treatments-xml/data/03/89/51/038951095C183D0C66B6FBD3A9FFFA47.xml
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<document ID-DOI="http://doi.org/10.5281/zenodo.6210783" ID-GBIF-Dataset="7485373c-0219-4ed7-bb5c-ac664acf6775" ID-GBIF-Taxon="114923689" ID-Zenodo-Dep="6210783" checkinTime="1427905504904" checkinUser="donat" docAuthor="Alberto Ballerio &amp; Munetoshi Maruyama" docDate="2010" docId="038951095C183D0C66B6FBD3A9FFFA47" docLanguage="en" docName="ZK_article_2140.pdf" docOrigin="Zookeys 34" docTitle="Madrasostes simplex Paulian 1989" docType="treatment" docVersion="8" lastPageNumber="93" masterDocId="FFB029715C083D1C663AFFC4A802FFD6" masterDocTitle="The Ceratocanthinae of Ulu Gombak: high species richness at a single site, with descriptions of three new species and an annotated checklist of the Ceratocanthinae of Western Malaysia and Singapore (Coleoptera, Scarabaeoidea, Hybosoridae)" masterLastPageNumber="104" masterPageNumber="77" pageId="16" pageNumber="93" updateTime="1645477292090" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The Ceratocanthinae of Ulu Gombak: high species richness at a single site, with descriptions of three new species and an annotated checklist of the Ceratocanthinae of Western Malaysia and Singapore (Coleoptera, Scarabaeoidea, Hybosoridae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Alberto Ballerio</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Munetoshi Maruyama</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zookeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2010</mods:date>
<mods:detail type="volume">
<mods:number>34</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>77</mods:start>
<mods:end>104</mods:end>
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</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.34.268</mods:identifier>
<mods:identifier type="ZooBank">553C91CE-FEA1-4867-929B-2865647E4B2E</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6210783" ID-GBIF-Taxon="114923689" ID-Zenodo-Dep="6210783" LSID="urn:lsid:plazi:treatment:038951095C183D0C66B6FBD3A9FFFA47" httpUri="http://treatment.plazi.org/id/038951095C183D0C66B6FBD3A9FFFA47" lastPageNumber="93" pageId="16" pageNumber="93">
<subSubSection box="[140,555,1047,1073]" pageId="16" pageNumber="93" type="nomenclature">
<paragraph box="[140,555,1047,1073]" pageId="16" pageNumber="93">
<heading bold="true" box="[140,555,1047,1073]" fontSize="11" level="1" pageId="16" pageNumber="93" reason="6">
<taxonomicName authority="Paulian, 1989" authorityName="Paulian" authorityYear="1989" box="[140,555,1047,1073]" class="Insecta" family="Hybosoridae" genus="Madrasostes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="93" phylum="Arthropoda" rank="species" species="simplex">
<emphasis bold="true" box="[140,555,1047,1073]" pageId="16" pageNumber="93">
<emphasis bold="true" box="[140,382,1047,1073]" italics="true" pageId="16" pageNumber="93">Madrasostes simplex</emphasis>
Paulian, 1989
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[185,957,1081,1108]" pageId="16" pageNumber="93" type="distribution">
<paragraph box="[185,957,1081,1108]" pageId="16" pageNumber="93">
Known distribution: East Malaysia (Sabah) (Paulian 1979) (
<figureCitation box="[864,925,1081,1108]" captionStart="Figure 1" captionStartId="13.[140,211,1590,1612]" captionTargetBox="[140,1107,211,1470]" captionTargetId="figure@13.[140,1108,204,1474]" captionTargetPageId="13" captionText="Figure 1. Ceratocanthinae of Ulu Gombak. A Madrasostes variolosum habitus dorsal B Madrasostes simplex habitus dorsal C Madrasostes hashimi sp. n. habitus dorsal D idem habitus lateral E Madrasostes sculpturatum habitus dorsal." pageId="16" pageNumber="93">Fig. 1</figureCitation>
B)
</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="93" type="discussion">
<paragraph pageId="16" pageNumber="93">
Remarks. First record for West Malaysia (Selangor).
<emphasis box="[766,980,1117,1143]" italics="true" pageId="16" pageNumber="93">
<taxonomicName box="[766,980,1117,1143]" class="Insecta" family="Hybosoridae" genus="Madrasostes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="93" phylum="Arthropoda" rank="species" species="simplex">Madrasostes simplex</taxonomicName>
</emphasis>
is extremely similar to
<emphasis box="[281,440,1152,1179]" italics="true" pageId="16" pageNumber="93">M. variolosum</emphasis>
and to
<taxonomicName authority="Lansberge, 1887" authorityName="Lansberge" authorityYear="1887" box="[527,908,1152,1179]" class="Insecta" family="Hybosoridae" genus="Madrasostes" genus.bestMatchDistance="144" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="93" phylum="Arthropoda" rank="species" species="reticulatum">
<emphasis box="[527,692,1152,1179]" italics="true" pageId="16" pageNumber="93">M. reticulatum</emphasis>
(Lansberge, 1887)
</taxonomicName>
.
<emphasis italics="true" pageId="16" pageNumber="93">
<taxonomicName box="[919,1047,1152,1179]" class="Insecta" family="Hybosoridae" genus="Madrasostes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="93" phylum="Arthropoda" rank="genus">Madrasostes</taxonomicName>
variolosum
</emphasis>
is a very common species and has a wide distribution, ranging from Thailand to Borneo (based on first authors unpublished data) and showing a clinal morphological variation in the sculpturing of elytra, while
<emphasis box="[677,938,1257,1284]" italics="true" pageId="16" pageNumber="93">
<taxonomicName box="[677,938,1257,1284]" class="Insecta" family="Hybosoridae" genus="Madrasostes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="93" phylum="Arthropoda" rank="species" species="reticulatum">Madrasostes reticulatum</taxonomicName>
</emphasis>
is restricted to Sulawesi and the Philippines (although the Philippine population shows intermediate characters between
<emphasis box="[359,517,1328,1355]" italics="true" pageId="16" pageNumber="93">M. variolosum</emphasis>
and
<emphasis box="[574,739,1328,1355]" italics="true" pageId="16" pageNumber="93">
<taxonomicName box="[574,739,1328,1355]" class="Insecta" family="Hybosoridae" genus="Madrasostes" genus.bestMatchDistance="22" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="93" phylum="Arthropoda" rank="species" species="reticulatum">M. reticulatum</taxonomicName>
</emphasis>
).
<emphasis box="[763,978,1328,1355]" italics="true" pageId="16" pageNumber="93">
<taxonomicName box="[763,978,1328,1355]" class="Insecta" family="Hybosoridae" genus="Madrasostes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="93" phylum="Arthropoda" rank="species" species="simplex">Madrasostes simplex</taxonomicName>
</emphasis>
can be easily distinguished from
<emphasis box="[400,560,1363,1390]" italics="true" pageId="16" pageNumber="93">M. variolosum</emphasis>
because of the sculpturing of elytra and pronotum and the shape of parameres.
</paragraph>
</subSubSection>
</treatment>
</document>