384 lines
53 KiB
XML
384 lines
53 KiB
XML
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<mods:title id="9EAB6CF1739B6E3D0AC61033F124FAC1">A new genus and two new species of hymenosomatid crabs (Crustacea: Brachyura: Hymenosomatidae) from the southwestern Atlantic and eastern Australia</mods:title>
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<mods:namePart id="606AD9901C1A620036A5E62DEEE95C33">Santana, William</mods:namePart>
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<mods:title id="07667D0D9818014F0D23633C0019767F">Zootaxa</mods:title>
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<mods:part id="A66E5CC71352ABF16D3B3108A4388C1C">
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<mods:date id="8FC52A09CDE279E5A3EBF6C7C519A271">2015</mods:date>
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<mods:number id="85E6464FE0910B8E2D58910B7E06684A">3905</mods:number>
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<treatment id="03B787CBFFA4FFAC80A95CCCFA7C08AC" ID-DOI="http://doi.org/10.5281/zenodo.3510534" ID-GBIF-Taxon="119587480" ID-Zenodo-Dep="3510534" LSID="urn:lsid:plazi:treatment:03B787CBFFA4FFAC80A95CCCFA7C08AC" httpUri="http://treatment.plazi.org/id/03B787CBFFA4FFAC80A95CCCFA7C08AC" lastPageId="5" lastPageNumber="96" pageId="3" pageNumber="94">
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<subSubSection id="C3046556FFA4FFAA80A95CCCFDB70A4B" pageId="3" pageNumber="94" type="nomenclature">
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<paragraph id="8BA136DDFFA4FFAA80A95CCCFD970DAA" blockId="3.[151,631,972,1031]" box="[151,631,972,998]" pageId="3" pageNumber="94">
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<heading id="D0E981B1FFA4FFAA80A95CCCFD970DAA" bold="true" box="[151,631,972,998]" fontSize="11" level="1" pageId="3" pageNumber="94" reason="1">
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<emphasis id="B96AEACFFFA4FFAA80A95CCCFD970DAA" bold="true" box="[151,631,972,998]" pageId="3" pageNumber="94">
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<taxonomicName id="4C1E4D5EFFA4FFAA80A95CCCFE2C0DAA" ID-CoL="55DDT" box="[151,460,972,998]" class="Malacostraca" family="Hymenosomatidae" genus="Teramnonotus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="94" phylum="Arthropoda" rank="species" species="johnlucasi" status="gen. nov.">
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<emphasis id="B96AEACFFFA4FFAA80A95CCCFE2C0DAA" bold="true" box="[151,460,972,998]" italics="true" pageId="3" pageNumber="94">Teramnonotus johnlucasi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25957B4FFA4FFAA81ED5CCCFDC60DAA" box="[467,550,972,998]" pageId="3" pageNumber="94" rank="genus">n. gen.</taxonomicNameLabel>
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, n. sp.
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BA136DDFFA4FFAA80A95CEEFDB70A4B" blockId="3.[151,631,972,1031]" box="[151,599,1006,1031]" pageId="3" pageNumber="94">
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(
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<figureCitation id="13252A58FFA4FFAA80A15CEEFEC30A4B" box="[159,291,1006,1031]" captionStart="FIGURE 1. A – D" captionStartId="4.[151,250,1669,1691]" captionTargetBox="[151,1436,177,1648]" captionTargetId="figure@4.[151,1436,177,1649]" captionTargetPageId="4" captionText="FIGURE 1. A – D, habitus, dorsal and ventral views. A, B, Teramnonotus johnlucasi n. gen., n. sp., ovigerous female holotype cl. 2.8 mm, cw. 2.7 mm (AM P. 12208). C, D, Teramnonotus monodi n. gen., n. sp., ovigerous female paratype cl. 4.3 mm, cw. 3.6 mm (MZUSP 10273). Note (arrows) rostrum integrated into the carapace outline in A (versus rostrum well detached from the carapace outline in C). A – D, stained with methylene blue. Scale bars: A – D, 1 mm." httpUri="https://zenodo.org/record/239543/files/figure.png" pageId="3" pageNumber="94">Figs. 1A–B</figureCitation>
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,
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<figureCitation id="13252A58FFA4FFAA81115CEFFEB20A4B" box="[303,338,1007,1031]" captionStart="FIGURE 4. A – D" captionStartId="9.[151,250,1923,1945]" captionTargetBox="[173,1416,417,1894]" captionTargetId="figure@9.[166,1420,409,1902]" captionTargetPageId="9" captionText="FIGURE 4. A – D, habitus, dorsal view. A, Teramnonotus gordonae (Monod, 1956) n. gen., n. comb., holotype (after Monod 1956: fig. 629). B, Teramnonotus johnlucasi n. gen., n. sp., ovigerous female holotype cl. 2.8 mm, cw. 2.7 mm (AM P. 12208). C, Teramnonotus monodi n. gen., n. sp., ovigerous female paratype cl. 4.3 mm, cw. 3.6 mm (MZUSP 10273). D, Elamena mathoei (Desmarest, 1823), female (ZRC 1994.4235). Scales: A – D, 1 mm." httpUri="https://zenodo.org/record/239546/files/figure.png" pageId="3" pageNumber="94">4B</figureCitation>
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,
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<figureCitation id="13252A58FFA4FFAA81605CEEFE600A4B" box="[350,384,1006,1031]" captionStart="FIGURE 5. A – D" captionStartId="10.[151,250,1240,1262]" captionTargetBox="[171,1412,216,1190]" captionTargetId="figure@10.[167,1419,211,1196]" captionTargetPageId="10" captionText="FIGURE 5. A – D, anteroventral view of the cephalothorax. A, Teramnonotus gordonae (Monod, 1956) n. gen., n. comb., holotype (after Monod 1956: fig. 630 bis). B, Teramnonotus johnlucasi n. gen., n. sp., ovigerous female holotype cl. 2.8 mm, cw. 2.7 mm (AM P. 12208). C, Teramnonotus monodi n. gen., n. sp., ovigerous female paratype cl. 4.3 mm, cw. 3.6 mm (MZUSP 10273). D, Elamena mathoei (Desmarest, 1823), female (ZRC 1994.4235). Arrows indicate the rostrum, which is integrated into the carapace outline in B (rostrum well detached from the carapace outline in C). Scales: A – D, 1 mm." httpUri="https://zenodo.org/record/239547/files/figure.png" pageId="3" pageNumber="94">5B</figureCitation>
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,
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<figureCitation id="13252A58FFA4FFAA81B35CEEFE520A4B" box="[397,434,1006,1031]" captionStart="FIGURE 6. A – C" captionStartId="11.[151,250,1487,1509]" captionTargetBox="[160,1432,590,1436]" captionTargetId="figure@11.[151,1436,581,1439]" captionTargetPageId="11" captionText="FIGURE 6. A – C, lateroanterior view of carapace. D, F, posterior view of the cephalothorax and abdomen. E, posterodorsal view of cephalothorax. A, Teramnonotus johnlucasi n. gen., n. sp., ovigerous female holotype cl. 2.8 mm, cw. 2.7 mm (AM P. 12208). B, D, Teramnonotus monodi n. gen., n. sp., ovigerous female paratype cl. 4.3 mm, cw. 3.6 mm (MZUSP 10273). C, F, Elamena mathoei (Desmarest, 1823), female (ZRC 1994.4235). E, Trigonoplax unguiformis H. Milne Edwards, 1853, female (RMNH). Arrows indicate the absence of the rostral keel in A – B (distinct vertical keel on ventral surface of rostrum in C). Ab 1, Ab 2, first and second abdominal somites. CxP 5, coxa of the pereopod 5. St 8, thoracic sternite 8 exposed. Scales: A – D, 1 mm. E – F, 2 mm." httpUri="https://zenodo.org/record/239548/files/figure.png" pageId="3" pageNumber="94">6A</figureCitation>
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,
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<figureCitation id="13252A58FFA4FFAA81835CEFFDE20A4B" box="[445,514,1007,1031]" captionStart="FIGURE 7. A, C" captionStartId="12.[151,250,1791,1813]" captionTargetBox="[166,1417,815,1736]" captionTargetId="figure@12.[158,1428,806,1743]" captionTargetPageId="12" captionText="FIGURE 7. A, C. Female abdominal somites and pleotelson, dorsal view. A, B, Teramnonotus johnlucasi n. gen., n. sp., ovigerous female holotype cl. 2.8 mm, cw. 2.7 mm (AM P. 12208). C, D, Teramnonotus monodi n. gen., n. sp., ovigerous female paratype cl. 4.3 mm, cw. 3.6 mm (MZUSP 10273). Ab 1, Ab 2, first and second abdominal somites. Ab 3 – Ab 5, third, fourth and fifth abdominal somites fused together. PlT, pleotelson (sixth abdominal somite and telson fused together). Scales: A – D, 1 mm." httpUri="https://zenodo.org/record/239549/files/figure.png" pageId="3" pageNumber="94">7A–B</figureCitation>
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,
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<figureCitation id="13252A58FFA4FFAA82305CEFFDFA0A4B" box="[526,538,1007,1031]" captionStart="FIGURE 8. A – C" captionStartId="13.[151,250,1232,1254]" captionTargetBox="[157,1423,225,1168]" captionTargetId="figure@13.[151,1436,217,1174]" captionTargetPageId="13" captionText="FIGURE 8. A – C, Teramnonotus gordonae (Monod, 1956) n. gen., n. comb. A, ovigerous female holotype cl. 3.1 mm, cw. 2.6 mm (MNHN-B 25803). B – C, ovigerous female paratype cl. 2.4 mm, cw. 2.2 mm (MNHN-B 25802) (after Monod 1956: figs. 634 and 636). D – F, Teramnonotus johnlucasi n. gen., n. sp., ovigerous female paratype cl. 2.5 mm, cw. 2.4 mm (MZUSP 32909). G – I, Teramnonotus monodi n. gen., n. sp., ovigerous female paratype cl. 4.3 mm, cw. 3.6 mm (MZUSP 10273). A, D, G, right cheliped, lateral view. B, E, H, right P 2, lateral view. C, F, I, left P 5, lateral view. Scales: 1 mm." httpUri="https://zenodo.org/record/239550/files/figure.png" pageId="3" pageNumber="94">8</figureCitation>
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D–F)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3046556FFA4FFAA80A95B36FE5E0AC6" pageId="3" pageNumber="94" type="reference_group">
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<paragraph id="8BA136DDFFA4FFAA80A95B36FE5E0AC6" blockId="3.[151,1437,1078,1162]" pageId="3" pageNumber="94">
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<treatmentCitationGroup id="AB0E11F3FFA4FFAA80A95B36FE5E0AC6" pageId="3" pageNumber="94">
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||
<taxonomicName id="4C1E4D5EFFA4FFAA80A95B36FEB80A00" ID-CoL="38YGH" box="[151,344,1078,1100]" class="Malacostraca" family="Hymenosomatidae" genus="Elamena" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="94" phylum="Arthropoda" rank="species" species="gordonae">
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<emphasis id="B96AEACFFFA4FFAA80A95B36FEB80A00" box="[151,344,1078,1100]" italics="true" pageId="3" pageNumber="94">Elamena gordonae</emphasis>
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</taxonomicName>
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||
—
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<treatmentCitation id="0ABF10CCFFA4FFAA814C5B37FDFD0A00" author="Lucas" box="[370,541,1078,1100]" page="174" pageId="3" pageNumber="94" year="1980">
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<bibRefCitation id="EF8F4B2CFFA4FFAA814C5B37FE0A0A00" author="Lucas" box="[370,490,1078,1100]" pageId="3" pageNumber="94" refString="Lucas, J. S. (1980) Spider crabs of the Family Hymenosomatidae (Crustacea: Brachyura) with particular reference to Australian species: Systematics and biology. Records of the Australian Museum, 33 (4), 148 - 247." type="journal article" year="1980">Lucas 1980</bibRefCitation>
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: 174
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</treatmentCitation>
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||
, figs. 2B, 6E,
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<treatmentCitation id="0ABF10CCFFA4FFAA82915B37FC730A00" author="Ng" box="[687,915,1078,1100]" page="6" pageId="3" pageNumber="94" year="1996">
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<bibRefCitation id="EF8F4B2CFFA4FFAA82915B37FC9B0A00" author="Ng" box="[687,891,1078,1100]" pageId="3" pageNumber="94" refString="Ng, P. K. L. & Chuang, C. T. N. (1996) The Hymenosomatidae (Crustacea: Decapoda: Brachyura) of Southeast Asia, with notes on other species. Raffles Bulletin of Zoology, 3 (Supplement), 1 - 82." type="journal article" year="1996">Ng & Chuang 1996</bibRefCitation>
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||
: 6
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</treatmentCitation>
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[pro parte],
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||
<treatmentCitation id="0ABF10CCFFA4FFAA842D5B36FB5F0A00" author="Davie" box="[1043,1215,1078,1100]" page="243" pageId="3" pageNumber="94" year="2002">
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<bibRefCitation id="EF8F4B2CFFA4FFAA842D5B36FB6D0A00" author="Davie" box="[1043,1165,1078,1100]" pageId="3" pageNumber="94" refString="Davie, P. J. F. (2002) Crustacea: Malacostraca: Eucarida (Part 2: Anomura, Brachyura). In: Wells, A. & Houston, W. W. K. (Eds.), Zoological Catalogue of Australia. Vol. 19.3 b. CSIRO Publishing, Melbourne, 641 pp." type="book" year="2002">Davie 2002</bibRefCitation>
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: 243
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||
</treatmentCitation>
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,
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<treatmentCitation id="0ABF10CCFFA4FFAA84F45B36FF000A27" ID-DOI="http://doi.org/10.5281/zenodo.3506966" ID-Zenodo-Dep="3506966" author="Almeida" httpUri="http://treatment.plazi.org/id/E218B402FFEF5169FF73F9B44DBDCF2A" page="29" pageId="3" pageNumber="94" year="2007">
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<bibRefCitation id="EF8F4B2CFFA4FFAA84F45B36FA790A00" author="Almeida" box="[1226,1433,1078,1100]" pageId="3" pageNumber="94" refString="Almeida, A. O., Guerrazzi, M. C. & Coelho, P. A. (2007) Stomatopod and decapod crustaceans from Camamu Bay, state of Bahia, Brazil. Zootaxa, 1553, 1 - 45." type="journal article" year="2007">
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Almeida
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<emphasis id="B96AEACFFFA4FFAA85165B37FAB50A00" box="[1320,1365,1078,1100]" italics="true" pageId="3" pageNumber="94">et al</emphasis>
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. 2007
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</bibRefCitation>
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||
: 29
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||
</treatmentCitation>
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||
[pro parte], Almeida & Coelho 2008: 197 [pro parte], Coelho
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<emphasis id="B96AEACFFFA4FFAA835D5B56FC700A27" box="[867,912,1109,1131]" italics="true" pageId="3" pageNumber="94">et al</emphasis>
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||
. 2008: 18 [pro parte]. [Not
|
||
<taxonomicName id="4C1E4D5EFFA4FFAA84905B55FE580AC6" ID-CoL="38YGH" authority="Monod, 1956" authorityName="Monod" authorityYear="1956" class="Malacostraca" family="Hymenosomatidae" genus="Elamena" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="94" phylum="Arthropoda" rank="species" species="gordonae" subGenus="Trigonoplax">
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<emphasis id="B96AEACFFFA4FFAA84905B55FAE80A27" box="[1198,1288,1109,1131]" italics="true" pageId="3" pageNumber="94">Elamena</emphasis>
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||
(
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<emphasis id="B96AEACFFFA4FFAA85265B55FA740A27" box="[1304,1428,1109,1131]" italics="true" pageId="3" pageNumber="94">Trigonoplax</emphasis>
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)
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<emphasis id="B96AEACFFFA4FFAA80F95B74FEC70AC6" box="[199,295,1140,1162]" italics="true" pageId="3" pageNumber="94">gordonae</emphasis>
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||
<bibRefCitation id="EF8F4B2CFFA4FFAA81135B74FE580AC6" author="Monod" box="[301,440,1140,1162]" pageId="3" pageNumber="94" refString="Monod, T. (1956) Hippidea et Brachyura ouest-africains. Memoires de l'Institut francais d'Afrique noire, 45, 1 - 674." type="journal article" year="1956">Monod, 1956</bibRefCitation>
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</taxonomicName>
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||
]
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3046556FFA4FFAA80A95BB6FBC80B11" pageId="3" pageNumber="94" type="materials_examined">
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<paragraph id="8BA136DDFFA4FFAA80A95BB6FEBB0B76" blockId="3.[151,1437,1206,1981]" pageId="3" pageNumber="94">
|
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<emphasis id="B96AEACFFFA4FFAA80A95BB6FEA70A83" bold="true" box="[151,327,1206,1231]" pageId="3" pageNumber="94">
|
||
<typeStatus id="54A5887FFFA4FFAA80A95BB6FF320A83" box="[151,210,1206,1231]" pageId="3" pageNumber="94">Type</typeStatus>
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material.
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</emphasis>
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Ovigerous female
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<typeStatus id="54A5887FFFA4FFAA82185BB6FD680A83" box="[550,648,1206,1231]" pageId="3" pageNumber="94" type="holotype">holotype</typeStatus>
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||
cl.
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<quantity id="4CE69B38FFA4FFAA82895BB7FCF20A82" box="[695,786,1207,1231]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="3" pageNumber="94" unit="mm" value="2.8">2.8 mm</quantity>
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||
, cw.
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||
<quantity id="4CE69B38FFA4FFAA836F5BB7FC4C0A82" box="[849,940,1207,1231]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="3" pageNumber="94" unit="mm" value="2.7">2.7 mm</quantity>
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||
, P1–P5 detached from the body,
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<collectingCountry id="F309764DFFA4FFAA85115BB6FA780A83" box="[1327,1432,1206,1231]" name="Australia" pageId="3" pageNumber="94">Australia</collectingCountry>
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||
, Queensland, Tyroom Roads, Gt. Sandy Strait, J. S. Hynd coll.,
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<date id="FFA0101DFFA4FFAA83575BDAFC220ABF" box="[873,962,1242,1267]" pageId="3" pageNumber="94" value="1952-03">iii.1952</date>
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||
, from a spiny alcyonarian from
|
||
<quantity id="4CE69B38FFA4FFAA850D5BDAFA8E0ABE" box="[1331,1390,1242,1266]" metricMagnitude="1" metricUnit="m" metricValue="1.8" pageId="3" pageNumber="94" unit="m" value="18.0">18 m</quantity>
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||
(10 fms) (AM P.12208); ovigerous female
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||
<typeStatus id="54A5887FFFA4FFAA82775BFFFD490B5B" box="[585,681,1279,1303]" pageId="3" pageNumber="94" type="paratype">paratype</typeStatus>
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cl.
|
||
<quantity id="4CE69B38FFA4FFAA82ED5BFDFCCB0B5A" box="[723,811,1277,1302]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="94" unit="mm" value="2.5">2.5 mm</quantity>
|
||
, cw.
|
||
<quantity id="4CE69B38FFA4FFAA835D5BFEFC5C0B5A" box="[867,956,1278,1302]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="3" pageNumber="94" unit="mm" value="2.4">2.4 mm</quantity>
|
||
, P1–P5 detached from the body, same data (
|
||
<collectionCode id="ED0FAE18FFA4FFAA80A05A22FF1D0B76" box="[158,253,1314,1338]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="94">MZUSP</collectionCode>
|
||
32909).
|
||
</paragraph>
|
||
<paragraph id="8BA136DDFFA4FFAA80F95A45FBC80B11" blockId="3.[151,1437,1206,1981]" box="[199,1064,1349,1374]" pageId="3" pageNumber="94">
|
||
<emphasis id="B96AEACFFFA4FFAA80F95A45FE850B12" bold="true" box="[199,357,1349,1374]" pageId="3" pageNumber="94">
|
||
<typeStatus id="54A5887FFFA4FFAA80F95A45FEE20B12" box="[199,258,1349,1374]" pageId="3" pageNumber="94">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<collectingCountry id="F309764DFFA4FFAA81525A45FE360B12" box="[364,470,1349,1374]" name="Australia" pageId="3" pageNumber="94">Australia</collectingCountry>
|
||
, Queensland, Tyroom Roads, Gt. Sandy Strait,
|
||
<specimenCount id="9D18FD54FFA4FFAA83D95A46FBC80B11" box="[999,1064,1350,1374]" pageId="3" pageNumber="94" type="generic">
|
||
<quantity id="4CE69B38FFA4FFAA83D95A46FBC30B11" box="[999,1059,1350,1374]" metricMagnitude="1" metricUnit="m" metricValue="1.8" pageId="3" pageNumber="94" unit="m" value="18.0">18 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3046556FFA4FFAA80F95A69FE270878" pageId="3" pageNumber="94" type="diagnosis">
|
||
<paragraph id="8BA136DDFFA4FFAA80F95A69FE270878" blockId="3.[151,1437,1206,1981]" pageId="3" pageNumber="94">
|
||
<emphasis id="B96AEACFFFA4FFAA80F95A69FEA10BCE" bold="true" box="[199,321,1385,1410]" pageId="3" pageNumber="94">Diagnosis.</emphasis>
|
||
Rostrum integrated into carapace outline, short, not outreaching second antennular article when fully extended in dorsal view, apex rounded. Tuft of setae on metagastric region of carapace absent. Inconspicuous postocular teeth. Antennules visible dorsally when folded. Antennular basal article with distinct but short anteroexternal tooth. Subhepatic region with 2 pronounced lobes of about same size. Mxp3 exopod nearly reaching distal margin of mxp3 merus. Mxp3 dactylus longer than propodus. Teeth of cutting edges of cheliped small, equal. Two pterygostomian lobes.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3046556FFA4FFAC80F9593FFEBD0C71" lastPageId="5" lastPageNumber="96" pageId="3" pageNumber="94" type="description">
|
||
<paragraph id="8BA136DDFFA4FFAA80F9593FFBA209F1" blockId="3.[151,1437,1206,1981]" pageId="3" pageNumber="94">
|
||
<emphasis id="B96AEACFFFA4FFAA80F9593FFEB90814" bold="true" box="[199,345,1599,1624]" pageId="3" pageNumber="94">Description.</emphasis>
|
||
Carapace pear-shaped, longer than wide, slightly convex; regions undefined, except for poorly marked gastrocardiac groove; dorsal surface with faint longitudinal ridge, without setae. Carapace anterolateral, posterolateral angles almost imperceptible, posterior margin rounded. Narrow sharp edged rim around carapace, rostrum. Rostrum straight, directed forward, broadly triangular, short, without subrostral keel; with minute setae on lateral margin, ventral surface. Cornea visible dorsally. Postocular tooth short. Carapace without orbits. Eyestalk immovable, with protuberant anteromesial lobe. Antennules short, stout, visible dorsally when folded, 3 peduncular articles subequal in length; basal article subquadrangular, with acute anteroexternal tooth; second somewhat longer, slender than first; distal article thinner. Interantennular septum small. Antenna very slender, short, concealed by rostrum in dorsal view. Epistome as long as wide. Subhepatic region with 2 marked lobes of approximately same size. Subtriangular pterygostomian region large, with 2 lobes; first prominent, acute, second smaller. Anterolateral angle of buccal frame projected anteroventrally. Setae absent.
|
||
</paragraph>
|
||
<caption id="DF616655FFA3FFAD80A95985FBE308BB" httpUri="https://zenodo.org/record/239543/files/figure.png" pageId="4" pageNumber="95" targetBox="[151,1436,177,1648]" targetPageId="4">
|
||
<paragraph id="8BA136DDFFA3FFAD80A95985FBE308BB" blockId="4.[151,1436,1669,1783]" pageId="4" pageNumber="95">
|
||
<emphasis id="B96AEACFFFA3FFAD80A95985FEF308D6" bold="true" box="[151,275,1669,1691]" pageId="4" pageNumber="95">FIGURE 1.</emphasis>
|
||
A–D, habitus, dorsal and ventral views. A, B,
|
||
<taxonomicName id="4C1E4D5EFFA3FFAD82D35986FC0E08D7" box="[749,1006,1669,1691]" class="Malacostraca" family="Hymenosomatidae" genus="Teramnonotus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="95" phylum="Arthropoda" rank="species" species="johnlucasi" status="gen. nov.">
|
||
<emphasis id="B96AEACFFFA3FFAD82D35986FC0E08D7" box="[749,1006,1669,1691]" italics="true" pageId="4" pageNumber="95">Teramnonotus johnlucasi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96AEACFFFA3FFAD83CA5985FB6108D6" bold="true" box="[1012,1153,1669,1690]" pageId="4" pageNumber="95">
|
||
<taxonomicNameLabel id="A25957B4FFA3FFAD83CA5985FBDC08D6" box="[1012,1084,1669,1690]" pageId="4" pageNumber="95" rank="genus">n. gen.</taxonomicNameLabel>
|
||
, n. sp.
|
||
</emphasis>
|
||
, ovigerous female holotype cl. 2.8 mm, cw. 2.7 mm (AM P.12208). C, D,
|
||
<taxonomicName id="4C1E4D5EFFA3FFAD825759A5FCAB08F6" box="[617,843,1700,1722]" class="Malacostraca" family="Hymenosomatidae" genus="Teramnonotus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="95" phylum="Arthropoda" rank="species" species="monodi" status="gen. nov.">
|
||
<emphasis id="B96AEACFFFA3FFAD825759A5FCAB08F6" box="[617,843,1700,1722]" italics="true" pageId="4" pageNumber="95">Teramnonotus monodi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96AEACFFFA3FFAD836C59A4FC3F08F5" bold="true" box="[850,991,1700,1721]" pageId="4" pageNumber="95">
|
||
<taxonomicNameLabel id="A25957B4FFA3FFAD836C59A4FC7A08F5" box="[850,922,1700,1721]" pageId="4" pageNumber="95" rank="genus">n. gen.</taxonomicNameLabel>
|
||
, n. sp.
|
||
</emphasis>
|
||
, ovigerous female paratype cl. 4.3 mm, cw. 3.6 mm (MZUSP 10273). Note (arrows) rostrum integrated into the carapace outline in A (versus rostrum well detached from the carapace outline in C). A–D, stained with methylene blue. Scale bars: A–D, 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA136DDFFA3FFAD80F95824FF0B09BC" blockId="4.[151,1436,1827,2032]" pageId="4" pageNumber="95">Mxp3 exopod long, reaching distal margin of merus, with sparse setae on inner margin. Ischium as wide as long, with dense row of stout, short setae on inner margin. Merus slightly wider than long, heart-shaped, with dense row of long setae on inner margin. Carpus articulating almost on lateral lobe of merus, with setae distally on inner margin. Dactylus longer than propodus; dactylus, propodus covered with setae, mostly on inner margin; dactylus fairly stout, bluntly pointed. Inhalant water opening bordered by long setae. Basal portion of epipod strongly curved.</paragraph>
|
||
<paragraph id="8BA136DDFFA2FFAC80F95F98FEBD0C71" blockId="5.[151,1437,151,1760]" pageId="5" pageNumber="96">P1 slender. Ischium short, narrow proximally, with few setae, without projection over merus. Merus long, slender, without spines or tooth, sparsely setous. Carpus short, without projections, with few setae. Palm narrower than long, hardly expanded laterally, vaguely shorter than merus. Fingers slender, slightly longer than palm, rounded, with very small gap in proximal third; small, equal teeth on both fingers, evenly distributed. Pereiopods similar, slightly decreasing in size from P2–P5. Ischium rounded. Merus with long spine on anterodorsal margin, longer than remaining articles, with few setae. Carpus shorter, without anterior projections, sparsely setate. Dactylus shorter than propodus, curved distally, 2 strong subterminal spines almost of same size on ventral margin. Propodus, dactylus flattened laterally, with few long setae, sparse short hooked setae. Ventral margin of dactylus with row of setae. Female abdomen much wider than long, somites 1, 2 free, somites 3–5 fused, somite 6 separate from previous somites, fused with telson. Abdominal margins with short setae; setae sparsely distributed over abdominal surface. Pleopods 2–5 well developed, projecting laterally beyond abdomen providing greater volume for carrying eggs.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3046556FFA2FFAC80F95D47FC010CC9" pageId="5" pageNumber="96" type="etymology">
|
||
<paragraph id="8BA136DDFFA2FFAC80F95D47FC010CC9" blockId="5.[151,1437,151,1760]" pageId="5" pageNumber="96">
|
||
<emphasis id="B96AEACFFFA2FFAC80F95D47FEAE0C2C" bold="true" box="[199,334,583,608]" pageId="5" pageNumber="96">Etymology.</emphasis>
|
||
The new species is named for John S. Lucas (James Cook University, Townsville,
|
||
<collectingCountry id="F309764DFFA2FFAC85335D47FA960C2C" box="[1293,1398,583,608]" name="Australia" pageId="5" pageNumber="96">Australia</collectingCountry>
|
||
) in recognition of his contributions to the knowledge of hymenosomatid crabs.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3046556FFA2FFAC80F95D8FFAC40AA4" pageId="5" pageNumber="96" type="discussion">
|
||
<paragraph id="8BA136DDFFA2FFAC80F95D8FFAC40AA4" blockId="5.[151,1437,151,1760]" pageId="5" pageNumber="96">
|
||
<emphasis id="B96AEACFFFA2FFAC80F95D8FFEDB0CE4" bold="true" box="[199,315,655,680]" pageId="5" pageNumber="96">Remarks.</emphasis>
|
||
The two ovigerous females from eastern
|
||
<collectingCountry id="F309764DFFA2FFAC831A5D8FFC6B0CE4" box="[804,907,655,680]" name="Australia" pageId="5" pageNumber="96">Australia</collectingCountry>
|
||
that form the basis for the description of the present new species were formerly considered by
|
||
<bibRefCitation id="EF8F4B2CFFA2FFAC82EC5DB5FC3A0C81" author="Lucas" box="[722,986,692,717]" pageId="5" pageNumber="96" refString="Lucas, J. S. (1980) Spider crabs of the Family Hymenosomatidae (Crustacea: Brachyura) with particular reference to Australian species: Systematics and biology. Records of the Australian Museum, 33 (4), 148 - 247." type="journal article" year="1980">Lucas (1980: 174–175)</bibRefCitation>
|
||
as conspecific with
|
||
<taxonomicName id="4C1E4D5EFFA2FFAC84FB5DB4FED10CBC" authority="Monod, 1956" authorityName="Monod" authorityYear="1956" class="Malacostraca" family="Hymenosomatidae" genus="Elamena" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="96" phylum="Arthropoda" rank="species" species="gordonae">
|
||
<emphasis id="B96AEACFFFA2FFAC84FB5DB4FA7C0C81" box="[1221,1436,692,717]" italics="true" pageId="5" pageNumber="96">Elamena gordonae</emphasis>
|
||
<bibRefCitation id="EF8F4B2CFFA2FFAC80A95DD7FED10CBC" author="Monod" box="[151,305,727,752]" pageId="5" pageNumber="96" refString="Monod, T. (1956) Hippidea et Brachyura ouest-africains. Memoires de l'Institut francais d'Afrique noire, 45, 1 - 674." type="journal article" year="1956">Monod, 1956</bibRefCitation>
|
||
</taxonomicName>
|
||
, from the eastern Atlantic. The
|
||
<typeStatus id="54A5887FFFA2FFAC82AA5DD8FD240CBC" box="[660,708,728,752]" pageId="5" pageNumber="96">type</typeStatus>
|
||
material of
|
||
<taxonomicName id="4C1E4D5EFFA2FFAC83725DD9FC350CBC" box="[844,981,727,752]" class="Malacostraca" family="Hymenosomatidae" genus="Elamena" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="96" phylum="Arthropoda" rank="species" species="gordonae">
|
||
<emphasis id="B96AEACFFFA2FFAC83725DD9FC350CBC" box="[844,981,727,752]" italics="true" pageId="5" pageNumber="96">E. gordonae</emphasis>
|
||
</taxonomicName>
|
||
was not available at that time for him to compare. A direct comparison between the
|
||
<typeStatus id="54A5887FFFA2FFAC82405DFDFD590D59" box="[638,697,765,789]" pageId="5" pageNumber="96">types</typeStatus>
|
||
of
|
||
<taxonomicName id="4C1E4D5EFFA2FFAC82E05DFDFC870D59" box="[734,871,764,789]" class="Malacostraca" family="Hymenosomatidae" genus="Elamena" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="96" phylum="Arthropoda" rank="species" species="gordonae">
|
||
<emphasis id="B96AEACFFFA2FFAC82E05DFDFC870D59" box="[734,871,764,789]" italics="true" pageId="5" pageNumber="96">E. gordonae</emphasis>
|
||
</taxonomicName>
|
||
and the Australian specimens has now shown that the Australian material should be assigned to a new species in
|
||
<taxonomicName id="4C1E4D5EFFA2FFAC83425C21FBFB0D74" box="[892,1051,801,824]" class="Malacostraca" family="Hymenosomatidae" genus="Teramnonotus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="96" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B96AEACFFFA2FFAC83425C21FBFB0D74" box="[892,1051,801,824]" italics="true" pageId="5" pageNumber="96">Teramnonotus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25957B4FFA2FFAC841B5C1FFB980D74" box="[1061,1144,799,824]" pageId="5" pageNumber="96" rank="genus">
|
||
<emphasis id="B96AEACFFFA2FFAC841B5C1FFB980D74" bold="true" box="[1061,1144,799,824]" pageId="5" pageNumber="96">n. gen.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, along with
|
||
<taxonomicName id="4C1E4D5EFFA2FFAC85315C21FED00D11" authority="Monod, 1956" authorityName="Monod" authorityYear="1956" class="Malacostraca" family="Hymenosomatidae" genus="Elamena" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="96" phylum="Arthropoda" rank="species" species="gordonae">
|
||
<emphasis id="B96AEACFFFA2FFAC85315C21FA7C0D74" box="[1295,1436,799,824]" italics="true" pageId="5" pageNumber="96">E. gordonae</emphasis>
|
||
<bibRefCitation id="EF8F4B2CFFA2FFAC80A95C44FED00D11" author="Monod" box="[151,304,836,861]" pageId="5" pageNumber="96" refString="Monod, T. (1956) Hippidea et Brachyura ouest-africains. Memoires de l'Institut francais d'Afrique noire, 45, 1 - 674." type="journal article" year="1956">Monod, 1956</bibRefCitation>
|
||
</taxonomicName>
|
||
(see above).
|
||
<taxonomicName id="4C1E4D5EFFA2FFAC81F35C45FD0B0D11" box="[461,747,836,861]" class="Malacostraca" family="Hymenosomatidae" genus="Teramnonotus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="96" phylum="Arthropoda" rank="species" species="johnlucasi" status="gen. nov.">
|
||
<emphasis id="B96AEACFFFA2FFAC81F35C45FD0B0D11" box="[461,747,836,861]" italics="true" pageId="5" pageNumber="96">Teramnonotus johnlucasi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96AEACFFFA2FFAC82CA5C44FC780D11" bold="true" box="[756,920,836,861]" pageId="5" pageNumber="96">
|
||
<taxonomicNameLabel id="A25957B4FFA2FFAC82CA5C44FCA60D11" box="[756,838,836,861]" pageId="5" pageNumber="96" rank="genus">n. gen.</taxonomicNameLabel>
|
||
, n. sp.
|
||
</emphasis>
|
||
can be separated from
|
||
<taxonomicName id="4C1E4D5EFFA2FFAC84975C45FF380DCC" authority="Monod, 1956" authorityName="Monod" authorityYear="1956" class="Malacostraca" family="Hymenosomatidae" genus="Elamena" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="96" phylum="Arthropoda" rank="species" species="gordonae" status="comb. nov.">
|
||
<emphasis id="B96AEACFFFA2FFAC84975C45FAD20D11" box="[1193,1330,836,861]" italics="true" pageId="5" pageNumber="96">T. gordonae</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8F4B2CFFA2FFAC857C5C44FF300DCC" author="Monod" pageId="5" pageNumber="96" refString="Monod, T. (1956) Hippidea et Brachyura ouest-africains. Memoires de l'Institut francais d'Afrique noire, 45, 1 - 674." type="journal article" year="1956">Monod, 1956</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25957B4FFA2FFAC80DA5C67FEAF0DCC" box="[228,335,871,896]" pageId="5" pageNumber="96">
|
||
<emphasis id="B96AEACFFFA2FFAC80DA5C67FEAF0DCC" bold="true" box="[228,335,871,896]" pageId="5" pageNumber="96">n. comb.</emphasis>
|
||
</taxonomicNameLabel>
|
||
in having: (i) rostrum integrated into the carapace outline, short, not outreaching the second antennular article when fully extended in dorsal view, apex rounded (
|
||
<figureCitation id="13252A58FFA2FFAC83875C8CFBC00DE8" box="[953,1056,908,933]" captionStart="FIGURE 1. A – D" captionStartId="4.[151,250,1669,1691]" captionTargetBox="[151,1436,177,1648]" captionTargetId="figure@4.[151,1436,177,1649]" captionTargetPageId="4" captionText="FIGURE 1. A – D, habitus, dorsal and ventral views. A, B, Teramnonotus johnlucasi n. gen., n. sp., ovigerous female holotype cl. 2.8 mm, cw. 2.7 mm (AM P. 12208). C, D, Teramnonotus monodi n. gen., n. sp., ovigerous female paratype cl. 4.3 mm, cw. 3.6 mm (MZUSP 10273). Note (arrows) rostrum integrated into the carapace outline in A (versus rostrum well detached from the carapace outline in C). A – D, stained with methylene blue. Scale bars: A – D, 1 mm." httpUri="https://zenodo.org/record/239543/files/figure.png" pageId="5" pageNumber="96">Figs. 1A</figureCitation>
|
||
; 4B; 5B) (rostrum well detached from the carapace outline, long, outreaching by far the second antennular article, apex sharper in
|
||
<taxonomicName id="4C1E4D5EFFA2FFAC84EF5CB1FAB60D84" box="[1233,1366,943,968]" class="Malacostraca" family="Hymenosomatidae" genus="Elamena" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="96" phylum="Arthropoda" rank="species" species="gordonae">
|
||
<emphasis id="B96AEACFFFA2FFAC84EF5CB1FAB60D84" box="[1233,1366,943,968]" italics="true" pageId="5" pageNumber="96">T. gordonae</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<figureCitation id="13252A58FFA2FFAC855A5CAFFF5C0DA0" captionStart="FIGURE 4. A – D" captionStartId="9.[151,250,1923,1945]" captionTargetBox="[173,1416,417,1894]" captionTargetId="figure@9.[166,1420,409,1902]" captionTargetPageId="9" captionText="FIGURE 4. A – D, habitus, dorsal view. A, Teramnonotus gordonae (Monod, 1956) n. gen., n. comb., holotype (after Monod 1956: fig. 629). B, Teramnonotus johnlucasi n. gen., n. sp., ovigerous female holotype cl. 2.8 mm, cw. 2.7 mm (AM P. 12208). C, Teramnonotus monodi n. gen., n. sp., ovigerous female paratype cl. 4.3 mm, cw. 3.6 mm (MZUSP 10273). D, Elamena mathoei (Desmarest, 1823), female (ZRC 1994.4235). Scales: A – D, 1 mm." httpUri="https://zenodo.org/record/239546/files/figure.png" pageId="5" pageNumber="96">Figs. 4A</figureCitation>
|
||
; 5A); (ii) pterygostomial lobes small, first rounded, second weak (
|
||
<figureCitation id="13252A58FFA2FFAC83F85CD4FBFF0DA1" box="[966,1055,980,1005]" captionStart="FIGURE 5. A – D" captionStartId="10.[151,250,1240,1262]" captionTargetBox="[171,1412,216,1190]" captionTargetId="figure@10.[167,1419,211,1196]" captionTargetPageId="10" captionText="FIGURE 5. A – D, anteroventral view of the cephalothorax. A, Teramnonotus gordonae (Monod, 1956) n. gen., n. comb., holotype (after Monod 1956: fig. 630 bis). B, Teramnonotus johnlucasi n. gen., n. sp., ovigerous female holotype cl. 2.8 mm, cw. 2.7 mm (AM P. 12208). C, Teramnonotus monodi n. gen., n. sp., ovigerous female paratype cl. 4.3 mm, cw. 3.6 mm (MZUSP 10273). D, Elamena mathoei (Desmarest, 1823), female (ZRC 1994.4235). Arrows indicate the rostrum, which is integrated into the carapace outline in B (rostrum well detached from the carapace outline in C). Scales: A – D, 1 mm." httpUri="https://zenodo.org/record/239547/files/figure.png" pageId="5" pageNumber="96">Fig. 5B</figureCitation>
|
||
) (pterygostomial lobes distinctly large, first ending in an acute tip, second much smaller but clearly discernible in
|
||
<taxonomicName id="4C1E4D5EFFA2FFAC84285CF9FB7B0A5C" box="[1046,1179,1015,1040]" class="Malacostraca" family="Hymenosomatidae" genus="Elamena" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="96" phylum="Arthropoda" rank="species" species="gordonae">
|
||
<emphasis id="B96AEACFFFA2FFAC84285CF9FB7B0A5C" box="[1046,1179,1015,1040]" italics="true" pageId="5" pageNumber="96">T. gordonae</emphasis>
|
||
</taxonomicName>
|
||
); (iii) subhepatic lobes about the same size (
|
||
<figureCitation id="13252A58FFA2FFAC81B35B1CFE050A79" box="[397,485,1052,1077]" captionStart="FIGURE 5. A – D" captionStartId="10.[151,250,1240,1262]" captionTargetBox="[171,1412,216,1190]" captionTargetId="figure@10.[167,1419,211,1196]" captionTargetPageId="10" captionText="FIGURE 5. A – D, anteroventral view of the cephalothorax. A, Teramnonotus gordonae (Monod, 1956) n. gen., n. comb., holotype (after Monod 1956: fig. 630 bis). B, Teramnonotus johnlucasi n. gen., n. sp., ovigerous female holotype cl. 2.8 mm, cw. 2.7 mm (AM P. 12208). C, Teramnonotus monodi n. gen., n. sp., ovigerous female paratype cl. 4.3 mm, cw. 3.6 mm (MZUSP 10273). D, Elamena mathoei (Desmarest, 1823), female (ZRC 1994.4235). Arrows indicate the rostrum, which is integrated into the carapace outline in B (rostrum well detached from the carapace outline in C). Scales: A – D, 1 mm." httpUri="https://zenodo.org/record/239547/files/figure.png" pageId="5" pageNumber="96">Fig. 5B</figureCitation>
|
||
) (subhepatic lobes clearly of distinct sizes, first much larger in
|
||
<taxonomicName id="4C1E4D5EFFA2FFAC84F15B1DFAB80A79" box="[1231,1368,1052,1077]" class="Malacostraca" family="Hymenosomatidae" genus="Elamena" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="96" phylum="Arthropoda" rank="species" species="gordonae">
|
||
<emphasis id="B96AEACFFFA2FFAC84F15B1DFAB80A79" box="[1231,1368,1052,1077]" italics="true" pageId="5" pageNumber="96">T. gordonae</emphasis>
|
||
</taxonomicName>
|
||
); (iv) antennules visible dorsally when folded (
|
||
<figureCitation id="13252A58FFA2FFAC825D5B3FFD590A14" box="[611,697,1087,1112]" captionStart="FIGURE 5. A – D" captionStartId="10.[151,250,1240,1262]" captionTargetBox="[171,1412,216,1190]" captionTargetId="figure@10.[167,1419,211,1196]" captionTargetPageId="10" captionText="FIGURE 5. A – D, anteroventral view of the cephalothorax. A, Teramnonotus gordonae (Monod, 1956) n. gen., n. comb., holotype (after Monod 1956: fig. 630 bis). B, Teramnonotus johnlucasi n. gen., n. sp., ovigerous female holotype cl. 2.8 mm, cw. 2.7 mm (AM P. 12208). C, Teramnonotus monodi n. gen., n. sp., ovigerous female paratype cl. 4.3 mm, cw. 3.6 mm (MZUSP 10273). D, Elamena mathoei (Desmarest, 1823), female (ZRC 1994.4235). Arrows indicate the rostrum, which is integrated into the carapace outline in B (rostrum well detached from the carapace outline in C). Scales: A – D, 1 mm." httpUri="https://zenodo.org/record/239547/files/figure.png" pageId="5" pageNumber="96">Fig. 5B</figureCitation>
|
||
) (antennules not visible dorsally in
|
||
<taxonomicName id="4C1E4D5EFFA2FFAC84745B41FB2F0A14" box="[1098,1231,1087,1112]" class="Malacostraca" family="Hymenosomatidae" genus="Elamena" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="96" phylum="Arthropoda" rank="species" species="gordonae">
|
||
<emphasis id="B96AEACFFFA2FFAC84745B41FB2F0A14" box="[1098,1231,1087,1112]" italics="true" pageId="5" pageNumber="96">T. gordonae</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<figureCitation id="13252A58FFA2FFAC84E05B3FFAD80A14" box="[1246,1336,1087,1112]" captionStart="FIGURE 5. A – D" captionStartId="10.[151,250,1240,1262]" captionTargetBox="[171,1412,216,1190]" captionTargetId="figure@10.[167,1419,211,1196]" captionTargetPageId="10" captionText="FIGURE 5. A – D, anteroventral view of the cephalothorax. A, Teramnonotus gordonae (Monod, 1956) n. gen., n. comb., holotype (after Monod 1956: fig. 630 bis). B, Teramnonotus johnlucasi n. gen., n. sp., ovigerous female holotype cl. 2.8 mm, cw. 2.7 mm (AM P. 12208). C, Teramnonotus monodi n. gen., n. sp., ovigerous female paratype cl. 4.3 mm, cw. 3.6 mm (MZUSP 10273). D, Elamena mathoei (Desmarest, 1823), female (ZRC 1994.4235). Arrows indicate the rostrum, which is integrated into the carapace outline in B (rostrum well detached from the carapace outline in C). Scales: A – D, 1 mm." httpUri="https://zenodo.org/record/239547/files/figure.png" pageId="5" pageNumber="96">Fig. 5A</figureCitation>
|
||
); and (v) no tuft of setae on metagastric region of the carapace (
|
||
<figureCitation id="13252A58FFA2FFAC83285B64FC8F0A30" box="[790,879,1124,1149]" captionStart="FIGURE 4. A – D" captionStartId="9.[151,250,1923,1945]" captionTargetBox="[173,1416,417,1894]" captionTargetId="figure@9.[166,1420,409,1902]" captionTargetPageId="9" captionText="FIGURE 4. A – D, habitus, dorsal view. A, Teramnonotus gordonae (Monod, 1956) n. gen., n. comb., holotype (after Monod 1956: fig. 629). B, Teramnonotus johnlucasi n. gen., n. sp., ovigerous female holotype cl. 2.8 mm, cw. 2.7 mm (AM P. 12208). C, Teramnonotus monodi n. gen., n. sp., ovigerous female paratype cl. 4.3 mm, cw. 3.6 mm (MZUSP 10273). D, Elamena mathoei (Desmarest, 1823), female (ZRC 1994.4235). Scales: A – D, 1 mm." httpUri="https://zenodo.org/record/239546/files/figure.png" pageId="5" pageNumber="96">Fig. 4B</figureCitation>
|
||
) (small tuft of setae on metagastric region in
|
||
<taxonomicName id="4C1E4D5EFFA2FFAC85B95B65FEE10AEC" class="Malacostraca" family="Hymenosomatidae" genus="Elamena" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="96" phylum="Arthropoda" rank="species" species="gordonae">
|
||
<emphasis id="B96AEACFFFA2FFAC85B95B65FEE10AEC" italics="true" pageId="5" pageNumber="96">T. gordonae</emphasis>
|
||
</taxonomicName>
|
||
). It also appears that
|
||
<taxonomicName id="4C1E4D5EFFA2FFAC81CD5B89FD630AEC" box="[499,643,1159,1184]" class="Malacostraca" family="Hymenosomatidae" genus="Teramnonotus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="96" phylum="Arthropoda" rank="species" species="johnlucasi">
|
||
<emphasis id="B96AEACFFFA2FFAC81CD5B89FD630AEC" box="[499,643,1159,1184]" italics="true" pageId="5" pageNumber="96">T. johnlucasi</emphasis>
|
||
</taxonomicName>
|
||
is smaller in overall size than
|
||
<taxonomicName id="4C1E4D5EFFA2FFAC83E45B89FB800AEC" box="[986,1120,1159,1184]" class="Malacostraca" family="Hymenosomatidae" genus="Elamena" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="96" phylum="Arthropoda" rank="species" species="gordonae">
|
||
<emphasis id="B96AEACFFFA2FFAC83E45B89FB800AEC" box="[986,1120,1159,1184]" italics="true" pageId="5" pageNumber="96">T. gordonae</emphasis>
|
||
</taxonomicName>
|
||
, the smallest mature female being only
|
||
<quantity id="4CE69B38FFA2FFAC81295BACFE8F0A88" box="[279,367,1196,1221]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="96" unit="mm" value="2.5">2.5 mm</quantity>
|
||
(cl) (cl.
|
||
<quantity id="4CE69B38FFA2FFAC81EC5BACFDC90A88" box="[466,553,1196,1220]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="5" pageNumber="96" unit="mm" value="3.1">3.1 mm</quantity>
|
||
in
|
||
<taxonomicName id="4C1E4D5EFFA2FFAC82715BADFD360A89" box="[591,726,1196,1221]" class="Malacostraca" family="Hymenosomatidae" genus="Elamena" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="96" phylum="Arthropoda" rank="species" species="gordonae">
|
||
<emphasis id="B96AEACFFFA2FFAC82715BADFD360A89" box="[591,726,1196,1221]" italics="true" pageId="5" pageNumber="96">T. gordonae</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<bibRefCitation id="EF8F4B2CFFA2FFAC82D05BADFC5D0A89" author="Lucas" box="[750,957,1196,1221]" pageId="5" pageNumber="96" refString="Lucas, J. S. (1980) Spider crabs of the Family Hymenosomatidae (Crustacea: Brachyura) with particular reference to Australian species: Systematics and biology. Records of the Australian Museum, 33 (4), 148 - 247." type="journal article" year="1980">Lucas (1980: 175)</bibRefCitation>
|
||
had noticed its minute size: "
|
||
<taxonomicName id="4C1E4D5EFFA2FFAC852F5BADFA7C0A89" box="[1297,1436,1196,1221]" class="Malacostraca" family="Hymenosomatidae" genus="Elamena" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="96" phylum="Arthropoda" rank="species" species="gordonae">
|
||
<emphasis id="B96AEACFFFA2FFAC852F5BADFA7C0A89" box="[1297,1436,1196,1221]" italics="true" pageId="5" pageNumber="96">E. gordonae</emphasis>
|
||
</taxonomicName>
|
||
[presently
|
||
<taxonomicName id="4C1E4D5EFFA2FFAC81305BD1FE7D0AA4" box="[270,413,1231,1256]" class="Malacostraca" family="Hymenosomatidae" genus="Teramnonotus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="96" phylum="Arthropoda" rank="species" species="johnlucasi">
|
||
<emphasis id="B96AEACFFFA2FFAC81305BD1FE7D0AA4" box="[270,413,1231,1256]" italics="true" pageId="5" pageNumber="96">T. johnlucasi</emphasis>
|
||
</taxonomicName>
|
||
] is smaller than the other 'triangular'
|
||
<taxonomicName id="4C1E4D5EFFA2FFAC837E5BCFFC440AA4" box="[832,932,1231,1256]" class="Malacostraca" family="Hymenosomatidae" genus="Elamena" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="96" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96AEACFFFA2FFAC837E5BCFFC440AA4" box="[832,932,1231,1256]" italics="true" pageId="5" pageNumber="96">Elamena</emphasis>
|
||
</taxonomicName>
|
||
species, except
|
||
<taxonomicName id="4C1E4D5EFFA2FFAC84645BD1FB360AA4" box="[1114,1238,1233,1256]" class="Malacostraca" family="Hymenosomatidae" genus="Elamena" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="96" phylum="Arthropoda" rank="species" species="umerata">
|
||
<emphasis id="B96AEACFFFA2FFAC84645BD1FB360AA4" box="[1114,1238,1233,1256]" italics="true" pageId="5" pageNumber="96">E. umerata</emphasis>
|
||
</taxonomicName>
|
||
n. sp".
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3046556FFA2FFAC80F95BF5FE8208F0" pageId="5" pageNumber="96" type="reference_group">
|
||
<paragraph id="8BA136DDFFA2FFAC80F95BF5FE8208F0" blockId="5.[151,1437,151,1760]" pageId="5" pageNumber="96">
|
||
In
|
||
<taxonomicName id="4C1E4D5EFFA2FFAC80DA5BF5FE940B41" ID-CoL="55DDT" box="[228,372,1268,1293]" class="Malacostraca" family="Hymenosomatidae" genus="Teramnonotus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="96" phylum="Arthropoda" rank="species" species="johnlucasi">
|
||
<emphasis id="B96AEACFFFA2FFAC80DA5BF5FE940B41" box="[228,372,1268,1293]" italics="true" pageId="5" pageNumber="96">T. johnlucasi</emphasis>
|
||
</taxonomicName>
|
||
, the P2–P4 dactyli are armed with two subterminal teeth of about the same size. The number of subterminal teeth in the P5 seems variable. Although in the
|
||
<typeStatus id="54A5887FFFA2FFAC83095A19FC870B7D" box="[823,871,1305,1329]" pageId="5" pageNumber="96">type</typeStatus>
|
||
specimens the P1–P5 are detached from the body, the P5 is readily recognizable because of its smaller size. One of the P5 pairs has only one subterminal tooth, the distal one, whereas the other pair has two subterminal teeth, though the proximal tooth is tiny and hardly recognizable. The examination of additional specimens of
|
||
<taxonomicName id="4C1E4D5EFFA2FFAC83185A85FC570BD1" ID-CoL="55DDT" box="[806,951,1412,1437]" class="Malacostraca" family="Hymenosomatidae" genus="Teramnonotus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="96" phylum="Arthropoda" rank="species" species="johnlucasi">
|
||
<emphasis id="B96AEACFFFA2FFAC83185A85FC570BD1" box="[806,951,1412,1437]" italics="true" pageId="5" pageNumber="96">T. johnlucasi</emphasis>
|
||
</taxonomicName>
|
||
will be necessary to determine whether the bidentate P5 is another distinctive character between
|
||
<taxonomicName id="4C1E4D5EFFA2FFAC82C05AA9FC710B8C" ID-CoL="55DDT" box="[766,913,1447,1472]" class="Malacostraca" family="Hymenosomatidae" genus="Teramnonotus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="96" phylum="Arthropoda" rank="species" species="johnlucasi">
|
||
<emphasis id="B96AEACFFFA2FFAC82C05AA9FC710B8C" box="[766,913,1447,1472]" italics="true" pageId="5" pageNumber="96">T. johnlucasi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1E4D5EFFA2FFAC83F35AA9FBB50B8C" ID-CoL="38YGH" box="[973,1109,1447,1472]" class="Malacostraca" family="Hymenosomatidae" genus="Elamena" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="96" phylum="Arthropoda" rank="species" species="gordonae">
|
||
<emphasis id="B96AEACFFFA2FFAC83F35AA9FBB50B8C" box="[973,1109,1447,1472]" italics="true" pageId="5" pageNumber="96">T. gordonae</emphasis>
|
||
</taxonomicName>
|
||
or just variation. It must be noted that
|
||
<bibRefCitation id="EF8F4B2CFFA2FFAC81265ACCFE210BA9" author="Monod" box="[280,449,1484,1509]" pageId="5" pageNumber="96" refString="Monod, T. (1956) Hippidea et Brachyura ouest-africains. Memoires de l'Institut francais d'Afrique noire, 45, 1 - 674." type="journal article" year="1956">Monod (1956)</bibRefCitation>
|
||
figured the
|
||
<typeStatus id="54A5887FFFA2FFAC82645ACCFD5C0BA9" box="[602,700,1484,1509]" pageId="5" pageNumber="96" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C1E4D5EFFA2FFAC82D25ACDFC9C0BA9" ID-CoL="38YGH" box="[748,892,1484,1509]" class="Malacostraca" family="Hymenosomatidae" genus="Elamena" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="96" phylum="Arthropoda" rank="species" species="gordonae">
|
||
<emphasis id="B96AEACFFFA2FFAC82D25ACDFC9C0BA9" box="[748,892,1484,1509]" italics="true" pageId="5" pageNumber="96">E. gordonae</emphasis>
|
||
</taxonomicName>
|
||
with tridentate P2–P4, while the P5 lack the subterminal tooth (
|
||
<bibRefCitation id="EF8F4B2CFFA2FFAC81545AEFFD7E0844" author="Monod" box="[362,670,1519,1544]" pageId="5" pageNumber="96" refString="Monod, T. (1956) Hippidea et Brachyura ouest-africains. Memoires de l'Institut francais d'Afrique noire, 45, 1 - 674." type="journal article" year="1956">Monod 1956: 470, fig. 630</bibRefCitation>
|
||
). The P5, however, is tridentate in his figure of the
|
||
<typeStatus id="54A5887FFFA2FFAC84E55AF1FADB0845" box="[1243,1339,1521,1545]" pageId="5" pageNumber="96" type="paratype">paratype</typeStatus>
|
||
(
|
||
<bibRefCitation id="EF8F4B2CFFA2FFAC85775AEFFE930861" author="Monod" pageId="5" pageNumber="96" refString="Monod, T. (1956) Hippidea et Brachyura ouest-africains. Memoires de l'Institut francais d'Afrique noire, 45, 1 - 674." type="journal article" year="1956">Monod 1956: 471, fig. 636</bibRefCitation>
|
||
).
|
||
<bibRefCitation id="EF8F4B2CFFA2FFAC81B85914FD820860" author="Monod" box="[390,610,1556,1581]" pageId="5" pageNumber="96" refString="Monod, T. (1956) Hippidea et Brachyura ouest-africains. Memoires de l'Institut francais d'Afrique noire, 45, 1 - 674." type="journal article" year="1956">Monod (1956: 472)</bibRefCitation>
|
||
maintained in his description that the P2–P5 are tridentate: "
|
||
<emphasis id="B96AEACFFFA2FFAC852E5914FB06081C" italics="true" pageId="5" pageNumber="96">Péréiopodes grêles, avec au bord supéro-distal du mérus une forte saillie tuberculiforme; dactyles 3-dentés</emphasis>
|
||
(
|
||
<emphasis id="B96AEACFFFA2FFAC84C45938FE9F0838" italics="true" pageId="5" pageNumber="96">1 apicale + 2 subapicales internes</emphasis>
|
||
)". Because the left P5 dactylus of the
|
||
<typeStatus id="54A5887FFFA2FFAC8301595CFC410839" box="[831,929,1628,1653]" pageId="5" pageNumber="96" type="holotype">holotype</typeStatus>
|
||
is broken and the right P5 missing, and the dissected appendages from the
|
||
<typeStatus id="54A5887FFFA2FFAC81C95980FDB708D4" box="[503,599,1664,1688]" pageId="5" pageNumber="96" type="paratype">paratype</typeStatus>
|
||
are also missing, it is not possible to verify if the P5 is bi- or tridentate in
|
||
<taxonomicName id="4C1E4D5EFFA2FFAC80A959A5FEC108F1" ID-CoL="38YGH" box="[151,289,1700,1725]" class="Malacostraca" family="Hymenosomatidae" genus="Elamena" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="96" phylum="Arthropoda" rank="species" sensu="stricto" species="gordonae">
|
||
<emphasis id="B96AEACFFFA2FFAC80A959A5FEC108F1" box="[151,289,1700,1725]" italics="true" pageId="5" pageNumber="96">E. gordonae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25957B4FFA2FFAC811959A5FE8208F0" box="[295,354,1701,1724]" pageId="5" pageNumber="96" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3046556FFA2FFAC80F959C7FA7C08AC" box="[199,1436,1735,1760]" pageId="5" pageNumber="96" type="distribution">
|
||
<paragraph id="8BA136DDFFA2FFAC80F959C7FA7C08AC" blockId="5.[151,1437,151,1760]" box="[199,1436,1735,1760]" pageId="5" pageNumber="96">
|
||
<emphasis id="B96AEACFFFA2FFAC80F959C7FE8008AC" bold="true" box="[199,352,1735,1760]" pageId="5" pageNumber="96">Distribution.</emphasis>
|
||
So far known from eastern
|
||
<collectingCountry id="F309764DFFA2FFAC82A759C7FD1F08AC" box="[665,767,1735,1760]" name="Australia" pageId="5" pageNumber="96">Australia</collectingCountry>
|
||
(Queensland, Tyroom Roads, Gt. Sandy Strait,
|
||
<quantity id="4CE69B38FFA2FFAC852859C8FAAF0893" box="[1302,1359,1736,1760]" metricMagnitude="1" metricUnit="m" metricValue="1.8" pageId="5" pageNumber="96" unit="m" value="18.0">18 m</quantity>
|
||
deep).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |