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<mods:title id="6D4DCAA24C46A20F121B7BB3459DDAF0">Two new species of Hetereleotris (Perciformes: Gobiidae) from the Red Sea</mods:title>
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<subSubSection id="C37A651A5F7EFFB2FF78F995FBD1F938" box="[151,1144,1660,1687]" pageId="2" pageNumber="503" type="nomenclature">
<paragraph id="8BDF36915F7EFFB2FF78F995FBD1F938" blockId="2.[151,1144,1660,1687]" box="[151,1144,1660,1687]" pageId="2" pageNumber="503">
<heading id="D09781FD5F7EFFB2FF78F995FBD1F938" bold="true" box="[151,1144,1660,1687]" fontSize="11" level="1" pageId="2" pageNumber="503" reason="1">
<emphasis id="B914EA835F7EFFB2FF78F995FBD1F938" bold="true" box="[151,1144,1660,1687]" pageId="2" pageNumber="503">
Key to Described Species of
<taxonomicName id="4C604D125F7EFFB2FE17F995FD3DF939" ID-CoL="62V3V" ID-ENA="2067727" authorityName="Bleeker" authorityYear="1874" box="[504,660,1660,1687]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="503" phylum="Chordata" rank="genus">
<emphasis id="B914EA835F7EFFB2FE17F995FD3DF939" bold="true" box="[504,660,1660,1687]" italics="true" pageId="2" pageNumber="503">Hetereleotris</emphasis>
</taxonomicName>
(modified from
<bibRefCitation id="EFF14B605F7EFFB2FC8FF995FBC6F938" author="Hoese, D. F. &amp; Larson, H. K." box="[864,1135,1660,1686]" pageId="2" pageNumber="503" pagination="1 - 16" refId="ref13143" refString="Hoese, D. F. &amp; Larson, H. K. (2005) Description of two new species of Hetereleotris from the south Pacific, with a revised key to species and synonymization of the genus Pascua with Hetereleotris. Zootaxa, 1096 (1), 1 - 16. https: // doi. org / 10.11646 / zootaxa. 1096.1.1" type="journal article" year="2005">Hoese &amp; Larson 2005</bibRefCitation>
)
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C37A651A5F7EFFB3FF78F928FA35F980" lastPageId="3" lastPageNumber="504" pageId="2" pageNumber="503" type="key">
<keyStep id="30942E345F7EFFB2FF78F928FA35F95D" pageId="2" pageNumber="503">
<paragraph id="8BDF36915F7EFFB2FF78F928FA35F978" blockId="2.[151,1436,1729,2044]" box="[151,1436,1729,1751]" pageId="2" pageNumber="503">
<keyLead id="309195A45F7EFFB2FF78F928FA35F978" box="[151,1436,1729,1751]" pageId="2" pageNumber="503">
1a. An enlarged tentacle over eye (western Indian Ocean)..............................................
<taxonomicName id="4C604D125F7EFFB2FAFCF928FA35F978" ID-CoL="3L7J9" baseAuthorityName="Smith" baseAuthorityYear="1958" box="[1299,1436,1729,1750]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="503" phylum="Chordata" rank="species" species="tentaculata">
<emphasis id="B914EA835F7EFFB2FAFCF928FA35F978" box="[1299,1436,1729,1750]" italics="true" pageId="2" pageNumber="503">H. tentaculata</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BDF36915F7EFFB2FF78F937FA35F95D" blockId="2.[151,1436,1729,2044]" box="[151,1436,1758,1780]" pageId="2" pageNumber="503">
<keyLead id="309195A45F7EFFB2FF78F937FA35F95D" box="[151,1436,1758,1780]" pageId="2" pageNumber="503">1b. No tentacle over eye................................................................................... 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="30942E345F7EFFB2FF78F912FA35F881" pageId="2" pageNumber="503">
<paragraph id="8BDF36915F7EFFB2FF78F912FA35F8BE" blockId="2.[151,1436,1729,2044]" box="[151,1436,1787,1809]" pageId="2" pageNumber="503">
<keyLead id="309195A45F7EFFB2FF78F912FA35F8BE" box="[151,1436,1787,1809]" pageId="2" pageNumber="503">2a. Body densely scaled or at least with scales on caudal peduncle................................................. 3</keyLead>
</paragraph>
<paragraph id="8BDF36915F7EFFB2FF78F8F0FA35F881" blockId="2.[151,1436,1729,2044]" box="[151,1436,1817,1839]" pageId="2" pageNumber="503">
<keyLead id="309195A45F7EFFB2FF78F8F0FA35F881" box="[151,1436,1817,1839]" pageId="2" pageNumber="503">2b. Body completely scaleless............................................................................. 15</keyLead>
</paragraph>
</keyStep>
<keyStep id="30942E345F7EFFB2FF78F8DFFA35F8C6" pageId="2" pageNumber="503">
<paragraph id="8BDF36915F7EFFB2FF78F8DFFA35F8E5" blockId="2.[151,1436,1729,2044]" box="[151,1436,1846,1868]" pageId="2" pageNumber="503">
<keyLead id="309195A45F7EFFB2FF78F8DFFA35F8E5" box="[151,1436,1846,1868]" pageId="2" pageNumber="503">3a. Scales ctenoid, at least posteriorly on body................................................................. 4</keyLead>
</paragraph>
<paragraph id="8BDF36915F7EFFB2FF78F8BAFA35F8C6" blockId="2.[151,1436,1729,2044]" box="[151,1436,1875,1897]" pageId="2" pageNumber="503">
<keyLead id="309195A45F7EFFB2FF78F8BAFA35F8C6" box="[151,1436,1875,1897]" pageId="2" pageNumber="503">3b. Scales entirely cycloid................................................................................. 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="30942E345F7EFFB2FF78F898FA35F80A" pageId="2" pageNumber="503">
<paragraph id="8BDF36915F7EFFB2FF78F898FA35F829" blockId="2.[151,1436,1729,2044]" box="[151,1436,1905,1927]" pageId="2" pageNumber="503">
<keyLead id="309195A45F7EFFB2FF78F898FA35F829" box="[151,1436,1905,1927]" pageId="2" pageNumber="503">4a. Body with scales restricted to caudal peduncle; no head pores.................................................. 5</keyLead>
</paragraph>
<paragraph id="8BDF36915F7EFFB2FF78F867FA35F80A" blockId="2.[151,1436,1729,2044]" box="[151,1436,1934,1956]" pageId="2" pageNumber="503">
<keyLead id="309195A45F7EFFB2FF78F867FA35F80A" box="[151,1436,1934,1956]" pageId="2" pageNumber="503">4b. Body completely or partly scaled, longitudinal scale count 1030; head pores present............................... 6</keyLead>
</paragraph>
</keyStep>
<paragraph id="8BDF36915F7EFFB2FF78F842FA35F870" blockId="2.[151,1436,1729,2044]" pageId="2" pageNumber="503">
<keyStep id="30942E345F7EFFB2FF78F842FA35F870" pageId="2" pageNumber="503">
<keyLead id="309195A45F7EFFB2FF78F842FA35F870" pageId="2" pageNumber="503">
5a. Body with 45 rows of scales restricted to caudal peduncle; operculum with 2 spines dorsally; body dusky grey with diffuse dark bars on upper half (western Indian Ocean).......................................................
<taxonomicName id="4C604D125F7EFFB2FAA9F820FA35F870" ID-CoL="3L7HP" baseAuthorityName="Hoese &amp; Winterbottom" baseAuthorityYear="1979" box="[1350,1436,1993,2014]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="503" phylum="Chordata" rank="species" species="apora">
<emphasis id="B914EA835F7EFFB2FAA9F820FA35F870" box="[1350,1436,1993,2014]" italics="true" pageId="2" pageNumber="503">H. apora</emphasis>
</taxonomicName>
</keyLead>
</keyStep>
</paragraph>
<paragraph id="8BDF36915F7EFFB3FF78F80FFA35FF47" blockId="2.[151,1436,1729,2044]" lastBlockId="3.[151,1437,153,1583]" lastPageId="3" lastPageNumber="504" pageId="2" pageNumber="503">
<keyLead id="309195A45F7EFFB3FF78F80FFF57FF01" lastPageId="3" lastPageNumber="504" pageId="2" pageNumber="503">5b. Body with squamation reduced to a few scales on caudal peduncle at caudal-fin base; operculum without spines; body yellowish</keyLead>
orange, with five vertical brown bars (Red Sea).........................................
<taxonomicName id="4C604D125F7FFFB3FB2CFF73FAE0FF01" authority="Kovačić &amp; Bogorodsky &amp; Mal, 2019" authorityName="Kovačić &amp; Bogorodsky &amp; Mal" authorityYear="2019" box="[1219,1353,154,175]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="504" phylum="Chordata" rank="species" species="aurantiaca" status="sp. nov.">
<emphasis id="B914EA835F7FFFB3FB2CFF73FB7DFF01" box="[1219,1236,154,175]" italics="true" pageId="3" pageNumber="504">H</emphasis>
.
<emphasis id="B914EA835F7FFFB3FB0FFF73FAE0FF01" box="[1248,1353,154,175]" italics="true" pageId="3" pageNumber="504">aurantiaca</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22757F85F7FFFB3FABFFF73FA35FF01" box="[1360,1436,154,175]" pageId="3" pageNumber="504" rank="species">
<emphasis id="B914EA835F7FFFB3FABFFF73FA35FF01" bold="true" box="[1360,1436,154,175]" pageId="3" pageNumber="504">
sp.
<date id="FFDE10515F7FFFB3FA9DFF73FA35FF01" box="[1394,1436,154,175]" pageId="3" pageNumber="504">nov.</date>
</emphasis>
</taxonomicNameLabel>
<keyStep id="30942E345F7FFFB3FF78FF5EFA35FF47" pageId="3" pageNumber="504">
<keyLead id="309195A45F7FFFB3FF78FF5EFA35FF47" pageId="3" pageNumber="504">
6a. Scales restricted to posterior body, longitudinal scale count 1013; six suborbital transverse papillae rows (
<collectingCountry id="F37776015F7FFFB3FB0CFF5EFAEAFF63" box="[1251,1347,183,205]" name="Mauritius" pageId="3" pageNumber="504">Mauritius</collectingCountry>
)...............................................................................................
<taxonomicName id="4C604D125F7FFFB3FAF9FF3CFA35FF47" ID-CoL="3L7HZ" authorityName="Gill" authorityYear="1998" box="[1302,1436,212,234]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="504" phylum="Chordata" rank="species" species="georgegilli">
<emphasis id="B914EA835F7FFFB3FAF9FF3CFA35FF47" box="[1302,1436,212,234]" italics="true" pageId="3" pageNumber="504">H. georgegilli</emphasis>
</taxonomicName>
</keyLead>
</keyStep>
</paragraph>
<paragraph id="8BDF36915F7FFFB3FF78FF18FA35FEA9" blockId="3.[151,1437,153,1583]" box="[151,1436,241,263]" pageId="3" pageNumber="504">
<keyLead id="309195A45F7FFFB3FF78FF18FA35FEA9" box="[151,1436,241,263]" pageId="3" pageNumber="504">6b. Scales entire on body, longitudinal scale count 2530; four suborbital transverse papillae rows........................ 7</keyLead>
</paragraph>
<keyStep id="30942E345F7FFFB3FF78FEE6FA35FE19" pageId="3" pageNumber="504">
<paragraph id="8BDF36915F7FFFB3FF78FEE6FA35FEF0" blockId="3.[151,1437,153,1583]" pageId="3" pageNumber="504">
<keyLead id="309195A45F7FFFB3FF78FEE6FA35FEF0" pageId="3" pageNumber="504">
7a. Second dorsal-fin rays I,9; anal-fin rays I,8; pelvic-fin rays I,5; body scales mainly ctenoid; pelvic fins usually united to form a disc; pore β present in anterior oculoscapular head canal (lateral canal pore LC1 of
<bibRefCitation id="EFF14B605F7FFFB3FBE3FEC4FB2BFEEC" author="Hoese, D. F." box="[1036,1154,300,322]" pageId="3" pageNumber="504" pagination="1 - 25" refId="ref13090" refString="Hoese, D. F. (1986) Descriptions of two new species of Hetereleotris (Pisces: Gobiidae) from the western Indian Ocean, with discussion of related species. J. L. B. Smith Institute of Ichthyology, Special Publication, 41, 1 - 25." type="journal article" year="1986">Hoese 1986</bibRefCitation>
); belly naked (western Indian Ocean)..................................................................................
<taxonomicName id="4C604D125F7FFFB3FB12FEA3FA35FEF0" authorityName="Smith" authorityYear="1958" box="[1277,1436,329,351]" family="Gobiidae" genus="Satulinus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="504" phylum="Chordata" rank="species" species="zanzibarensis">
<emphasis id="B914EA835F7FFFB3FB12FEA3FA35FEF0" box="[1277,1436,329,351]" italics="true" pageId="3" pageNumber="504">H. zanzibarensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BDF36915F7FFFB3FF78FE8EFA35FE19" blockId="3.[151,1437,153,1583]" pageId="3" pageNumber="504">
<keyLead id="309195A45F7FFFB3FF78FE8EFA35FE19" pageId="3" pageNumber="504">
7b. Second dorsal-fin rays I,10; anal-fin rays I,9; pelvic-fin rays usually I,4, fifth ray sometimes present as a rudiment; ctenoid scales from caudal-fin base to below middle of second dorsal fin; pelvic fins separated; pore β absent in anterior oculoscapular head canal; belly scaled (western Indian Ocean)......................................................
<taxonomicName id="4C604D125F7FFFB3FAD5FE4BFA35FE19" ID-CoL="3L7JB" authorityName="Hoese" authorityYear="1986" box="[1338,1436,418,439]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="504" phylum="Chordata" rank="species" species="vinsoni">
<emphasis id="B914EA835F7FFFB3FAD5FE4BFA35FE19" box="[1338,1436,418,439]" italics="true" pageId="3" pageNumber="504">H. vinsoni</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30942E345F7FFFB3FF78FE56FA35FE5F" pageId="3" pageNumber="504">
<paragraph id="8BDF36915F7FFFB3FF78FE56FA35FE7A" blockId="3.[151,1437,153,1583]" box="[151,1436,447,469]" pageId="3" pageNumber="504">
<keyLead id="309195A45F7FFFB3FF78FE56FA35FE7A" box="[151,1436,447,469]" pageId="3" pageNumber="504">8a. Black spot subequal to pupil diameter above rear edge of opercle............................................... 9</keyLead>
</paragraph>
<paragraph id="8BDF36915F7FFFB3FF78FE35FA35FE5F" blockId="3.[151,1437,153,1583]" box="[151,1436,476,498]" pageId="3" pageNumber="504">
<keyLead id="309195A45F7FFFB3FF78FE35FA35FE5F" box="[151,1436,476,498]" pageId="3" pageNumber="504">8a. No black spot above rear edge of opercle................................................................. 10</keyLead>
</paragraph>
</keyStep>
<keyStep id="30942E345F7FFFB3FF78FE13FA35FD0C" pageId="3" pageNumber="504">
<paragraph id="8BDF36915F7FFFB3FF78FE13FA34FDE7" blockId="3.[151,1437,153,1583]" pageId="3" pageNumber="504">
<keyLead id="309195A45F7FFFB3FF78FE13FA34FDE7" pageId="3" pageNumber="504">
9a. Second dorsal-fin rays I,1213; anal-fin rays I,1112; pectoral-fin rays 15; a black spot dorsally at posterior end of caudal peduncle sometimes present; body scales small, longitudinal scale count 4052; body slender, depth at anal-fin origin 6.77.1 in SL (north-western Indian Ocean).............................................................
<taxonomicName id="4C604D125F7FFFB3FAF7FDDCFA34FDE7" ID-CoL="3L7HR" authorityName="Tortonese" authorityYear="1976" box="[1304,1437,564,586]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="504" phylum="Chordata" rank="species" species="bipunctata">
<emphasis id="B914EA835F7FFFB3FAF7FDDCFA34FDE7" box="[1304,1437,564,586]" italics="true" pageId="3" pageNumber="504">H. bipunctata</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BDF36915F7FFFB3FF78FDB8FA35FD0C" blockId="3.[151,1437,153,1583]" pageId="3" pageNumber="504">
<keyLead id="309195A45F7FFFB3FF78FDB8FA35FD0C" pageId="3" pageNumber="504">
9b. Second dorsal-fin rays I,11; anal-fin rays I,10; pectoral-fin rays 17; no black spot dorsally at end of caudal peduncle; body scales large, longitudinal scale count 2830; body more robust, depth at anal-fin origin 5.36.3 in SL (western Indian Ocean)........................................................................................
<taxonomicName id="4C604D125F7FFFB3FAFEFD64FA35FD0C" ID-CoL="3L7J3" authorityName="Hoese" authorityYear="1986" box="[1297,1436,653,674]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="504" phylum="Chordata" rank="species" species="margaretae">
<emphasis id="B914EA835F7FFFB3FAFEFD64FA35FD0C" box="[1297,1436,653,674]" italics="true" pageId="3" pageNumber="504">H. margaretae</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30942E345F7FFFB3FF78FD43FA35FD57" pageId="3" pageNumber="504">
<paragraph id="8BDF36915F7FFFB3FF78FD43FA35FD72" blockId="3.[151,1437,153,1583]" pageId="3" pageNumber="504">
<keyLead id="309195A45F7FFFB3FF78FD43FA35FD72" pageId="3" pageNumber="504">
10a. Scales restricted to caudal peduncle, longitudinal scale count 79; body covered with small dark spots (western Indian Ocean)......................................................................................
<taxonomicName id="4C604D125F7FFFB3FB1DFD2EFA35FD72" ID-CoL="3L7J4" baseAuthorityName="Smith" baseAuthorityYear="1958" box="[1266,1436,711,732]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="504" phylum="Chordata" rank="species" species="nebulofasciata">
<emphasis id="B914EA835F7FFFB3FB1DFD2EFA35FD72" box="[1266,1436,711,732]" italics="true" pageId="3" pageNumber="504">H. nebulofasciata</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BDF36915F7FFFB3FF78FD0DFA35FD57" blockId="3.[151,1437,153,1583]" box="[151,1436,740,762]" pageId="3" pageNumber="504">
<keyLead id="309195A45F7FFFB3FF78FD0DFA35FD57" box="[151,1436,740,762]" pageId="3" pageNumber="504">10b. Body variably scaled but not restricted to caudal peduncle, longitudinal scale count 1448; body without spots.......... 11</keyLead>
</paragraph>
</keyStep>
<keyStep id="30942E345F7FFFB3FF78FCEBFA35FCFF" pageId="3" pageNumber="504">
<paragraph id="8BDF36915F7FFFB3FF78FCEBFA35FC9A" blockId="3.[151,1437,153,1583]" pageId="3" pageNumber="504">
<keyLead id="309195A45F7FFFB3FF78FCEBFA35FC9A" pageId="3" pageNumber="504">11a. Body with obvious brown bar below first dorsal fin; head with brown band across interorbital area, continuing diagonally from eye to corner of opercle............................................................................... 12</keyLead>
</paragraph>
<paragraph id="8BDF36915F7FFFB3FF78FCD5FA35FCFF" blockId="3.[151,1437,153,1583]" box="[151,1436,828,850]" pageId="3" pageNumber="504">
<keyLead id="309195A45F7FFFB3FF78FCD5FA35FCFF" box="[151,1436,828,850]" pageId="3" pageNumber="504">11b. Body without bar below first dorsal fin; no band on head..................................................... 14</keyLead>
</paragraph>
</keyStep>
<keyStep id="30942E345F7FFFB3FF78FCB3FA35FC69" pageId="3" pageNumber="504">
<paragraph id="8BDF36915F7FFFB3FF78FCB3FA35FC22" blockId="3.[151,1437,153,1583]" pageId="3" pageNumber="504">
<keyLead id="309195A45F7FFFB3FF78FCB3FA35FC22" pageId="3" pageNumber="504">
12a. Second dorsal-fin rays I,11; head pores absent; scale pattern a long triangle tapering to nearly pectoral-fin base, longitudinal scale count 29; moderately broad brown bar on caudal-fin base (
<collectingRegion id="49A4F8735F7FFFB3FCA4FC9EFC34FC23" box="[843,925,887,909]" country="Egypt" name="Al Bahr al Ahmar" pageId="3" pageNumber="504">Red Sea</collectingRegion>
)....................
<taxonomicName id="4C604D125F7FFFB3FB4CFC9EFAE0FC22" authority="Kovačić &amp; Bogorodsky &amp; Mal, 2019" authorityName="Kovačić &amp; Bogorodsky &amp; Mal" authorityYear="2019" box="[1187,1353,887,908]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="504" phylum="Chordata" rank="species" species="semisquamata" status="sp. nov.">
<emphasis id="B914EA835F7FFFB3FB4CFC9EFAE0FC22" box="[1187,1353,887,908]" italics="true" pageId="3" pageNumber="504">H. semisquamata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22757F85F7FFFB3FABFFC9EFA35FC22" box="[1360,1436,887,908]" pageId="3" pageNumber="504" rank="species">
<emphasis id="B914EA835F7FFFB3FABFFC9EFA35FC22" bold="true" box="[1360,1436,887,908]" pageId="3" pageNumber="504">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BDF36915F7FFFB3FF78FC7DFA35FC69" blockId="3.[151,1437,153,1583]" pageId="3" pageNumber="504">
<keyLead id="309195A45F7FFFB3FF78FC7DFA35FC69" pageId="3" pageNumber="504">12b. Second dorsal-fin rays I,1213; head pores present; scale pattern not tapering, covering most of body, longitudinal scale count 3248; no bar on caudal-fin base........................................................................ 13</keyLead>
</paragraph>
</keyStep>
<keyStep id="30942E345F7FFFB3FF78FC26FA35FB92" pageId="3" pageNumber="504">
<paragraph id="8BDF36915F7FFFB3FF78FC26FA34FBAF" blockId="3.[151,1437,153,1583]" pageId="3" pageNumber="504">
<keyLead id="309195A45F7FFFB3FF78FC26FA34FBAF" pageId="3" pageNumber="504">
13a. Scales moderately large, longitudinal scale count 3233; head with brown stripe from eye to middle of upper jaw (western Indian Ocean).................................................................................
<taxonomicName id="4C604D125F7FFFB3FAD3FC04FA34FBAF" ID-CoL="3L7J2" authorityName="Smith" authorityYear="1958" box="[1340,1437,1004,1026]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="504" phylum="Chordata" rank="species" species="kenyae">
<emphasis id="B914EA835F7FFFB3FAD3FC04FA34FBAF" box="[1340,1437,1004,1026]" italics="true" pageId="3" pageNumber="504">H. kenyae</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BDF36915F7FFFB3FF78FBE0FA35FB92" blockId="3.[151,1437,153,1583]" pageId="3" pageNumber="504">
<keyLead id="309195A45F7FFFB3FF78FBE0FA35FB92" pageId="3" pageNumber="504">
13b. Scales small, longitudinal scale count 3548; no stripe between eye and upper jaw (western to central Indian Ocean)...................................................................................................
<taxonomicName id="4C604D125F7FFFB3FAD0FBCEFA35FB92" ID-CoL="3L7JF" baseAuthorityName="Fowler" baseAuthorityYear="1934" box="[1343,1436,1063,1084]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="504" phylum="Chordata" rank="species" species="zonata">
<emphasis id="B914EA835F7FFFB3FAD0FBCEFA35FB92" box="[1343,1436,1063,1084]" italics="true" pageId="3" pageNumber="504">H. zonata</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30942E345F7FFFB3FF78FBADFA34FB1F" pageId="3" pageNumber="504">
<paragraph id="8BDF36915F7FFFB3FF78FBADFA34FBD8" blockId="3.[151,1437,153,1583]" pageId="3" pageNumber="504">
<keyLead id="309195A45F7FFFB3FF78FBADFA34FBD8" pageId="3" pageNumber="504">
14a. Second dorsal-fin rays usually I,911; anal-fin rays usually I,89; longitudinal scale count 3225; head pores present (western Indian Ocean to
<collectingCountry id="F37776015F7FFFB3FE6EFB8BFE10FBD9" box="[385,441,1122,1143]" name="Japan" pageId="3" pageNumber="504">Japan</collectingCountry>
)..........................................................................
<taxonomicName id="4C604D125F7FFFB3FAD7FB8BFA34FBD8" authorityName="Fowler" authorityYear="1946" box="[1336,1437,1121,1143]" family="Gobiidae" genus="Riukiuia" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="504" phylum="Chordata" rank="species" species="poecila">
<emphasis id="B914EA835F7FFFB3FAD7FB8BFA34FBD8" box="[1336,1437,1121,1143]" italics="true" pageId="3" pageNumber="504">H. poecila</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BDF36915F7FFFB3FF78FB96FA34FB1F" blockId="3.[151,1437,153,1583]" pageId="3" pageNumber="504">
<keyLead id="309195A45F7FFFB3FF78FB96FA34FB1F" pageId="3" pageNumber="504">
14b. Second dorsal-fin rays usually I,12; anal-fin rays usually I,11; scales restricted to posterior body, longitudinal scale count 1415; head pores absent (
<collectingCountry id="F37776015F7FFFB3FE0DFB75FDB2FB1C" box="[482,539,1180,1202]" name="Japan" pageId="3" pageNumber="504">Japan</collectingCountry>
)...................................................................
<taxonomicName id="4C604D125F7FFFB3FAA3FB74FA34FB1F" ID-CoL="3L7HY" authorityName="Shibukawa" authorityYear="2010" box="[1356,1437,1180,1202]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="504" phylum="Chordata" rank="species" species="exilis">
<emphasis id="B914EA835F7FFFB3FAA3FB74FA34FB1F" box="[1356,1437,1180,1202]" italics="true" pageId="3" pageNumber="504">H. exilis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30942E345F7FFFB3FF78FB50FA35FB42" pageId="3" pageNumber="504">
<paragraph id="8BDF36915F7FFFB3FF78FB50FA35FB61" blockId="3.[151,1437,153,1583]" box="[151,1436,1209,1231]" pageId="3" pageNumber="504">
<keyLead id="309195A45F7FFFB3FF78FB50FA35FB61" box="[151,1436,1209,1231]" pageId="3" pageNumber="504">15a. Head pores present................................................................................... 16</keyLead>
</paragraph>
<paragraph id="8BDF36915F7FFFB3FF78FB3EFA35FB42" blockId="3.[151,1437,153,1583]" box="[151,1436,1239,1261]" pageId="3" pageNumber="504">
<keyLead id="309195A45F7FFFB3FF78FB3EFA35FB42" box="[151,1436,1239,1261]" pageId="3" pageNumber="504">15b. Head pores absent.................................................................................... 17</keyLead>
</paragraph>
</keyStep>
<keyStep id="30942E345F7FFFB3FF78FB1DFA34FACF" pageId="3" pageNumber="504">
<paragraph id="8BDF36915F7FFFB3FF78FB1DFA35FA89" blockId="3.[151,1437,153,1583]" pageId="3" pageNumber="504">
<keyLead id="309195A45F7FFFB3FF78FB1DFA35FA89" pageId="3" pageNumber="504">
16a. Posterior nasal tube elongate, about twice length of anterior nasal tube; second dorsal-fin rays usually I,12; pectoral-fin rays usually 1718; posterior opercular margin with dark bar (western Indian Ocean)..........................
<taxonomicName id="4C604D125F7FFFB3FAC9FAFBFA35FA89" ID-CoL="3L7HS" baseAuthorityName="Smith" baseAuthorityYear="1958" box="[1318,1436,1298,1319]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="504" phylum="Chordata" rank="species" species="caminata">
<emphasis id="B914EA835F7FFFB3FAC9FAFBFA35FA89" box="[1318,1436,1298,1319]" italics="true" pageId="3" pageNumber="504">H. caminata</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BDF36915F7FFFB3FF78FAC6FA34FACF" blockId="3.[151,1437,153,1583]" pageId="3" pageNumber="504">
<keyLead id="309195A45F7FFFB3FF78FAC6FA34FACF" pageId="3" pageNumber="504">
16b. Posterior nasal tube subequal to anterior tube; second dorsal-fin rays usually I,11; pectoral-fin rays usually 1516; posterior opercular margin with light edge (western to central Indian Ocean)......................................
<taxonomicName id="4C604D125F7FFFB3FADFFAA4FA34FACF" ID-CoL="3L7JD" baseAuthorityName="Klunzinger" baseAuthorityYear="1871" box="[1328,1437,1356,1378]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="504" phylum="Chordata" rank="species" species="vulgaris">
<emphasis id="B914EA835F7FFFB3FADFFAA4FA34FACF" box="[1328,1437,1356,1378]" italics="true" pageId="3" pageNumber="504">H. vulgaris</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30942E345F7FFFB3FF78FA83FA35FA17" pageId="3" pageNumber="504">
<paragraph id="8BDF36915F7FFFB3FF78FA83FA35FA32" blockId="3.[151,1437,153,1583]" pageId="3" pageNumber="504">
<keyLead id="309195A45F7FFFB3FF78FA83FA35FA32" pageId="3" pageNumber="504">
17a. Posterior nasal tube slightly shorter than anterior nasal tube; transverse dark band below first dorsal fin (
<collectingRegion id="49A4F8735F7FFFB3FB24FA80FAB4FA2E" box="[1227,1309,1385,1408]" country="Egypt" name="Al Bahr al Ahmar" pageId="3" pageNumber="504">Red Sea</collectingRegion>
)..................................................................................................
<taxonomicName id="4C604D125F7FFFB3FAF5FA6EFA35FA32" ID-CoL="3L7HW" baseAuthorityName="Ruppell" baseAuthorityYear="1830" box="[1306,1436,1415,1436]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="504" phylum="Chordata" rank="species" species="diademata">
<emphasis id="B914EA835F7FFFB3FAF5FA6EFA35FA32" box="[1306,1436,1415,1436]" italics="true" pageId="3" pageNumber="504">H. diademata</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BDF36915F7FFFB3FF78FA4DFA35FA17" blockId="3.[151,1437,153,1583]" box="[151,1436,1444,1466]" pageId="3" pageNumber="504">
<keyLead id="309195A45F7FFFB3FF78FA4DFA35FA17" box="[151,1436,1444,1466]" pageId="3" pageNumber="504">17b. Posterior nasal tube 1/10-1/2 of length of anterior nasal tube; no transverse dark band below first dorsal fin............. 18</keyLead>
</paragraph>
</keyStep>
<keyStep id="30942E345F7FFFB3FF78FA2BFA35F980" pageId="3" pageNumber="504">
<paragraph id="8BDF36915F7FFFB3FF78FA2BFA35FA5A" blockId="3.[151,1437,153,1583]" pageId="3" pageNumber="504">
<keyLead id="309195A45F7FFFB3FF78FA2BFA35FA5A" pageId="3" pageNumber="504">
18a. Dorsal fins with longitudinal dark submarginal band; second dorsal-fin rays I,12; anal-fin rays I,11 (
<collectingRegion id="49A4F8735F7FFFB3FB4EFA28FB5DFA76" box="[1185,1268,1473,1496]" country="Egypt" name="Al Bahr al Ahmar" pageId="3" pageNumber="504">Red Sea</collectingRegion>
).....................................................................................................
<taxonomicName id="4C604D125F7FFFB3FAE4FA36FA35FA5A" authorityName="Kovacic &amp; Bogorodsky" authorityYear="2014" box="[1291,1436,1503,1524]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="504" phylum="Chordata" rank="species" species="dorsovittata">
<emphasis id="B914EA835F7FFFB3FAE4FA36FA35FA5A" box="[1291,1436,1503,1524]" italics="true" pageId="3" pageNumber="504">H. dorsovittata</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BDF36915F7FFFB3FF78FA15FA35F980" blockId="3.[151,1437,153,1583]" pageId="3" pageNumber="504">
<keyLead id="309195A45F7FFFB3FF78FA15FA35F980" pageId="3" pageNumber="504">
18b. No longitudinal dark submarginal band on dorsal fins; second dorsal-fin rays I,10; anal-fin rays I,9 (
<collectingRegion id="49A4F8735F7FFFB3FB4DFA15FB5DF9BC" box="[1186,1268,1532,1554]" country="Egypt" name="Al Bahr al Ahmar" pageId="3" pageNumber="504">Red Sea</collectingRegion>
)....................................................................................................
<taxonomicName id="4C604D125F7FFFB3FB11F9F3FA35F980" ID-CoL="3L7J6" authorityName="Kovacic &amp; Bogorodsky" authorityYear="2014" box="[1278,1436,1561,1583]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="504" phylum="Chordata" rank="species" species="psammophila">
<emphasis id="B914EA835F7FFFB3FB11F9F3FA35F980" box="[1278,1436,1561,1583]" italics="true" pageId="3" pageNumber="504">H. psammophila</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>