treatments-xml/data/03/C9/48/03C948605B13FFB028C919EAFB5F6C19.xml
2024-06-21 12:22:17 +02:00

959 lines
106 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="0C9054FA026B0280EAB2561AFF7FC8AC" ID-CLB-Dataset="296802" ID-DOI="10.5281/zenodo.11281013" ID-GBIF-Dataset="4c4497ef-d299-4263-a5b7-45da80aeeb0d" ID-ISSN="2189-6720" ID-Zenodo-Dep="11281013" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1714415915295" checkinUser="felipe" docAuthor="Sugawara, Hirotaka, Naito, Jun-ichi, Iwata, Takayuki &amp; Nagano, Masahiro" docDate="2022" docId="03C948605B13FFB028C919EAFB5F6C19" docLanguage="en" docName="BullKanagawaPrefectMus.51.35-46.pdf" docOrigin="Bulletin of the Kanagawa Prefectural Museum (Natural Science) 51" docStyle="DocumentStyle:020B98DC1285EBBD02620724ECB44EDE.1:BullKanagawaPrefectMus.2013-.journal_article.english" docStyleId="020B98DC1285EBBD02620724ECB44EDE" docStyleName="BullKanagawaPrefectMus.2013-.journal_article.english" docStyleVersion="1" docTitle="Hynobius geiyoensis Sugawara &amp; Naito &amp; Iwata &amp; Nagano 2022, sp. nov." docType="treatment" docVersion="3" lastPageNumber="42" masterDocId="FFF030185B17FFB72B261B7CFF8B6E21" masterDocTitle="Molecular Phylogenetic and Morphological Problems of the Aki Salamander Hynobius akiensis: Description of Two New Species from Chugoku, Japan" masterLastPageNumber="46" masterPageNumber="35" pageNumber="39" updateTime="1716555535773" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="41E3C0F33384270D0DDB939B458CB084" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6DB36023D3D4CBED6D6C30C400573EF6">
<mods:title id="762F4AB82746A1D5963AB8FA4C7CEE42">Molecular Phylogenetic and Morphological Problems of the Aki Salamander Hynobius akiensis: Description of Two New Species from Chugoku, Japan</mods:title>
</mods:titleInfo>
<mods:name id="873DBC73607BAEE11C132F88E8CA9FAC" type="personal">
<mods:role id="E9ACF134C86157DB610146E75C096310">
<mods:roleTerm id="ACB4358631DCCDC3F3E76097BB21E8DC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CFAC73CD3273FE0FE8FA463EDE33F221">Sugawara, Hirotaka</mods:namePart>
<mods:affiliation id="AD2438FBFE32C05495F523194C933270">Faculty of Science and Technology, Kochi University, 2 - 5 - 1 Akebonocho, Kochi-shi, Kochi 780 - 8520, Japan</mods:affiliation>
<mods:nameIdentifier id="A3CEE5EE9251E047CAFD62FD889FFFAC" type="email">chiropterotriton@yahoo·co·jp</mods:nameIdentifier>
</mods:name>
<mods:name id="5CD384D40F48CF4EF88AC26AADEDDC13" type="personal">
<mods:role id="9A91843DC0493D08B1E20972854B5F19">
<mods:roleTerm id="2DE26BBB536F928AA2E1A17C4B8A41E1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8B50782D32FC503F5F28741469105C6E">Naito, Jun-ichi</mods:namePart>
<mods:affiliation id="7CEEF025C57E9A9B67FA476441A530D2">Natural History Society of Nishi-Chugoku Sanchi, 10119 - 1 Higashiyawatahara, Kitahiroshima-cho, Hiroshima 731 - 2551, Japan</mods:affiliation>
</mods:name>
<mods:name id="98B5EAF697C5D61265471AFD14FE39FC" type="personal">
<mods:role id="7695376118E50F05D5B6DE05A1F993EB">
<mods:roleTerm id="1CA008F49DC00EA468C13606DF87BB59">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BAFC1585D6D869DECD16F3DCA7BB133A">Iwata, Takayuki</mods:namePart>
<mods:affiliation id="9C85D668C0578BEAFCE2428CA27F9693">Yasugi City Hall, 878 - 2, Yasugicho, Yasugi-shi, Shimane 692 - 8686, Japan</mods:affiliation>
</mods:name>
<mods:name id="2445B342598FBA09F8B891F916E19D8A" type="personal">
<mods:role id="CB37F69C1E6E527D7132BDD28AF4E4B0">
<mods:roleTerm id="54F38E99529614E51685F71543C315E7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DD30C66DAB1E166F249F8510C7D1EA6C">Nagano, Masahiro</mods:namePart>
<mods:affiliation id="F34E577BC931208C499388CB800F9DDD">Faculty of Science and Technology, Oita University, 700 Dannoharu, Oita-shi, Oita 870 - 1192, Japan</mods:affiliation>
</mods:name>
<mods:typeOfResource id="B4466B3945039D29335A8C1E7462D242">text</mods:typeOfResource>
<mods:relatedItem id="B6B16941BDDBDF8BE93DCC00EA0E3581" type="host">
<mods:titleInfo id="A7D4B72B81248D4C9466DBC65BA030B0">
<mods:title id="ABCDE215DFCD2729826C755EDB3DAA39">Bulletin of the Kanagawa Prefectural Museum (Natural Science)</mods:title>
</mods:titleInfo>
<mods:part id="EE5E03F836643591FBDC08BB71576BC4">
<mods:date id="54E9993501A769290012E30373410D4E">2022</mods:date>
<mods:detail id="EDA264F1169513B316E494331E1F5EF1" type="pubDate">
<mods:number id="8B0775233A20BA142FE59048FC252B3B">2022-03-29</mods:number>
</mods:detail>
<mods:detail id="F61DD184E214245AF661E724BB29D41D" type="volume">
<mods:number id="450CE2AFDDD33D08183C54B624F3BCC6">51</mods:number>
</mods:detail>
<mods:extent id="2F0C291D02FB4509BD54D928DADEAAFF" unit="page">
<mods:start id="A9D26DEABF1A5E08D42E7F154F9F4AD9">35</mods:start>
<mods:end id="B7D631699CF03115E7EBF4023B58C7B9">46</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="1232BE04610583D4EAE01DA86F44FE21">journal article</mods:classification>
<mods:identifier id="43F1EFFF5E78E29AC04DE4C67FA521B9" type="CLB-Dataset">296802</mods:identifier>
<mods:identifier id="E00274FE0A59223D5707048633BAD374" type="DOI">10.5281/zenodo.11281013</mods:identifier>
<mods:identifier id="B63A0071A0ED6076D616CBAC0CEB8BDE" type="GBIF-Dataset">4c4497ef-d299-4263-a5b7-45da80aeeb0d</mods:identifier>
<mods:identifier id="4F05B44EE5B0560CB39D631E7F79EDEF" type="ISSN">2189-6720</mods:identifier>
<mods:identifier id="80C0C7B7A3FE424A8D875B7EA8510EE0" type="Zenodo-Dep">11281013</mods:identifier>
<mods:identifier id="A4D247649BA3735842D0F9D864D7FFEB" type="ZooBank">2861D048-4805-4B5C-90E0-274A313A16EE</mods:identifier>
</mods:mods>
<treatment id="03C948605B13FFB028C919EAFB5F6C19" ID-DOI="http://doi.org/10.5281/zenodo.11281023" ID-Zenodo-Dep="11281023" LSID="urn:lsid:plazi:treatment:03C948605B13FFB028C919EAFB5F6C19" httpUri="http://treatment.plazi.org/id/03C948605B13FFB028C919EAFB5F6C19" lastPageId="7" lastPageNumber="42" pageId="4" pageNumber="39">
<subSubSection id="C37AAAFD5B13FFB328C919EAFA9C6C8E" box="[1007,1303,662,687]" pageId="4" pageNumber="39" type="nomenclature">
<paragraph id="8BDFF9765B13FFB328C919EAFA9C6C8E" blockId="4.[883,1403,662,765]" box="[1007,1303,662,687]" pageId="4" pageNumber="39">
<heading id="D0974E1A5B13FFB328C919EAFA9C6C8E" box="[1007,1303,662,687]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="39" reason="6">
<emphasis id="B91425645B13FFB328C919EAFA9C6C8E" bold="true" box="[1007,1303,662,687]" pageId="4" pageNumber="39">
<taxonomicName id="4C6082F55B13FFB328C919EAFB496C8F" authority="Sugawara &amp; Naito &amp; Iwata &amp; Nagano, 2022" authorityName="Sugawara &amp; Naito &amp; Iwata &amp; Nagano" authorityYear="2022" box="[1007,1218,662,686]" class="Amphibia" family="Hynobiidae" genus="Hynobius" kingdom="Animalia" order="Caudata" pageId="4" pageNumber="39" phylum="Chordata" rank="species" species="geiyoensis" status="sp. nov.">
<emphasis id="B91425645B13FFB328C919EAFB496C8F" bold="true" box="[1007,1218,662,686]" italics="true" pageId="4" pageNumber="39">Hynobius geiyoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A227981F5B13FFB32FEE19EBFA9C6C8E" box="[1224,1303,663,687]" pageId="4" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37AAAFD5B13FFB3285519C2FAF06CF7" box="[883,1403,701,726]" pageId="4" pageNumber="39" type="vernacular_names">
<paragraph id="8BDFF9765B13FFB3285519C2FAF06CF7" blockId="4.[883,1403,662,765]" box="[883,1403,701,726]" pageId="4" pageNumber="39">
(New standard Japanese name:
<emphasis id="B91425645B13FFB32FE519C1FAF96CF4" box="[1219,1394,701,725]" italics="true" pageId="4" pageNumber="39">Geiyo-sanshouo</emphasis>
)
</paragraph>
</subSubSection>
<subSubSection id="C37AAAFD5B13FFB32F181998FB3A6CDD" box="[1086,1201,740,765]" pageId="4" pageNumber="39" type="description">
<paragraph id="8BDFF9765B13FFB32F181998FB3A6CDD" blockId="4.[883,1403,662,765]" box="[1086,1201,740,765]" pageId="4" pageNumber="39">
(
<figureCitation id="135BE5F35B13FFB32F601998FB226CDD" box="[1094,1193,740,765]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="6.[851,886,1402,1424]" captionStartId-1="8.[189,225,1023,1045]" captionTargetBox-0="[856,1436,156,1378]" captionTargetBox-1="[189,775,156,1002]" captionTargetId-0="figure-423@6.[850,1436,151,1378]" captionTargetId-1="figure-6@8.[189,775,156,1002]" captionTargetPageId-0="6" captionTargetPageId-1="8" captionText-0="Fig. 4. Holotype of Hynobius geiyoensis sp. nov. (HMNH- AM-101, adult male, 58.4 mm SVL) from the (A) dorsal and (B) ventral perspective; holotype of H. sumidai sp. nov. (HMNH-AM-102, adult male, 48.2 mm SVL) from the (C) dorsal and (D) ventral perspective; and a specimen from the type locality (topotype) of H. akiensis (KPM- NFA 946, adult male, 55.3 mm) from the (E) dorsal and (F) ventral perspective." captionText-1="Fig. 5. Larvae, egg sacs, and habitat at the type locality of Hynobius geiyoensis sp. nov. (A, C, and E, respectively) and H. sumidai sp. nov. (B, D, and F, respectively)." figureDoi-0="http://doi.org/10.5281/zenodo.11281025" figureDoi-1="http://doi.org/10.5281/zenodo.11281027" httpUri-0="https://zenodo.org/record/11281025/files/figure.png" httpUri-1="https://zenodo.org/record/11281027/files/figure.png" pageId="4" pageNumber="39">Figs. 45</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37AAAFD5B13FFB32875184EFB2D6DB9" pageId="4" pageNumber="39" type="reference_group">
<paragraph id="8BDFF9765B13FFB32875184EFAE66D6B" blockId="4.[851,1434,818,920]" box="[851,1389,818,842]" pageId="4" pageNumber="39">
<treatmentCitationGroup id="AB70DE585B13FFB32875184EFAE66D6B" box="[851,1389,818,842]" pageId="4" pageNumber="39">
<taxonomicName id="4C6082F55B13FFB32875184EFBA86D6B" ID-CoL="6MQKZ" baseAuthorityName="Temminck &amp; Schlegel" baseAuthorityYear="1838" box="[851,1059,818,842]" class="Amphibia" family="Hynobiidae" genus="Hynobius" kingdom="Animalia" order="Caudata" pageId="4" pageNumber="39" phylum="Chordata" rank="species" species="nebulosus">
<emphasis id="B91425645B13FFB32875184EFBA86D6B" box="[851,1059,818,842]" italics="true" pageId="4" pageNumber="39">Hynobius nebulosus</emphasis>
</taxonomicName>
:
<treatmentCitation id="0AC1DF675B13FFB32F16184EFA916D6B" author="Okawa, H. &amp; T. Utsunomiya &amp; Y. Utsunomiya &amp; J. Naito" box="[1072,1306,818,842]" page="50" pageId="4" pageNumber="39" year="1990">
<bibRefCitation id="EFF184875B13FFB32F16184EFA916D6B" author="Okawa, H. &amp; T. Utsunomiya &amp; Y. Utsunomiya &amp; J. Naito" box="[1072,1306,818,842]" pageId="4" pageNumber="39" pagination="49 - 53" refId="ref10359" refString="Okawa, H., T. Utsunomiya, Y. Utsunomiya &amp; J. Naito, 1990. Hynobius nebulosus from Hiroshima Prefecture. Hibakagaku, 146: 49 - 53. (In Japanese)." type="journal article" year="1990">
Okawa
<emphasis id="B91425645B13FFB32F58184FFB396D6B" box="[1150,1202,818,842]" italics="true" pageId="4" pageNumber="39">et al.</emphasis>
, 1990: 50
</bibRefCitation>
</treatmentCitation>
, in part.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDFF9765B13FFB328751825FB2D6DB9" blockId="4.[851,1434,818,920]" pageId="4" pageNumber="39">
<treatmentCitationGroup id="AB70DE585B13FFB328751825FA116D50" box="[851,1434,857,881]" pageId="4" pageNumber="39">
<taxonomicName id="4C6082F55B13FFB328751825FAF66D50" ID-CoL="3NLSY" authority="Matsui, Okawa and Nishikawa" authorityName="Matsui, Okawa and Nishikawa" box="[851,1405,857,881]" class="Amphibia" family="Hynobiidae" genus="Hynobius" kingdom="Animalia" order="Caudata" pageId="4" pageNumber="39" phylum="Chordata" rank="species" species="akiensis">
<emphasis id="B91425645B13FFB328751825FB966D50" box="[851,1053,857,881]" italics="true" pageId="4" pageNumber="39">Hynobius akiensis</emphasis>
Matsui, Okawa and Nishikawa
</taxonomicName>
in
</treatmentCitationGroup>
<bibRefCitation id="EFF184875B13FFB3284D18FCFBD86DB9" author="Matsui, M. &amp; H. Okawa &amp; K. Nishikawa &amp; G. Aoki &amp; K. Eto &amp; N. Yoshikawa &amp; S. Tanabe &amp; Y. Misawa &amp; A. Tominaga" box="[875,1107,896,920]" pageId="4" pageNumber="39" pagination="32 - 90" refId="ref10175" refString="Matsui, M., H. Okawa, K. Nishikawa, G. Aoki, K. Eto, N. Yoshikawa, S. Tanabe, Y. Misawa &amp; A. Tominaga, 2019. Systematics of the widely distributed Japanese clouded salamander, Hynobius nebulosus (Amphibia: Caudata: Hynobiidae), and its closest relatives. Current Herpetology, 38 (1): 32 - 90." type="journal article" year="2019">
Matsui
<emphasis id="B91425645B13FFB3289E18FDFC606DB9" box="[952,1003,896,920]" italics="true" pageId="4" pageNumber="39">et al.</emphasis>
, 2019: 75
</bibRefCitation>
, in part.
</paragraph>
</subSubSection>
<caption id="DF1FA9FE5B13FFB32B9A18D2FE066B3C" ID-DOI="http://doi.org/10.5281/zenodo.11281017" ID-Zenodo-Dep="11281017" httpUri="https://zenodo.org/record/11281017/files/figure.png" pageId="4" pageNumber="39" startId="4.[188,222,942,964]" targetBox="[220,742,155,883]" targetPageId="4" targetType="figure">
<paragraph id="8BDFF9765B13FFB32B9A18D2FE066B3C" blockId="4.[188,773,942,1309]" pageId="4" pageNumber="39">
Fig. 2. Phylogenetic tree produced by Bayesian inference using 630-bp cytochrome b genes.
<taxonomicName id="4C6082F55B13FFB3292518B1FC8F6DC3" authorityName="Dybowski" authorityYear="1870" box="[515,772,973,994]" class="Amphibia" family="Hynobiidae" genus="Salamandrella" kingdom="Animalia" order="Caudata" pageId="4" pageNumber="39" phylum="Chordata" rank="species" species="keyserlingii">
<emphasis id="B91425645B13FFB3292518B1FC8F6DC3" box="[515,772,973,994]" italics="true" pageId="4" pageNumber="39">Salamandrella keyserlingii</emphasis>
</taxonomicName>
was used as an outgroup. Scale bar shows genetic distance (expected changes per site). Numbers located near the nodes are posterior probabilities (PP) for Bayesian inference and bootstrap (BS) values for maximum likelihood estimation. Values appearing in parentheses after the haplotype names correspond to population localities as indicated in Table 1 and Fig. 1. Asterisks after the parentheses (Pops. 1, 8, and 54) indicate the type locality of the three species. The labels covered by shaded boxes indicate the transition type of
<taxonomicName id="4C6082F55B13FFB32BC31E7BFE0C6B3D" box="[229,391,1287,1308]" class="Amphibia" family="Hynobiidae" genus="Hynobius" kingdom="Animalia" order="Caudata" pageId="4" pageNumber="39" phylum="Chordata" rank="species" species="akiensis">
<emphasis id="B91425645B13FFB32BC31E7BFE0C6B3D" box="[229,391,1287,1308]" italics="true" pageId="4" pageNumber="39">Hynobius akiensis</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="C37AAAFD5B13FFB3284D18B2FBE56AA0" pageId="4" pageNumber="39" type="etymology">
<paragraph id="8BDFF9765B13FFB3284D18B2FBE56AA0" blockId="4.[851,1436,974,2088]" pageId="4" pageNumber="39">
<emphasis id="B91425645B13FFB3284D18B2FC7D6DC7" bold="true" box="[875,1014,974,998]" pageId="4" pageNumber="39">Etymology.</emphasis>
The specific name is derived from “
<emphasis id="B91425645B13FFB328791889FC2B6A2D" box="[863,928,1013,1036]" italics="true" pageId="4" pageNumber="39">Geiyo</emphasis>
.” In ancient times, the combined area of Aki (= central to the western part of
<collectingRegion id="49A437945B13FFB32F971F60FA1F6A15" box="[1201,1428,1052,1076]" country="Japan" name="Hiroshima" pageId="4" pageNumber="39">Hiroshima Prefecture</collectingRegion>
) and Iyo (=
<collectingRegion id="49A437945B13FFB328F61F3FFB196A7A" box="[976,1170,1091,1115]" country="Japan" name="Ehime" pageId="4" pageNumber="39">Ehime Prefecture</collectingRegion>
), where the new species occurred, was called
<emphasis id="B91425645B13FFB32F0E1F16FBED6AA0" box="[1064,1126,1130,1153]" italics="true" pageId="4" pageNumber="39">Geiyo</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="C37AAAFD5B13FFB3284D1FEDFB136882" pageId="4" pageNumber="39" type="materials_examined">
<paragraph id="8BDFF9765B13FFB3284D1FEDFC396B4A" blockId="4.[851,1436,974,2088]" pageId="4" pageNumber="39">
<materialsCitation id="3B08F32B5B13FFB3284D1FEDFC396B4A" collectingDate="2019-02-23" collectorName="Jun-ichi Naito" country="Japan" county="Chugoku" elevation="150" latitude="34.216667" location="An" longLatPrecision="1200" longitude="132.88333" municipality="Osakikamijima-cho" pageId="4" pageNumber="39" specimenCount="1" stateProvince="Hiroshima" typeStatus="holotype">
<emphasis id="B91425645B13FFB3284D1FEDFC5E6A88" bold="true" box="[875,981,1169,1193]" pageId="4" pageNumber="39">
<typeStatus id="54DB47D45B13FFB3284D1FEDFC5A6A88" box="[875,977,1169,1193]" pageId="4" pageNumber="39" type="holotype">Holotype</typeStatus>
.
</emphasis>
<location id="8EBFAFAD5B13FFB328FA1FEDFC776A88" LSID="urn:lsid:plazi:treatment:03C948605B13FFB028C919EAFB5F6C19:8EBFAFAD5B13FFB328FA1FEDFC776A88" box="[988,1020,1169,1193]" country="Japan" county="Chugoku" latitude="34.216667" longLatPrecision="1200" longitude="132.88333" municipality="Osakikamijima-cho" name="An" pageId="4" pageNumber="39" stateProvince="Hiroshima">An</location>
adult male (specimen number: HMNH-AM-101) from
<location id="8EBFAFAD5B13FFB328D11FC4FBC16AF1" LSID="urn:lsid:plazi:treatment:03C948605B13FFB028C919EAFB5F6C19:8EBFAFAD5B13FFB328D11FC4FBC16AF1" box="[1015,1098,1208,1232]" country="Japan" county="Chugoku" latitude="34.216667" longLatPrecision="1200" longitude="132.88333" municipality="Osakikamijima-cho" name="Nakano" pageId="4" pageNumber="39" stateProvince="Hiroshima">Nakano</location>
,
<collectingMunicipality id="6BBB630C5B13FFB32F701FC4FAA86AF1" box="[1110,1315,1208,1232]" pageId="4" pageNumber="39">Osakikamijima-cho</collectingMunicipality>
,
<collectingRegion id="49A437945B13FFB32E091FC4FC436AD6" country="Japan" name="Hiroshima" pageId="4" pageNumber="39">Hiroshima Prefecture</collectingRegion>
,
<collectingCounty id="62BE81FA5B13FFB328F31FA3FBB76AD6" box="[981,1084,1247,1271]" pageId="4" pageNumber="39">Chugoku</collectingCounty>
,
<collectingCountry id="F377B9E65B13FFB32F6C1FA3FB036AD7" box="[1098,1160,1247,1270]" name="Japan" pageId="4" pageNumber="39">Japan</collectingCountry>
[
<geoCoordinate id="EE549FB15B13FFB32FBC1FA3FA866AD7" box="[1178,1293,1245,1271]" degrees="34" direction="north" minutes="13" orientation="latitude" pageId="4" pageNumber="39" precision="925" value="34.216667">34° 13 N</geoCoordinate>
,
<geoCoordinate id="EE549FB15B13FFB32E3D1FA3FA1D6AD7" box="[1307,1430,1245,1271]" degrees="132" direction="east" minutes="53" orientation="longitude" pageId="4" pageNumber="39" precision="925" value="132.88333">132° 53 E</geoCoordinate>
; elevation =
<elevation id="004D1E455B13FFB328F51E79FB4F6B3C" box="[979,1220,1285,1309]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="39" unit="m" value="150.0">
<quantity id="4C9854935B13FFB328F51E79FB936B3C" box="[979,1048,1285,1309]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="39" unit="m" value="150.0">150 m</quantity>
above sea level
</elevation>
(a.s.l.); in all cases, datum = WGS84], collected by
<collectorName id="26959CA05B13FFB32FA91E50FA966B65" box="[1167,1309,1324,1348]" pageId="4" pageNumber="39">Jun-ichi Naito</collectorName>
on
<date id="FFDEDFB65B13FFB32E671E50FC256B4A" pageId="4" pageNumber="39" value="2019-02-23">
<collectingDate id="EF9A265E5B13FFB32E671E50FC256B4A" pageId="4" pageNumber="39" value="2019-02-23">February 23, 2019</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8BDFF9765B13FFB3284D1E06FB136882" blockId="4.[851,1436,974,2088]" pageId="4" pageNumber="39">
<materialsCitation id="3B08F32B5B13FFB3284D1E06FC026874" collectingDate="2019-02-23" collectorName="Jun-ichi Naito" country="Japan" county="Chugoku" elevation="30" latitude="34.233334" location="An" longLatPrecision="1200" longitude="132.86667" municipality="Osakikamijima-cho" pageId="4" pageNumber="39" specimenCode="KPM-NFA 943" specimenCount="1" stateProvince="Hiroshima" typeStatus="paratype">
<emphasis id="B91425645B13FFB3284D1E06FC5C6BB3" bold="true" box="[875,983,1402,1426]" pageId="4" pageNumber="39">
<typeStatus id="54DB47D45B13FFB3284D1E06FC5A6BB3" box="[875,977,1402,1426]" pageId="4" pageNumber="39" type="paratype">Paratype</typeStatus>
.
</emphasis>
<location id="8EBFAFAD5B13FFB328FA1E07FC776BB2" LSID="urn:lsid:plazi:treatment:03C948605B13FFB028C919EAFB5F6C19:8EBFAFAD5B13FFB328FA1E07FC776BB2" box="[988,1020,1403,1427]" country="Japan" county="Chugoku" latitude="34.233334" longLatPrecision="1200" longitude="132.86667" municipality="Osakikamijima-cho" name="An" pageId="4" pageNumber="39" stateProvince="Hiroshima">An</location>
adult female (specimen number:
<specimenCode id="DBC6510D5B13FFB32E7D1E07FC3F6B98" collectionCode="KPM-NFA" pageId="4" pageNumber="39">KPM-NFA 943</specimenCode>
) from
<location id="8EBFAFAD5B13FFB328DB1EDDFBC16B98" LSID="urn:lsid:plazi:treatment:03C948605B13FFB028C919EAFB5F6C19:8EBFAFAD5B13FFB328DB1EDDFBC16B98" box="[1021,1098,1441,1465]" country="Japan" county="Chugoku" latitude="34.233334" longLatPrecision="1200" longitude="132.86667" municipality="Osakikamijima-cho" name="Harada" pageId="4" pageNumber="39" stateProvince="Hiroshima">Harada</location>
,
<collectingMunicipality id="6BBB630C5B13FFB32F701EDDFAA86B98" box="[1110,1315,1441,1465]" pageId="4" pageNumber="39">Osakikamijima-cho</collectingMunicipality>
,
<collectingRegion id="49A437945B13FFB32E091EDDFC436BC1" country="Japan" name="Hiroshima" pageId="4" pageNumber="39">Hiroshima Prefecture</collectingRegion>
,
<collectingCounty id="62BE81FA5B13FFB328F31EB4FBB76BC1" box="[981,1084,1480,1504]" pageId="4" pageNumber="39">Chugoku</collectingCounty>
,
<collectingCountry id="F377B9E65B13FFB32F6C1EB5FB036BC1" box="[1098,1160,1481,1504]" name="Japan" pageId="4" pageNumber="39">Japan</collectingCountry>
[
<geoCoordinate id="EE549FB15B13FFB32FBC1EB4FA866BC1" box="[1178,1293,1479,1504]" degrees="34" direction="north" minutes="14" orientation="latitude" pageId="4" pageNumber="39" precision="925" value="34.233334">34° 14 N</geoCoordinate>
,
<geoCoordinate id="EE549FB15B13FFB32E3D1EB4FA1D6BC1" box="[1307,1430,1479,1504]" degrees="132" direction="east" minutes="52" orientation="longitude" pageId="4" pageNumber="39" precision="925" value="132.86667">132° 52 E</geoCoordinate>
; elevation =
<elevation id="004D1E455B13FFB328E11E93FB136826" box="[967,1176,1519,1543]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="39" unit="m" value="30.0">
<quantity id="4C9854935B13FFB328E11E93FC706826" box="[967,1019,1519,1543]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="39" unit="m" value="30.0">30 m</quantity>
above sea level
</elevation>
(a.s.l.); in all cases, datum = WGS84], collected by
<collectorName id="26959CA05B13FFB32F7A1D6AFB79680F" box="[1116,1266,1558,1582]" pageId="4" pageNumber="39">Jun-ichi Naito</collectorName>
on
<date id="FFDEDFB65B13FFB32E3A1D6AFC026874" pageId="4" pageNumber="39" value="2019-02-23">
<collectingDate id="EF9A265E5B13FFB32E3A1D6AFC026874" pageId="4" pageNumber="39" value="2019-02-23">23 February 2019</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B08F32B5B13FFB328B41D41FB1F6882" collectingDate="2019-02-23" collectorName="Jun-ichi Naito" country="Japan" county="Chugoku" elevation="150" latitude="34.216667" location="An" longLatPrecision="1200" longitude="132.88333" municipality="Osakikamijima-cho" pageId="4" pageNumber="39" specimenCode="KPM-NFA 944" specimenCount="1" stateProvince="Hiroshima" typeStatus="paratype">
An adult male (specimen number:
<specimenCode id="DBC6510D5B13FFB32FD01D41FA1E6874" box="[1270,1429,1597,1621]" collectionCode="KPM-NFA" pageId="4" pageNumber="39">KPM-NFA 944</specimenCode>
) from the same locality as the holotype, collected by
<collectorName id="26959CA05B13FFB32E561D18FC3D6882" pageId="4" pageNumber="39">Jun-ichi Naito</collectorName>
on
<date id="FFDEDFB65B13FFB328FC1DF7FB1F6882" box="[986,1172,1675,1699]" pageId="4" pageNumber="39" value="2019-02-23">
<collectingDate id="EF9A265E5B13FFB328FC1DF7FB1F6882" box="[986,1172,1675,1699]" pageId="4" pageNumber="39" value="2019-02-23">February 23, 2019</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37AAAFD5B13FFB2284D1DCEFD5A691C" lastPageId="5" lastPageNumber="40" pageId="4" pageNumber="39" type="diagnosis">
<paragraph id="8BDFF9765B13FFB2284D1DCEFD5A691C" blockId="4.[851,1436,974,2088]" lastBlockId="5.[150,738,1790,2087]" lastPageId="5" lastPageNumber="40" pageId="4" pageNumber="39">
<emphasis id="B91425645B13FFB3284D1DCEFC6E68EB" bold="true" box="[875,997,1714,1738]" pageId="4" pageNumber="39">Diagnosis.</emphasis>
A comparatively large species (with a mean snoutvent length of
<quantity id="4C9854935B13FFB32F4D1DA5FB4368D1" box="[1131,1224,1753,1777]" metricMagnitude="-2" metricUnit="m" metricValue="6.01" pageId="4" pageNumber="39" unit="mm" value="60.1">60.1 mm</quantity>
in males) within the Japanese lentic
<taxonomicName id="4C6082F55B13FFB328D71C7CFBDA6939" authorityName="Tschudi" authorityYear="1838" box="[1009,1105,1792,1816]" class="Amphibia" family="Hynobiidae" genus="Hynobius" kingdom="Animalia" order="Caudata" pageId="4" pageNumber="39" phylum="Chordata" rank="genus">
<emphasis id="B91425645B13FFB328D71C7CFBDA6939" box="[1009,1105,1792,1816]" italics="true" pageId="4" pageNumber="39">Hynobius</emphasis>
</taxonomicName>
: snoutvent length usually more than
<quantity id="4C9854935B13FFB328A01C5AFC59691F" box="[902,978,1830,1854]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="39" unit="mm" value="50.0">50 mm</quantity>
; distinct white dots on the ventral side of the body always present; distinct brownishyellow line on the dorsal edge of the tail almost always present; distinct white dots on the lateral side of the body usually present; distinct yellow stripe on the ventral edge of the tail always absent; distinct gular mottling almost always absent in males; fifth toe of hindlimb always present; V-shaped vomerine teeth series; 12 (rarely 11) costal grooves; the number of costal folds between adpressed limbs always 2 to
<quantity id="4C9854935B12FFB229771C59FD01691C" box="[593,650,1829,1853]" metricMagnitude="-2" metricUnit="m" metricValue="1.27" pageId="5" pageNumber="40" unit="in" value="0.5">0.5 in</quantity>
males.
</paragraph>
</subSubSection>
<caption id="DF1FA9FE5B13FFB32B9B1C01FE266630" ID-DOI="http://doi.org/10.5281/zenodo.11281021" ID-Zenodo-Dep="11281021" httpUri="https://zenodo.org/record/11281021/files/figure.png" pageId="4" pageNumber="39" startId="4.[189,226,1917,1939]" targetBox="[199,756,1404,1867]" targetPageId="4" targetType="figure">
<paragraph id="8BDFF9765B13FFB32B9B1C01FE266630" blockId="4.[189,773,1917,2065]" pageId="4" pageNumber="39">Fig. 3. Two-dimensional plots of canonical discriminant analysis in males. The x and y axes show discriminant score 1 (DS1) and discriminant score 2 (DS2), respectively. The contribution ratios of DS1 and DS2 were 88.03 % and 11.97 %, respectively.</paragraph>
</caption>
<caption id="DF1FA9FE5B12FFB22A171BDAFE6B6EFA" ID-Table-UUID="DF1FA9FE5B12FFB22A171BDAFE6B6EFA" httpUri="http://table.plazi.org/id/DF1FA9FE5B12FFB22A171BDAFE6B6EFA" pageId="5" pageNumber="40" startId="5.[305,356,166,188]" targetBox="[310,1215,240,1691]" targetIsTable="true" targetPageId="5" targetType="table">
<paragraph id="8BDFF9765B12FFB22A171BDAFE6B6EFA" blockId="5.[305,1259,166,219]" pageId="5" pageNumber="40">Table 2. Measurements (mm) of SVL and character ratios (R = %SVL) of TRL to LJL (ranges are shown in parentheses)</paragraph>
</caption>
<paragraph id="8BDFF9765B12FFB22AED1B8DFB3468BB" pageId="5" pageNumber="40">
<table id="F9600BD65B1200482A101B8CFB3468BA" box="[310,1215,240,1691]" gridcols="7" gridrows="46" pageId="5" pageNumber="40">
<tr id="3550FB345B1200482A101B8CFB346F22" box="[310,1215,240,259]" gridrow="0" pageId="5" pageNumber="40" rowspan-0="1">
<th id="768192485B1200482A861B8CFD0F6F22" box="[416,644,240,259]" colspan="2" colspanRight="1" gridcol="1" gridrow="0" pageId="5" pageNumber="40">
<taxonomicName id="4C6082F55B12FFB22AED1B8DFDB26F22" authority="Sugawara &amp; Naito &amp; Iwata &amp; Nagano, 2022" authorityName="Sugawara &amp; Naito &amp; Iwata &amp; Nagano" authorityYear="2022" box="[459,569,241,259]" class="Amphibia" family="Hynobiidae" genus="Hynobius" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="40" phylum="Chordata" rank="species" species="geiyoensis" status="sp. nov.">
<emphasis id="B91425645B12FFB22AED1B8DFDB26F22" box="[459,569,241,259]" italics="true" pageId="5" pageNumber="40">H. geiyoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A227981F5B12FFB229641B8CFD0F6F22" box="[578,644,240,259]" pageId="5" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</th>
<th id="768192485B12004829E61B8CFC2A6F22" box="[704,929,240,259]" colspan="2" colspanRight="1" gridcol="3" gridrow="0" pageId="5" pageNumber="40">
<taxonomicName id="4C6082F55B12FFB229D61B8DFCC16F22" authority="Sugawara &amp; Naito &amp; Iwata &amp; Nagano, 2022" authorityName="Sugawara &amp; Naito &amp; Iwata &amp; Nagano" authorityYear="2022" box="[752,842,240,259]" class="Amphibia" family="Hynobiidae" genus="Hynobius" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="40" phylum="Chordata" rank="species" species="sumidai" status="sp. nov.">
<emphasis id="B91425645B12FFB229D61B8DFCC16F22" box="[752,842,240,259]" italics="true" pageId="5" pageNumber="40">H. sumidai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A227981F5B12FFB228741B8CFC1F6F22" box="[850,916,240,259]" pageId="5" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</th>
<th id="768192485B12004828FB1B8CFB346F22" box="[989,1215,240,259]" colspan="2" colspanRight="1" gridcol="5" gridrow="0" pageId="5" pageNumber="40">
<taxonomicName id="4C6082F55B12FFB22F161B8DFB006F22" authority="Trait" authorityName="Trait" box="[1072,1163,240,259]" class="Amphibia" family="Hynobiidae" genus="Hynobius" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="40" phylum="Chordata" rank="species" species="akiensis">
<emphasis id="B91425645B12FFB22F161B8DFB006F22" box="[1072,1163,240,259]" italics="true" pageId="5" pageNumber="40">H. akiensis</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="3550FB345B1200482A101A6DFB346F05" box="[310,1215,273,292]" gridrow="1" pageId="5" pageNumber="40">
<th id="768192485B1200482A101A6DFE0F6F05" box="[310,388,273,292]" gridcol="0" gridrow="1" pageId="5" pageNumber="40">Trait</th>
<td id="768192485B1200482A861A6DFE676F05" box="[416,492,273,292]" gridcol="1" gridrow="1" pageId="5" pageNumber="40">Holotype</td>
<td id="768192485B12004829011A6DFD0F6F05" box="[551,644,273,292]" gridcol="2" gridrow="1" pageId="5" pageNumber="40">
<emphasis id="B91425645B12FFB2291F1A6DFDC86F02" box="[569,579,273,291]" italics="true" pageId="5" pageNumber="40">n</emphasis>
= 19
</td>
<td id="768192485B12004829E61A6DFC876F05" box="[704,780,273,292]" gridcol="3" gridrow="1" pageId="5" pageNumber="40">Holotype</td>
<td id="768192485B12004828631A6DFC2A6F05" box="[837,929,273,292]" gridcol="4" gridrow="1" pageId="5" pageNumber="40">
<emphasis id="B91425645B12FFB2287A1A6DFCED6F02" box="[860,870,273,291]" italics="true" pageId="5" pageNumber="40">n</emphasis>
= 7
</td>
<td id="768192485B12004828FB1A6DFBA06F05" box="[989,1067,273,292]" gridcol="5" gridrow="1" pageId="5" pageNumber="40">Topotype</td>
<td id="768192485B1200482F451A6DFB346F05" box="[1123,1215,273,292]" gridcol="6" gridrow="1" pageId="5" pageNumber="40">
<emphasis id="B91425645B12FFB22F531A6DFBF46F02" box="[1141,1151,273,291]" italics="true" pageId="5" pageNumber="40">n</emphasis>
= 28
</td>
</tr>
<tr id="3550FB345B1200482A101A4CFB346F65" box="[310,1215,304,324]" gridrow="2" pageId="5" pageNumber="40">
<th id="768192485B1200482A101A4CFE0F6F65" box="[310,388,304,324]" gridcol="0" gridrow="2" pageId="5" pageNumber="40">SVL</th>
<td id="768192485B1200482A861A4CFE676F65" box="[416,492,304,324]" gridcol="1" gridrow="2" pageId="5" pageNumber="40">58.4</td>
<td id="768192485B12004829011A4CFD0F6F65" box="[551,644,304,324]" gridcol="2" gridrow="2" pageId="5" pageNumber="40">60.1±3.23</td>
<td id="768192485B12004829E61A4CFC876F65" box="[704,780,304,324]" gridcol="3" gridrow="2" pageId="5" pageNumber="40">48.2</td>
<td id="768192485B12004828631A4CFC2A6F65" box="[837,929,304,324]" gridcol="4" gridrow="2" pageId="5" pageNumber="40">47.2±2.23</td>
<td id="768192485B12004828FB1A4CFBA06F65" box="[989,1067,304,324]" gridcol="5" gridrow="2" pageId="5" pageNumber="40">55.3</td>
<td id="768192485B1200482F451A4CFB346F65" box="[1123,1215,304,324]" gridcol="6" gridrow="2" pageId="5" pageNumber="40">51.2±4.07</td>
</tr>
<tr id="3550FB345B1200482A101A2CFB346F42" box="[310,1215,336,355]" gridrow="3" pageId="5" pageNumber="40" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-5="1">
<td id="768192485B12004829011A2CFD0F6F42" box="[551,644,336,355]" gridcol="2" gridrow="3" pageId="5" pageNumber="40">(50.5-64.5)</td>
<td id="768192485B12004828631A2CFC2A6F42" box="[837,929,336,355]" gridcol="4" gridrow="3" pageId="5" pageNumber="40">(44.9-49.7)</td>
<td id="768192485B1200482F451A2CFB346F42" box="[1123,1215,336,355]" gridcol="6" gridrow="3" pageId="5" pageNumber="40">(42.6-60.5)</td>
</tr>
<tr id="3550FB345B1200482A101A0CFB346FA2" box="[310,1215,368,387]" gridrow="4" pageId="5" pageNumber="40">
<th id="768192485B1200482A101A0CFE0F6FA2" box="[310,388,368,387]" gridcol="0" gridrow="4" pageId="5" pageNumber="40">RTRL</th>
<td id="768192485B1200482A861A0CFE676FA2" box="[416,492,368,387]" gridcol="1" gridrow="4" pageId="5" pageNumber="40">78.3</td>
<td id="768192485B12004829011A0CFD0F6FA2" box="[551,644,368,387]" gridcol="2" gridrow="4" pageId="5" pageNumber="40">77.2±0.80</td>
<td id="768192485B12004829E61A0CFC876FA2" box="[704,780,368,387]" gridcol="3" gridrow="4" pageId="5" pageNumber="40">75.9</td>
<td id="768192485B12004828631A0CFC2A6FA2" box="[837,929,368,387]" gridcol="4" gridrow="4" pageId="5" pageNumber="40">76.9±1.26</td>
<td id="768192485B12004828FB1A0CFBA06FA2" box="[989,1067,368,387]" gridcol="5" gridrow="4" pageId="5" pageNumber="40">77.8</td>
<td id="768192485B1200482F451A0CFB346FA2" box="[1123,1215,368,387]" gridcol="6" gridrow="4" pageId="5" pageNumber="40">77.0±1.14</td>
</tr>
<tr id="3550FB345B1200482A101AECFB346F82" box="[310,1215,400,419]" gridrow="5" pageId="5" pageNumber="40" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-5="1">
<td id="768192485B12004829011AECFD0F6F82" box="[551,644,400,419]" gridcol="2" gridrow="5" pageId="5" pageNumber="40">(75.5-78.4)</td>
<td id="768192485B12004828631AECFC2A6F82" box="[837,929,400,419]" gridcol="4" gridrow="5" pageId="5" pageNumber="40">(75.3-78.3)</td>
<td id="768192485B1200482F451AECFB346F82" box="[1123,1215,400,419]" gridcol="6" gridrow="5" pageId="5" pageNumber="40">(74.6-78.9)</td>
</tr>
<tr id="3550FB345B1200482A101AD3FB346FE2" box="[310,1215,431,451]" gridrow="6" pageId="5" pageNumber="40">
<th id="768192485B1200482A101AD3FE0F6FE2" box="[310,388,431,451]" gridcol="0" gridrow="6" pageId="5" pageNumber="40">RAGD</th>
<td id="768192485B1200482A861AD3FE676FE2" box="[416,492,431,451]" gridcol="1" gridrow="6" pageId="5" pageNumber="40">54.1</td>
<td id="768192485B12004829011AD3FD0F6FE2" box="[551,644,431,451]" gridcol="2" gridrow="6" pageId="5" pageNumber="40">52.5±1.32</td>
<td id="768192485B12004829E61AD3FC876FE2" box="[704,780,431,451]" gridcol="3" gridrow="6" pageId="5" pageNumber="40">53.9</td>
<td id="768192485B12004828631AD3FC2A6FE2" box="[837,929,431,451]" gridcol="4" gridrow="6" pageId="5" pageNumber="40">53.5±1.78</td>
<td id="768192485B12004828FB1AD3FBA06FE2" box="[989,1067,431,451]" gridcol="5" gridrow="6" pageId="5" pageNumber="40">50.3</td>
<td id="768192485B1200482F451AD3FB346FE2" box="[1123,1215,431,451]" gridcol="6" gridrow="6" pageId="5" pageNumber="40">52.9±1.41</td>
</tr>
<tr id="3550FB345B1200482A101AB3FB346FC3" box="[310,1215,463,482]" gridrow="7" pageId="5" pageNumber="40" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-5="1">
<td id="768192485B12004829011AB3FD0F6FC3" box="[551,644,463,482]" gridcol="2" gridrow="7" pageId="5" pageNumber="40">(49.8-54.4)</td>
<td id="768192485B12004828631AB3FC2A6FC3" box="[837,929,463,482]" gridcol="4" gridrow="7" pageId="5" pageNumber="40">(51.0-56.8)</td>
<td id="768192485B1200482F451AB3FB346FC3" box="[1123,1215,463,482]" gridcol="6" gridrow="7" pageId="5" pageNumber="40">(49.7-55.3)</td>
</tr>
<tr id="3550FB345B1200482A101A93FB346C22" box="[310,1215,495,515]" gridrow="8" pageId="5" pageNumber="40">
<th id="768192485B1200482A101A93FE0F6C22" box="[310,388,495,515]" gridcol="0" gridrow="8" pageId="5" pageNumber="40">RHL</th>
<td id="768192485B1200482A861A93FE676C22" box="[416,492,495,515]" gridcol="1" gridrow="8" pageId="5" pageNumber="40">21.4</td>
<td id="768192485B12004829011A93FD0F6C22" box="[551,644,495,515]" gridcol="2" gridrow="8" pageId="5" pageNumber="40">23.1±0.86</td>
<td id="768192485B12004829E61A93FC876C22" box="[704,780,495,515]" gridcol="3" gridrow="8" pageId="5" pageNumber="40">23.4</td>
<td id="768192485B12004828631A93FC2A6C22" box="[837,929,495,515]" gridcol="4" gridrow="8" pageId="5" pageNumber="40">24.3±1.54</td>
<td id="768192485B12004828FB1A93FBA06C22" box="[989,1067,495,515]" gridcol="5" gridrow="8" pageId="5" pageNumber="40">23.1</td>
<td id="768192485B1200482F451A93FB346C22" box="[1123,1215,495,515]" gridcol="6" gridrow="8" pageId="5" pageNumber="40">24.2±0.82</td>
</tr>
<tr id="3550FB345B1200482A101973FB346C03" box="[310,1215,527,546]" gridrow="9" pageId="5" pageNumber="40" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-5="1">
<td id="768192485B12004829011973FD0F6C03" box="[551,644,527,546]" gridcol="2" gridrow="9" pageId="5" pageNumber="40">(21.3-24.9)</td>
<td id="768192485B12004828631973FC2A6C03" box="[837,929,527,546]" gridcol="4" gridrow="9" pageId="5" pageNumber="40">(22.6-27.4)</td>
<td id="768192485B1200482F451973FB346C03" box="[1123,1215,527,546]" gridcol="6" gridrow="9" pageId="5" pageNumber="40">(23.1-26.0)</td>
</tr>
<tr id="3550FB345B1200482A101953FB346C63" box="[310,1215,559,578]" gridrow="10" pageId="5" pageNumber="40">
<th id="768192485B1200482A101953FE0F6C63" box="[310,388,559,578]" gridcol="0" gridrow="10" pageId="5" pageNumber="40">RTAL</th>
<td id="768192485B1200482A861953FE676C63" box="[416,492,559,578]" gridcol="1" gridrow="10" pageId="5" pageNumber="40">85.6</td>
<td id="768192485B12004829011953FD0F6C63" box="[551,644,559,578]" gridcol="2" gridrow="10" pageId="5" pageNumber="40">75.7±6.46</td>
<td id="768192485B12004829E61953FC876C63" box="[704,780,559,578]" gridcol="3" gridrow="10" pageId="5" pageNumber="40">71.8</td>
<td id="768192485B12004828631953FC2A6C63" box="[837,929,559,578]" gridcol="4" gridrow="10" pageId="5" pageNumber="40">68.9±7.47</td>
<td id="768192485B12004828FB1953FBA06C63" box="[989,1067,559,578]" gridcol="5" gridrow="10" pageId="5" pageNumber="40">58.2</td>
<td id="768192485B1200482F451953FB346C63" box="[1123,1215,559,578]" gridcol="6" gridrow="10" pageId="5" pageNumber="40">68.2±5.91</td>
</tr>
<tr id="3550FB345B1200482A101932FB346C40" box="[310,1215,590,609]" gridrow="11" pageId="5" pageNumber="40" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-5="1">
<td id="768192485B12004829011932FD0F6C40" box="[551,644,590,609]" gridcol="2" gridrow="11" pageId="5" pageNumber="40">(60.5-85.7)</td>
<td id="768192485B12004828631932FC2A6C40" box="[837,929,590,609]" gridcol="4" gridrow="11" pageId="5" pageNumber="40">(55.5-76.2)</td>
<td id="768192485B1200482F451932FB346C40" box="[1123,1215,590,609]" gridcol="6" gridrow="11" pageId="5" pageNumber="40">(57.2-78.4)</td>
</tr>
<tr id="3550FB345B1200482A101912FB346CA0" box="[310,1215,622,641]" gridrow="12" pageId="5" pageNumber="40">
<th id="768192485B1200482A101912FE0F6CA0" box="[310,388,622,641]" gridcol="0" gridrow="12" pageId="5" pageNumber="40">RMTAW</th>
<td id="768192485B1200482A861912FE676CA0" box="[416,492,622,641]" gridcol="1" gridrow="12" pageId="5" pageNumber="40">6.8</td>
<td id="768192485B12004829011912FD0F6CA0" box="[551,644,622,641]" gridcol="2" gridrow="12" pageId="5" pageNumber="40">6.0±0.78</td>
<td id="768192485B12004829E61912FC876CA0" box="[704,780,622,641]" gridcol="3" gridrow="12" pageId="5" pageNumber="40">5.0</td>
<td id="768192485B12004828631912FC2A6CA0" box="[837,929,622,641]" gridcol="4" gridrow="12" pageId="5" pageNumber="40">5.9±0.53</td>
<td id="768192485B12004828FB1912FBA06CA0" box="[989,1067,622,641]" gridcol="5" gridrow="12" pageId="5" pageNumber="40">4.9</td>
<td id="768192485B1200482F451912FB346CA0" box="[1123,1215,622,641]" gridcol="6" gridrow="12" pageId="5" pageNumber="40">6.5±0.86</td>
</tr>
<tr id="3550FB345B1200482A1019F2FB346C80" box="[310,1215,654,673]" gridrow="13" pageId="5" pageNumber="40" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-5="1">
<td id="768192485B120048290119F2FD0F6C80" box="[551,644,654,673]" gridcol="2" gridrow="13" pageId="5" pageNumber="40">(4.7-7.5)</td>
<td id="768192485B120048286319F2FC2A6C80" box="[837,929,654,673]" gridcol="4" gridrow="13" pageId="5" pageNumber="40">(5.0-6.7)</td>
<td id="768192485B1200482F4519F2FB346C80" box="[1123,1215,654,673]" gridcol="6" gridrow="13" pageId="5" pageNumber="40">(4.2-8.1)</td>
</tr>
<tr id="3550FB345B1200482A1019D2FB346CE0" box="[310,1215,686,705]" gridrow="14" pageId="5" pageNumber="40">
<th id="768192485B1200482A1019D2FE0F6CE0" box="[310,388,686,705]" gridcol="0" gridrow="14" pageId="5" pageNumber="40">RMTAH</th>
<td id="768192485B1200482A8619D2FE676CE0" box="[416,492,686,705]" gridcol="1" gridrow="14" pageId="5" pageNumber="40">14.0</td>
<td id="768192485B120048290119D2FD0F6CE0" box="[551,644,686,705]" gridcol="2" gridrow="14" pageId="5" pageNumber="40">11.7±1.00</td>
<td id="768192485B12004829E619D2FC876CE0" box="[704,780,686,705]" gridcol="3" gridrow="14" pageId="5" pageNumber="40">9.8</td>
<td id="768192485B120048286319D2FC2A6CE0" box="[837,929,686,705]" gridcol="4" gridrow="14" pageId="5" pageNumber="40">10.7±1.19</td>
<td id="768192485B12004828FB19D2FBA06CE0" box="[989,1067,686,705]" gridcol="5" gridrow="14" pageId="5" pageNumber="40">10.1</td>
<td id="768192485B1200482F4519D2FB346CE0" box="[1123,1215,686,705]" gridcol="6" gridrow="14" pageId="5" pageNumber="40">10.4±1.18</td>
</tr>
<tr id="3550FB345B1200482A1019B2FB346CC0" box="[310,1215,718,737]" gridrow="15" pageId="5" pageNumber="40" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-5="1">
<td id="768192485B120048290119B2FD0F6CC0" box="[551,644,718,737]" gridcol="2" gridrow="15" pageId="5" pageNumber="40">(9.5-14.0)</td>
<td id="768192485B120048286319B2FC2A6CC0" box="[837,929,718,737]" gridcol="4" gridrow="15" pageId="5" pageNumber="40">(9.3-12.6)</td>
<td id="768192485B1200482F4519B2FB346CC0" box="[1123,1215,718,737]" gridcol="6" gridrow="15" pageId="5" pageNumber="40">(7.7-12.7)</td>
</tr>
<tr id="3550FB345B1200482A101991FB346D20" box="[310,1215,749,769]" gridrow="16" pageId="5" pageNumber="40">
<th id="768192485B1200482A101991FE0F6D20" box="[310,388,749,769]" gridcol="0" gridrow="16" pageId="5" pageNumber="40">RVTL</th>
<td id="768192485B1200482A861991FE676D20" box="[416,492,749,769]" gridcol="1" gridrow="16" pageId="5" pageNumber="40">5.4</td>
<td id="768192485B12004829011991FD0F6D20" box="[551,644,749,769]" gridcol="2" gridrow="16" pageId="5" pageNumber="40">4.9±0.48</td>
<td id="768192485B12004829E61991FC876D20" box="[704,780,749,769]" gridcol="3" gridrow="16" pageId="5" pageNumber="40">5.4</td>
<td id="768192485B12004828631991FC2A6D20" box="[837,929,749,769]" gridcol="4" gridrow="16" pageId="5" pageNumber="40">5.2±0.49</td>
<td id="768192485B12004828FB1991FBA06D20" box="[989,1067,749,769]" gridcol="5" gridrow="16" pageId="5" pageNumber="40">4.7</td>
<td id="768192485B1200482F451991FB346D20" box="[1123,1215,749,769]" gridcol="6" gridrow="16" pageId="5" pageNumber="40">4.7±0.38</td>
</tr>
<tr id="3550FB345B1200482A101871FB346D01" box="[310,1215,781,800]" gridrow="17" pageId="5" pageNumber="40" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-5="1">
<td id="768192485B12004829011871FD0F6D01" box="[551,644,781,800]" gridcol="2" gridrow="17" pageId="5" pageNumber="40">(3.9-5.9)</td>
<td id="768192485B12004828631871FC2A6D01" box="[837,929,781,800]" gridcol="4" gridrow="17" pageId="5" pageNumber="40">(4.3-5.6)</td>
<td id="768192485B1200482F451871FB346D01" box="[1123,1215,781,800]" gridcol="6" gridrow="17" pageId="5" pageNumber="40">(3.3-5.3)</td>
</tr>
<tr id="3550FB345B1200482A101851FB346D60" box="[310,1215,813,833]" gridrow="18" pageId="5" pageNumber="40">
<th id="768192485B1200482A101851FE0F6D60" box="[310,388,813,833]" gridcol="0" gridrow="18" pageId="5" pageNumber="40">RVTW</th>
<td id="768192485B1200482A861851FE676D60" box="[416,492,813,833]" gridcol="1" gridrow="18" pageId="5" pageNumber="40">4.8</td>
<td id="768192485B12004829011851FD0F6D60" box="[551,644,813,833]" gridcol="2" gridrow="18" pageId="5" pageNumber="40">5.3±0.26</td>
<td id="768192485B12004829E61851FC876D60" box="[704,780,813,833]" gridcol="3" gridrow="18" pageId="5" pageNumber="40">5.2</td>
<td id="768192485B12004828631851FC2A6D60" box="[837,929,813,833]" gridcol="4" gridrow="18" pageId="5" pageNumber="40">5.4±0.24</td>
<td id="768192485B12004828FB1851FBA06D60" box="[989,1067,813,833]" gridcol="5" gridrow="18" pageId="5" pageNumber="40">5.6</td>
<td id="768192485B1200482F451851FB346D60" box="[1123,1215,813,833]" gridcol="6" gridrow="18" pageId="5" pageNumber="40">5.5±0.35</td>
</tr>
<tr id="3550FB345B1200482A101831FB346D41" box="[310,1215,845,864]" gridrow="19" pageId="5" pageNumber="40" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-5="1">
<td id="768192485B12004829011831FD0F6D41" box="[551,644,845,864]" gridcol="2" gridrow="19" pageId="5" pageNumber="40">(4.8-5.7)</td>
<td id="768192485B12004828631831FC2A6D41" box="[837,929,845,864]" gridcol="4" gridrow="19" pageId="5" pageNumber="40">(5.1-5.7)</td>
<td id="768192485B1200482F451831FB346D41" box="[1123,1215,845,864]" gridcol="6" gridrow="19" pageId="5" pageNumber="40">(4.9-6.1)</td>
</tr>
<tr id="3550FB345B1200482A101811FB346DA1" box="[310,1215,877,896]" gridrow="20" pageId="5" pageNumber="40">
<th id="768192485B1200482A101811FE0F6DA1" box="[310,388,877,896]" gridcol="0" gridrow="20" pageId="5" pageNumber="40">RHW</th>
<td id="768192485B1200482A861811FE676DA1" box="[416,492,877,896]" gridcol="1" gridrow="20" pageId="5" pageNumber="40">17.5</td>
<td id="768192485B12004829011811FD0F6DA1" box="[551,644,877,896]" gridcol="2" gridrow="20" pageId="5" pageNumber="40">16.4±0.60</td>
<td id="768192485B12004829E61811FC876DA1" box="[704,780,877,896]" gridcol="3" gridrow="20" pageId="5" pageNumber="40">16.6</td>
<td id="768192485B12004828631811FC2A6DA1" box="[837,929,877,896]" gridcol="4" gridrow="20" pageId="5" pageNumber="40">16.6±0.58</td>
<td id="768192485B12004828FB1811FBA06DA1" box="[989,1067,877,896]" gridcol="5" gridrow="20" pageId="5" pageNumber="40">17.7</td>
<td id="768192485B1200482F451811FB346DA1" box="[1123,1215,877,896]" gridcol="6" gridrow="20" pageId="5" pageNumber="40">16.7±0.61</td>
</tr>
<tr id="3550FB345B1200482A1018F0FB346DBE" box="[310,1215,908,927]" gridrow="21" pageId="5" pageNumber="40" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-5="1">
<td id="768192485B120048290118F0FD0F6DBE" box="[551,644,908,927]" gridcol="2" gridrow="21" pageId="5" pageNumber="40">(15.3-17.8)</td>
<td id="768192485B120048286318F0FC2A6DBE" box="[837,929,908,927]" gridcol="4" gridrow="21" pageId="5" pageNumber="40">(15.7-17.4)</td>
<td id="768192485B1200482F4518F0FB346DBE" box="[1123,1215,908,927]" gridcol="6" gridrow="21" pageId="5" pageNumber="40">(15.4-17.8)</td>
</tr>
<tr id="3550FB345B1200482A1018D0FB346DE1" box="[310,1215,940,960]" gridrow="22" pageId="5" pageNumber="40">
<th id="768192485B1200482A1018D0FE0F6DE1" box="[310,388,940,960]" gridcol="0" gridrow="22" pageId="5" pageNumber="40">RFLL</th>
<td id="768192485B1200482A8618D0FE676DE1" box="[416,492,940,960]" gridcol="1" gridrow="22" pageId="5" pageNumber="40">23.6</td>
<td id="768192485B120048290118D0FD0F6DE1" box="[551,644,940,960]" gridcol="2" gridrow="22" pageId="5" pageNumber="40">23.0±1.29</td>
<td id="768192485B12004829E618D0FC876DE1" box="[704,780,940,960]" gridcol="3" gridrow="22" pageId="5" pageNumber="40">24.5</td>
<td id="768192485B120048286318D0FC2A6DE1" box="[837,929,940,960]" gridcol="4" gridrow="22" pageId="5" pageNumber="40">25.1±1.42</td>
<td id="768192485B12004828FB18D0FBA06DE1" box="[989,1067,940,960]" gridcol="5" gridrow="22" pageId="5" pageNumber="40">22.8</td>
<td id="768192485B1200482F4518D0FB346DE1" box="[1123,1215,940,960]" gridcol="6" gridrow="22" pageId="5" pageNumber="40">25.1±1.22</td>
</tr>
<tr id="3550FB345B1200482A1018B0FB346DFE" box="[310,1215,972,991]" gridrow="23" pageId="5" pageNumber="40" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-5="1">
<td id="768192485B120048290118B0FD0F6DFE" box="[551,644,972,991]" gridcol="2" gridrow="23" pageId="5" pageNumber="40">(20.6-25.0)</td>
<td id="768192485B120048286318B0FC2A6DFE" box="[837,929,972,991]" gridcol="4" gridrow="23" pageId="5" pageNumber="40">(23.9-28.1)</td>
<td id="768192485B1200482F4518B0FB346DFE" box="[1123,1215,972,991]" gridcol="6" gridrow="23" pageId="5" pageNumber="40">(22.7-27.5)</td>
</tr>
<tr id="3550FB345B1200482A101890FB346DDE" box="[310,1215,1004,1023]" gridrow="24" pageId="5" pageNumber="40">
<th id="768192485B1200482A101890FE0F6DDE" box="[310,388,1004,1023]" gridcol="0" gridrow="24" pageId="5" pageNumber="40">RHLL</th>
<td id="768192485B1200482A861890FE676DDE" box="[416,492,1004,1023]" gridcol="1" gridrow="24" pageId="5" pageNumber="40">30.5</td>
<td id="768192485B12004829011890FD0F6DDE" box="[551,644,1004,1023]" gridcol="2" gridrow="24" pageId="5" pageNumber="40">30.6±1.03</td>
<td id="768192485B12004829E61890FC876DDE" box="[704,780,1004,1023]" gridcol="3" gridrow="24" pageId="5" pageNumber="40">29.3</td>
<td id="768192485B12004828631890FC2A6DDE" box="[837,929,1004,1023]" gridcol="4" gridrow="24" pageId="5" pageNumber="40">30.1±1.80</td>
<td id="768192485B12004828FB1890FBA06DDE" box="[989,1067,1004,1023]" gridcol="5" gridrow="24" pageId="5" pageNumber="40">31.8</td>
<td id="768192485B1200482F451890FB346DDE" box="[1123,1215,1004,1023]" gridcol="6" gridrow="24" pageId="5" pageNumber="40">30.6±1.29</td>
</tr>
<tr id="3550FB345B1200482A101F70FB346A3E" box="[310,1215,1036,1055]" gridrow="25" pageId="5" pageNumber="40" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-5="1">
<td id="768192485B12004829011F70FD0F6A3E" box="[551,644,1036,1055]" gridcol="2" gridrow="25" pageId="5" pageNumber="40">(28.4-32.7)</td>
<td id="768192485B12004828631F70FC2A6A3E" box="[837,929,1036,1055]" gridcol="4" gridrow="25" pageId="5" pageNumber="40">(27.6-32.7)</td>
<td id="768192485B1200482F451F70FB346A3E" box="[1123,1215,1036,1055]" gridcol="6" gridrow="25" pageId="5" pageNumber="40">(27.2-32.9)</td>
</tr>
<tr id="3550FB345B1200482A101F57FB346A1E" box="[310,1215,1067,1087]" gridrow="26" pageId="5" pageNumber="40">
<th id="768192485B1200482A101F57FE0F6A1E" box="[310,388,1067,1087]" gridcol="0" gridrow="26" pageId="5" pageNumber="40">R2FL</th>
<td id="768192485B1200482A861F57FE676A1E" box="[416,492,1067,1087]" gridcol="1" gridrow="26" pageId="5" pageNumber="40">5.5</td>
<td id="768192485B12004829011F57FD0F6A1E" box="[551,644,1067,1087]" gridcol="2" gridrow="26" pageId="5" pageNumber="40">5.0±0.54</td>
<td id="768192485B12004829E61F57FC876A1E" box="[704,780,1067,1087]" gridcol="3" gridrow="26" pageId="5" pageNumber="40">4.4</td>
<td id="768192485B12004828631F57FC2A6A1E" box="[837,929,1067,1087]" gridcol="4" gridrow="26" pageId="5" pageNumber="40">4.8±0.27</td>
<td id="768192485B12004828FB1F57FBA06A1E" box="[989,1067,1067,1087]" gridcol="5" gridrow="26" pageId="5" pageNumber="40">5.8</td>
<td id="768192485B1200482F451F57FB346A1E" box="[1123,1215,1067,1087]" gridcol="6" gridrow="26" pageId="5" pageNumber="40">4.9±0.58</td>
</tr>
<tr id="3550FB345B1200482A101F37FB346A7F" box="[310,1215,1099,1118]" gridrow="27" pageId="5" pageNumber="40" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-5="1">
<td id="768192485B12004829011F37FD0F6A7F" box="[551,644,1099,1118]" gridcol="2" gridrow="27" pageId="5" pageNumber="40">(4.2-6.1)</td>
<td id="768192485B12004828631F37FC2A6A7F" box="[837,929,1099,1118]" gridcol="4" gridrow="27" pageId="5" pageNumber="40">(4.4-5.1)</td>
<td id="768192485B1200482F451F37FB346A7F" box="[1123,1215,1099,1118]" gridcol="6" gridrow="27" pageId="5" pageNumber="40">(3.8-5.8)</td>
</tr>
<tr id="3550FB345B1200482A101F17FB346A5F" box="[310,1215,1131,1150]" gridrow="28" pageId="5" pageNumber="40">
<th id="768192485B1200482A101F17FE0F6A5F" box="[310,388,1131,1150]" gridcol="0" gridrow="28" pageId="5" pageNumber="40">R3FL</th>
<td id="768192485B1200482A861F17FE676A5F" box="[416,492,1131,1150]" gridcol="1" gridrow="28" pageId="5" pageNumber="40">5.3</td>
<td id="768192485B12004829011F17FD0F6A5F" box="[551,644,1131,1150]" gridcol="2" gridrow="28" pageId="5" pageNumber="40">4.5±0.64</td>
<td id="768192485B12004829E61F17FC876A5F" box="[704,780,1131,1150]" gridcol="3" gridrow="28" pageId="5" pageNumber="40">3.9</td>
<td id="768192485B12004828631F17FC2A6A5F" box="[837,929,1131,1150]" gridcol="4" gridrow="28" pageId="5" pageNumber="40">3.8±0.21</td>
<td id="768192485B12004828FB1F17FBA06A5F" box="[989,1067,1131,1150]" gridcol="5" gridrow="28" pageId="5" pageNumber="40">5.2</td>
<td id="768192485B1200482F451F17FB346A5F" box="[1123,1215,1131,1150]" gridcol="6" gridrow="28" pageId="5" pageNumber="40">4.2±0.43</td>
</tr>
<tr id="3550FB345B1200482A101FF7FB346ABF" box="[310,1215,1163,1182]" gridrow="29" pageId="5" pageNumber="40" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-5="1">
<td id="768192485B12004829011FF7FD0F6ABF" box="[551,644,1163,1182]" gridcol="2" gridrow="29" pageId="5" pageNumber="40">(3.0-5.3)</td>
<td id="768192485B12004828631FF7FC2A6ABF" box="[837,929,1163,1182]" gridcol="4" gridrow="29" pageId="5" pageNumber="40">(3.6-4.2)</td>
<td id="768192485B1200482F451FF7FB346ABF" box="[1123,1215,1163,1182]" gridcol="6" gridrow="29" pageId="5" pageNumber="40">(3.4-5.3)</td>
</tr>
<tr id="3550FB345B1200482A101FD6FB346A9F" box="[310,1215,1194,1214]" gridrow="30" pageId="5" pageNumber="40">
<th id="768192485B1200482A101FD6FE0F6A9F" box="[310,388,1194,1214]" gridcol="0" gridrow="30" pageId="5" pageNumber="40">R3TL</th>
<td id="768192485B1200482A861FD6FE676A9F" box="[416,492,1194,1214]" gridcol="1" gridrow="30" pageId="5" pageNumber="40">9.1</td>
<td id="768192485B12004829011FD6FD0F6A9F" box="[551,644,1194,1214]" gridcol="2" gridrow="30" pageId="5" pageNumber="40">8.2±0.49</td>
<td id="768192485B12004829E61FD6FC876A9F" box="[704,780,1194,1214]" gridcol="3" gridrow="30" pageId="5" pageNumber="40">6.8</td>
<td id="768192485B12004828631FD6FC2A6A9F" box="[837,929,1194,1214]" gridcol="4" gridrow="30" pageId="5" pageNumber="40">7.5±0.72</td>
<td id="768192485B12004828FB1FD6FBA06A9F" box="[989,1067,1194,1214]" gridcol="5" gridrow="30" pageId="5" pageNumber="40">8.9</td>
<td id="768192485B1200482F451FD6FB346A9F" box="[1123,1215,1194,1214]" gridcol="6" gridrow="30" pageId="5" pageNumber="40">7.9±0.69</td>
</tr>
<tr id="3550FB345B1200482A101FB6FB346AFC" box="[310,1215,1226,1245]" gridrow="31" pageId="5" pageNumber="40" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-5="1">
<td id="768192485B12004829011FB6FD0F6AFC" box="[551,644,1226,1245]" gridcol="2" gridrow="31" pageId="5" pageNumber="40">(7.4-9.3)</td>
<td id="768192485B12004828631FB6FC2A6AFC" box="[837,929,1226,1245]" gridcol="4" gridrow="31" pageId="5" pageNumber="40">(6.7-8.7)</td>
<td id="768192485B1200482F451FB6FB346AFC" box="[1123,1215,1226,1245]" gridcol="6" gridrow="31" pageId="5" pageNumber="40">(6.3-9.5)</td>
</tr>
<tr id="3550FB345B1200482A101F96FB346ADC" box="[310,1215,1258,1277]" gridrow="32" pageId="5" pageNumber="40">
<th id="768192485B1200482A101F96FE0F6ADC" box="[310,388,1258,1277]" gridcol="0" gridrow="32" pageId="5" pageNumber="40">R5TL</th>
<td id="768192485B1200482A861F96FE676ADC" box="[416,492,1258,1277]" gridcol="1" gridrow="32" pageId="5" pageNumber="40">2.7</td>
<td id="768192485B12004829011F96FD0F6ADC" box="[551,644,1258,1277]" gridcol="2" gridrow="32" pageId="5" pageNumber="40">2.4±0.44</td>
<td id="768192485B12004829E61F96FC876ADC" box="[704,780,1258,1277]" gridcol="3" gridrow="32" pageId="5" pageNumber="40">1.7</td>
<td id="768192485B12004828631F96FC2A6ADC" box="[837,929,1258,1277]" gridcol="4" gridrow="32" pageId="5" pageNumber="40">1.3±0.37</td>
<td id="768192485B12004828FB1F96FBA06ADC" box="[989,1067,1258,1277]" gridcol="5" gridrow="32" pageId="5" pageNumber="40">2.2</td>
<td id="768192485B1200482F451F96FB346ADC" box="[1123,1215,1258,1277]" gridcol="6" gridrow="32" pageId="5" pageNumber="40">1.3±0.79</td>
</tr>
<tr id="3550FB345B1200482A101E76FB346B3C" box="[310,1215,1290,1309]" gridrow="33" pageId="5" pageNumber="40" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-5="1">
<td id="768192485B12004829011E76FD0F6B3C" box="[551,644,1290,1309]" gridcol="2" gridrow="33" pageId="5" pageNumber="40">(1.7-3.4)</td>
<td id="768192485B12004828631E76FC2A6B3C" box="[837,929,1290,1309]" gridcol="4" gridrow="33" pageId="5" pageNumber="40">(0.8-1.8)</td>
<td id="768192485B1200482F451E76FB346B3C" box="[1123,1215,1290,1309]" gridcol="6" gridrow="33" pageId="5" pageNumber="40">(0.0-2.6)</td>
</tr>
<tr id="3550FB345B1200482A101E56FB346B1C" box="[310,1215,1322,1341]" gridrow="34" pageId="5" pageNumber="40">
<th id="768192485B1200482A101E56FE0F6B1C" box="[310,388,1322,1341]" gridcol="0" gridrow="34" pageId="5" pageNumber="40">RIND</th>
<td id="768192485B1200482A861E56FE676B1C" box="[416,492,1322,1341]" gridcol="1" gridrow="34" pageId="5" pageNumber="40">5.0</td>
<td id="768192485B12004829011E56FD0F6B1C" box="[551,644,1322,1341]" gridcol="2" gridrow="34" pageId="5" pageNumber="40">5.4±0.43</td>
<td id="768192485B12004829E61E56FC876B1C" box="[704,780,1322,1341]" gridcol="3" gridrow="34" pageId="5" pageNumber="40">4.8</td>
<td id="768192485B12004828631E56FC2A6B1C" box="[837,929,1322,1341]" gridcol="4" gridrow="34" pageId="5" pageNumber="40">5.2±0.35</td>
<td id="768192485B12004828FB1E56FBA06B1C" box="[989,1067,1322,1341]" gridcol="5" gridrow="34" pageId="5" pageNumber="40">5.2</td>
<td id="768192485B1200482F451E56FB346B1C" box="[1123,1215,1322,1341]" gridcol="6" gridrow="34" pageId="5" pageNumber="40">5.3±0.47</td>
</tr>
<tr id="3550FB345B1200482A101E35FB346B7C" box="[310,1215,1353,1373]" gridrow="35" pageId="5" pageNumber="40" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-5="1">
<td id="768192485B12004829011E35FD0F6B7C" box="[551,644,1353,1373]" gridcol="2" gridrow="35" pageId="5" pageNumber="40">(4.7-6.1)</td>
<td id="768192485B12004828631E35FC2A6B7C" box="[837,929,1353,1373]" gridcol="4" gridrow="35" pageId="5" pageNumber="40">(4.7-5.7)</td>
<td id="768192485B1200482F451E35FB346B7C" box="[1123,1215,1353,1373]" gridcol="6" gridrow="35" pageId="5" pageNumber="40">(4.1-6.2)</td>
</tr>
<tr id="3550FB345B1200482A101E15FB346B5D" box="[310,1215,1385,1404]" gridrow="36" pageId="5" pageNumber="40">
<th id="768192485B1200482A101E15FE0F6B5D" box="[310,388,1385,1404]" gridcol="0" gridrow="36" pageId="5" pageNumber="40">RIOD</th>
<td id="768192485B1200482A861E15FE676B5D" box="[416,492,1385,1404]" gridcol="1" gridrow="36" pageId="5" pageNumber="40">6.5</td>
<td id="768192485B12004829011E15FD0F6B5D" box="[551,644,1385,1404]" gridcol="2" gridrow="36" pageId="5" pageNumber="40">5.8±0.31</td>
<td id="768192485B12004829E61E15FC876B5D" box="[704,780,1385,1404]" gridcol="3" gridrow="36" pageId="5" pageNumber="40">6.2</td>
<td id="768192485B12004828631E15FC2A6B5D" box="[837,929,1385,1404]" gridcol="4" gridrow="36" pageId="5" pageNumber="40">6.4±0.62</td>
<td id="768192485B12004828FB1E15FBA06B5D" box="[989,1067,1385,1404]" gridcol="5" gridrow="36" pageId="5" pageNumber="40">6.0</td>
<td id="768192485B1200482F451E15FB346B5D" box="[1123,1215,1385,1404]" gridcol="6" gridrow="36" pageId="5" pageNumber="40">6.3±0.33</td>
</tr>
<tr id="3550FB345B1200482A101EF5FB346BBD" box="[310,1215,1417,1436]" gridrow="37" pageId="5" pageNumber="40" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-5="1">
<td id="768192485B12004829011EF5FD0F6BBD" box="[551,644,1417,1436]" gridcol="2" gridrow="37" pageId="5" pageNumber="40">(5.1-6.6)</td>
<td id="768192485B12004828631EF5FC2A6BBD" box="[837,929,1417,1436]" gridcol="4" gridrow="37" pageId="5" pageNumber="40">(5.6-7.6)</td>
<td id="768192485B1200482F451EF5FB346BBD" box="[1123,1215,1417,1436]" gridcol="6" gridrow="37" pageId="5" pageNumber="40">(5.7-7.0)</td>
</tr>
<tr id="3550FB345B1200482A101ED5FB346B9D" box="[310,1215,1449,1468]" gridrow="38" pageId="5" pageNumber="40">
<th id="768192485B1200482A101ED5FE0F6B9D" box="[310,388,1449,1468]" gridcol="0" gridrow="38" pageId="5" pageNumber="40">RUEW</th>
<td id="768192485B1200482A861ED5FE676B9D" box="[416,492,1449,1468]" gridcol="1" gridrow="38" pageId="5" pageNumber="40">2.7</td>
<td id="768192485B12004829011ED5FD0F6B9D" box="[551,644,1449,1468]" gridcol="2" gridrow="38" pageId="5" pageNumber="40">2.9±0.20</td>
<td id="768192485B12004829E61ED5FC876B9D" box="[704,780,1449,1468]" gridcol="3" gridrow="38" pageId="5" pageNumber="40">3.3</td>
<td id="768192485B12004828631ED5FC2A6B9D" box="[837,929,1449,1468]" gridcol="4" gridrow="38" pageId="5" pageNumber="40">3.4±0.19</td>
<td id="768192485B12004828FB1ED5FBA06B9D" box="[989,1067,1449,1468]" gridcol="5" gridrow="38" pageId="5" pageNumber="40">2.7</td>
<td id="768192485B1200482F451ED5FB346B9D" box="[1123,1215,1449,1468]" gridcol="6" gridrow="38" pageId="5" pageNumber="40">3.4±0.35</td>
</tr>
<tr id="3550FB345B1200482A101EB5FB346BFD" box="[310,1215,1481,1500]" gridrow="39" pageId="5" pageNumber="40" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-5="1">
<td id="768192485B12004829011EB5FD0F6BFD" box="[551,644,1481,1500]" gridcol="2" gridrow="39" pageId="5" pageNumber="40">(2.6-3.4)</td>
<td id="768192485B12004828631EB5FC2A6BFD" box="[837,929,1481,1500]" gridcol="4" gridrow="39" pageId="5" pageNumber="40">(3.2-3.8)</td>
<td id="768192485B1200482F451EB5FB346BFD" box="[1123,1215,1481,1500]" gridcol="6" gridrow="39" pageId="5" pageNumber="40">(2.6-4.0)</td>
</tr>
<tr id="3550FB345B1200482A101E94FB346BDD" box="[310,1215,1512,1532]" gridrow="40" pageId="5" pageNumber="40">
<th id="768192485B1200482A101E94FE0F6BDD" box="[310,388,1512,1532]" gridcol="0" gridrow="40" pageId="5" pageNumber="40">RSL</th>
<td id="768192485B1200482A861E94FE676BDD" box="[416,492,1512,1532]" gridcol="1" gridrow="40" pageId="5" pageNumber="40">6.8</td>
<td id="768192485B12004829011E94FD0F6BDD" box="[551,644,1512,1532]" gridcol="2" gridrow="40" pageId="5" pageNumber="40">6.3±0.36</td>
<td id="768192485B12004829E61E94FC876BDD" box="[704,780,1512,1532]" gridcol="3" gridrow="40" pageId="5" pageNumber="40">7.1</td>
<td id="768192485B12004828631E94FC2A6BDD" box="[837,929,1512,1532]" gridcol="4" gridrow="40" pageId="5" pageNumber="40">6.2±0.47</td>
<td id="768192485B12004828FB1E94FBA06BDD" box="[989,1067,1512,1532]" gridcol="5" gridrow="40" pageId="5" pageNumber="40">7.2</td>
<td id="768192485B1200482F451E94FB346BDD" box="[1123,1215,1512,1532]" gridcol="6" gridrow="40" pageId="5" pageNumber="40">6.5±0.47</td>
</tr>
<tr id="3550FB345B1200482A101D74FB34683A" box="[310,1215,1544,1563]" gridrow="41" pageId="5" pageNumber="40" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-5="1">
<td id="768192485B12004829011D74FD0F683A" box="[551,644,1544,1563]" gridcol="2" gridrow="41" pageId="5" pageNumber="40">(5.7-6.8)</td>
<td id="768192485B12004828631D74FC2A683A" box="[837,929,1544,1563]" gridcol="4" gridrow="41" pageId="5" pageNumber="40">(5.7-7.1)</td>
<td id="768192485B1200482F451D74FB34683A" box="[1123,1215,1544,1563]" gridcol="6" gridrow="41" pageId="5" pageNumber="40">(5.4-7.3)</td>
</tr>
<tr id="3550FB345B1200482A101D54FB34681D" box="[310,1215,1576,1596]" gridrow="42" pageId="5" pageNumber="40">
<th id="768192485B1200482A101D54FE0F681D" box="[310,388,1576,1596]" gridcol="0" gridrow="42" pageId="5" pageNumber="40">RUEL</th>
<td id="768192485B1200482A861D54FE67681D" box="[416,492,1576,1596]" gridcol="1" gridrow="42" pageId="5" pageNumber="40">4.3</td>
<td id="768192485B12004829011D54FD0F681D" box="[551,644,1576,1596]" gridcol="2" gridrow="42" pageId="5" pageNumber="40">3.9±0.21</td>
<td id="768192485B12004829E61D54FC87681D" box="[704,780,1576,1596]" gridcol="3" gridrow="42" pageId="5" pageNumber="40">5.0</td>
<td id="768192485B12004828631D54FC2A681D" box="[837,929,1576,1596]" gridcol="4" gridrow="42" pageId="5" pageNumber="40">4.7±0.35</td>
<td id="768192485B12004828FB1D54FBA0681D" box="[989,1067,1576,1596]" gridcol="5" gridrow="42" pageId="5" pageNumber="40">4.2</td>
<td id="768192485B1200482F451D54FB34681D" box="[1123,1215,1576,1596]" gridcol="6" gridrow="42" pageId="5" pageNumber="40">4.7±0.31</td>
</tr>
<tr id="3550FB345B1200482A101D34FB34687A" box="[310,1215,1608,1627]" gridrow="43" pageId="5" pageNumber="40" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-5="1">
<td id="768192485B12004829011D34FD0F687A" box="[551,644,1608,1627]" gridcol="2" gridrow="43" pageId="5" pageNumber="40">(3.6-4.4)</td>
<td id="768192485B12004828631D34FC2A687A" box="[837,929,1608,1627]" gridcol="4" gridrow="43" pageId="5" pageNumber="40">(4.4-5.1)</td>
<td id="768192485B1200482F451D34FB34687A" box="[1123,1215,1608,1627]" gridcol="6" gridrow="43" pageId="5" pageNumber="40">(3.8-5.1)</td>
</tr>
<tr id="3550FB345B1200482A101D14FB34685A" box="[310,1215,1640,1659]" gridrow="44" pageId="5" pageNumber="40">
<th id="768192485B1200482A101D14FE0F685A" box="[310,388,1640,1659]" gridcol="0" gridrow="44" pageId="5" pageNumber="40">RLJL</th>
<td id="768192485B1200482A861D14FE67685A" box="[416,492,1640,1659]" gridcol="1" gridrow="44" pageId="5" pageNumber="40">12.5</td>
<td id="768192485B12004829011D14FD0F685A" box="[551,644,1640,1659]" gridcol="2" gridrow="44" pageId="5" pageNumber="40">13.4±0.61</td>
<td id="768192485B12004829E61D14FC87685A" box="[704,780,1640,1659]" gridcol="3" gridrow="44" pageId="5" pageNumber="40">14.3</td>
<td id="768192485B12004828631D14FC2A685A" box="[837,929,1640,1659]" gridcol="4" gridrow="44" pageId="5" pageNumber="40">13.9±0.62</td>
<td id="768192485B12004828FB1D14FBA0685A" box="[989,1067,1640,1659]" gridcol="5" gridrow="44" pageId="5" pageNumber="40">13.6</td>
<td id="768192485B1200482F451D14FB34685A" box="[1123,1215,1640,1659]" gridcol="6" gridrow="44" pageId="5" pageNumber="40">14.3±0.73</td>
</tr>
<tr id="3550FB345B1200482A101DFBFB3468BA" box="[310,1215,1671,1691]" gridrow="45" pageId="5" pageNumber="40" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-5="1">
<td id="768192485B12004829011DFBFD0F68BA" box="[551,644,1671,1691]" gridcol="2" gridrow="45" pageId="5" pageNumber="40">(12.5-14.4)</td>
<td id="768192485B12004828631DFBFC2A68BA" box="[837,929,1671,1691]" gridcol="4" gridrow="45" pageId="5" pageNumber="40">(13.0-14.9)</td>
<td id="768192485B1200482F451DFBFB3468BA" box="[1123,1215,1671,1691]" gridcol="6" gridrow="45" pageId="5" pageNumber="40">(13.1-15.8)</td>
</tr>
</table>
</paragraph>
<paragraph id="8BDFF9765B12FFB22A161DCDFC6B68E6" blockId="5.[304,992,1713,1735]" box="[304,992,1713,1735]" pageId="5" pageNumber="40">
<tableNote id="7686F8F85B12FFB22A161DCDFC6B68E6" box="[304,992,1713,1735]" pageId="5" pageNumber="40" targetBox="[310,1215,240,1691]" targetPageId="5">See Materials and Methods for definitions of morphological characteristics.</tableNote>
</paragraph>
<subSubSection id="C37AAAFD5B12FFB12B891C30FDD36873" lastPageId="6" lastPageNumber="41" pageId="5" pageNumber="40" type="description">
<paragraph id="8BDFF9765B12FFB12B891C30FDD36873" blockId="5.[150,738,1790,2087]" lastBlockId="6.[188,776,1322,2086]" lastPageId="6" lastPageNumber="41" pageId="5" pageNumber="40">
<emphasis id="B91425645B12FFB22B891C30FE256945" bold="true" box="[175,430,1868,1892]" pageId="5" pageNumber="40">
Description of
<typeStatus id="54DB47D45B12FFB22A6A1C30FE216945" box="[332,426,1868,1892]" pageId="5" pageNumber="40" type="holotype">holotype</typeStatus>
.
</emphasis>
A moderately large individual: HL larger than HW; TAL shorter than SVL; body almost cylindrical; rounded snout; gular fold present; tail gradually compressed toward the tip; clearly expanded cloaca; webbing between digits absent; four fingers on each forelimb, order of length II&gt; III&gt; IV&gt; I on both sides; five toes on each hindlimb, order of length III&gt; IV&gt; II&gt; I&gt; V on both sides; V-shaped vomerine teeth; skin smooth and shiny; DBDD present; DWDV and DWDL present (it became unclear after preservation); DTBYLD present; DTBYLV absent; DGM absent. The
<typeStatus id="54DB47D45B12FFB22FD21CE6FAC76993" box="[1268,1356,1946,1970]" pageId="5" pageNumber="40" type="holotype">holotype</typeStatus>
had the following measurements (in mm): SVL =58.4, TRL = 45.7, AGD = 31.6, HL = 12.5, TAL = 50.0, MTAW = 4.0, MTAH = 8.2, BTAW = 8.0, BTAH = 7.3, VTL = 3.2, VTW = 2.8, HW = 10.2, MXHW = 10.4, LFLL = 13.8, RFLL = 13.7, LHLL = 17.8, RHLL = 18.0, L1FL = 1.5, L2FL = 3.2, L3FL = 3.1, L4FL = 1.7, R1FL = 1.3, R2FL = 3.4, R3FL = 2.7, R4FL = 1.6, L1TL = 1.9, L2TL = 3.9, L3TL = 5.3, L4TL = 4.0, L5TL = 1.6, R1TL = 1.6, R2TL = 3.6, R3TL = 5.0, R4TL = 3.7, R5TL = 1.0, IND = 2.9, IOD = 3.8, LUEW = 1.6, RUEW = 1.4, SL =4.0, LUEL =2.5, RUEL = 2.7, LJL = 7.3, CGN = 12.
</paragraph>
</subSubSection>
<caption id="DF1FA9FE5B11FFB12B9B1BDAFE5C6EFA" ID-Table-UUID="DF1FA9FE5B11FFB12B9B1BDAFE5C6EFA" httpUri="http://table.plazi.org/id/DF1FA9FE5B11FFB12B9B1BDAFE5C6EFA" pageId="6" pageNumber="41" startId="6.[189,242,166,188]" targetBox="[199,739,239,1115]" targetIsTable="true" targetPageId="6" targetType="table">
<paragraph id="8BDFF9765B11FFB12B9B1BDAFE5C6EFA" blockId="6.[189,774,166,219]" pageId="6" pageNumber="41">Table 3. Significant values of the 22 morphological traits in males among three species</paragraph>
</caption>
<paragraph id="8BDFF9765B11FFB129221B93FD3F6A7A" pageId="6" pageNumber="41">
<table id="F9600BD65B1100482BE11B93FD686A7A" box="[199,739,239,1115]" gridcols="4" gridrows="29" pageId="6" pageNumber="41">
<tr id="3550FB345B1100482BE11B93FD686F21" box="[199,739,239,256]" gridrow="0" pageId="6" pageNumber="41">
<th id="768192485B1100482BE11B93FD686F21" box="[199,739,239,256]" colspan="4" colspanRight="3" gridcol="0" gridrow="0" pageId="6" pageNumber="41">Male</th>
</tr>
<tr id="3550FB345B1100482BE11A73FD686F01" box="[199,739,271,288]" gridrow="1" pageId="6" pageNumber="41">
<th id="768192485B1100482BE11A73FE9C6F01" box="[199,279,271,288]" gridcol="0" gridrow="1" pageId="6" pageNumber="41">Trait</th>
<td id="768192485B1100482A6D1A73FE3B6F01" box="[331,432,271,288]" gridcol="1" gridrow="1" pageId="6" pageNumber="41">GEI vs. SUM</td>
<td id="768192485B1100482ACE1A73FDCD6F01" box="[488,582,271,288]" gridcol="2" gridrow="1" pageId="6" pageNumber="41">GEI vs. AKI</td>
<td id="768192485B110048295B1A73FD686F01" box="[637,739,271,288]" gridcol="3" gridrow="1" pageId="6" pageNumber="41">SUM vs. AKI</td>
</tr>
<tr id="3550FB345B1100482BE11A51FD686F1F" box="[199,739,301,318]" gridrow="2" pageId="6" pageNumber="41">
<th id="768192485B1100482BE11A51FE9C6F1F" box="[199,279,301,318]" gridcol="0" gridrow="2" pageId="6" pageNumber="41">SVL</th>
<td id="768192485B1100482A6D1A51FE3B6F1F" box="[331,432,301,318]" gridcol="1" gridrow="2" pageId="6" pageNumber="41">
<emphasis id="B91425645B11FFB12A7F1A51FE2A6F1F" bold="true" box="[345,417,301,318]" pageId="6" pageNumber="41">P &lt;0.001</emphasis>
</td>
<td id="768192485B1100482ACE1A51FDCD6F1F" box="[488,582,301,318]" gridcol="2" gridrow="2" pageId="6" pageNumber="41">
<emphasis id="B91425645B11FFB12AC81A51FDB46F1F" bold="true" box="[494,575,301,318]" pageId="6" pageNumber="41">P &lt;0.0001</emphasis>
</td>
<td id="768192485B110048295B1A51FD686F1F" box="[637,739,301,318]" gridcol="3" gridrow="2" pageId="6" pageNumber="41">P &lt;0.05</td>
</tr>
<tr id="3550FB345B1100482BE11A30FD686F7C" box="[199,739,332,349]" gridrow="3" pageId="6" pageNumber="41">
<th id="768192485B1100482BE11A30FE9C6F7C" box="[199,279,332,349]" gridcol="0" gridrow="3" pageId="6" pageNumber="41">RTRL</th>
<td id="768192485B1100482A6D1A30FE3B6F7C" box="[331,432,332,349]" gridcol="1" gridrow="3" pageId="6" pageNumber="41">NS</td>
<td id="768192485B1100482ACE1A30FDCD6F7C" box="[488,582,332,349]" gridcol="2" gridrow="3" pageId="6" pageNumber="41">NS</td>
<td id="768192485B110048295B1A30FD686F7C" box="[637,739,332,349]" gridcol="3" gridrow="3" pageId="6" pageNumber="41">NS</td>
</tr>
<tr id="3550FB345B1100482BE11A17FD686F5D" box="[199,739,363,380]" gridrow="4" pageId="6" pageNumber="41">
<th id="768192485B1100482BE11A17FE9C6F5D" box="[199,279,363,380]" gridcol="0" gridrow="4" pageId="6" pageNumber="41">RAGD</th>
<td id="768192485B1100482A6D1A17FE3B6F5D" box="[331,432,363,380]" gridcol="1" gridrow="4" pageId="6" pageNumber="41">NS</td>
<td id="768192485B1100482ACE1A17FDCD6F5D" box="[488,582,363,380]" gridcol="2" gridrow="4" pageId="6" pageNumber="41">NS</td>
<td id="768192485B110048295B1A17FD686F5D" box="[637,739,363,380]" gridcol="3" gridrow="4" pageId="6" pageNumber="41">NS</td>
</tr>
<tr id="3550FB345B1100482BE11AF5FD686FBB" box="[199,739,393,410]" gridrow="5" pageId="6" pageNumber="41">
<th id="768192485B1100482BE11AF5FE9C6FBB" box="[199,279,393,410]" gridcol="0" gridrow="5" pageId="6" pageNumber="41">RHL</th>
<td id="768192485B1100482A6D1AF5FE3B6FBB" box="[331,432,393,410]" gridcol="1" gridrow="5" pageId="6" pageNumber="41">P &lt;0.05</td>
<td id="768192485B1100482ACE1AF5FDCD6FBB" box="[488,582,393,410]" gridcol="2" gridrow="5" pageId="6" pageNumber="41">P &lt;0.01</td>
<td id="768192485B110048295B1AF5FD686FBB" box="[637,739,393,410]" gridcol="3" gridrow="5" pageId="6" pageNumber="41">NS</td>
</tr>
<tr id="3550FB345B1100482BE11AD4FD686F98" box="[199,739,424,441]" gridrow="6" pageId="6" pageNumber="41">
<th id="768192485B1100482BE11AD4FE9C6F98" box="[199,279,424,441]" gridcol="0" gridrow="6" pageId="6" pageNumber="41">RTAL</th>
<td id="768192485B1100482A6D1AD4FE3B6F98" box="[331,432,424,441]" gridcol="1" gridrow="6" pageId="6" pageNumber="41">P &lt;0.05</td>
<td id="768192485B1100482ACE1AD4FDCD6F98" box="[488,582,424,441]" gridcol="2" gridrow="6" pageId="6" pageNumber="41">
<emphasis id="B91425645B11FFB12AD51AD4FDB16F98" bold="true" box="[499,570,424,441]" pageId="6" pageNumber="41">P &lt;0.001</emphasis>
</td>
<td id="768192485B110048295B1AD4FD686F98" box="[637,739,424,441]" gridcol="3" gridrow="6" pageId="6" pageNumber="41">NS</td>
</tr>
<tr id="3550FB345B1100482BE11ABAFD686FF6" box="[199,739,454,471]" gridrow="7" pageId="6" pageNumber="41">
<th id="768192485B1100482BE11ABAFE9C6FF6" box="[199,279,454,471]" gridcol="0" gridrow="7" pageId="6" pageNumber="41">RMTAW</th>
<td id="768192485B1100482A6D1ABAFE3B6FF6" box="[331,432,454,471]" gridcol="1" gridrow="7" pageId="6" pageNumber="41">NS</td>
<td id="768192485B1100482ACE1ABAFDCD6FF6" box="[488,582,454,471]" gridcol="2" gridrow="7" pageId="6" pageNumber="41">NS</td>
<td id="768192485B110048295B1ABAFD686FF6" box="[637,739,454,471]" gridcol="3" gridrow="7" pageId="6" pageNumber="41">NS</td>
</tr>
<tr id="3550FB345B1100482BE11A99FD686FD7" box="[199,739,485,502]" gridrow="8" pageId="6" pageNumber="41">
<th id="768192485B1100482BE11A99FE9C6FD7" box="[199,279,485,502]" gridcol="0" gridrow="8" pageId="6" pageNumber="41">RMTAH</th>
<td id="768192485B1100482A6D1A99FE3B6FD7" box="[331,432,485,502]" gridcol="1" gridrow="8" pageId="6" pageNumber="41">NS</td>
<td id="768192485B1100482ACE1A99FDCD6FD7" box="[488,582,485,502]" gridcol="2" gridrow="8" pageId="6" pageNumber="41">
<emphasis id="B91425645B11FFB12AD51A99FDB16FD7" bold="true" box="[499,570,485,502]" pageId="6" pageNumber="41">P &lt;0.001</emphasis>
</td>
<td id="768192485B110048295B1A99FD686FD7" box="[637,739,485,502]" gridcol="3" gridrow="8" pageId="6" pageNumber="41">NS</td>
</tr>
<tr id="3550FB345B1100482BE11978FD686C34" box="[199,739,516,533]" gridrow="9" pageId="6" pageNumber="41">
<th id="768192485B1100482BE11978FE9C6C34" box="[199,279,516,533]" gridcol="0" gridrow="9" pageId="6" pageNumber="41">RVTL</th>
<td id="768192485B1100482A6D1978FE3B6C34" box="[331,432,516,533]" gridcol="1" gridrow="9" pageId="6" pageNumber="41">NS</td>
<td id="768192485B1100482ACE1978FDCD6C34" box="[488,582,516,533]" gridcol="2" gridrow="9" pageId="6" pageNumber="41">NS</td>
<td id="768192485B110048295B1978FD686C34" box="[637,739,516,533]" gridcol="3" gridrow="9" pageId="6" pageNumber="41">P &lt;0.05</td>
</tr>
<tr id="3550FB345B1100482BE1195EFD686C12" box="[199,739,546,563]" gridrow="10" pageId="6" pageNumber="41">
<th id="768192485B1100482BE1195EFE9C6C12" box="[199,279,546,563]" gridcol="0" gridrow="10" pageId="6" pageNumber="41">RVTW</th>
<td id="768192485B1100482A6D195EFE3B6C12" box="[331,432,546,563]" gridcol="1" gridrow="10" pageId="6" pageNumber="41">NS</td>
<td id="768192485B1100482ACE195EFDCD6C12" box="[488,582,546,563]" gridcol="2" gridrow="10" pageId="6" pageNumber="41">NS</td>
<td id="768192485B110048295B195EFD686C12" box="[637,739,546,563]" gridcol="3" gridrow="10" pageId="6" pageNumber="41">NS</td>
</tr>
<tr id="3550FB345B1100482BE1193DFD686C73" box="[199,739,577,594]" gridrow="11" pageId="6" pageNumber="41">
<th id="768192485B1100482BE1193DFE9C6C73" box="[199,279,577,594]" gridcol="0" gridrow="11" pageId="6" pageNumber="41">RHW</th>
<td id="768192485B1100482A6D193DFE3B6C73" box="[331,432,577,594]" gridcol="1" gridrow="11" pageId="6" pageNumber="41">NS</td>
<td id="768192485B1100482ACE193DFDCD6C73" box="[488,582,577,594]" gridcol="2" gridrow="11" pageId="6" pageNumber="41">NS</td>
<td id="768192485B110048295B193DFD686C73" box="[637,739,577,594]" gridcol="3" gridrow="11" pageId="6" pageNumber="41">NS</td>
</tr>
<tr id="3550FB345B1100482BE1191CFD686C50" box="[199,739,608,625]" gridrow="12" pageId="6" pageNumber="41">
<th id="768192485B1100482BE1191CFE9C6C50" box="[199,279,608,625]" gridcol="0" gridrow="12" pageId="6" pageNumber="41">RFLL</th>
<td id="768192485B1100482A6D191CFE3B6C50" box="[331,432,608,625]" gridcol="1" gridrow="12" pageId="6" pageNumber="41">P &lt;0.01</td>
<td id="768192485B1100482ACE191CFDCD6C50" box="[488,582,608,625]" gridcol="2" gridrow="12" pageId="6" pageNumber="41">
<emphasis id="B91425645B11FFB12AC8191CFDB46C50" bold="true" box="[494,575,608,625]" pageId="6" pageNumber="41">P &lt;0.0001</emphasis>
</td>
<td id="768192485B110048295B191CFD686C50" box="[637,739,608,625]" gridcol="3" gridrow="12" pageId="6" pageNumber="41">NS</td>
</tr>
<tr id="3550FB345B1100482BE11902FD686CAE" box="[199,739,638,655]" gridrow="13" pageId="6" pageNumber="41">
<th id="768192485B1100482BE11902FE9C6CAE" box="[199,279,638,655]" gridcol="0" gridrow="13" pageId="6" pageNumber="41">RHLL</th>
<td id="768192485B1100482A6D1902FE3B6CAE" box="[331,432,638,655]" gridcol="1" gridrow="13" pageId="6" pageNumber="41">NS</td>
<td id="768192485B1100482ACE1902FDCD6CAE" box="[488,582,638,655]" gridcol="2" gridrow="13" pageId="6" pageNumber="41">NS</td>
<td id="768192485B110048295B1902FD686CAE" box="[637,739,638,655]" gridcol="3" gridrow="13" pageId="6" pageNumber="41">NS</td>
</tr>
<tr id="3550FB345B1100482BE119E1FD686C8F" box="[199,739,669,686]" gridrow="14" pageId="6" pageNumber="41">
<th id="768192485B1100482BE119E1FE9C6C8F" box="[199,279,669,686]" gridcol="0" gridrow="14" pageId="6" pageNumber="41">R2FL</th>
<td id="768192485B1100482A6D19E1FE3B6C8F" box="[331,432,669,686]" gridcol="1" gridrow="14" pageId="6" pageNumber="41">NS</td>
<td id="768192485B1100482ACE19E1FDCD6C8F" box="[488,582,669,686]" gridcol="2" gridrow="14" pageId="6" pageNumber="41">NS</td>
<td id="768192485B110048295B19E1FD686C8F" box="[637,739,669,686]" gridcol="3" gridrow="14" pageId="6" pageNumber="41">NS</td>
</tr>
<tr id="3550FB345B1100482BE119C0FD686CEC" box="[199,739,700,717]" gridrow="15" pageId="6" pageNumber="41">
<th id="768192485B1100482BE119C0FE9C6CEC" box="[199,279,700,717]" gridcol="0" gridrow="15" pageId="6" pageNumber="41">R3FL</th>
<td id="768192485B1100482A6D19C0FE3B6CEC" box="[331,432,700,717]" gridcol="1" gridrow="15" pageId="6" pageNumber="41">P &lt;0.01</td>
<td id="768192485B1100482ACE19C0FDCD6CEC" box="[488,582,700,717]" gridcol="2" gridrow="15" pageId="6" pageNumber="41">NS</td>
<td id="768192485B110048295B19C0FD686CEC" box="[637,739,700,717]" gridcol="3" gridrow="15" pageId="6" pageNumber="41">P &lt;0.01</td>
</tr>
<tr id="3550FB345B1100482BE119A6FD686CCA" box="[199,739,730,747]" gridrow="16" pageId="6" pageNumber="41">
<th id="768192485B1100482BE119A6FE9C6CCA" box="[199,279,730,747]" gridcol="0" gridrow="16" pageId="6" pageNumber="41">R3TL</th>
<td id="768192485B1100482A6D19A6FE3B6CCA" box="[331,432,730,747]" gridcol="1" gridrow="16" pageId="6" pageNumber="41">P &lt;0.05</td>
<td id="768192485B1100482ACE19A6FDCD6CCA" box="[488,582,730,747]" gridcol="2" gridrow="16" pageId="6" pageNumber="41">NS</td>
<td id="768192485B110048295B19A6FD686CCA" box="[637,739,730,747]" gridcol="3" gridrow="16" pageId="6" pageNumber="41">NS</td>
</tr>
<tr id="3550FB345B1100482BE11985FD686D2B" box="[199,739,761,778]" gridrow="17" pageId="6" pageNumber="41">
<th id="768192485B1100482BE11985FE9C6D2B" box="[199,279,761,778]" gridcol="0" gridrow="17" pageId="6" pageNumber="41">R5TL</th>
<td id="768192485B1100482A6D1985FE3B6D2B" box="[331,432,761,778]" gridcol="1" gridrow="17" pageId="6" pageNumber="41">
<emphasis id="B91425645B11FFB12A731985FE2D6D2B" bold="true" box="[341,422,761,778]" pageId="6" pageNumber="41">P &lt;0.0001</emphasis>
</td>
<td id="768192485B1100482ACE1985FDCD6D2B" box="[488,582,761,778]" gridcol="2" gridrow="17" pageId="6" pageNumber="41">
<emphasis id="B91425645B11FFB12AC81985FDB46D2B" bold="true" box="[494,575,761,778]" pageId="6" pageNumber="41">P &lt;0.0001</emphasis>
</td>
<td id="768192485B110048295B1985FD686D2B" box="[637,739,761,778]" gridcol="3" gridrow="17" pageId="6" pageNumber="41">NS</td>
</tr>
<tr id="3550FB345B1100482BE1186BFD686D09" box="[199,739,791,808]" gridrow="18" pageId="6" pageNumber="41">
<th id="768192485B1100482BE1186BFE9C6D09" box="[199,279,791,808]" gridcol="0" gridrow="18" pageId="6" pageNumber="41">RIND</th>
<td id="768192485B1100482A6D186BFE3B6D09" box="[331,432,791,808]" gridcol="1" gridrow="18" pageId="6" pageNumber="41">NS</td>
<td id="768192485B1100482ACE186BFDCD6D09" box="[488,582,791,808]" gridcol="2" gridrow="18" pageId="6" pageNumber="41">NS</td>
<td id="768192485B110048295B186BFD686D09" box="[637,739,791,808]" gridcol="3" gridrow="18" pageId="6" pageNumber="41">NS</td>
</tr>
<tr id="3550FB345B1100482BE1184AFD686D66" box="[199,739,822,839]" gridrow="19" pageId="6" pageNumber="41">
<th id="768192485B1100482BE1184AFE9C6D66" box="[199,279,822,839]" gridcol="0" gridrow="19" pageId="6" pageNumber="41">RIOD</th>
<td id="768192485B1100482A6D184AFE3B6D66" box="[331,432,822,839]" gridcol="1" gridrow="19" pageId="6" pageNumber="41">NS</td>
<td id="768192485B1100482ACE184AFDCD6D66" box="[488,582,822,839]" gridcol="2" gridrow="19" pageId="6" pageNumber="41">
<emphasis id="B91425645B11FFB12AC8184AFDB46D66" bold="true" box="[494,575,822,839]" pageId="6" pageNumber="41">P &lt;0.0001</emphasis>
</td>
<td id="768192485B110048295B184AFD686D66" box="[637,739,822,839]" gridcol="3" gridrow="19" pageId="6" pageNumber="41">NS</td>
</tr>
<tr id="3550FB345B1100482BE11829FD686D47" box="[199,739,853,870]" gridrow="20" pageId="6" pageNumber="41">
<th id="768192485B1100482BE11829FE9C6D47" box="[199,279,853,870]" gridcol="0" gridrow="20" pageId="6" pageNumber="41">RUEW</th>
<td id="768192485B1100482A6D1829FE3B6D47" box="[331,432,853,870]" gridcol="1" gridrow="20" pageId="6" pageNumber="41">
<emphasis id="B91425645B11FFB12A7F1829FE2A6D47" bold="true" box="[345,417,853,870]" pageId="6" pageNumber="41">P &lt;0.001</emphasis>
</td>
<td id="768192485B1100482ACE1829FDCD6D47" box="[488,582,853,870]" gridcol="2" gridrow="20" pageId="6" pageNumber="41">
<emphasis id="B91425645B11FFB12AC81829FDB46D47" bold="true" box="[494,575,853,870]" pageId="6" pageNumber="41">P &lt;0.0001</emphasis>
</td>
<td id="768192485B110048295B1829FD686D47" box="[637,739,853,870]" gridcol="3" gridrow="20" pageId="6" pageNumber="41">NS</td>
</tr>
<tr id="3550FB345B1100482BE1180FFD686DA5" box="[199,739,883,900]" gridrow="21" pageId="6" pageNumber="41">
<th id="768192485B1100482BE1180FFE9C6DA5" box="[199,279,883,900]" gridcol="0" gridrow="21" pageId="6" pageNumber="41">RSL</th>
<td id="768192485B1100482A6D180FFE3B6DA5" box="[331,432,883,900]" gridcol="1" gridrow="21" pageId="6" pageNumber="41">NS</td>
<td id="768192485B1100482ACE180FFDCD6DA5" box="[488,582,883,900]" gridcol="2" gridrow="21" pageId="6" pageNumber="41">NS</td>
<td id="768192485B110048295B180FFD686DA5" box="[637,739,883,900]" gridcol="3" gridrow="21" pageId="6" pageNumber="41">NS</td>
</tr>
<tr id="3550FB345B1100482BE118EEFD686D82" box="[199,739,914,931]" gridrow="22" pageId="6" pageNumber="41">
<th id="768192485B1100482BE118EEFE9C6D82" box="[199,279,914,931]" gridcol="0" gridrow="22" pageId="6" pageNumber="41">RUEL</th>
<td id="768192485B1100482A6D18EEFE3B6D82" box="[331,432,914,931]" gridcol="1" gridrow="22" pageId="6" pageNumber="41">
<emphasis id="B91425645B11FFB12A7F18EEFE2A6D82" bold="true" box="[345,417,914,931]" pageId="6" pageNumber="41">P &lt;0.001</emphasis>
</td>
<td id="768192485B1100482ACE18EEFDCD6D82" box="[488,582,914,931]" gridcol="2" gridrow="22" pageId="6" pageNumber="41">
<emphasis id="B91425645B11FFB12AC818EEFDB46D82" bold="true" box="[494,575,914,931]" pageId="6" pageNumber="41">P &lt;0.0001</emphasis>
</td>
<td id="768192485B110048295B18EEFD686D82" box="[637,739,914,931]" gridcol="3" gridrow="22" pageId="6" pageNumber="41">NS</td>
</tr>
<tr id="3550FB345B1100482BE118CDFD686DE3" box="[199,739,945,962]" gridrow="23" pageId="6" pageNumber="41">
<th id="768192485B1100482BE118CDFE9C6DE3" box="[199,279,945,962]" gridcol="0" gridrow="23" pageId="6" pageNumber="41">RLJL</th>
<td id="768192485B1100482A6D18CDFE3B6DE3" box="[331,432,945,962]" gridcol="1" gridrow="23" pageId="6" pageNumber="41">NS</td>
<td id="768192485B1100482ACE18CDFDCD6DE3" box="[488,582,945,962]" gridcol="2" gridrow="23" pageId="6" pageNumber="41">
<emphasis id="B91425645B11FFB12AD418CDFDB16DE3" bold="true" box="[498,570,945,962]" pageId="6" pageNumber="41">P &lt;0.001</emphasis>
</td>
<td id="768192485B110048295B18CDFD686DE3" box="[637,739,945,962]" gridcol="3" gridrow="23" pageId="6" pageNumber="41">NS</td>
</tr>
<tr id="3550FB345B1100482BE118B3FD686DC1" box="[199,739,975,992]" gridrow="24" pageId="6" pageNumber="41">
<th id="768192485B1100482BE118B3FE9C6DC1" box="[199,279,975,992]" gridcol="0" gridrow="24" pageId="6" pageNumber="41">P &lt;0.05</th>
<td id="768192485B1100482A6D18B3FE3B6DC1" box="[331,432,975,992]" gridcol="1" gridrow="24" pageId="6" pageNumber="41">3</td>
<td id="768192485B1100482ACE18B3FDCD6DC1" box="[488,582,975,992]" gridcol="2" gridrow="24" pageId="6" pageNumber="41">0</td>
<td id="768192485B110048295B18B3FD686DC1" box="[637,739,975,992]" gridcol="3" gridrow="24" pageId="6" pageNumber="41">2</td>
</tr>
<tr id="3550FB345B1100482BE11892FD686DDE" box="[199,739,1006,1023]" gridrow="25" pageId="6" pageNumber="41">
<th id="768192485B1100482BE11892FE9C6DDE" box="[199,279,1006,1023]" gridcol="0" gridrow="25" pageId="6" pageNumber="41">P &lt;0.01</th>
<td id="768192485B1100482A6D1892FE3B6DDE" box="[331,432,1006,1023]" gridcol="1" gridrow="25" pageId="6" pageNumber="41">2</td>
<td id="768192485B1100482ACE1892FDCD6DDE" box="[488,582,1006,1023]" gridcol="2" gridrow="25" pageId="6" pageNumber="41">1</td>
<td id="768192485B110048295B1892FD686DDE" box="[637,739,1006,1023]" gridcol="3" gridrow="25" pageId="6" pageNumber="41">1</td>
</tr>
<tr id="3550FB345B1100482BE11F71FD686A3F" box="[199,739,1037,1054]" gridrow="26" pageId="6" pageNumber="41">
<th id="768192485B1100482BE11F71FE9C6A3F" box="[199,279,1037,1054]" gridcol="0" gridrow="26" pageId="6" pageNumber="41">P &lt;0.001</th>
<td id="768192485B1100482A6D1F71FE3B6A3F" box="[331,432,1037,1054]" gridcol="1" gridrow="26" pageId="6" pageNumber="41">3</td>
<td id="768192485B1100482ACE1F71FDCD6A3F" box="[488,582,1037,1054]" gridcol="2" gridrow="26" pageId="6" pageNumber="41">3</td>
<td id="768192485B110048295B1F71FD686A3F" box="[637,739,1037,1054]" gridcol="3" gridrow="26" pageId="6" pageNumber="41">0</td>
</tr>
<tr id="3550FB345B1100482BE11F57FD686A1D" box="[199,739,1067,1084]" gridrow="27" pageId="6" pageNumber="41">
<th id="768192485B1100482BE11F57FE9C6A1D" box="[199,279,1067,1084]" gridcol="0" gridrow="27" pageId="6" pageNumber="41">P &lt;0.0001</th>
<td id="768192485B1100482A6D1F57FE3B6A1D" box="[331,432,1067,1084]" gridcol="1" gridrow="27" pageId="6" pageNumber="41">1</td>
<td id="768192485B1100482ACE1F57FDCD6A1D" box="[488,582,1067,1084]" gridcol="2" gridrow="27" pageId="6" pageNumber="41">6</td>
<td id="768192485B110048295B1F57FD686A1D" box="[637,739,1067,1084]" gridcol="3" gridrow="27" pageId="6" pageNumber="41">0</td>
</tr>
<tr id="3550FB345B1100482BE11F36FD686A7A" box="[199,739,1098,1115]" gridrow="28" pageId="6" pageNumber="41">
<th id="768192485B1100482BE11F36FE9C6A7A" box="[199,279,1098,1115]" gridcol="0" gridrow="28" pageId="6" pageNumber="41">Total</th>
<td id="768192485B1100482A6D1F36FE3B6A7A" box="[331,432,1098,1115]" gridcol="1" gridrow="28" pageId="6" pageNumber="41">9</td>
<td id="768192485B1100482ACE1F36FDCD6A7A" box="[488,582,1098,1115]" gridcol="2" gridrow="28" pageId="6" pageNumber="41">10</td>
<td id="768192485B110048295B1F36FD686A7A" box="[637,739,1098,1115]" gridcol="3" gridrow="28" pageId="6" pageNumber="41">3</td>
</tr>
</table>
</paragraph>
<paragraph id="8BDFF9765B11FFB12B981F0FFECD6AD4" blockId="6.[189,775,1138,1269]" pageId="6" pageNumber="41">
<tableNote id="7686F8F85B11FFB12B981F0FFECD6AD4" pageId="6" pageNumber="41" targetBox="[199,739,239,1115]" targetPageId="6">
GEI, SUM, and AKI are abbreviations for
<taxonomicName id="4C6082F55B11FFB1296D1F0EFC8C6AA9" authority="Sugawara &amp; Naito &amp; Iwata &amp; Nagano, 2022" authorityName="Sugawara &amp; Naito &amp; Iwata &amp; Nagano" authorityYear="2022" box="[587,775,1138,1160]" class="Amphibia" family="Hynobiidae" genus="Hynobius" kingdom="Animalia" order="Caudata" pageId="6" pageNumber="41" phylum="Chordata" rank="species" species="geiyoensis" status="sp. nov.">
<emphasis id="B91425645B11FFB1296D1F0EFC8C6AA9" box="[587,775,1138,1160]" italics="true" pageId="6" pageNumber="41">Hynobius geiyoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A227981F5B11FFB12B9B1FF2FE866A82" box="[189,269,1166,1187]" pageId="6" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C6082F55B11FFB12A3C1FF2FE0E6A83" authority="Sugawara &amp; Naito &amp; Iwata &amp; Nagano, 2022" authorityName="Sugawara &amp; Naito &amp; Iwata &amp; Nagano" authorityYear="2022" box="[282,389,1165,1187]" class="Amphibia" family="Hynobiidae" genus="Hynobius" kingdom="Animalia" order="Caudata" pageId="6" pageNumber="41" phylum="Chordata" rank="species" species="sumidai" status="sp. nov.">
<emphasis id="B91425645B11FFB12A3C1FF2FE0E6A83" box="[282,389,1165,1187]" italics="true" pageId="6" pageNumber="41">H. sumidai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A227981F5B11FFB12AAA1FF2FE506A82" box="[396,475,1166,1187]" pageId="6" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
, and
<taxonomicName id="4C6082F55B11FFB129321FF2FDF46A83" box="[532,639,1165,1187]" class="Amphibia" family="Hynobiidae" genus="Hynobius" kingdom="Animalia" order="Caudata" pageId="6" pageNumber="41" phylum="Chordata" rank="species" species="akiensis">
<emphasis id="B91425645B11FFB129321FF2FDF46A83" box="[532,639,1165,1187]" italics="true" pageId="6" pageNumber="41">H. akiensis</emphasis>
</taxonomicName>
, respectively. Significant differences (P &lt;0.001) are shown in bold. See Materials and Methods for definitions of morphological characteristics.
</tableNote>
</paragraph>
<caption id="DF1FA9FE5B11FFB128751E06FBBB684D" ID-DOI="http://doi.org/10.5281/zenodo.11281025" ID-Zenodo-Dep="11281025" httpUri="https://zenodo.org/record/11281025/files/figure.png" pageId="6" pageNumber="41" startId="6.[851,886,1402,1424]" targetBox="[856,1436,156,1378]" targetPageId="6" targetType="figure">
<paragraph id="8BDFF9765B11FFB128751E06FBBB684D" blockId="6.[851,1437,1401,1644]" pageId="6" pageNumber="41">
Fig. 4. Holotype of
<taxonomicName id="4C6082F55B11FFB12F3B1E05FB6D6BAE" authority="Sugawara &amp; Naito &amp; Iwata &amp; Nagano, 2022" authorityName="Sugawara &amp; Naito &amp; Iwata &amp; Nagano" authorityYear="2022" box="[1053,1254,1401,1423]" class="Amphibia" family="Hynobiidae" genus="Hynobius" kingdom="Animalia" order="Caudata" pageId="6" pageNumber="41" phylum="Chordata" rank="species" species="geiyoensis" status="sp. nov.">
<emphasis id="B91425645B11FFB12F3B1E05FB6D6BAE" box="[1053,1254,1401,1423]" italics="true" pageId="6" pageNumber="41">Hynobius geiyoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A227981F5B11FFB12FCA1E06FAB16BAE" box="[1260,1338,1402,1423]" pageId="6" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
(HMNH-AM-101, adult male, 58.4 mm SVL) from the (A) dorsal and (B) ventral perspective; holotype of
<taxonomicName id="4C6082F55B11FFB12E291EC5FAF16BEC" authority="Sugawara &amp; Naito &amp; Iwata &amp; Nagano, 2022" authorityName="Sugawara &amp; Naito &amp; Iwata &amp; Nagano" authorityYear="2022" box="[1295,1402,1464,1486]" class="Amphibia" family="Hynobiidae" genus="Hynobius" kingdom="Animalia" order="Caudata" pageId="6" pageNumber="41" phylum="Chordata" rank="species" species="sumidai" status="sp. nov.">
<emphasis id="B91425645B11FFB12E291EC5FAF16BEC" box="[1295,1402,1464,1486]" italics="true" pageId="6" pageNumber="41">H. sumidai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A227981F5B11FFB12EA71EC5FC2A6BCF" pageId="6" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
(HMNH-AM-102, adult male, 48.2 mm SVL) from the (C) dorsal and (D) ventral perspective; and a specimen from the type locality (topotype) of
<taxonomicName id="4C6082F55B11FFB12FC21D6BFADA680D" box="[1252,1361,1559,1580]" class="Amphibia" family="Hynobiidae" genus="Hynobius" kingdom="Animalia" order="Caudata" pageId="6" pageNumber="41" phylum="Chordata" rank="species" species="akiensis">
<emphasis id="B91425645B11FFB12FC21D6BFADA680D" box="[1252,1361,1559,1580]" italics="true" pageId="6" pageNumber="41">H. akiensis</emphasis>
</taxonomicName>
(KPM-NFA 946, adult male, 55.3 mm) from the (E) dorsal and (F) ventral perspective.
</paragraph>
</caption>
<subSubSection id="C37AAAFD5B11FFB12BF31D1DFBFB68E8" pageId="6" pageNumber="41" type="diagnosis">
<paragraph id="8BDFF9765B11FFB12BF31D1DFBFB68E8" blockId="6.[188,776,1322,2086]" lastBlockId="6.[850,1436,1712,2087]" pageId="6" pageNumber="41">
<emphasis id="B91425645B11FFB12BF31D1DFEE06858" bold="true" box="[213,363,1633,1657]" pageId="6" pageNumber="41">Comparisons.</emphasis>
The new species statistically differs from
<taxonomicName id="4C6082F55B11FFB12B9A1DF5FEA76881" box="[188,300,1672,1696]" class="Amphibia" family="Hynobiidae" genus="Hynobius" kingdom="Animalia" order="Caudata" pageId="6" pageNumber="41" phylum="Chordata" rank="species" species="akiensis">
<emphasis id="B91425645B11FFB12B9A1DF5FEA76881" box="[188,300,1672,1696]" italics="true" pageId="6" pageNumber="41">H. akiensis</emphasis>
</taxonomicName>
in terms of the following length measurements in males: SVL, RHL, RTAL, RMTAH, RFLL, R5TL, RIOD, RUEW, RUEL, and RLJL; except for SVL, RTAL, RMTAH, and R5TL, these lengths were significantly shorter in
<taxonomicName id="4C6082F55B11FFB12A391C59FE2E691D" authority="Sugawara &amp; Naito &amp; Iwata &amp; Nagano, 2022" authorityName="Sugawara &amp; Naito &amp; Iwata &amp; Nagano" authorityYear="2022" box="[287,421,1829,1852]" class="Amphibia" family="Hynobiidae" genus="Hynobius" kingdom="Animalia" order="Caudata" pageId="6" pageNumber="41" phylum="Chordata" rank="species" species="geiyoensis" status="sp. nov.">
<emphasis id="B91425645B11FFB12A391C59FE2E691D" box="[287,421,1829,1852]" italics="true" pageId="6" pageNumber="41">H. geiyoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A227981F5B11FFB12A8D1C58FE7C691A" box="[427,503,1828,1851]" pageId="6" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
relative to the lengths in
<taxonomicName id="4C6082F55B11FFB129CB1C59FE856942" class="Amphibia" family="Hynobiidae" genus="Hynobius" kingdom="Animalia" order="Caudata" pageId="6" pageNumber="41" phylum="Chordata" rank="species" species="akiensis">
<emphasis id="B91425645B11FFB129CB1C59FE856942" italics="true" pageId="6" pageNumber="41">H. akiensis</emphasis>
</taxonomicName>
. The new species differs from
<taxonomicName id="4C6082F55B11FFB1296A1C30FD356942" box="[588,702,1867,1891]" class="Amphibia" family="Hynobiidae" genus="Hynobius" kingdom="Animalia" order="Caudata" pageId="6" pageNumber="41" phylum="Chordata" rank="species" species="akiensis">
<emphasis id="B91425645B11FFB1296A1C30FD356942" box="[588,702,1867,1891]" italics="true" pageId="6" pageNumber="41">H. akiensis</emphasis>
</taxonomicName>
by the following characters: SVL&gt;
<quantity id="4C9854935B11FFB12AFA1C0EFDBD69A8" box="[476,566,1906,1930]" metricMagnitude="-2" metricUnit="m" metricValue="5.71" pageId="6" pageNumber="41" unit="mm" value="57.1">57.1 mm</quantity>
(18/19 = 94.7 %) vs. SVL &lt;
<quantity id="4C9854935B11FFB12A211CE5FEE96991" box="[263,354,1945,1969]" metricMagnitude="-2" metricUnit="m" metricValue="5.72" pageId="6" pageNumber="41" unit="mm" value="57.2">57.2 mm</quantity>
(26/28 = 92.9 %); RUEW &lt;3.2 % (18/19 = 94.7 %) vs. RUEW&gt; 3.1 % (23/28 = 82.1 %); RUEL &lt;4.2 % (17/19 = 89.5 %) vs. RUEL&gt; 4.1 % (27/28 = 96.4 %); usually have DTBYLD (17/19 = 89.5 %) vs. usually lack DTBYLD (25/28 = 89.3 %).
</paragraph>
</subSubSection>
<subSubSection id="C37AAAFD5B11FFB0284D1DABFE646E92" lastPageId="7" lastPageNumber="42" pageId="6" pageNumber="41" type="description">
<paragraph id="8BDFF9765B11FFB0284D1DABFE646E92" blockId="6.[850,1436,1712,2087]" lastBlockId="7.[151,738,155,997]" lastPageId="7" lastPageNumber="42" pageId="6" pageNumber="41">
<emphasis id="B91425645B11FFB1284D1DABFC5868CE" bold="true" box="[875,979,1751,1775]" pageId="6" pageNumber="41">Variation.</emphasis>
Morphometric measurements and observations are presented in
<tableCitation id="C6E2CCCD5B11FFB128DC1D82FBDB6936" box="[1018,1104,1790,1815]" captionStart="Table 2" captionStartId="5.[305,356,166,188]" captionTargetPageId="5" captionText="Table 2. Measurements (mm) of SVL and character ratios (R = %SVL) of TRL to LJL (ranges are shown in parentheses)" httpUri="http://table.plazi.org/id/DF1FA9FE5B12FFB22A171BDAFE6B6EFA" pageId="6" pageNumber="41" tableUuid="DF1FA9FE5B12FFB22A171BDAFE6B6EFA">Tables 2</tableCitation>
and
<tableCitation id="C6E2CCCD5B11FFB12FA21D83FB186936" box="[1156,1171,1791,1815]" captionStart="Table 4" captionStartId="7.[325,377,1057,1079]" captionTargetPageId="7" captionText="Table 4. Characteristics among the three species of Hynobius" httpUri="http://table.plazi.org/id/DF1FA9FE5B10FFB02A631F5DFCF66A17" pageId="6" pageNumber="41" tableUuid="DF1FA9FE5B10FFB02A631F5DFCF66A17">4</tableCitation>
, respectively. The dorsum is uniformly yellowish brown or darkish brown. In the new species, DBDD are sometimes present (8/19 = 42.1 %), DWDL (3/19 = 15.8 %) and DTBYLD (2/19 = 10.5 %) are rarely absent, DGM is rarely present (1/19 = 5.3 %), CGN rarely 11 (2/19 = 10.5 %), and CFBALN rarely more than 0 (1/19 = 5.3 %). The iris is dark brown. When preserved, the dorsal coloration tends to fade to dark gray and DTBYLD becomes unclear after preservation.
</paragraph>
</subSubSection>
<subSubSection id="C37AAAFD5B10FFB02B891BBEFDBA6D51" pageId="7" pageNumber="42" type="distribution">
<paragraph id="8BDFF9765B10FFB02B891BBEFDBA6D51" blockId="7.[151,738,155,997]" pageId="7" pageNumber="42">
<emphasis id="B91425645B10FFB02B891BBEFEED6EFB" bold="true" box="[175,358,194,218]" pageId="7" pageNumber="42">Distribution.</emphasis>
This new species is known from Higashihiroshima-shi (including the former Higashihiroshima-shi and Kurose-cho), Takehara-shi, and Kure-shi (including the former Kure-shi) in
<collectingRegion id="49A437945B10FFB029541A4BFE8B6F57" country="Japan" name="Hiroshima" pageId="7" pageNumber="42">Hiroshima Prefecture</collectingRegion>
, and Imabari-shi (including the former Imabari-shi and Namikata-cho) in
<collectingRegion id="49A437945B10FFB02ABC1AF9FDC06FBC" box="[410,587,389,413]" country="Japan" name="Ehime" pageId="7" pageNumber="42">Ehime Prefecture</collectingRegion>
. DNA samples from the former Kurose-cho is not included in this study; however, based on previous field surveys (
<bibRefCitation id="EFF184875B10FFB029501AAFFF716C33" author="Okawa, H. &amp; T. Utsunomiya &amp; Y. Utsunomiya &amp; J. Naito" pageId="7" pageNumber="42" pagination="49 - 53" refId="ref10359" refString="Okawa, H., T. Utsunomiya, Y. Utsunomiya &amp; J. Naito, 1990. Hynobius nebulosus from Hiroshima Prefecture. Hibakagaku, 146: 49 - 53. (In Japanese)." type="journal article" year="1990">
Okawa
<emphasis id="B91425645B10FFB029EB1AAFFF386C33" italics="true" pageId="7" pageNumber="42">et al.</emphasis>
, 1990
</bibRefCitation>
), this species may be distributed in this area. This new species may also occur in the former Akitsucho (currently part of Higashihiroshima-shi) but there is currently no evidence to support this hypothesis. It was reported in 2018 that the populations of
<bibRefCitation id="EFF184875B10FFB0291A19E9FD6B6C8C" author="Kure-shi &amp; In Japanse" box="[572,736,661,685]" pageId="7" pageNumber="42" refId="ref10101" refString="Kure-shi, 2018. Red Data Book Kure. Kure-Kankyo- Shiminnokai. Hiroshima Prefecture, Japan. (In Japanse)." type="book" year="2018">Kure-shi (2018)</bibRefCitation>
might already be extinct as. However, sufficient field surveys with objective evidences to support its extinction in Kure-shi have not been performed. Therefore, further field surveys are needed to clarify the current distribution range of the new species in greater detail.
</paragraph>
</subSubSection>
<subSubSection id="C37AAAFD5B10FFB02B891803FB186FBC" pageId="7" pageNumber="42" type="biology_ecology">
<paragraph id="8BDFF9765B10FFB02B891803FB186FBC" blockId="7.[151,738,155,997]" lastBlockId="7.[813,1399,155,568]" pageId="7" pageNumber="42">
<emphasis id="B91425645B10FFB02B891803FEE96DB6" bold="true" box="[175,354,895,919]" pageId="7" pageNumber="42">Natural History.</emphasis>
The dominant vegetation
<typeStatus id="54DB47D45B10FFB029551803FD146DB6" box="[627,671,895,919]" pageId="7" pageNumber="42">type</typeStatus>
of the
<typeStatus id="54DB47D45B10FFB02BB118DAFF486D9F" box="[151,195,934,958]" pageId="7" pageNumber="42">type</typeStatus>
locality is a mixed forest of evergreen
<taxonomicName id="4C6082F55B10FFB0296818DAFD276D9F" box="[590,684,934,958]" class="Magnoliopsida" family="Fagaceae" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="42" phylum="Tracheophyta" rank="family">Fagaceae</taxonomicName>
trees (i.e.,
<taxonomicName id="4C6082F55B10FFB02BEB18B1FEAF6DC5" box="[205,292,973,996]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="42" phylum="Tracheophyta" rank="genus">
<emphasis id="B91425645B10FFB02BEB18B1FEAF6DC5" box="[205,292,973,996]" italics="true" pageId="7" pageNumber="42">Quercus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6082F55B10FFB02A7C18B1FD816DC4" box="[346,522,973,997]" class="Magnoliopsida" family="Fagaceae" genus="Castanopsis" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="42" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B91425645B10FFB02A7C18B1FE536DC5" box="[346,472,973,996]" italics="true" pageId="7" pageNumber="42">Castanopsis</emphasis>
spp.
</taxonomicName>
) and Japanese cedar (
<taxonomicName id="4C6082F55B10FFB028131BE0FBAD6E92" box="[821,1062,156,179]" class="Pinopsida" family="Cupressaceae" genus="Cryptomeria" kingdom="Plantae" order="Pinales" pageId="7" pageNumber="42" phylum="Tracheophyta" rank="species" species="japonica">
<emphasis id="B91425645B10FFB028131BE0FBAD6E92" box="[821,1062,156,179]" italics="true" pageId="7" pageNumber="42">Cryptomeria japonica</emphasis>
</taxonomicName>
) (
<figureCitation id="135BE5F35B10FFB02F661BE7FB086E92" box="[1088,1155,155,179]" captionStart="Fig" captionStartId="8.[189,225,1023,1045]" captionTargetBox="[189,775,156,1002]" captionTargetId="figure-6@8.[189,775,156,1002]" captionTargetPageId="8" captionText="Fig. 5. Larvae, egg sacs, and habitat at the type locality of Hynobius geiyoensis sp. nov. (A, C, and E, respectively) and H. sumidai sp. nov. (B, D, and F, respectively)." figureDoi="http://doi.org/10.5281/zenodo.11281027" httpUri="https://zenodo.org/record/11281027/files/figure.png" pageId="7" pageNumber="42">Fig. 5</figureCitation>
). Larvae have distinct black dots on the lateral sides of the tail, whereas claws on the tips of the fingers and toes are absent. One pair of balancers is present during the early developmental stages of the larvae. Egg sacs are coil-shaped and are attached to fallen branches or leaves in ponds, puddles, or swamps at forest edges from February to April.
</paragraph>
</subSubSection>
<subSubSection id="C37AAAFD5B10FFB028631AD0FB5F6C19" pageId="7" pageNumber="42" type="discussion">
<paragraph id="8BDFF9765B10FFB028631AD0FB5F6C19" blockId="7.[813,1399,155,568]" pageId="7" pageNumber="42">
<emphasis id="B91425645B10FFB028631AD0FC256FE5" bold="true" box="[837,942,428,452]" pageId="7" pageNumber="42">Remarks.</emphasis>
The new species forms a monophyletic group with the
<collectingRegion id="49A437945B10FFB028AC1AAFFC616FCA" box="[906,1002,467,491]" country="United Kingdom" name="Highland" pageId="7" pageNumber="42">Highland</collectingRegion>
and Aki groups (
<bibRefCitation id="EFF184875B10FFB02F861AAFFAE06FCA" author="Matsui, M. &amp; H. Okawa &amp; K. Nishikawa &amp; G. Aoki &amp; K. Eto &amp; N. Yoshikawa &amp; S. Tanabe &amp; Y. Misawa &amp; A. Tominaga" box="[1184,1387,467,491]" pageId="7" pageNumber="42" pagination="32 - 90" refId="ref10175" refString="Matsui, M., H. Okawa, K. Nishikawa, G. Aoki, K. Eto, N. Yoshikawa, S. Tanabe, Y. Misawa &amp; A. Tominaga, 2019. Systematics of the widely distributed Japanese clouded salamander, Hynobius nebulosus (Amphibia: Caudata: Hynobiidae), and its closest relatives. Current Herpetology, 38 (1): 32 - 90." type="journal article" year="2019">
Matsui
<emphasis id="B91425645B10FFB02FD51AAFFAAC6FCA" box="[1267,1319,467,491]" italics="true" pageId="7" pageNumber="42">et al.</emphasis>
, 2019
</bibRefCitation>
). The morphology of females is unclear; thus, further studies to clarify female morphology are required.
</paragraph>
</subSubSection>
</treatment>
</document>