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<document id="00E519541B11E6A73A7A0F23CD8B41F4" ID-CLB-Dataset="46245" ID-DOI="10.5281/zenodo.278844" ID-GBIF-Dataset="da2ca486-83fa-4e10-99eb-f3374d9a2a51" ID-ISSN="1175-5326" ID-Zenodo-Dep="278844" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460395166422" checkinUser="plazi" docAuthor="Jäger, Peter" docDate="2011" docId="03895E31152F995BFF72887D9291FBBF" docLanguage="en" docName="zt03046p038.pdf" docOrigin="Zootaxa 3046" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Sphedanus undatus Thorell 1877" docType="treatment" docVersion="9" lastPageNumber="10" masterDocId="FFB0264915269952FFE58912907EFF9D" masterDocTitle="Revision of the spider genera Nilus O. Pickard-Cambridge 1876, Sphedanus Thorell 1877 and Dendrolycosa Doleschall 1859 (Araneae: Pisauridae)" masterLastPageNumber="38" masterPageNumber="1" pageNumber="10" updateTime="1698298638739" updateUser="ExternalLinkService">
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<mods:title id="945EE51EA574EDD08893661CA58C4636">Revision of the spider genera Nilus O. Pickard-Cambridge 1876, Sphedanus Thorell 1877 and Dendrolycosa Doleschall 1859 (Araneae: Pisauridae)</mods:title>
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<treatment id="03895E31152F995BFF72887D9291FBBF" ID-DOI="http://doi.org/10.5281/zenodo.6186796" ID-GBIF-Taxon="119417967" ID-Zenodo-Dep="6186796" LSID="urn:lsid:plazi:treatment:03895E31152F995BFF72887D9291FBBF" httpUri="http://treatment.plazi.org/id/03895E31152F995BFF72887D9291FBBF" lastPageNumber="10" pageId="9" pageNumber="10">
<subSubSection id="C33ABCAC152F995BFF72887D9117FE37" pageId="9" pageNumber="10" type="nomenclature">
<paragraph id="8B9FEF27152F995BFF72887D9255FE14" blockId="9.[151,555,367,426]" box="[151,555,367,393]" pageId="9" pageNumber="10">
<heading id="D0D7584B152F995BFF72887D9255FE14" bold="true" box="[151,555,367,393]" fontSize="11" level="1" pageId="9" pageNumber="10" reason="1">
<taxonomicName id="4C2094A4152F995BFF72887D9255FE14" ID-CoL="6YXFZ" authority="Thorell 1877" authorityName="Thorell" authorityYear="1877" box="[151,555,367,393]" class="Arachnida" family="Pisauridae" genus="Sphedanus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="undatus">
<emphasis id="B9543335152F995BFF72887D9255FE14" bold="true" box="[151,555,367,393]" pageId="9" pageNumber="10">
<emphasis id="B9543335152F995BFF72887D91FBFE14" bold="true" box="[151,389,367,393]" italics="true" pageId="9" pageNumber="10">Sphedanus undatus</emphasis>
<bibRefCitation id="EFB192D6152F995BFE69887D9255FE14" author="Thorell" box="[396,555,367,393]" pageId="9" pageNumber="10" refString="Thorell, T. (1877) Studi sui Ragni Malesi e Papuani. I. Ragni di Selebes raccolti nel 1874 dal Dott. O. Beccari. Annali del Museo civico di storia naturale di Genova, 10, 341 - 637." type="journal article" year="1877">Thorell 1877</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B9FEF27152F995BFF7288809117FE37" blockId="9.[151,555,367,426]" box="[151,361,402,426]" pageId="9" pageNumber="10">
<figureCitation id="131BF3A2152F995BFF7288809090FE37" box="[151,238,402,426]" captionStart="FIGURES 16 19. 16 17" captionStartId="7.[151,264,1757,1780]" captionTargetBox="[154,1433,599,1736]" captionTargetId="figure@7.[151,1436,589,1737]" captionTargetPageId="7" captionText="FIGURES 16 19. 16 17 Nilus amazonicus Simon 1898, holotype, juvenile, from Brazil, habitus (16 dorsal view; 17 ventral view). 18 Records of Nilus spp. (1 Egypt, Alexandria; 2 Ethiopia, detailed locality unclear; 3 India, Andhra Pradesh; 4 Andaman Islands; asterisk denotes type species of the genus). 19 Records of Sphedanus spp. (1 China, Yunnan, Menglun; 2 Laos, Luang Nam Tha Province: Ban Tavan, Nam Ha Protected Area; 3 Laos, Luang Prabang Province: Ban Mout, Ban Kengkoung, Tad Se; 4 Laos, Vientiane Province: Vang Vieng; 5 Laos, Champasak Province: Tha Hou; 6 Thailand, Trat Province: Koh Chang; 7 Malaysia: Genting [from Murphy &amp; Murphy 2000]; 8 Singapore; 9 Brunei; 10 Indonesia, Sulawesi: Kandari; asterisks denote type localities of particular species)." httpUri="https://zenodo.org/record/278847/files/figure.png" pageId="9" pageNumber="10">Figs 19</figureCitation>
: 10, 2023
</paragraph>
</subSubSection>
<subSubSection id="C33ABCAC152F995BFF7288C89277FDB0" pageId="9" pageNumber="10" type="reference_group">
<paragraph id="8B9FEF27152F995BFF7288C891F3FD92" blockId="9.[151,1436,474,558]" pageId="9" pageNumber="10">
<treatmentCitationGroup id="AB30C809152F995BFF7288C891F3FD92" pageId="9" pageNumber="10">
<taxonomicName id="4C2094A4152F995BFF7288C89265FE6D" ID-CoL="6YXFZ" authority="Thorell 1877: 523" authorityName="Thorell" authorityPageNumber="523" authorityYear="1877" box="[151,539,474,496]" class="Arachnida" family="Pisauridae" genus="Sphedanus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="undatus">
<emphasis id="B9543335152F995BFF7288C89122FE6D" box="[151,348,474,496]" italics="true" pageId="9" pageNumber="10">Sphedanus undatus</emphasis>
<treatmentCitation id="0A81C936152F995BFE8688C89265FE6D" author="Thorell" box="[355,539,474,496]" page="523" pageId="9" pageNumber="10" year="1877">
<bibRefCitation id="EFB192D6152F995BFE8688C89194FE6D" author="Thorell" box="[355,490,474,496]" pageId="9" pageNumber="10" refString="Thorell, T. (1877) Studi sui Ragni Malesi e Papuani. I. Ragni di Selebes raccolti nel 1874 dal Dott. O. Beccari. Annali del Museo civico di storia naturale di Genova, 10, 341 - 637." type="journal article" year="1877">Thorell 1877</bibRefCitation>
: 523
</treatmentCitation>
</taxonomicName>
(Description of female; female
<typeStatus id="549B5185152F995BFC8688C893C2FE6D" box="[867,956,474,496]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
from Sulawesi, Kandari,
<collectingCountry id="F337AFB7152F995BFB2788C994A9FE6D" box="[1218,1239,475,496]" name="Iceland" pageId="9" pageNumber="10">O</collectingCountry>
. Beccari leg. 1874, MCSN, examined).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9FEF27152F995BFF728B0A9277FDB0" blockId="9.[151,1436,474,558]" box="[151,521,536,558]" pageId="9" pageNumber="10">
<treatmentCitationGroup id="AB30C809152F995BFF728B0A9277FDB0" box="[151,521,536,558]" pageId="9" pageNumber="10">
<taxonomicName id="4C2094A4152F995BFF728B0A915DFDB3" box="[151,291,536,558]" class="Arachnida" family="Pisauridae" genus="Nilus" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="undatus">
<emphasis id="B9543335152F995BFF728B0A915DFDB3" box="[151,291,536,558]" italics="true" pageId="9" pageNumber="10">Nilus undatus</emphasis>
</taxonomicName>
,—
<treatmentCitation id="0A81C936152F995BFEAD8B0A927BFDB0" author="Simon" box="[328,517,536,558]" page="289" pageId="9" pageNumber="10" year="1898">
<bibRefCitation id="EFB192D6152F995BFEAD8B0A91ADFDB0" author="Simon" box="[328,467,536,558]" pageId="9" pageNumber="10" refString="Simon, E. (1898 a) Histoire naturelle des araignees. L. Mulo, Paris, 2, 193 - 380." type="journal article" year="1898" yearSuffix="a">Simon 1898a</bibRefCitation>
: 289
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C33ABCAC152F995BFF728B489478FC9F" pageId="9" pageNumber="10" type="diagnosis">
<paragraph id="8B9FEF27152F995BFF728B489478FC9F" blockId="9.[151,1437,602,1059]" pageId="9" pageNumber="10">
<emphasis id="B9543335152F995BFF728B48916FFDEE" bold="true" box="[151,273,602,627]" pageId="9" pageNumber="10">Diagnosis.</emphasis>
Epigyne similar to
<taxonomicName id="4C2094A4152F995BFE158B4992B5FDEE" box="[496,715,603,627]" class="Arachnida" family="Pisauridae" genus="Sphedanus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="quadrimaculatus" status="comb. nov.">
<emphasis id="B9543335152F995BFE158B4992B5FDEE" box="[496,715,603,627]" italics="true" pageId="9" pageNumber="10">S. quadrimaculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2678E4E152F995BFD378B489331FDEF" box="[722,847,602,626]" pageId="9" pageNumber="10">comb. nov.</taxonomicNameLabel>
(
<figureCitation id="131BF3A2152F995BFCBB8B4893A0FDEF" box="[862,990,602,626]" captionStart="FIGURES 20 29" captionStartId="8.[151,264,1922,1945]" captionTargetBox="[151,1436,183,1901]" captionTargetId="figure@8.[151,1436,183,1901]" captionTargetPageId="8" captionText="FIGURES 20 29. Sphedanus spp., female copulatory organs (20, 24, 27 epigyne, ventral view; 21, 25, 28 vulva, dorsal view; 22 schematic course of internal duct system; 23, 26, 29 epigyne, posterior view). 20 23 Sphedanus undatus Thorell 1877, holotype, female from Indonesia, Sulawesi. 24 26 Sphedanus quadrimaculatus (Thorell 1897), female syntype from Singapore. 27 29 Sphedanus banna (Zhang, Zhu &amp; Song 2004), female from Laos." httpUri="https://zenodo.org/record/278848/files/figure.png" pageId="9" pageNumber="10">Figs 2426</figureCitation>
) and
<taxonomicName id="4C2094A4152F995BFBFB8B4994FEFDEE" box="[1054,1152,603,627]" class="Arachnida" family="Pisauridae" genus="Sphedanus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="banna" status="comb. nov.">
<emphasis id="B9543335152F995BFBFB8B4994FEFDEE" box="[1054,1152,603,627]" italics="true" pageId="9" pageNumber="10">S. banna</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2678E4E152F995BFB6D8B48957BFDEF" box="[1160,1285,602,626]" pageId="9" pageNumber="10">comb. nov.</taxonomicNameLabel>
(
<figureCitation id="131BF3A2152F995BFAF68B4895EAFDEF" box="[1299,1428,602,626]" captionStart="FIGURES 20 29" captionStartId="8.[151,264,1922,1945]" captionTargetBox="[151,1436,183,1901]" captionTargetId="figure@8.[151,1436,183,1901]" captionTargetPageId="8" captionText="FIGURES 20 29. Sphedanus spp., female copulatory organs (20, 24, 27 epigyne, ventral view; 21, 25, 28 vulva, dorsal view; 22 schematic course of internal duct system; 23, 26, 29 epigyne, posterior view). 20 23 Sphedanus undatus Thorell 1877, holotype, female from Indonesia, Sulawesi. 24 26 Sphedanus quadrimaculatus (Thorell 1897), female syntype from Singapore. 27 29 Sphedanus banna (Zhang, Zhu &amp; Song 2004), female from Laos." httpUri="https://zenodo.org/record/278848/files/figure.png" pageId="9" pageNumber="10">Figs 2729</figureCitation>
) but distinguished by the atria in dorsal view covering the copulatory ducts almost completely. It is more similar to
<taxonomicName id="4C2094A4152F995BFF728BB19086FD26" box="[151,248,675,699]" class="Arachnida" family="Pisauridae" genus="Sphedanus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="banna" status="comb. nov.">
<emphasis id="B9543335152F995BFF728BB19086FD26" box="[151,248,675,699]" italics="true" pageId="9" pageNumber="10">S. banna</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2678E4E152F995BFF1A8BB09104FD27" box="[255,378,674,698]" pageId="9" pageNumber="10">comb. nov.</taxonomicNameLabel>
in having short and posteriorly situated head of spermathecae, a long median ridge of epigynal field extending into posterior half and in exhibiting only short ridges in the lateral lobes. Median margins of lateral lobes undulating and diverging posteriorly as in
<taxonomicName id="4C2094A4152F995BFD518BF993FDFC9E" box="[692,899,747,771]" class="Arachnida" family="Pisauridae" genus="Eurychoera" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="quadrimaculata" status="comb. nov.">
<emphasis id="B9543335152F995BFD518BF993FDFC9E" box="[692,899,747,771]" italics="true" pageId="9" pageNumber="10">S. quadrimaculata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2678E4E152F995BFC6F8BF89478FC9F" box="[906,1030,746,770]" pageId="9" pageNumber="10">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C33ABCAC152F995BFF208A1C91F8FC63" pageId="9" pageNumber="10" type="description">
<paragraph id="8B9FEF27152F995BFF208A1C90B4FCF3" blockId="9.[151,1437,602,1059]" pageId="9" pageNumber="10">
<emphasis id="B9543335152F995BFF208A1C910DFCBA" bold="true" box="[197,371,782,807]" pageId="9" pageNumber="10">Redescription.</emphasis>
PL 3.1, PW 2.6, AW 1.5, OL 4.6, OW 2.2. AME 0.21, ALE 0.21, PME 0.19, PLE 0.21, Chelicerae with 3 anterior and 3 posterior teeth, furrow without denticles. Leg I measurements: 14.4 (4.0, 1.4, 4.0, 3.3, 1.7).
</paragraph>
<paragraph id="8B9FEF27152F995BFF208A689252FC47" blockId="9.[151,1437,602,1059]" pageId="9" pageNumber="10">
Epigyne in general similar to
<taxonomicName id="4C2094A4152F995BFDEB8A699211FC0E" box="[526,623,891,915]" class="Arachnida" family="Pisauridae" genus="Sphedanus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="banna" status="comb. nov.">
<emphasis id="B9543335152F995BFDEB8A699211FC0E" box="[526,623,891,915]" italics="true" pageId="9" pageNumber="10">S. banna</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2678E4E152F995BFD938A68928FFC0F" box="[630,753,890,914]" pageId="9" pageNumber="10">comb. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C2094A4152F995BFCC38A69938AFC0E" box="[806,1012,891,915]" class="Arachnida" family="Pisauridae" genus="Eurychoera" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="quadrimaculata" status="comb. nov.">
<emphasis id="B9543335152F995BFCC38A69938AFC0E" box="[806,1012,891,915]" italics="true" pageId="9" pageNumber="10">S. quadrimaculata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2678E4E152F995BFC1F8A689408FC0F" box="[1018,1142,890,914]" pageId="9" pageNumber="10">comb. nov.</taxonomicNameLabel>
but distinguished by traits listed in the diagnosis. Posterior part of median lobes similarly large as in
<taxonomicName id="4C2094A4152F995BFC2B8A8D944EFC2B" box="[974,1072,926,950]" class="Arachnida" family="Pisauridae" genus="Sphedanus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="banna" status="comb. nov.">
<emphasis id="B9543335152F995BFC2B8A8D944EFC2B" box="[974,1072,926,950]" italics="true" pageId="9" pageNumber="10">S. banna</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2678E4E152F995BFBD38A8C94CAFC2B" box="[1078,1204,926,950]" pageId="9" pageNumber="10">comb. nov.</taxonomicNameLabel>
, but more prominent (outline) in posterior view (
<figureCitation id="131BF3A2152F995BFE2E8AD09261FC47" box="[459,543,962,986]" captionStart="FIGURES 20 29" captionStartId="8.[151,264,1922,1945]" captionTargetBox="[151,1436,183,1901]" captionTargetId="figure@8.[151,1436,183,1901]" captionTargetPageId="8" captionText="FIGURES 20 29. Sphedanus spp., female copulatory organs (20, 24, 27 epigyne, ventral view; 21, 25, 28 vulva, dorsal view; 22 schematic course of internal duct system; 23, 26, 29 epigyne, posterior view). 20 23 Sphedanus undatus Thorell 1877, holotype, female from Indonesia, Sulawesi. 24 26 Sphedanus quadrimaculatus (Thorell 1897), female syntype from Singapore. 27 29 Sphedanus banna (Zhang, Zhu &amp; Song 2004), female from Laos." httpUri="https://zenodo.org/record/278848/files/figure.png" pageId="9" pageNumber="10">Fig. 23</figureCitation>
).
</paragraph>
<paragraph id="8B9FEF27152F995BFF208AF591F8FC63" blockId="9.[151,1437,602,1059]" box="[197,390,998,1023]" pageId="9" pageNumber="10">
<emphasis id="B9543335152F995BFF208AF59177FC62" bold="true" box="[197,265,999,1023]" pageId="9" pageNumber="10">
<collectingCountry id="F337AFB7152F995BFF208AF5917AFC62" box="[197,260,999,1023]" name="Malta" pageId="9" pageNumber="10">Male</collectingCountry>
.
</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C33ABCAC152F995BFF208D189291FBBF" box="[197,751,1034,1059]" pageId="9" pageNumber="10" type="distribution">
<paragraph id="8B9FEF27152F995BFF208D189291FBBF" blockId="9.[151,1437,602,1059]" box="[197,751,1034,1059]" pageId="9" pageNumber="10">
<emphasis id="B9543335152F995BFF208D189120FBBE" bold="true" box="[197,350,1034,1059]" pageId="9" pageNumber="10">Distribution.</emphasis>
Known only from the
<typeStatus id="549B5185152F995BFDBB8D1992F0FBBE" box="[606,654,1035,1059]" pageId="9" pageNumber="10">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>