treatments-xml/data/03/89/51/038951095C0E3D1A66B6FEBEA99FFB4D.xml

<document ID-DOI="http://doi.org/10.5281/zenodo.6210739" ID-GBIF-Dataset="7485373c-0219-4ed7-bb5c-ac664acf6775" ID-GBIF-Taxon="114923653" ID-Zenodo-Dep="6210739" checkinTime="1427905504904" checkinUser="donat" docAuthor="Alberto Ballerio &amp; Munetoshi Maruyama" docDate="2010" docId="038951095C0E3D1A66B6FEBEA99FFB4D" docLanguage="en" docName="ZK_article_2140.pdf" docOrigin="Zookeys 34" docTitle="Besuchetostes jaccoudi Paulian 1977" docType="treatment" docVersion="8" lastPageNumber="83" masterDocId="FFB029715C083D1C663AFFC4A802FFD6" masterDocTitle="The Ceratocanthinae of Ulu Gombak: high species richness at a single site, with descriptions of three new species and an annotated checklist of the Ceratocanthinae of Western Malaysia and Singapore (Coleoptera, Scarabaeoidea, Hybosoridae)" masterLastPageNumber="104" masterPageNumber="77" pageId="6" pageNumber="83" updateTime="1645477292090" updateUser="ExternalLinkService">
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<mods:title>The Ceratocanthinae of Ulu Gombak: high species richness at a single site, with descriptions of three new species and an annotated checklist of the Ceratocanthinae of Western Malaysia and Singapore (Coleoptera, Scarabaeoidea, Hybosoridae)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Alberto Ballerio</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Munetoshi Maruyama</mods:namePart>
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<mods:title>Zookeys</mods:title>
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<mods:date>2010</mods:date>
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<mods:number>34</mods:number>
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<mods:start>77</mods:start>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.34.268</mods:identifier>
<mods:identifier type="ZooBank">553C91CE-FEA1-4867-929B-2865647E4B2E</mods:identifier>
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<taxonomicName authority="Paulian, 1977" authorityName="Paulian" authorityYear="1977" box="[140,577,378,404]" class="Insecta" family="Hybosoridae" genus="Besuchetostes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="83" phylum="Arthropoda" rank="species" species="jaccoudi">
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<emphasis bold="true" box="[140,404,378,404]" italics="true" pageId="6" pageNumber="83">Besuchetostes jaccoudi</emphasis>
Paulian, 1977
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<subSubSection box="[185,879,413,439]" pageId="6" pageNumber="83" type="distribution">
<paragraph box="[185,879,413,439]" pageId="6" pageNumber="83">Known distribution: West Malaysia (Pahang) (Paulian 1977).</paragraph>
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Remarks. This is another flightless species known up to now by the holotype (in the collection of the Museum National d’Histoire naturelle, Paris) and a few further specimens from the type locality (MHNG, ABCB). Examination of all available material revealed that this species does not belong to 
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<taxonomicName box="[779,920,553,580]" class="Insecta" family="Hybosoridae" genus="Besuchetostes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="83" phylum="Arthropoda" rank="genus">Besuchetostes</taxonomicName>
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. Paulian placed this species in the genus 
<taxonomicName box="[434,575,589,615]" class="Insecta" family="Hybosoridae" genus="Besuchetostes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis box="[434,575,589,615]" italics="true" pageId="6" pageNumber="83">Besuchetostes</emphasis>
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mainly because of the lack of genal canthus/ dorsal ocular area, a character to which he gave much importance. However, according to a preliminary analysis carried out by the first author, the presence/absence of a genal canthus seems often the result of the fusion of the genal canthus with the occipital area of the head surface, a character that could be related to an adaptation to live in dark environments (leaf litter, dead logs), so it is highly homoplastic in the group and not particularly reliable for defining genera. Based on the aforesaid analysis the genus 
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<taxonomicName box="[356,496,835,862]" class="Insecta" family="Hybosoridae" genus="Besuchetostes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="83" phylum="Arthropoda" rank="genus">Besuchetostes</taxonomicName>
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is now characterized by a combination of characters involving mouthparts morphology, pronotum posteriorly swollen and protruding backwards, male genitalia and, above all, the shape of antennal scape and pedicellus. In 
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<emphasis box="[175,316,941,967]" italics="true" pageId="6" pageNumber="83">Besuchetostes</emphasis>
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the antennal scape is regularly gradually swollen proximad and the pedicellus is very large (about as wide as the apical portion of scape), whereas in 
<taxonomicName class="Insecta" family="Hybosoridae" genus="Besuchetostes" genus.bestMatchDistance="257" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="83" phylum="Arthropoda" rank="species" species="jaccoudi">
<emphasis italics="true" pageId="6" pageNumber="83">B. jaccoudi</emphasis>
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the scape is securiform, with pedicellus smaller than the apical portion of the scape. The overall morphology of 
<taxonomicName box="[576,700,1046,1073]" class="Insecta" family="Hybosoridae" genus="Besuchetostes" genus.bestMatchDistance="280" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="83" phylum="Arthropoda" rank="species" species="jaccoudi">
<emphasis box="[576,700,1046,1073]" italics="true" pageId="6" pageNumber="83">B. jaccoudi</emphasis>
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would suggest its placement in the 
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<taxonomicName box="[140,290,1082,1108]" class="Insecta" family="Hybosoridae" genus="Perignamptus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="83" phylum="Arthropoda" rank="genus">Perignamptus</taxonomicName>
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genus group, as defined by Ballerio (2009), although currently it is not possible to assign it to any given genus, due to the messy systematics of the genera belonging to the group.
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