treatments-xml/data/03/90/CE/0390CE542535FFDBFF6CAF10C58F7F15.xml

<document id="9C3B19515F28AC876DED57AEDBB98AC0" ID-CLB-Dataset="22538" ID-DOI="10.11646/zootaxa.4657.2.9" ID-GBIF-Dataset="298f6dc6-661e-43d0-b37d-722967c2be93" ID-ISSN="1175-5326" ID-Zenodo-Dep="3772362" ID-ZooBank="92516F1D-F525-473E-BB75-798372C3B39F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588057393436" checkinUser="plazi" docAuthor="Suwannarat, Nannaphat, Malicky, Hans, Morse, John C. &amp; Laudee, Pongsak" docDate="2019" docId="0390CE542535FFDBFF6CAF10C58F7F15" docLanguage="en" docName="zootaxa.4657.2.9.pdf" docOrigin="Zootaxa 4657 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhyacophilidae Suwannarat, Malicky, Morse &amp; Laudee, 2019, n. sp." docType="treatment" docVersion="7" lastPageNumber="375" masterDocId="FFA9B62C2534FFDDFFFBAD35C71F7D76" masterDocTitle="Four new species of Rhyacophila Pictet 1834 (Trichoptera: Rhyacophilidae) from Southeast Asia" masterLastPageNumber="376" masterPageNumber="369" pageNumber="370" updateTime="1698837625868" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="F5AD410C1A0ADA507230FFAD78ABA147">Four new species of Rhyacophila Pictet 1834 (Trichoptera: Rhyacophilidae) from Southeast Asia</mods:title>
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<mods:namePart id="7A122D6413095B42BC6E0F512039C294">Suwannarat, Nannaphat</mods:namePart>
<mods:affiliation id="CF0509AB138ADA1FCBF82814B76A767C">Department of Fishery and Coastal Resources, Faculty of Science and Industrial Technology, Prince of Songkla University, Surat Thani Campus, Muang District, Surat Thani Province, Thailand 84100.</mods:affiliation>
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<mods:namePart id="0C568F36C42442C915365D30E65850F1">Malicky, Hans</mods:namePart>
<mods:affiliation id="38228E84386608312CF2433237E46F47">Sonnengasse 13, A- 3293 Lunz am See, Austria</mods:affiliation>
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<mods:namePart id="19AE00E43F114025BD95AC04CFC48B3F">Morse, John C.</mods:namePart>
<mods:affiliation id="6ACEAFAFC483136A402E466FF99CFD03">Department of Plant &amp; Environmental Sciences, Clemson University, Clemson, South Carolina 29634 - 0310 USA</mods:affiliation>
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<mods:namePart id="607E52573AF009A3C01767554D6B08C7">Laudee, Pongsak</mods:namePart>
<mods:affiliation id="C4FC0BA4EF40C2C5FB2521599EE2FF6D">Department of Fishery and Coastal Resources, Faculty of Science and Industrial Technology, Prince of Songkla University, Surat Thani Campus, Muang District, Surat Thani Province, Thailand 84100.</mods:affiliation>
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<taxonomicName id="4C3904C12535FFDCFF6CAF10C6447F49" ID-CoL="FR8" box="[151,347,549,575]" class="Insecta" family="Rhyacophilidae" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="370" phylum="Arthropoda" rank="family">
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<taxonomicName id="4C3904C12535FFDCFF6CAF58C6D17FF1" authority="Suwannarat &amp; Malicky &amp; Morse &amp; Laudee, 2019" authorityName="Suwannarat &amp; Malicky &amp; Morse &amp; Laudee" authorityYear="2019" box="[151,462,621,647]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="370" phylum="Arthropoda" rank="species" species="longicaudata" status="sp. nov.">
<emphasis id="B94DA3502535FFDCFF6CAF58C6D17FF1" box="[151,462,621,647]" italics="true" pageId="1" pageNumber="370">Rhyacophila longicaudata</emphasis>
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Suwannarat &amp; Malicky
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<figureCitation id="130263C72535FFDCFF6CAFA4C7C07FDD" box="[151,223,657,683]" captionStart="FIGURES 1" captionStartId="1.[151,264,1845,1870]" captionTargetBox="[166,1416,1006,1783]" captionTargetId="figure@1.[151,1436,990,1818]" captionTargetPageId="1" captionText="FIGURES 1A–1E. Male genitalia of Rhyacophila longicaudata n. sp. 1A, left lateral, with left inferior appendage omitted. 1B, dorsal. 1C, ventral. 1D, phallus, ventral. 1E, phallus, left lateral. IX = segment IX, Pre = preanal appendages, Api IX = apicodorsal lobe of segment IX, Ana = anal sclerites, Api B = apical band, Sag = sagittal appendage, Ter = tergal band, Pha = phallus, Bas Seg = basal segment of an inferior appendage (paired), Api Seg = apical segment of an inferior appendage (paired), Aed = aedeagus, Phal= phallotheca, End = endotheca." figureDoi="http://doi.org/10.5281/zenodo.3772364" httpUri="https://zenodo.org/record/3772364/files/figure.png" pageId="1" pageNumber="370">Figs. 1</figureCitation>
A–1E
</paragraph>
<paragraph id="8B867F422535FFDCFF6CAFECC3EE7E61" blockId="1.[151,1436,729,971]" pageId="1" pageNumber="370">
Type material. 
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male (
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). 
<collectingCountry id="F32E3FD22535FFDCFD73AFECC4027F85" box="[648,797,729,755]" name="Thailand" pageId="1" pageNumber="370">THAILAND</collectingCountry>
: 
<collectingRegion id="49FDB1A02535FFDCFCD2AFECC3037F85" box="[809,1052,729,755]" country="Thailand" name="Surat Thani" pageId="1" pageNumber="370">Surat Thani Province</collectingRegion>
, 
<collectingMunicipality id="6BE2E5382535FFDCFBD3AFECC27A7F85" box="[1064,1381,729,755]" pageId="1" pageNumber="370">Tai Rom Yen National Park</collectingMunicipality>
, 
<location id="8EE629992535FFDCFA89AFECC7EC7E61" LSID="urn:lsid:plazi:treatment:0390CE542535FFDBFF6CAF10C58F7F15:8EE629992535FFDCFA89AFECC7EC7E61" country="Thailand" latitude="8.711945" longLatPrecision="21" longitude="99.51" municipality="Tai Rom Yen National Park" name="Pha San Yen" pageId="1" pageNumber="370" stateProvince="Surat Thani">Pha San Yen</location>
stream, 
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, 
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, ca. 
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<quantity id="4CC1D2A72535FFDCFD74AFC8C5FC7E61" box="[655,739,765,791]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="370" unit="m" value="1100.0">1100 m</quantity>
a.s.l.
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, 
<date id="FF8759822535FFDCFCD7AFC8C4E97E61" box="[812,1014,765,791]" pageId="1" pageNumber="370" value="2017-02-12">
<collectingDate id="EFC3A06A2535FFDCFCD7AFC8C4E97E61" box="[812,1014,765,791]" pageId="1" pageNumber="370" value="2017-02-12">12 February 2017</collectingDate>
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, leg. 
<collectorName id="26CC1A942535FFDCFBCAAFC8C3EE7E61" box="[1073,1265,765,791]" pageId="1" pageNumber="370">Pongsak Laudee.</collectorName>
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<paragraph id="8B867F422535FFDCFF3CAE14C4327E4D" blockId="1.[151,1436,729,971]" box="[199,813,801,827]" pageId="1" pageNumber="370">
<materialsCitation id="3B51751F2535FFDCFF3CAE14C4357E4D" ID-GBIF-Occurrence="2609011280" box="[199,810,801,827]" collectionCode="PSUNHM" pageId="1" pageNumber="370" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="5482C1E02535FFDCFF3CAE14C6277E4D" box="[199,312,801,827]" pageId="1" pageNumber="370" type="paratype">Paratypes</typeStatus>
: Same data as 
<typeStatus id="5482C1E02535FFDCFE27AE14C55E7E4D" box="[476,577,801,827]" pageId="1" pageNumber="370" type="holotype">holotype</typeStatus>
, 
<specimenCount id="9D3FB4CB2535FFDCFDB7AE14C5887E4D" box="[588,663,801,827]" pageId="1" pageNumber="370" type="male">1 male</specimenCount>
(
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)
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.
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<paragraph id="8B867F422535FFDCFF3CAE70C43A7EF5" blockId="1.[151,1436,729,971]" pageId="1" pageNumber="370">Etymology. The species epithet comes from Latin adjectives longus, -a, -um and caudatus, -a, -um, meaning “long tailed,” in reference to the long inferior appendages.</paragraph>
<paragraph id="8B867F422535FFDCFF3CAEB8C6767EBD" blockId="1.[151,1436,729,971]" pageId="1" pageNumber="370">Description. Length of each male forewing 7.0 mm (n = 2); specimens in alcohol with head, thorax, and forewings dark brown.</paragraph>
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<paragraph id="8B867F422535FFDCFF6CAA00C5697AA8" blockId="1.[151,1436,1845,2014]" pageId="1" pageNumber="370">
<emphasis id="B94DA3502535FFDCFF6CAA00C6427A38" bold="true" box="[151,349,1845,1870]" pageId="1" pageNumber="370">FIGURES 1A–1E.</emphasis>
Male genitalia of 
<taxonomicName id="4C3904C12535FFDCFDEEAA03C43E7A38" authority="Suwannarat &amp; Malicky &amp; Morse &amp; Laudee, 2019" authorityName="Suwannarat &amp; Malicky &amp; Morse &amp; Laudee" authorityYear="2019" box="[533,801,1846,1870]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="370" phylum="Arthropoda" rank="species" species="longicaudata" status="sp. nov.">
<emphasis id="B94DA3502535FFDCFDEEAA03C43E7A38" box="[533,801,1846,1870]" italics="true" pageId="1" pageNumber="370">Rhyacophila longicaudata</emphasis>
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<taxonomicNameLabel id="A27E1E2B2535FFDCFCDDAA02C4427A38" box="[806,861,1847,1870]" pageId="1" pageNumber="370" rank="species">n. sp.</taxonomicNameLabel>
1A, left lateral, with left inferior appendage omitted. 1B, dorsal. 1C, ventral. 1D, phallus, ventral. 1E, phallus, left lateral. IX = segment IX, Pre = preanal appendages, Api IX = apicodorsal lobe of segment IX, Ana = anal sclerites, Api B = apical band, Sag = sagittal appendage, Ter = tergal band, Pha = phallus, Bas Seg = basal segment of an inferior appendage (paired), Api Seg = apical segment of an inferior appendage (paired), Aed = aedeagus, Phal= phallotheca, End = endotheca.
</paragraph>
</caption>
<paragraph id="8B867F422536FFDFFF3CADACC65F7E61" blockId="2.[151,1437,152,1079]" pageId="2" pageNumber="371">
Male genitalia (
<figureCitation id="130263C72536FFDFFE87ADACC6FA7DC5" box="[380,485,152,179]" captionStart="FIGURES 1" captionStartId="1.[151,264,1845,1870]" captionTargetBox="[166,1416,1006,1783]" captionTargetId="figure@1.[151,1436,990,1818]" captionTargetPageId="1" captionText="FIGURES 1A–1E. Male genitalia of Rhyacophila longicaudata n. sp. 1A, left lateral, with left inferior appendage omitted. 1B, dorsal. 1C, ventral. 1D, phallus, ventral. 1E, phallus, left lateral. IX = segment IX, Pre = preanal appendages, Api IX = apicodorsal lobe of segment IX, Ana = anal sclerites, Api B = apical band, Sag = sagittal appendage, Ter = tergal band, Pha = phallus, Bas Seg = basal segment of an inferior appendage (paired), Api Seg = apical segment of an inferior appendage (paired), Aed = aedeagus, Phal= phallotheca, End = endotheca." figureDoi="http://doi.org/10.5281/zenodo.3772364" httpUri="https://zenodo.org/record/3772364/files/figure.png" pageId="2" pageNumber="371">Figures 1</figureCitation>
A–1E5). In lateral view (
<figureCitation id="130263C72536FFDFFCF7ADACC4787DC5" box="[780,871,153,179]" captionStart="FIGURES 1" captionStartId="1.[151,264,1845,1870]" captionTargetBox="[166,1416,1006,1783]" captionTargetId="figure@1.[151,1436,990,1818]" captionTargetPageId="1" captionText="FIGURES 1A–1E. Male genitalia of Rhyacophila longicaudata n. sp. 1A, left lateral, with left inferior appendage omitted. 1B, dorsal. 1C, ventral. 1D, phallus, ventral. 1E, phallus, left lateral. IX = segment IX, Pre = preanal appendages, Api IX = apicodorsal lobe of segment IX, Ana = anal sclerites, Api B = apical band, Sag = sagittal appendage, Ter = tergal band, Pha = phallus, Bas Seg = basal segment of an inferior appendage (paired), Api Seg = apical segment of an inferior appendage (paired), Aed = aedeagus, Phal= phallotheca, End = endotheca." figureDoi="http://doi.org/10.5281/zenodo.3772364" httpUri="https://zenodo.org/record/3772364/files/figure.png" pageId="2" pageNumber="371">Fig. 1A</figureCitation>
), segment IX subrectangular with broad posterolateral excision below midline, its apicodorsal lobe very long, lancet-like with short setae subapicodorsally, acute apically; in dorsal view (
<figureCitation id="130263C72536FFDFFE48ADD4C5157D8D" box="[435,522,225,251]" captionStart="FIGURES 1" captionStartId="1.[151,264,1845,1870]" captionTargetBox="[166,1416,1006,1783]" captionTargetId="figure@1.[151,1436,990,1818]" captionTargetPageId="1" captionText="FIGURES 1A–1E. Male genitalia of Rhyacophila longicaudata n. sp. 1A, left lateral, with left inferior appendage omitted. 1B, dorsal. 1C, ventral. 1D, phallus, ventral. 1E, phallus, left lateral. IX = segment IX, Pre = preanal appendages, Api IX = apicodorsal lobe of segment IX, Ana = anal sclerites, Api B = apical band, Sag = sagittal appendage, Ter = tergal band, Pha = phallus, Bas Seg = basal segment of an inferior appendage (paired), Api Seg = apical segment of an inferior appendage (paired), Aed = aedeagus, Phal= phallotheca, End = endotheca." figureDoi="http://doi.org/10.5281/zenodo.3772364" httpUri="https://zenodo.org/record/3772364/files/figure.png" pageId="2" pageNumber="371">Fig. 1B</figureCitation>
), annular portion of segment IX transversely straight anteriorly, appearing separated by concave suture from its apicodorsal lobe (
<bibRefCitation id="EFA802B32536FFDFFD2BAC30C4747C69" author="Schmid, F." box="[720,875,261,287]" pageId="2" pageNumber="371" refId="ref4292" refString="Schmid, F. (1970) Le genre Rhyacophila et la Famille des Rhyacophilidae (Trichoptera). Memoires de la Societe Entomologique du Canada, 66, 230 pp. https: // doi. org / 10.4039 / entm 10266 fv" type="book" year="1970">Schmid 1970</bibRefCitation>
), apicodorsal lobe of segment IX very long, hourglass-like with very deep incision in apical 3/4 of posterior margin, apices acute. Preanal appendages located at posterodorsal margin of annular portion of segment IX, very short, black, wart-like. Segment X reduced, obscure, except anal sclerites, the latter oval in lateral view, rounded posteriorly, positioned underneath anterior portion of apicodorsal lobe of segment IX. Inferior appendages very long, more than twice as long as apicodorsal lobe of segment IX, cylindrical; each with basal segment about six times as long as apical segment, dorsal and ventral margins parallel in lateral view (
<figureCitation id="130263C72536FFDFFE5DACE8C51F7C81" box="[422,512,477,503]" captionStart="FIGURES 1" captionStartId="1.[151,264,1845,1870]" captionTargetBox="[166,1416,1006,1783]" captionTargetId="figure@1.[151,1436,990,1818]" captionTargetPageId="1" captionText="FIGURES 1A–1E. Male genitalia of Rhyacophila longicaudata n. sp. 1A, left lateral, with left inferior appendage omitted. 1B, dorsal. 1C, ventral. 1D, phallus, ventral. 1E, phallus, left lateral. IX = segment IX, Pre = preanal appendages, Api IX = apicodorsal lobe of segment IX, Ana = anal sclerites, Api B = apical band, Sag = sagittal appendage, Ter = tergal band, Pha = phallus, Bas Seg = basal segment of an inferior appendage (paired), Api Seg = apical segment of an inferior appendage (paired), Aed = aedeagus, Phal= phallotheca, End = endotheca." figureDoi="http://doi.org/10.5281/zenodo.3772364" httpUri="https://zenodo.org/record/3772364/files/figure.png" pageId="2" pageNumber="371">Fig. 1A</figureCitation>
), broader basally in ventral view (
<figureCitation id="130263C72536FFDFFC7DACE8C4C27C81" box="[902,989,477,503]" captionStart="FIGURES 1" captionStartId="1.[151,264,1845,1870]" captionTargetBox="[166,1416,1006,1783]" captionTargetId="figure@1.[151,1436,990,1818]" captionTargetPageId="1" captionText="FIGURES 1A–1E. Male genitalia of Rhyacophila longicaudata n. sp. 1A, left lateral, with left inferior appendage omitted. 1B, dorsal. 1C, ventral. 1D, phallus, ventral. 1E, phallus, left lateral. IX = segment IX, Pre = preanal appendages, Api IX = apicodorsal lobe of segment IX, Ana = anal sclerites, Api B = apical band, Sag = sagittal appendage, Ter = tergal band, Pha = phallus, Bas Seg = basal segment of an inferior appendage (paired), Api Seg = apical segment of an inferior appendage (paired), Aed = aedeagus, Phal= phallotheca, End = endotheca." figureDoi="http://doi.org/10.5281/zenodo.3772364" httpUri="https://zenodo.org/record/3772364/files/figure.png" pageId="2" pageNumber="371">Fig. 1C</figureCitation>
), with longitudinal ridge dorsomesally; apical segment short and ovoid in lateral and ventral views, with numerous stout subapicomesal setae. Tergal band from dorsal base of phallotheca terminating in sagittal appendage articulating with apical band connected with root of anal sclerite. In dorsal and ventral views (
<figureCitation id="130263C72536FFDFFD76AF7CC43E7F15" box="[653,801,585,611]" captionStart="FIGURES 1" captionStartId="1.[151,264,1845,1870]" captionTargetBox="[166,1416,1006,1783]" captionTargetId="figure@1.[151,1436,990,1818]" captionTargetPageId="1" captionText="FIGURES 1A–1E. Male genitalia of Rhyacophila longicaudata n. sp. 1A, left lateral, with left inferior appendage omitted. 1B, dorsal. 1C, ventral. 1D, phallus, ventral. 1E, phallus, left lateral. IX = segment IX, Pre = preanal appendages, Api IX = apicodorsal lobe of segment IX, Ana = anal sclerites, Api B = apical band, Sag = sagittal appendage, Ter = tergal band, Pha = phallus, Bas Seg = basal segment of an inferior appendage (paired), Api Seg = apical segment of an inferior appendage (paired), Aed = aedeagus, Phal= phallotheca, End = endotheca." figureDoi="http://doi.org/10.5281/zenodo.3772364" httpUri="https://zenodo.org/record/3772364/files/figure.png" pageId="2" pageNumber="371">Figs. 1A, 1D</figureCitation>
), retracted phallus clavate, with phallotheca rectangular, endotheca thicker and oval with numerous spines, and aedeagus long and spoon-like with two long internal spines and five curved hooks apically (only four visible ventrally); in lateral view (
<figureCitation id="130263C72536FFDFFC16AFA4C35E7FDD" box="[1005,1089,657,683]" captionStart="FIGURES 1" captionStartId="1.[151,264,1845,1870]" captionTargetBox="[166,1416,1006,1783]" captionTargetId="figure@1.[151,1436,990,1818]" captionTargetPageId="1" captionText="FIGURES 1A–1E. Male genitalia of Rhyacophila longicaudata n. sp. 1A, left lateral, with left inferior appendage omitted. 1B, dorsal. 1C, ventral. 1D, phallus, ventral. 1E, phallus, left lateral. IX = segment IX, Pre = preanal appendages, Api IX = apicodorsal lobe of segment IX, Ana = anal sclerites, Api B = apical band, Sag = sagittal appendage, Ter = tergal band, Pha = phallus, Bas Seg = basal segment of an inferior appendage (paired), Api Seg = apical segment of an inferior appendage (paired), Aed = aedeagus, Phal= phallotheca, End = endotheca." figureDoi="http://doi.org/10.5281/zenodo.3772364" httpUri="https://zenodo.org/record/3772364/files/figure.png" pageId="2" pageNumber="371">Fig. 1E</figureCitation>
) protracted phallus long, phallotheca cylindrical, endotheca with numerous spines basally and membranous apically; aedeagus slender and cylindrical with two long internal spines of which dorsal spine about twice as long as ventral spine and with five long hooks apically.
</paragraph>
<paragraph id="8B867F422536FFDFFF3CAE14C6307941" blockId="2.[151,1437,152,1079]" pageId="2" pageNumber="371">
Diagnosis. 
<taxonomicName id="4C3904C12536FFDFFEBCAE14C56D7E4D" authority="Suwannarat &amp; Malicky &amp; Morse &amp; Laudee, 2019" authorityName="Suwannarat &amp; Malicky &amp; Morse &amp; Laudee" authorityYear="2019" box="[327,626,801,827]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="longicaudata" status="sp. nov.">
<emphasis id="B94DA3502536FFDFFEBCAE14C56D7E4D" box="[327,626,801,827]" italics="true" pageId="2" pageNumber="371">Rhyacophila longicaudata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27E1E2B2536FFDFFD80AE14C5A67E4D" box="[635,697,801,827]" pageId="2" pageNumber="371" rank="species">n. sp.</taxonomicNameLabel>
is a member of 
<taxonomicName id="4C3904C12536FFDFFC83AE17C3327E4D" authorityName="Iwata" authorityYear="1927" box="[888,1069,801,827]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="nigrocephala">
<emphasis id="B94DA3502536FFDFFC83AE17C4967E4D" box="[888,905,802,827]" italics="true" pageId="2" pageNumber="371">R</emphasis>
. 
<emphasis id="B94DA3502536FFDFFC62AE14C3327E4D" box="[921,1069,801,827]" italics="true" pageId="2" pageNumber="371">nigrocephala</emphasis>
</taxonomicName>
Group (
<bibRefCitation id="EFA802B32536FFDFFB75AE14C2377E4D" author="Schmid, F." box="[1166,1320,801,827]" pageId="2" pageNumber="371" refId="ref4292" refString="Schmid, F. (1970) Le genre Rhyacophila et la Famille des Rhyacophilidae (Trichoptera). Memoires de la Societe Entomologique du Canada, 66, 230 pp. https: // doi. org / 10.4039 / entm 10266 fv" type="book" year="1970">Schmid 1970</bibRefCitation>
) in which there is a long apicodorsal lobe of segment IX extending caudad from the annular portion of that segment, the fused anal sclerites are reduced, apical and tergal bands are present and sclerotized, and in most species (including this new species) the phallic parameres are absent and the basal segments of the inferior appendages are fused with each other at their extreme bases. This species is similar to R. 
<emphasis id="B94DA3502536FFDFFCD2AE84C4877EBD" box="[809,920,945,971]" italics="true" pageId="2" pageNumber="371">drosampa</emphasis>
<bibRefCitation id="EFA802B32536FFDFFC5AAE84C3257EBD" author="Schmid, F." box="[929,1082,945,971]" pageId="2" pageNumber="371" refId="ref4292" refString="Schmid, F. (1970) Le genre Rhyacophila et la Famille des Rhyacophilidae (Trichoptera). Memoires de la Societe Entomologique du Canada, 66, 230 pp. https: // doi. org / 10.4039 / entm 10266 fv" type="book" year="1970">Schmid 1970</bibRefCitation>
, R. 
<emphasis id="B94DA3502536FFDFFB91AE84C3AF7EBD" box="[1130,1200,945,971]" italics="true" pageId="2" pageNumber="371">laocai</emphasis>
<bibRefCitation id="EFA802B32536FFDFFB4CAE84C7CD7E99" author="Armitage, B. J. &amp; Arefina, T." pageId="2" pageNumber="371" pagination="95 - 124" refId="ref3533" refString="Armitage, B. J. &amp; Arefina, T. (2003) The genus Rhyacophila Pictet (Trichoptera: Rhyacophilidae) in Vietnam. Aquatic Insects, 25, 95 - 124. https: // doi. org / 10.1076 / aqin. 25.2.95.14037" type="journal article" year="2003">Armitage &amp; Arefina 2003</bibRefCitation>
, 
<taxonomicName id="4C3904C12536FFDFFF26AEE3C6C67E99" authority="Schmid 1970" authorityName="Schmid" authorityYear="1970" box="[221,473,981,1007]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="lhabu">
<emphasis id="B94DA3502536FFDFFF26AEE3C7F17E99" box="[221,238,982,1007]" italics="true" pageId="2" pageNumber="371">R</emphasis>
. 
<emphasis id="B94DA3502536FFDFFF00AEE0C6247E99" box="[251,315,981,1007]" italics="true" pageId="2" pageNumber="371">lhabu</emphasis>
<bibRefCitation id="EFA802B32536FFDFFEB9AEE0C6C67E99" author="Schmid, F." box="[322,473,981,1007]" pageId="2" pageNumber="371" refId="ref4292" refString="Schmid, F. (1970) Le genre Rhyacophila et la Famille des Rhyacophilidae (Trichoptera). Memoires de la Societe Entomologique du Canada, 66, 230 pp. https: // doi. org / 10.4039 / entm 10266 fv" type="book" year="1970">Schmid 1970</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C3904C12536FFDFFE1FAEE3C5F67E99" authority="Mey 1996" authorityName="Mey" authorityYear="1996" box="[484,745,981,1007]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="paratecta">
<emphasis id="B94DA3502536FFDFFE1FAEE3C6EA7E99" box="[484,501,982,1007]" italics="true" pageId="2" pageNumber="371">R</emphasis>
. 
<emphasis id="B94DA3502536FFDFFDF9AEE3C5727E99" box="[514,621,982,1007]" italics="true" pageId="2" pageNumber="371">paratecta</emphasis>
<bibRefCitation id="EFA802B32536FFDFFD8FAEE0C5F67E99" author="Mey, W." box="[628,745,981,1007]" pageId="2" pageNumber="371" pagination="39 - 65" refId="ref4103" refString="Mey, W. (1996) Die Kocherfliegenfauna des Fan Si Pan-Massivs in NordVietnam. Beitrage zur Entomologie, 46 (1), 39 - 65." type="journal article" year="1996">Mey 1996</bibRefCitation>
</taxonomicName>
, and 
<taxonomicName id="4C3904C12536FFDFFCD8AEE3C3507E99" authority="Malicky 1991" authorityName="Malicky" authorityYear="1991" box="[803,1103,981,1007]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="mayestril">
<emphasis id="B94DA3502536FFDFFCD8AEE3C42B7E99" box="[803,820,982,1007]" italics="true" pageId="2" pageNumber="371">R</emphasis>
. 
<emphasis id="B94DA3502536FFDFFCBAAEE0C4B67E99" box="[833,937,981,1007]" italics="true" pageId="2" pageNumber="371">mayestril</emphasis>
<bibRefCitation id="EFA802B32536FFDFFC4BAEE0C3507E99" author="Malicky, H." box="[944,1103,981,1007]" pageId="2" pageNumber="371" pagination="381 - 384" refId="ref3894" refString="Malicky, H. (1991) Some unusual caddisflies (Trichoptera) from southeastern Asia (Studies on caddisflies of Thailand, No. 5). In: Tomaszewski, C. (Ed.), Proceedings of the 6 th International Symposium on Trichoptera. Adam Mickiewicz University Press, Poznan, Poland, pp. 381 - 384." type="book chapter" year="1991">Malicky 1991</bibRefCitation>
</taxonomicName>
. However, R. 
<taxonomicName id="4C3904C12536FFDFFB16AEE0C29E7E99" authority="Suwannarat &amp; Malicky &amp; Morse &amp; Laudee, 2019" authorityName="Suwannarat &amp; Malicky &amp; Morse &amp; Laudee" authorityYear="2019" box="[1261,1409,981,1007]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="longicaudata" status="sp. nov.">
<emphasis id="B94DA3502536FFDFFB16AEE0C29E7E99" box="[1261,1409,981,1007]" italics="true" pageId="2" pageNumber="371">longicaudata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27E1E2B2536FFDFFA7CAEE0C7A87965" pageId="2" pageNumber="371" rank="species">n. sp.</taxonomicNameLabel>
differs from those by the unusually long inferior appendages and by the presence of long hooks at the apex of the aedeagus.
</paragraph>
<paragraph id="8B867F422536FFDFFF6CA9BCC5E179D5" blockId="2.[151,766,1161,1223]" box="[151,766,1161,1187]" pageId="2" pageNumber="371">
<taxonomicName id="4C3904C12536FFDFFF6CA9BCC6DD79D5" authority="Suwannarat &amp; Malicky &amp; Morse &amp; Laudee, 2019" authorityName="Suwannarat &amp; Malicky &amp; Morse &amp; Laudee" authorityYear="2019" box="[151,450,1161,1187]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="aksornkoaei" status="sp. nov.">
<emphasis id="B94DA3502536FFDFFF6CA9BCC6DD79D5" box="[151,450,1161,1187]" italics="true" pageId="2" pageNumber="371">Rhyacophila aksornkoaei</emphasis>
</taxonomicName>
<emphasis id="B94DA3502536FFDFFE32A9BCC5E179D5" bold="true" box="[457,766,1161,1187]" pageId="2" pageNumber="371">
<taxonomicNameLabel id="A27E1E2B2536FFDFFE32A9BCC51379D5" box="[457,524,1161,1187]" pageId="2" pageNumber="371" rank="species">sp. n.</taxonomicNameLabel>
Laudee &amp; Malicky
</emphasis>
</paragraph>
<paragraph id="8B867F422536FFDFFF6CA998C63979B1" blockId="2.[151,766,1161,1223]" box="[151,294,1197,1223]" pageId="2" pageNumber="371">
<figureCitation id="130263C72536FFDFFF6CA998C7FD79B1" box="[151,226,1197,1223]" captionStart="FIGURES 2" captionStartId="3.[151,264,1238,1263]" captionTargetBox="[151,1436,181,1213]" captionTargetId="figure@3.[151,1436,181,1213]" captionTargetPageId="3" captionText="FIGURES 2A–2C. Male genitalia of Rhyacophila aksornkoaei n. sp. 2A, left lateral, with left inferior appendage omitted. 2B, dorsal. 2C, ventral. Par = parameres, Dor = dorsal process of the phallic apparatus, Ven = ventral plate of the aedeagus." figureDoi="http://doi.org/10.5281/zenodo.3772366" httpUri="https://zenodo.org/record/3772366/files/figure.png" pageId="2" pageNumber="371">Figs. 2</figureCitation>
A–2C
</paragraph>
<paragraph id="8B867F422536FFDFFF6CA9C0C3AE7845" blockId="2.[151,1437,1269,1979]" pageId="2" pageNumber="371">
Type material. 
<materialsCitation id="3B51751F2536FFDFFEC5A9C0C3AE7845" ID-GBIF-Occurrence="2609011276" collectingDate="2018-03-31" collectionCode="PSUNHM" collectorName="Nannaphat Suwannarat." country="Thailand" elevation="802" latitude="8.479444" location="Tha Di" longLatPrecision="21" longitude="99.751945" municipality="Khao Luang National Park" pageId="2" pageNumber="371" specimenCount="1" stateProvince="Surat Thani" typeStatus="holotype">
<typeStatus id="5482C1E02536FFDFFEC5A9C0C6B97879" box="[318,422,1269,1295]" pageId="2" pageNumber="371" type="holotype">Holotype</typeStatus>
male (
<collectionCode id="ED28E7872536FFDFFE0AA9C0C5727879" box="[497,621,1269,1295]" pageId="2" pageNumber="371">PSUNHM</collectionCode>
). 
<collectingCountry id="F32E3FD22536FFDFFD85A9C0C40C7879" box="[638,787,1269,1295]" name="Thailand" pageId="2" pageNumber="371">THAILAND</collectingCountry>
: 
<collectingRegion id="49FDB1A02536FFDFFCE6A9C0C3147879" box="[797,1035,1269,1295]" country="Thailand" name="Surat Thani" pageId="2" pageNumber="371">Surat Thani Province</collectingRegion>
, 
<collectingMunicipality id="6BE2E5382536FFDFFBEDA9C0C25B7879" box="[1046,1348,1269,1295]" pageId="2" pageNumber="371">Khao Luang National Park</collectingMunicipality>
, 
<location id="8EE629992536FFDFFAB5A9C0C2837879" LSID="urn:lsid:plazi:treatment:0390CE542535FFDBFF6CAF10C58F7F15:8EE629992536FFDFFAB5A9C0C2837879" box="[1358,1436,1269,1295]" country="Thailand" latitude="8.479444" longLatPrecision="21" longitude="99.751945" municipality="Khao Luang National Park" name="Tha Di" pageId="2" pageNumber="371" stateProvince="Surat Thani">Tha Di</location>
stream, 
<geoCoordinate id="EE0D19852536FFDFFF0BA82DC6767845" box="[240,361,1304,1331]" degrees="8" direction="north" minutes="28" orientation="latitude" pageId="2" pageNumber="371" precision="15" seconds="46" value="8.479444">8°28′46″N</geoCoordinate>
, 
<geoCoordinate id="EE0D19852536FFDFFE8EA82DC6E67845" box="[373,505,1304,1331]" degrees="99" direction="east" minutes="45" orientation="longitude" pageId="2" pageNumber="371" precision="15" seconds="07" value="99.751945">99°45′07″E</geoCoordinate>
, ca. 
<elevation id="001498712536FFDFFDD7A82CC5B07845" box="[556,687,1305,1331]" metricMagnitude="2" metricUnit="m" metricValue="8.02" pageId="2" pageNumber="371" unit="m" value="802.0">
<quantity id="4CC1D2A72536FFDFFDD7A82CC56C7845" box="[556,627,1305,1331]" metricMagnitude="2" metricUnit="m" metricValue="8.02" pageId="2" pageNumber="371" unit="m" value="802.0">802 m</quantity>
a.s.l.
</elevation>
, 
<date id="FF8759822536FFDFFD47A82CC4757845" box="[700,874,1305,1331]" pageId="2" pageNumber="371" value="2018-03-31">
<collectingDate id="EFC3A06A2536FFDFFD47A82CC4757845" box="[700,874,1305,1331]" pageId="2" pageNumber="371" value="2018-03-31">31 March 2018</collectingDate>
</date>
, leg. 
<collectorName id="26CC1A942536FFDFFC5EA82CC3AE7845" box="[933,1201,1305,1331]" pageId="2" pageNumber="371">Nannaphat Suwannarat.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8B867F422536FFDFFF3CA808C4327821" blockId="2.[151,1437,1269,1979]" box="[199,813,1340,1367]" pageId="2" pageNumber="371">
<materialsCitation id="3B51751F2536FFDFFF3CA808C4357821" ID-GBIF-Occurrence="2609011281" box="[199,810,1340,1367]" collectionCode="PSUNHM" pageId="2" pageNumber="371" specimenCount="5" specimenCount-male="5" typeStatus="paratype">
<typeStatus id="5482C1E02536FFDFFF3CA808C6277821" box="[199,312,1341,1367]" pageId="2" pageNumber="371" type="paratype">Paratypes</typeStatus>
: Same data as 
<typeStatus id="5482C1E02536FFDFFE27A808C55E7821" box="[476,577,1341,1367]" pageId="2" pageNumber="371" type="holotype">holotype</typeStatus>
, 
<specimenCount id="9D3FB4CB2536FFDFFDB7A809C5887821" box="[588,663,1340,1367]" pageId="2" pageNumber="371" type="male">5 male</specimenCount>
(
<collectionCode id="ED28E7872536FFDFFD5DA808C43D7821" box="[678,802,1341,1367]" pageId="2" pageNumber="371">PSUNHM</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B867F422536FFDFFF3CA854C25B780D" blockId="2.[151,1437,1269,1979]" box="[199,1348,1377,1403]" pageId="2" pageNumber="371">Etymology. The species is dedicated to Prof. Dr. Sanit Aksornkoae, who is a prominent Thai ecologist.</paragraph>
<paragraph id="8B867F422536FFDFFF3CA8B0C67678B5" blockId="2.[151,1437,1269,1979]" pageId="2" pageNumber="371">Description. Length of each male forewing 11.0 mm (n = 2); specimens in alcohol with head, thorax, and forewings dark brown.</paragraph>
<paragraph id="8B867F422536FFDFFF3CA8F8C49B7ACD" blockId="2.[151,1437,1269,1979]" pageId="2" pageNumber="371">
Male genitalia (
<figureCitation id="130263C72536FFDFFE8EA8F8C6C07891" box="[373,479,1485,1511]" captionStart="FIGURES 2" captionStartId="3.[151,264,1238,1263]" captionTargetBox="[151,1436,181,1213]" captionTargetId="figure@3.[151,1436,181,1213]" captionTargetPageId="3" captionText="FIGURES 2A–2C. Male genitalia of Rhyacophila aksornkoaei n. sp. 2A, left lateral, with left inferior appendage omitted. 2B, dorsal. 2C, ventral. Par = parameres, Dor = dorsal process of the phallic apparatus, Ven = ventral plate of the aedeagus." figureDoi="http://doi.org/10.5281/zenodo.3772366" httpUri="https://zenodo.org/record/3772366/files/figure.png" pageId="2" pageNumber="371">Figures 2</figureCitation>
A–2C). Segment IX anterior margins convex dorsally and concave ventrally, posterior margins convex below its apicodorsal lobe, longitudinally longest at 2/3 height, anterodorsal margin concave except for small triangular mesal projection, its apicodorsal lobe about as long as annular portion of segment, divided by deep V-shaped apical incision nearly half its length mesally, slightly concave laterally, dark brown apically and subapically. Preanal appendages drop-like, shorter than apicodorsal lobe of segment IX. Segment X trapezoid, hidden below apicodorsal lobe of segment IX and between preanal appendages. Anal sclerite flat oval, oblique in lateral view. Inferior appendages each with basal segment shaped as pararellogram in lateral view, in ventral view slender in middle; apical segment mitten-shaped in mesal and ventral views, with short triangular “thumb” and with stout marginal and submarginal setae along subapicomesal and apicomesal margins. Phallus complex; phallobase short, tubular, its dorsal process somewhat triangular, convex dorsally, acute apically; ventral process tubular with expanded base and blunt apex; parameres long, tubular, expanded and bent slightly upward, each with tuft of bristles subapicomesally and apically, slightly expanded and truncate in lateral view and slightly convergent and blunt in ventral view; aedeagus as long as parameres, needle-like, expanded basally and upturned apically in lateral view (
<figureCitation id="130263C72536FFDFFF5BAA94C7E57ACD" box="[160,250,1953,1979]" captionStart="FIGURES 2" captionStartId="3.[151,264,1238,1263]" captionTargetBox="[151,1436,181,1213]" captionTargetId="figure@3.[151,1436,181,1213]" captionTargetPageId="3" captionText="FIGURES 2A–2C. Male genitalia of Rhyacophila aksornkoaei n. sp. 2A, left lateral, with left inferior appendage omitted. 2B, dorsal. 2C, ventral. Par = parameres, Dor = dorsal process of the phallic apparatus, Ven = ventral plate of the aedeagus." figureDoi="http://doi.org/10.5281/zenodo.3772366" httpUri="https://zenodo.org/record/3772366/files/figure.png" pageId="2" pageNumber="371">Fig. 2A</figureCitation>
), uniformly slender and straight in ventral view (
<figureCitation id="130263C72536FFDFFCE5AA94C4687ACD" box="[798,887,1953,1979]" captionStart="FIGURES 2" captionStartId="3.[151,264,1238,1263]" captionTargetBox="[151,1436,181,1213]" captionTargetId="figure@3.[151,1436,181,1213]" captionTargetPageId="3" captionText="FIGURES 2A–2C. Male genitalia of Rhyacophila aksornkoaei n. sp. 2A, left lateral, with left inferior appendage omitted. 2B, dorsal. 2C, ventral. Par = parameres, Dor = dorsal process of the phallic apparatus, Ven = ventral plate of the aedeagus." figureDoi="http://doi.org/10.5281/zenodo.3772366" httpUri="https://zenodo.org/record/3772366/files/figure.png" pageId="2" pageNumber="371">Fig. 2C</figureCitation>
).
</paragraph>
<caption id="DF462FCA2537FFDEFF6CA9E3C2447865" ID-DOI="http://doi.org/10.5281/zenodo.3772366" ID-Zenodo-Dep="3772366" httpUri="https://zenodo.org/record/3772366/files/figure.png" pageId="3" pageNumber="372" startId="3.[151,264,1238,1263]" targetBox="[151,1436,181,1213]" targetPageId="3">
<paragraph id="8B867F422537FFDEFF6CA9E3C2447865" blockId="3.[151,1436,1238,1299]" pageId="3" pageNumber="372">
<emphasis id="B94DA3502537FFDEFF6CA9E3C6407999" bold="true" box="[151,351,1238,1263]" pageId="3" pageNumber="372">FIGURES 2A–2C.</emphasis>
Male genitalia of 
<taxonomicName id="4C3904C12537FFDEFDE2A9E2C4047999" authority="Suwannarat &amp; Malicky &amp; Morse &amp; Laudee, 2019" authorityName="Suwannarat &amp; Malicky &amp; Morse &amp; Laudee" authorityYear="2019" box="[537,795,1239,1263]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="species" species="aksornkoaei" status="sp. nov.">
<emphasis id="B94DA3502537FFDEFDE2A9E2C4047999" box="[537,795,1239,1263]" italics="true" pageId="3" pageNumber="372">Rhyacophila aksornkoaei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27E1E2B2537FFDEFCDAA9E2C4477998" box="[801,856,1239,1262]" pageId="3" pageNumber="372" rank="species">n. sp.</taxonomicNameLabel>
2A, left lateral, with left inferior appendage omitted. 2B, dorsal. 2C, ventral. Par = parameres, Dor = dorsal process of the phallic apparatus, Ven = ventral plate of the aedeagus.
</paragraph>
</caption>
<paragraph id="8B867F422537FFDEFF3CA874C54D7A46" blockId="3.[151,1437,1345,1840]" pageId="3" pageNumber="372">
Diagnosis. This new species is a member of 
<taxonomicName id="4C3904C12537FFDEFD41A874C4BB782D" authorityName="Morton" authorityYear="1900" box="[698,932,1345,1371]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="species" species="anatina">
<emphasis id="B94DA3502537FFDEFD41A874C4BB782D" box="[698,932,1345,1371]" italics="true" pageId="3" pageNumber="372">Rhyacophila anatina</emphasis>
</taxonomicName>
Group (
<bibRefCitation id="EFA802B32537FFDEFBFAA874C387782D" author="Schmid, F." box="[1025,1176,1345,1371]" pageId="3" pageNumber="372" refId="ref4292" refString="Schmid, F. (1970) Le genre Rhyacophila et la Famille des Rhyacophilidae (Trichoptera). Memoires de la Societe Entomologique du Canada, 66, 230 pp. https: // doi. org / 10.4039 / entm 10266 fv" type="book" year="1970">Schmid 1970</bibRefCitation>
) and is similar to other species of this Group found in Asia. According to 
<bibRefCitation id="EFA802B32537FFDEFD3EA850C45A78F6" author="Sun, C." box="[709,837,1381,1408]" pageId="3" pageNumber="372" pagination="273 - 278" refId="ref4336" refString="Sun, C. (2016) Two new species of the Rhyacophila anatina Species Group from China (Trichoptera: Rhyacophilidae). Zootaxa, 4085 (2), 273 - 278. https: // doi. org / 10.11646 / zootaxa. 4085.2.7" type="journal article" year="2016">Sun (2016)</bibRefCitation>
, the males of this Group can be diagnosed as follows: “segment IX is short ventrally; the apicodorsal lobe of segment IX is strong and bilobed; the preanal appendages are large and closely joined to the apicodorsal lobe, segment X is vertical; the anal sclerites are usually paired, without roots; the apical band is strongly sclerotized and with its two lateral arms not joined to each other at the base; the tergal band is short and membranous; the basal segment of the inferior appendages are usually long, and the apical segments are short, the phallotheca is cylindrical and the endotheca well developed; the aedeagus is slender, tubular; parameres are slender, with the distal ends heavily bristled.” 
<taxonomicName id="4C3904C12537FFDEFC63AB08C3AA7B21" authority="Suwannarat &amp; Malicky &amp; Morse &amp; Laudee, 2019" authorityName="Suwannarat &amp; Malicky &amp; Morse &amp; Laudee" authorityYear="2019" box="[920,1205,1597,1623]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="species" species="aksornkoaei" status="sp. nov.">
<emphasis id="B94DA3502537FFDEFC63AB08C3AA7B21" box="[920,1205,1597,1623]" italics="true" pageId="3" pageNumber="372">Rhyacophila aksornkoaei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27E1E2B2537FFDEFB40AB0BC3E87B2E" box="[1211,1271,1598,1624]" pageId="3" pageNumber="372" rank="species">sp. n.</taxonomicNameLabel>
is very similar to 
<taxonomicName id="4C3904C12537FFDEFF4FAB57C5837B0D" authority="Laudee &amp; Malicky 2014" authorityName="Laudee &amp; Malicky" authorityYear="2014" box="[180,668,1633,1660]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="species" species="suratthaniensis">
R. 
<emphasis id="B94DA3502537FFDEFF2EAB54C69F7B0D" box="[213,384,1633,1659]" italics="true" pageId="3" pageNumber="372">suratthaniensis</emphasis>
<bibRefCitation id="EFA802B32537FFDEFE73AB54C5837B0D" author="Laudee, P. &amp; Malicky, H." box="[392,668,1633,1659]" pageId="3" pageNumber="372" pagination="161 - 169" refId="ref3828" refString="Laudee, P. &amp; Malicky, H. (2014) Trichoptera fauna from Nakhon Si Thammarat Range (southern Thailand), with the description of a new species of Rhyacophila Pictet, 1834 (Trichoptera: Rhyacophilidae). Aquatic Insects, 36 (3 - 4), 161 - 169. https: // doi. org / 10.1080 / 01650424.2015.1064965" type="journal article" year="2014">Laudee &amp; Malicky 2014</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C3904C12537FFDEFD28AB57C38B7B0D" authority="Schmid &amp; Denning 1971" authorityName="Schmid &amp; Denning" authorityYear="1971" box="[723,1172,1633,1660]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="species" species="petersorum">
<emphasis id="B94DA3502537FFDEFD28AB57C5FB7B0D" box="[723,740,1634,1659]" italics="true" pageId="3" pageNumber="372">R</emphasis>
. 
<emphasis id="B94DA3502537FFDEFD08AB57C46C7B0D" box="[755,883,1634,1659]" italics="true" pageId="3" pageNumber="372">petersorum</emphasis>
Schmid &amp; 
<bibRefCitation id="EFA802B32537FFDEFC0FAB54C38B7B0D" author="Denning, D. G. &amp; Schmid, F." box="[1012,1172,1633,1659]" pageId="3" pageNumber="372" pagination="1553 - 1556" refId="ref3703" refString="Denning, D. G. &amp; Schmid, F. (1971) Descriptions of four new Rhyacophila (Trichoptera: Rhyacophilidae). Canadian Entomologist, 103, 1553 - 1556. https: // doi. org / 10.4039 / Ent 1031553 - 11" type="journal article" year="1971">Denning 1971</bibRefCitation>
</taxonomicName>
(in 
<bibRefCitation id="EFA802B32537FFDEFB39AB54C7CD7BE9" author="Denning, D. G. &amp; Schmid, F." pageId="3" pageNumber="372" pagination="1553 - 1556" refId="ref3703" refString="Denning, D. G. &amp; Schmid, F. (1971) Descriptions of four new Rhyacophila (Trichoptera: Rhyacophilidae). Canadian Entomologist, 103, 1553 - 1556. https: // doi. org / 10.4039 / Ent 1031553 - 11" type="journal article" year="1971">Denning &amp; Schmid 1971</bibRefCitation>
), but can be distinguished in that (1) the phallus of the new species is armed with a ventral plate that is absent in 
<taxonomicName id="4C3904C12537FFDEFF4FAB9FC5847BB5" authority="Laudee &amp; Malicky 2014" authorityName="Laudee &amp; Malicky" authorityYear="2014" box="[180,667,1705,1732]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="species" species="suratthaniensis">
<emphasis id="B94DA3502537FFDEFF4FAB9FC7DA7BB5" box="[180,197,1706,1731]" italics="true" pageId="3" pageNumber="372">R</emphasis>
. 
<emphasis id="B94DA3502537FFDEFF2FAB9CC6607BB5" box="[212,383,1705,1731]" italics="true" pageId="3" pageNumber="372">suratthaniensis</emphasis>
<bibRefCitation id="EFA802B32537FFDEFE7CAB9CC5847BB5" author="Laudee, P. &amp; Malicky, H." box="[391,667,1705,1731]" pageId="3" pageNumber="372" pagination="161 - 169" refId="ref3828" refString="Laudee, P. &amp; Malicky, H. (2014) Trichoptera fauna from Nakhon Si Thammarat Range (southern Thailand), with the description of a new species of Rhyacophila Pictet, 1834 (Trichoptera: Rhyacophilidae). Aquatic Insects, 36 (3 - 4), 161 - 169. https: // doi. org / 10.1080 / 01650424.2015.1064965" type="journal article" year="2014">Laudee &amp; Malicky 2014</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C3904C12537FFDEFD29AB9FC38C7BB5" authority="Schmid &amp; Denning 1971" authorityName="Schmid &amp; Denning" authorityYear="1971" box="[722,1171,1705,1732]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="species" species="petersorum">
<emphasis id="B94DA3502537FFDEFD29AB9FC5FC7BB5" box="[722,739,1706,1731]" italics="true" pageId="3" pageNumber="372">R</emphasis>
. 
<emphasis id="B94DA3502537FFDEFD09AB9FC46D7BB5" box="[754,882,1706,1731]" italics="true" pageId="3" pageNumber="372">petersorum</emphasis>
Schmid &amp; 
<bibRefCitation id="EFA802B32537FFDEFC08AB9CC38C7BB5" author="Denning, D. G. &amp; Schmid, F." box="[1011,1171,1705,1731]" pageId="3" pageNumber="372" pagination="1553 - 1556" refId="ref3703" refString="Denning, D. G. &amp; Schmid, F. (1971) Descriptions of four new Rhyacophila (Trichoptera: Rhyacophilidae). Canadian Entomologist, 103, 1553 - 1556. https: // doi. org / 10.4039 / Ent 1031553 - 11" type="journal article" year="1971">Denning 1971</bibRefCitation>
</taxonomicName>
(in 
<bibRefCitation id="EFA802B32537FFDEFB3AAB9CC7CD7B91" author="Denning, D. G. &amp; Schmid, F." pageId="3" pageNumber="372" pagination="1553 - 1556" refId="ref3703" refString="Denning, D. G. &amp; Schmid, F. (1971) Descriptions of four new Rhyacophila (Trichoptera: Rhyacophilidae). Canadian Entomologist, 103, 1553 - 1556. https: // doi. org / 10.4039 / Ent 1031553 - 11" type="journal article" year="1971">Denning &amp; Schmid 1971</bibRefCitation>
), (2) the distal ends of the parameres of 
<taxonomicName id="4C3904C12537FFDEFD68ABF8C4AE7B91" authority="Suwannarat &amp; Malicky &amp; Morse &amp; Laudee, 2019" authorityName="Suwannarat &amp; Malicky &amp; Morse &amp; Laudee" authorityYear="2019" box="[659,945,1741,1767]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="species" species="aksornkoaei" status="sp. nov.">
<emphasis id="B94DA3502537FFDEFD68ABF8C4AE7B91" box="[659,945,1741,1767]" italics="true" pageId="3" pageNumber="372">Rhyacophila aksornkoaei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27E1E2B2537FFDEFC4CABFBC4EC7B9E" box="[951,1011,1742,1768]" pageId="3" pageNumber="372" rank="species">sp. n.</taxonomicNameLabel>
are bent upward and bristled apically, but bristles are arranged like a toothbrush in 
<taxonomicName id="4C3904C12537FFDEFD6BABC7C4447A7D" authorityName="Laudee &amp; Malicky" authorityYear="2014" box="[656,859,1777,1804]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="species" species="suratthaniensis">
<emphasis id="B94DA3502537FFDEFD6BABC7C5BE7A7D" box="[656,673,1778,1803]" italics="true" pageId="3" pageNumber="372">R</emphasis>
. 
<emphasis id="B94DA3502537FFDEFD4BABC4C4447A7D" box="[688,859,1777,1803]" italics="true" pageId="3" pageNumber="372">suratthaniensis</emphasis>
</taxonomicName>
and the paramere apices bear several thin and long spines directed distad in 
<taxonomicName id="4C3904C12537FFDEFE57AA23C5547A59" authorityName="Schmid &amp; Denning" authorityYear="1971" box="[428,587,1814,1840]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="species" species="petersorum">
<emphasis id="B94DA3502537FFDEFE57AA23C6A27A59" box="[428,445,1814,1839]" italics="true" pageId="3" pageNumber="372">R</emphasis>
. 
<emphasis id="B94DA3502537FFDEFE30AA23C5547A59" box="[459,587,1814,1839]" italics="true" pageId="3" pageNumber="372">petersorum</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B867F422530FFD9FF6CADAFC5C57DC2" blockId="4.[151,730,154,216]" box="[151,730,154,180]" pageId="4" pageNumber="373">
<taxonomicName id="4C3904C12530FFD9FF6CADAFC6807DC2" authority="Suwannarat &amp; Malicky &amp; Morse &amp; Laudee, 2019" authorityName="Suwannarat &amp; Malicky &amp; Morse &amp; Laudee" authorityYear="2019" box="[151,415,154,180]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="species" species="limsakuli" status="sp. nov.">
<emphasis id="B94DA3502530FFD9FF6CADAFC6807DC2" box="[151,415,154,180]" italics="true" pageId="4" pageNumber="373">Rhyacophila limsakuli</emphasis>
</taxonomicName>
<emphasis id="B94DA3502530FFD9FE5DADAFC5C57DC2" bold="true" box="[422,730,154,180]" pageId="4" pageNumber="373">
<taxonomicNameLabel id="A27E1E2B2530FFD9FE5DADAFC6F77DC2" box="[422,488,154,180]" pageId="4" pageNumber="373" rank="species">sp. n.</taxonomicNameLabel>
Laudee &amp; Malicky
</emphasis>
</paragraph>
<paragraph id="8B867F422530FFD9FF6CAD8BC6377DAE" blockId="4.[151,730,154,216]" box="[151,296,190,216]" pageId="4" pageNumber="373">
<figureCitation id="130263C72530FFD9FF6CAD8BC7FE7DAE" box="[151,225,190,216]" captionStart="FIGURES 3" captionStartId="4.[151,264,1974,1999]" captionTargetBox="[151,1436,922,1921]" captionTargetId="figure@4.[151,1436,922,1950]" captionTargetPageId="4" captionText="FIGURES 3A–3D. Male genitalia of Rhyacophila limsakuli n. sp. 3A, left lateral, with left inferior appendage and phallus omitted. 3B, dorsal. 3C, ventral. 3D, phallus, left lateral. Ana = anal sclerites." figureDoi="http://doi.org/10.5281/zenodo.3772368" httpUri="https://zenodo.org/record/3772368/files/figure.png" pageId="4" pageNumber="373">Figs. 3</figureCitation>
A–3D
</paragraph>
<paragraph id="8B867F422530FFD9FF6CAC33C3847C32" blockId="4.[151,1437,262,900]" pageId="4" pageNumber="373">
Type material. 
<materialsCitation id="3B51751F2530FFD9FEC6AC33C3847C32" ID-GBIF-Occurrence="2609011275" collectingDate="2017-06-01" collectionCode="PSUNHM" collectorName="Pongsak Laudee." country="Thailand" elevation="352" latitude="12.798334" location="Ban Klang" longLatPrecision="21" longitude="99.4525" municipality="Kaeng Krachan National Park" pageId="4" pageNumber="373" specimenCount="1" stateProvince="Phetchaburi" typeStatus="holotype">
<typeStatus id="5482C1E02530FFD9FEC6AC33C6BA7C56" box="[317,421,262,288]" pageId="4" pageNumber="373" type="holotype">Holotype</typeStatus>
male (
<collectionCode id="ED28E7872530FFD9FE14AC33C5747C56" box="[495,619,262,288]" pageId="4" pageNumber="373">PSUNHM</collectionCode>
). 
<collectingCountry id="F32E3FD22530FFD9FD80AC33C40F7C56" box="[635,784,262,288]" name="Thailand" pageId="4" pageNumber="373">THAILAND</collectingCountry>
: 
<collectingRegion id="49FDB1A02530FFD9FCE2AC33C3147C56" box="[793,1035,262,288]" country="Thailand" name="Phetchaburi" pageId="4" pageNumber="373">Phetchaburi Province</collectingRegion>
, 
<collectingMunicipality id="6BE2E5382530FFD9FBEEAC33C27A7C56" box="[1045,1381,262,288]" pageId="4" pageNumber="373">Kaeng Krachan National Park</collectingMunicipality>
, 
<location id="8EE629992530FFD9FA94AC33C7C47C32" LSID="urn:lsid:plazi:treatment:0390CE542535FFDBFF6CAF10C58F7F15:8EE629992530FFD9FA94AC33C7C47C32" country="Thailand" latitude="12.798334" longLatPrecision="21" longitude="99.4525" municipality="Kaeng Krachan National Park" name="Ban Klang" pageId="4" pageNumber="373" stateProvince="Phetchaburi">Ban Klang</location>
stream, 
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, 
<geoCoordinate id="EE0D19852530FFD9FE35AC1CC54C7C32" box="[462,595,297,324]" degrees="99" direction="east" minutes="27" orientation="longitude" pageId="4" pageNumber="373" precision="15" seconds="09" value="99.4525">99°27′09″E</geoCoordinate>
, ca. 
<elevation id="001498712530FFD9FD7EAC1CC4167C32" box="[645,777,297,324]" metricMagnitude="2" metricUnit="m" metricValue="3.52" pageId="4" pageNumber="373" unit="m" value="352.0">
<quantity id="4CC1D2A72530FFD9FD7EAC1CC5D37C32" box="[645,716,297,324]" metricMagnitude="2" metricUnit="m" metricValue="3.52" pageId="4" pageNumber="373" unit="m" value="352.0">352 m</quantity>
a.s.l.
</elevation>
, 
<date id="FF8759822530FFD9FCEEAC1FC4BF7C32" box="[789,928,298,324]" pageId="4" pageNumber="373" value="2017-06-01">
<collectingDate id="EFC3A06A2530FFD9FCEEAC1FC4BF7C32" box="[789,928,298,324]" pageId="4" pageNumber="373" value="2017-06-01">1 June 2017</collectingDate>
</date>
, leg. 
<collectorName id="26CC1A942530FFD9FC20AC1FC3847C32" box="[987,1179,298,324]" pageId="4" pageNumber="373">Pongsak Laudee.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8B867F422530FFD9FF3CAC7BC4327C1E" blockId="4.[151,1437,262,900]" box="[199,813,334,360]" pageId="4" pageNumber="373">
<materialsCitation id="3B51751F2530FFD9FF3CAC7BC4357C1E" ID-GBIF-Occurrence="2609011277" box="[199,810,334,360]" collectionCode="PSUNHM" pageId="4" pageNumber="373" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="5482C1E02530FFD9FF3CAC7BC6277C1E" box="[199,312,334,360]" pageId="4" pageNumber="373" type="paratype">Paratypes</typeStatus>
: Same data as 
<typeStatus id="5482C1E02530FFD9FE27AC7BC55E7C1E" box="[476,577,334,360]" pageId="4" pageNumber="373" type="holotype">holotype</typeStatus>
, 
<specimenCount id="9D3FB4CB2530FFD9FDB7AC7BC5887C1E" box="[588,663,334,360]" pageId="4" pageNumber="373" type="male">1 male</specimenCount>
(
<collectionCode id="ED28E7872530FFD9FD5DAC7BC43D7C1E" box="[678,802,334,360]" pageId="4" pageNumber="373">PSUNHM</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B867F422530FFD9FF3CAC47C5527CC6" blockId="4.[151,1437,262,900]" pageId="4" pageNumber="373">
Etymology. The species is dedicated to Assoc. Prof. Dr. Chusak Limsakul, who was the President of Prince of Songkla University during 
<date id="FF8759822530FFD9FE3CACA3C5567CC6" box="[455,585,406,432]" pageId="4" pageNumber="373" value="2013" valueMax="2018">2013–2018</date>
.
</paragraph>
<paragraph id="8B867F422530FFD9FF3CAC8FC6767C8E" blockId="4.[151,1437,262,900]" pageId="4" pageNumber="373">Description. Length of each male forewing 7.0 mm (n = 2); specimens in alcohol with head, thorax, and forewings dark brown.</paragraph>
<paragraph id="8B867F422530FFD9FF3CAF37C3437EF2" blockId="4.[151,1437,262,900]" pageId="4" pageNumber="373">
Male genitalia (
<figureCitation id="130263C72530FFD9FE8EAF37C6C17F6A" box="[373,478,514,540]" captionStart="FIGURES 3" captionStartId="4.[151,264,1974,1999]" captionTargetBox="[151,1436,922,1921]" captionTargetId="figure@4.[151,1436,922,1950]" captionTargetPageId="4" captionText="FIGURES 3A–3D. Male genitalia of Rhyacophila limsakuli n. sp. 3A, left lateral, with left inferior appendage and phallus omitted. 3B, dorsal. 3C, ventral. 3D, phallus, left lateral. Ana = anal sclerites." figureDoi="http://doi.org/10.5281/zenodo.3772368" httpUri="https://zenodo.org/record/3772368/files/figure.png" pageId="4" pageNumber="373">Figures 3</figureCitation>
A–3D). Anterolateral margins of segment IX convex subdorsally and concave subventrally, posterolateral margins convex; apicodorsal lobe of segment IX about as long as preanal appendages, convex dorsally and apically blunt in lateral view; in dorsal view slightly concave laterally and with V-shaped incision in apicomesal 1/3. Preanal appendages as long as apicodorsal lobe of segment IX, oval in lateral view, blunt in dorsal view. Anal sclerite flat, oval in lateral view, oriented obliquely dorsad from root. Basal segment of each inferior appendage tubular, tapered anteriorly, rounded posteriorly, and with longitudinal ridge subdorsally in apical half, apical segment nearly rectangular with broad and shallow invagination apically; in ventral view basal segment of each inferior appendage cylindrical and apical segment peanut-like with subapicomesal setae. Phallus complex; phallotheca tubular with lateral tenons; in lateral view dorsal process somewhat bird-head-like with round “crown” dorsally and long “beak” apically; aedeagus as long as parameres, needle-like; parameres fused basally, thick especially basally, their apices horizontally flattened and hooked ventromesad and caudad.
</paragraph>
<caption id="DF462FCA2530FFD9FF6CAA83C4667A85" ID-DOI="http://doi.org/10.5281/zenodo.3772368" ID-Zenodo-Dep="3772368" httpUri="https://zenodo.org/record/3772368/files/figure.png" pageId="4" pageNumber="373" startId="4.[151,264,1974,1999]" targetBox="[151,1436,922,1921]" targetPageId="4">
<paragraph id="8B867F422530FFD9FF6CAA83C4667A85" blockId="4.[151,1436,1974,2035]" pageId="4" pageNumber="373">
<emphasis id="B94DA3502530FFD9FF6CAA83C6417AB9" bold="true" box="[151,350,1974,1999]" pageId="4" pageNumber="373">FIGURES 3A–3D.</emphasis>
Male genitalia of 
<taxonomicName id="4C3904C12530FFD9FDE8AA82C5EA7AB9" authority="Suwannarat &amp; Malicky &amp; Morse &amp; Laudee, 2019" authorityName="Suwannarat &amp; Malicky &amp; Morse &amp; Laudee" authorityYear="2019" box="[531,757,1975,1999]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="species" species="limsakuli" status="sp. nov.">
<emphasis id="B94DA3502530FFD9FDE8AA82C5EA7AB9" box="[531,757,1975,1999]" italics="true" pageId="4" pageNumber="373">Rhyacophila limsakuli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27E1E2B2530FFD9FD00AA8DC42F7AB9" box="[763,816,1976,1999]" pageId="4" pageNumber="373" rank="species">n. sp.</taxonomicNameLabel>
3A, left lateral, with left inferior appendage and phallus omitted. 3B, dorsal. 3C, ventral. 3D, phallus, left lateral. Ana = anal sclerites.
</paragraph>
</caption>
<paragraph id="8B867F422531FFD8FF3CADACC3867CFD" blockId="5.[151,1436,153,395]" pageId="5" pageNumber="374">
Diagnosis. This new species is a member of 
<taxonomicName id="4C3904C12531FFD8FD35ADAFC4597DC5" authorityName="Morton" authorityYear="1900" box="[718,838,154,179]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="anatina">
<emphasis id="B94DA3502531FFD8FD35ADAFC4597DC5" box="[718,838,154,179]" italics="true" pageId="5" pageNumber="374">R. anatina</emphasis>
</taxonomicName>
Group (
<bibRefCitation id="EFA802B32531FFD8FC53ADACC35A7DC5" author="Schmid, F." box="[936,1093,153,179]" pageId="5" pageNumber="374" refId="ref4292" refString="Schmid, F. (1970) Le genre Rhyacophila et la Famille des Rhyacophilidae (Trichoptera). Memoires de la Societe Entomologique du Canada, 66, 230 pp. https: // doi. org / 10.4039 / entm 10266 fv" type="book" year="1970">Schmid 1970</bibRefCitation>
). It is similar to other species from Asia and can be diagnosed by the characters mentioned by 
<bibRefCitation id="EFA802B32531FFD8FC8DAD88C4E87DAE" author="Sun, C." box="[886,1015,189,216]" pageId="5" pageNumber="374" pagination="273 - 278" refId="ref4336" refString="Sun, C. (2016) Two new species of the Rhyacophila anatina Species Group from China (Trichoptera: Rhyacophilidae). Zootaxa, 4085 (2), 273 - 278. https: // doi. org / 10.11646 / zootaxa. 4085.2.7" type="journal article" year="2016">Sun (2016)</bibRefCitation>
. 
<taxonomicName id="4C3904C12531FFD8FBF8AD88C21F7DA1" authority="Suwannarat &amp; Malicky &amp; Morse &amp; Laudee, 2019" authorityName="Suwannarat &amp; Malicky &amp; Morse &amp; Laudee" authorityYear="2019" box="[1027,1280,189,215]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="limsakuli" status="sp. nov.">
<emphasis id="B94DA3502531FFD8FBF8AD88C21F7DA1" box="[1027,1280,189,215]" italics="true" pageId="5" pageNumber="374">Rhyacophila limsakuli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27E1E2B2531FFD8FAF3AD88C25A7DA1" box="[1288,1349,189,215]" pageId="5" pageNumber="374" rank="species">sp. n.</taxonomicNameLabel>
closely resembles 
<taxonomicName id="4C3904C12531FFD8FEEBADD7C4017D8D" authority="Malicky &amp; Chantaramongkol 1993" authorityName="Malicky &amp; Chantaramongkol" authorityYear="1993" box="[272,798,225,251]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="noeibia">
<emphasis id="B94DA3502531FFD8FEEBADD7C69A7D8D" box="[272,389,225,251]" italics="true" pageId="5" pageNumber="374">R. noeibia</emphasis>
<bibRefCitation id="EFA802B32531FFD8FE76ADD4C4017D8D" author="Malicky, H. &amp; Chantaramongkol, P." box="[397,798,225,251]" pageId="5" pageNumber="374" pagination="433 - 487" refId="ref4009" refString="Malicky, H. &amp; Chantaramongkol, P. (1993) Neue Trichopteren aus Thailand. Rhyacophilidae, Hydrobiosidae, Philopotamidae, Polycentropodidae, Ecnomidae, Psychomyiidae, Arctopsychidae, Hydropsychidae (Arbeiten uber thailandische Kocherfliegen Nr. 12) (Fortsetzung). Linzer Biologische Beitrage, 25 (1), 433 - 487." type="journal article" year="1993">Malicky &amp; Chantaramongkol 1993</bibRefCitation>
</taxonomicName>
, but the two species can be distinguished in that (1) the basal segment of each inferior appendage of 
<taxonomicName id="4C3904C12531FFD8FD74AC33C4097C69" authority="Suwannarat &amp; Malicky &amp; Morse &amp; Laudee, 2019" authorityName="Suwannarat &amp; Malicky &amp; Morse &amp; Laudee" authorityYear="2019" box="[655,790,261,287]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="limsakuli" status="sp. nov.">
<emphasis id="B94DA3502531FFD8FD74AC33C5BF7C69" box="[655,672,262,287]" italics="true" pageId="5" pageNumber="374">R</emphasis>
. 
<emphasis id="B94DA3502531FFD8FD54AC30C4097C69" box="[687,790,261,287]" italics="true" pageId="5" pageNumber="374">limsakuli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27E1E2B2531FFD8FCE6AC30C4467C69" box="[797,857,261,287]" pageId="5" pageNumber="374" rank="species">sp. n.</taxonomicNameLabel>
is 4.5 times as long as the apical segment, but only 2.5 times as long in 
<taxonomicName id="4C3904C12531FFD8FE81AC1FC6F17C35" authorityName="Malicky &amp; Chantaramongkol" authorityYear="1993" box="[378,494,297,323]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="noeibia">
<emphasis id="B94DA3502531FFD8FE81AC1FC6947C35" box="[378,395,298,323]" italics="true" pageId="5" pageNumber="374">R</emphasis>
. 
<emphasis id="B94DA3502531FFD8FE61AC1CC6F17C35" box="[410,494,297,323]" italics="true" pageId="5" pageNumber="374">noeibia</emphasis>
</taxonomicName>
, (2) the apical segment of each inferior appendage of 
<taxonomicName id="4C3904C12531FFD8FBB4AC1FC3DD7C35" authorityName="Malicky &amp; Chantaramongkol" authorityYear="1993" box="[1103,1218,297,323]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="noeibia">
<emphasis id="B94DA3502531FFD8FBB4AC1FC3DD7C35" box="[1103,1218,297,323]" italics="true" pageId="5" pageNumber="374">R. noeibia</emphasis>
</taxonomicName>
is boot-shaped and projected dorsad, but that of 
<taxonomicName id="4C3904C12531FFD8FE2CAC7BC5427C11" authority="Suwannarat &amp; Malicky &amp; Morse &amp; Laudee, 2019" authorityName="Suwannarat &amp; Malicky &amp; Morse &amp; Laudee" authorityYear="2019" box="[471,605,333,359]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="limsakuli" status="sp. nov.">
<emphasis id="B94DA3502531FFD8FE2CAC7BC6F77C11" box="[471,488,334,359]" italics="true" pageId="5" pageNumber="374">R</emphasis>
. 
<emphasis id="B94DA3502531FFD8FE0DAC78C5427C11" box="[502,605,333,359]" italics="true" pageId="5" pageNumber="374">limsakuli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27E1E2B2531FFD8FD98AC78C5807C11" box="[611,671,333,359]" pageId="5" pageNumber="374" rank="species">sp. n.</taxonomicNameLabel>
is rectangular and lacks a dorsal projection, and (3) the apices of the parameres of 
<taxonomicName id="4C3904C12531FFD8FECBAC47C6BC7CFD" authorityName="Malicky &amp; Chantaramongkol" authorityYear="1993" box="[304,419,369,395]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="noeibia">
<emphasis id="B94DA3502531FFD8FECBAC47C65E7CFD" box="[304,321,370,395]" italics="true" pageId="5" pageNumber="374">R</emphasis>
. 
<emphasis id="B94DA3502531FFD8FEB4AC44C6BC7CFD" box="[335,419,369,395]" italics="true" pageId="5" pageNumber="374">noeibia</emphasis>
</taxonomicName>
are bent upward, but in 
<taxonomicName id="4C3904C12531FFD8FD4FAC47C4257CFD" authority="Suwannarat &amp; Malicky &amp; Morse &amp; Laudee, 2019" authorityName="Suwannarat &amp; Malicky &amp; Morse &amp; Laudee" authorityYear="2019" box="[692,826,369,395]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="limsakuli" status="sp. nov.">
<emphasis id="B94DA3502531FFD8FD4FAC47C5DA7CFD" box="[692,709,370,395]" italics="true" pageId="5" pageNumber="374">R</emphasis>
. 
<emphasis id="B94DA3502531FFD8FD28AC44C4257CFD" box="[723,826,369,395]" italics="true" pageId="5" pageNumber="374">limsakuli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27E1E2B2531FFD8FCBBAC44C4637CFD" box="[832,892,369,395]" pageId="5" pageNumber="374" rank="species">sp. n.</taxonomicNameLabel>
they are bent downward.
</paragraph>
<paragraph id="8B867F422531FFD8FF6CACE8C4177C81" blockId="5.[151,776,477,539]" box="[151,776,477,503]" pageId="5" pageNumber="374">
<taxonomicName id="4C3904C12531FFD8FF6CACE8C6C67C81" authority="Suwannarat &amp; Malicky &amp; Morse &amp; Laudee, 2019" authorityName="Suwannarat &amp; Malicky &amp; Morse &amp; Laudee" authorityYear="2019" box="[151,473,477,503]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="kengtungensis" status="sp. nov.">
<emphasis id="B94DA3502531FFD8FF6CACE8C6C67C81" box="[151,473,477,503]" italics="true" pageId="5" pageNumber="374">Rhyacophila kengtungensis</emphasis>
</taxonomicName>
<emphasis id="B94DA3502531FFD8FE1BACE8C4177C81" bold="true" box="[480,776,477,503]" pageId="5" pageNumber="374">
<taxonomicNameLabel id="A27E1E2B2531FFD8FE1BACE8C53D7C81" box="[480,546,477,503]" pageId="5" pageNumber="374" rank="species">sp. n.</taxonomicNameLabel>
Morse &amp; Malicky
</emphasis>
</paragraph>
<paragraph id="8B867F422531FFD8FF6CAF34C6397F6D" blockId="5.[151,776,477,539]" box="[151,294,513,539]" pageId="5" pageNumber="374">
<figureCitation id="130263C72531FFD8FF6CAF34C7FC7F6D" box="[151,227,513,539]" captionStart="FIGURES 4" captionStartId="5.[151,264,1731,1756]" captionTargetBox="[151,1436,813,1695]" captionTargetId="figure@5.[151,1436,813,1706]" captionTargetPageId="5" captionText="FIGURES 4A–4C. Male genitalia of Rhyacophila kengtungensis n. sp. 4A, left lateral, with left inferior appendage and phallus omitted. 4B, dorsal. 4C, phallus, left lateral. Pre = preanal appendages, Api IX = apicodorsal lobe of segment IX, Ana = anal sclerites." figureDoi="http://doi.org/10.5281/zenodo.3772370" httpUri="https://zenodo.org/record/3772370/files/figure.png" pageId="5" pageNumber="374">Figs. 4</figureCitation>
A–4C
</paragraph>
<paragraph id="8B867F422531FFD8FF6CAF7CC3367FF1" blockId="5.[151,1436,585,791]" pageId="5" pageNumber="374">
Type material. 
<materialsCitation id="3B51751F2531FFD8FEB3AF7CC3367FF1" ID-GBIF-Occurrence="2609011282" collectingDate="2015-01-13" collectionCode="PSUNHM" collectorName="Pongsak Laudee." country="Myanmar" elevation="977" latitude="21.443611" location="Pin Tao Waterfall" longLatPrecision="21" longitude="99.57833" municipality="Keng Tung Province" pageId="5" pageNumber="374" specimenCount="1" stateProvince="Shan" typeStatus="holotype">
<typeStatus id="5482C1E02531FFD8FEB3AF7CC6AF7F15" box="[328,432,585,611]" pageId="5" pageNumber="374" type="holotype">Holotype</typeStatus>
male (
<collectionCode id="ED28E7872531FFD8FDFFAF7CC59F7F15" box="[516,640,585,611]" pageId="5" pageNumber="374">PSUNHM</collectionCode>
). 
<collectingCountry id="F32E3FD22531FFD8FD6DAF7CC4327F15" box="[662,813,585,611]" name="Myanmar" pageId="5" pageNumber="374">MYANMAR</collectingCountry>
: 
<collectingRegion id="49FDB1A02531FFD8FCC0AF7CC4A67F15" box="[827,953,585,611]" country="Myanmar" name="Shan" pageId="5" pageNumber="374">Shan State</collectingRegion>
, 
<collectingMunicipality id="6BE2E5382531FFD8FC33AF7CC3A57F15" box="[968,1210,585,611]" pageId="5" pageNumber="374">Keng Tung Province</collectingMunicipality>
, 
<location id="8EE629992531FFD8FB31AF7CC2867F15" LSID="urn:lsid:plazi:treatment:0390CE542535FFDBFF6CAF10C58F7F15:8EE629992531FFD8FB31AF7CC2867F15" box="[1226,1433,585,611]" country="Myanmar" latitude="21.443611" longLatPrecision="21" longitude="99.57833" municipality="Keng Tung Province" name="Pin Tao Waterfall" pageId="5" pageNumber="374" stateProvince="Shan">Pin Tao Waterfall</location>
, 
<geoCoordinate id="EE0D19852531FFD8FF6CAF59C6007FF1" box="[151,287,620,647]" degrees="21" direction="north" minutes="26" orientation="latitude" pageId="5" pageNumber="374" precision="15" seconds="37" value="21.443611">21°26′37″N</geoCoordinate>
, 
<geoCoordinate id="EE0D19852531FFD8FED1AF59C6B07FF1" box="[298,431,620,647]" degrees="99" direction="east" minutes="34" orientation="longitude" pageId="5" pageNumber="374" precision="15" seconds="42" value="99.57833">99°34′42″E</geoCoordinate>
, ca. 
<elevation id="001498712531FFD8FE1AAF58C57A7FF1" box="[481,613,621,647]" metricMagnitude="2" metricUnit="m" metricValue="9.77" pageId="5" pageNumber="374" unit="m" value="977.0">
<quantity id="4CC1D2A72531FFD8FE1AAF58C5377FF1" box="[481,552,621,647]" metricMagnitude="2" metricUnit="m" metricValue="9.77" pageId="5" pageNumber="374" unit="m" value="977.0">977 m</quantity>
a.s.l.
</elevation>
, 
<date id="FF8759822531FFD8FD8AAF58C4327FF1" box="[625,813,620,647]" pageId="5" pageNumber="374" value="2015-01-13">
<collectingDate id="EFC3A06A2531FFD8FD8AAF58C4327FF1" box="[625,813,620,647]" pageId="5" pageNumber="374" value="2015-01-13">13 January 2015</collectingDate>
</date>
, leg. 
<collectorName id="26CC1A942531FFD8FC92AF58C3367FF1" box="[873,1065,621,647]" pageId="5" pageNumber="374">Pongsak Laudee.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8B867F422531FFD8FF3CAFA4C4197FDD" blockId="5.[151,1436,585,791]" box="[199,774,657,683]" pageId="5" pageNumber="374">
<materialsCitation id="3B51751F2531FFD8FF3CAFA4C41C7FDD" ID-GBIF-Occurrence="2609011279" box="[199,771,657,683]" collectionCode="CUAC" pageId="5" pageNumber="374" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="5482C1E02531FFD8FF3CAFA4C6277FDD" box="[199,312,657,683]" pageId="5" pageNumber="374" type="paratype">Paratypes</typeStatus>
: Same data as 
<typeStatus id="5482C1E02531FFD8FE27AFA4C55E7FDD" box="[476,577,657,683]" pageId="5" pageNumber="374" type="holotype">holotype</typeStatus>
, 
<specimenCount id="9D3FB4CB2531FFD8FDB7AFA4C5887FDD" box="[588,663,657,683]" pageId="5" pageNumber="374" type="male">1 male</specimenCount>
(
<collectionCode id="ED28E7872531FFD8FD5DAFA4C5E57FDD" box="[678,762,657,683]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" pageId="5" pageNumber="374" type="University or college">CUAC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B867F422531FFD8FF3CAF80C3BF7FB9" blockId="5.[151,1436,585,791]" box="[199,1184,693,719]" pageId="5" pageNumber="374">Etymology. The species epithet refers to the province where the new species was found.</paragraph>
<paragraph id="8B867F422531FFD8FF3CAFECC6727E61" blockId="5.[151,1436,585,791]" pageId="5" pageNumber="374">Description. Length of each male forewing 5.0 mm (n = 2); specimens in alcohol with head, thorax, and forewings light yellow.</paragraph>
<caption id="DF462FCA2531FFD8FF6CABF6C7EE7A52" ID-DOI="http://doi.org/10.5281/zenodo.3772370" ID-Zenodo-Dep="3772370" httpUri="https://zenodo.org/record/3772370/files/figure.png" pageId="5" pageNumber="374" startId="5.[151,264,1731,1756]" targetBox="[151,1436,813,1695]" targetPageId="5">
<paragraph id="8B867F422531FFD8FF6CABF6C7EE7A52" blockId="5.[151,1436,1731,1828]" pageId="5" pageNumber="374">
<emphasis id="B94DA3502531FFD8FF6CABF6C6407BAA" bold="true" box="[151,351,1731,1756]" pageId="5" pageNumber="374">FIGURES 4A–4C.</emphasis>
Male genitalia of 
<taxonomicName id="4C3904C12531FFD8FDEDABF6C4337BAD" authority="Suwannarat &amp; Malicky &amp; Morse &amp; Laudee, 2019" authorityName="Suwannarat &amp; Malicky &amp; Morse &amp; Laudee" authorityYear="2019" box="[534,812,1731,1755]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="kengtungensis" status="sp. nov.">
<emphasis id="B94DA3502531FFD8FDEDABF6C4337BAD" box="[534,812,1731,1755]" italics="true" pageId="5" pageNumber="374">Rhyacophila kengtungensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27E1E2B2531FFD8FCCAABF1C4787BAD" box="[817,871,1732,1755]" pageId="5" pageNumber="374" rank="species">n. sp.</taxonomicNameLabel>
4A, left lateral, with left inferior appendage and phallus omitted. 4B, dorsal. 4C, phallus, left lateral. Pre = preanal appendages, Api IX = apicodorsal lobe of segment IX, Ana = anal sclerites.
</paragraph>
</caption>
<paragraph id="8B867F422531FFDBFF3CAA67C4587C69" blockId="5.[151,1437,1874,2044]" lastBlockId="6.[151,1437,152,611]" lastPageId="6" lastPageNumber="375" pageId="5" pageNumber="374">
Male genitalia (
<figureCitation id="130263C72531FFD8FE84AA67C6F17A1A" box="[383,494,1874,1900]" captionStart="FIGURES 4" captionStartId="5.[151,264,1731,1756]" captionTargetBox="[151,1436,813,1695]" captionTargetId="figure@5.[151,1436,813,1706]" captionTargetPageId="5" captionText="FIGURES 4A–4C. Male genitalia of Rhyacophila kengtungensis n. sp. 4A, left lateral, with left inferior appendage and phallus omitted. 4B, dorsal. 4C, phallus, left lateral. Pre = preanal appendages, Api IX = apicodorsal lobe of segment IX, Ana = anal sclerites." figureDoi="http://doi.org/10.5281/zenodo.3772370" httpUri="https://zenodo.org/record/3772370/files/figure.png" pageId="5" pageNumber="374">Figures 4</figureCitation>
A–4C). Segment IX anterolateral margins nearly straight, posterolateral margins sinuous below apicodorsal lobe, apicodorsal lobe short, half as long as inferior appendages, and nearly as wide as annular portion of segment, tubular. Preanal appendages strongly sclerotized, extending beyond apicodorsal lobe of segment IX, their mesal surfaces covered with short dark brown setae, oval in lateral view, subtriangular in dorsal view. Anal sclerites small, beneath preanal appendages. Inferior appendages long, cylindrical, each with basal seg- ment twice as long as apical segment; apical segment subtriangular, its dorsal margin forming 50° angle with dorsal margin of basal segment, blunt apically, mesal surface covered with numerous setae. Phallus (
<figureCitation id="130263C72532FFDBFB51AD88C3E07DA1" box="[1194,1279,189,215]" captionStart="FIGURES 4" captionStartId="5.[151,264,1731,1756]" captionTargetBox="[151,1436,813,1695]" captionTargetId="figure@5.[151,1436,813,1706]" captionTargetPageId="5" captionText="FIGURES 4A–4C. Male genitalia of Rhyacophila kengtungensis n. sp. 4A, left lateral, with left inferior appendage and phallus omitted. 4B, dorsal. 4C, phallus, left lateral. Pre = preanal appendages, Api IX = apicodorsal lobe of segment IX, Ana = anal sclerites." figureDoi="http://doi.org/10.5281/zenodo.3772370" httpUri="https://zenodo.org/record/3772370/files/figure.png" pageId="6" pageNumber="375">Fig. 4C</figureCitation>
) small, dorsal appendage of phallic apparatus cylindrical, aedeagus short, apically acute; pair of ventral appendages of phallic apparatus very large, curved upward, covered with strong setae.
</paragraph>
<paragraph id="8B867F422532FFDBFF3CAC1CC58F7F15" blockId="6.[151,1437,152,611]" pageId="6" pageNumber="375">
Diagnosis. 
<taxonomicName id="4C3904C12532FFDBFEBEAC1CC5667C35" authority="Suwannarat &amp; Malicky &amp; Morse &amp; Laudee, 2019" authorityName="Suwannarat &amp; Malicky &amp; Morse &amp; Laudee" authorityYear="2019" box="[325,633,297,323]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="375" phylum="Arthropoda" rank="species" species="kengtungensis" status="sp. nov.">
<emphasis id="B94DA3502532FFDBFEBEAC1CC5667C35" box="[325,633,297,323]" italics="true" pageId="6" pageNumber="375">Rhyacophila kengtungensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27E1E2B2532FFDBFD84AC1CC5A47C35" box="[639,699,297,323]" pageId="6" pageNumber="375" rank="species">sp. n.</taxonomicNameLabel>
is a member of the 
<taxonomicName id="4C3904C12532FFDBFC62AC1FC3117C35" authorityName="Groups. Sun" authorityYear="2016" box="[921,1038,297,323]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="375" phylum="Arthropoda" rank="species" species="yishepa">
<emphasis id="B94DA3502532FFDBFC62AC1FC4B57C35" box="[921,938,298,323]" italics="true" pageId="6" pageNumber="375">R</emphasis>
. 
<emphasis id="B94DA3502532FFDBFC43AC1CC3117C35" box="[952,1038,297,323]" italics="true" pageId="6" pageNumber="375">yishepa</emphasis>
</taxonomicName>
Group, in which males each have a short and broad apicodorsal lobe of segment IX, a reduced anal sclerite, and no parameres (
<bibRefCitation id="EFA802B32532FFDBFB71AC78C23B7C11" author="Schmid, F." box="[1162,1316,333,359]" pageId="6" pageNumber="375" refId="ref4292" refString="Schmid, F. (1970) Le genre Rhyacophila et la Famille des Rhyacophilidae (Trichoptera). Memoires de la Societe Entomologique du Canada, 66, 230 pp. https: // doi. org / 10.4039 / entm 10266 fv" type="book" year="1970">Schmid 1970</bibRefCitation>
). This species is close to 
<taxonomicName id="4C3904C12532FFDBFEBBAC47C5B67CFD" authority="Malicky &amp; Sun 2002" authorityName="Malicky &amp; Sun" authorityYear="2002" box="[320,681,369,395]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="375" phylum="Arthropoda" rank="species" species="ligulata">
<emphasis id="B94DA3502532FFDBFEBBAC47C64E7CFD" box="[320,337,370,395]" italics="true" pageId="6" pageNumber="375">R</emphasis>
. 
<emphasis id="B94DA3502532FFDBFEA4AC44C6A97CFD" box="[351,438,369,395]" italics="true" pageId="6" pageNumber="375">ligulata</emphasis>
<bibRefCitation id="EFA802B32532FFDBFE45AC44C5B67CFD" author="Malicky, H. &amp; Sun, C." box="[446,681,369,395]" pageId="6" pageNumber="375" pagination="541 - 561" refId="ref4067" refString="Malicky, H. &amp; Sun, C. (2002) 25 new species of Rhyacophilidae (Trichoptera) from China. Linzer Biologische Beitrage, 34 (1), 541 - 561." type="journal article" year="2002">Malicky &amp; Sun 2002</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C3904C12532FFDBFD24AC47C3DC7CFD" authority="Malicky &amp; Chantaramongkol 2009" authorityName="Malicky &amp; Chantaramongkol" authorityYear="2009" box="[735,1219,369,395]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="375" phylum="Arthropoda" rank="species" species="suah">
<emphasis id="B94DA3502532FFDBFD24AC47C5EF7CFD" box="[735,752,370,395]" italics="true" pageId="6" pageNumber="375">R</emphasis>
. 
<emphasis id="B94DA3502532FFDBFD05AC44C42C7CFD" box="[766,819,369,395]" italics="true" pageId="6" pageNumber="375">suah</emphasis>
Malicky &amp; Chantaramongkol 2009
</taxonomicName>
(in 
<bibRefCitation id="EFA802B32532FFDBFB0BAC44C28E7CFD" author="Malicky, H." box="[1264,1425,369,395]" pageId="6" pageNumber="375" pagination="11 - 58" refId="ref3958" refString="Malicky, H. (2009) Beitrage zur Kenntnis asiatischer Trichopteren. Braueria, 36, 11 - 58." type="journal article" year="2009">Malicky 2009</bibRefCitation>
). However, 
<taxonomicName id="4C3904C12532FFDBFEF7ACA3C6D47CD9" authority="Suwannarat &amp; Malicky &amp; Morse &amp; Laudee, 2019" authorityName="Suwannarat &amp; Malicky &amp; Morse &amp; Laudee" authorityYear="2019" box="[268,459,405,431]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="375" phylum="Arthropoda" rank="species" species="kengtungensis" status="sp. nov.">
<emphasis id="B94DA3502532FFDBFEF7ACA3C6027CD9" box="[268,285,406,431]" italics="true" pageId="6" pageNumber="375">R</emphasis>
. 
<emphasis id="B94DA3502532FFDBFED7ACA0C6D47CD9" box="[300,459,405,431]" italics="true" pageId="6" pageNumber="375">kengtungensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27E1E2B2532FFDBFE28ACA0C50F7CD9" box="[467,528,405,431]" pageId="6" pageNumber="375" rank="species">sp. n.</taxonomicNameLabel>
differs from those species by (1) the subtriangular shape of the preanal appendages in dorsal view in 
<taxonomicName id="4C3904C12532FFDBFE6AAC8FC5507CA5" authority="Suwannarat &amp; Malicky &amp; Morse &amp; Laudee, 2019" authorityName="Suwannarat &amp; Malicky &amp; Morse &amp; Laudee" authorityYear="2019" box="[401,591,441,467]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="375" phylum="Arthropoda" rank="species" species="kengtungensis" status="sp. nov.">
<emphasis id="B94DA3502532FFDBFE6AAC8FC5507CA5" box="[401,591,441,467]" italics="true" pageId="6" pageNumber="375">R. kengtungensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27E1E2B2532FFDBFDAEAC8CC58D7CA5" box="[597,658,441,467]" pageId="6" pageNumber="375" rank="species">sp. n.</taxonomicNameLabel>
, but hooked in 
<taxonomicName id="4C3904C12532FFDBFCC5AC8FC4AB7CA5" authorityName="Malicky &amp; Sun" authorityYear="2002" box="[830,948,441,467]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="375" phylum="Arthropoda" rank="species" species="ligulata">
<emphasis id="B94DA3502532FFDBFCC5AC8FC4507CA5" box="[830,847,442,467]" italics="true" pageId="6" pageNumber="375">R</emphasis>
. 
<emphasis id="B94DA3502532FFDBFCA6AC8CC4AB7CA5" box="[861,948,441,467]" italics="true" pageId="6" pageNumber="375">ligulata</emphasis>
</taxonomicName>
and bilobed in 
<taxonomicName id="4C3904C12532FFDBFB9AAC8FC3AA7CA5" authorityName="Malicky &amp; Chantaramongkol" authorityYear="2009" box="[1121,1205,441,467]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="375" phylum="Arthropoda" rank="species" species="suah">
<emphasis id="B94DA3502532FFDBFB9AAC8FC36D7CA5" box="[1121,1138,442,467]" italics="true" pageId="6" pageNumber="375">R</emphasis>
. 
<emphasis id="B94DA3502532FFDBFB7BAC8CC3AA7CA5" box="[1152,1205,441,467]" italics="true" pageId="6" pageNumber="375">suah</emphasis>
</taxonomicName>
; (2) the right-angled triangular shape of the apical segment of each inferior appendage in 
<taxonomicName id="4C3904C12532FFDBFC5AACEBC37F7C81" authority="Suwannarat &amp; Malicky &amp; Morse &amp; Laudee, 2019" authorityName="Suwannarat &amp; Malicky &amp; Morse &amp; Laudee" authorityYear="2019" box="[929,1120,477,503]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="375" phylum="Arthropoda" rank="species" species="kengtungensis" status="sp. nov.">
<emphasis id="B94DA3502532FFDBFC5AACEBC37F7C81" box="[929,1120,477,503]" italics="true" pageId="6" pageNumber="375">R. kengtungensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27E1E2B2532FFDBFB93ACE8C3B97C81" box="[1128,1190,477,503]" pageId="6" pageNumber="375" rank="species">sp. n.</taxonomicNameLabel>
, contrasting with the oval apical segment of each inferior appendage in 
<taxonomicName id="4C3904C12532FFDBFD37AF37C45D7F6D" authorityName="Malicky &amp; Sun" authorityYear="2002" box="[716,834,513,539]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="375" phylum="Arthropoda" rank="species" species="ligulata">
<emphasis id="B94DA3502532FFDBFD37AF37C5C27F6D" box="[716,733,514,539]" italics="true" pageId="6" pageNumber="375">R</emphasis>
. 
<emphasis id="B94DA3502532FFDBFD10AF34C45D7F6D" box="[747,834,513,539]" italics="true" pageId="6" pageNumber="375">ligulata</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3904C12532FFDBFC82AF37C4D27F6D" authorityName="Malicky &amp; Chantaramongkol" authorityYear="2009" box="[889,973,513,539]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="375" phylum="Arthropoda" rank="species" species="suah">
<emphasis id="B94DA3502532FFDBFC82AF37C4957F6D" box="[889,906,514,539]" italics="true" pageId="6" pageNumber="375">R</emphasis>
. 
<emphasis id="B94DA3502532FFDBFC63AF34C4D27F6D" box="[920,973,513,539]" italics="true" pageId="6" pageNumber="375">suah</emphasis>
</taxonomicName>
; and (3) the ventral plate of the aedeagus of 
<taxonomicName id="4C3904C12532FFDBFF4EAF13C67B7F49" box="[181,356,549,575]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="375" phylum="Arthropoda" rank="species" species="kentungensis">
<emphasis id="B94DA3502532FFDBFF4EAF13C7D97F49" box="[181,198,550,575]" italics="true" pageId="6" pageNumber="375">R</emphasis>
. 
<emphasis id="B94DA3502532FFDBFF28AF10C67B7F49" box="[211,356,549,575]" italics="true" pageId="6" pageNumber="375">kentungensis</emphasis>
</taxonomicName>
is obscured by the setose dorsal appendages of the phallic apparatus, but is conspicuous, slender, tubular, and acute in 
<taxonomicName id="4C3904C12532FFDBFE79AF7FC6E77F15" authorityName="Malicky &amp; Sun" authorityYear="2002" box="[386,504,585,611]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="375" phylum="Arthropoda" rank="species" species="ligulata">
<emphasis id="B94DA3502532FFDBFE79AF7FC68C7F15" box="[386,403,586,611]" italics="true" pageId="6" pageNumber="375">R</emphasis>
. 
<emphasis id="B94DA3502532FFDBFE5AAF7CC6E77F15" box="[417,504,585,611]" italics="true" pageId="6" pageNumber="375">ligulata</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3904C12532FFDBFDD5AF7FC59D7F15" authorityName="Malicky &amp; Chantaramongkol" authorityYear="2009" box="[558,642,585,611]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="375" phylum="Arthropoda" rank="species" species="suah">
<emphasis id="B94DA3502532FFDBFDD5AF7FC5207F15" box="[558,575,586,611]" italics="true" pageId="6" pageNumber="375">R</emphasis>
. 
<emphasis id="B94DA3502532FFDBFDB6AF7CC59D7F15" box="[589,642,585,611]" italics="true" pageId="6" pageNumber="375">suah</emphasis>
</taxonomicName>
..
</paragraph>
</subSubSection>
</treatment>
</document>