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<document id="D143E0331143E277A640FE2A598379D6" ID-CLB-Dataset="24345" ID-DOI="10.11646/zootaxa.4722.3.4" ID-GBIF-Dataset="03d67507-fe7a-44e3-bf0f-3b842d7d2e72" ID-ISSN="1175-5326" ID-Zenodo-Dep="3607103" ID-ZooBank="B7AEB4DA-FB6B-4B7A-8539-075E39AB7050" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1578986109188" checkinUser="plazi" docAuthor="Saxton, Natalie A., Powell, Gareth S., Martin, Gavin J. &amp; Bybee, Seth M." docDate="2020" docId="03FF4B3BFF8DFFEAFF02FF07FCC58CFE" docLanguage="en" docName="zootaxa.4722.3.4.pdf" docOrigin="Zootaxa 4722 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Atyphella marigenous , Saxton &amp; Bybee 2020, sp. nov." docType="treatment" docVersion="7" lastPageNumber="275" masterDocId="FFC63343FF8EFFEFFF95FF9EFFD48E79" masterDocTitle="Two new species of coastal Atyphella Olliff (Lampyridae: Luciolinae)" masterLastPageNumber="276" masterPageNumber="270" pageNumber="273" updateTime="1698777403341" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="0CEE285D5DAC740F77E01C25A1FC1EC0" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="64773F1C1D779708AD082683E99E0C6D">
<mods:title id="C37EEA3BFF4D414318EA1BB9358CBD5A">Two new species of coastal Atyphella Olliff (Lampyridae: Luciolinae)</mods:title>
</mods:titleInfo>
<mods:name id="7DD43E4BAACE9082361E19D25E884A1F" type="personal">
<mods:role id="72AD5B9BAAC8F2ED58521F3B1B884528">
<mods:roleTerm id="37BF1A989E711F56FB1AF9B67177AD6A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="83DDB01883C8153704F1230E12F68C06">Saxton, Natalie A.</mods:namePart>
</mods:name>
<mods:name id="A9EE12634888B674C30DCFA00A0F8F2D" type="personal">
<mods:role id="765C70165261EF5AB79E3EC87D413AB7">
<mods:roleTerm id="2A505A265A4DCB64118B71BD805A153F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A15A5A96CA4047C6C9DEC97FA8EC8EB4">Powell, Gareth S.</mods:namePart>
</mods:name>
<mods:name id="E75791C5C7922EC0F04DB0DA8D763BB2" type="personal">
<mods:role id="A5BA9C4537CAC935A5B463956D74671B">
<mods:roleTerm id="D88A73AA58BFBE18A061185CFFF4E9B3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2CB034D7F831099BDB7B4F07CD6FADC5">Martin, Gavin J.</mods:namePart>
</mods:name>
<mods:name id="4273B698F24190373FF7C92F88EF430F" type="personal">
<mods:role id="D035A3A4617E73FDAAD8BF4A93D96D7B">
<mods:roleTerm id="4AD95B4FD669FCC8B1B738BF36D0A75F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4A533487FF909630EBC2099C21942653">Bybee, Seth M.</mods:namePart>
</mods:name>
<mods:typeOfResource id="5A5ADEEB66C9206A52B8DCF172CD0FD3">text</mods:typeOfResource>
<mods:relatedItem id="483F86064B0B93728498DEE8E0223133" type="host">
<mods:titleInfo id="484C0B1029FCBE1B4BFB0E289619BCA9">
<mods:title id="7E4A21B63F3DBCF3DA53F8220E8D4370">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="8DFA5CCD1AE89F118C8CA8436BC8633F">
<mods:date id="D9A93FEB73F99138E7C6DE33DD85639B">2020</mods:date>
<mods:detail id="FE313B51866DD1EEB92633A734694DCD" type="pubDate">
<mods:number id="ED080FA6A064A366E8F2B65C0FB5E069">2020-01-14</mods:number>
</mods:detail>
<mods:detail id="BE27775BFC906A649E789F4B004D1DC6" type="volume">
<mods:number id="F4FD1BCDABB62B77C7743C2993451AC6">4722</mods:number>
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<mods:number id="1146A69EB0EF4043B4FD51DEC9993408">3</mods:number>
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<mods:start id="8FDC14739A81FC81A0EBD071723AF146">270</mods:start>
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<mods:classification id="66DFBEBC3AEC0F9A75E980006E0064D5">journal article</mods:classification>
<mods:identifier id="C0B985AC6839C1E40D2E81564ADEF802" type="CLB-Dataset">24345</mods:identifier>
<mods:identifier id="3A51C84707517B704CAA61CFCF5F2EEA" type="DOI">10.11646/zootaxa.4722.3.4</mods:identifier>
<mods:identifier id="40D7AAF8CF7A5690AEB475AF1B7D8A0E" type="GBIF-Dataset">03d67507-fe7a-44e3-bf0f-3b842d7d2e72</mods:identifier>
<mods:identifier id="8AA7CBC7D8F77FF800B9B4CDECFA665A" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="D74A089AE3174892AAB3BC7BF5D83F33" type="Zenodo-Dep">3607103</mods:identifier>
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<treatment id="03FF4B3BFF8DFFEAFF02FF07FCC58CFE" ID-DOI="http://doi.org/10.5281/zenodo.5920489" ID-GBIF-Taxon="161678580" ID-Zenodo-Dep="5920489" LSID="urn:lsid:plazi:treatment:03FF4B3BFF8DFFEAFF02FF07FCC58CFE" httpUri="http://treatment.plazi.org/id/03FF4B3BFF8DFFEAFF02FF07FCC58CFE" lastPageId="5" lastPageNumber="275" pageId="3" pageNumber="273">
<subSubSection id="C34CA9A6FF8DFFECFF02FF07FD338ECD" box="[151,743,153,180]" pageId="3" pageNumber="273" type="nomenclature">
<paragraph id="8BE9FA2DFF8DFFECFF02FF07FD338ECD" blockId="3.[151,743,153,216]" box="[151,743,153,180]" pageId="3" pageNumber="273">
<heading id="D0A14D41FF8DFFECFF02FF07FD338ECD" bold="true" box="[151,743,153,180]" fontSize="11" level="1" pageId="3" pageNumber="273" reason="1">
<emphasis id="B922263FFF8DFFECFF02FF07FD338ECD" bold="true" box="[151,743,153,180]" pageId="3" pageNumber="273">
<taxonomicName id="4C5681AEFF8DFFECFF02FF07FDA88ECD" ID-CoL="8NC37" authority="Saxton &amp; Bybee" authorityName=", Saxton &amp; Bybee" authorityYear="2020" box="[151,636,153,180]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="273" phylum="Arthropoda" rank="species" species="marigenous" status="sp. nov.">
<emphasis id="B922263FFF8DFFECFF02FF07FE758ECA" bold="true" box="[151,417,153,180]" italics="true" pageId="3" pageNumber="273">Atyphella marigenous</emphasis>
, Saxton &amp; Bybee
</taxonomicName>
,
<taxonomicNameLabel id="A2119B44FF8DFFECFD1CFF04FD338ECD" box="[649,743,154,180]" pageId="3" pageNumber="273" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34CA9A6FF8DFFECFF02FF23FEBF8EAE" box="[151,363,189,216]" pageId="3" pageNumber="273" type="description">
<paragraph id="8BE9FA2DFF8DFFECFF02FF23FEBF8EAE" blockId="3.[151,743,153,216]" box="[151,363,189,216]" pageId="3" pageNumber="273">
(
<figureCitation id="136DE6A8FF8DFFECFF0AFF23FEE18EA1" box="[159,309,189,216]" captionStart="FIGURE 1" captionStartId="3.[151,250,1906,1931]" captionTargetBox="[151,1436,410,1886]" captionTargetId="figure@3.[151,1436,410,1886]" captionTargetPageId="3" captionText="FIGURE 1. A–I) Atyphella spp. holotypes. A–C) dorsal pronotum: A) A. aphrogeneia, B) A. maritimus, C) A. marigenous; D–F) scutellum: D) A. aphrogeneia, E) A. maritimus, F) A. marigenous; G–I) left elytron, notice extent of costae G) A. aphrogeneia, H) A. maritimus, I) A. marigenous. Atyphella aphrogeneia holotype images are the property of © CSIRO." figureDoi="http://doi.org/10.5281/zenodo.3607105" httpUri="https://zenodo.org/record/3607105/files/figure.png" pageId="3" pageNumber="273">Figs. 1C, F, I</figureCitation>
;
<figureCitation id="136DE6A8FF8DFFECFED4FF23FEB68EAE" box="[321,354,189,215]" captionStart="FIGURE 2" captionStartId="4.[151,250,1230,1255]" captionTargetBox="[151,1436,409,1210]" captionTargetId="figure@4.[151,1436,409,1210]" captionTargetPageId="4" captionText="FIGURE 2. A–C) Atyphella spp. dorsal view of aedeagus. A) A. aphrogeneia holotype, B) A. maritimus paratype, C) A. marigenous holotype. Atyphella aphrogeneia holotype images are the property of © CSIRO." figureDoi="http://doi.org/10.5281/zenodo.3607107" httpUri="https://zenodo.org/record/3607107/files/figure.png" pageId="3" pageNumber="273">2C</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34CA9A6FF8DFFECFF02FE98FC0A8FF2" pageId="3" pageNumber="273" type="materials_examined">
<paragraph id="8BE9FA2DFF8DFFECFF02FE98FC0A8FF2" blockId="3.[151,1436,261,396]" pageId="3" pageNumber="273">
<emphasis id="B922263FFF8DFFECFF02FE98FE9C8F59" bold="true" box="[151,328,261,288]" pageId="3" pageNumber="273">Type material.</emphasis>
<materialsCitation id="3B3EF070FF8DFFECFEC6FE98FD688F3A" ID-GBIF-Occurrence="2549146465" collectingDate="2018-06-12" collectionCode="BYU" collectorName="Efate Is. &amp; S. Bybee &amp; G. Powell" country="Vanuatu" latitude="-17.6841" location="Devil's Point Rd." longLatPrecision="7" longitude="168.2536" pageId="3" pageNumber="273" specimenCount="1" typeStatus="holotype">
<emphasis id="B922263FFF8DFFECFEC6FE98FDD08F59" bold="true" box="[339,516,262,288]" pageId="3" pageNumber="273">
<typeStatus id="54ED448FFF8DFFECFEC6FE98FE148F59" box="[339,448,262,288]" pageId="3" pageNumber="273" type="holotype">Holotype</typeStatus>
male
</emphasis>
(
<figureCitation id="136DE6A8FF8DFFECFD83FE9BFDA78F66" box="[534,627,261,287]" captionStart="FIGURE 1" captionStartId="3.[151,250,1906,1931]" captionTargetBox="[151,1436,410,1886]" captionTargetId="figure@3.[151,1436,410,1886]" captionTargetPageId="3" captionText="FIGURE 1. A–I) Atyphella spp. holotypes. A–C) dorsal pronotum: A) A. aphrogeneia, B) A. maritimus, C) A. marigenous; D–F) scutellum: D) A. aphrogeneia, E) A. maritimus, F) A. marigenous; G–I) left elytron, notice extent of costae G) A. aphrogeneia, H) A. maritimus, I) A. marigenous. Atyphella aphrogeneia holotype images are the property of © CSIRO." figureDoi="http://doi.org/10.5281/zenodo.3607105" httpUri="https://zenodo.org/record/3607105/files/figure.png" pageId="3" pageNumber="273">Fig. 1D</figureCitation>
): //
<collectingCountry id="F341BABDFF8DFFECFD30FE98FCD18F59" box="[677,773,262,288]" name="Vanuatu" pageId="3" pageNumber="273">Vanuatu</collectingCountry>
:
<collectorName id="26A39FFBFF8DFFECFC87FE9BFCA78F59" box="[786,883,261,288]" pageId="3" pageNumber="273">Efate Is.</collectorName>
/ 1999
<location id="8E89ACF6FF8DFFECFC47FE9BFB498F66" LSID="urn:lsid:plazi:treatment:03FF4B3BFF8DFFEAFF02FF07FCC58CFE:8E89ACF6FF8DFFECFC47FE9BFB498F66" box="[978,1181,261,287]" country="Vanuatu" latitude="-17.6841" longLatPrecision="7" longitude="168.2536" name="Devil's Point Rd." pageId="3" pageNumber="273">Devil’s Point Rd.</location>
,
<geoCoordinate id="EE629CEAFF8DFFECFB3EFE9BFAC48F66" box="[1195,1296,261,287]" degrees="17.6841" direction="south" orientation="latitude" pageId="3" pageNumber="273" precision="5" value="-17.6841">-17.6841</geoCoordinate>
/
<geoCoordinate id="EE629CEAFF8DFFECFAB9FE9BFA4C8F59" box="[1324,1432,261,288]" degrees="168.2536" direction="east" orientation="longitude" pageId="3" pageNumber="273" precision="5" value="168.2536">168.2536</geoCoordinate>
,
<date id="FFE8DCEDFF8DFFECFF02FEB7FEDB8F3A" box="[151,271,297,323]" pageId="3" pageNumber="273" value="2018-06-12">
<collectingDate id="EFAC2505FF8DFFECFF02FEB7FEDB8F3A" box="[151,271,297,323]" pageId="3" pageNumber="273" value="2018-06-12">12.vi.2018</collectingDate>
</date>
/ coll.
<collectorName id="26A39FFBFF8DFFECFECFFEB7FE6B8F3A" box="[346,447,297,323]" pageId="3" pageNumber="273">S. Bybee</collectorName>
&amp;
<collectorName id="26A39FFBFF8DFFECFE75FEB7FD848F3A" box="[480,592,297,323]" pageId="3" pageNumber="273">G. Powell</collectorName>
// (
<collectionCode id="ED4762E8FF8DFFECFDE6FEB7FD608F3A" box="[627,692,297,323]" country="USA" httpUri="http://grbio.org/cool/0am8-rdhc" name="Monte L. Bean Life Science Museum" pageId="3" pageNumber="273" type="University or college">BYU</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B3EF070FF8DFFECFD53FEB4FB258F3A" ID-GBIF-Occurrence="2549146467" box="[710,1265,297,324]" collectionCode="ANIC, BYU, USNM" country="Vanuatu" pageId="3" pageNumber="273" specimenCount="7" specimenCount-male="7" typeStatus="paratype">
<emphasis id="B922263FFF8DFFECFD53FEB4FCEA8F3D" bold="true" box="[710,830,298,324]" pageId="3" pageNumber="273">
<typeStatus id="54ED448FFF8DFFECFD53FEB4FCEA8F3D" box="[710,830,298,324]" pageId="3" pageNumber="273" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<specimenCount id="9D5031A4FF8DFFECFCD8FEB7FC7C8F3D" box="[845,936,297,324]" pageId="3" pageNumber="273" type="male">7 males</specimenCount>
): (2
<collectionCode id="ED4762E8FF8DFFECFC43FEB7FBC98F3A" box="[982,1053,297,323]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="3" pageNumber="273">ANIC</collectionCode>
, 2
<collectionCode id="ED4762E8FF8DFFECFBAEFEB4FBAD8F3D" box="[1083,1145,298,324]" country="USA" httpUri="http://grbio.org/cool/0am8-rdhc" name="Monte L. Bean Life Science Museum" pageId="3" pageNumber="273" type="University or college">BYU</collectionCode>
, 1
<collectionCode id="ED4762E8FF8DFFECFB0DFEB7FB3F8F3A" box="[1176,1259,297,323]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="273" type="Museum">USNM</collectionCode>
)
</materialsCitation>
<materialsCitation id="3B3EF070FF8DFFECFB62FEB4FB278F11" ID-GBIF-Occurrence="2549146464" collectingDate="2018-06-12" collectionCode="NHM" collectorName="Efate Is. &amp; S. Bybee &amp; G. Powell" country="Vanuatu" latitude="-17.6841" location="Devil's Point Rd." longLatPrecision="7" longitude="168.2536" pageId="3" pageNumber="273" specimenCount="1" typeStatus="paratype">
<collectingCountry id="F341BABDFF8DFFECFB62FEB4FA838F3D" box="[1271,1367,298,324]" name="Vanuatu" pageId="3" pageNumber="273">Vanuatu</collectingCountry>
:
<collectorName id="26A39FFBFF8DFFECFAF4FEB7FF668F11" pageId="3" pageNumber="273">Efate Is.</collectorName>
/ 1999
<location id="8E89ACF6FF8DFFECFE96FED3FE118F1E" LSID="urn:lsid:plazi:treatment:03FF4B3BFF8DFFEAFF02FF07FCC58CFE:8E89ACF6FF8DFFECFE96FED3FE118F1E" box="[259,453,333,359]" country="Vanuatu" latitude="-17.6841" longLatPrecision="7" longitude="168.2536" name="Devil's Point Rd." pageId="3" pageNumber="273">Devil’s Point Rd.</location>
,
<geoCoordinate id="EE629CEAFF8DFFECFE5AFED3FDE78F1E" box="[463,563,333,359]" degrees="17.6841" direction="south" orientation="latitude" pageId="3" pageNumber="273" precision="5" value="-17.6841">-17.6841</geoCoordinate>
/
<geoCoordinate id="EE629CEAFF8DFFECFDD3FED3FD668F11" box="[582,690,333,360]" degrees="168.2536" direction="east" orientation="longitude" pageId="3" pageNumber="273" precision="5" value="168.2536">168.2536</geoCoordinate>
,
<date id="FFE8DCEDFF8DFFECFD29FED3FCE08F1E" box="[700,820,333,359]" pageId="3" pageNumber="273" value="2018-06-12">
<collectingDate id="EFAC2505FF8DFFECFD29FED3FCE08F1E" box="[700,820,333,359]" pageId="3" pageNumber="273" value="2018-06-12">12.vi.2018</collectingDate>
</date>
/ coll.
<collectorName id="26A39FFBFF8DFFECFCE8FED3FC358F1E" box="[893,993,333,359]" pageId="3" pageNumber="273">S. Bybee</collectorName>
&amp;
<collectorName id="26A39FFBFF8DFFECFB97FED3FBA58F1E" box="[1026,1137,333,359]" pageId="3" pageNumber="273">G. Powell</collectorName>
// (2
<collectionCode id="ED4762E8FF8DFFECFB3DFED0FB3F8F11" box="[1192,1259,334,360]" pageId="3" pageNumber="273">NHM</collectionCode>
)
</materialsCitation>
<materialsCitation id="3B3EF070FF8DFFECFB6DFED0FC0A8FF2" ID-GBIF-Occurrence="2549146462" collectingDate="2018-06-13" collectorName="Efate Is. &amp; S. Bybee &amp; G. Powell" country="Vanuatu" latitude="-17.5714" location="Epule" longLatPrecision="7" longitude="168.4702" pageId="3" pageNumber="273" specimenCount="1" typeStatus="paratype">
<collectingCountry id="F341BABDFF8DFFECFB6DFED0FA8C8F11" box="[1272,1368,334,360]" name="Vanuatu" pageId="3" pageNumber="273">Vanuatu</collectingCountry>
:
<collectorName id="26A39FFBFF8DFFECFAF4FED3FF668FF5" pageId="3" pageNumber="273">Efate Is.</collectorName>
/
<location id="8E89ACF6FF8DFFECFF5DFEEFFED88FF2" LSID="urn:lsid:plazi:treatment:03FF4B3BFF8DFFEAFF02FF07FCC58CFE:8E89ACF6FF8DFFECFF5DFEEFFED88FF2" box="[200,268,369,395]" country="Vanuatu" latitude="-17.5714" longLatPrecision="7" longitude="168.4702" name="Epule" pageId="3" pageNumber="273">Epule</location>
,
<geoCoordinate id="EE629CEAFF8DFFECFE8DFEEFFEA88FF5" box="[280,380,369,396]" degrees="17.5714" direction="south" orientation="latitude" pageId="3" pageNumber="273" precision="5" value="-17.5714">-17.5714</geoCoordinate>
/
<geoCoordinate id="EE629CEAFF8DFFECFE07FEEFFE2A8FF2" box="[402,510,369,395]" degrees="168.4702" direction="east" orientation="longitude" pageId="3" pageNumber="273" precision="5" value="168.4702">168.4702</geoCoordinate>
,
<date id="FFE8DCEDFF8DFFECFD9CFEEFFD548FF2" box="[521,640,369,395]" pageId="3" pageNumber="273" value="2018-06-13">
<collectingDate id="EFAC2505FF8DFFECFD9CFEEFFD548FF2" box="[521,640,369,395]" pageId="3" pageNumber="273" value="2018-06-13">13.vi.2018</collectingDate>
</date>
/ coll.
<collectorName id="26A39FFBFF8DFFECFD5BFEEFFCE08FF2" box="[718,820,369,395]" pageId="3" pageNumber="273">S. Bybee</collectorName>
&amp;
<collectorName id="26A39FFBFF8DFFECFCC2FEEFFC138FF2" box="[855,967,369,395]" pageId="3" pageNumber="273">G. Powell</collectorName>
//
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="DF29AAA5FF8DFFECFF02F8ECFACE89AA" ID-DOI="http://doi.org/10.5281/zenodo.3607105" ID-Zenodo-Dep="3607105" httpUri="https://zenodo.org/record/3607105/files/figure.png" pageId="3" pageNumber="273" startId="3.[151,250,1906,1931]" targetBox="[151,1436,410,1886]" targetPageId="3">
<paragraph id="8BE9FA2DFF8DFFECFF02F8ECFACE89AA" blockId="3.[151,1436,1906,2003]" pageId="3" pageNumber="273">
<emphasis id="B922263FFF8DFFECFF02F8ECFE9989F2" bold="true" box="[151,333,1906,1931]" pageId="3" pageNumber="273">FIGURE 1. A–I)</emphasis>
<taxonomicName id="4C5681AEFF8DFFECFEC0F8EDFE6289F2" box="[341,438,1907,1931]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="273" phylum="Arthropoda" rank="genus">
<emphasis id="B922263FFF8DFFECFEC0F8EDFE6289F2" box="[341,438,1907,1931]" italics="true" pageId="3" pageNumber="273">Atyphella</emphasis>
</taxonomicName>
spp. holotypes.
<emphasis id="B922263FFF8DFFECFDF4F8ECFD4E89F2" bold="true" box="[609,666,1906,1931]" pageId="3" pageNumber="273">A–C)</emphasis>
dorsal pronotum:
<emphasis id="B922263FFF8DFFECFCCFF8ECFCA089F2" bold="true" box="[858,884,1906,1931]" pageId="3" pageNumber="273">A)</emphasis>
<taxonomicName id="4C5681AEFF8DFFECFCE8F8EDFBCE89F2" baseAuthorityName="Ballantyne" baseAuthorityYear="1979" box="[893,1050,1907,1931]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="273" phylum="Arthropoda" rank="species" species="aphrogeneia">
<emphasis id="B922263FFF8DFFECFCE8F8EDFBCE89F2" box="[893,1050,1907,1931]" italics="true" pageId="3" pageNumber="273">A. aphrogeneia</emphasis>
</taxonomicName>
,
<emphasis id="B922263FFF8DFFECFBBDF8ECFB9689F2" bold="true" box="[1064,1090,1906,1931]" pageId="3" pageNumber="273">B)</emphasis>
<taxonomicName id="4C5681AEFF8DFFECFBDFF8EDFB1B89F3" authorityName="Saxton &amp; Powell" authorityYear="2020" box="[1098,1231,1907,1930]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="273" phylum="Arthropoda" rank="species" species="maritimus">
<emphasis id="B922263FFF8DFFECFBDFF8EDFB1B89F3" box="[1098,1231,1907,1930]" italics="true" pageId="3" pageNumber="273">A. maritimus</emphasis>
</taxonomicName>
,
<emphasis id="B922263FFF8DFFECFB48F8ECFB2C89F2" bold="true" box="[1245,1272,1906,1931]" pageId="3" pageNumber="273">C)</emphasis>
<taxonomicName id="4C5681AEFF8DFFECFA95F8EDFA4289F3" authorityName=", Saxton &amp; Bybee" authorityYear="2020" box="[1280,1430,1907,1930]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="273" phylum="Arthropoda" rank="species" species="marigenous">
<emphasis id="B922263FFF8DFFECFA95F8EDFA4289F3" box="[1280,1430,1907,1930]" italics="true" pageId="3" pageNumber="273">A. marigenous</emphasis>
</taxonomicName>
;
<emphasis id="B922263FFF8DFFECFF02F808FF1A89D6" bold="true" box="[151,206,1942,1967]" pageId="3" pageNumber="273">D–F)</emphasis>
scutellum:
<emphasis id="B922263FFF8DFFECFED1F808FE8A89D6" bold="true" box="[324,350,1942,1967]" pageId="3" pageNumber="273">D)</emphasis>
<taxonomicName id="4C5681AEFF8DFFECFEF0F809FDD489D6" baseAuthorityName="Ballantyne" baseAuthorityYear="1979" box="[357,512,1943,1967]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="273" phylum="Arthropoda" rank="species" species="aphrogeneia">
<emphasis id="B922263FFF8DFFECFEF0F809FDD489D6" box="[357,512,1943,1967]" italics="true" pageId="3" pageNumber="273">A. aphrogeneia</emphasis>
</taxonomicName>
,
<emphasis id="B922263FFF8DFFECFD98F808FDF189D6" bold="true" box="[525,549,1942,1967]" pageId="3" pageNumber="273">E)</emphasis>
<taxonomicName id="4C5681AEFF8DFFECFDB9F809FD7B89D7" authorityName="Saxton &amp; Powell" authorityYear="2020" box="[556,687,1943,1966]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="273" phylum="Arthropoda" rank="species" species="maritimus">
<emphasis id="B922263FFF8DFFECFDB9F809FD7B89D7" box="[556,687,1943,1966]" italics="true" pageId="3" pageNumber="273">A. maritimus</emphasis>
</taxonomicName>
,
<emphasis id="B922263FFF8DFFECFD2EF808FD0089D6" bold="true" box="[699,724,1942,1967]" pageId="3" pageNumber="273">F)</emphasis>
<emphasis id="B922263FFF8DFFECFD4CF809FCA189D7" box="[729,885,1943,1966]" italics="true" pageId="3" pageNumber="273">
<taxonomicName id="4C5681AEFF8DFFECFD4CF809FCBB89D7" authorityName=", Saxton &amp; Bybee" authorityYear="2020" box="[729,879,1943,1966]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="273" phylum="Arthropoda" rank="species" species="marigenous">A. marigenous</taxonomicName>
;
</emphasis>
<emphasis id="B922263FFF8DFFECFCE9F808FC7A89D6" bold="true" box="[892,942,1942,1967]" pageId="3" pageNumber="273">G–I)</emphasis>
left elytron, notice extent of costae
<emphasis id="B922263FFF8DFFECFA8FF808FAE289D6" bold="true" box="[1306,1334,1942,1967]" pageId="3" pageNumber="273">G)</emphasis>
<taxonomicName id="4C5681AEFF8DFFECFAA9F809FF0F89AB" baseAuthorityName="Ballantyne" baseAuthorityYear="1979" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="273" phylum="Arthropoda" rank="species" species="aphrogeneia">
<emphasis id="B922263FFF8DFFECFAA9F809FF0F89AB" italics="true" pageId="3" pageNumber="273">A. aphrogeneia</emphasis>
</taxonomicName>
,
<emphasis id="B922263FFF8DFFECFF72F824FED789AA" bold="true" box="[231,259,1978,2003]" pageId="3" pageNumber="273">H)</emphasis>
<taxonomicName id="4C5681AEFF8DFFECFE9FF825FE5989AB" authorityName="Saxton &amp; Powell" authorityYear="2020" box="[266,397,1979,2002]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="273" phylum="Arthropoda" rank="species" species="maritimus">
<emphasis id="B922263FFF8DFFECFE9FF825FE5989AB" box="[266,397,1979,2002]" italics="true" pageId="3" pageNumber="273">A. maritimus</emphasis>
</taxonomicName>
,
<emphasis id="B922263FFF8DFFECFE0CF824FE7889AA" bold="true" box="[409,428,1978,2003]" pageId="3" pageNumber="273">I)</emphasis>
<taxonomicName id="4C5681AEFF8DFFECFE27F825FD9289AB" authorityName=", Saxton &amp; Bybee" authorityYear="2020" box="[434,582,1979,2002]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="273" phylum="Arthropoda" rank="species" species="marigenous">
<emphasis id="B922263FFF8DFFECFE27F825FD9289AB" box="[434,582,1979,2002]" italics="true" pageId="3" pageNumber="273">A. marigenous</emphasis>
</taxonomicName>
.
<taxonomicName id="4C5681AEFF8DFFECFDC7F825FCED89AA" baseAuthorityName="Ballantyne" baseAuthorityYear="1979" box="[594,825,1979,2003]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="273" phylum="Arthropoda" rank="species" species="aphrogeneia">
<emphasis id="B922263FFF8DFFECFDC7F825FCED89AA" box="[594,825,1979,2003]" italics="true" pageId="3" pageNumber="273">Atyphella aphrogeneia</emphasis>
</taxonomicName>
holotype images are the property of © CSIRO.
</paragraph>
</caption>
<subSubSection id="C34CA9A6FF8AFFEBFF52FF06FBA48FF2" pageId="4" pageNumber="274" type="diagnosis">
<paragraph id="8BE9FA2DFF8AFFEBFF52FF06FBA48FF2" blockId="4.[151,1437,152,395]" pageId="4" pageNumber="274">
<emphasis id="B922263FFF8AFFEBFF52FF06FE968ECA" bold="true" box="[199,322,152,179]" pageId="4" pageNumber="274">Diagnosis.</emphasis>
<taxonomicName id="4C5681AEFF8AFFEBFEDFFF07FD968ECA" authorityName=", Saxton &amp; Bybee" authorityYear="2020" box="[330,578,153,179]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="274" phylum="Arthropoda" rank="species" species="marigenous">
<emphasis id="B922263FFF8AFFEBFEDFFF07FD968ECA" box="[330,578,153,179]" italics="true" pageId="4" pageNumber="274">Atyphella marigenous</emphasis>
</taxonomicName>
is distinguished from all other South Pacific
<emphasis id="B922263FFF8AFFEBFBDCFF07FB688ECA" box="[1097,1212,153,179]" italics="true" pageId="4" pageNumber="274">
<taxonomicName id="4C5681AEFF8AFFEBFBDCFF07FB638ECA" box="[1097,1207,153,179]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="274" phylum="Arthropoda" rank="genus">Atyphella</taxonomicName>
,
</emphasis>
except
<taxonomicName id="4C5681AEFF8AFFEBFA83FF07FF138EAE" baseAuthorityName="Ballantyne" baseAuthorityYear="1979" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="274" phylum="Arthropoda" rank="species" species="aphrogeneia">
<emphasis id="B922263FFF8AFFEBFA83FF07FF138EAE" italics="true" pageId="4" pageNumber="274">A. aphrogeneia</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5681AEFF8AFFEBFF6AFF23FE448EAE" authorityName="Saxton &amp; Powell" authorityYear="2020" box="[255,400,189,215]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="274" phylum="Arthropoda" rank="species" species="maritimus">
<emphasis id="B922263FFF8AFFEBFF6AFF23FE448EAE" box="[255,400,189,215]" italics="true" pageId="4" pageNumber="274">A. maritimus</emphasis>
</taxonomicName>
by the thickened elytral apices in conjunction with a dark median marking on the pronotum (see
<bibRefCitation id="EFC787DCFF8AFFEBFF59FF7FFDF18E82" author="Ballantyne, L. A. &amp; Lambkin, C." box="[204,549,225,251]" pageId="4" pageNumber="274" pagination="1 - 188" refId="ref4464" refString="Ballantyne, L. A. &amp; Lambkin, C. (2009) Systematics of Indo-Pacific fireflies with a redefinition of Australasian Atyphella Olliff, Madagascan Photuroluciola Pic, and description of seven new genera from the Luciolinae (Coleoptera: Lampyridae). Zootaxa, 1997 (1), 1 - 188. https: // doi. org / 10.11646 / zootaxa. 1997.1.1" type="journal article" year="2009">Ballantyne and Lambkin 2009</bibRefCitation>
).
<taxonomicName id="4C5681AEFF8AFFEBFDADFF7FFCFB8E82" authorityName=", Saxton &amp; Bybee" authorityYear="2020" box="[568,815,225,251]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="274" phylum="Arthropoda" rank="species" species="marigenous">
<emphasis id="B922263FFF8AFFEBFDADFF7FFCFB8E82" box="[568,815,225,251]" italics="true" pageId="4" pageNumber="274">Atyphella marigenous</emphasis>
</taxonomicName>
resembles
<taxonomicName id="4C5681AEFF8AFFEBFC38FF7FFBEB8E82" authorityName="Saxton &amp; Powell" authorityYear="2020" box="[941,1087,225,251]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="274" phylum="Arthropoda" rank="species" species="maritimus">
<emphasis id="B922263FFF8AFFEBFC38FF7FFBEB8E82" box="[941,1087,225,251]" italics="true" pageId="4" pageNumber="274">A. maritimus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5681AEFF8AFFEBFBE0FF7FFAF68E82" baseAuthorityName="Ballantyne" baseAuthorityYear="1979" box="[1141,1314,225,251]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="274" phylum="Arthropoda" rank="species" species="aphrogeneia">
<emphasis id="B922263FFF8AFFEBFBE0FF7FFAF68E82" box="[1141,1314,225,251]" italics="true" pageId="4" pageNumber="274">A. aphrogeneia</emphasis>
</taxonomicName>
but differs in the following ways: frons is deeply excavated (shallower in
<taxonomicName id="4C5681AEFF8AFFEBFCD8FE9BFC0A8F66" authorityName="Saxton &amp; Powell" authorityYear="2020" box="[845,990,261,287]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="274" phylum="Arthropoda" rank="species" species="maritimus">
<emphasis id="B922263FFF8AFFEBFCD8FE9BFC0A8F66" box="[845,990,261,287]" italics="true" pageId="4" pageNumber="274">A. maritimus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5681AEFF8AFFEBFB87FE9BFB698F66" baseAuthorityName="Ballantyne" baseAuthorityYear="1979" box="[1042,1213,261,287]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="274" phylum="Arthropoda" rank="species" species="aphrogeneia">
<emphasis id="B922263FFF8AFFEBFB87FE9BFB698F66" box="[1042,1213,261,287]" italics="true" pageId="4" pageNumber="274">A. aphrogeneia</emphasis>
</taxonomicName>
); pronotum broadly and continuously rounded (pronotum with weakly obtuse and anterior angles in
<taxonomicName id="4C5681AEFF8AFFEBFB80FEB7FB148F3A" baseAuthorityName="Ballantyne" baseAuthorityYear="1979" box="[1045,1216,297,323]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="274" phylum="Arthropoda" rank="species" species="aphrogeneia">
<emphasis id="B922263FFF8AFFEBFB80FEB7FB148F3A" box="[1045,1216,297,323]" italics="true" pageId="4" pageNumber="274">A. aphrogeneia</emphasis>
</taxonomicName>
); base of scutellum with less dense and larger punctation (apex of scutellum with dense punctation at the base in
<taxonomicName id="4C5681AEFF8AFFEBFB39FED3FAE98F1E" authorityName="Saxton &amp; Powell" authorityYear="2020" box="[1196,1341,333,359]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="274" phylum="Arthropoda" rank="species" species="maritimus">
<emphasis id="B922263FFF8AFFEBFB39FED3FAE98F1E" box="[1196,1341,333,359]" italics="true" pageId="4" pageNumber="274">A. maritimus</emphasis>
</taxonomicName>
); elytral costae pronounced (elytral costae less pronounced in
<taxonomicName id="4C5681AEFF8AFFEBFD78FEEFFC4D8FF2" baseAuthorityName="Ballantyne" baseAuthorityYear="1979" box="[749,921,369,395]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="274" phylum="Arthropoda" rank="species" species="aphrogeneia">
<emphasis id="B922263FFF8AFFEBFD78FEEFFC4D8FF2" box="[749,921,369,395]" italics="true" pageId="4" pageNumber="274">A. aphrogeneia</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5681AEFF8AFFEBFC5AFEEFFBB48FF2" authorityName="Saxton &amp; Powell" authorityYear="2020" box="[975,1120,369,395]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="274" phylum="Arthropoda" rank="species" species="maritimus">
<emphasis id="B922263FFF8AFFEBFC5AFEEFFBB48FF2" box="[975,1120,369,395]" italics="true" pageId="4" pageNumber="274">A. maritimus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<caption id="DF29AAA5FF8AFFEBFF02FB50FBD28B72" ID-DOI="http://doi.org/10.5281/zenodo.3607107" ID-Zenodo-Dep="3607107" httpUri="https://zenodo.org/record/3607107/files/figure.png" pageId="4" pageNumber="274" startId="4.[151,250,1230,1255]" targetBox="[151,1436,409,1210]" targetPageId="4">
<paragraph id="8BE9FA2DFF8AFFEBFF02FB50FBD28B72" blockId="4.[151,1436,1230,1291]" pageId="4" pageNumber="274">
<emphasis id="B922263FFF8AFFEBFF02FB50FE868A9E" bold="true" box="[151,338,1230,1255]" pageId="4" pageNumber="274">FIGURE 2. A–C)</emphasis>
<taxonomicName id="4C5681AEFF8AFFEBFEC2FB50FE6C8A9F" box="[343,440,1230,1254]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="274" phylum="Arthropoda" rank="genus">
<emphasis id="B922263FFF8AFFEBFEC2FB50FE6C8A9F" box="[343,440,1230,1254]" italics="true" pageId="4" pageNumber="274">Atyphella</emphasis>
</taxonomicName>
spp. dorsal view of aedeagus.
<emphasis id="B922263FFF8AFFEBFD79FB50FCD28A9E" bold="true" box="[748,774,1230,1255]" pageId="4" pageNumber="274">A)</emphasis>
<taxonomicName id="4C5681AEFF8AFFEBFC99FB51FC738A9F" baseAuthorityName="Ballantyne" baseAuthorityYear="1979" box="[780,935,1230,1254]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="274" phylum="Arthropoda" rank="species" species="aphrogeneia">
<emphasis id="B922263FFF8AFFEBFC99FB51FC738A9F" box="[780,935,1230,1254]" italics="true" pageId="4" pageNumber="274">A. aphrogeneia</emphasis>
</taxonomicName>
holotype,
<emphasis id="B922263FFF8AFFEBFB84FB50FBFF8A9E" bold="true" box="[1041,1067,1230,1255]" pageId="4" pageNumber="274">B)</emphasis>
<taxonomicName id="4C5681AEFF8AFFEBFBA5FB51FB678A9F" authorityName="Saxton &amp; Powell" authorityYear="2020" box="[1072,1203,1231,1254]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="274" phylum="Arthropoda" rank="species" species="maritimus">
<emphasis id="B922263FFF8AFFEBFBA5FB51FB678A9F" box="[1072,1203,1231,1254]" italics="true" pageId="4" pageNumber="274">A. maritimus</emphasis>
</taxonomicName>
paratype,
<emphasis id="B922263FFF8AFFEBFA8EFB50FAE28A9E" bold="true" box="[1307,1334,1230,1255]" pageId="4" pageNumber="274">C)</emphasis>
<taxonomicName id="4C5681AEFF8AFFEBFAA9FB51FF068B73" authorityName=", Saxton &amp; Bybee" authorityYear="2020" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="274" phylum="Arthropoda" rank="species" species="marigenous">
<emphasis id="B922263FFF8AFFEBFAA9FB51FF068B73" italics="true" pageId="4" pageNumber="274">A. marigenous</emphasis>
</taxonomicName>
holotype.
<taxonomicName id="4C5681AEFF8AFFEBFEABFB6CFDF08B73" baseAuthorityName="Ballantyne" baseAuthorityYear="1979" box="[318,548,1266,1290]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="274" phylum="Arthropoda" rank="species" species="aphrogeneia">
<emphasis id="B922263FFF8AFFEBFEABFB6CFDF08B73" box="[318,548,1266,1290]" italics="true" pageId="4" pageNumber="274">Atyphella aphrogeneia</emphasis>
</taxonomicName>
holotype images are the property of © CSIRO.
</paragraph>
</caption>
<subSubSection id="C34CA9A6FF8AFFEAFF52FAA6FADB8FF2" lastPageId="5" lastPageNumber="275" pageId="4" pageNumber="274" type="description">
<paragraph id="8BE9FA2DFF8AFFEAFF52FAA6FA978F3A" blockId="4.[151,1437,1336,2011]" lastBlockId="5.[151,1437,152,647]" lastPageId="5" lastPageNumber="275" pageId="4" pageNumber="274">
<emphasis id="B922263FFF8AFFEBFF52FAA6FE028B2A" bold="true" box="[199,470,1336,1363]" pageId="4" pageNumber="274">
Description.
<typeStatus id="54ED448FFF8AFFEBFEF7FAA7FE058B2A" box="[354,465,1337,1363]" pageId="4" pageNumber="274" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male. Overall, body parallel-sided, length
<quantity id="4CAE57C8FF8AFFEBFC5DFAA7FBC78B2A" box="[968,1043,1337,1363]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="274" unit="mm" value="10.0">10mm</quantity>
, width measured at elytral humeri
<quantity id="4CAE57C8FF8AFFEBFF02FAC3FF3E8B0E" box="[151,234,1373,1399]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="4" pageNumber="274" unit="mm" value="3.4">3.4mm</quantity>
.
<emphasis id="B922263FFF8AFFEBFF60FAC2FEAB8B0E" bold="true" box="[245,383,1372,1399]" pageId="4" pageNumber="274">Coloration:</emphasis>
Elytra and head dark brown to black, pronotum notably lighter, light brown with disc darker (
<figureCitation id="136DE6A8FF8AFFEBFF35FA1FFEDA8BE2" box="[160,270,1409,1435]" captionStart="FIGURE 1" captionStartId="3.[151,250,1906,1931]" captionTargetBox="[151,1436,410,1886]" captionTargetId="figure@3.[151,1436,410,1886]" captionTargetPageId="3" captionText="FIGURE 1. A–I) Atyphella spp. holotypes. A–C) dorsal pronotum: A) A. aphrogeneia, B) A. maritimus, C) A. marigenous; D–F) scutellum: D) A. aphrogeneia, E) A. maritimus, F) A. marigenous; G–I) left elytron, notice extent of costae G) A. aphrogeneia, H) A. maritimus, I) A. marigenous. Atyphella aphrogeneia holotype images are the property of © CSIRO." figureDoi="http://doi.org/10.5281/zenodo.3607105" httpUri="https://zenodo.org/record/3607105/files/figure.png" pageId="4" pageNumber="274">Fig. 1C, I</figureCitation>
). Venter lighter than pronotal margins, becoming darker apically but before the light organ. Sculpturing on dorsal body surfaces moderately shiny beneath pubescence; pubescence fine and gold in color. Legs at base tan, becoming darker distally.
<emphasis id="B922263FFF8AFFEBFE2FFA57FDD78B9A" bold="true" box="[442,515,1481,1507]" pageId="4" pageNumber="274">Head:</emphasis>
<quantity id="4CAE57C8FF8AFFEBFD9EFA57FD8D8B9A" box="[523,601,1481,1507]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="4" pageNumber="274" unit="mm" value="2.3">2.3mm</quantity>
at widest point (center of eyes), eyes large, black; frons deeply depressed, depression narrow and deep, gradually widening and becoming shallow towards clypeus, small, red apunctate region at narrowest point of depression, setae long and golden on frons; antennae filiform, light brown at scape and second antennomere and becoming darker distally; scape about
<quantity id="4CAE57C8FF8AFFEBFC9FF9ABFC8D8836" box="[778,857,1589,1615]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="274" unit="mm" value="0.3">0.3mm</quantity>
long, antennomeres 2–11 broadly uniform in length and about half as long as scape; mandibles broad, reddish brown, darker toward pointed apices; apical maxillary palpomere enlarged, widest almost at base, apex rounded, longer than wide; apical labial palpomere longer than wide, similar to that of apical maxillary palpomere but smaller and thinner, half as long as apical maxillary palpomere.
<emphasis id="B922263FFF8AFFEBFF02F95BFECD88A6" bold="true" box="[151,281,1733,1759]" pageId="4" pageNumber="274">Pronotum:</emphasis>
<quantity id="4CAE57C8FF8AFFEBFEB4F95BFEBB88A6" box="[289,367,1733,1759]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="4" pageNumber="274" unit="mm" value="2.2">2.2mm</quantity>
long,
<quantity id="4CAE57C8FF8AFFEBFE23F95BFDD188A6" box="[438,517,1733,1759]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="4" pageNumber="274" unit="mm" value="3.6">3.6mm</quantity>
wide, pronotal width slightly less than humeral width; margins light brown, basal third subparallel, broadly rounded anteriorly, weakly explanate, hind angles almost at right angles; disc dark brown, dorsal surface with dense punctation, with dark median region not reaching either anterior or posterior margins, median area of dorsal surface somewhat convex.
<emphasis id="B922263FFF8AFFEBFD52F8AEFC9C8932" bold="true" box="[711,840,1840,1867]" pageId="4" pageNumber="274">Scutellum:</emphasis>
trapezoidal with dense punctation, distance between punctures increasing anteriorly; brown with lighter lateral and posterior margins.
<emphasis id="B922263FFF8AFFEBFBBCF8CBFBAB8916" bold="true" box="[1065,1151,1877,1903]" pageId="4" pageNumber="274">Elytra:</emphasis>
<quantity id="4CAE57C8FF8AFFEBFB13F8CBFB008916" box="[1158,1236,1877,1903]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" pageId="4" pageNumber="274" unit="mm" value="7.9">7.9mm</quantity>
long, dark brown almost black, parallel–sided; punctures dense, not as large as that on pronotum, unevenly spaced; costae epipleuron developed as ridge along margin, widest at humeri, narrowing posteriorly, weakly developed towards apex; Costae well pronounced, becoming less distinct apically.
<emphasis id="B922263FFF8AFFEBFD28F85EFCEF89A2" bold="true" box="[701,827,1984,2011]" pageId="4" pageNumber="274">Abdomen:</emphasis>
Ventrites 1–4 tan, becoming darker anteriorly, fat body visible at margins of ventrite 2 to ventrite 4; light organ occupying all of ventrite 5–6.
<emphasis id="B922263FFF8BFFEAFBC2FF07FB4C8ECA" bold="true" box="[1111,1176,153,179]" pageId="5" pageNumber="275">Legs:</emphasis>
Dark brown except for pale coxae, trochanters, and basal half of femora; dark marking present on coxae 3.
<emphasis id="B922263FFF8BFFEAFBA2FF22FB608EAE" bold="true" box="[1079,1204,188,215]" pageId="5" pageNumber="275">Aedeagus:</emphasis>
<quantity id="4CAE57C8FF8BFFEAFB2FFF23FADD8EAE" box="[1210,1289,189,215]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="5" pageNumber="275" unit="mm" value="1.4">1.4mm</quantity>
long,
<quantity id="4CAE57C8FF8BFFEAFADBFF23FA488EAE" box="[1358,1436,189,215]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="275" unit="mm" value="0.6">0.6mm</quantity>
wide, slender, (
<figureCitation id="136DE6A8FF8BFFEAFED1FF7FFE4A8E82" box="[324,414,225,251]" captionStart="FIGURE 2" captionStartId="4.[151,250,1230,1255]" captionTargetBox="[151,1436,409,1210]" captionTargetId="figure@4.[151,1436,409,1210]" captionTargetPageId="4" captionText="FIGURE 2. A–C) Atyphella spp. dorsal view of aedeagus. A) A. aphrogeneia holotype, B) A. maritimus paratype, C) A. marigenous holotype. Atyphella aphrogeneia holotype images are the property of © CSIRO." figureDoi="http://doi.org/10.5281/zenodo.3607107" httpUri="https://zenodo.org/record/3607107/files/figure.png" pageId="5" pageNumber="275">Fig. 2C</figureCitation>
); median lobe slightly longer than lateral lobes; posterior end of lateral lobes run parallel; lateral margins of lateral lobes symmetrical, tapering to apex, and rounded, diverging anteriorly; interior margins of lateral lobes sclerotized from midpoint to apex; median lobe apex rounded and deeply ventrally excavated.
</paragraph>
<paragraph id="8BE9FA2DFF8BFFEAFF52FED2FADB8FF2" blockId="5.[151,1437,152,647]" pageId="5" pageNumber="275">
<emphasis id="B922263FFF8BFFEAFF52FED2FEEB8F1E" bold="true" box="[199,319,332,359]" pageId="5" pageNumber="275">Variation.</emphasis>
Males: Length
<quantity id="4CAE57C8FF8BFFEAFE7BFED3FDA88F1E" box="[494,636,333,359]" metricMagnitude="-3" metricUnit="m" metricValue="9.899999999999999" metricValueMax="10.2" metricValueMin="9.6" pageId="5" pageNumber="275" unit="mm" value="9.899999999999999" valueMax="10.2" valueMin="9.6">9.6–10.2mm</quantity>
(n=6), width
<quantity id="4CAE57C8FF8BFFEAFC80FED3FC408F1E" box="[789,916,333,359]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="3.4" metricValueMin="3.1" pageId="5" pageNumber="275" unit="mm" value="3.25" valueMax="3.4" valueMin="3.1">3.1–3.4mm</quantity>
(n=6). Pronotum of some specimens lighter in color, some with wider disc. Antennae color in some specimens generally uniform. Female: unknown.
</paragraph>
</subSubSection>
<subSubSection id="C34CA9A6FF8BFFEAFF52FE0BFC6C8FAA" pageId="5" pageNumber="275" type="etymology">
<paragraph id="8BE9FA2DFF8BFFEAFF52FE0BFC6C8FAA" blockId="5.[151,1437,152,647]" pageId="5" pageNumber="275">
<emphasis id="B922263FFF8BFFEAFF52FE0BFE9A8FD6" bold="true" box="[199,334,405,431]" pageId="5" pageNumber="275">Etymology.</emphasis>
In reference to this species habitat, the specific epithet for this species is derived from the Latin roots mar- referring to “the sea” and -genous meaning “the product of.”
</paragraph>
</subSubSection>
<subSubSection id="C34CA9A6FF8BFFEAFF52FE42FCC58CFE" pageId="5" pageNumber="275" type="biology_ecology">
<paragraph id="8BE9FA2DFF8BFFEAFF52FE42FCC58CFE" blockId="5.[151,1437,152,647]" pageId="5" pageNumber="275">
<emphasis id="B922263FFF8BFFEAFF52FE42FEF38F8E" bold="true" box="[199,295,476,503]" pageId="5" pageNumber="275">Biology.</emphasis>
<taxonomicName id="4C5681AEFF8BFFEAFEB8FE43FDF68F8E" authorityName=", Saxton &amp; Bybee" authorityYear="2020" box="[301,546,477,503]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="275" phylum="Arthropoda" rank="species" species="marigenous">
<emphasis id="B922263FFF8BFFEAFEB8FE43FDF68F8E" box="[301,546,477,503]" italics="true" pageId="5" pageNumber="275">Atyphella marigenous</emphasis>
</taxonomicName>
was collected on volcanic rock outcroppings extremely similar to those
<taxonomicName id="4C5681AEFF8BFFEAFAD5FE43FF008C62" authorityName="Saxton &amp; Powell" authorityYear="2020" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="275" phylum="Arthropoda" rank="species" species="maritimus">
<emphasis id="B922263FFF8BFFEAFAD5FE43FF008C62" italics="true" pageId="5" pageNumber="275">A. maritimus</emphasis>
</taxonomicName>
inhabit. Larvae were found in crevices on the rocks despite the frequent waves from the ocean. Larvae of all sizes were collected suggesting that this species reproduces year-round. Adult males were caught flying over these rocks, but no females were observed. The similar biology and morphology of this species to both
<taxonomicName id="4C5681AEFF8BFFEAFB49FDD7FAB98C1A" authorityName="Saxton &amp; Powell" authorityYear="2020" box="[1244,1389,585,611]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="275" phylum="Arthropoda" rank="species" species="maritimus">
<emphasis id="B922263FFF8BFFEAFB49FDD7FAB98C1A" box="[1244,1389,585,611]" italics="true" pageId="5" pageNumber="275">A. maritimus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5681AEFF8BFFEAFF02FDF3FE978CFE" baseAuthorityName="Ballantyne" baseAuthorityYear="1979" box="[151,323,621,647]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="275" phylum="Arthropoda" rank="species" species="aphrogeneia">
<emphasis id="B922263FFF8BFFEAFF02FDF3FE978CFE" box="[151,323,621,647]" italics="true" pageId="5" pageNumber="275">A. aphrogeneia</emphasis>
</taxonomicName>
suggest these species are closely related.
</paragraph>
</subSubSection>
</treatment>
</document>
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