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<document id="F0AFAD2B45C04B5458FF98556B91B7EA" ID-DOI="10.1649/0010-065X-73.3.687" ID-ISSN="1938-4394" ID-Zenodo-Dep="5402498" ID-ZooBank="urn:lsid:zoobank.org:pub:6044D8D0-770C-4904-90CF-E0184D4F3BD5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630337023334" checkinUser="valdenar" docAuthor="Delgado, Leonardo &amp; Mora-Aguilar, Eder F." docDate="2019" docId="03A66664FF80FF88A8E4FAFAFB1AFD13" docLanguage="en" docName="0010-065X-73.3.687.pdf" docOrigin="The Coleopterists Bulletin 73 (3)" docSource="http://dx.doi.org/10.1649/0010-065x-73.3.687" docStyle="DocumentStyle:F824235CA89CB83023C7D5789122F870.3:TheColeopBull.2017-.journal_article.1cover" docStyleId="F824235CA89CB83023C7D5789122F870" docStyleName="TheColeopBull.2017-.journal_article.1cover" docStyleVersion="3" docTitle="Verodes carinatus" docType="treatment" docVersion="3" lastPageNumber="690" masterDocId="FF9F1E1CFF84FF8CA835FFCEFF9BFF93" masterDocTitle="A New Mexican Species and Distributional Comments of Verodes Casey (Coleoptera: Zopheridae: Zopherinae)" masterLastPageNumber="692" masterPageNumber="687" pageNumber="690" updateTime="1699234001971" updateUser="plazi">
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<mods:title id="042CE2D24C42BA1AC5BFB03B90AE3657">A New Mexican Species and Distributional Comments of Verodes Casey (Coleoptera: Zopheridae: Zopherinae)</mods:title>
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<mods:namePart id="EB80BD22241FFD2F119D8B38845DB807">Delgado, Leonardo</mods:namePart>
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<mods:namePart id="1117B13C2A943CFF2DEECD4602D36D39">Mora-Aguilar, Eder F.</mods:namePart>
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<mods:title id="D3923E7BC1DEFD6BBF6D6FAC63A383DF">The Coleopterists Bulletin</mods:title>
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<mods:date id="D614C084D3F9732433F66733B0787119">2019</mods:date>
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<mods:number id="C2C1FF8DA8F0A694F8D5F209A3D227CF">2019-09-22</mods:number>
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<mods:classification id="45BE57231B3365AB41C555F485BFD8C1">journal article</mods:classification>
<mods:identifier id="14F8E7FE63CEF91A79B182187F027A01" type="DOI">10.1649/0010-065X-73.3.687</mods:identifier>
<mods:identifier id="96C514991490697F1CF84EAC69AF8D79" type="ISSN">1938-4394</mods:identifier>
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<treatment id="03A66664FF80FF88A8E4FAFAFB1AFD13" LSID="urn:lsid:plazi:treatment:03A66664FF80FF88A8E4FAFAFB1AFD13" httpUri="http://treatment.plazi.org/id/03A66664FF80FF88A8E4FAFAFB1AFD13" lastPageNumber="690" pageId="4" pageNumber="690">
<subSubSection id="C31584F9FF80FF88A8E4FAFAFEFDF9C0" pageId="4" pageNumber="690" type="nomenclature">
<paragraph id="8BB0D772FF80FF88A8E4FAFAFDDEFADA" blockId="4.[151,638,1332,1619]" box="[209,581,1332,1353]" pageId="4" pageNumber="690">
<heading id="D0F8601EFF80FF88A8E4FAFAFDDEFADA" box="[209,581,1332,1353]" centered="true" fontSize="9" level="2" pageId="4" pageNumber="690" reason="2">
<taxonomicName id="4C0FACF1FF80FF88A8E4FAFAFDDEFADA" authority="(Champion, 1884)" baseAuthorityName="Champion" baseAuthorityYear="1884" box="[209,581,1332,1353]" class="Insecta" family="Zopheridae" genus="Verodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="690" phylum="Arthropoda" rank="species" species="carinatus">
<emphasis id="B97B0B60FF80FF88A8E4FAFAFDDEFADA" bold="true" box="[209,581,1332,1353]" pageId="4" pageNumber="690">
<emphasis id="B97B0B60FF80FF88A8E4FAFAFE1EFADA" bold="true" box="[209,389,1332,1353]" italics="true" pageId="4" pageNumber="690">Verodes carinatus</emphasis>
(
<bibRefCitation id="EF9EAA83FF80FF88A9A6FAFAFDA5FADA" author="Champion, G. C." box="[403,574,1332,1353]" pageId="4" pageNumber="690" pagination="1 - 88" refId="ref3775" refString="Champion, G. C. 1884. Fam. Tenebrionidae (part) [pp. 1 - 88]. In: Biologia Centrali-Americana. Insecta. Coleoptera. Vol. IV, Part 1 (F. D. Godman and O. Salvin, editors). Porter, London, UK." type="book chapter" year="1884">Champion, 1884</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BB0D772FF80FF88A89AFA81FEFDF9C0" blockId="4.[151,638,1332,1619]" pageId="4" pageNumber="690">
This species is endemic to
<collectingRegion id="49CB1990FF80FF88A98DFA81FD99FAF7" box="[440,514,1359,1380]" country="Mexico" name="Mexico" pageId="4" pageNumber="690">Mexico</collectingRegion>
in the states of
<collectingRegion id="49CB1990FF80FF88A881FAA7FF64FAED" box="[180,255,1385,1406]" country="Mexico" name="Oaxaca" pageId="4" pageNumber="690">Oaxaca</collectingRegion>
,
<collectingRegion id="49CB1990FF80FF88A93EFAA7FED4FAED" box="[267,335,1385,1406]" country="Mexico" name="Puebla" pageId="4" pageNumber="690">Puebla</collectingRegion>
, and
<collectingRegion id="49CB1990FF80FF88A9B2FAA7FE46FAED" box="[391,477,1385,1406]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="4" pageNumber="690">Veracruz</collectingRegion>
(Foley and Ivie 2008;
<bibRefCitation id="EF9EAA83FF80FF88A8E3FA4AFE03FA0A" author="Baselga, A. &amp; Recuero, G." box="[214,408,1411,1433]" pageId="4" pageNumber="690" pagination="5060 - 5073" refId="ref3648" refString="Baselga, A., E. Recuero, G. Parra-Olea, and M. Garc'ia Par'is. 2011. Phylogenetic patterns in zopherine beetles are related to ecological niche width and dispersal limitation. Molecular Ecology 20: 5060 - 5073." type="journal article" year="2011">
Baselga
<emphasis id="B97B0B60FF80FF88A91FFA4AFEC6FA0A" box="[298,349,1411,1433]" italics="true" pageId="4" pageNumber="690">et al.</emphasis>
2011
</bibRefCitation>
). We add the following record: “
<collectingCountry id="F31897E2FF80FF88A8C2FA51FEC1FA27" box="[247,346,1439,1460]" name="Mexico" pageId="4" pageNumber="690">MEXICO</collectingCountry>
:
<collectingRegion id="49CB1990FF80FF88A959FA51FE5EFA27" box="[364,453,1439,1460]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="4" pageNumber="690">Veracruz</collectingRegion>
, Tlaquilpa, Vista Hermosa,
<date id="FFB1F1B2FF80FF88A8CEFA77FEFEFA5D" box="[251,357,1465,1486]" pageId="4" pageNumber="690" value="2013-05-11">11-V-2013</date>
,
<quantity id="4CF77A97FF80FF88A95AFA77FE5AFA5D" box="[367,449,1465,1486]" metricMagnitude="3" metricUnit="m" metricValue="2.4899999999999998" pageId="4" pageNumber="690" unit="m" value="2490.0">2,490 m</quantity>
, pine-oak forest, L. Delgado, E. Mora-A. cols.” (
<specimenCount id="9D091CFBFF80FF88A987FA1AFE48FA7A" box="[434,467,1491,1513]" pageId="4" pageNumber="690" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D091CFBFF80FF88A9EBFA1AFE65FA7A" box="[478,510,1491,1513]" pageId="4" pageNumber="690" type="female">1 ♀</specimenCount>
, IEXA). The record from Refugio in the state of
<collectingRegion id="49CB1990FF80FF88AA1CFA20FDE5F990" box="[553,638,1518,1539]" country="Mexico" name="Durango" pageId="4" pageNumber="690">Durango</collectingRegion>
(
<bibRefCitation id="EF9EAA83FF80FF88A8ABF9C7FEDAF98D" author="Champion, G. C." box="[158,321,1545,1566]" pageId="4" pageNumber="690" pagination="477 - 524" refId="ref3832" refString="Champion, G. C. 1892. Supplement to Heteromera [pp. 477 - 524]. In: Biologia Centrali-Americana. Insecta. Coleoptera. Vol. IV, Part 1 (F. D. Godman and O. Salvin, editors). Porter, London, UK." type="book chapter" year="1892">Champion 1892</bibRefCitation>
) needs confirmation due to its remoteness from the other states and the lack of intermediate records.
</paragraph>
</subSubSection>
<subSubSection id="C31584F9FF80FF88A8FEF9B9FB1AFD13" pageId="4" pageNumber="690" type="reference_group">
<paragraph id="8BB0D772FF80FF88A8FEF9B9FDE6F951" blockId="4.[151,638,1655,1730]" pageId="4" pageNumber="690">
<taxonomicName id="4C0FACF1FF80FF88A8FEF9B9FF42F934" authority="(Champion, 1884) This" authorityName="This" baseAuthorityName="Champion" baseAuthorityYear="1884" class="Insecta" family="Zopheridae" genus="Verodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="690" phylum="Arthropoda" rank="species" species="exsculptus">
<emphasis id="B97B0B60FF80FF88A8FEF9B9FDD1F91E" bold="true" box="[203,586,1655,1677]" pageId="4" pageNumber="690">
<emphasis id="B97B0B60FF80FF88A8FEF9B9FE10F91F" bold="true" box="[203,395,1655,1676]" italics="true" pageId="4" pageNumber="690">Verodes exsculptus</emphasis>
(
<bibRefCitation id="EF9EAA83FF80FF88A9ADF9B6FDD8F91E" author="Champion, G. C." box="[408,579,1656,1677]" pageId="4" pageNumber="690" pagination="1 - 88" refId="ref3775" refString="Champion, G. C. 1884. Fam. Tenebrionidae (part) [pp. 1 - 88]. In: Biologia Centrali-Americana. Insecta. Coleoptera. Vol. IV, Part 1 (F. D. Godman and O. Salvin, editors). Porter, London, UK." type="book chapter" year="1884">Champion, 1884</bibRefCitation>
)
</emphasis>
This
</taxonomicName>
species and its synonym,
<taxonomicName id="4C0FACF1FF80FF88A9FCF95CFF66F951" authority="Champion" authorityName="Champion" class="Insecta" family="Zopheridae" genus="Nosoderma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="690" phylum="Arthropoda" rank="species" species="anceps">
<emphasis id="B97B0B60FF80FF88A9FCF95CFDE5F93B" box="[457,638,1682,1704]" italics="true" pageId="4" pageNumber="690">Nosoderma anceps</emphasis>
Champion
</taxonomicName>
, have been cited from
<collectingRegion id="49CB1990FF80FF88A9E5F963FDE6F951" box="[464,637,1709,1730]" country="Mexico" name="Distrito Federal" pageId="4" pageNumber="690">Ciudad de México</collectingRegion>
</paragraph>
<paragraph id="8BB0D772FF80FF88AA98FF0CFB1AFD13" blockId="4.[685,1173,194,640]" pageId="4" pageNumber="690">
and the states of Estado de
<collectingCountry id="F31897E2FF80FF88AB9DFF0CFC68FF44" box="[936,1011,194,215]" name="Mexico" pageId="4" pageNumber="690">México</collectingCountry>
,
<collectingRegion id="49CB1990FF80FF88ABCEFF0CFBDFFF44" box="[1019,1092,194,215]" country="Mexico" name="Oaxaca" pageId="4" pageNumber="690">Oaxaca</collectingRegion>
,
<collectingRegion id="49CB1990FF80FF88AC78FF0CFB0BFF44" box="[1101,1168,194,215]" country="Mexico" name="Puebla" pageId="4" pageNumber="690">Puebla</collectingRegion>
, and
<collectingRegion id="49CB1990FF80FF88AAEBFF12FCAFFF62" box="[734,820,220,241]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="4" pageNumber="690">Veracruz</collectingRegion>
, at elevations between 2,500 and
<quantity id="4CF77A97FF80FF88AA98FF39FD66FE9F" box="[685,765,247,268]" metricMagnitude="3" metricUnit="m" metricValue="3.4" pageId="4" pageNumber="690" unit="m" value="3400.0">3,400 m</quantity>
, and associated with pine forest (
<bibRefCitation id="EF9EAA83FF80FF88AC1AFF39FD44FEB5" author="Champion, G. C." pageId="4" pageNumber="690" pagination="1 - 88" refId="ref3775" refString="Champion, G. C. 1884. Fam. Tenebrionidae (part) [pp. 1 - 88]. In: Biologia Centrali-Americana. Insecta. Coleoptera. Vol. IV, Part 1 (F. D. Godman and O. Salvin, editors). Porter, London, UK." type="book chapter" year="1884">Champion 1884</bibRefCitation>
,
<bibRefCitation id="EF9EAA83FF80FF88AADDFEDFFC80FEB5" author="Champion, G. C." box="[744,795,273,294]" pageId="4" pageNumber="690" pagination="477 - 524" refId="ref3832" refString="Champion, G. C. 1892. Supplement to Heteromera [pp. 477 - 524]. In: Biologia Centrali-Americana. Insecta. Coleoptera. Vol. IV, Part 1 (F. D. Godman and O. Salvin, editors). Porter, London, UK." type="book chapter" year="1892">1892</bibRefCitation>
;
<bibRefCitation id="EF9EAA83FF80FF88AB11FEDFFB9FFEB5" author="Garc'ia-Par'is, M." box="[804,1028,272,295]" pageId="4" pageNumber="690" pagination="215 - 230" refId="ref4107" refString="Garc'ia-Par'is, M., M. M. Coca-Abia, and G. Parra- Olea. 2006. Re-evaluation of the genera Phloeodes, Noserus, and Nosoderma (Coleoptera: Zopheridae) with description of a new species of Nosoderma from northern Mexico. Annales de la Societe entomologique de France (n. s.) 42 (2): 215 - 230." type="journal article" year="2006">
Garc´ıa-Par´ıs
<emphasis id="B97B0B60FF80FF88ABABFEDFFC56FEB5" box="[926,973,272,294]" italics="true" pageId="4" pageNumber="690">et al.</emphasis>
2006
</bibRefCitation>
; Foley and Ivie 2008;
<bibRefCitation id="EF9EAA83FF80FF88AAD9FEE2FC36FED2" author="Baselga, A. &amp; Recuero, G." box="[748,941,299,321]" pageId="4" pageNumber="690" pagination="5060 - 5073" refId="ref3648" refString="Baselga, A., E. Recuero, G. Parra-Olea, and M. Garc'ia Par'is. 2011. Phylogenetic patterns in zopherine beetles are related to ecological niche width and dispersal limitation. Molecular Ecology 20: 5060 - 5073." type="journal article" year="2011">
Baselga
<emphasis id="B97B0B60FF80FF88AB75FEE2FCE9FED2" box="[832,882,299,321]" italics="true" pageId="4" pageNumber="690">et al.</emphasis>
2011
</bibRefCitation>
). We add the following records: “
<collectingCountry id="F31897E2FF80FF88AB3CFE88FCF1FEC8" box="[777,874,326,347]" name="Mexico" pageId="4" pageNumber="690">MEXICO</collectingCountry>
: Estado de
<collectingCountry id="F31897E2FF80FF88ABE7FE88FB86FEC8" box="[978,1053,326,347]" name="Mexico" pageId="4" pageNumber="690">México</collectingCountry>
, Km 24 carr. Amecameca-Tlamacas,
<date id="FFB1F1B2FF80FF88ABACFEAFFC75FEE5" box="[921,1006,353,374]" pageId="4" pageNumber="690" value="2006-01-01">1-I-2006</date>
,
<quantity id="4CF77A97FF80FF88AC35FEAFFBC2FEE5" box="[1024,1113,353,374]" metricMagnitude="3" metricUnit="m" metricValue="3.3" pageId="4" pageNumber="690" unit="m" value="3300.0">3,300 m</quantity>
, bajo corteza, L. Delgado col.” (
<specimenCount id="9D091CFBFF80FF88ABF3FEB5FC64FE02" box="[966,1023,379,401]" pageId="4" pageNumber="690" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="9D091CFBFF80FF88AC27FEB5FBA1FE02" box="[1042,1082,379,401]" pageId="4" pageNumber="690" type="female">1 ♀</specimenCount>
, IEXA); “
<collectingCountry id="F31897E2FF80FF88AA8DFE58FC82FE38" box="[696,793,406,427]" name="Mexico" pageId="4" pageNumber="690">MEXICO</collectingCountry>
:
<collectingRegion id="49CB1990FF80FF88AB17FE58FCF0FE38" box="[802,875,406,427]" country="Mexico" name="Oaxaca" pageId="4" pageNumber="690">Oaxaca</collectingRegion>
, Santiago Comaltepec,
<quantity id="4CF77A97FF80FF88AC79FE58FBE7FE38" box="[1100,1148,406,427]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="690" unit="km" value="4.0">4 km</quantity>
al N de la desviación a Yolox,
<date id="FFB1F1B2FF80FF88AB9BFE7FFBBAFE55" box="[942,1057,433,454]" pageId="4" pageNumber="690" value="2014-11-22">22-XI-2014</date>
,
<quantity id="4CF77A97FF80FF88AC1CFE7FFBE3FE55" box="[1065,1144,433,454]" metricMagnitude="3" metricUnit="m" metricValue="2.85" pageId="4" pageNumber="690" unit="m" value="2850.0">2,850 m</quantity>
, L. Delgado, E. Mora-A. cols.” (
<specimenCount id="9D091CFBFF80FF88ABEEFE05FB99FE72" box="[987,1026,459,481]" pageId="4" pageNumber="690" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D091CFBFF80FF88AC21FE05FBA1FE72" box="[1044,1082,459,481]" pageId="4" pageNumber="690" type="female">1 ♀</specimenCount>
, IEXA); “
<collectingCountry id="F31897E2FF80FF88AA8DFE28FC82FE68" box="[696,793,486,507]" name="Mexico" pageId="4" pageNumber="690">MEXICO</collectingCountry>
:
<collectingRegion id="49CB1990FF80FF88AB11FE28FCE1FE68" box="[804,890,486,507]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="4" pageNumber="690">Veracruz</collectingRegion>
, Calcahualco,
<quantity id="4CF77A97FF80FF88AC3FFE28FBA7FE68" box="[1034,1084,486,507]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="690" unit="km" value="1.0">1 km</quantity>
antes de Nueva Vaquer´ıa,
<date id="FFB1F1B2FF80FF88AB62FDCEFC5BFD86" box="[855,960,512,533]" pageId="4" pageNumber="690" value="1992-02-28">28-II-1992</date>
,
<quantity id="4CF77A97FF80FF88ABF8FDCEFBBAFD86" box="[973,1057,512,533]" metricMagnitude="3" metricUnit="m" metricValue="2.67" pageId="4" pageNumber="690" unit="m" value="2670.0">2,670 m</quantity>
, Bosque de pino-encino, dentro de troncos, R. Novelo, F. Capistrán, L. Delgado cols” (
<specimenCount id="9D091CFBFF80FF88ABE2FDFBFB90FDD8" box="[983,1035,565,587]" pageId="4" pageNumber="690" type="female">2 ♀♀</specimenCount>
exs., IEXA);
<emphasis id="B97B0B60FF80FF88AA98FD81FD40FDF6" box="[685,731,591,613]" italics="true" pageId="4" pageNumber="690">idem</emphasis>
except: Camino al Albergue alpino,
<date id="FFB1F1B2FF80FF88AC08FD9EFB0BFDF6" box="[1085,1168,592,613]" pageId="4" pageNumber="690" value="1992-06">VI-1992</date>
,
<quantity id="4CF77A97FF80FF88AA98FDA5FC9BFD13" box="[685,768,619,640]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="690" unit="m" value="3000.0">3,000 m</quantity>
, L. Delgado col.” (
<specimenCount id="9D091CFBFF80FF88AB89FDA5FC75FD13" box="[956,1006,618,640]" pageId="4" pageNumber="690" type="male">5 ♂♂</specimenCount>
,
<specimenCount id="9D091CFBFF80FF88ABC9FDA5FBB7FD13" box="[1020,1068,618,640]" pageId="4" pageNumber="690" type="female">2 ♀♀</specimenCount>
, IEXA).
</paragraph>
</subSubSection>
</treatment>
</document>