treatments-xml/data/03/B7/80/03B78039FFABA71EFF27FA45C5BC062D.xml

<document id="BEC41232E4168007A65DE8F9C9E03258" ID-DOI="10.5252/g2012n3a7" ID-ISSN="1638-9395" ID-Zenodo-Dep="5378120" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630284107944" checkinUser="marcus" docAuthor="Mohamed Abdelhamid, Marouf Abdel-Aty &amp; Moustafa Azab, Mahmoud" docDate="2012" docId="03B78039FFABA71EFF27FA45C5BC062D" docLanguage="en" docName="Geodiversitas.2012.34.3.575-615.pdf" docOrigin="Geodiversitas 34 (3)" docSource="http://dx.doi.org/10.5252/g2012n3a7" docStyle="DocumentStyle:DF8BFA78045876A77BB343B212438C9A.3:Geodiversitas.2011-2012.journal_article.type1" docStyleId="DF8BFA78045876A77BB343B212438C9A" docStyleName="Geodiversitas.2011-2012.journal_article.type1" docStyleVersion="3" docTitle="Rachiosoma irregulare Fourtau 1921" docType="treatment" docVersion="4" lastPageNumber="590" masterDocId="FF8EF841FFA4A711FF96FFAFC105036A" masterDocTitle="Turonian-Santonian echinoids from Egypt" masterLastPageNumber="615" masterPageNumber="575" pageNumber="590" updateTime="1699213872600" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="B66DFE3353274ECD3A0D25367627E2A3">Turonian-Santonian echinoids from Egypt</mods:title>
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<mods:namePart id="9FBBA287EC94011DDE1AAC742F246563">Mohamed Abdelhamid, Marouf Abdel-Aty</mods:namePart>
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<mods:namePart id="DB0B5900E88C5AE08B15603A04181539">Moustafa Azab, Mahmoud</mods:namePart>
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<treatment id="03B78039FFABA71EFF27FA45C5BC062D" LSID="urn:lsid:plazi:treatment:03B78039FFABA71EFF27FA45C5BC062D" httpUri="http://treatment.plazi.org/id/03B78039FFABA71EFF27FA45C5BC062D" lastPageNumber="590" pageId="15" pageNumber="590">
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<paragraph id="8BA1312FFFABA71EFF27FA45C0D0054F" blockId="15.[177,583,1514,1573]" pageId="15" pageNumber="590">
<heading id="D0E98643FFABA71EFF27FA45C0D0054F" centered="true" fontSize="11" level="2" pageId="15" pageNumber="590" reason="2">
<taxonomicName id="4C1E4AACFFABA71EFF27FA45C342056F" authority="Fourtau, 1921" authorityName="Fourtau" authorityYear="1921" box="[177,583,1514,1541]" class="Echinoidea" family="Glyphocyphidae" genus="Rachiosoma" kingdom="Animalia" order="Camarodonta" pageId="15" pageNumber="590" phylum="Echinodermata" rank="species" species="irregulare">
<emphasis id="B96AED3DFFABA71EFF27FA45C0A6056F" bold="true" box="[177,419,1514,1541]" italics="true" pageId="15" pageNumber="590">Rachiosoma irregulare</emphasis>
<bibRefCitation id="EF8F4CDEFFABA71EFE3CFA44C342056F" author="FOURTAU R." box="[426,583,1515,1541]" pageId="15" pageNumber="590" refId="ref18766" refString="FOURTAU R. 1921. - Catalogue des invertebres fossiles de l'Egypte. Terrains Cretaces, 3, Echinodermes (supplement). Geological Survey of Egypt, Paleontological Serie s 5: 101 p." type="book" year="1921">Fourtau, 1921</bibRefCitation>
</taxonomicName>
(
<figureCitation id="13252DAAFFABA71EFEBDF9A5C080054E" box="[299,389,1546,1572]" captionStart="FIG" captionStartId="12.[113,124,1544,1561]" captionTargetBox="[190,1150,216,1478]" captionTargetId="figure-17@12.[166,1157,203,1488]" captionTargetPageId="12" captionText="FIG. 7. — Camera lucida drawings: A, B, plating, tuberculation and poring in Phymosoma baylei (Cotteau, 1864) from the Turonian of Wadi Abu Qaada; A, ambital ambulacral plates; B, ambital interambulacral plate; C, D, plating, tuberculation and poring in Phymosoma beadnelli (Gregory, 1906) from the Coniacian-Santonian of Abu Roash; C, adapical ambulacral plates; D, ambital interambulacral plates; E, F, plating, tuberculation and poring in Phymosoma thevestense (Péron &amp; Gauthier, 1879) from the Coniacian-Santonian of Abu Roash; E, ambital interambulacral plates; F, subambital ambulacral plates; G, plating and tuberculation of ambital interambulacral plates in Rachiosoma irregulare Fourtau, 1921 from the Turonian of Wadi Abu Qaada. Scale bars: A, 0.8 mm; B, 0.9 mm; C, D, 1 mm; E, F, 0.5 mm; G, 0.7 mm." figureDoi="http://doi.org/10.5281/zenodo.5378126" httpUri="https://zenodo.org/record/5378126/files/figure.png" pageId="15" pageNumber="590">Figs 7G</figureCitation>
; 
<figureCitation id="13252DAAFFABA71EFE06F9A5C0C9054F" box="[400,460,1546,1573]" captionStart="FIG" captionStartId="13.[113,124,1498,1515]" captionText="FIG. 8. — A, B, Phymosoma roachensis (Fourtau, 1914), Abu Roash, Coniacian-Santonian, Ostrea and Plicatula unit, REH4, adapical and side views; C, D, Phymosoma thevestense (Péron &amp; Gauthier, 1879), Abu Roash, Coniacian-Santonian, Ostrea and Plicatula unit, REH4, adapical and side views; E, F, Rachiosoma irregulare Fourtau, 1921, Wadi Abu Qaada, Turonian, Wata Formation, QEH4, adapical and side views; G, Rachiosoma rectilineatum (Péron &amp; Gauthier, 1881),Wadi Dakhl, Coniacian-Santonian, Matulla Formation, DEH3, adapical view; H, I, Thylechinus (T.) sinaiensis n. sp., Wadi Abu Qaada, Turonian, Wata Formation, QEH3, (holotype) adapical and side views; J, K, Thylechinus (O.) quincuncialis Gregory, 1906, Wadi Abu Qaada, Turonian, Wata Formation, QEH2, adapical and side views; L, M, Goniopygus innesi Gauthier,1901,Abu Roash, Coniacian-Santonian,Ostrea and Plicatula unit,REH4, adapical view and a spine; N, Trochalosoma (T.) gharamulensis Abdelhamid &amp; Azab, 2003, Wadi Abu Qada, Turonian, Wata Formation, QEH1. Scale bars: A, B, D, 2 mm; C, H. I, M, 3.3 mm; E, 4 mm; F, G, 5 mm; J, 10 mm; K, 6.7 mm; L, 2.5 mm; N, 5 mm." pageId="15" pageNumber="590">8E, F</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30462A4FFABA71EFFE7F9F9C0A905C8" pageId="15" pageNumber="590" type="reference_group">
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<treatmentCitationGroup id="AB0E1601FFABA71EFFE7F9F9C0A905C8" pageId="15" pageNumber="590">
<treatmentCitation id="0ABF173EFFABA71EFFE7F9F9C0E70504" author="FOURTAU R." box="[113,482,1622,1646]" page="39" pageId="15" pageNumber="590" year="1921">
<taxonomicName id="4C1E4AACFFABA71EFFE7F9F9C0E70504" authority="Fourtau, 1921: 39" authorityName="Fourtau" authorityPageNumber="39" authorityYear="1921" box="[113,482,1622,1646]" class="Echinoidea" family="Glyphocyphidae" genus="Rachiosoma" kingdom="Animalia" order="Camarodonta" pageId="15" pageNumber="590" phylum="Echinodermata" rank="species" species="irregulare">
<emphasis id="B96AED3DFFABA71EFFE7F9F9C0310504" box="[113,308,1622,1646]" italics="true" pageId="15" pageNumber="590">Rachiosoma irregulare</emphasis>
<bibRefCitation id="EF8F4CDEFFABA71EFEAEF9F8C0E70504" author="FOURTAU R." box="[312,482,1622,1646]" pageId="15" pageNumber="590" refId="ref18766" refString="FOURTAU R. 1921. - Catalogue des invertebres fossiles de l'Egypte. Terrains Cretaces, 3, Echinodermes (supplement). Geological Survey of Egypt, Paleontological Serie s 5: 101 p." type="book" year="1921">Fourtau, 1921: 39</bibRefCitation>
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, pl. 5, figs3, 4. — 
<treatmentCitation id="0ABF173EFFABA71EFFF9F9DEC0A905E3" author="ABDELHAMID M. A. M. &amp; EL QOT G. M." box="[111,428,1649,1673]" page="18" pageId="15" pageNumber="590" year="2001">
<bibRefCitation id="EF8F4CDEFFABA71EFFF9F9DEC0A905E3" author="ABDELHAMID M. A. M. &amp; EL QOT G. M." box="[111,428,1649,1673]" pageId="15" pageNumber="590" pagination="1 - 31" refId="ref17779" refString="ABDELHAMID M. A. M. &amp; EL QOT G. M. 2001. - Upper Cretaceous echinoids from Gebel El-Hamra and Gebel El-Minsherah, north Sinai, Egypt. Middle East Research Center, Earth Science Series 15: 1 - 31." type="book chapter" year="2001">Abdelhamid &amp; El Qot 2001: 18</bibRefCitation>
</treatmentCitation>
, fig. 5D, E. — 
<treatmentCitation id="0ABF173EFFABA71EFDD5F9DEC1DC05C9" author="EL QOT G. M." page="137" pageId="15" pageNumber="590" year="2006">
<bibRefCitation id="EF8F4CDEFFABA71EFDD5F9DEC1DC05C9" author="EL QOT G. M." pageId="15" pageNumber="590" refId="ref18478" refString="EL QOT G. M. 2006. - Late Cretaceous macrofossils from Sinai. Beringeria 36, 163 p, 34 pls." type="book" year="2006">El Qot 2006: 137</bibRefCitation>
</treatmentCitation>
, pl. 31, figs 8, 10a, b.
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</paragraph>
</subSubSection>
<subSubSection id="C30462A4FFABA71EFD33FE19C5BF0169" pageId="15" pageNumber="590" type="materials_examined">
<paragraph id="8BA1312FFFABA71EFD33FE19C5BF0169" blockId="15.[677,1211,438,515]" pageId="15" pageNumber="590">
MATERIAL EXAMINED. — Three specimens from the Turonian of Wadi Abu Qaada (Wata Formation), 
<collectionCode id="ED0FA9EAFFABA71EFB01FE7FC3E80168" pageId="15" pageNumber="590">AS- UAQE</collectionCode>
5-7, Wadi Abu Qaada echinoid horizon 4 (
<collectionCode id="ED0FA9EAFFABA71EFBFCFE44C5A70169" box="[1130,1186,491,515]" pageId="15" pageNumber="590">QEH</collectionCode>
4).
</paragraph>
</subSubSection>
<subSubSection id="C30462A4FFABA71EFD33FD9AC5BD0080" pageId="15" pageNumber="590" type="description">
<paragraph id="8BA1312FFFABA71EFD33FD9AC2300127" blockId="15.[674,1212,565,1003]" box="[677,821,565,591]" pageId="15" pageNumber="590">DESCRIPTION</paragraph>
<paragraph id="8BA1312FFFABA71EFD33FDFAC5BD0080" blockId="15.[674,1212,565,1003]" pageId="15" pageNumber="590">
Test diameter attaining 
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. Adapical surface convex, high (H/D = 0.47-0.54). Outline of apical disc pentagonal, caducous (
<figureCitation id="13252DAAFFABA71EFB90FD3CC55001C7" box="[1030,1109,659,685]" captionStart="FIG" captionStartId="13.[113,124,1498,1515]" captionText="FIG. 8. — A, B, Phymosoma roachensis (Fourtau, 1914), Abu Roash, Coniacian-Santonian, Ostrea and Plicatula unit, REH4, adapical and side views; C, D, Phymosoma thevestense (Péron &amp; Gauthier, 1879), Abu Roash, Coniacian-Santonian, Ostrea and Plicatula unit, REH4, adapical and side views; E, F, Rachiosoma irregulare Fourtau, 1921, Wadi Abu Qaada, Turonian, Wata Formation, QEH4, adapical and side views; G, Rachiosoma rectilineatum (Péron &amp; Gauthier, 1881),Wadi Dakhl, Coniacian-Santonian, Matulla Formation, DEH3, adapical view; H, I, Thylechinus (T.) sinaiensis n. sp., Wadi Abu Qaada, Turonian, Wata Formation, QEH3, (holotype) adapical and side views; J, K, Thylechinus (O.) quincuncialis Gregory, 1906, Wadi Abu Qaada, Turonian, Wata Formation, QEH2, adapical and side views; L, M, Goniopygus innesi Gauthier,1901,Abu Roash, Coniacian-Santonian,Ostrea and Plicatula unit,REH4, adapical view and a spine; N, Trochalosoma (T.) gharamulensis Abdelhamid &amp; Azab, 2003, Wadi Abu Qada, Turonian, Wata Formation, QEH1. Scale bars: A, B, D, 2 mm; C, H. I, M, 3.3 mm; E, 4 mm; F, G, 5 mm; J, 10 mm; K, 6.7 mm; L, 2.5 mm; N, 5 mm." pageId="15" pageNumber="590">Fig. 8E</figureCitation>
) (La/D = 0.43-0.47). Ambulacral area wide (Wa/Wi = 0.62- 0.75). Poriferous zones uniserial. Major ambulacral plates quadrigeminate, but they are trigeminate in three plates adjacent to peristome. Two columns of granules occur around the perradial suture. Interambulacral area with two columns of imperforate,crenulated, and nonconfluent primary tubercles. Adradial extrascrobicular surface wide and occupied by very small tubercles (
<figureCitation id="13252DAAFFABA71EFCD0FC1FC29000A1" box="[838,917,944,971]" captionStart="FIG" captionStartId="12.[113,124,1544,1561]" captionTargetBox="[190,1150,216,1478]" captionTargetId="figure-17@12.[166,1157,203,1488]" captionTargetPageId="12" captionText="FIG. 7. — Camera lucida drawings: A, B, plating, tuberculation and poring in Phymosoma baylei (Cotteau, 1864) from the Turonian of Wadi Abu Qaada; A, ambital ambulacral plates; B, ambital interambulacral plate; C, D, plating, tuberculation and poring in Phymosoma beadnelli (Gregory, 1906) from the Coniacian-Santonian of Abu Roash; C, adapical ambulacral plates; D, ambital interambulacral plates; E, F, plating, tuberculation and poring in Phymosoma thevestense (Péron &amp; Gauthier, 1879) from the Coniacian-Santonian of Abu Roash; E, ambital interambulacral plates; F, subambital ambulacral plates; G, plating and tuberculation of ambital interambulacral plates in Rachiosoma irregulare Fourtau, 1921 from the Turonian of Wadi Abu Qaada. Scale bars: A, 0.8 mm; B, 0.9 mm; C, D, 1 mm; E, F, 0.5 mm; G, 0.7 mm." figureDoi="http://doi.org/10.5281/zenodo.5378126" httpUri="https://zenodo.org/record/5378126/files/figure.png" pageId="15" pageNumber="590">Fig.7G</figureCitation>
). Interradial extrascrobicular surface occupied by two columns of smaller granules.
</paragraph>
</subSubSection>
<subSubSection id="C30462A4FFABA71EFD33FBBFC5BC062D" pageId="15" pageNumber="590" type="discussion">
<paragraph id="8BA1312FFFABA71EFD33FBBFC20C0742" blockId="15.[675,1212,1040,1351]" box="[677,777,1040,1066]" pageId="15" pageNumber="590">REMARKS</paragraph>
<paragraph id="8BA1312FFFABA71EFD33FB80C5BC062D" blockId="15.[675,1212,1040,1351]" pageId="15" pageNumber="590">
This species is distinguished from 
<taxonomicName id="4C1E4AACFFABA71EFB85FB9FC2B60703" authority="(Peron &amp; Gauthier, 1881)" baseAuthorityName="Peron &amp; Gauthier" baseAuthorityYear="1881" class="Echinoidea" family="Glyphocyphidae" genus="Rachiosoma" kingdom="Animalia" order="Camarodonta" pageId="15" pageNumber="590" phylum="Echinodermata" rank="species" species="rectilineatum">
<emphasis id="B96AED3DFFABA71EFB85FB9FC5260723" box="[1043,1059,1072,1097]" italics="true" pageId="15" pageNumber="590">R</emphasis>
. 
<emphasis id="B96AED3DFFABA71EFBA6FB80C5BC0723" box="[1072,1209,1071,1097]" italics="true" pageId="15" pageNumber="590">rectilineatum</emphasis>
(Péron&amp; Gauthier, 1881)
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in having wider ambulacra at the apex, well developed ambulacral tubercles on adapical surface, relatively higher test (H/D = 0.47-0.54 against 
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<taxonomicName id="4C1E4AACFFABA71EFB96FB00C5AF07A2" baseAuthorityName="Peron &amp; Gauthier" baseAuthorityYear="1881" box="[1024,1194,1198,1224]" class="Echinoidea" family="Glyphocyphidae" genus="Rachiosoma" kingdom="Animalia" order="Camarodonta" pageId="15" pageNumber="590" phylum="Echinodermata" rank="species" species="rectilineatum">
<emphasis id="B96AED3DFFABA71EFB96FB00C50A07A2" box="[1024,1039,1199,1224]" italics="true" pageId="15" pageNumber="590">R</emphasis>
. 
<emphasis id="B96AED3DFFABA71EFBB4FB01C5AF07A2" box="[1058,1194,1198,1224]" italics="true" pageId="15" pageNumber="590">rectilineatum</emphasis>
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), and the mamelonated granules on the adradial extrascrobicular surface reach both the apical disc and peristome whereas in 
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<emphasis id="B96AED3DFFABA71EFC2FFAA1C2CC064D" box="[953,969,1294,1319]" italics="true" pageId="15" pageNumber="590">R</emphasis>
. 
<emphasis id="B96AED3DFFABA71EFC40FAA2C565064D" box="[982,1120,1293,1319]" italics="true" pageId="15" pageNumber="590">rectilineatum</emphasis>
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they are more developed but confined to the ambital area.
</paragraph>
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</treatment>
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