFFE8E" collectingDate="2022-07-26" collectionCode="IZISU" collectorName="D. Antic &amp; E. Kiria &amp; L. Shavadze" country="Georgia" county="Chiatura Municipality" elevation="691" location="Zemo Imereti plateau karst massif" pageId="11" pageNumber="50" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Imereti Region" typeStatus="paratype">
<specimenCount id="9D4EFD63FFC2696EFB4B0B3BFB3CFEA1" box="[1209,1256,298,326]" count="1" pageId="11" pageNumber="50" type="male">1 ♂</specimenCount>
, 
<specimenCount id="9D4EFD63FFC2696EFB0B0B3BFAFDFEA1" box="[1273,1321,300,326]" count="1" pageId="11" pageNumber="50" type="female">1 ♀</specimenCount>
; same collection data as for holotype; 
<collectionCode id="ED59AE2FFFC2696EFDC20B58FDAFFE8E" box="[560,635,335,361]" collectionName="IZISU" pageId="11" pageNumber="50">IZISU</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B203CB7FFC2696EFD660B58FABDFE8D" box="[660,1385,335,362]" collectingDate="2022-07-26" collectionCode="NHMW" collectorName="D. Antic &amp; E. Kiria &amp; L. Shavadze" country="Georgia" county="Chiatura Municipality" elevation="691" location="Zemo Imereti plateau karst massif" pageId="11" pageNumber="50" specimenCode="MY10365" specimenCount="1" specimenCount-female="1" stateProvince="Imereti Region" typeStatus="paratype">
<specimenCount id="9D4EFD63FFC2696EFD660B58FD15FE8E" box="[660,705,335,361]" count="1" pageId="11" pageNumber="50" type="female">1 ♀</specimenCount>
; same collection data as for holotype; 
<collectionCode id="ED59AE2FFFC2696EFB770B58FB31FE8E" box="[1157,1253,335,361]" collectionName="Austria, Wien, Naturhistorisches Museum Wien" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="11" pageNumber="50" type="Museum">NHMW</collectionCode>
<specimenCode id="DBEE9E91FFC2696EFB1E0B58FABDFE8D" box="[1260,1385,335,362]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="11" pageNumber="50" type="Museum">MY10365</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BF736EAFFC2696EFF180B87FE35FE4D" blockId="11.[189,1297,400,462]" box="[234,481,400,426]" pageId="11" pageNumber="50">
<emphasis id="B93CEAF8FFC2696EFF180B87FE35FE4D" bold="true" box="[234,481,400,426]" pageId="11" pageNumber="50">Additional material</emphasis>
</paragraph>
<paragraph id="8BF736EAFFC2696EFF4F0BA3FAC5FE29" blockId="11.[189,1297,400,462]" box="[189,1297,433,462]" pageId="11" pageNumber="50">
<materialsCitation id="3B203CB7FFC2696EFF4F0BA3FAD9FE29" box="[189,1293,433,462]" collectingDate="2018-11-23" collectionCode="IZISU" collectorName="G. Nebieridze" country="Georgia" county="Chiatura Municipality" elevation="691" location="GEORGIA" pageId="11" pageNumber="50" specimenCount="6" specimenCount-male="1" stateProvince="Imereti Region" typeStatus="holotype">
<collectingCountry id="F35F767AFFC2696EFF4F0BA3FEEBFE29" box="[189,319,436,462]" name="Georgia" pageId="11" pageNumber="50">GEORGIA</collectingCountry>
• 
<specimenCount id="9D4EFD63FFC2696EFEAA0BA3FE50FE2A" box="[344,388,433,462]" count="1" pageId="11" pageNumber="50" type="male">1 ♂</specimenCount>
, 
<specimenCount id="9D4EFD63FFC2696EFE610BA4FE0AFE2A" box="[403,478,435,462]" pageId="11" pageNumber="50">5 juvs</specimenCount>
; same cave as for holotype; 
<date id="FFF6102AFFC2696EFCDB0BA3FC1FFE29" box="[809,971,436,462]" pageId="11" pageNumber="50" value="2018-11-23">
<collectingDate id="EFB2E9C2FFC2696EFCDB0BA3FC1FFE29" box="[809,971,436,462]" pageId="11" pageNumber="50" value="2018-11-23">23 Nov. 2018</collectingDate>
</date>
; 
<collectorName id="26BD533CFFC2696EFC2A0BA3FBA8FE2A" box="[984,1148,435,462]" pageId="11" pageNumber="50">G. Nebieridze</collectorName>
leg.; 
<collectionCode id="ED59AE2FFFC2696EFB4C0BA3FAD9FE29" box="[1214,1293,436,462]" collectionName="IZISU" pageId="11" pageNumber="50">IZISU</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3526561FFC26974FF4F0BFCFDE1FC29" lastPageId="17" lastPageNumber="56" pageId="11" pageNumber="50" type="description">
<paragraph id="8BF736EAFFC2696EFF4F0BFCFE9AFDE2" blockId="11.[189,334,491,517]" box="[189,334,491,517]" pageId="11" pageNumber="50">
<heading id="D0BF8186FFC2696EFF4F0BFCFE9AFDE2" bold="true" box="[189,334,491,517]" fontSize="11" level="3" pageId="11" pageNumber="50" reason="3">
<emphasis id="B93CEAF8FFC2696EFF4F0BFCFE9AFDE2" bold="true" box="[189,334,491,517]" pageId="11" pageNumber="50">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BF736EAFFC2696EFF4F080EFD41FD7E" blockId="11.[189,1399,536,666]" pageId="11" pageNumber="50">
<smallCapsWord id="8D11A036FFC2696EFF4F080EFF39FDD6" baselines="556,556" box="[189,237,537,563]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Size" pageId="11" pageNumber="50">SIZE</smallCapsWord>
<smallCapsWord id="8D11A036FFC2696EFF0A0809FEF1FDD6" baselines="556" box="[248,293,542,561]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="and" pageId="11" pageNumber="50">AND</smallCapsWord>
<smallCapsWord id="8D11A036FFC2696EFEDD0809FE5CFDD6" baselines="556" box="[303,392,542,561]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="number" pageId="11" pageNumber="50">NUMBER</smallCapsWord>
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. Body with 31 rings (including telson). 
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male 
<quantity id="4CB09B0FFFC2696EFB2F080EFAE5FDD4" box="[1245,1329,537,563]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="11" pageNumber="50" unit="mm" value="11.0">11 mm</quantity>
long, vertical diameter of the largest ring 
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. 
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males 
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long, vertical diameter of the largest ring 
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, respectively. 
<typeStatus id="54F38848FFC2696EFD5D084AFCC2FD90" box="[687,790,605,631]" pageId="11" pageNumber="50" type="paratype">Paratype</typeStatus>
females 
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long, vertical diameter of the largest ring 
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, respectively.
</paragraph>
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(
<figureCitation id="13732A6FFFC2696EFEAB08A8FE05FD3D" box="[345,465,703,730]" captionStart="Fig" captionStartId="1.[189,232,1857,1883]" captionTargetBox="[189,1398,420,1810]" captionTargetId="figure-57@1.[189,1398,420,1810]" captionTargetPageId="1" captionText="Fig. 1. Troglobiotic members of the genus Acanthophorella Antić &amp; Makarov, 2016 in situ. A–B. Acanthophorella barjadzei Antić &amp; Makarov, 2016, ♂ from Nikortsminda Sakinule Cave. C–D. Acanthophorella barjadzei Antić &amp; Makarov, 2016, ♂ from Muradi Cave. E–F. Acanthophorella devi Antić sp. nov., types, ♂♂. G–H. Acanthophorella valerii Antić sp. nov., holotype, ♂ (NHMW MY10366). Photos by Dragan Antić." figureDoi="http://doi.org/10.5281/zenodo.10149300" httpUri="https://zenodo.org/record/10149300/files/figure.png" pageId="11" pageNumber="50">Figs 1E–F</figureCitation>
, 
<figureCitation id="13732A6FFFC2696EFE1208A8FE25FD3E" box="[480,497,703,729]" captionStart="Fig" captionStartId="12.[189,232,1879,1905]" captionTargetBox="[189,1398,270,1829]" captionTargetId="figure-16@12.[189,1398,270,1829]" captionTargetPageId="12" captionText="Fig. 6. Acanthophorella devi Antić sp. nov. A–B. Paratype, ♀ (IZB). C–E. Holotype, ♂ (NHMW MY10364). A. Habitus, lateral view. B. Head and collum, lateral view. C. Habitus, lateral view. D. Midbody rings, dorsal view. E. Midbody rings, lateral view. Scale bars: A, C–E = 1 mm; B = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10149312" httpUri="https://zenodo.org/record/10149312/files/figure.png" pageId="11" pageNumber="50">6</figureCitation>
). Living animals white with pale brownish ommatidia.
</paragraph>
<paragraph id="8BF736EAFFC2696EFF4F0917FABEFBAA" blockId="11.[188,1400,767,1102]" pageId="11" pageNumber="50">
<smallCapsWord id="8D11A036FFC2696EFF4F0917FF28FCFF" baselines="787,787" box="[189,252,768,794]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Head" pageId="11" pageNumber="50">HEAD</smallCapsWord>
(
<figureCitation id="13732A6FFFC2696EFEFC0917FE44FCFD" box="[270,400,767,794]" captionStart="Fig" captionStartId="12.[189,232,1879,1905]" captionTargetBox="[189,1398,270,1829]" captionTargetId="figure-16@12.[189,1398,270,1829]" captionTargetPageId="12" captionText="Fig. 6. Acanthophorella devi Antić sp. nov. A–B. Paratype, ♀ (IZB). C–E. Holotype, ♂ (NHMW MY10364). A. Habitus, lateral view. B. Head and collum, lateral view. C. Habitus, lateral view. D. Midbody rings, dorsal view. E. Midbody rings, lateral view. Scale bars: A, C–E = 1 mm; B = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10149312" httpUri="https://zenodo.org/record/10149312/files/figure.png" pageId="11" pageNumber="50">Figs 6A–B</figureCitation>
, 
<figureCitation id="13732A6FFFC2696EFE6C0917FE32FCFD" box="[414,486,768,794]" captionStart="Fig" captionStartId="13.[189,232,1770,1796]" captionTargetBox="[189,1398,265,1700]" captionTargetId="figure-11@13.[189,1398,265,1700]" captionTargetPageId="13" captionText="Fig. 7. Acanthophorella devi Antić sp. nov., paratype, ♂ (IZB). A. Head, anterior view. B. Labrum, anterior view. C. Right side of head showing field with ommatidia, organ of Tömösvary and group of papilliform outgrowths, lateral view. D. Right field with ommatidia, anterior view. E. Ring 16, dorsal view.F. Ring 16, dorsolateral view. G. Rings 14–16, posterior view. H. Left interior macrochaeta of ring 15, lateral view. Scale bars: A–B, E–G = 0.1 mm; C, H = 0.05 mm; D = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.10149314" httpUri="https://zenodo.org/record/10149314/files/figure.png" pageId="11" pageNumber="50">7A–D</figureCitation>
). Setose, roundly convex in females, in males with labral and frontal surfaces flat with a convexity between and with a pair of lateral lobes, each below antennal sockets. Labrum with three medial teeth and 5+5 labral and 2+2 supralabral setae. Promentum triangular, without setae. Lamellae linguales with 7+6 setae. Stipites with ca 25 setae each. Antennae 
<quantity id="4CB09B0FFFC2696EFBD90971FBB9FC66" box="[1067,1133,870,897]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="50" unit="mm" value="2.0">2 mm</quantity>
long in 
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male. Length of antennomeres (in mm): I (0.1), II (0.2), III (0.55), IV (0.27), V (0.5), VI (0.17), VII (0.17) and VIII (0.04). Length/breadth ratios of antennomeres I–VII: I (1), II (1.5), III (5.5), IV (2.7), V (3.5), VI (1.1) and VII (1.5). Antennomeres II, IV, V, VI and VII with one, three, one, four and one long sensillum trichoideum, respectively. Antennomere 7 with one rather bacilliform sensillum (sensillum basiconicum?) curved distad, located below sensillum trichodeum. Lateral to antennal sockets, a group of papilliform outgrowths present. Number of ommatidia: 
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2–3 irregular rows (
<figureCitation id="13732A6FFFC2696EFB570E24FAD3FBAA" box="[1189,1287,1075,1101]" captionStart="Fig" captionStartId="12.[189,232,1879,1905]" captionTargetBox="[189,1398,270,1829]" captionTargetId="figure-16@12.[189,1398,270,1829]" captionTargetPageId="12" captionText="Fig. 6. Acanthophorella devi Antić sp. nov. A–B. Paratype, ♀ (IZB). C–E. Holotype, ♂ (NHMW MY10364). A. Habitus, lateral view. B. Head and collum, lateral view. C. Habitus, lateral view. D. Midbody rings, dorsal view. E. Midbody rings, lateral view. Scale bars: A, C–E = 1 mm; B = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10149312" httpUri="https://zenodo.org/record/10149312/files/figure.png" pageId="11" pageNumber="50">Figs 6A</figureCitation>
, 
<figureCitation id="13732A6FFFC2696EFAE10E24FA8AFBAA" box="[1299,1374,1075,1101]" captionStart="Fig" captionStartId="13.[189,232,1770,1796]" captionTargetBox="[189,1398,265,1700]" captionTargetId="figure-11@13.[189,1398,265,1700]" captionTargetPageId="13" captionText="Fig. 7. Acanthophorella devi Antić sp. nov., paratype, ♂ (IZB). A. Head, anterior view. B. Labrum, anterior view. C. Right side of head showing field with ommatidia, organ of Tömösvary and group of papilliform outgrowths, lateral view. D. Right field with ommatidia, anterior view. E. Ring 16, dorsal view.F. Ring 16, dorsolateral view. G. Rings 14–16, posterior view. H. Left interior macrochaeta of ring 15, lateral view. Scale bars: A–B, E–G = 0.1 mm; C, H = 0.05 mm; D = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.10149314" httpUri="https://zenodo.org/record/10149314/files/figure.png" pageId="11" pageNumber="50">7C–D</figureCitation>
).
</paragraph>
<paragraph id="8BF736EAFFC2696EFF4F0E64FDEBFB57" blockId="11.[189,1398,1139,1200]" pageId="11" pageNumber="50">
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. Narrower than head, with six macrochaetae as all body rings. Anterior edge semi-circular, posterior margin gently concave.
</paragraph>
<paragraph id="8BF736EAFFC2696EFF4F0EC1FB43FAD3" blockId="11.[189,1399,1238,1333]" pageId="11" pageNumber="50">
<smallCapsWord id="8D11A036FFC2696EFF4F0EC1FF2AFB09" baselines="1257,1257" box="[189,254,1238,1264]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Body" pageId="11" pageNumber="50">BODY</smallCapsWord>
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(
<figureCitation id="13732A6FFFC2696EFE970EC1FDC3FB17" box="[357,535,1238,1264]" captionStart="Fig" captionStartId="12.[189,232,1879,1905]" captionTargetBox="[189,1398,270,1829]" captionTargetId="figure-16@12.[189,1398,270,1829]" captionTargetPageId="12" captionText="Fig. 6. Acanthophorella devi Antić sp. nov. A–B. Paratype, ♀ (IZB). C–E. Holotype, ♂ (NHMW MY10364). A. Habitus, lateral view. B. Head and collum, lateral view. C. Habitus, lateral view. D. Midbody rings, dorsal view. E. Midbody rings, lateral view. Scale bars: A, C–E = 1 mm; B = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10149312" httpUri="https://zenodo.org/record/10149312/files/figure.png" pageId="11" pageNumber="50">Figs 6A, C–E</figureCitation>
, 
<figureCitation id="13732A6FFFC2696EFDDC0EC1FDACFB17" box="[558,632,1238,1264]" captionStart="Fig" captionStartId="13.[189,232,1770,1796]" captionTargetBox="[189,1398,265,1700]" captionTargetId="figure-11@13.[189,1398,265,1700]" captionTargetPageId="13" captionText="Fig. 7. Acanthophorella devi Antić sp. nov., paratype, ♂ (IZB). A. Head, anterior view. B. Labrum, anterior view. C. Right side of head showing field with ommatidia, organ of Tömösvary and group of papilliform outgrowths, lateral view. D. Right field with ommatidia, anterior view. E. Ring 16, dorsal view.F. Ring 16, dorsolateral view. G. Rings 14–16, posterior view. H. Left interior macrochaeta of ring 15, lateral view. Scale bars: A–B, E–G = 0.1 mm; C, H = 0.05 mm; D = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.10149314" httpUri="https://zenodo.org/record/10149314/files/figure.png" pageId="11" pageNumber="50">7E–H</figureCitation>
). With well-developed lateral keels, rounded in dorsal view. Macrochaetae rather short and rodlike, with longitudinal ribs, ending with spikes (
<figureCitation id="13732A6FFFC2696EFB0D0EEFFABDFAF5" box="[1279,1385,1272,1298]" captionStart="Fig" captionStartId="13.[189,232,1770,1796]" captionTargetBox="[189,1398,265,1700]" captionTargetId="figure-11@13.[189,1398,265,1700]" captionTargetPageId="13" captionText="Fig. 7. Acanthophorella devi Antić sp. nov., paratype, ♂ (IZB). A. Head, anterior view. B. Labrum, anterior view. C. Right side of head showing field with ommatidia, organ of Tömösvary and group of papilliform outgrowths, lateral view. D. Right field with ommatidia, anterior view. E. Ring 16, dorsal view.F. Ring 16, dorsolateral view. G. Rings 14–16, posterior view. H. Left interior macrochaeta of ring 15, lateral view. Scale bars: A–B, E–G = 0.1 mm; C, H = 0.05 mm; D = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.10149314" httpUri="https://zenodo.org/record/10149314/files/figure.png" pageId="11" pageNumber="50">Fig. 7H</figureCitation>
). CIX (ring 15) = 0.6; MIX (ring 15) ~ 1.5; PIX (ring 15) = 0.6; MA (ring 15) ~ 100°.
</paragraph>
<paragraph id="8BF736EAFFC2696EFF4F0F4CFC29FA70" blockId="11.[189,1398,1370,1431]" pageId="11" pageNumber="50">
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. Epiproct with a pair of spinnerets and 3+3 setae (1+1 paramedian, 2+2 marginal). Hypoproct with 1+1 distal setae. Paraprocts with 3+3 marginal setae in distal part.
</paragraph>
<paragraph id="8BF736EAFFC2696EFF4F0FAAFF2BFA1E" blockId="11.[189,1398,1469,1529]" pageId="11" pageNumber="50">
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1 
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2. In both sexes with tarsal combs; femora, postfemora and tibiae with long and robust setae.
</paragraph>
<paragraph id="8BF736EAFFC2696EFF4F0C37FBBCF8AD" blockId="11.[188,1401,1567,1867]" pageId="11" pageNumber="50">
<smallCapsWord id="8D11A036FFC2696EFF4F0C37FF2BF9DF" baselines="1587,1587" box="[189,255,1568,1594]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Male" pageId="11" pageNumber="50">MALE</smallCapsWord>
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(
<figureCitation id="13732A6FFFC2696EFE080C08FDA6F9DD" box="[506,626,1567,1594]" captionStart="Fig" captionStartId="1.[189,232,1857,1883]" captionTargetBox="[189,1398,420,1810]" captionTargetId="figure-57@1.[189,1398,420,1810]" captionTargetPageId="1" captionText="Fig. 1. Troglobiotic members of the genus Acanthophorella Antić &amp; Makarov, 2016 in situ. A–B. Acanthophorella barjadzei Antić &amp; Makarov, 2016, ♂ from Nikortsminda Sakinule Cave. C–D. Acanthophorella barjadzei Antić &amp; Makarov, 2016, ♂ from Muradi Cave. E–F. Acanthophorella devi Antić sp. nov., types, ♂♂. G–H. Acanthophorella valerii Antić sp. nov., holotype, ♂ (NHMW MY10366). Photos by Dragan Antić." figureDoi="http://doi.org/10.5281/zenodo.10149300" httpUri="https://zenodo.org/record/10149300/files/figure.png" pageId="11" pageNumber="50">Figs 1E–F</figureCitation>
, 
<figureCitation id="13732A6FFFC2696EFD700C08FD11F9DD" box="[642,709,1567,1594]" captionStart="Fig" captionStartId="14.[189,232,1892,1918]" captionTargetBox="[229,1359,265,1842]" captionTargetId="figure-17@14.[229,1359,265,1842]" captionTargetPageId="14" captionText="Fig. 8. Acanthophorella devi Antić sp. nov. A. Paratype, ♀ (IZB), B–G. Paratype, ♂ (IZB). A. Legpair 2, anterior view. B. Leg-pair 2, anterior view. C. Right leg 4, anterior view, white arrows indicate distoventral pads. D. Leg-pair 7, posterior view. E. Leg-pair 10, anterior view. F. Anterior gonopods, anterior view. G. Posterior gonopods, posterior view. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10149316" httpUri="https://zenodo.org/record/10149316/files/figure.png" pageId="11" pageNumber="50">8B–E</figureCitation>
). Gonopores mesally on coxae 2 (
<figureCitation id="13732A6FFFC2696EFBAC0C08FB6FF9DE" box="[1118,1211,1567,1593]" captionStart="Fig" captionStartId="14.[189,232,1892,1918]" captionTargetBox="[229,1359,265,1842]" captionTargetId="figure-17@14.[229,1359,265,1842]" captionTargetPageId="14" captionText="Fig. 8. Acanthophorella devi Antić sp. nov. A. Paratype, ♀ (IZB), B–G. Paratype, ♂ (IZB). A. Legpair 2, anterior view. B. Leg-pair 2, anterior view. C. Right leg 4, anterior view, white arrows indicate distoventral pads. D. Leg-pair 7, posterior view. E. Leg-pair 10, anterior view. F. Anterior gonopods, anterior view. G. Posterior gonopods, posterior view. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10149316" httpUri="https://zenodo.org/record/10149316/files/figure.png" pageId="11" pageNumber="50">Fig. 8B</figureCitation>
). Leg-pairs 3–7 enlarged, especially leg-pairs 3, 4 and 7 (
<figureCitation id="13732A6FFFC2696EFD6F0C56FCC3F9BB" box="[669,791,1601,1628]" captionStart="Fig" captionStartId="1.[189,232,1857,1883]" captionTargetBox="[189,1398,420,1810]" captionTargetId="figure-57@1.[189,1398,420,1810]" captionTargetPageId="1" captionText="Fig. 1. Troglobiotic members of the genus Acanthophorella Antić &amp; Makarov, 2016 in situ. A–B. Acanthophorella barjadzei Antić &amp; Makarov, 2016, ♂ from Nikortsminda Sakinule Cave. C–D. Acanthophorella barjadzei Antić &amp; Makarov, 2016, ♂ from Muradi Cave. E–F. Acanthophorella devi Antić sp. nov., types, ♂♂. G–H. Acanthophorella valerii Antić sp. nov., holotype, ♂ (NHMW MY10366). Photos by Dragan Antić." figureDoi="http://doi.org/10.5281/zenodo.10149300" httpUri="https://zenodo.org/record/10149300/files/figure.png" pageId="11" pageNumber="50">Fig. 1E–F</figureCitation>
). Leg-pairs 3 and 4 very thick, each with a proximal lateral protrusion on prefemora; prefemora and femora strong, rectangular; tarsi shorter and thicker compared to other legs; femora, postfemora and tibiae each with a distoventral pad (
<figureCitation id="13732A6FFFC2696EFB950C91FB6AF947" box="[1127,1214,1670,1696]" captionStart="Fig" captionStartId="14.[189,232,1892,1918]" captionTargetBox="[229,1359,265,1842]" captionTargetId="figure-17@14.[229,1359,265,1842]" captionTargetPageId="14" captionText="Fig. 8. Acanthophorella devi Antić sp. nov. A. Paratype, ♀ (IZB), B–G. Paratype, ♂ (IZB). A. Legpair 2, anterior view. B. Leg-pair 2, anterior view. C. Right leg 4, anterior view, white arrows indicate distoventral pads. D. Leg-pair 7, posterior view. E. Leg-pair 10, anterior view. F. Anterior gonopods, anterior view. G. Posterior gonopods, posterior view. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10149316" httpUri="https://zenodo.org/record/10149316/files/figure.png" pageId="11" pageNumber="50">Fig. 8C</figureCitation>
, arrows indicate distoventral pads). Leg-pair 5 with a proximal, somewhat elongated, anterior, triangular, coxal protrusion (
<figureCitation id="13732A6FFFC2696EFF370CDDFECAF903" box="[197,286,1738,1764]" captionStart="Fig" captionStartId="1.[189,232,1857,1883]" captionTargetBox="[189,1398,420,1810]" captionTargetId="figure-57@1.[189,1398,420,1810]" captionTargetPageId="1" captionText="Fig. 1. Troglobiotic members of the genus Acanthophorella Antić &amp; Makarov, 2016 in situ. A–B. Acanthophorella barjadzei Antić &amp; Makarov, 2016, ♂ from Nikortsminda Sakinule Cave. C–D. Acanthophorella barjadzei Antić &amp; Makarov, 2016, ♂ from Muradi Cave. E–F. Acanthophorella devi Antić sp. nov., types, ♂♂. G–H. Acanthophorella valerii Antić sp. nov., holotype, ♂ (NHMW MY10366). Photos by Dragan Antić." figureDoi="http://doi.org/10.5281/zenodo.10149300" httpUri="https://zenodo.org/record/10149300/files/figure.png" pageId="11" pageNumber="50">Fig. 1E</figureCitation>
). Leg-pair 6 without peculiarities. Leg-pair 7 robust; coxae with wide, well-developed, flattened posterior processes, covered with long setae anteriorly and with a lateral lobe giving the impression of a subtriangular appereance (
<figureCitation id="13732A6FFFC2696EFE1D0D19FD98F8CF" box="[495,588,1806,1832]" captionStart="Fig" captionStartId="14.[189,232,1892,1918]" captionTargetBox="[229,1359,265,1842]" captionTargetId="figure-17@14.[229,1359,265,1842]" captionTargetPageId="14" captionText="Fig. 8. Acanthophorella devi Antić sp. nov. A. Paratype, ♀ (IZB), B–G. Paratype, ♂ (IZB). A. Legpair 2, anterior view. B. Leg-pair 2, anterior view. C. Right leg 4, anterior view, white arrows indicate distoventral pads. D. Leg-pair 7, posterior view. E. Leg-pair 10, anterior view. F. Anterior gonopods, anterior view. G. Posterior gonopods, posterior view. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10149316" httpUri="https://zenodo.org/record/10149316/files/figure.png" pageId="11" pageNumber="50">Fig. 8D</figureCitation>
). Leg-pair 10 with coxal glands and a poorly developed or completely absent protrusion (
<figureCitation id="13732A6FFFC2696EFE670D27FE3AF8AD" box="[405,494,1840,1866]" captionStart="Fig" captionStartId="14.[189,232,1892,1918]" captionTargetBox="[229,1359,265,1842]" captionTargetId="figure-17@14.[229,1359,265,1842]" captionTargetPageId="14" captionText="Fig. 8. Acanthophorella devi Antić sp. nov. A. Paratype, ♀ (IZB), B–G. Paratype, ♂ (IZB). A. Legpair 2, anterior view. B. Leg-pair 2, anterior view. C. Right leg 4, anterior view, white arrows indicate distoventral pads. D. Leg-pair 7, posterior view. E. Leg-pair 10, anterior view. F. Anterior gonopods, anterior view. G. Posterior gonopods, posterior view. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10149316" httpUri="https://zenodo.org/record/10149316/files/figure.png" pageId="11" pageNumber="50">Fig. 8E</figureCitation>
). Leg-pair 11 with coxal glands, no other peculiarities.
</paragraph>
<paragraph id="8BF736EAFFC26974FF4F0D66FD3AFE17" blockId="11.[189,1399,1905,2000]" lastBlockId="17.[188,1399,265,496]" lastPageId="17" lastPageNumber="56" pageId="11" pageNumber="50">
<smallCapsWord id="8D11A036FFC2696EFF4F0D66FEF8F86E" baselines="1924,1924" box="[189,300,1905,1931]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Anterior" pageId="11" pageNumber="50">ANTERIOR</smallCapsWord>
<smallCapsWord id="8D11A036FFC2696EFEC00D61FE77F86E" baselines="1924" box="[306,419,1910,1929]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="gonopods" pageId="11" pageNumber="50">GONOPODS</smallCapsWord>
(
<figureCitation id="13732A6FFFC2696EFE410D66FDDCF86C" box="[435,520,1905,1931]" captionStart="Fig" captionStartId="14.[189,232,1892,1918]" captionTargetBox="[229,1359,265,1842]" captionTargetId="figure-17@14.[229,1359,265,1842]" captionTargetPageId="14" captionText="Fig. 8. Acanthophorella devi Antić sp. nov. A. Paratype, ♀ (IZB), B–G. Paratype, ♂ (IZB). A. Legpair 2, anterior view. B. Leg-pair 2, anterior view. C. Right leg 4, anterior view, white arrows indicate distoventral pads. D. Leg-pair 7, posterior view. E. Leg-pair 10, anterior view. F. Anterior gonopods, anterior view. G. Posterior gonopods, posterior view. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10149316" httpUri="https://zenodo.org/record/10149316/files/figure.png" pageId="11" pageNumber="50">Figs 8F</figureCitation>
, 
<figureCitation id="13732A6FFFC2696EFDE40D66FD83F86C" box="[534,599,1905,1931]" captionStart="Fig" captionStartId="15.[189,232,1787,1813]" captionTargetBox="[282,1305,265,1742]" captionTargetId="figure-12@15.[282,1305,265,1742]" captionTargetPageId="15" captionText="Fig. 9. Acanthophorella devi Antić sp. nov. A–F. Paratype, ♂ (IZB). G–I. Paratype, ♀ (IZB). A. Anterior gonopods, anterior view. B. Anterior gonopods, anterodistal view. C. Anterior gonopods, distal view. D. Anterior gonopods, lateral view. E. Anterior gonopods, lateral lamella and tuft, posterodistal view. F. Anterior gonopods, posterodistal view. G. Left vulva, lateral view. H. Vulvae, distal view. I. Left vulva, anterior view.Abbreviations: a = angiocoxite; ap = anterior process; b = bursa; ll = lateral lamella; mp = medial part; o = operculum; pl = posterior lobe; s = sternum; sa = synangiocoxal base; sl = sternal lamella; tf = angiocoxal tuft. Scale bars: A–D, F–I = 0.1 mm; E = 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.10149318" httpUri="https://zenodo.org/record/10149318/files/figure.png" pageId="11" pageNumber="50">9A–F</figureCitation>
, 
<figureCitation id="13732A6FFFC2696EFD960D66FD68F86C" box="[612,700,1905,1931]" captionStart="Fig" captionStartId="16.[189,232,1787,1813]" captionTargetBox="[293,1287,282,1718]" captionTargetId="figure-17@16.[287,1300,265,1743]" captionTargetPageId="16" captionText="Fig. 10. Acanthophorella devi Antić sp. nov. A–E. Paratype, ♂ (IZB). F–G. Paratype, ♀ (IZB). A. Anterior gonopods, anterior view. B. Anterior gonopods, posterior view. C. Anterior gonopods, lateral view. D. Posterior gonopods, posterior view. E. Left posterior gonopod, lateral view. F. Right vulva, distal view. G. Right vulva, lateral view. Abbreviations: a = angiocoxite; ap = anterior process; b = bursa; c = colpocoxite; ll = lateral lamella; mp = medial part; o = operculum; pl = posterior lobe; s = sternum; sa = synangiocoxal base; sl = sternal lamella; t = telopodite; tf = angiocoxal tuft. Drawings by DA (A–E) and MŠ (F–G). Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10149320" httpUri="https://zenodo.org/record/10149320/files/figure.png" pageId="11" pageNumber="50">10A–C</figureCitation>
, 
<figureCitation id="13732A6FFFC2696EFD350D66FD2DF86C" box="[711,761,1905,1931]" captionStart="Fig" captionStartId="33.[189,232,1822,1848]" captionTargetBox="[202,1378,705,1770]" captionTargetId="figure-170@33.[189,1398,686,1780]" captionTargetPageId="33" captionText="Fig. 21. Anterior gonopods in the genera Acanthophorella Antić &amp; Makarov, 2016 and PseudoflagellophorellaAntić &amp; Makarov, 2016, anterior views. A. A. auritaAntić sp. nov. B. A. barjadzei Antić &amp; Makarov, 2016. C. A. chegemi Antić &amp; Makarov, 2016. D. A. devi Antić sp. nov. E. A. irystoni Antić &amp; Makarov, 2016. F. A. valerii Antić sp. nov. G. P. eskovi Antić &amp; Makarov, 2016. H. P. mirabilis Antić &amp; Makarov, 2016. I. P. papilioformis Antić &amp; Makarov, 2016. B–C, E, G–I after Antić &amp; Makarov (2016). Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10149345" httpUri="https://zenodo.org/record/10149345/files/figure.png" pageId="11" pageNumber="50">21D</figureCitation>
). Gonopodal sternum (s) wide, medially with a poorlydeveloped, low, fimbriate lamella (sl) on anterior side. Angiocoxites (a) consisting of a medial part (mp), lateral lamellae (ll) and a synangiocoxal base with anterior processes (ap). Medial parts well-developed, together forming a heartlike structure, divided, but appressed to each other, shieldlike, distomesal and lateral margins folded posteriad; each medial part with subdistal posterior lobe (pl), triangular in posterior view; angiocoxites posteroproximally with a pair of tufts (tf) with hairlike outgrowths and a rounded lobe, covered by spiculiform outgrowths, from which a ridge, also covered with spiculiform outgrowths, continues and fuses with the posterior side of the medial part. Lateral lamellae low, rather wide in lateral view, with slightly denticulate margins. Anterior processes (ap) tapering distad, acuminate, twice as high as lateral lamellae. Coxal vesicles not observed.
</paragraph>
<caption id="DF376662FFC56969FF4F0D40FAA3F85F" ID-DOI="http://doi.org/10.5281/zenodo.10149312" ID-Zenodo-Dep="10149312" httpUri="https://zenodo.org/record/10149312/files/figure.png" pageId="12" pageNumber="51" startId="12.[189,232,1879,1905]" targetBox="[189,1398,270,1829]" targetPageId="12">
<paragraph id="8BF736EAFFC56969FF4F0D40FAA3F85F" blockId="12.[189,1399,1877,1976]" pageId="12" pageNumber="51">
<emphasis id="B93CEAF8FFC56969FF4F0D40FEC4F896" bold="true" box="[189,272,1879,1905]" pageId="12" pageNumber="51">Fig. 6.</emphasis>
<taxonomicName id="4C484D69FFC56969FEEF0D40FDBDF896" authority="Antic" authorityName="Antic" authorityYear="2023" box="[285,617,1879,1905]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="12" pageNumber="51" phylum="Arthropoda" rank="species" species="devi" status="sp. nov.">
<emphasis id="B93CEAF8FFC56969FEEF0D40FDCAF896" box="[285,542,1879,1905]" italics="true" pageId="12" pageNumber="51">Acanthophorella devi</emphasis>
Antić
</taxonomicName>
<taxonomicNameLabel id="A20F5783FFC56969FD840D40FD01F896" box="[630,725,1879,1905]" pageId="12" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B93CEAF8FFC56969FD130D40FCCDF896" bold="true" box="[737,793,1879,1905]" pageId="12" pageNumber="51">A–B</emphasis>
. Paratype, ♀ (IZB). 
<emphasis id="B93CEAF8FFC56969FBED0D40FB83F896" bold="true" box="[1055,1111,1879,1905]" pageId="12" pageNumber="51">C–E</emphasis>
. Holotype, ♂ (NHMW MY10364). 
<emphasis id="B93CEAF8FFC56969FEA60D6DFEBDF873" bold="true" box="[340,361,1914,1940]" pageId="12" pageNumber="51">A</emphasis>
. Habitus, lateral view. 
<emphasis id="B93CEAF8FFC56969FD600D6DFD72F873" bold="true" box="[658,678,1914,1940]" pageId="12" pageNumber="51">B</emphasis>
. Head and collum, lateral view. 
<emphasis id="B93CEAF8FFC56969FBB50D6DFB88F873" bold="true" box="[1095,1116,1914,1940]" pageId="12" pageNumber="51">C</emphasis>
. Habitus, lateral view. 
<emphasis id="B93CEAF8FFC56969FF4F0D8AFF06F850" bold="true" box="[189,210,1949,1975]" pageId="12" pageNumber="51">D</emphasis>
. Midbody rings, dorsal view. 
<emphasis id="B93CEAF8FFC56969FDDE0D8AFD94F850" bold="true" box="[556,576,1949,1975]" pageId="12" pageNumber="51">E</emphasis>
. Midbody rings, lateral view. Scale bars: A, C–E = 1 mm; B = 0.5 mm.
</paragraph>
</caption>
<caption id="DF376662FFC46968FF4F0CFDFBAFF875" ID-DOI="http://doi.org/10.5281/zenodo.10149314" ID-Zenodo-Dep="10149314" httpUri="https://zenodo.org/record/10149314/files/figure.png" pageId="13" pageNumber="52" startId="13.[189,232,1770,1796]" targetBox="[189,1398,265,1700]" targetPageId="13">
<paragraph id="8BF736EAFFC46968FF4F0CFDFBAFF875" blockId="13.[188,1399,1768,1938]" pageId="13" pageNumber="52">
<emphasis id="B93CEAF8FFC46968FF4F0CFDFEDAF8E2" bold="true" box="[189,270,1770,1797]" pageId="13" pageNumber="52">Fig. 7.</emphasis>
<taxonomicName id="4C484D69FFC46968FEEB0CFDFDB6F8E3" authority="Antic" authorityName="Antic" authorityYear="2023" box="[281,610,1770,1796]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="13" pageNumber="52" phylum="Arthropoda" rank="species" species="devi" status="sp. nov.">
<emphasis id="B93CEAF8FFC46968FEEB0CFDFDCDF8E3" box="[281,537,1770,1796]" italics="true" pageId="13" pageNumber="52">Acanthophorella devi</emphasis>
Antić
</taxonomicName>
<taxonomicNameLabel id="A20F5783FFC46968FD9F0CFCFD19F8E2" box="[621,717,1771,1797]" pageId="13" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
, paratype, ♂ (IZB). 
<emphasis id="B93CEAF8FFC46968FC3A0CFDFC09F8E3" bold="true" box="[968,989,1770,1796]" pageId="13" pageNumber="52">A</emphasis>
. Head, anterior view. 
<emphasis id="B93CEAF8FFC46968FB1F0CFCFAD5F8E2" bold="true" box="[1261,1281,1771,1797]" pageId="13" pageNumber="52">B</emphasis>
. Labrum, anterior view. 
<emphasis id="B93CEAF8FFC46968FE9B0D1AFEA9F8C0" bold="true" box="[361,381,1805,1831]" pageId="13" pageNumber="52">C</emphasis>
. Right side of head showing field with ommatidia, organ of Tömösvary and group of papilliform outgrowths, lateral view. 
<emphasis id="B93CEAF8FFC46968FD880D26FD5AF8AC" bold="true" box="[634,654,1841,1867]" pageId="13" pageNumber="52">D</emphasis>
. Right field with ommatidia, anterior view. 
<emphasis id="B93CEAF8FFC46968FB690D26FB7BF8AC" bold="true" box="[1179,1199,1841,1867]" pageId="13" pageNumber="52">E</emphasis>
. Ring 16, dorsal view. 
<emphasis id="B93CEAF8FFC46968FEF30D43FEC7F889" bold="true" box="[257,275,1876,1902]" pageId="13" pageNumber="52">F</emphasis>
. Ring 16, dorsolateral view. 
<emphasis id="B93CEAF8FFC46968FDAC0D43FDA1F889" bold="true" box="[606,629,1876,1902]" pageId="13" pageNumber="52">G</emphasis>
. Rings 14–16, posterior view. 
<emphasis id="B93CEAF8FFC46968FC240D43FC39F889" bold="true" box="[982,1005,1876,1902]" pageId="13" pageNumber="52">H</emphasis>
. Left interior macrochaeta of ring 15, lateral view. Scale bars: A–B, E–G = 0.1 mm; C, H = 0.05 mm; D = 0.01 mm.
</paragraph>
</caption>
<caption id="DF376662FFC7696BFF4F0D73FBFFF80E" ID-DOI="http://doi.org/10.5281/zenodo.10149316" ID-Zenodo-Dep="10149316" httpUri="https://zenodo.org/record/10149316/files/figure.png" pageId="14" pageNumber="53" startId="14.[189,232,1892,1918]" targetBox="[229,1359,265,1842]" targetPageId="14">
<paragraph id="8BF736EAFFC7696BFF4F0D73FBFFF80E" blockId="14.[189,1399,1891,2025]" pageId="14" pageNumber="53">
<emphasis id="B93CEAF8FFC7696BFF4F0D73FEDAF898" bold="true" box="[189,270,1892,1919]" pageId="14" pageNumber="53">Fig. 8.</emphasis>
<taxonomicName id="4C484D69FFC7696BFEEA0D72FDB4F898" authority="Antic" authorityName="Antic" authorityYear="2023" box="[280,608,1893,1919]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="14" pageNumber="53" phylum="Arthropoda" rank="species" species="devi" status="sp. nov.">
<emphasis id="B93CEAF8FFC7696BFEEA0D72FDC3F898" box="[280,535,1893,1919]" italics="true" pageId="14" pageNumber="53">Acanthophorella devi</emphasis>
Antić
</taxonomicName>
<taxonomicNameLabel id="A20F5783FFC7696BFD980D72FD13F898" box="[618,711,1893,1919]" pageId="14" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B93CEAF8FFC7696BFD230D73FD32F899" bold="true" box="[721,742,1892,1918]" pageId="14" pageNumber="53">A</emphasis>
. Paratype, ♀ (IZB), 
<emphasis id="B93CEAF8FFC7696BFC110D73FBCFF899" bold="true" box="[995,1051,1892,1918]" pageId="14" pageNumber="53">B–G</emphasis>
. Paratype, ♂ (IZB). 
<emphasis id="B93CEAF8FFC7696BFAEB0D73FAFAF899" bold="true" box="[1305,1326,1892,1918]" pageId="14" pageNumber="53">A</emphasis>
. Legpair 2, anterior view. 
<emphasis id="B93CEAF8FFC7696BFE4F0D9FFE04F845" bold="true" box="[445,464,1928,1954]" pageId="14" pageNumber="53">B</emphasis>
. Leg-pair 2, anterior view. 
<emphasis id="B93CEAF8FFC7696BFCE40D9FFCFEF845" bold="true" box="[790,810,1928,1954]" pageId="14" pageNumber="53">C</emphasis>
. Right leg 4, anterior view, white arrows indicate distoventral pads. 
<emphasis id="B93CEAF8FFC7696BFE6B0DBCFE78F822" bold="true" box="[409,428,1963,1989]" pageId="14" pageNumber="53">D</emphasis>
. Leg-pair 7, posterior view. 
<emphasis id="B93CEAF8FFC7696BFCF70DBCFCC3F822" bold="true" box="[773,791,1963,1989]" pageId="14" pageNumber="53">E</emphasis>
. Leg-pair 10, anterior view. 
<emphasis id="B93CEAF8FFC7696BFB830DBCFB55F822" bold="true" box="[1137,1153,1963,1989]" pageId="14" pageNumber="53">F</emphasis>
. Anterior gonopods, anterior view. 
<emphasis id="B93CEAF8FFC7696BFE970DD9FEA8F80F" bold="true" box="[357,380,1998,2024]" pageId="14" pageNumber="53">G</emphasis>
. Posterior gonopods, posterior view. Scale bars = 0.2 mm.
</paragraph>
</caption>
<caption id="DF376662FFC6696AFF4F0CECFBE7F80E" ID-DOI="http://doi.org/10.5281/zenodo.10149318" ID-Zenodo-Dep="10149318" httpUri="https://zenodo.org/record/10149318/files/figure.png" pageId="15" pageNumber="54" startId="15.[189,232,1787,1813]" targetBox="[282,1305,265,1742]" targetPageId="15">
<paragraph id="8BF736EAFFC6696AFF4F0CECFBE7F80E" blockId="15.[186,1399,1785,2025]" pageId="15" pageNumber="54">
<emphasis id="B93CEAF8FFC6696AFF4F0CECFEDEF8F2" bold="true" box="[189,266,1787,1813]" pageId="15" pageNumber="54">Fig. 9.</emphasis>
<taxonomicName id="4C484D69FFC6696AFEFD0CECFD9BF8F2" authority="Antic" authorityName="Antic" authorityYear="2023" box="[271,591,1787,1813]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="15" pageNumber="54" phylum="Arthropoda" rank="species" species="devi" status="sp. nov.">
<emphasis id="B93CEAF8FFC6696AFEFD0CECFDDEF8F2" box="[271,522,1787,1813]" italics="true" pageId="15" pageNumber="54">Acanthophorella devi</emphasis>
Antić
</taxonomicName>
<taxonomicNameLabel id="A20F5783FFC6696AFDA70CEBFD7AF8F1" box="[597,686,1788,1814]" pageId="15" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B93CEAF8FFC6696AFD410CECFD3DF8F2" bold="true" box="[691,745,1787,1813]" pageId="15" pageNumber="54">A–F</emphasis>
. Paratype, ♂ (IZB). 
<emphasis id="B93CEAF8FFC6696AFC270CECFBD2F8F2" bold="true" box="[981,1030,1787,1813]" pageId="15" pageNumber="54">G–I</emphasis>
. Paratype, ♀ (IZB). 
<emphasis id="B93CEAF8FFC6696AFB000CECFAD3F8F2" bold="true" box="[1266,1287,1787,1813]" pageId="15" pageNumber="54">A</emphasis>
. Anterior gonopods, anterior view. 
<emphasis id="B93CEAF8FFC6696AFE1D0D09FDD4F8DF" bold="true" box="[495,512,1822,1848]" pageId="15" pageNumber="54">B</emphasis>
. Anterior gonopods, anterodistal view. 
<emphasis id="B93CEAF8FFC6696AFC2F0D09FC24F8DF" bold="true" box="[989,1008,1822,1848]" pageId="15" pageNumber="54">C</emphasis>
. Anterior gonopods, distal view. 
<emphasis id="B93CEAF8FFC6696AFF4D0D55FF06F8BB" bold="true" box="[191,210,1858,1884]" pageId="15" pageNumber="54">D</emphasis>
. Anterior gonopods, lateral view. 
<emphasis id="B93CEAF8FFC6696AFD990D55FDA9F8BB" bold="true" box="[619,637,1858,1884]" pageId="15" pageNumber="54">E</emphasis>
. Anterior gonopods, lateral lamella and tuft, posterodistal view. 
<emphasis id="B93CEAF8FFC6696AFF4D0D72FF1BF898" bold="true" box="[191,207,1893,1919]" pageId="15" pageNumber="54">F</emphasis>
. Anterior gonopods, posterodistal view. 
<emphasis id="B93CEAF8FFC6696AFD4A0D73FD1BF899" bold="true" box="[696,719,1892,1918]" pageId="15" pageNumber="54">G</emphasis>
. Left vulva, lateral view. 
<emphasis id="B93CEAF8FFC6696AFBF80D72FBF5F898" bold="true" box="[1034,1057,1893,1919]" pageId="15" pageNumber="54">H</emphasis>
. Vulvae, distal view. 
<emphasis id="B93CEAF8FFC6696AFADB0D72FAE0F898" bold="true" box="[1321,1332,1893,1919]" pageId="15" pageNumber="54">I</emphasis>
. Left vulva, anterior view.Abbreviations: a = angiocoxite; ap = anterior process; b = bursa; ll = lateral lamella; mp = medial part; o = operculum; pl = posterior lobe; s = sternum; sa = synangiocoxal base; sl = sternal lamella; tf = angiocoxal tuft. Scale bars: A–D, F–I = 0.1 mm; E = 0.02 mm.
</paragraph>
</caption>
<caption id="DF376662FFD96975FF4F0CECFD24F80E" ID-DOI="http://doi.org/10.5281/zenodo.10149320" ID-Zenodo-Dep="10149320" httpUri="https://zenodo.org/record/10149320/files/figure.png" pageId="16" pageNumber="55" startId="16.[189,232,1787,1813]" targetBox="[293,1287,282,1718]" targetPageId="16">
<paragraph id="8BF736EAFFD96975FF4F0CECFD24F80E" blockId="16.[189,1399,1785,2027]" pageId="16" pageNumber="55">
<emphasis id="B93CEAF8FFD96975FF4F0CECFEF5F8F2" bold="true" box="[189,289,1787,1813]" pageId="16" pageNumber="55">Fig. 10.</emphasis>
<taxonomicName id="4C484D69FFD96975FEDD0CECFDABF8F2" authority="Antic" authorityName="Antic" authorityYear="2023" box="[303,639,1787,1813]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="16" pageNumber="55" phylum="Arthropoda" rank="species" species="devi" status="sp. nov.">
<emphasis id="B93CEAF8FFD96975FEDD0CECFDE6F8F2" box="[303,562,1787,1813]" italics="true" pageId="16" pageNumber="55">Acanthophorella devi</emphasis>
Antić
</taxonomicName>
<taxonomicNameLabel id="A20F5783FFD96975FD7F0CEBFD3AF8F1" box="[653,750,1788,1814]" pageId="16" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B93CEAF8FFD96975FD090CECFCE7F8F2" bold="true" box="[763,819,1787,1813]" pageId="16" pageNumber="55">A–E</emphasis>
. Paratype, ♂ (IZB). 
<emphasis id="B93CEAF8FFD96975FBB20CECFBA3F8F2" bold="true" box="[1088,1143,1787,1813]" pageId="16" pageNumber="55">F–G</emphasis>
. Paratype, ♀ (IZB). 
<emphasis id="B93CEAF8FFD96975FF4F0D09FF06F8DF" bold="true" box="[189,210,1822,1848]" pageId="16" pageNumber="55">A</emphasis>
. Anterior gonopods, anterior view. 
<emphasis id="B93CEAF8FFD96975FD7E0D09FD74F8DF" bold="true" box="[652,672,1822,1848]" pageId="16" pageNumber="55">B</emphasis>
. Anterior gonopods, posterior view. 
<emphasis id="B93CEAF8FFD96975FB940D09FBAFF8DF" bold="true" box="[1126,1147,1822,1848]" pageId="16" pageNumber="55">C</emphasis>
. Anterior gonopods, lateral view. 
<emphasis id="B93CEAF8FFD96975FEA80D55FEB9F8BB" bold="true" box="[346,365,1858,1884]" pageId="16" pageNumber="55">D</emphasis>
. Posterior gonopods, posterior view. 
<emphasis id="B93CEAF8FFD96975FCC30D55FC96F8BB" bold="true" box="[817,834,1858,1884]" pageId="16" pageNumber="55">E</emphasis>
. Left posterior gonopod, lateral view. 
<emphasis id="B93CEAF8FFD96975FAE70D55FAF1F8BB" bold="true" box="[1301,1317,1858,1884]" pageId="16" pageNumber="55">F</emphasis>
. Right vulva, distal view. 
<emphasis id="B93CEAF8FFD96975FE6C0D73FE60F899" bold="true" box="[414,436,1892,1918]" pageId="16" pageNumber="55">G</emphasis>
. Right vulva, lateral view. Abbreviations: a = angiocoxite; ap = anterior process; b = bursa; c = colpocoxite; ll = lateral lamella; mp = medial part; o = operculum; pl = posterior lobe; s = sternum; sa = synangiocoxal base; sl = sternal lamella; t = telopodite; tf = angiocoxal tuft. Drawings by DA (A–E) and MŠ (F–G). Scale bars = 0.2 mm.
</paragraph>
</caption>
<paragraph id="8BF736EAFFD86974FF4F080FFD83FD59" blockId="17.[189,1400,535,703]" pageId="17" pageNumber="56">
<smallCapsWord id="8D11A036FFD86974FF4F080FFEE6FDD7" baselines="555,555" box="[189,306,536,562]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Posterior" pageId="17" pageNumber="56">POSTERIOR</smallCapsWord>
<smallCapsWord id="8D11A036FFD86974FECC080AFE7BFDD7" baselines="555" box="[318,431,541,560]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="gonopods" pageId="17" pageNumber="56">GONOPODS</smallCapsWord>
(
<figureCitation id="13732A6FFFD86974FE360800FDF2FDD6" box="[452,550,535,561]" captionStart="Fig" captionStartId="14.[189,232,1892,1918]" captionTargetBox="[229,1359,265,1842]" captionTargetId="figure-17@14.[229,1359,265,1842]" captionTargetPageId="14" captionText="Fig. 8. Acanthophorella devi Antić sp. nov. A. Paratype, ♀ (IZB), B–G. Paratype, ♂ (IZB). A. Legpair 2, anterior view. B. Leg-pair 2, anterior view. C. Right leg 4, anterior view, white arrows indicate distoventral pads. D. Leg-pair 7, posterior view. E. Leg-pair 10, anterior view. F. Anterior gonopods, anterior view. G. Posterior gonopods, posterior view. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10149316" httpUri="https://zenodo.org/record/10149316/files/figure.png" pageId="17" pageNumber="56">Figs 8G</figureCitation>
, 
<figureCitation id="13732A6FFFD86974FDCB0800FD59FDD5" box="[569,653,535,562]" captionStart="Fig" captionStartId="16.[189,232,1787,1813]" captionTargetBox="[293,1287,282,1718]" captionTargetId="figure-17@16.[287,1300,265,1743]" captionTargetPageId="16" captionText="Fig. 10. Acanthophorella devi Antić sp. nov. A–E. Paratype, ♂ (IZB). F–G. Paratype, ♀ (IZB). A. Anterior gonopods, anterior view. B. Anterior gonopods, posterior view. C. Anterior gonopods, lateral view. D. Posterior gonopods, posterior view. E. Left posterior gonopod, lateral view. F. Right vulva, distal view. G. Right vulva, lateral view. Abbreviations: a = angiocoxite; ap = anterior process; b = bursa; c = colpocoxite; ll = lateral lamella; mp = medial part; o = operculum; pl = posterior lobe; s = sternum; sa = synangiocoxal base; sl = sternal lamella; t = telopodite; tf = angiocoxal tuft. Drawings by DA (A–E) and MŠ (F–G). Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10149320" httpUri="https://zenodo.org/record/10149320/files/figure.png" pageId="17" pageNumber="56">10D–E</figureCitation>
). Gonopodal sternum (s) wide, well-developed. Angiocoxites (a) positioned posteriorly, well-developed, slender, curved anterolaterad, longer than colpocoxites (c). Colpocoxites in anterior and posterior views wide in proximal half, then abruptly narrow to half their width in the distal half; fused with basal half of angiocoxites. Telopodites (t) small, rounded, placed posteriorly at base of angiocoxites.
</paragraph>
<paragraph id="8BF736EAFFD86974FF4F08F1FED5FCC4" blockId="17.[189,1398,741,803]" pageId="17" pageNumber="56">
<smallCapsWord id="8D11A036FFD86974FF4F08F1FF3FFD19" baselines="761,761" box="[189,235,742,768]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Leg" pageId="17" pageNumber="56">LEG</smallCapsWord>
<smallCapsWord id="8D11A036FFD86974FF0608FCFEF5FD19" baselines="761" box="[244,289,747,766]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="pair" pageId="17" pageNumber="56">PAIR</smallCapsWord>
2 
<smallCapsWord id="8D11A036FFD86974FEB008FCFE8CFD19" baselines="761" box="[322,344,747,766]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="in" pageId="17" pageNumber="56">IN</smallCapsWord>
<smallCapsWord id="8D11A036FFD86974FE9008FCFE14FD19" baselines="761" box="[354,448,747,766]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="females" pageId="17" pageNumber="56">FEMALES</smallCapsWord>
(
<figureCitation id="13732A6FFFD86974FE2108F2FDE6FCE7" box="[467,562,741,768]" captionStart="Fig" captionStartId="14.[189,232,1892,1918]" captionTargetBox="[229,1359,265,1842]" captionTargetId="figure-17@14.[229,1359,265,1842]" captionTargetPageId="14" captionText="Fig. 8. Acanthophorella devi Antić sp. nov. A. Paratype, ♀ (IZB), B–G. Paratype, ♂ (IZB). A. Legpair 2, anterior view. B. Leg-pair 2, anterior view. C. Right leg 4, anterior view, white arrows indicate distoventral pads. D. Leg-pair 7, posterior view. E. Leg-pair 10, anterior view. F. Anterior gonopods, anterior view. G. Posterior gonopods, posterior view. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10149316" httpUri="https://zenodo.org/record/10149316/files/figure.png" pageId="17" pageNumber="56">Fig. 8A</figureCitation>
). Coxae with distomesal protrusions covered with small tubercles and setae.
</paragraph>
<paragraph id="8BF736EAFFD86974FF4F095CFDE1FC29" blockId="17.[189,1400,842,974]" pageId="17" pageNumber="56">
<smallCapsWord id="8D11A036FFD86974FF4F095CFEC0FC84" baselines="862,862" box="[189,276,843,869]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Vulvae" pageId="17" pageNumber="56">VULVAE</smallCapsWord>
(
<figureCitation id="13732A6FFFD86974FED5095DFE74FC82" box="[295,416,842,869]" captionStart="Fig" captionStartId="15.[189,232,1787,1813]" captionTargetBox="[282,1305,265,1742]" captionTargetId="figure-12@15.[282,1305,265,1742]" captionTargetPageId="15" captionText="Fig. 9. Acanthophorella devi Antić sp. nov. A–F. Paratype, ♂ (IZB). G–I. Paratype, ♀ (IZB). A. Anterior gonopods, anterior view. B. Anterior gonopods, anterodistal view. C. Anterior gonopods, distal view. D. Anterior gonopods, lateral view. E. Anterior gonopods, lateral lamella and tuft, posterodistal view. F. Anterior gonopods, posterodistal view. G. Left vulva, lateral view. H. Vulvae, distal view. I. Left vulva, anterior view.Abbreviations: a = angiocoxite; ap = anterior process; b = bursa; ll = lateral lamella; mp = medial part; o = operculum; pl = posterior lobe; s = sternum; sa = synangiocoxal base; sl = sternal lamella; tf = angiocoxal tuft. Scale bars: A–D, F–I = 0.1 mm; E = 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.10149318" httpUri="https://zenodo.org/record/10149318/files/figure.png" pageId="17" pageNumber="56">Figs 9G–I</figureCitation>
, 
<figureCitation id="13732A6FFFD86974FE43095DFDD7FC83" box="[433,515,842,869]" captionStart="Fig" captionStartId="16.[189,232,1787,1813]" captionTargetBox="[293,1287,282,1718]" captionTargetId="figure-17@16.[287,1300,265,1743]" captionTargetPageId="16" captionText="Fig. 10. Acanthophorella devi Antić sp. nov. A–E. Paratype, ♂ (IZB). F–G. Paratype, ♀ (IZB). A. Anterior gonopods, anterior view. B. Anterior gonopods, posterior view. C. Anterior gonopods, lateral view. D. Posterior gonopods, posterior view. E. Left posterior gonopod, lateral view. F. Right vulva, distal view. G. Right vulva, lateral view. Abbreviations: a = angiocoxite; ap = anterior process; b = bursa; c = colpocoxite; ll = lateral lamella; mp = medial part; o = operculum; pl = posterior lobe; s = sternum; sa = synangiocoxal base; sl = sternal lamella; t = telopodite; tf = angiocoxal tuft. Drawings by DA (A–E) and MŠ (F–G). Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10149320" httpUri="https://zenodo.org/record/10149320/files/figure.png" pageId="17" pageNumber="56">10F–G</figureCitation>
). Slightly longer than wide. Operculum (o) well-developed, bilobed, with 5+5 setae (4+4 lateral shorter setae and 1+1 mesal longer setae). Bursa (b) with strongly thickened anteroproximal lips on which the operculum rests. Lateral valve with six setae only in anterior part, mesal valve with 11 or 12 setae.
</paragraph>
</subSubSection>
<subSubSection id="C3526561FFD86974FF4F09E2FC12FA65" pageId="17" pageNumber="56" type="biology_ecology">
<paragraph id="8BF736EAFFD86974FF4F09E2FE6BFBE8" blockId="17.[189,447,1013,1039]" box="[189,447,1013,1039]" pageId="17" pageNumber="56">
<heading id="D0BF8186FFD86974FF4F09E2FE6BFBE8" bold="true" box="[189,447,1013,1039]" fontSize="11" level="3" pageId="17" pageNumber="56" reason="3">
<emphasis id="B93CEAF8FFD86974FF4F09E2FE6BFBE8" bold="true" box="[189,447,1013,1039]" pageId="17" pageNumber="56">Locality and ecology</emphasis>
</heading>
</paragraph>
<paragraph id="8BF736EAFFD86974FF4F0E33FCDCFB63" blockId="17.[188,1399,1059,1157]" pageId="17" pageNumber="56">
Devis Khvreli is a small horizontal cave 
<quantity id="4CB09B0FFFD86974FD650E34FD2AFBD9" box="[663,766,1059,1086]" metricMagnitude="1" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="17" pageNumber="56" unit="m" value="55.0" valueMax="60.0" valueMin="50.0">50–60 m</quantity>
long, but as the entrance is relatively small, the whole cave being dark. The cave consists of two parts, a first dry part connected to a second wet part by a narrow passage. This cave is poor in speleothems.
</paragraph>
<paragraph id="8BF736EAFFD86974FF4F0EB5FD78FAC1" blockId="17.[189,1399,1186,1318]" pageId="17" pageNumber="56">
The specimens were found crawling on the substrate or on dead wood in the second, dark and wet part of the cave. Besides the new species, this cave is inhabited by two other millipedes, the troglobiotic 
<taxonomicName id="4C484D69FFD86974FF4F0EFDFD4FFAE4" authority="Antic &amp; Reip, 2020" authorityName="Antic &amp; Reip" authorityYear="2020" box="[189,667,1257,1283]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="17" pageNumber="56" phylum="Arthropoda" rank="species" species="gioi">
<emphasis id="B93CEAF8FFD86974FF4F0EFDFE4DFAE4" box="[189,409,1258,1283]" italics="true" pageId="17" pageNumber="56">Leucogeorgia gioi</emphasis>
<bibRefCitation id="EFD94B1BFFD86974FE560EFEFD4FFAE4" author="Antic D. Z. &amp; Reip H. S." box="[420,667,1257,1283]" pageId="17" pageNumber="56" pagination="1 - 106" refId="ref14480" refString="Antic D. Z. &amp; Reip H. S. 2020. The millipede genus Leucogeorgia Verhoeff, 1930 in the Caucasus, with descriptions of eleven new species, erection of a new monotypic genus and notes on the tribe Leucogeorgiini (Diplopoda: Julida: Julidae). European Journal of Taxonomy 713: 1 - 106. https: // doi. org / 10.5852 / ejt. 2020.713" type="journal article" year="2020">Antić &amp; Reip, 2020</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="EFD94B1BFFD86974FD5D0EFEFC76FAE4" author="Antic D. Z. &amp; Reip H. S." box="[687,930,1257,1283]" pageId="17" pageNumber="56" pagination="1 - 106" refId="ref14480" refString="Antic D. Z. &amp; Reip H. S. 2020. The millipede genus Leucogeorgia Verhoeff, 1930 in the Caucasus, with descriptions of eleven new species, erection of a new monotypic genus and notes on the tribe Leucogeorgiini (Diplopoda: Julida: Julidae). European Journal of Taxonomy 713: 1 - 106. https: // doi. org / 10.5852 / ejt. 2020.713" type="journal article" year="2020">Antić &amp; Reip 2020</bibRefCitation>
) and the troglophilous 
<taxonomicName id="4C484D69FFD86974FB340EFEFE2DFAC1" authority="Golovatch, 1976" authorityName="Golovatch" authorityYear="1976" class="Diplopoda" family="Glomeridae" genus="Trachysphaera" kingdom="Animalia" order="Glomerida" pageId="17" pageNumber="56" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="B93CEAF8FFD86974FB340EFEFEF9FAC1" italics="true" pageId="17" pageNumber="56">Trachysphaera orientalis</emphasis>
Golovatch, 1976
</taxonomicName>
(unpubl. data).
</paragraph>
<paragraph id="8BF736EAFFD86974FF4F0F52FC12FA65" blockId="17.[189,1398,1349,1410]" pageId="17" pageNumber="56">
Based on the ecology, the completely unpigmented body and the reduced number of pale brownish ommatidia, 
<taxonomicName id="4C484D69FFD86974FEBB0F7FFE4FFA65" authority="Antić &amp; Šević &amp; Barjadze &amp; Makarov, 2023" authorityName="Antić &amp; Šević &amp; Barjadze &amp; Makarov" authorityYear="2023" box="[329,411,1384,1410]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="17" pageNumber="56" phylum="Arthropoda" rank="species" species="devi" status="sp. nov.">
<emphasis id="B93CEAF8FFD86974FEBB0F7FFE4FFA65" box="[329,411,1384,1410]" italics="true" pageId="17" pageNumber="56">A. devi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20F5783FFD86974FE500F7FFE29FA65" box="[418,509,1384,1410]" pageId="17" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
can be considered as a neotroglobiont.
</paragraph>
</subSubSection>
<subSubSection id="C3526561FFD86974FF4F0FB7FDB8F9EB" pageId="17" pageNumber="56" type="distribution">
<paragraph id="8BF736EAFFD86974FF4F0FB7FE82FA5D" blockId="17.[189,342,1440,1466]" box="[189,342,1440,1466]" pageId="17" pageNumber="56">
<heading id="D0BF8186FFD86974FF4F0FB7FE82FA5D" bold="true" box="[189,342,1440,1466]" fontSize="11" level="3" pageId="17" pageNumber="56" reason="3">
<emphasis id="B93CEAF8FFD86974FF4F0FB7FE82FA5D" bold="true" box="[189,342,1440,1466]" pageId="17" pageNumber="56">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BF736EAFFD86974FF4F0FD8FDB8F9EB" blockId="17.[187,1398,1487,1548]" pageId="17" pageNumber="56">
A Georgian endemic known only from its 
<typeStatus id="54F38848FFD86974FD530FC7FD00FA0D" box="[673,724,1488,1514]" pageId="17" pageNumber="56">type</typeStatus>
locality, Devis Khvreli Cave, near Rgani village (Chiatura Municipality) (
<figureCitation id="13732A6FFFD86974FE9F0FE5FE10F9EB" box="[365,452,1522,1548]" captionStart="Fig" captionStartId="35.[189,232,1998,2024]" captionTargetBox="[189,1398,1153,1959]" captionTargetId="figure-355@35.[189,1398,1150,1959]" captionTargetPageId="35" captionText="Fig. 22. Distribution of the genus Acanthophorella Antić &amp; Makarov, 2016." figureDoi="http://doi.org/10.5281/zenodo.10149347" httpUri="https://zenodo.org/record/10149347/files/figure.png" pageId="17" pageNumber="56">Fig. 22</figureCitation>
, violet circle).
</paragraph>
</subSubSection>
</treatment>
</document>