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<document id="792E376CCB69A4DD69EB9836C1F6E3C9" ID-CLB-Dataset="21477" ID-DOI="10.3161/1733-5329(2007)9[13:ANSOHC]2.0.CO;2" ID-GBIF-Dataset="ead95ba6-be5d-41bd-bdbf-18e73cd3a7eb" ID-ISSN="1733-5329" ID-Zenodo-Dep="3944948" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1594741757959" checkinUser="valdenar" docAuthor="Bates, Paul J. J., Rossiter, Stephen J., Suyanto, Augustinus &amp; Kingston, Tigga" docDate="2007" docId="03FF33468E15FF8BFD52EC28FCE7FBE7" docLanguage="en" docName="ActaChiropterologica.9.1.13-26.pdf.imf" docOrigin="Acta Chiropterologica 9 (1)" docStyle="DocumentStyle:D17EC282BABB8B0847763DCC9F0C3514.3:ActaChiropterol.2016-.journal_article" docStyleId="D17EC282BABB8B0847763DCC9F0C3514" docStyleName="ActaChiropterol.2016-.journal_article" docStyleVersion="3" docTitle="Hipposideros boeadii Bates &amp; Rossiter &amp; Suyanto &amp; Kingston 2007, sp. nov." docType="treatment" docVersion="9" lastPageNumber="24" masterDocId="FFC64B3E8E17FF87FFAEE951FF80FF90" masterDocTitle="A new species of Hipposideros (Chiroptera: Hipposideridae) from Sulawesi" masterLastPageNumber="26" masterPageNumber="13" pageNumber="14" updateTime="1698843657427" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="CC99D4584E09D1E1AC8952156B8A41C7" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D7DC8F0E044B42E7EFA8921A6B67468E">
<mods:title id="3F9BAFF40E2DB396E384AC288F915AFB">A new species of Hipposideros (Chiroptera: Hipposideridae) from Sulawesi</mods:title>
</mods:titleInfo>
<mods:name id="675FDAFB8B04FB25B6A32CC032FF19CE" type="personal">
<mods:role id="497FD4CA078B9DC32D0A10E9EEBAA62B">
<mods:roleTerm id="DDC5D7B415F694F23D9C6E8C8C2D51DE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F6630A4A6BA7AB79CBBB391673DBE710">Bates, Paul J. J.</mods:namePart>
<mods:affiliation id="21FE5F7EF96209B7EE9BA88C40F0ED56">Harrison Institute, Centre for Systematics and Biodiversity Research, Bowerwood House, St. Botolphs Road, Sevenoaks, Kent, TN 13 3 AQ, United Kingdom; E-mail: harrisoninstitute @ btopenworld. com</mods:affiliation>
</mods:name>
<mods:name id="70DC669D740F97653AF9AB74F7728E5C" type="personal">
<mods:role id="F27A57AAAD76A820AF19B62D98045293">
<mods:roleTerm id="6016F6D3ED7800483431349D7175CA99">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AFFDC00135B3A00A23B2DD5B9D9E5E47">Rossiter, Stephen J.</mods:namePart>
<mods:affiliation id="3C9C29CDFAA5EAA5E21B27AD8C3CE69F">School of Biological and Chemical Sciences, Queen Mary, University of London, London, E 1 4 NS, United Kingdom</mods:affiliation>
</mods:name>
<mods:name id="C1A6B554669EB03A6C3730025BC76C06" type="personal">
<mods:role id="B0D8EEB5AC2D58BDA6F468D305835E80">
<mods:roleTerm id="4724783046CFEED85D448FF30F3EE1DC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5A49B82C4A0D11CC1AA382CF8A785402">Suyanto, Augustinus</mods:namePart>
<mods:affiliation id="0E2BC8B40850E5AE7C3FB62BA5F507E0">Museum Zoologicum Bogoriense, Research Centre for Biology, Indonesian Institute of Sciences (LIPI), Widyasatwaloka Building, Jl. Raya Bogor Km 46, Cibinong 16911, Indonesia</mods:affiliation>
</mods:name>
<mods:name id="806CD50E6C13F18725D84845FBE34B93" type="personal">
<mods:role id="00132891574C7B2C42D2DE82142234D5">
<mods:roleTerm id="9654CFE33E1858E55F1DC202475C5ACE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7CAED429FB1B612C6ED7F8BE4CE398C8">Kingston, Tigga</mods:namePart>
<mods:affiliation id="9C00C5F28D1399210EDD6B1C429E3175">Department of Biological Sciences, Texas Tech University, Lubbock, TX 79409, USA</mods:affiliation>
</mods:name>
<mods:typeOfResource id="575283D10698027B63B28004EB732EB1">text</mods:typeOfResource>
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<mods:titleInfo id="6B1C463914D3282EE22B59B2D9FA9E1C">
<mods:title id="84FB8E72EF89FC9E7D9F453C026CAE90">Acta Chiropterologica</mods:title>
</mods:titleInfo>
<mods:part id="FB5A04FB4F14CFAE9BB25C9F82EA15D6">
<mods:date id="347FA7BE37B49AC4305BDB9311E3EFA5">2007</mods:date>
<mods:detail id="55388448392FEA91661961B959F2689B" type="pubDate">
<mods:number id="1CBF4D9A4E21C580A9BB95697C244907">2007-04-01</mods:number>
</mods:detail>
<mods:detail id="679AEEC08131013714C6FF0BC98AF4CA" type="volume">
<mods:number id="859AD67BCC52D241D9321B800FD80A73">9</mods:number>
</mods:detail>
<mods:detail id="46263E654F5EF1253C564E26D209201C" type="issue">
<mods:number id="FD7BBE748CAE0887C2390D682B3EB8AA">1</mods:number>
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<mods:start id="249C1FC66160A0C31CFA7AE292098C15">13</mods:start>
<mods:end id="D6F1CC050CA2BD3C759E37CEDA3D8A41">26</mods:end>
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<mods:classification id="FFF0AB0C7C068585C5FBB320AD4811E4">journal article</mods:classification>
<mods:identifier id="C4ACE7AB07E3683A4DB38446DA209EF7" type="CLB-Dataset">21477</mods:identifier>
<mods:identifier id="28CC501A6338EFE3A1B5AAEEE26F98B2" type="DOI">10.3161/1733-5329(2007)9[13:ANSOHC]2.0.CO;2</mods:identifier>
<mods:identifier id="3A1611C413EDD941EBD952EBDC891223" type="GBIF-Dataset">ead95ba6-be5d-41bd-bdbf-18e73cd3a7eb</mods:identifier>
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<mods:identifier id="569D59BEA582A0EB8D2883C0F71A6092" type="Zenodo-Dep">3944948</mods:identifier>
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<treatment id="03FF33468E15FF8BFD52EC28FCE7FBE7" ID-DOI="http://doi.org/10.5281/zenodo.4323904" ID-GBIF-Taxon="165089388" ID-Zenodo-Dep="4323904" LSID="urn:lsid:plazi:treatment:03FF33468E15FF8BFD52EC28FCE7FBE7" httpUri="http://treatment.plazi.org/id/03FF33468E15FF8BFD52EC28FCE7FBE7" lastPageId="12" lastPageNumber="24" pageId="2" pageNumber="14">
<subSubSection id="C34CD1DB8E15FF85FD52EC28FBDDFA04" box="[764,1117,1401,1428]" pageId="2" pageNumber="14" type="nomenclature">
<paragraph id="8BE982508E15FF85FD52EC28FBDDFA04" blockId="2.[764,1117,1401,1428]" box="[764,1117,1401,1428]" pageId="2" pageNumber="14">
<heading id="D0A1353C8E15FF85FD52EC28FBDDFA04" box="[764,1117,1401,1428]" pageId="2" pageNumber="14">
<taxonomicName id="4C56F9D38E15FF85FD52EC28FC7AFA03" ID-CoL="3M367" authority="Bates &amp; Rossiter &amp; Suyanto &amp; Kingston, 2007" authorityName="Bates &amp; Rossiter &amp; Suyanto &amp; Kingston" authorityYear="2007" box="[764,1018,1401,1427]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="2" pageNumber="14" phylum="Chordata" rank="species" species="boeadii" status="sp. nov.">
<emphasis id="B9225E428E15FF85FD52EC28FC7AFA03" box="[764,1018,1401,1427]" italics="true" pageId="2" pageNumber="14">Hipposideros boeadii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A211E3398E15FF85FBACEC2BFBDDFA04" box="[1026,1117,1402,1428]" pageId="2" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34CD1DB8E15FF85FD05ECE9FBCAF9CB" pageId="2" pageNumber="14" type="materials_examined">
<paragraph id="8BE982508E15FF85FD05ECE9FC97FA42" blockId="2.[683,1199,1464,1730]" box="[683,791,1464,1490]" pageId="2" pageNumber="14">
<materialsCitation id="3B3E880D8E15FF85FD05ECE9FC97FA42" ID-GBIF-Occurrence="2813291301" box="[683,791,1464,1490]" collectingDate="3 August" pageId="2" pageNumber="14" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis id="B9225E428E15FF85FD05ECE9FC97FA42" box="[683,791,1464,1490]" italics="true" pageId="2" pageNumber="14">
<typeStatus id="54ED3CF28E15FF85FD05ECE9FC97FA42" box="[683,791,1464,1490]" pageId="2" pageNumber="14" type="holotype">Holotype</typeStatus>
</emphasis>
</materialsCitation>
</paragraph>
<paragraph id="8BE982508E15FF85FD7FEC8AFBCAF9CB" blockId="2.[683,1199,1464,1730]" pageId="2" pageNumber="14">
Museum Zoologicum Bogoriense M28253 (Field No. TK000803.2), adult
<specimenCount id="9D5049D98E15FF85FB21ECABFB27F987" box="[1167,1191,1530,1559]" pageId="2" pageNumber="14" type="female"></specimenCount>
, body in alcohol, skull extracted, collected by TK and SJR on
<date id="FFE8A4908E15FF85FC20EF10FC7BF9CB" box="[910,1019,1601,1627]" pageId="2" pageNumber="14">
<collectingDate id="EFAC5D788E15FF85FC20EF10FC7BF9CB" box="[910,1019,1601,1627]" pageId="2" pageNumber="14">3 August</collectingDate>
</date>
, 2000.
</paragraph>
</subSubSection>
<subSubSection id="C34CD1DB8E15FF84FD7FEF35FDF1FC8E" lastPageId="3" lastPageNumber="15" pageId="2" pageNumber="14" type="description">
<paragraph id="8BE982508E15FF85FD7FEF35FB2FF952" blockId="2.[683,1199,1464,1730]" pageId="2" pageNumber="14">Measurements (in mm) — forearm (FA): 40.7; fifth (5MET), fourth (4MET) and third (3MET) metacarpal lengths</paragraph>
<paragraph id="8BE982508E14FF84FF2CE9F2FDF1FC8E" blockId="3.[130,643,163,1731]" pageId="3" pageNumber="15">
respectively: 28.7, 32.0, 32.3; length of first and second phalanges of third digit (3D1P and 3D2P) respectively: 13.6, 14.0; tail (TAIL): 16.0; foot (HF): 8.7; tibia (TIB): 17.0; ear (E): 18.0; body mass (WGT)
<quantity id="4CAE2FB58E14FF84FDF0E87AFF4DFEF7" metricMagnitude="-3" metricUnit="kg" metricValue="8.5" pageId="3" pageNumber="15" unit="g" value="8.5">8.5 grams</quantity>
; greatest skull length (GTL): 18.0; condylobasal length (CBL): 15.6; condylo- canine length (CCL): 15.2; zygomatic breadth (ZB): 10.2; breadth of braincase (BB): 7.7; mastoid width (MW): 9.0; rostral width (RW) 5.0; postorbital constriction (PC): 2.9; upper toothrow length (CM
<superScript id="7C232F188E14FF84FDC8EB42FDF0FDB6" attach="left" box="[614,624,531,550]" fontSize="8" pageId="3" pageNumber="15">3</superScript>
): 6.7; breadth of palate (M
<superScript id="7C232F188E14FF84FE1EEB65FE3AFDD7" attach="left" box="[432,442,564,583]" fontSize="8" pageId="3" pageNumber="15">3</superScript>
M
<superScript id="7C232F188E14FF84FE4DEB65FE6DFDD7" attach="left" box="[483,493,564,583]" fontSize="8" pageId="3" pageNumber="15">3</superScript>
): 7.3; lower toothrow length (CM
<subScript id="17D280158E14FF84FE0AEB38FE2EFDEC" attach="left" box="[420,430,617,636]" fontSize="8" pageId="3" pageNumber="15">3</subScript>
): 7.2; mandible length (M): 11.9; first upper premolar width (P
<superScript id="7C232F188E14FF84FF32EBCBFF26FD3D" attach="both" box="[156,166,666,685]" fontSize="8" pageId="3" pageNumber="15">2</superScript>
W): 0.50 and length (P
<superScript id="7C232F188E14FF84FE63EBCBFE57FD3D" attach="both" box="[461,471,666,685]" fontSize="8" pageId="3" pageNumber="15">2</superScript>
L): 0.45; first lower premolar width (P
<subScript id="17D280158E14FF84FE68EB9EFE50FD72" attach="both" box="[454,464,719,738]" fontSize="8" pageId="3" pageNumber="15">2</subScript>
W): 0.45 and length (P
<subScript id="17D280158E14FF84FF5EEBA0FF7AFC94" attach="both" box="[240,250,753,772]" fontSize="8" pageId="3" pageNumber="15">2</subScript>
L): 0.45; second lower premolar width (P
<subScript id="17D280158E14FF84FF49EA43FF71FCB5" attach="both" box="[231,241,786,805]" fontSize="8" pageId="3" pageNumber="15">4</subScript>
W): 0.80 and length (P
<subScript id="17D280158E14FF84FDAEEA43FD8AFCB5" attach="both" box="[512,522,786,805]" fontSize="8" pageId="3" pageNumber="15">4</subScript>
L): 0.90.
</paragraph>
</subSubSection>
<caption id="DF29D2D88E14FF84FD00EBD9FB15FD2A" ID-DOI="http://doi.org/10.5281/zenodo.3944950" ID-Zenodo-Dep="3944950" httpUri="https://zenodo.org/record/3944950/files/figure.png" pageId="3" pageNumber="15" startId="3.[686,699,648,669]" targetBox="[683,1201,161,625]" targetPageId="3">
<paragraph id="8BE982508E14FF84FD00EBD9FB15FD2A" blockId="3.[686,1199,647,698]" pageId="3" pageNumber="15">
<subScript id="17D280158E14FF84FD00EBD9FB15FD2A" attach="right" fontSize="9" pageId="3" pageNumber="15">
FIG. 1.
<taxonomicName id="4C56F9D38E14FF84FCA8EBD6FC65FD0C" authorityName="Bates &amp; Rossiter &amp; Suyanto &amp; Kingston" authorityYear="2007" box="[774,997,647,668]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="3" pageNumber="15" phylum="Chordata" rank="species" species="boeadii">
<emphasis id="B9225E428E14FF84FCA8EBD6FC65FD0C" box="[774,997,647,668]" italics="true" pageId="3" pageNumber="15">Hipposideros boeadii</emphasis>
</taxonomicName>
from Rawa Aopa Watumohai National Park, South-East
<collectingRegion id="49924CB28E14FF84FB90EBF5FB15FD2A" box="[1086,1173,676,698]" country="Indonesia" name="Sulawesi Utara" pageId="3" pageNumber="15">Sulawesi</collectingRegion>
</subScript>
</paragraph>
</caption>
<subSubSection id="C34CD1DB8E14FF84FF2CEA1EFE84FC5E" pageId="3" pageNumber="15" type="distribution">
<paragraph id="8BE982508E14FF84FF2CEA1EFEA1FCF8" blockId="3.[130,643,163,1731]" box="[130,289,846,872]" pageId="3" pageNumber="15">
<emphasis id="B9225E428E14FF84FF2CEA1EFEA1FCF8" box="[130,289,846,872]" italics="true" pageId="3" pageNumber="15">
<typeStatus id="54ED3CF28E14FF84FF2CEA1EFF39FCF8" box="[130,185,847,872]" pageId="3" pageNumber="15">Type</typeStatus>
Locality
</emphasis>
</paragraph>
<paragraph id="8BE982508E14FF84FF06EA21FE84FC5E" blockId="3.[130,643,163,1731]" pageId="3" pageNumber="15">
Lanowulu, Rawa Aopa Watumohai Na- tional Park, South-East
<collectingRegion id="49924CB28E14FF84FE30EAC0FD88FC3B" box="[414,520,913,939]" country="Indonesia" name="Sulawesi Utara" pageId="3" pageNumber="15">Sulawesi</collectingRegion>
(
<geoCoordinate id="EE62E4978E14FF84FDB4EAC3FDFDFC3C" box="[538,637,914,940]" degrees="04" direction="south" minutes="22" orientation="latitude" pageId="3" pageNumber="15" precision="925" value="-4.366667">04°22S</geoCoordinate>
,
<geoCoordinate id="EE62E4978E14FF84FF2CEAE5FF77FC5E" box="[130,247,948,974]" degrees="121" direction="east" minutes="44" orientation="longitude" pageId="3" pageNumber="15" precision="925" value="121.73333">121°44E</geoCoordinate>
).
</paragraph>
</subSubSection>
<subSubSection id="C34CD1DB8E14FF84FF2CEAAFFD00FAD8" pageId="3" pageNumber="15" type="materials_examined">
<paragraph id="8BE982508E14FF84FF2CEAAFFF79FB87" blockId="3.[130,643,163,1731]" box="[130,249,1022,1047]" pageId="3" pageNumber="15">
<emphasis id="B9225E428E14FF84FF2CEAAFFF79FB87" box="[130,249,1022,1047]" italics="true" pageId="3" pageNumber="15">
<typeStatus id="54ED3CF28E14FF84FF2CEAAFFF79FB87" box="[130,249,1022,1047]" pageId="3" pageNumber="15" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BE982508E14FF84FF06ED71FD00FAD8" blockId="3.[130,643,163,1731]" pageId="3" pageNumber="15">
Museum Zoologicum Bogoriense M22935 (
<collectorName id="26A3E7868E14FF84FF57ED10FE20FBCB" box="[249,416,1089,1116]" pageId="3" pageNumber="15">Field No. TK</collectorName>
000804.2) adult ♁ (4
<collectorName id="26A3E7868E14FF84FF02ED32FE88FBED" box="[172,264,1123,1149]" pageId="3" pageNumber="15">August</collectorName>
, 2000);
<accessionNumber id="94051FB38E14FF84FE2FED32FE68FBED" box="[385,488,1123,1149]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/M22937" pageId="3" pageNumber="15" type="EnaNcbi">M22937</accessionNumber>
(
<collectorName id="26A3E7868E14FF84FDADED32FD03FBED" box="[515,643,1123,1149]" pageId="3" pageNumber="15">Field No.</collectorName>
TK000809.2) adult ♁ (9
<collectorName id="26A3E7868E14FF84FE64EDD4FDA2FB0F" box="[458,546,1157,1183]" pageId="3" pageNumber="15">August</collectorName>
, 2000); M22934 (
<collectorName id="26A3E7868E14FF84FF57EDF7FE21FB51" box="[249,417,1190,1217]" pageId="3" pageNumber="15">Field No. TK</collectorName>
000804.1) adult
<specimenCount id="9D5049D98E14FF84FDC4EDF6FD02FB51" box="[618,642,1191,1217]" pageId="3" pageNumber="15" type="female"></specimenCount>
(4
<collectorName id="26A3E7868E14FF84FF0EED99FF78FB72" box="[160,248,1224,1250]" pageId="3" pageNumber="15">August</collectorName>
, 2000); all bodies in alcohol with skulls extracted collected at
<collectingMunicipality id="6B8D182A8E14FF84FDACEDBBFDFEFA94" box="[514,638,1258,1284]" pageId="3" pageNumber="15">Lanowulu</collectingMunicipality>
,
<location id="8E89D48B8E14FF84FF2CEC5DFDFFFAB6" LSID="urn:lsid:plazi:treatment:03FF33468E15FF8BFD52EC28FCE7FBE7:8E89D48B8E14FF84FF2CEC5DFDFFFAB6" box="[130,639,1292,1318]" county="Description" latitude="-4.366667" longLatPrecision="1306" longitude="121.73333" municipality="Lanowulu" name="Rawa Aopa Watumo-hai National Park" pageId="3" pageNumber="15" stateProvince="Sulawesi Utara">Rawa Aopa Watumo-hai National Park</location>
,
<location id="8E89D48B8E14FF84FF2CEC7FFE83FAD8" LSID="urn:lsid:plazi:treatment:03FF33468E15FF8BFD52EC28FCE7FBE7:8E89D48B8E14FF84FF2CEC7FFE83FAD8" box="[130,259,1326,1352]" county="Description" latitude="-4.366667" longLatPrecision="1306" longitude="121.73333" municipality="Lanowulu" name="South-East" pageId="3" pageNumber="15" stateProvince="Sulawesi Utara">South-East</location>
<collectingRegion id="49924CB28E14FF84FEA5EC7FFEF8FAD8" box="[267,376,1326,1352]" country="Indonesia" name="Sulawesi Utara" pageId="3" pageNumber="15">Sulawesi</collectingRegion>
, (
<geoCoordinate id="EE62E4978E14FF84FE23EC7FFE71FAD8" box="[397,497,1326,1352]" degrees="04" direction="south" minutes="22" orientation="latitude" pageId="3" pageNumber="15" precision="925" value="-4.366667">04°22S</geoCoordinate>
,
<geoCoordinate id="EE62E4978E14FF84FE50EC7FFDF2FAD8" box="[510,626,1326,1352]" degrees="121" direction="east" minutes="44" orientation="longitude" pageId="3" pageNumber="15" precision="925" value="121.73333">121°44E</geoCoordinate>
).
</paragraph>
</subSubSection>
<subSubSection id="C34CD1DB8E14FF84FF2CEC29FCEEFB30" pageId="3" pageNumber="15" type="diagnosis">
<paragraph id="8BE982508E14FF84FF2CEC29FF79FA01" blockId="3.[130,643,163,1731]" box="[130,249,1400,1425]" pageId="3" pageNumber="15">Diagnosis</paragraph>
<paragraph id="8BE982508E14FF84FF06ECC8FCEEFB30" blockId="3.[130,643,163,1731]" lastBlockId="3.[686,1199,747,1184]" pageId="3" pageNumber="15">
This is a medium-small hipposiderid bat with a forearm length of
<quantity id="4CAE2FB58E14FF84FE0AECEAFDCEFA46" box="[420,590,1467,1494]" metricMagnitude="-2" metricUnit="m" metricValue="4.16" metricValueMax="4.27" metricValueMin="4.05" pageId="3" pageNumber="15" unit="mm" value="41.6" valueMax="42.7" valueMin="40.5">40.542.7 mm</quantity>
tak- en from specimens preserved in alcohol (
<tableCitation id="C6D4B7EB8E14FF84FF24ECAEFF64F989" box="[138,228,1535,1561]" captionStart="TABLE" captionStartId="4.[86,99,647,674]" captionTargetPageId="4" captionText="TABLE 1. External measurements (in mm) and body mass (in g) of four species of Hipposideros recently collected in Sulawesi. Used abbreviations: FA — forearm length; 5MET, 4MET, 3MET — fifth, fourth, and third metacarpal lengths; 5MET/FA× 100 — % length of fifth metacarpal relative to forearm length; 3D1P, 3D2P — length of first and second phalanges of the third digit; 3D1P/3MET × 100 — % length of first phalanx of the third digit relative to third metacarpal length; TAIL — tail length; HF — foot; TIBIA — tibia; E — ear and WGT — body mass. Sample sizes differing from those reported under n are given in parentheses. Mean and SD are included when sample size is ≥ 3" httpUri="http://table.plazi.org/id/DF29D2D88E13FF83FFF8EBD6FD8AFC83" pageId="3" pageNumber="15" tableUuid="DF29D2D88E13FF83FFF8EBD6FD8AFC83">Table 1</tableCitation>
). The ears have rounded tips. The noseleaf has two supplementary leaflets; the inner is short; the outer virtually reach- es the midline anteriorly (
<figureCitation id="136D9ED58E14FF84FE15EF35FD8FF9EE" box="[443,527,1636,1662]" captionStart="FIG" captionStartId="3.[686,699,648,669]" captionTargetBox="[683,1201,161,625]" captionTargetId="figure@3.[683,1202,159,625]" captionTargetPageId="3" captionText="FIG. 1. Hipposideros boeadii from Rawa Aopa Watumohai National Park, South-East Sulawesi" figureDoi="http://doi.org/10.5281/zenodo.3944950" httpUri="https://zenodo.org/record/3944950/files/figure.png" pageId="3" pageNumber="15">Figs. 1</figureCitation>
and
<figureCitation id="136D9ED58E14FF84FDE0EF35FDF5F9EE" box="[590,629,1636,1662]" captionStart="FIG" captionStartId="5.[132,145,1490,1511]" captionTargetBox="[129,652,940,1452]" captionTargetId="figure@5.[355,658,938,1463]" captionTargetPageId="5" captionText="FIG. 2. Noseleaves of four species of Hipposideros (with hairs omitted). A — H. boeadii (holotype), M28253, Rawa Aopa Watumohai National Park, South-East Sulawesi; B — H. papua, M16951, Biak Island, Papua Province; C — H. galeritus, M28250, Kakenauwe Nature Reserve, Buton Island, South- East Sulawesi; D — H. cervinus, M28249, Lambasango Wildlife Reserve, Buton Island, South- East Sulawesi. Scale = 5 mm" figureDoi="http://doi.org/10.5281/zenodo.3944954" httpUri="https://zenodo.org/record/3944954/files/figure.png" pageId="3" pageNumber="15">2A</figureCitation>
). In the wing, the fifth metacarpal is be- tween 7075% of forearm length and the first phalanx of the third digit is short (
<quantity id="4CAE2FB58E14FF84FD18EA5CFCE9FCB8" box="[694,873,781,808]" metricMagnitude="-2" metricUnit="m" metricValue="1.375" metricValueMax="1.39" metricValueMin="1.3599999999999999" pageId="3" pageNumber="15" unit="mm" value="13.75" valueMax="13.9" valueMin="13.6">13.613.9 mm</quantity>
). The tail is also very short (16.021.0 mm). The skull, with a condylo- canine length of
<quantity id="4CAE2FB58E14FF84FCDFEA00FB9BFCFC" box="[881,1051,849,876]" metricMagnitude="-2" metricUnit="m" metricValue="1.535" metricValueMax="1.55" metricValueMin="1.52" pageId="3" pageNumber="15" unit="mm" value="15.35" valueMax="15.5" valueMin="15.2">15.215.5 mm</quantity>
(
<tableCitation id="C6D4B7EB8E14FF84FB84EA00FB05FCFC" box="[1066,1157,849,876]" captionStart="TABLE" captionStartId="9.[89,102,641,668]" captionText="TABLE 2. Cranial and dental measurements (in mm) of four species of Hipposideros recently collected in Sulawesi. Used abbreviations: GTL — greatest length of skull; CBL — condylobasal length; CCL — condylo-canine length; ZB — zygomatic breadth; BB — breadth of braincase; MW — mastoid width; ZB/MW × 100 — % breadth of zygomata relative to mastoid breadth; RW — rostral width; PC — postorbital constriction; CM3 — upper toothrow length; CM3/CCL × 100 — % length of upper toothrow relative to condylo-canine length; M3M3 — palatal breadth; CM — lower toothrow length; M — mandible length. Sample sizes 3 differing from those reported under n are given in parentheses. Mean and SD are included when sample size is ≥ 3" pageId="3" pageNumber="15">Table 2</tableCitation>
), is large in comparison to forearm length. The zygomata are robust, broad and consider- ably exceed mastoid width (
<figureCitation id="136D9ED58E14FF84FC59EAE9FBD5FC42" box="[1015,1109,952,978]" captionStart="FIG" captionStartId="6.[130,143,1651,1672]" captionTargetBox="[288,1018,552,1640]" captionTargetId="figure@6.[244,1035,543,1637]" captionTargetPageId="6" captionText="FIG. 3. Dorsal (above) and ventral (below) views of the skull and dorsal view of the right half mandible of two species of Hipposideros. A — H. boeadii (holotype), M28253, Rawa Aopa Watumohai National Park, South- East Sulawesi, and B — H. papua, M16951, Biak Island, Papua Province. Scale = 5 mm" figureDoi="http://doi.org/10.5281/zenodo.3944958" httpUri="https://zenodo.org/record/3944958/files/figure.png" pageId="3" pageNumber="15">Fig. 3A</figureCitation>
). There is a well developed dorsal projection on the jugal bone (
<figureCitation id="136D9ED58E14FF84FC99EAADFC16FB86" box="[823,918,1020,1046]" captionStart="FIG" captionStartId="7.[130,143,1486,1507]" captionTargetBox="[146,639,341,1457]" captionTargetId="figure@7.[125,645,615,920]" captionTargetPageId="7" captionText="FIG. 4. Lateral view of the skulls and mandibles of four species of Hipposideros. A — H. boeadii (holo- type), M28253, Rawa Aopa Watumohai National Park, South-East Sulawesi; B — H. papua, M16951, Biak Island, Papua Province; C — H. galeritus, M28250, Kakenauwe Nature Reserve, Buton Island, South-East Sulawesi; D — H. cervinus, M28249, Lambasango Wildlife Reserve, Buton Island, South- East Sulawesi. Scale = 5 mm" figureDoi="http://doi.org/10.5281/zenodo.3944962" httpUri="https://zenodo.org/record/3944962/files/figure.png" pageId="3" pageNumber="15">Fig. 4A</figureCitation>
). The rostrum is moder- ately inflated with a slight depression in the midline and with the anterior and postero- medial rostral chambers about equally in- flated (
<figureCitation id="136D9ED58E14FF84FCAFEDD4FCE0FB30" box="[769,864,1157,1184]" captionStart="FIG" captionStartId="8.[130,143,1626,1647]" captionTargetBox="[273,1107,833,1601]" captionTargetId="figure@8.[235,1126,829,1604]" captionTargetPageId="8" captionText="FIG. 5. Dorsal view of the rostrum of four species of Hipposideros. A — H. boeadii (holotype), M28253, Rawa Aopa Watumohai National Park, South-East Sulawesi; B — H. papua, M16951, Biak Island, Papua Province; C — H. galeritus, M28250, Kakenauwe Nature Reserve, Buton Island, South-East Sulawesi; D —" figureDoi="http://doi.org/10.5281/zenodo.3944964" httpUri="https://zenodo.org/record/3944964/files/figure.png" pageId="3" pageNumber="15">Fig. 5A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34CD1DB8E14FF80FD00ED9BFBE5FA3D" lastPageId="7" lastPageNumber="19" pageId="3" pageNumber="15" type="description">
<paragraph id="8BE982508E14FF84FD00ED9BFCB9FB73" blockId="3.[686,1198,1226,1731]" box="[686,825,1226,1251]" pageId="3" pageNumber="15">
<emphasis id="B9225E428E14FF84FD00ED9BFCB9FB73" box="[686,825,1226,1251]" italics="true" pageId="3" pageNumber="15">
<collectingCounty id="6288FADC8E14FF84FD00ED9BFCB9FB73" box="[686,825,1226,1251]" pageId="3" pageNumber="15">Description</collectingCounty>
</emphasis>
</paragraph>
<paragraph id="8BE982508E14FF84FD7DEDBDFB2EF952" blockId="3.[686,1198,1226,1731]" pageId="3" pageNumber="15">
<location id="8E89D48B8E14FF84FD7DEDBDFC8CFA96" LSID="urn:lsid:plazi:treatment:03FF33468E15FF8BFD52EC28FCE7FBE7:8E89D48B8E14FF84FD7DEDBDFC8CFA96" box="[723,780,1260,1286]" county="Description" latitude="-4.366667" longLatPrecision="1306" longitude="121.73333" municipality="Lanowulu" name="With" pageId="3" pageNumber="15" stateProvince="Sulawesi Utara">With</location>
a body mass of
<quantity id="4CAE2FB58E14FF84FC7DEDBDFBFBFA96" box="[979,1147,1260,1287]" metricMagnitude="-3" metricUnit="kg" metricValue="7.65" metricValueMax="8.5" metricValueMin="6.8" pageId="3" pageNumber="15" unit="g" value="7.65" valueMax="8.5" valueMin="6.8">6.88.5 grams</quantity>
and a forearm length of
<quantity id="4CAE2FB58E14FF84FC38EC5FFBC0FAB8" box="[918,1088,1294,1320]" metricMagnitude="-2" metricUnit="m" metricValue="4.165000000000001" metricValueMax="4.3100000000000005" metricValueMin="4.0200000000000005" pageId="3" pageNumber="15" unit="mm" value="41.65" valueMax="43.1" valueMin="40.2">40.243.1 mm</quantity>
(live and alcohol preserved specimen data combined,
<emphasis id="B9225E428E14FF84FD00EC02FD3DFAFC" box="[686,701,1363,1388]" italics="true" pageId="3" pageNumber="15">n</emphasis>
= 15), this is a medium-small
<emphasis id="B9225E428E14FF84FBF5EC02FC82FA1E" italics="true" pageId="3" pageNumber="15">Hippo- sideros</emphasis>
(
<tableCitation id="C6D4B7EB8E14FF84FCBFEC25FCE9FA1F" box="[785,873,1396,1423]" captionStart="TABLE" captionStartId="4.[86,99,647,674]" captionTargetPageId="4" captionText="TABLE 1. External measurements (in mm) and body mass (in g) of four species of Hipposideros recently collected in Sulawesi. Used abbreviations: FA — forearm length; 5MET, 4MET, 3MET — fifth, fourth, and third metacarpal lengths; 5MET/FA× 100 — % length of fifth metacarpal relative to forearm length; 3D1P, 3D2P — length of first and second phalanges of the third digit; 3D1P/3MET × 100 — % length of first phalanx of the third digit relative to third metacarpal length; TAIL — tail length; HF — foot; TIBIA — tibia; E — ear and WGT — body mass. Sample sizes differing from those reported under n are given in parentheses. Mean and SD are included when sample size is ≥ 3" httpUri="http://table.plazi.org/id/DF29D2D88E13FF83FFF8EBD6FD8AFC83" pageId="3" pageNumber="15" tableUuid="DF29D2D88E13FF83FFF8EBD6FD8AFC83">Table 1</tableCitation>
). Pelage colour is uniform- ly reddish-brown. The wing and tail membranes are dark brown to blackish, as are the ears, noseleaf and lateral leaflets. The tail is very short, on average slightly exceeding the tibiae in length. In the wing (based on four alcohol preserved specimens), the fifth metarcarpal is between 5.710.3% shorter than the fourth ([4MET-5MET]/4MET) and is 7075% of forearm
</paragraph>
<caption id="DF29D2D88E13FF83FFF8EBD6FD8AFC83" ID-Table-UUID="DF29D2D88E13FF83FFF8EBD6FD8AFC83" httpUri="http://table.plazi.org/id/DF29D2D88E13FF83FFF8EBD6FD8AFC83" pageId="4" pageNumber="16" targetBox="[86,1653,812,1244]" targetIsTable="true" targetPageId="4">
<paragraph id="8BE982508E13FF83FFF8EBD6FD8AFC83" blockId="4.[86,1654,647,787]" pageId="4" pageNumber="16">
TABLE 1. External measurements (in mm) and body mass (in g) of four species of
<taxonomicName id="4C56F9D38E13FF83FCC1EBD6FC6FFD32" authorityName="Bates &amp; Rossiter &amp; Suyanto &amp; Kingston" authorityYear="2007" box="[879,1007,647,674]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="4" pageNumber="16" phylum="Chordata" rank="genus">
<emphasis id="B9225E428E13FF83FCC1EBD6FC6FFD32" box="[879,1007,647,674]" italics="true" pageId="4" pageNumber="16">Hipposideros</emphasis>
</taxonomicName>
recently collected in Sulawesi. Used abbreviations: FA — forearm length; 5MET, 4MET, 3MET — fifth, fourth, and third metacarpal lengths; 5MET/FA × 100 — % length of fifth metacarpal relative to forearm length; 3D1P, 3D2P — length of first and second phalanges of the third digit; 3D1P/3MET × 100 — % length of first phalanx of the third digit relative to third metacarpal length; TAIL — tail length; HF — foot; TIBIA — tibia; E — ear and WGT — body mass. Sample sizes differing from those reported under
<emphasis id="B9225E428E13FF83FA9AEB8DFAC0FD67" box="[1332,1344,732,759]" italics="true" pageId="4" pageNumber="16">n</emphasis>
are given in parentheses. Mean and SD are included when sample size is ≥ 3
</paragraph>
</caption>
<footnote id="E84D9E5E8E13FF83FFF8EA69FD8CFB4C" pageId="4" pageNumber="16">
<paragraph id="8BE982508E13FF83FFF8EA69FD8CFB4C" pageId="4" pageNumber="16">
<table id="F95670F08E130078FFF8EA7DF9F5FB4C" box="[86,1653,812,1244]" gridcols="12" gridrows="7" pageId="4" pageNumber="16">
<tr id="356680128E130078FFF8EA7DF9F5FCCD" box="[86,1653,812,861]" gridrow="0" pageId="4" pageNumber="16">
<th id="76B7E96E8E130078FFF8EA7DFF14FCCD" box="[86,148,812,861]" gridcol="0" gridrow="0" pageId="4" pageNumber="16">
<emphasis id="B9225E428E13FF83FFF8EA69FFE2FCC3" box="[86,98,824,851]" italics="true" pageId="4" pageNumber="16">n</emphasis>
sex
</th>
<th id="76B7E96E8E130078FF07EA7DFE8EFCCD" box="[169,270,812,861]" gridcol="1" gridrow="0" pageId="4" pageNumber="16">FA</th>
<th id="76B7E96E8E130078FEB0EA7DFEFFFCCD" box="[286,383,812,861]" gridcol="2" gridrow="0" pageId="4" pageNumber="16">5MET</th>
<th id="76B7E96E8E130078FE3EEA7DFE70FCCD" box="[400,496,812,861]" gridcol="3" gridrow="0" pageId="4" pageNumber="16">4MET</th>
<th id="76B7E96E8E130078FDAFEA7DFDE2FCCD" box="[513,610,812,861]" gridcol="4" gridrow="0" pageId="4" pageNumber="16">3MET</th>
<th id="76B7E96E8E130078FDDCEA7DFD57FCCD" box="[626,727,812,861]" gridcol="5" gridrow="0" pageId="4" pageNumber="16">5MET/FA × 100</th>
<th id="76B7E96E8E130078FD4AEA7DFBBEFCCD" box="[740,1086,812,861]" gridcol="6" gridrow="0" pageId="4" pageNumber="16">3D1P 3D2P 3D1P/3MET × 100</th>
<th id="76B7E96E8E130078FBE1EA7DFB2FFCCD" box="[1103,1199,812,861]" gridcol="7" gridrow="0" pageId="4" pageNumber="16">TAIL</th>
<th id="76B7E96E8E130078FB62EA7DFA95FCCD" box="[1228,1301,812,861]" gridcol="8" gridrow="0" pageId="4" pageNumber="16">HF</th>
<th id="76B7E96E8E130078FA9FEA7DFA12FCCD" box="[1329,1426,812,861]" gridcol="9" gridrow="0" pageId="4" pageNumber="16">TIBIA</th>
<th id="76B7E96E8E130078FA0DEA7DF983FCCD" box="[1443,1539,812,861]" gridcol="10" gridrow="0" pageId="4" pageNumber="16">E</th>
<th id="76B7E96E8E130078F98EEA7DF9F5FCCD" box="[1568,1653,812,861]" gridcol="11" gridrow="0" pageId="4" pageNumber="16">WGT</th>
</tr>
<tr id="356680128E130078FFF8EA39F9F5FC2C" box="[86,1653,872,956]" gridrow="1" pageId="4" pageNumber="16">
<th id="76B7E96E8E130078FFF8EA39FF14FC2C" box="[86,148,872,956]" gridcol="0" gridrow="1" pageId="4" pageNumber="16">
2 Ƌ
<specimenCount id="9D5049D98E13FF83FFF8EAF1FF09FC2C" box="[86,137,928,956]" pageId="4" pageNumber="16" type="female">2 ♀</specimenCount>
</th>
<td id="76B7E96E8E130078FF07EA39FE8EFC2C" box="[169,270,872,956]" gridcol="1" gridrow="1" pageId="4" pageNumber="16">40.5, 40.6 40.7, 42.7</td>
<td id="76B7E96E8E130078FEB0EA39FEFFFC2C" box="[286,383,872,956]" gridcol="2" gridrow="1" pageId="4" pageNumber="16">29.5, 30.4 28.7, 30.3</td>
<td id="76B7E96E8E130078FE3EEA39FE70FC2C" box="[400,496,872,956]" gridcol="3" gridrow="1" pageId="4" pageNumber="16">32.0, 32.2 32.0, 32.9</td>
<td id="76B7E96E8E130078FDAFEA39FDE2FC2C" box="[513,610,872,956]" gridcol="4" gridrow="1" pageId="4" pageNumber="16">32.0, 32.7 32.3, 33.1</td>
<td id="76B7E96E8E130078FDDCEA39FD57FC2C" box="[626,727,872,956]" gridcol="5" gridrow="1" pageId="4" pageNumber="16">72.8, 74.8 70.5, 71.0</td>
<td id="76B7E96E8E130078FD4AEA39FBBEFC2C" box="[740,1086,872,956]" gridcol="6" gridrow="1" pageId="4" pageNumber="16">
<taxonomicName id="4C56F9D38E13FF83FD50EA39FC4DFC13" authorityName="Bates &amp; Rossiter &amp; Suyanto &amp; Kingston" authorityYear="2007" box="[766,973,872,899]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="4" pageNumber="16" phylum="Chordata" rank="species" species="boeadii">
<emphasis id="B9225E428E13FF83FD50EA39FC4DFC13" box="[766,973,872,899]" italics="true" pageId="4" pageNumber="16">Hipposideros boeadii</emphasis>
</taxonomicName>
13.7, 13.9 14.1, 14.1 43.2, 44.4 13.6, 13.9 14.0, 14.4 43.3, 43.6
</td>
<td id="76B7E96E8E130078FBE1EA39FB2FFC2C" box="[1103,1199,872,956]" gridcol="7" gridrow="1" pageId="4" pageNumber="16">19.5, 21.0 16.0, 21.0</td>
<td id="76B7E96E8E130078FB62EA39FA95FC2C" box="[1228,1301,872,956]" gridcol="8" gridrow="1" pageId="4" pageNumber="16">6.9, 7.1 8.6, 8.7</td>
<td id="76B7E96E8E130078FA9FEA39FA12FC2C" box="[1329,1426,872,956]" gridcol="9" gridrow="1" pageId="4" pageNumber="16">17.0, 18.0 17.0, 18.3</td>
<td id="76B7E96E8E130078FA0DEA39F983FC2C" box="[1443,1539,872,956]" gridcol="10" gridrow="1" pageId="4" pageNumber="16">17.6, 18.0 16.5, 18.0</td>
<td id="76B7E96E8E130078F98EEA39F9F5FC2C" box="[1568,1653,872,956]" gridcol="11" gridrow="1" pageId="4" pageNumber="16">6.8, 7.5 8.5, 8.5</td>
</tr>
<tr id="356680128E130078FFF8EA92F9F5FC4E" box="[86,1653,963,990]" gridrow="2" pageId="4" pageNumber="16">
<th id="76B7E96E8E130078FFF8EA92F9F5FC4E" box="[86,1653,963,990]" colspan="12" colspanRight="11" gridcol="0" gridrow="2" pageId="4" pageNumber="16">
<taxonomicName id="4C56F9D38E13FF83FC94EA92FC14FC4E" box="[826,916,963,990]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="4" pageNumber="16" phylum="Chordata" rank="species" species="papua">
<emphasis id="B9225E428E13FF83FC94EA92FC14FC4E" box="[826,916,963,990]" italics="true" pageId="4" pageNumber="16">H. papua</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="356680128E130078FFF8EAB7F9F5FBA9" box="[86,1653,998,1081]" gridrow="3" pageId="4" pageNumber="16">
<th id="76B7E96E8E130078FFF8EAB7FF14FBA9" box="[86,148,998,1081]" gridcol="0" gridrow="3" pageId="4" pageNumber="16">
2 Ƌ*
<specimenCount id="9D5049D98E13FF83FFF8ED5CFF09FBB8" box="[86,137,1037,1064]" pageId="4" pageNumber="16" type="female">5 ♀</specimenCount>
*
</th>
<td id="76B7E96E8E130078FF07EAB7FE8EFBA9" box="[169,270,998,1081]" gridcol="1" gridrow="3" pageId="4" pageNumber="16">49.6, 52.6 52.0, 1.4 50.153.6</td>
<td id="76B7E96E8E130078FEB0EAB7FEFFFBA9" box="[286,383,998,1081]" gridcol="2" gridrow="3" pageId="4" pageNumber="16"> (2) 32.7, 35.2</td>
<td id="76B7E96E8E130078FE3EEAB7FE70FBA9" box="[400,496,998,1081]" gridcol="3" gridrow="3" pageId="4" pageNumber="16"> (2) 35.2, 40.5</td>
<td id="76B7E96E8E130078FDAFEAB7FDE2FBA9" box="[513,610,998,1081]" gridcol="4" gridrow="3" pageId="4" pageNumber="16"> (2) 37.7, 41.7</td>
<td id="76B7E96E8E130078FDDCEAB7FD57FBA9" box="[626,727,998,1081]" gridcol="5" gridrow="3" pageId="4" pageNumber="16"> (2) 65.1, 65.6</td>
<td id="76B7E96E8E130078FD4AEAB7FBBEFBA9" box="[740,1086,998,1081]" gridcol="6" gridrow="3" pageId="4" pageNumber="16"> (2) (2) (2) 15.7, 17.2 17.1, 19.1 45.4, 45.9</td>
<td id="76B7E96E8E130078FBE1EAB7FB2FFBA9" box="[1103,1199,998,1081]" gridcol="7" gridrow="3" pageId="4" pageNumber="16"> (1) 27.7</td>
<td id="76B7E96E8E130078FB62EAB7FA95FBA9" box="[1228,1301,998,1081]" gridcol="8" gridrow="3" pageId="4" pageNumber="16"> (2) 6.5, 7.7</td>
<td id="76B7E96E8E130078FA9FEAB7FA12FBA9" box="[1329,1426,998,1081]" gridcol="9" gridrow="3" pageId="4" pageNumber="16"> (2) 18.2, 20.7</td>
<td id="76B7E96E8E130078FA0DEAB7F983FBA9" box="[1443,1539,998,1081]" gridcol="10" gridrow="3" pageId="4" pageNumber="16"> (2) 10.5, 10.5</td>
<td id="76B7E96E8E130078F98EEAB7F9F5FBA9" box="[1568,1653,998,1081]" gridcol="11" gridrow="3" pageId="4" pageNumber="16"> (2) 9.3, 10.0</td>
</tr>
<tr id="356680128E130078FFF8ED10F9F5FBE8" box="[86,1653,1089,1144]" gridrow="4" pageId="4" pageNumber="16">
<th id="76B7E96E8E130078FFF8ED10FF14FBE8" box="[86,148,1089,1144]" gridcol="0" gridrow="4" pageId="4" pageNumber="16">
<specimenCount id="9D5049D98E13FF83FFF8ED0CFF09FBE8" box="[86,137,1117,1144]" pageId="4" pageNumber="16" type="female">2 ♀</specimenCount>
</th>
<td id="76B7E96E8E130078FF07ED10FE8EFBE8" box="[169,270,1089,1144]" gridcol="1" gridrow="4" pageId="4" pageNumber="16">50.8, 52.1</td>
<td id="76B7E96E8E130078FEB0ED10FEFFFBE8" box="[286,383,1089,1144]" gridcol="2" gridrow="4" pageId="4" pageNumber="16">33.4, 33.6</td>
<td id="76B7E96E8E130078FE3EED10FE70FBE8" box="[400,496,1089,1144]" gridcol="3" gridrow="4" pageId="4" pageNumber="16">38.6, 39.3</td>
<td id="76B7E96E8E130078FDAFED10FDE2FBE8" box="[513,610,1089,1144]" gridcol="4" gridrow="4" pageId="4" pageNumber="16">39.0, 39.9</td>
<td id="76B7E96E8E130078FDDCED10FD57FBE8" box="[626,727,1089,1144]" gridcol="5" gridrow="4" pageId="4" pageNumber="16">64.1, 66.1</td>
<td id="76B7E96E8E130078FD4AED10FBBEFBE8" box="[740,1086,1089,1144]" gridcol="6" gridrow="4" pageId="4" pageNumber="16">
<taxonomicName id="4C56F9D38E13FF83FC81ED10FC22FBCC" authorityName="Cantor" authorityYear="1846" box="[815,930,1089,1116]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="4" pageNumber="16" phylum="Chordata" rank="species" species="galeritus">
<emphasis id="B9225E428E13FF83FC81ED10FC22FBCC" box="[815,930,1089,1116]" italics="true" pageId="4" pageNumber="16">H. galeritus</emphasis>
</taxonomicName>
15.4, 16.0 19.6, 20.5 50.3, 51.4
</td>
<td id="76B7E96E8E130078FBE1ED10FB2FFBE8" box="[1103,1199,1089,1144]" gridcol="7" gridrow="4" pageId="4" pageNumber="16">39.0, 40.0</td>
<td id="76B7E96E8E130078FB62ED10FA95FBE8" box="[1228,1301,1089,1144]" gridcol="8" gridrow="4" pageId="4" pageNumber="16">6.0, 6.1</td>
<td id="76B7E96E8E130078FA9FED10FA12FBE8" box="[1329,1426,1089,1144]" gridcol="9" gridrow="4" pageId="4" pageNumber="16">20.0, 21.0</td>
<td id="76B7E96E8E130078FA0DED10F983FBE8" box="[1443,1539,1089,1144]" gridcol="10" gridrow="4" pageId="4" pageNumber="16">14.0, 14.5</td>
<td id="76B7E96E8E130078F98EED10F9F5FBE8" box="[1568,1653,1089,1144]" gridcol="11" gridrow="4" pageId="4" pageNumber="16">7.0, 8.0</td>
</tr>
<tr id="356680128E130078FFF8EDD1F9F5FB27" box="[86,1653,1152,1207]" gridrow="5" pageId="4" pageNumber="16">
<th id="76B7E96E8E130078FFF8EDD1FF14FB27" box="[86,148,1152,1207]" gridcol="0" gridrow="5" pageId="4" pageNumber="16">
<specimenCount id="9D5049D98E13FF83FFF8EDCDFF09FB27" box="[86,137,1180,1207]" pageId="4" pageNumber="16" type="female">1 ♀</specimenCount>
</th>
<td id="76B7E96E8E130078FF07EDD1FE8EFB27" box="[169,270,1152,1207]" gridcol="1" gridrow="5" pageId="4" pageNumber="16">44.8</td>
<td id="76B7E96E8E130078FEB0EDD1FEFFFB27" box="[286,383,1152,1207]" gridcol="2" gridrow="5" pageId="4" pageNumber="16">28.8</td>
<td id="76B7E96E8E130078FE3EEDD1FE70FB27" box="[400,496,1152,1207]" gridcol="3" gridrow="5" pageId="4" pageNumber="16">32.3</td>
<td id="76B7E96E8E130078FDAFEDD1FDE2FB27" box="[513,610,1152,1207]" gridcol="4" gridrow="5" pageId="4" pageNumber="16">34.3</td>
<td id="76B7E96E8E130078FDDCEDD1FD57FB27" box="[626,727,1152,1207]" gridcol="5" gridrow="5" pageId="4" pageNumber="16">64.3</td>
<td id="76B7E96E8E130078FD4AEDD1FBBEFB27" box="[740,1086,1152,1207]" gridcol="6" gridrow="5" pageId="4" pageNumber="16">
<taxonomicName id="4C56F9D38E13FF83FC9FEDD1FC20FB0B" box="[817,928,1152,1179]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="4" pageNumber="16" phylum="Chordata" rank="species" species="cervinus">
<emphasis id="B9225E428E13FF83FC9FEDD1FC20FB0B" box="[817,928,1152,1179]" italics="true" pageId="4" pageNumber="16">H. cervinus</emphasis>
</taxonomicName>
15.0 17.2 50.3
</td>
<td id="76B7E96E8E130078FBE1EDD1FB2FFB27" box="[1103,1199,1152,1207]" gridcol="7" gridrow="5" pageId="4" pageNumber="16">31.0</td>
<td id="76B7E96E8E130078FB62EDD1FA95FB27" box="[1228,1301,1152,1207]" gridcol="8" gridrow="5" pageId="4" pageNumber="16">6.0</td>
<td id="76B7E96E8E130078FA9FEDD1FA12FB27" box="[1329,1426,1152,1207]" gridcol="9" gridrow="5" pageId="4" pageNumber="16">17.0</td>
<td id="76B7E96E8E130078FA0DEDD1F983FB27" box="[1443,1539,1152,1207]" gridcol="10" gridrow="5" pageId="4" pageNumber="16">14.0</td>
<td id="76B7E96E8E130078F98EEDD1F9F5FB27" box="[1568,1653,1152,1207]" gridcol="11" gridrow="5" pageId="4" pageNumber="16">5.0</td>
</tr>
<tr id="356680128E130078FFF8ED95F9F5FB4C" box="[86,1653,1220,1244]" gridrow="6" pageId="4" pageNumber="16">
<th id="76B7E96E8E130078FFF8ED95F9F5FB4C" box="[86,1653,1220,1244]" colspan="12" colspanRight="11" gridcol="0" gridrow="6" pageId="4" pageNumber="16">* — includes data from Hill and Rozendaal (1989)</th>
</tr>
</table>
</paragraph>
</footnote>
<paragraph id="8BE982508E12FF82FF2AE9F4FD04FCE0" blockId="5.[132,645,165,881]" pageId="5" pageNumber="17">
length. The first phalanx of the third digit is short, between 43.2 and 44.4% of third metacarpal length. The ear is 16.518.0 mm in length, with a rounded tip and a slight concavity on the posterior border. The an- titragus has a fold on its upper border (
<figureCitation id="136D9ED58E12FF82FF22E825FF51FE1E" box="[140,209,372,398]" captionStart="FIG" captionStartId="3.[686,699,648,669]" captionTargetBox="[683,1201,161,625]" captionTargetId="figure@3.[683,1202,159,625]" captionTargetPageId="3" captionText="FIG. 1. Hipposideros boeadii from Rawa Aopa Watumohai National Park, South-East Sulawesi" figureDoi="http://doi.org/10.5281/zenodo.3944950" httpUri="https://zenodo.org/record/3944950/files/figure.png" pageId="5" pageNumber="17">Fig. 1</figureCitation>
); when flattened, the fold appears as a concavity which divides the antitragus in to two parts. In the noseleaf (
<figureCitation id="136D9ED58E12FF82FE46E8E8FDC9FE43" box="[488,585,441,467]" captionStart="FIG" captionStartId="5.[132,145,1490,1511]" captionTargetBox="[129,652,940,1452]" captionTargetId="figure@5.[355,658,938,1463]" captionTargetPageId="5" captionText="FIG. 2. Noseleaves of four species of Hipposideros (with hairs omitted). A — H. boeadii (holotype), M28253, Rawa Aopa Watumohai National Park, South-East Sulawesi; B — H. papua, M16951, Biak Island, Papua Province; C — H. galeritus, M28250, Kakenauwe Nature Reserve, Buton Island, South- East Sulawesi; D — H. cervinus, M28249, Lambasango Wildlife Reserve, Buton Island, South- East Sulawesi. Scale = 5 mm" figureDoi="http://doi.org/10.5281/zenodo.3944954" httpUri="https://zenodo.org/record/3944954/files/figure.png" pageId="5" pageNumber="17">Fig. 2A</figureCitation>
), the internarial septum is slightly swollen anteri- orly and the narial lappets have a smoothly rounded outline. The intermediate leaf is simple and slightly inflated medially. It is subequal in width to the anterior leaf and distinctly narrower than the posterior one. Although its medial part is essentially naked, it has a number of hairs of various lengths on the lateral lappets. The anterior noseleaf is a simple horseshoe shape with a small notch in the mid-part of the anterior border. The upper part of the anterior leaf
</paragraph>
<paragraph id="8BE982508E12FF82FD00E9F2FD7AFE5F" blockId="5.[686,1199,163,1730]" pageId="5" pageNumber="17">is attached to the intermediate leaf. The posterior leaf has an evenly rounded upper margin and four well-defined concavities. A well-developed frontal sac is present in males on the forehead behind the posterior leaf; it is not present in female specimens. Small projections on the forehead above each eye are present in both males and females.</paragraph>
<paragraph id="8BE982508E12FF82FD7DE889FBA5FC3F" blockId="5.[686,1199,163,1730]" pageId="5" pageNumber="17">
There are two supplementary lateral leaflets (
<figureCitation id="136D9ED58E12FF82FCBAE8ABFCE7FD84" box="[788,871,506,532]" captionStart="FIG" captionStartId="3.[686,699,648,669]" captionTargetBox="[683,1201,161,625]" captionTargetId="figure@3.[683,1202,159,625]" captionTargetPageId="3" captionText="FIG. 1. Hipposideros boeadii from Rawa Aopa Watumohai National Park, South-East Sulawesi" figureDoi="http://doi.org/10.5281/zenodo.3944950" httpUri="https://zenodo.org/record/3944950/files/figure.png" pageId="5" pageNumber="17">Figs. 1</figureCitation>
and
<figureCitation id="136D9ED58E12FF82FC0DE8ABFC4BFD84" box="[931,971,506,532]" captionStart="FIG" captionStartId="5.[132,145,1490,1511]" captionTargetBox="[129,652,940,1452]" captionTargetId="figure@5.[355,658,938,1463]" captionTargetPageId="5" captionText="FIG. 2. Noseleaves of four species of Hipposideros (with hairs omitted). A — H. boeadii (holotype), M28253, Rawa Aopa Watumohai National Park, South-East Sulawesi; B — H. papua, M16951, Biak Island, Papua Province; C — H. galeritus, M28250, Kakenauwe Nature Reserve, Buton Island, South- East Sulawesi; D — H. cervinus, M28249, Lambasango Wildlife Reserve, Buton Island, South- East Sulawesi. Scale = 5 mm" figureDoi="http://doi.org/10.5281/zenodo.3944954" httpUri="https://zenodo.org/record/3944954/files/figure.png" pageId="5" pageNumber="17">2A</figureCitation>
). The first is about half the length of the anterior leaf and its upper part is attached to the posterior leaf. The second supplementary leaflet nearly reaches the midline anteriorly; there is an indication of the leaflet as far as the midline, although at this point the leaflet is attached completely to the muzzle. Its upper part stops about
<quantity id="4CAE2FB58E12FF82FCDCEA5DFC4EFCB6" box="[882,974,780,806]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="5" pageNumber="17" unit="mm" value="1.3">1.3 mm</quantity>
below the eye and is not attached to the posterior leaf. Both supplementary leaflets are lightly covered with bristly hairs of different lengths (these have been omitted from
<figureCitation id="136D9ED58E12FF82FC61EAC4FB97FC3F" box="[975,1047,917,943]" captionStart="FIG" captionStartId="5.[132,145,1490,1511]" captionTargetBox="[129,652,940,1452]" captionTargetId="figure@5.[355,658,938,1463]" captionTargetPageId="5" captionText="FIG. 2. Noseleaves of four species of Hipposideros (with hairs omitted). A — H. boeadii (holotype), M28253, Rawa Aopa Watumohai National Park, South-East Sulawesi; B — H. papua, M16951, Biak Island, Papua Province; C — H. galeritus, M28250, Kakenauwe Nature Reserve, Buton Island, South- East Sulawesi; D — H. cervinus, M28249, Lambasango Wildlife Reserve, Buton Island, South- East Sulawesi. Scale = 5 mm" figureDoi="http://doi.org/10.5281/zenodo.3944954" httpUri="https://zenodo.org/record/3944954/files/figure.png" pageId="5" pageNumber="17">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="8BE982508E12FF82FD7DEAE6FCA3FA43" blockId="5.[686,1199,163,1730]" pageId="5" pageNumber="17">
The skull has a condylo-canine length of
<quantity id="4CAE2FB58E12FF82FD00EA88FCD9FC64" box="[686,857,985,1012]" metricMagnitude="-2" metricUnit="m" metricValue="1.535" metricValueMax="1.55" metricValueMin="1.52" pageId="5" pageNumber="17" unit="mm" value="15.35" valueMax="15.5" valueMin="15.2">15.215.5 mm</quantity>
(
<tableCitation id="C6D4B7EB8E12FF82FCC6EA88FC44FC63" box="[872,964,985,1011]" captionStart="TABLE" captionStartId="9.[89,102,641,668]" captionText="TABLE 2. Cranial and dental measurements (in mm) of four species of Hipposideros recently collected in Sulawesi. Used abbreviations: GTL — greatest length of skull; CBL — condylobasal length; CCL — condylo-canine length; ZB — zygomatic breadth; BB — breadth of braincase; MW — mastoid width; ZB/MW × 100 — % breadth of zygomata relative to mastoid breadth; RW — rostral width; PC — postorbital constriction; CM3 — upper toothrow length; CM3/CCL × 100 — % length of upper toothrow relative to condylo-canine length; M3M3 — palatal breadth; CM — lower toothrow length; M — mandible length. Sample sizes 3 differing from those reported under n are given in parentheses. Mean and SD are included when sample size is ≥ 3" pageId="5" pageNumber="17">Table 2</tableCitation>
). The zygomata are robust and broad, considerably exceed- ing mastoid width (
<figureCitation id="136D9ED58E12FF82FC3EED4CFC6EFBA8" box="[912,1006,1053,1080]" captionStart="FIG" captionStartId="6.[130,143,1651,1672]" captionTargetBox="[288,1018,552,1640]" captionTargetId="figure@6.[244,1035,543,1637]" captionTargetPageId="6" captionText="FIG. 3. Dorsal (above) and ventral (below) views of the skull and dorsal view of the right half mandible of two species of Hipposideros. A — H. boeadii (holotype), M28253, Rawa Aopa Watumohai National Park, South- East Sulawesi, and B — H. papua, M16951, Biak Island, Papua Province. Scale = 5 mm" figureDoi="http://doi.org/10.5281/zenodo.3944958" httpUri="https://zenodo.org/record/3944958/files/figure.png" pageId="5" pageNumber="17">Fig. 3A</figureCitation>
). There is a well developed dorsal projection on the jugal bone; the upper border of which is evenly rounded, almost semi-circular in outline (
<figureCitation id="136D9ED58E12FF82FD18EDF7FC97FB51" box="[694,791,1190,1217]" captionStart="FIG" captionStartId="7.[130,143,1486,1507]" captionTargetBox="[146,639,341,1457]" captionTargetId="figure@7.[125,645,615,920]" captionTargetPageId="7" captionText="FIG. 4. Lateral view of the skulls and mandibles of four species of Hipposideros. A — H. boeadii (holo- type), M28253, Rawa Aopa Watumohai National Park, South-East Sulawesi; B — H. papua, M16951, Biak Island, Papua Province; C — H. galeritus, M28250, Kakenauwe Nature Reserve, Buton Island, South-East Sulawesi; D — H. cervinus, M28249, Lambasango Wildlife Reserve, Buton Island, South- East Sulawesi. Scale = 5 mm" figureDoi="http://doi.org/10.5281/zenodo.3944962" httpUri="https://zenodo.org/record/3944962/files/figure.png" pageId="5" pageNumber="17">Fig. 4A</figureCitation>
). The anterorbital foramen, which has a thin narrow bar of bone above it, is long, narrow and elliptical in shape. The rostrum is moderately inflated with a slight depression in the midline; the anterior chambers are about equally inflated to the postero-median chambers; the postero- lateral chambers are not greatly inflated (
<figureCitation id="136D9ED58E12FF82FD18ECE9FC96FA43" box="[694,790,1464,1491]" captionStart="FIG" captionStartId="8.[130,143,1626,1647]" captionTargetBox="[273,1107,833,1601]" captionTargetId="figure@8.[235,1126,829,1604]" captionTargetPageId="8" captionText="FIG. 5. Dorsal view of the rostrum of four species of Hipposideros. A — H. boeadii (holotype), M28253, Rawa Aopa Watumohai National Park, South-East Sulawesi; B — H. papua, M16951, Biak Island, Papua Province; C — H. galeritus, M28250, Kakenauwe Nature Reserve, Buton Island, South-East Sulawesi; D —" figureDoi="http://doi.org/10.5281/zenodo.3944964" httpUri="https://zenodo.org/record/3944964/files/figure.png" pageId="5" pageNumber="17">Fig. 5A</figureCitation>
).
</paragraph>
<caption id="DF29D2D88E12FF82FF2AEC83FD90F959" ID-DOI="http://doi.org/10.5281/zenodo.3944954" ID-Zenodo-Dep="3944954" httpUri="https://zenodo.org/record/3944954/files/figure.png" pageId="5" pageNumber="17" startId="5.[132,145,1490,1511]" targetBox="[129,652,940,1452]" targetPageId="5">
<paragraph id="8BE982508E12FF82FF2AEC83FD90F959" blockId="5.[132,645,1490,1738]" pageId="5" pageNumber="17">
FIG. 2. Noseleaves of four species of
<emphasis id="B9225E428E12FF82FDAAEC83FD04FA77" box="[516,644,1490,1511]" italics="true" pageId="5" pageNumber="17">Hipposideros</emphasis>
(with hairs omitted). A —
<taxonomicName id="4C56F9D38E12FF82FE0CECBEFD8EF993" authorityName="Bates &amp; Rossiter &amp; Suyanto &amp; Kingston" authorityYear="2007" box="[418,526,1518,1540]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="5" pageNumber="17" phylum="Chordata" rank="species" species="boeadii">
<emphasis id="B9225E428E12FF82FE0CECBEFD8EF993" box="[418,526,1518,1540]" italics="true" pageId="5" pageNumber="17">H. boeadii</emphasis>
</taxonomicName>
(
<typeStatus id="54ED3CF28E12FF82FD8FECBFFDFAF994" box="[545,634,1518,1540]" pageId="5" pageNumber="17" type="holotype">holotype</typeStatus>
), M28253,
<location id="8E89D48B8E12FF82FF44EF5AFD01F9B0" LSID="urn:lsid:plazi:treatment:03FF33468E15FF8BFD52EC28FCE7FBE7:8E89D48B8E12FF82FF44EF5AFD01F9B0" box="[234,641,1546,1568]" county="South-East Sulawesi" municipality="It" name="Rawa Aopa Watumohai National Park" pageId="5" pageNumber="17">Rawa Aopa Watumohai National Park</location>
,
<collectingCounty id="6288FADC8E12FF82FF2AEF76FECEF9AD" box="[132,334,1575,1597]" pageId="5" pageNumber="17">South-East Sulawesi</collectingCounty>
; B —
<emphasis id="B9225E428E12FF82FE3EEF76FE71F9AC" box="[400,497,1575,1596]" italics="true" pageId="5" pageNumber="17">
<taxonomicName id="4C56F9D38E12FF82FE3EEF76FE6DF9AC" box="[400,493,1575,1596]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="5" pageNumber="17" phylum="Chordata" rank="species" species="papua">H. papua</taxonomicName>
,
</emphasis>
M16951,
<location id="8E89D48B8E12FF82FDF9EF76FD04F9AD" LSID="urn:lsid:plazi:treatment:03FF33468E15FF8BFD52EC28FCE7FBE7:8E89D48B8E12FF82FDF9EF76FD04F9AD" box="[599,644,1575,1597]" country="Iceland" municipality="Papua Province" name="Biak" pageId="5" pageNumber="17">Biak</location>
<collectingCountry id="F341C2C08E12FF82FF2AEF12FF41F9C9" box="[132,193,1603,1625]" name="Iceland" pageId="5" pageNumber="17">Island</collectingCountry>
,
<collectingMunicipality id="6B8D182A8E12FF82FF62EF12FEE7F9C9" box="[204,359,1603,1625]" pageId="5" pageNumber="17">Papua Province</collectingMunicipality>
; C —
<taxonomicName id="4C56F9D38E12FF82FE04EF15FD9FF9C8" authorityName="Cantor" authorityYear="1846" box="[426,543,1603,1625]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="5" pageNumber="17" phylum="Chordata" rank="species" species="galeritus">
<emphasis id="B9225E428E12FF82FE04EF15FD9FF9C8" box="[426,543,1603,1625]" italics="true" pageId="5" pageNumber="17">H. galeritus</emphasis>
</taxonomicName>
, M28250,
<collectingMunicipality id="6B8D182A8E12FF82FF2AEF0EFE20F9E5" box="[132,416,1631,1653]" pageId="5" pageNumber="17">Kakenauwe Nature Reserve</collectingMunicipality>
,
<collectingCounty id="6288FADC8E12FF82FE01EF31FDB4F9E5" box="[431,564,1631,1653]" pageId="5" pageNumber="17">Buton Island</collectingCounty>
, South- East
<collectingRegion id="49924CB28E12FF82FF6DEF2AFEA1F901" box="[195,289,1659,1681]" country="Indonesia" name="Sulawesi Utara" pageId="5" pageNumber="17">Sulawesi</collectingRegion>
; D —
<taxonomicName id="4C56F9D38E12FF82FE23EF2DFD8FF901" box="[397,527,1660,1681]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="5" pageNumber="17" phylum="Chordata" rank="species" species="cervinus">
<emphasis id="B9225E428E12FF82FE23EF2DFD8FF901" box="[397,527,1660,1681]" italics="true" pageId="5" pageNumber="17">H. cervinus</emphasis>
</taxonomicName>
, M28249, Lambasango Wildlife Reserve, Buton Island, South- East
<collectingRegion id="49924CB28E12FF82FE89EFE5FE01F95A" box="[295,385,1716,1738]" country="Indonesia" name="Sulawesi Utara" pageId="5" pageNumber="17">Sulawesi</collectingRegion>
. Scale =
<quantity id="4CAE2FB58E12FF82FE77EFE5FD90F959" box="[473,528,1716,1738]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="17" unit="mm" value="5.0">5 mm</quantity>
</paragraph>
</caption>
<paragraph id="8BE982508E12FF82FD7DEC8BFB2EF952" blockId="5.[686,1199,163,1730]" pageId="5" pageNumber="17">
<taxonomicName id="4C56F9D38E12FF82FD7DEC8BFC81FA64" authority="The" authorityName="The" box="[723,769,1498,1524]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="5" pageNumber="17" phylum="Chordata" rank="genus">The</taxonomicName>
sagittal crest is well defined ante- riorly.
<collectingMunicipality id="6B8D182A8E12FF82FCA8ECACFC98F987" box="[774,792,1533,1559]" pageId="5" pageNumber="17">It</collectingMunicipality>
bifurcates into the supraorbital ridges which are indicated but not devel- oped.
<location id="8E89D48B8E12FF82FD5BEF10FCA3F9CB" LSID="urn:lsid:plazi:treatment:03FF33468E15FF8BFD52EC28FCE7FBE7:8E89D48B8E12FF82FD5BEF10FCA3F9CB" box="[757,803,1601,1627]" country="Iceland" county="Buton Island" municipality="Kakenauwe Nature Reserve" name="The" pageId="5" pageNumber="17">The</location>
postorbital constriction, although narrower than the rostrum, is not conspicu- ously constricted. The mastoid is inflated. The supraoccipital and foramen magnum
</paragraph>
<paragraph id="8BE982508E11FF81FF2CE9F5FCA3FE3C" blockId="6.[130,643,163,497]" lastBlockId="6.[686,1199,163,496]" pageId="6" pageNumber="18">
are directed downwards and only slightly backwards. The palate is emarginated ante- riorly to a point corresponding to the ante- rior border of the first upper molar (M
<superScript id="7C232F188E11FF81FDC0E854FDF8FE88" attach="left" box="[622,632,261,280]" fontSize="8" pageId="6" pageNumber="18">1</superScript>
) (
<figureCitation id="136D9ED58E11FF81FF25E87DFF6CFED6" box="[139,236,300,326]" captionStart="FIG" captionStartId="6.[130,143,1651,1672]" captionTargetBox="[288,1018,552,1640]" captionTargetId="figure@6.[244,1035,543,1637]" captionTargetPageId="6" captionText="FIG. 3. Dorsal (above) and ventral (below) views of the skull and dorsal view of the right half mandible of two species of Hipposideros. A — H. boeadii (holotype), M28253, Rawa Aopa Watumohai National Park, South- East Sulawesi, and B — H. papua, M16951, Biak Island, Papua Province. Scale = 5 mm" figureDoi="http://doi.org/10.5281/zenodo.3944958" httpUri="https://zenodo.org/record/3944958/files/figure.png" pageId="6" pageNumber="18">Fig. 3A</figureCitation>
). Posteriorly, it is emarginated to the mid-point of the third upper molar (M
<superScript id="7C232F188E11FF81FDC9E81BFDF1FECD" attach="left" box="[615,625,330,349]" fontSize="8" pageId="6" pageNumber="18">3</superScript>
). The anterior border of the mesopterygo- id space is essentially U-shaped and there is no bony projection (spicule) from the midpoint of the posterior palate. There is a shallow sphenoidal depression. The width of each cochlea exceeds the width between the cochleae. Each tympanic bulla covers less than half of its respective cochlea. Each half mandible has a robust horizontal ramus and a well developed coronoid process, which exceeds the lower canine in height (
<figureCitation id="136D9ED58E11FF81FD18E8C3FC96FE3C" box="[694,790,402,428]" captionStart="FIG" captionStartId="7.[130,143,1486,1507]" captionTargetBox="[146,639,341,1457]" captionTargetId="figure@7.[125,645,615,920]" captionTargetPageId="7" captionText="FIG. 4. Lateral view of the skulls and mandibles of four species of Hipposideros. A — H. boeadii (holo- type), M28253, Rawa Aopa Watumohai National Park, South-East Sulawesi; B — H. papua, M16951, Biak Island, Papua Province; C — H. galeritus, M28250, Kakenauwe Nature Reserve, Buton Island, South-East Sulawesi; D — H. cervinus, M28249, Lambasango Wildlife Reserve, Buton Island, South- East Sulawesi. Scale = 5 mm" figureDoi="http://doi.org/10.5281/zenodo.3944962" httpUri="https://zenodo.org/record/3944962/files/figure.png" pageId="6" pageNumber="18">Fig. 4A</figureCitation>
).
</paragraph>
<paragraph id="8BE982508E11FF81FD7DE8E5FB2EFE60" blockId="6.[686,1199,163,496]" pageId="6" pageNumber="18">
Upper toothrow length is
<quantity id="4CAE2FB58E11FF81FBBBE8E5FB2AFE5E" box="[1045,1194,436,462]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" metricValueMax="6.9" metricValueMin="6.7" pageId="6" pageNumber="18" unit="mm" value="6.8" valueMax="6.9" valueMin="6.7">6.76.9 mm</quantity>
. The upper incisor (I
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) is very small and
</paragraph>
<caption id="DF29D2D88E11FF81FF2CEF22FBC5F950" ID-DOI="http://doi.org/10.5281/zenodo.3944956" ID-Zenodo-Dep="3944956" httpUri="https://zenodo.org/record/3944956/files/figure.png" pageId="6" pageNumber="18" startId="6.[130,143,1651,1672]" targetBox="[288,1018,552,1640]" targetPageId="6">
<paragraph id="8BE982508E11FF81FF2CEF22FBC5F950" blockId="6.[130,1198,1650,1729]" pageId="6" pageNumber="18">
FIG. 3. Dorsal (above) and ventral (below) views of the skull and dorsal view of the right half mandible of two species of
<emphasis id="B9225E428E11FF81FF44EFDFFEF2F933" box="[234,370,1678,1699]" italics="true" pageId="6" pageNumber="18">
<taxonomicName id="4C56F9D38E11FF81FF44EFDFFEEEF933" box="[234,366,1678,1699]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="6" pageNumber="18" phylum="Chordata" rank="genus">Hipposideros</taxonomicName>
.
</emphasis>
A —
<taxonomicName id="4C56F9D38E11FF81FE01EFDEFD96F933" authorityName="Bates &amp; Rossiter &amp; Suyanto &amp; Kingston" authorityYear="2007" box="[431,534,1678,1700]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="6" pageNumber="18" phylum="Chordata" rank="species" species="boeadii">
<emphasis id="B9225E428E11FF81FE01EFDEFD96F933" box="[431,534,1678,1700]" italics="true" pageId="6" pageNumber="18">H. boeadii</emphasis>
</taxonomicName>
(holotype), M28253, Rawa Aopa Watumohai National Park, South- East
<collectingRegion id="49924CB28E11FF81FEB4EFFAFEF3F951" box="[282,371,1707,1729]" country="Indonesia" name="Sulawesi Utara" pageId="6" pageNumber="18">Sulawesi</collectingRegion>
, and B —
<emphasis id="B9225E428E11FF81FE72EFFAFDBDF950" box="[476,573,1707,1728]" italics="true" pageId="6" pageNumber="18">
<taxonomicName id="4C56F9D38E11FF81FE72EFFAFDB8F950" box="[476,568,1707,1728]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="6" pageNumber="18" phylum="Chordata" rank="species" species="papua">H. papua</taxonomicName>
,
</emphasis>
M16951, Biak Island,
<collectingRegion id="49924CB28E11FF81FCB2EFFAFC36F951" box="[796,950,1707,1729]" country="Indonesia" name="Papua Barat" pageId="6" pageNumber="18">Papua Province</collectingRegion>
. Scale =
<quantity id="4CAE2FB58E11FF81FBA0EFFBFBC5F950" box="[1038,1093,1706,1728]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="18" unit="mm" value="5.0">5 mm</quantity>
</paragraph>
</caption>
<paragraph id="8BE982508E10FF80FF2CE9F5FD02FE95" blockId="7.[130,642,160,262]" pageId="7" pageNumber="19">
wedge shaped. The upper canine (C
<superScript id="7C232F188E10FF80FD9EE9F1FDBAFF23" attach="left" box="[560,570,160,179]" fontSize="8" pageId="7" pageNumber="19">1</superScript>
) is of moderate size, with the faintest indication of a groove on its antero-lateral surface and
</paragraph>
<paragraph id="8BE982508E10FF80FD00E9F2FCFEFBA0" blockId="7.[686,1199,163,1072]" pageId="7" pageNumber="19">
with an ill-defined cingulum. The first up- per premolar (P
<superScript id="7C232F188E10FF80FCC0E990FCF8FF44" attach="left" box="[878,888,193,212]" fontSize="8" pageId="7" pageNumber="19">2</superScript>
) is minute and displaced externally (
<figureCitation id="136D9ED58E10FF80FC96E9B6FC1BFE91" box="[824,923,231,257]" captionStart="FIG" captionStartId="6.[130,143,1651,1672]" captionTargetBox="[288,1018,552,1640]" captionTargetId="figure@6.[244,1035,543,1637]" captionTargetPageId="6" captionText="FIG. 3. Dorsal (above) and ventral (below) views of the skull and dorsal view of the right half mandible of two species of Hipposideros. A — H. boeadii (holotype), M28253, Rawa Aopa Watumohai National Park, South- East Sulawesi, and B — H. papua, M16951, Biak Island, Papua Province. Scale = 5 mm" figureDoi="http://doi.org/10.5281/zenodo.3944958" httpUri="https://zenodo.org/record/3944958/files/figure.png" pageId="7" pageNumber="19">Fig. 3A</figureCitation>
). The canine (C
<superScript id="7C232F188E10FF80FBCAE9B2FBEEFF66" attach="left" box="[1124,1134,227,246]" fontSize="8" pageId="7" pageNumber="19">1</superScript>
) and second upper premolar (P
<superScript id="7C232F188E10FF80FC45E854FC75FE88" attach="left" box="[1003,1013,261,280]" fontSize="8" pageId="7" pageNumber="19">4</superScript>
) are almost in contact. All the cheekteeth are robust. The first (M
<superScript id="7C232F188E10FF80FCA1E819FC99FECB" attach="left" box="[783,793,328,347]" fontSize="8" pageId="7" pageNumber="19">1</superScript>
) and second upper molars (M
<superScript id="7C232F188E10FF80FB34E819FB24FECB" attach="left" box="[1178,1188,328,347]" fontSize="8" pageId="7" pageNumber="19">2</superScript>
) have the typical W-shaped cusp pattern with parastyle and metastyle more developed than the mesostyle; the paracone and metacone are about equal in size. The third upper molar (M
<superScript id="7C232F188E10FF80FC17E8A3FC43FD95" attach="left" box="[953,963,498,517]" fontSize="8" pageId="7" pageNumber="19">3</superScript>
) is reduced; it includes a parastyle and mesostyle and two commissures, with a vestige of a third commissure posteriorly. Both the lower incisors are tricuspidate. The second (I
<subScript id="17D280158E10FF80FB86EBDDFBB2FD0F" attach="right" box="[1064,1074,652,671]" fontSize="8" pageId="7" pageNumber="19">2</subScript>
) is partly situated behind the first (I
<subScript id="17D280158E10FF80FC4AEBFFFC6EFD51" attach="right" box="[996,1006,686,705]" fontSize="8" pageId="7" pageNumber="19">1</subScript>
). The lower canine (C
<subScript id="17D280158E10FF80FCA4EB81FC94FD73" attach="left" box="[778,788,720,739]" fontSize="8" pageId="7" pageNumber="19">1</subScript>
) is not especially tall (
<figureCitation id="136D9ED58E10FF80FB93EB93FB21FD4C" box="[1085,1185,706,732]" captionStart="FIG" captionStartId="7.[130,143,1486,1507]" captionTargetBox="[146,639,341,1457]" captionTargetId="figure@7.[125,645,615,920]" captionTargetPageId="7" captionText="FIG. 4. Lateral view of the skulls and mandibles of four species of Hipposideros. A — H. boeadii (holo- type), M28253, Rawa Aopa Watumohai National Park, South-East Sulawesi; B — H. papua, M16951, Biak Island, Papua Province; C — H. galeritus, M28250, Kakenauwe Nature Reserve, Buton Island, South-East Sulawesi; D — H. cervinus, M28249, Lambasango Wildlife Reserve, Buton Island, South- East Sulawesi. Scale = 5 mm" figureDoi="http://doi.org/10.5281/zenodo.3944962" httpUri="https://zenodo.org/record/3944962/files/figure.png" pageId="7" pageNumber="19">Fig. 4A</figureCitation>
). The first lower premolar (P
<subScript id="17D280158E10FF80FC54EBA3FB84FC95" attach="left" box="[1018,1028,754,773]" fontSize="8" pageId="7" pageNumber="19">2</subScript>
) is about two thirds the crown area of the second (P
<subScript id="17D280158E10FF80FB3DEA45FB1DFCB7" attach="left" box="[1171,1181,788,807]" fontSize="8" pageId="7" pageNumber="19">4</subScript>
). The second lower molar (M
<subScript id="17D280158E10FF80FC52EA67FB86FCD9" attach="left" box="[1020,1030,822,841]" fontSize="8" pageId="7" pageNumber="19">2</subScript>
) very slightly exceeds the first (M
<subScript id="17D280158E10FF80FC00EA09FC38FCFB" attach="left" box="[942,952,856,875]" fontSize="8" pageId="7" pageNumber="19">1</subScript>
) in crown area; in the third (M
<subScript id="17D280158E10FF80FCE5EA2BFCD5FC1D" attach="left" box="[843,853,890,909]" fontSize="8" pageId="7" pageNumber="19">3</subScript>
), the talonid is reduced in size. The baculum is small (
<quantity id="4CAE2FB58E10FF80FBAAEADCFBE2FC38" box="[1028,1122,909,936]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="7" pageNumber="19" unit="mm" value="1.9">1.9 mm</quantity>
in to- tal length) with a horseshoe shaped tip, short shaft and a broad base, which is hollowed underneath and with a small keel ventrally (
<figureCitation id="136D9ED58E10FF80FC89ED44FCEFFBA0" box="[807,879,1045,1072]" captionStart="FIG" captionStartId="10.[130,143,513,534]" captionTargetBox="[183,574,164,492]" captionTargetId="figure@10.[126,647,159,483]" captionTargetPageId="10" captionText="FIG. 6. Baculum of H. boeadii, M22937, Rawa Aopa Watumohai National Park, South-East Sulawesi. Scale = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3944966" httpUri="https://zenodo.org/record/3944966/files/figure.png" pageId="7" pageNumber="19">Fig. 6</figureCitation>
).
</paragraph>
<paragraph id="8BE982508E10FF80FD00ED33FCCAFBEC" blockId="7.[686,1199,1122,1453]" box="[686,842,1122,1148]" pageId="7" pageNumber="19">
<emphasis id="B9225E428E10FF80FD00ED33FCCAFBEC" box="[686,842,1122,1148]" italics="true" pageId="7" pageNumber="19">Echolocation</emphasis>
</paragraph>
<paragraph id="8BE982508E10FF80FD7DEDD5FBE5FA3D" blockId="7.[686,1199,1122,1453]" pageId="7" pageNumber="19">
The echolocation calls of
<taxonomicName id="4C56F9D38E10FF80FBABEDD5FB02FB0E" authorityName="Bates &amp; Rossiter &amp; Suyanto &amp; Kingston" authorityYear="2007" box="[1029,1154,1156,1182]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="7" pageNumber="19" phylum="Chordata" rank="species" species="boeadii">
<emphasis id="B9225E428E10FF80FBABEDD5FB02FB0E" box="[1029,1154,1156,1182]" italics="true" pageId="7" pageNumber="19">H. boeadii</emphasis>
</taxonomicName>
are typical of the
<taxonomicName id="4C56F9D38E10FF80FCE3EDF4FB83FB2F" box="[845,1027,1189,1215]" class="Mammalia" family="Hipposideridae" kingdom="Animalia" order="Chiroptera" pageId="7" pageNumber="19" phylum="Chordata" rank="family">Hipposideridae</taxonomicName>
, with most energy in a constant frequency (CF) component of approximately 2.54.0 ms, terminating in a steep frequency modulated tail (1.01.5 ms duration). Mean value for the frequency of maximum energy of the CF portion of the call is 116.6 kHz, with a range of 114.2118.3 kHz (
<emphasis id="B9225E428E10FF80FBA6ECC5FB97FA3D" box="[1032,1047,1428,1453]" italics="true" pageId="7" pageNumber="19">n</emphasis>
= 13).
</paragraph>
</subSubSection>
<caption id="DF29D2D88E10FF80FF2CEC9FFD8EF955" ID-DOI="http://doi.org/10.5281/zenodo.3944960" ID-Zenodo-Dep="3944960" httpUri="https://zenodo.org/record/3944960/files/figure.png" pageId="7" pageNumber="19" startId="7.[130,143,1486,1507]" targetBox="[146,639,341,1457]" targetPageId="7">
<paragraph id="8BE982508E10FF80FF2CEC9FFD8EF955" blockId="7.[130,642,1486,1734]" pageId="7" pageNumber="19">
FIG. 4. Lateral view of the skulls and mandibles of four species of
<emphasis id="B9225E428E10FF80FEB9ECBBFE1FFA6F" box="[279,415,1514,1535]" italics="true" pageId="7" pageNumber="19">
<taxonomicName id="4C56F9D38E10FF80FEB9ECBBFE1BFA6F" box="[279,411,1514,1535]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="7" pageNumber="19" phylum="Chordata" rank="genus">Hipposideros</taxonomicName>
.
</emphasis>
A —
<taxonomicName id="4C56F9D38E10FF80FE74ECBAFDC1FA6F" authorityName="Bates &amp; Rossiter &amp; Suyanto &amp; Kingston" authorityYear="2007" box="[474,577,1514,1536]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="7" pageNumber="19" phylum="Chordata" rank="species" species="boeadii">
<emphasis id="B9225E428E10FF80FE74ECBAFDC1FA6F" box="[474,577,1514,1536]" italics="true" pageId="7" pageNumber="19">H. boeadii</emphasis>
</taxonomicName>
(holo-
<typeStatus id="54ED3CF28E10FF80FF2CEF56FF2FF98C" box="[130,175,1543,1564]" pageId="7" pageNumber="19">type</typeStatus>
), M28253,
<collectingCounty id="6288FADC8E10FF80FE83EF56FD02F98C" box="[301,642,1542,1564]" pageId="7" pageNumber="19">Rawa Aopa Watumohai National</collectingCounty>
<collectingMunicipality id="6B8D182A8E10FF80FF2CEF72FF30F9A9" box="[130,176,1571,1593]" pageId="7" pageNumber="19">Park</collectingMunicipality>
,
<location id="8E89D48B8E10FF80FF14EF72FEA3F9A9" LSID="urn:lsid:plazi:treatment:03FF33468E15FF8BFD52EC28FCE7FBE7:8E89D48B8E10FF80FF14EF72FEA3F9A9" box="[186,291,1571,1593]" county="Rawa Aopa Watumohai National" municipality="Park" name="South-East" pageId="7" pageNumber="19" stateProvince="Sulawesi Utara">South-East</location>
<collectingRegion id="49924CB28E10FF80FE84EF72FE04F9A9" box="[298,388,1571,1593]" country="Indonesia" name="Sulawesi Utara" pageId="7" pageNumber="19">Sulawesi</collectingRegion>
; B —
<emphasis id="B9225E428E10FF80FE6AEF72FDA4F9A8" box="[452,548,1571,1592]" italics="true" pageId="7" pageNumber="19">
<taxonomicName id="4C56F9D38E10FF80FE6AEF72FDA0F9A8" box="[452,544,1571,1592]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="7" pageNumber="19" phylum="Chordata" rank="species" species="papua">H. papua</taxonomicName>
,
</emphasis>
M16951,
<location id="8E89D48B8E10FF80FF2CEF6EFF7AF9C5" LSID="urn:lsid:plazi:treatment:03FF33468E15FF8BFD52EC28FCE7FBE7:8E89D48B8E10FF80FF2CEF6EFF7AF9C5" box="[130,250,1599,1621]" county="Indonesian" municipality="Boeadi" name="Biak Island" pageId="7" pageNumber="19" stateProvince="Papua Barat">Biak Island</location>
,
<collectingRegion id="49924CB28E10FF80FEA5EF6EFE2CF9C5" box="[267,428,1599,1621]" country="Indonesia" name="Papua Barat" pageId="7" pageNumber="19">Papua Province</collectingRegion>
; C —
<taxonomicName id="4C56F9D38E10FF80FDAFEF11FDFCF9C4" authorityName="Cantor" authorityYear="1846" box="[513,636,1599,1621]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="7" pageNumber="19" phylum="Chordata" rank="species" species="galeritus">
<emphasis id="B9225E428E10FF80FDAFEF11FDFCF9C4" box="[513,636,1599,1621]" italics="true" pageId="7" pageNumber="19">H. galeritus</emphasis>
</taxonomicName>
, M28250,
<collectingMunicipality id="6B8D182A8E10FF80FF4FEF0AFE73F9E1" box="[225,499,1627,1649]" pageId="7" pageNumber="19">Kakenauwe Nature Reserve</collectingMunicipality>
,
<location id="8E89D48B8E10FF80FE51EF0DFDFEF9E1" LSID="urn:lsid:plazi:treatment:03FF33468E15FF8BFD52EC28FCE7FBE7:8E89D48B8E10FF80FE51EF0DFDFEF9E1" box="[511,638,1627,1649]" municipality="Kakenauwe Nature Reserve" name="Buton Island" pageId="7" pageNumber="19" stateProvince="Papua Barat">Buton Island</location>
, South-East
<collectingRegion id="49924CB28E10FF80FF59EF26FED1F91D" box="[247,337,1655,1677]" country="Indonesia" name="Sulawesi Utara" pageId="7" pageNumber="19">Sulawesi</collectingRegion>
; D —
<taxonomicName id="4C56F9D38E10FF80FE0DEF29FD98F91D" box="[419,536,1656,1677]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="7" pageNumber="19" phylum="Chordata" rank="species" species="cervinus">
<emphasis id="B9225E428E10FF80FE0DEF29FD98F91D" box="[419,536,1656,1677]" italics="true" pageId="7" pageNumber="19">H. cervinus</emphasis>
</taxonomicName>
, M28249, Lambasango Wildlife Reserve, Buton Island, South- East
<collectingRegion id="49924CB28E10FF80FE8BEFE1FEFFF956" box="[293,383,1712,1734]" country="Indonesia" name="Sulawesi Utara" pageId="7" pageNumber="19">Sulawesi</collectingRegion>
. Scale =
<quantity id="4CAE2FB58E10FF80FE79EFE1FD8EF955" box="[471,526,1712,1734]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="19" unit="mm" value="5.0">5 mm</quantity>
</paragraph>
</caption>
<subSubSection id="C34CD1DB8E10FF80FD00ECB1FD63F955" pageId="7" pageNumber="19" type="etymology">
<paragraph id="8BE982508E10FF80FD00ECB1FCABFA6A" blockId="7.[686,1199,1504,1733]" box="[686,811,1504,1530]" pageId="7" pageNumber="19">
<emphasis id="B9225E428E10FF80FD00ECB1FCABFA6A" box="[686,811,1504,1530]" italics="true" pageId="7" pageNumber="19">Etymology</emphasis>
</paragraph>
<paragraph id="8BE982508E10FF80FD7DEF50FD63F955" blockId="7.[686,1199,1504,1733]" pageId="7" pageNumber="19">
<taxonomicName id="4C56F9D38E10FF80FD7DEF50FC5EF98B" authorityName="Bates &amp; Rossiter &amp; Suyanto &amp; Kingston" authorityYear="2007" box="[723,990,1537,1563]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="7" pageNumber="19" phylum="Chordata" rank="species" species="boeadii">
<emphasis id="B9225E428E10FF80FD7DEF50FC5EF98B" box="[723,990,1537,1563]" italics="true" pageId="7" pageNumber="19">Hipposideros boeadii</emphasis>
</taxonomicName>
is named after the
<collectingCounty id="6288FADC8E10FF80FD4CEF72FCE3F9AD" box="[738,867,1571,1597]" pageId="7" pageNumber="19">Indonesian</collectingCounty>
zoologist
<collectingMunicipality id="6B8D182A8E10FF80FC41EF72FBC5F9AD" box="[1007,1093,1571,1597]" pageId="7" pageNumber="19">Boeadi</collectingMunicipality>
, in rec- ognition of his long-standing contribu- tion to bat research in
<collectingCountry id="F341C2C08E10FF80FC63EF36FBC3F911" box="[973,1091,1639,1665]" name="Indonesia" pageId="7" pageNumber="19">Indonesia</collectingCountry>
. Its pro- posed English name is Boeadis roundleaf bat.
</paragraph>
</subSubSection>
<subSubSection id="C34CD1DB8E1FFF8CFF2EE9F5FC69FAF7" lastPageId="11" lastPageNumber="23" pageId="8" pageNumber="20" type="discussion">
<paragraph id="8BE982508E1FFF8FFF2EE9F5FE88FF2C" blockId="8.[128,264,164,190]" box="[128,264,164,190]" pageId="8" pageNumber="20">DISCUSSION</paragraph>
<paragraph id="8BE982508E1FFF8FFF2EE9BAFE59FE95" blockId="8.[128,273,235,261]" box="[128,473,235,261]" lastBlockId="8.[281,473,235,261]" pageId="8" pageNumber="20">
Comparison
<emphasis id="B9225E428E1FFF8FFEB7E9BAFE59FE95" box="[281,473,235,261]" italics="true" pageId="8" pageNumber="20">with Other Taxa</emphasis>
</paragraph>
<paragraph id="8BE982508E1FFF8FFF08E879FD03FC8A" blockId="8.[128,644,296,794]" pageId="8" pageNumber="20">
<taxonomicName id="4C56F9D38E1FFF8FFF08E879FE22FED2" authorityName="Bates &amp; Rossiter &amp; Suyanto &amp; Kingston" authorityYear="2007" box="[166,418,296,322]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="20" phylum="Chordata" rank="species" species="boeadii">
<emphasis id="B9225E428E1FFF8FFF08E879FE22FED2" box="[166,418,296,322]" italics="true" pageId="8" pageNumber="20">Hipposideros boeadii</emphasis>
</taxonomicName>
is one of a number of
<taxonomicName id="4C56F9D38E1FFF8FFF0FE818FEBEFEF3" box="[161,318,329,355]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="20" phylum="Chordata" rank="genus">
<emphasis id="B9225E428E1FFF8FFF0FE818FEBEFEF3" box="[161,318,329,355]" italics="true" pageId="8" pageNumber="20">Hipposideros</emphasis>
</taxonomicName>
taxa described from south- ern and South-East Asia, which have two supplementary leaflets in the noseleaf. In
<taxonomicName id="4C56F9D38E1FFF8FFF2EE8FEFF7FFE59" authorityName="Bates &amp; Rossiter &amp; Suyanto &amp; Kingston" authorityYear="2007" box="[128,255,431,457]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="20" phylum="Chordata" rank="species" species="boeadii">
<emphasis id="B9225E428E1FFF8FFF2EE8FEFF7FFE59" box="[128,255,431,457]" italics="true" pageId="8" pageNumber="20">H. boeadii</emphasis>
</taxonomicName>
, the inner leaflet is shorter than the outer. This distinguishes it from the two geographically widespread species,
<emphasis id="B9225E428E1FFF8FFD9AE8A2FF4BFDBE" italics="true" pageId="8" pageNumber="20">H. ga- leritus</emphasis>
<bibRefCitation id="EFC7FFA18E1FFF8FFF75EB45FEFBFDBE" author="CANTOR, T." box="[219,379,532,558]" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="8" pageNumber="20" pagination="171 - 203" part="15" refId="ref7434" refString="CANTOR, T. 1846. Catalogue of mammalia inhabiting the Malayan Peninsular and islands. Journal of the Asiatic Society of Bengal, 15: 171 - 203." title="Catalogue of mammalia inhabiting the Malayan Peninsular and islands" type="journal article" year="1846">Cantor, 1846</bibRefCitation>
(described from Pe- nang,
<collectingCountry id="F341C2C08E1FFF8FFF7FEB67FEBEFDC1" box="[209,318,566,593]" name="Malaysia" pageId="8" pageNumber="20">Malaysia</collectingCountry>
) and
<taxonomicName id="4C56F9D38E1FFF8FFE3CEB66FF3BFDE2" authority="Gould, 1854" authorityName="Gould" authorityYear="1854" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="20" phylum="Chordata" rank="species" species="cervinus">
<emphasis id="B9225E428E1FFF8FFE3CEB66FDA2FDC0" box="[402,546,567,592]" italics="true" pageId="8" pageNumber="20">H. cervinus</emphasis>
<bibRefCitation id="EFC7FFA18E1FFF8FFD9CEB67FF3BFDE2" author="GOULD, J." bookContentInfo="52 pp." journalOrPublisher="London" pageId="8" pageNumber="20" refId="ref7948" refString="GOULD, J. 1854. The mammals of Australia. 3. Pri- vately published, London, 52 pp. + 52 plates." title="The mammals of Australia. 3. Pri- vately published" type="book" year="1854">Gould, 1854</bibRefCitation>
</taxonomicName>
(described from Queensland, Austral- ia), both of which were also collected from Buton (South-East Sulawesi) during 2000. In both these latter taxa, the inner leaflet ex- ceeds the outer in length (
<figureCitation id="136D9ED58E1FFF8FFE19EB8EFDF6FD69" box="[439,630,735,761]" captionStart="FIG" captionStartId="5.[132,145,1490,1511]" captionTargetBox="[129,652,940,1452]" captionTargetId="figure@5.[355,658,938,1463]" captionTargetPageId="5" captionText="FIG. 2. Noseleaves of four species of Hipposideros (with hairs omitted). A — H. boeadii (holotype), M28253, Rawa Aopa Watumohai National Park, South-East Sulawesi; B — H. papua, M16951, Biak Island, Papua Province; C — H. galeritus, M28250, Kakenauwe Nature Reserve, Buton Island, South- East Sulawesi; D — H. cervinus, M28249, Lambasango Wildlife Reserve, Buton Island, South- East Sulawesi. Scale = 5 mm" figureDoi="http://doi.org/10.5281/zenodo.3944954" httpUri="https://zenodo.org/record/3944954/files/figure.png" pageId="8" pageNumber="20">Fig. 2C and 2D</figureCitation>
). Other characters that distinguish
<taxonomicName id="4C56F9D38E1FFF8FFDA9EA50FD03FC8A" authorityName="Bates &amp; Rossiter &amp; Suyanto &amp; Kingston" authorityYear="2007" box="[519,643,768,794]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="20" phylum="Chordata" rank="species" species="boeadii">
<emphasis id="B9225E428E1FFF8FFDA9EA50FD03FC8A" box="[519,643,768,794]" italics="true" pageId="8" pageNumber="20">H. boeadii</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BE982508E1FFF8FFD00E9F2FCACFD26" blockId="8.[686,1199,163,795]" pageId="8" pageNumber="20">
from
<taxonomicName id="4C56F9D38E1FFF8FFD5AE9F5FC07FF2D" authorityName="Cantor" authorityYear="1846" box="[756,903,163,189]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="20" phylum="Chordata" rank="species" species="galeritus">
<emphasis id="B9225E428E1FFF8FFD5AE9F5FC07FF2D" box="[756,903,163,189]" italics="true" pageId="8" pageNumber="20">H. galeritus</emphasis>
</taxonomicName>
and its associated taxa [
<emphasis id="B9225E428E1FFF8FFD19E994FCB9FF4F" box="[695,825,197,223]" italics="true" pageId="8" pageNumber="20">brachyotus</emphasis>
<bibRefCitation id="EFC7FFA18E1FFF8FFCEDE994FC69FF4F" author="DOBSON, G. E." box="[835,1001,197,223]" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="8" pageNumber="20" pagination="237 - 238" part="43" refId="ref7585" refString="DOBSON, G. E. 1874. Descriptions of new species of Chiroptera from India and Yunan. Journal of the Asiatic Society of Bengal, 43: 237 - 238." title="Descriptions of new species of Chiroptera from India and Yunan" type="journal article" year="1874">Dobson, 1874</bibRefCitation>
(described from Central
<collectingCountry id="F341C2C08E1FFF8FFCA3E9B6FCCEFE92" box="[781,846,231,258]" name="India" pageId="8" pageNumber="20">India</collectingCountry>
),
<emphasis id="B9225E428E1FFF8FFCCDE9B6FC43FE91" box="[867,963,231,257]" italics="true" pageId="8" pageNumber="20">insolens</emphasis>
<bibRefCitation id="EFC7FFA18E1FFF8FFC62E9B6FBD2FE91" author="LYON, M. W." box="[972,1106,231,257]" journalOrPublisher="Proceedings of the United States National Museum" pageId="8" pageNumber="20" pagination="53 - 146" part="40" refId="ref7618" refString="LYON, M. W. 1911. Mammals collected by Dr W. L. Abbott on Borneo and some of the small adjacent islands. Proceedings of the United States National Museum, 40: 53 - 146." title="Mammals collected by Dr W. L. Abbott on Borneo and some of the small adjacent islands" type="journal article" year="1911">Lyon, 1911</bibRefCitation>
(southeastern Kalimantan), and
<emphasis id="B9225E428E1FFF8FFBB7E859FB2FFEB2" box="[1049,1199,264,290]" italics="true" pageId="8" pageNumber="20">longicaudus</emphasis>
<bibRefCitation id="EFC7FFA18E1FFF8FFD00E87AFCC1FED4" author="PETERS, W." box="[686,833,298,325]" journalOrPublisher="Monatsberichte der Koniglichen Preussischen Akademie der Wissenschaften zu Berlin" pageId="8" pageNumber="20" pagination="706 - 712" part="1861" refId="ref7896" refString="PETERS, W. 1861. Uber die von Hrn. F. Jagor bisher auf Malacca, Borneo, Java und den Philippinen gesammelten Saugethiere aus den Ordnungen der Halbaffen, Pelzflatterer und Flederthiere. Monatsberichte der Koniglichen Preussischen Akademie der Wissenschaften zu Berlin, 1861: 706 - 712." title="Uber die von Hrn. F. Jagor bisher auf Malacca, Borneo, Java und den Philippinen gesammelten Saugethiere aus den Ordnungen der Halbaffen, Pelzflatterer und Flederthiere" type="journal article" year="1861">Peters, 1861</bibRefCitation>
(
<collectingRegion id="49924CB28E1FFF8FFCFCE878FC0BFED4" box="[850,907,297,324]" country="Indonesia" name="Jawa Timur" pageId="8" pageNumber="20">Java</collectingRegion>
)] are its shorter forearm length, larger skull, broader zygomata, more robust dentition and the absence of a greatly inflated postero-lateral chamber either side of the rostrum [
<tableCitation id="C6D4B7EB8E1FFF8FFC5FE8E1FBDAFE5B" box="[1009,1114,432,459]" captionStart="TABLE" captionStartId="4.[86,99,647,674]" captionTargetPageId="4" captionText="TABLE 1. External measurements (in mm) and body mass (in g) of four species of Hipposideros recently collected in Sulawesi. Used abbreviations: FA — forearm length; 5MET, 4MET, 3MET — fifth, fourth, and third metacarpal lengths; 5MET/FA× 100 — % length of fifth metacarpal relative to forearm length; 3D1P, 3D2P — length of first and second phalanges of the third digit; 3D1P/3MET × 100 — % length of first phalanx of the third digit relative to third metacarpal length; TAIL — tail length; HF — foot; TIBIA — tibia; E — ear and WGT — body mass. Sample sizes differing from those reported under n are given in parentheses. Mean and SD are included when sample size is ≥ 3" httpUri="http://table.plazi.org/id/DF29D2D88E13FF83FFF8EBD6FD8AFC83" pageId="8" pageNumber="20" tableUuid="DF29D2D88E13FF83FFF8EBD6FD8AFC83">Tables 1</tableCitation>
and
<tableCitation id="C6D4B7EB8E1FFF8FFB36E8E0FB29FE5B" box="[1176,1193,433,459]" captionStart="TABLE" captionStartId="9.[89,102,641,668]" captionText="TABLE 2. Cranial and dental measurements (in mm) of four species of Hipposideros recently collected in Sulawesi. Used abbreviations: GTL — greatest length of skull; CBL — condylobasal length; CCL — condylo-canine length; ZB — zygomatic breadth; BB — breadth of braincase; MW — mastoid width; ZB/MW × 100 — % breadth of zygomata relative to mastoid breadth; RW — rostral width; PC — postorbital constriction; CM3 — upper toothrow length; CM3/CCL × 100 — % length of upper toothrow relative to condylo-canine length; M3M3 — palatal breadth; CM — lower toothrow length; M — mandible length. Sample sizes 3 differing from those reported under n are given in parentheses. Mean and SD are included when sample size is ≥ 3" pageId="8" pageNumber="20">2</tableCitation>
,
<figureCitation id="136D9ED58E1FFF8FFD00E883FD7FFE7C" box="[686,767,466,492]" captionStart="FIG" captionStartId="7.[130,143,1486,1507]" captionTargetBox="[146,639,341,1457]" captionTargetId="figure@7.[125,645,615,920]" captionTargetPageId="7" captionText="FIG. 4. Lateral view of the skulls and mandibles of four species of Hipposideros. A — H. boeadii (holo- type), M28253, Rawa Aopa Watumohai National Park, South-East Sulawesi; B — H. papua, M16951, Biak Island, Papua Province; C — H. galeritus, M28250, Kakenauwe Nature Reserve, Buton Island, South-East Sulawesi; D — H. cervinus, M28249, Lambasango Wildlife Reserve, Buton Island, South- East Sulawesi. Scale = 5 mm" figureDoi="http://doi.org/10.5281/zenodo.3944962" httpUri="https://zenodo.org/record/3944962/files/figure.png" pageId="8" pageNumber="20">Figs. 4</figureCitation>
and
<figureCitation id="136D9ED58E1FFF8FFC97E883FCC8FE7D" box="[825,840,466,493]" captionStart="FIG" captionStartId="8.[130,143,1626,1647]" captionTargetBox="[273,1107,833,1601]" captionTargetId="figure@8.[235,1126,829,1604]" captionTargetPageId="8" captionText="FIG. 5. Dorsal view of the rostrum of four species of Hipposideros. A — H. boeadii (holotype), M28253, Rawa Aopa Watumohai National Park, South-East Sulawesi; B — H. papua, M16951, Biak Island, Papua Province; C — H. galeritus, M28250, Kakenauwe Nature Reserve, Buton Island, South-East Sulawesi; D —" figureDoi="http://doi.org/10.5281/zenodo.3944964" httpUri="https://zenodo.org/record/3944964/files/figure.png" pageId="8" pageNumber="20">5</figureCitation>
— for further details of these latter taxa, including measurements, see
<bibRefCitation id="EFC7FFA18E1FFF8FFD00EB44FCB1FDA0" author="HILL, J. E." box="[686,817,533,560]" journalOrPublisher="Bulletin of the British Museum (Natural History), Zoology" pageId="8" pageNumber="20" pagination="1 - 129" part="11" refId="ref8178" refString="HILL, J. E. 1963. A revison of the genus Hipposideros. Bulletin of the British Museum (Natural History), Zoology, 11: 1 - 129" title="A revison of the genus Hipposideros" type="journal article" year="1963">Hill (1963)</bibRefCitation>
and Jenkins and Hill (1981)]. In the noseleaf, the intermediate leaf is nar- rower than the posterior, whereas in
<taxonomicName id="4C56F9D38E1FFF8FFBCCEB08FD79FD04" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="20" phylum="Chordata" rank="species" species="gaia">
<emphasis id="B9225E428E1FFF8FFBCCEB08FD79FD04" italics="true" pageId="8" pageNumber="20">H. galeritus</emphasis>
</taxonomicName>
and its associated forms, it is broad- er (
<figureCitation id="136D9ED58E1FFF8FFD7BEBCDFC9EFD26" box="[725,798,668,694]" captionStart="FIG" captionStartId="5.[132,145,1490,1511]" captionTargetBox="[129,652,940,1452]" captionTargetId="figure@5.[355,658,938,1463]" captionTargetPageId="5" captionText="FIG. 2. Noseleaves of four species of Hipposideros (with hairs omitted). A — H. boeadii (holotype), M28253, Rawa Aopa Watumohai National Park, South-East Sulawesi; B — H. papua, M16951, Biak Island, Papua Province; C — H. galeritus, M28250, Kakenauwe Nature Reserve, Buton Island, South- East Sulawesi; D — H. cervinus, M28249, Lambasango Wildlife Reserve, Buton Island, South- East Sulawesi. Scale = 5 mm" figureDoi="http://doi.org/10.5281/zenodo.3944954" httpUri="https://zenodo.org/record/3944954/files/figure.png" pageId="8" pageNumber="20">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="8BE982508E1FFF8FFD7DEBEFFB2EFC8A" blockId="8.[686,1199,163,795]" pageId="8" pageNumber="20">
In comparison to
<taxonomicName id="4C56F9D38E1FFF8FFC05EBEFFBB5FD47" box="[939,1077,702,727]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="20" phylum="Chordata" rank="species" species="cervinus">
<emphasis id="B9225E428E1FFF8FFC05EBEFFBB5FD47" box="[939,1077,702,727]" italics="true" pageId="8" pageNumber="20">H. cervinus</emphasis>
</taxonomicName>
, the skull of
<taxonomicName id="4C56F9D38E1FFF8FFD7EEBB1FCCFFD69" authorityName="Bates &amp; Rossiter &amp; Suyanto &amp; Kingston" authorityYear="2007" box="[720,847,735,761]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="20" phylum="Chordata" rank="species" species="boeadii">
<emphasis id="B9225E428E1FFF8FFD7EEBB1FCCFFD69" box="[720,847,735,761]" italics="true" pageId="8" pageNumber="20">H. boeadii</emphasis>
</taxonomicName>
is longer relative to forearm length, the zygomata are broader, the cheek-
</paragraph>
<caption id="DF29D2D88E1FFF8FFF2CEF0BFB0EF954" ID-DOI="http://doi.org/10.5281/zenodo.3944964" ID-Zenodo-Dep="3944964" httpUri="https://zenodo.org/record/3944964/files/figure.png" pageId="8" pageNumber="20" startId="8.[130,143,1626,1647]" targetBox="[273,1107,833,1601]" targetPageId="8">
<paragraph id="8BE982508E1FFF8FFF2CEF0BFB0EF954" blockId="8.[130,1198,1625,1732]" pageId="8" pageNumber="20">
FIG. 5. Dorsal view of the rostrum of four species of
<emphasis id="B9225E428E1FFF8FFD0BEF08FCACF9FE" box="[677,812,1625,1646]" italics="true" pageId="8" pageNumber="20">
<taxonomicName id="4C56F9D38E1FFF8FFD0BEF08FCA9F9FE" box="[677,809,1625,1646]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="20" phylum="Chordata" rank="genus">Hipposideros</taxonomicName>
.
</emphasis>
A —
<taxonomicName id="4C56F9D38E1FFF8FFCDEEF0BFC59F9FE" authorityName="Bates &amp; Rossiter &amp; Suyanto &amp; Kingston" authorityYear="2007" box="[880,985,1625,1647]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="20" phylum="Chordata" rank="species" species="boeadii">
<emphasis id="B9225E428E1FFF8FFCDEEF0BFC59F9FE" box="[880,985,1625,1647]" italics="true" pageId="8" pageNumber="20">H. boeadii</emphasis>
</taxonomicName>
(holotype), M28253, Rawa Aopa Watumohai National Park, South-East Sulawesi; B —
<emphasis id="B9225E428E1FFF8FFC8AEF27FC08F91B" box="[804,904,1654,1675]" italics="true" pageId="8" pageNumber="20">
<taxonomicName id="4C56F9D38E1FFF8FFC8AEF27FC03F91B" box="[804,899,1654,1675]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="20" phylum="Chordata" rank="species" species="papua">H. papua</taxonomicName>
,
</emphasis>
M16951, Biak Island, Papua Province; C —
<taxonomicName id="4C56F9D38E1FFF8FFEB3EFC2FE11F937" authorityName="Cantor" authorityYear="1846" box="[285,401,1682,1704]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="20" phylum="Chordata" rank="species" species="galeritus">
<emphasis id="B9225E428E1FFF8FFEB3EFC2FE11F937" box="[285,401,1682,1704]" italics="true" pageId="8" pageNumber="20">H. galeritus</emphasis>
</taxonomicName>
, M28250, Kakenauwe Nature Reserve, Buton Island, South-East Sulawesi; D —
<taxonomicName id="4C56F9D38E1FFF8FFF0CEFFEFE91F954" box="[162,273,1711,1732]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="20" phylum="Chordata" rank="species" species="cervinus">
<emphasis id="B9225E428E1FFF8FFF0CEFFEFE91F954" box="[162,273,1711,1732]" italics="true" pageId="8" pageNumber="20">H. cervinus</emphasis>
</taxonomicName>
, M28249, Lambasango Wildlife Reserve, Buton Island, South-East
<collectingRegion id="49924CB28E1FFF8FFC0AEFFFFC7EF954" box="[932,1022,1710,1732]" country="Indonesia" name="Sulawesi Utara" pageId="8" pageNumber="20">Sulawesi</collectingRegion>
. Scale =
<quantity id="4CAE2FB58E1FFF8FFBF8EFFFFB0EF954" box="[1110,1166,1710,1732]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="20" unit="mm" value="5.0">5 mm</quantity>
</paragraph>
</caption>
<caption id="DF29D2D88E1EFF8EFFF7EBD0FB2EFC9D" pageId="9" pageNumber="21">
<paragraph id="8BE982508E1EFF8EFFF7EBD0FB2EFC9D" blockId="9.[89,1652,641,781]" pageId="9" pageNumber="21">
TABLE 2. Cranial and dental measurements (in mm) of four species of
<taxonomicName id="4C56F9D38E1EFF8EFD51EBD0FCFFFD0C" box="[767,895,641,668]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="9" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="B9225E428E1EFF8EFD51EBD0FCFFFD0C" box="[767,895,641,668]" italics="true" pageId="9" pageNumber="21">Hipposideros</emphasis>
</taxonomicName>
recently collected in Sulawesi. Used abbreviations: GTL — greatest length of skull; CBL — condylobasal length; CCL — condylo-canine length; ZB — zygomatic breadth; BB — breadth of braincase; MW — mastoid width; ZB/MW × 100 — % breadth of zygomata relative to mastoid breadth; RW — rostral width; PC — postorbital constriction; CM
<superScript id="7C232F188E1EFF8EFB34EBE6FB22FD5A" attach="left" box="[1178,1186,695,714]" fontSize="6" pageId="9" pageNumber="21">3</superScript>
— upper toothrow length; CM
<superScript id="7C232F188E1EFF8EFA75EBE6FA63FD5A" attach="left" box="[1499,1507,695,714]" fontSize="6" pageId="9" pageNumber="21">3</superScript>
/CCL × 100 — % length of upper toothrow relative to condylo-canine length; M
<superScript id="7C232F188E1EFF8EFD7DEB85FD5BFD77" attach="left" box="[723,731,724,743]" fontSize="6" pageId="9" pageNumber="21">3</superScript>
M
<superScript id="7C232F188E1EFF8EFD53EB85FC85FD77" attach="left" box="[765,773,724,743]" fontSize="6" pageId="9" pageNumber="21">3</superScript>
— palatal breadth; CM — lower toothrow length; M — mandible length. Sample sizes
<subScript id="17D280158E1EFF8EFC55EBB2FB83FD66" attach="left" box="[1019,1027,739,758]" fontSize="6" pageId="9" pageNumber="21">3</subScript>
differing from those reported under
<emphasis id="B9225E428E1EFF8EFE18EBA3FE42FC9D" box="[438,450,754,781]" italics="true" pageId="9" pageNumber="21">n</emphasis>
are given in parentheses. Mean and SD are included when sample size is ≥ 3
</paragraph>
</caption>
<paragraph id="8BE982508E1EFF8EFFF8EA68F9E2FB2B" pageId="9" pageNumber="21">
<table id="F95670F08E1E0078FFF8EA7DF9FCFB2B" box="[86,1660,812,1211]" gridcols="15" gridrows="12" pageId="9" pageNumber="21">
<tr id="356680128E1E0078FFF8EA7DF9FCFCF5" box="[86,1660,812,869]" gridrow="0" pageId="9" pageNumber="21">
<th id="76B7E96E8E1E0078FFF8EA7DFF0CFCF5" box="[86,140,812,869]" gridcol="0" gridrow="0" pageId="9" pageNumber="21">
<emphasis id="B9225E428E1EFF8EFFF8EA68FFE2FCC4" box="[86,98,825,852]" italics="true" pageId="9" pageNumber="21">n</emphasis>
sex
</th>
<th id="76B7E96E8E1E0078FF30EA7DFF7EFCF5" box="[158,254,812,869]" gridcol="1" gridrow="0" pageId="9" pageNumber="21">GTL</th>
<th id="76B7E96E8E1E0078FEA1EA7DFEF0FCF5" box="[271,368,812,869]" gridcol="2" gridrow="0" pageId="9" pageNumber="21">CBL</th>
<th id="76B7E96E8E1E0078FE2FEA7DFE61FCF5" box="[385,481,812,869]" gridcol="3" gridrow="0" pageId="9" pageNumber="21">CCL</th>
<th id="76B7E96E8E1E0078FE5CEA7DFDD3FCF5" box="[498,595,812,869]" gridcol="4" gridrow="0" pageId="9" pageNumber="21">ZB</th>
<th id="76B7E96E8E1E0078FDC6EA7DFD32FCF5" box="[616,690,812,869]" gridcol="5" gridrow="0" pageId="9" pageNumber="21">BB</th>
<th id="76B7E96E8E1E0078FD64EA7DFC93FCF5" box="[714,787,812,869]" gridcol="6" gridrow="0" pageId="9" pageNumber="21">MW</th>
<th id="76B7E96E8E1E0078FC82EA7DFC23FCF5" box="[812,931,812,869]" gridcol="7" gridrow="0" pageId="9" pageNumber="21">ZB/MW × 100</th>
<th id="76B7E96E8E1E0078FC19EA7DFB85FCF5" box="[951,1029,812,869]" gridcol="8" gridrow="0" pageId="9" pageNumber="21">RW</th>
<th id="76B7E96E8E1E0078FBB9EA7DFBDFFCF5" box="[1047,1119,812,869]" gridcol="9" gridrow="0" pageId="9" pageNumber="21">PC</th>
<th id="76B7E96E8E1E0078FBDFEA7DFB3AFCF5" box="[1137,1210,812,869]" gridcol="10" gridrow="0" pageId="9" pageNumber="21">CM3</th>
<th id="76B7E96E8E1E0078FB7BEA7DFAC3FCF5" box="[1237,1347,812,869]" gridcol="11" gridrow="0" pageId="9" pageNumber="21">CM3/CCL × 100</th>
<th id="76B7E96E8E1E0078FAF2EA7DFA25FCF5" box="[1372,1445,812,869]" gridcol="12" gridrow="0" pageId="9" pageNumber="21">M3M3</th>
<th id="76B7E96E8E1E0078FA10EA7DF989FCF5" box="[1470,1545,812,869]" gridcol="13" gridrow="0" pageId="9" pageNumber="21">CM3</th>
<th id="76B7E96E8E1E0078F9B2EA7DF9FCFCF5" box="[1564,1660,812,869]" gridcol="14" gridrow="0" pageId="9" pageNumber="21">M</th>
</tr>
<tr id="356680128E1E0078FFF8EA23F9FCFC1D" box="[86,1660,882,909]" gridrow="1" pageId="9" pageNumber="21" rowspan-0="1" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-9="1">
<td id="76B7E96E8E1E0078FD64EA23FB85FC1D" box="[714,1029,882,909]" colspan="3" colspanRight="2" gridcol="6" gridrow="1" pageId="9" pageNumber="21">
<taxonomicName id="4C56F9D38E1EFF8EFCAFEA23FC50FC1D" authorityName="Bates &amp; Rossiter &amp; Suyanto &amp; Kingston" authorityYear="2007" box="[769,976,882,909]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="9" pageNumber="21" phylum="Chordata" rank="species" species="boeadii">
<emphasis id="B9225E428E1EFF8EFCAFEA23FC50FC1D" box="[769,976,882,909]" italics="true" pageId="9" pageNumber="21">Hipposideros boeadii</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="356680128E1E0078FFF8EADEF9FCFC3A" box="[86,1660,911,938]" gridrow="2" pageId="9" pageNumber="21">
<th id="76B7E96E8E1E0078FFF8EADEFF0CFC3A" box="[86,140,911,938]" gridcol="0" gridrow="2" pageId="9" pageNumber="21">2 Ƌ</th>
<td id="76B7E96E8E1E0078FF30EADEFF7EFC3A" box="[158,254,911,938]" gridcol="1" gridrow="2" pageId="9" pageNumber="21">18.3, 18.3</td>
<td id="76B7E96E8E1E0078FEA1EADEFEF0FC3A" box="[271,368,911,938]" gridcol="2" gridrow="2" pageId="9" pageNumber="21">15.6, 15.6</td>
<td id="76B7E96E8E1E0078FE2FEADEFE61FC3A" box="[385,481,911,938]" gridcol="3" gridrow="2" pageId="9" pageNumber="21">15.3, 15.5</td>
<td id="76B7E96E8E1E0078FE5CEADEFDD3FC3A" box="[498,595,911,938]" gridcol="4" gridrow="2" pageId="9" pageNumber="21">10.0, 10.3</td>
<td id="76B7E96E8E1E0078FDC6EADEFD32FC3A" box="[616,690,911,938]" gridcol="5" gridrow="2" pageId="9" pageNumber="21">7.7, 7.9</td>
<td id="76B7E96E8E1E0078FD64EADEFC93FC3A" box="[714,787,911,938]" gridcol="6" gridrow="2" pageId="9" pageNumber="21">8.8, 9.1</td>
<td id="76B7E96E8E1E0078FC82EADEFC23FC3A" box="[812,931,911,938]" gridcol="7" gridrow="2" pageId="9" pageNumber="21">113.4, 113.5</td>
<td id="76B7E96E8E1E0078FC19EADEFB85FC3A" box="[951,1029,911,938]" gridcol="8" gridrow="2" pageId="9" pageNumber="21">5.1, 5.2</td>
<td id="76B7E96E8E1E0078FBB9EADEFBDFFC3A" box="[1047,1119,911,938]" gridcol="9" gridrow="2" pageId="9" pageNumber="21">2.6, 2.9</td>
<td id="76B7E96E8E1E0078FBDFEADEFB3AFC3A" box="[1137,1210,911,938]" gridcol="10" gridrow="2" pageId="9" pageNumber="21">6.8, 6.8</td>
<td id="76B7E96E8E1E0078FB7BEADEFAC3FC3A" box="[1237,1347,911,938]" gridcol="11" gridrow="2" pageId="9" pageNumber="21">44.0, 44.1</td>
<td id="76B7E96E8E1E0078FAF2EADEFA25FC3A" box="[1372,1445,911,938]" gridcol="12" gridrow="2" pageId="9" pageNumber="21">7.0, 7.5</td>
<td id="76B7E96E8E1E0078FA10EADEF989FC3A" box="[1470,1545,911,938]" gridcol="13" gridrow="2" pageId="9" pageNumber="21">7.2, 7.4</td>
<td id="76B7E96E8E1E0078F9B2EADEF9FCFC3A" box="[1564,1660,911,938]" gridcol="14" gridrow="2" pageId="9" pageNumber="21">11.7, 12.1</td>
</tr>
<tr id="356680128E1E0078FFF8EAFAF9FCFC56" box="[86,1660,939,966]" gridrow="3" pageId="9" pageNumber="21">
<th id="76B7E96E8E1E0078FFF8EAFAFF0CFC56" box="[86,140,939,966]" gridcol="0" gridrow="3" pageId="9" pageNumber="21">2 ♀</th>
<td id="76B7E96E8E1E0078FF30EAFAFF7EFC56" box="[158,254,939,966]" gridcol="1" gridrow="3" pageId="9" pageNumber="21">18.0, 18.1</td>
<td id="76B7E96E8E1E0078FEA1EAFAFEF0FC56" box="[271,368,939,966]" gridcol="2" gridrow="3" pageId="9" pageNumber="21">15.6, 15.8</td>
<td id="76B7E96E8E1E0078FE2FEAFAFE61FC56" box="[385,481,939,966]" gridcol="3" gridrow="3" pageId="9" pageNumber="21">15.2, 15.4</td>
<td id="76B7E96E8E1E0078FE5CEAFAFDD3FC56" box="[498,595,939,966]" gridcol="4" gridrow="3" pageId="9" pageNumber="21">10.2, 10.4</td>
<td id="76B7E96E8E1E0078FDC6EAFAFD32FC56" box="[616,690,939,966]" gridcol="5" gridrow="3" pageId="9" pageNumber="21">7.6, 7.7</td>
<td id="76B7E96E8E1E0078FD64EAFAFC93FC56" box="[714,787,939,966]" gridcol="6" gridrow="3" pageId="9" pageNumber="21">9.0, 9.0</td>
<td id="76B7E96E8E1E0078FC82EAFAFC23FC56" box="[812,931,939,966]" gridcol="7" gridrow="3" pageId="9" pageNumber="21">113.4, 115.5</td>
<td id="76B7E96E8E1E0078FC19EAFAFB85FC56" box="[951,1029,939,966]" gridcol="8" gridrow="3" pageId="9" pageNumber="21">5.0, 5.3</td>
<td id="76B7E96E8E1E0078FBB9EAFAFBDFFC56" box="[1047,1119,939,966]" gridcol="9" gridrow="3" pageId="9" pageNumber="21">2.9, 3.0</td>
<td id="76B7E96E8E1E0078FBDFEAFAFB3AFC56" box="[1137,1210,939,966]" gridcol="10" gridrow="3" pageId="9" pageNumber="21">6.7, 6.9</td>
<td id="76B7E96E8E1E0078FB7BEAFAFAC3FC56" box="[1237,1347,939,966]" gridcol="11" gridrow="3" pageId="9" pageNumber="21">44.2, 44.4</td>
<td id="76B7E96E8E1E0078FAF2EAFAFA25FC56" box="[1372,1445,939,966]" gridcol="12" gridrow="3" pageId="9" pageNumber="21">7.3, 7.4</td>
<td id="76B7E96E8E1E0078FA10EAFAF989FC56" box="[1470,1545,939,966]" gridcol="13" gridrow="3" pageId="9" pageNumber="21">7.2, 7.4</td>
<td id="76B7E96E8E1E0078F9B2EAFAF9FCFC56" box="[1564,1660,939,966]" gridcol="14" gridrow="3" pageId="9" pageNumber="21">11.9, 12.1</td>
</tr>
<tr id="356680128E1E0078FFF8EA9FF9FCFC79" box="[86,1660,974,1001]" gridrow="4" pageId="9" pageNumber="21" rowspan-0="1" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-8="1" rowspan-9="1">
<td id="76B7E96E8E1E0078FC82EA9FFC23FC79" box="[812,931,974,1001]" gridcol="7" gridrow="4" pageId="9" pageNumber="21">
<taxonomicName id="4C56F9D38E1EFF8EFC92EA9FFC16FC79" box="[828,918,974,1001]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="9" pageNumber="21" phylum="Chordata" rank="species" species="papua">
<emphasis id="B9225E428E1EFF8EFC92EA9FFC16FC79" box="[828,918,974,1001]" italics="true" pageId="9" pageNumber="21">H. papua</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="356680128E1E0078FFF8EABBF9FCFB95" box="[86,1660,1002,1029]" gridrow="5" pageId="9" pageNumber="21">
<th id="76B7E96E8E1E0078FFF8EABBFF0CFB95" box="[86,140,1002,1029]" gridcol="0" gridrow="5" pageId="9" pageNumber="21">1 Ƌ</th>
<td id="76B7E96E8E1E0078FF30EABBFF7EFB95" box="[158,254,1002,1029]" gridcol="1" gridrow="5" pageId="9" pageNumber="21">19.7</td>
<td id="76B7E96E8E1E0078FEA1EABBFEF0FB95" box="[271,368,1002,1029]" gridcol="2" gridrow="5" pageId="9" pageNumber="21">17.1</td>
<td id="76B7E96E8E1E0078FE2FEABBFE61FB95" box="[385,481,1002,1029]" gridcol="3" gridrow="5" pageId="9" pageNumber="21">17.0</td>
<td id="76B7E96E8E1E0078FE5CEABBFDD3FB95" box="[498,595,1002,1029]" gridcol="4" gridrow="5" pageId="9" pageNumber="21">10.8</td>
<td id="76B7E96E8E1E0078FDC6EABBFD32FB95" box="[616,690,1002,1029]" gridcol="5" gridrow="5" pageId="9" pageNumber="21">8.5</td>
<td id="76B7E96E8E1E0078FD64EABBFC93FB95" box="[714,787,1002,1029]" gridcol="6" gridrow="5" pageId="9" pageNumber="21">9.7</td>
<td id="76B7E96E8E1E0078FC82EABBFC23FB95" box="[812,931,1002,1029]" gridcol="7" gridrow="5" pageId="9" pageNumber="21">111.3</td>
<td id="76B7E96E8E1E0078FC19EABBFB85FB95" box="[951,1029,1002,1029]" gridcol="8" gridrow="5" pageId="9" pageNumber="21">6.0</td>
<td id="76B7E96E8E1E0078FBB9EABBFBDFFB95" box="[1047,1119,1002,1029]" gridcol="9" gridrow="5" pageId="9" pageNumber="21"></td>
<td id="76B7E96E8E1E0078FBDFEABBFB3AFB95" box="[1137,1210,1002,1029]" gridcol="10" gridrow="5" pageId="9" pageNumber="21">7.3</td>
<td id="76B7E96E8E1E0078FB7BEABBFAC3FB95" box="[1237,1347,1002,1029]" gridcol="11" gridrow="5" pageId="9" pageNumber="21">42.9</td>
<td id="76B7E96E8E1E0078FAF2EABBFA25FB95" box="[1372,1445,1002,1029]" gridcol="12" gridrow="5" pageId="9" pageNumber="21"></td>
<td id="76B7E96E8E1E0078FA10EABBF989FB95" box="[1470,1545,1002,1029]" gridcol="13" gridrow="5" pageId="9" pageNumber="21">8.0</td>
<td id="76B7E96E8E1E0078F9B2EABBF9FCFB95" box="[1564,1660,1002,1029]" gridcol="14" gridrow="5" pageId="9" pageNumber="21">13.2</td>
</tr>
<tr id="356680128E1E0078FFF8ED57F9FCFBB1" box="[86,1660,1030,1057]" gridrow="6" pageId="9" pageNumber="21">
<th id="76B7E96E8E1E0078FFF8ED57FF7EFBB1" box="[86,254,1030,1057]" colspan="2" colspanRight="1" gridcol="0" gridrow="6" pageId="9" pageNumber="21">3 ♀* 19.8, 0.7</th>
<td id="76B7E96E8E1E0078FEA1ED57FEF0FBB1" box="[271,368,1030,1057]" gridcol="2" gridrow="6" pageId="9" pageNumber="21">(2)</td>
<td id="76B7E96E8E1E0078FE2FED57FE61FBB1" box="[385,481,1030,1057]" gridcol="3" gridrow="6" pageId="9" pageNumber="21">17.2, 0.5</td>
<td id="76B7E96E8E1E0078FE5CED57FDD3FBB1" box="[498,595,1030,1057]" gridcol="4" gridrow="6" pageId="9" pageNumber="21">11.1, 0.3</td>
<td id="76B7E96E8E1E0078FDC6ED57FD32FBB1" box="[616,690,1030,1057]" gridcol="5" gridrow="6" pageId="9" pageNumber="21">8.2, 0.3</td>
<td id="76B7E96E8E1E0078FD64ED57FC93FBB1" box="[714,787,1030,1057]" gridcol="6" gridrow="6" pageId="9" pageNumber="21">9.8, 0.3</td>
<td id="76B7E96E8E1E0078FC82ED57FC23FBB1" box="[812,931,1030,1057]" gridcol="7" gridrow="6" pageId="9" pageNumber="21">113.8, 1.8</td>
<td id="76B7E96E8E1E0078FC19ED57FB85FBB1" box="[951,1029,1030,1057]" gridcol="8" gridrow="6" pageId="9" pageNumber="21">5.9, 0.4</td>
<td id="76B7E96E8E1E0078FBB9ED57FBDFFBB1" box="[1047,1119,1030,1057]" gridcol="9" gridrow="6" pageId="9" pageNumber="21">(2)</td>
<td id="76B7E96E8E1E0078FBDFED57FB3AFBB1" box="[1137,1210,1030,1057]" gridcol="10" gridrow="6" pageId="9" pageNumber="21">7.7, 0.3</td>
<td id="76B7E96E8E1E0078FB7BED57FAC3FBB1" box="[1237,1347,1030,1057]" gridcol="11" gridrow="6" pageId="9" pageNumber="21">44.8, 0.1</td>
<td id="76B7E96E8E1E0078FAF2ED57FA25FBB1" box="[1372,1445,1030,1057]" gridcol="12" gridrow="6" pageId="9" pageNumber="21">(2)</td>
<td id="76B7E96E8E1E0078FA10ED57F989FBB1" box="[1470,1545,1030,1057]" gridcol="13" gridrow="6" pageId="9" pageNumber="21">8.3, 0.1</td>
<td id="76B7E96E8E1E0078F9B2ED57F9FCFBB1" box="[1564,1660,1030,1057]" gridcol="14" gridrow="6" pageId="9" pageNumber="21">13.4, 0.6</td>
</tr>
<tr id="356680128E1E0078FFF8ED73F9FCFBAD" box="[86,1660,1058,1085]" gridrow="7" pageId="9" pageNumber="21" rowspan-0="1">
<td id="76B7E96E8E1E0078FF30ED73FF7EFBAD" box="[158,254,1058,1085]" gridcol="1" gridrow="7" pageId="9" pageNumber="21">19.220.5</td>
<td id="76B7E96E8E1E0078FEA1ED73FEF0FBAD" box="[271,368,1058,1085]" gridcol="2" gridrow="7" pageId="9" pageNumber="21">17.5, 17.9</td>
<td id="76B7E96E8E1E0078FE2FED73FE61FBAD" box="[385,481,1058,1085]" gridcol="3" gridrow="7" pageId="9" pageNumber="21">16.717.6</td>
<td id="76B7E96E8E1E0078FE5CED73FDD3FBAD" box="[498,595,1058,1085]" gridcol="4" gridrow="7" pageId="9" pageNumber="21">10.911.4</td>
<td id="76B7E96E8E1E0078FDC6ED73FD32FBAD" box="[616,690,1058,1085]" gridcol="5" gridrow="7" pageId="9" pageNumber="21">7.98.4</td>
<td id="76B7E96E8E1E0078FD64ED73FC23FBAD" box="[714,931,1058,1085]" colspan="2" colspanRight="1" gridcol="6" gridrow="7" pageId="9" pageNumber="21">9.410.1 112.2115.8</td>
<td id="76B7E96E8E1E0078FC19ED73FB85FBAD" box="[951,1029,1058,1085]" gridcol="8" gridrow="7" pageId="9" pageNumber="21">5.66.3</td>
<td id="76B7E96E8E1E0078FBB9ED73FBDFFBAD" box="[1047,1119,1058,1085]" gridcol="9" gridrow="7" pageId="9" pageNumber="21">2.9, 2.9</td>
<td id="76B7E96E8E1E0078FBDFED73FB3AFBAD" box="[1137,1210,1058,1085]" gridcol="10" gridrow="7" pageId="9" pageNumber="21">7.58.0</td>
<td id="76B7E96E8E1E0078FB7BED73FAC3FBAD" box="[1237,1347,1058,1085]" gridcol="11" gridrow="7" pageId="9" pageNumber="21">43.645.5</td>
<td id="76B7E96E8E1E0078FAF2ED73FA25FBAD" box="[1372,1445,1058,1085]" gridcol="12" gridrow="7" pageId="9" pageNumber="21">7.8, 8.3</td>
<td id="76B7E96E8E1E0078FA10ED73F989FBAD" box="[1470,1545,1058,1085]" gridcol="13" gridrow="7" pageId="9" pageNumber="21">8.28.4</td>
<td id="76B7E96E8E1E0078F9B2ED73F9FCFBAD" box="[1564,1660,1058,1085]" gridcol="14" gridrow="7" pageId="9" pageNumber="21">12.813.9</td>
</tr>
<tr id="356680128E1E0078FFF8ED14F9FCFBF0" box="[86,1660,1093,1120]" gridrow="8" pageId="9" pageNumber="21" rowspan-0="1" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-8="1" rowspan-9="1">
<td id="76B7E96E8E1E0078FC82ED14FC23FBF0" box="[812,931,1093,1120]" gridcol="7" gridrow="8" pageId="9" pageNumber="21">
<taxonomicName id="4C56F9D38E1EFF8EFC81ED14FC22FBF0" authorityName="Cantor" authorityYear="1846" box="[815,930,1093,1120]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="9" pageNumber="21" phylum="Chordata" rank="species" species="galeritus">
<emphasis id="B9225E428E1EFF8EFC81ED14FC22FBF0" box="[815,930,1093,1120]" italics="true" pageId="9" pageNumber="21">H. galeritus</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="356680128E1E0078FFF8ED30F9FCFBEC" box="[86,1660,1121,1148]" gridrow="9" pageId="9" pageNumber="21">
<th id="76B7E96E8E1E0078FFF8ED30FF0CFBEC" box="[86,140,1121,1148]" gridcol="0" gridrow="9" pageId="9" pageNumber="21">2 ♀</th>
<td id="76B7E96E8E1E0078FF30ED30FF7EFBEC" box="[158,254,1121,1148]" gridcol="1" gridrow="9" pageId="9" pageNumber="21">16.5, 16.7</td>
<td id="76B7E96E8E1E0078FEA1ED30FEF0FBEC" box="[271,368,1121,1148]" gridcol="2" gridrow="9" pageId="9" pageNumber="21">14.6, 14.6</td>
<td id="76B7E96E8E1E0078FE2FED30FE61FBEC" box="[385,481,1121,1148]" gridcol="3" gridrow="9" pageId="9" pageNumber="21">14.1, 14.2</td>
<td id="76B7E96E8E1E0078FE5CED30FDD3FBEC" box="[498,595,1121,1148]" gridcol="4" gridrow="9" pageId="9" pageNumber="21">7.8, 8.1</td>
<td id="76B7E96E8E1E0078FDC6ED30FD32FBEC" box="[616,690,1121,1148]" gridcol="5" gridrow="9" pageId="9" pageNumber="21">7.0, 7.0</td>
<td id="76B7E96E8E1E0078FD64ED30FC93FBEC" box="[714,787,1121,1148]" gridcol="6" gridrow="9" pageId="9" pageNumber="21">8.4, 8.7</td>
<td id="76B7E96E8E1E0078FC82ED30FC23FBEC" box="[812,931,1121,1148]" gridcol="7" gridrow="9" pageId="9" pageNumber="21">93.1, 93.8</td>
<td id="76B7E96E8E1E0078FC19ED30FB85FBEC" box="[951,1029,1121,1148]" gridcol="8" gridrow="9" pageId="9" pageNumber="21">4.4, 4.5</td>
<td id="76B7E96E8E1E0078FBB9ED30FBDFFBEC" box="[1047,1119,1121,1148]" gridcol="9" gridrow="9" pageId="9" pageNumber="21">2.3, 2.3</td>
<td id="76B7E96E8E1E0078FBDFED30FB3AFBEC" box="[1137,1210,1121,1148]" gridcol="10" gridrow="9" pageId="9" pageNumber="21">5.4, 5.5</td>
<td id="76B7E96E8E1E0078FB7BED30FAC3FBEC" box="[1237,1347,1121,1148]" gridcol="11" gridrow="9" pageId="9" pageNumber="21">38.3, 38.5</td>
<td id="76B7E96E8E1E0078FAF2ED30FA25FBEC" box="[1372,1445,1121,1148]" gridcol="12" gridrow="9" pageId="9" pageNumber="21">5.4, 5.4</td>
<td id="76B7E96E8E1E0078FA10ED30F989FBEC" box="[1470,1545,1121,1148]" gridcol="13" gridrow="9" pageId="9" pageNumber="21">5.6, 5.8</td>
<td id="76B7E96E8E1E0078F9B2ED30F9FCFBEC" box="[1564,1660,1121,1148]" gridcol="14" gridrow="9" pageId="9" pageNumber="21">9.8, 9.9</td>
</tr>
<tr id="356680128E1E0078FFF8EDD5F9FCFB0F" box="[86,1660,1156,1183]" gridrow="10" pageId="9" pageNumber="21" rowspan-0="1" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-8="1" rowspan-9="1">
<td id="76B7E96E8E1E0078FC82EDD5FC23FB0F" box="[812,931,1156,1183]" gridcol="7" gridrow="10" pageId="9" pageNumber="21">
<taxonomicName id="4C56F9D38E1EFF8EFC9FEDD5FC20FB0F" box="[817,928,1156,1183]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="9" pageNumber="21" phylum="Chordata" rank="species" species="cervinus">
<emphasis id="B9225E428E1EFF8EFC9FEDD5FC20FB0F" box="[817,928,1156,1183]" italics="true" pageId="9" pageNumber="21">H. cervinus</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="356680128E1E0078FFF8EDF1F9FCFB2B" box="[86,1660,1184,1211]" gridrow="11" pageId="9" pageNumber="21">
<th id="76B7E96E8E1E0078FFF8EDF1FF0CFB2B" box="[86,140,1184,1211]" gridcol="0" gridrow="11" pageId="9" pageNumber="21">1 ♀</th>
<td id="76B7E96E8E1E0078FF30EDF1FF7EFB2B" box="[158,254,1184,1211]" gridcol="1" gridrow="11" pageId="9" pageNumber="21">16.6</td>
<td id="76B7E96E8E1E0078FEA1EDF1FEF0FB2B" box="[271,368,1184,1211]" gridcol="2" gridrow="11" pageId="9" pageNumber="21">14.4</td>
<td id="76B7E96E8E1E0078FE2FEDF1FE61FB2B" box="[385,481,1184,1211]" gridcol="3" gridrow="11" pageId="9" pageNumber="21">13.9</td>
<td id="76B7E96E8E1E0078FE5CEDF1FDD3FB2B" box="[498,595,1184,1211]" gridcol="4" gridrow="11" pageId="9" pageNumber="21">8.7</td>
<td id="76B7E96E8E1E0078FDC6EDF1FD32FB2B" box="[616,690,1184,1211]" gridcol="5" gridrow="11" pageId="9" pageNumber="21">7.2</td>
<td id="76B7E96E8E1E0078FD64EDF1FC93FB2B" box="[714,787,1184,1211]" gridcol="6" gridrow="11" pageId="9" pageNumber="21">8.4</td>
<td id="76B7E96E8E1E0078FC82EDF1FC23FB2B" box="[812,931,1184,1211]" gridcol="7" gridrow="11" pageId="9" pageNumber="21">104.4</td>
<td id="76B7E96E8E1E0078FC19EDF1FB85FB2B" box="[951,1029,1184,1211]" gridcol="8" gridrow="11" pageId="9" pageNumber="21">4.3</td>
<td id="76B7E96E8E1E0078FBB9EDF1FBDFFB2B" box="[1047,1119,1184,1211]" gridcol="9" gridrow="11" pageId="9" pageNumber="21">2.5</td>
<td id="76B7E96E8E1E0078FBDFEDF1FB3AFB2B" box="[1137,1210,1184,1211]" gridcol="10" gridrow="11" pageId="9" pageNumber="21">5.7</td>
<td id="76B7E96E8E1E0078FB7BEDF1FAC3FB2B" box="[1237,1347,1184,1211]" gridcol="11" gridrow="11" pageId="9" pageNumber="21">40.8</td>
<td id="76B7E96E8E1E0078FAF2EDF1FA25FB2B" box="[1372,1445,1184,1211]" gridcol="12" gridrow="11" pageId="9" pageNumber="21">5.8</td>
<td id="76B7E96E8E1E0078FA10EDF1F989FB2B" box="[1470,1545,1184,1211]" gridcol="13" gridrow="11" pageId="9" pageNumber="21">6.1</td>
<td id="76B7E96E8E1E0078F9B2EDF1F9FCFB2B" box="[1564,1660,1184,1211]" gridcol="14" gridrow="11" pageId="9" pageNumber="21">10.4</td>
</tr>
</table>
</paragraph>
<footnote id="E84D9E5E8E1EFF8EFFF8ED97FDC3FB71" box="[86,579,1222,1249]" pageId="9" pageNumber="21">
<paragraph id="8BE982508E1EFF8EFFF8ED97FDC3FB71" blockId="9.[86,579,1222,1249]" box="[86,579,1222,1249]" pageId="9" pageNumber="21">* — includes data from Hill and Rozendaal (1989)</paragraph>
</footnote>
<caption id="DF29D2D88E1DFF8DFF2CEB50FE45FDDE" ID-DOI="http://doi.org/10.5281/zenodo.3944966" ID-Zenodo-Dep="3944966" httpUri="https://zenodo.org/record/3944966/files/figure.png" pageId="10" pageNumber="22" startId="10.[130,143,513,534]" targetBox="[183,574,164,492]" targetPageId="10">
<paragraph id="8BE982508E1DFF8DFF2CEB50FE45FDDE" blockId="10.[130,644,512,591]" pageId="10" pageNumber="22">
FIG. 6. Baculum of
<taxonomicName id="4C56F9D38E1DFF8DFEEFEB50FE27FD85" authorityName="Bates &amp; Rossiter &amp; Suyanto &amp; Kingston" authorityYear="2007" box="[321,423,512,534]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="22" phylum="Chordata" rank="species" species="boeadii">
<emphasis id="B9225E428E1DFF8DFEEFEB50FE27FD85" box="[321,423,512,534]" italics="true" pageId="10" pageNumber="22">H. boeadii</emphasis>
</taxonomicName>
,
<accessionNumber id="94051FB38E1DFF8DFE1AEB50FD88FD86" box="[436,520,513,534]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/M22937" pageId="10" pageNumber="22" type="EnaNcbi">M22937</accessionNumber>
, Rawa Aopa Watumohai National Park, South-East
<collectingRegion id="49924CB28E1DFF8DFD8DEB4CFDFFFDA3" box="[547,639,541,563]" country="Indonesia" name="Sulawesi Utara" pageId="10" pageNumber="22">Sulawesi</collectingRegion>
. Scale =
<quantity id="4CAE2FB58E1DFF8DFE20EB68FE45FDDE" box="[398,453,569,590]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="22" unit="mm" value="1.0">1 mm</quantity>
</paragraph>
</caption>
<paragraph id="8BE982508E1DFF8DFF2CEBD0FDE7FBA7" blockId="10.[130,643,641,1730]" pageId="10" pageNumber="22">
teeth less robust relative to skull size and the postero-lateral chambers of the rostrum not greatly inflated (
<figureCitation id="136D9ED58E1DFF8DFE29EB97FE56FD71" box="[391,470,710,737]" captionStart="FIG" captionStartId="8.[130,143,1626,1647]" captionTargetBox="[273,1107,833,1601]" captionTargetId="figure@8.[235,1126,829,1604]" captionTargetPageId="8" captionText="FIG. 5. Dorsal view of the rostrum of four species of Hipposideros. A — H. boeadii (holotype), M28253, Rawa Aopa Watumohai National Park, South-East Sulawesi; B — H. papua, M16951, Biak Island, Papua Province; C — H. galeritus, M28250, Kakenauwe Nature Reserve, Buton Island, South-East Sulawesi; D —" figureDoi="http://doi.org/10.5281/zenodo.3944964" httpUri="https://zenodo.org/record/3944964/files/figure.png" pageId="10" pageNumber="22">Fig. 5</figureCitation>
). Comparison was made with a further five taxa that have
<taxonomicName id="4C56F9D38E1DFF8DFF2CEA5BFF39FCB4" box="[130,185,778,804]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="22" phylum="Chordata" rank="species" species="galeribeen">
<emphasis id="B9225E428E1DFF8DFF2CEA5BFF39FCB4" box="[130,185,778,804]" italics="true" pageId="10" pageNumber="22">been</emphasis>
</taxonomicName>
referred to
<taxonomicName id="4C56F9D38E1DFF8DFEFCEA5AFE5DFCB4" box="[338,477,779,804]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="22" phylum="Chordata" rank="species" species="cervinus">
<emphasis id="B9225E428E1DFF8DFEFCEA5AFE5DFCB4" box="[338,477,779,804]" italics="true" pageId="10" pageNumber="22">H. cervinus</emphasis>
</taxonomicName>
(Jenkins and Hill, 1981). These are
<emphasis id="B9225E428E1DFF8DFE39EA7CFDA3FCD7" box="[407,547,813,839]" italics="true" pageId="10" pageNumber="22">labuanensis</emphasis>
<bibRefCitation id="EFC7FFA18E1DFF8DFD83EA7CFF3DFCFA" author="TOMES, R. F." journalOrPublisher="Proceed- ings of the Zoological Society of London" pageId="10" pageNumber="22" pagination="536 - 540" part="1858" refId="ref8215" refString="TOMES, R. F. 1859. Notice of five species of bats in the collections of L. L. Dillwyn, Esq., M. P.: col- lected in Labuan by Mr James Motley. Proceed- ings of the Zoological Society of London, 1858: 536 - 540." title="Notice of five species of bats in the collections of L. L. Dillwyn, Esq., M. P.: col- lected in Labuan by Mr James Motley" type="journal article" year="1859">Tomes, 1859</bibRefCitation>
(described from Labuan Island and
<collectingCountry id="F341C2C08E1DFF8DFF2CEA20FF6CFC1C" box="[130,236,881,908]" name="Malaysia" pageId="10" pageNumber="22">Sarawak</collectingCountry>
),
<emphasis id="B9225E428E1DFF8DFEAEEA20FE01FC1B" box="[256,385,881,907]" italics="true" pageId="10" pageNumber="22">batchianus</emphasis>
<bibRefCitation id="EFC7FFA18E1DFF8DFE27EA20FDBFFC1B" author="MATSCHIE, P." box="[393,575,881,907]" journalOrPublisher="Abhandlungen hrsg. Von der Senckenbergischen Naturforschenden Ge- selleschaft" pageId="10" pageNumber="22" pagination="247 - 296" part="25" refId="ref7685" refString="MATSCHIE, P. 1901. Die Saugethiere der von Kukenthal auf Halmahera, Batjan und Nord Celebes gemachten Ausbeute. Abhandlungen hrsg. Von der Senckenbergischen Naturforschenden Ge- selleschaft, 25: 247 - 296." title="Die Saugethiere der von Kukenthal auf Halmahera, Batjan und Nord Celebes gemachten Ausbeute" type="journal article" year="1901">Matschie, 1901</bibRefCitation>
(Bat- jan
<collectingCountry id="F341C2C08E1DFF8DFF1FEAC2FF7CFC3D" box="[177,252,915,941]" name="Iceland" pageId="10" pageNumber="22">Island</collectingCountry>
, northern
<collectingCountry id="F341C2C08E1DFF8DFED7EAC2FE70FC3E" box="[377,496,915,942]" name="Indonesia" pageId="10" pageNumber="22">Moluccas</collectingCountry>
),
<emphasis id="B9225E428E1DFF8DFDA9EAC2FF16FC40" italics="true" pageId="10" pageNumber="22">schneider- si</emphasis>
(misprint for
<emphasis id="B9225E428E1DFF8DFEE9EAE7FE41FC40" box="[327,449,950,976]" italics="true" pageId="10" pageNumber="22">schneideri</emphasis>
) Thomas, 1904 (Sukaranda,
<collectingRegion id="49924CB28E1DFF8DFE8FEA89FE07FC62" box="[289,391,984,1010]" country="Indonesia" name="Sumatera Utara" pageId="10" pageNumber="22">Sumatra</collectingRegion>
),
<emphasis id="B9225E428E1DFF8DFE0BEA88FDA1FC62" box="[421,545,985,1010]" italics="true" pageId="10" pageNumber="22">misorensis</emphasis>
<bibRefCitation id="EFC7FFA18E1DFF8DFD9DEA88FF3DFB84" author="PETERS, W." bookContentInfo="70 plates" journalOrPublisher="Berlin" pageId="10" pageNumber="22" refId="ref8026" refString="PETERS, W. 1906. Museum Zoologicum Berolinense (Plates). Berlin, 70 plates." title="Museum Zoologicum Berolinense (Plates)" type="book" year="1906">Peters, 1906</bibRefCitation>
(Biak Island, Papua Province) and
<emphasis id="B9225E428E1DFF8DFDF1EAAAFF61FBA6" italics="true" pageId="10" pageNumber="22">ce- lebensis</emphasis>
<bibRefCitation id="EFC7FFA18E1DFF8DFF47ED4DFEEDFBA6" author="Sody" box="[233,365,1052,1078]" firstAuthor="Sody" pageId="10" pageNumber="22" pagination="42 - 55" refId="ref8318" refString="SODY, H. J. V. 1936. Seventeen new generic, specific gischer Forschungsreisen in Sumatra. Zoologi- and subspecific names for Dutch East Indian scher Anzeiger, 27: 722 - 724. mammals. Natuurkundig Tijdshrift voor Neder- THOMAS, O. 1913. On new mammals obtained by the landsch-Indie, 96: 42 - 55. Utkawa Expedition to Dutch New Guinea. Annals" type="journal article" year="1936">Sody, 1936</bibRefCitation>
(southern Sulawesi).
</paragraph>
<paragraph id="8BE982508E1DFF8DFF06ED6EFD03F952" blockId="10.[130,643,641,1730]" pageId="10" pageNumber="22">
<taxonomicName id="4C56F9D38E1DFF8DFF06ED6EFF56FBC9" authority="The" authorityName="The" box="[168,214,1087,1113]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="22" phylum="Chordata" rank="species" species="stenotis">The</taxonomicName>
noseleaf morphology of the holo- type of
<emphasis id="B9225E428E1DFF8DFF75ED30FEE7FBEB" box="[219,359,1121,1147]" italics="true" pageId="10" pageNumber="22">labuanensis</emphasis>
is difficult to determine since it is a dry skin. The forearm is longer (
<quantity id="4CAE2FB58E1DFF8DFF24EDF7FF77FB50" box="[138,247,1190,1216]" metricMagnitude="-2" metricUnit="m" metricValue="4.779999999999999" pageId="10" pageNumber="22" unit="mm" value="47.8">47.8 mm</quantity>
) but the dentition of the damaged skull is similar in size to that of
<emphasis id="B9225E428E1DFF8DFE51ED98FD03FB72" box="[511,643,1224,1250]" italics="true" pageId="10" pageNumber="22">
<taxonomicName id="4C56F9D38E1DFF8DFE51ED98FDFFFB72" authorityName="Bates &amp; Rossiter &amp; Suyanto &amp; Kingston" authorityYear="2007" box="[511,639,1224,1250]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="22" phylum="Chordata" rank="species" species="boeadii">H. boeadii</taxonomicName>
.
</emphasis>
However, unlike
<taxonomicName id="4C56F9D38E1DFF8DFECCEDBAFE69FA94" authorityName="Bates &amp; Rossiter &amp; Suyanto &amp; Kingston" authorityYear="2007" box="[354,489,1258,1284]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="22" phylum="Chordata" rank="species" species="boeadii">
<emphasis id="B9225E428E1DFF8DFECCEDBAFE69FA94" box="[354,489,1258,1284]" italics="true" pageId="10" pageNumber="22">H. boeadii</emphasis>
</taxonomicName>
, the lateral compartments of the rostrum in
<emphasis id="B9225E428E1DFF8DFDA2EC5DFF21FAD9" italics="true" pageId="10" pageNumber="22">labuanen- sis</emphasis>
are inflated (typical of
<taxonomicName id="4C56F9D38E1DFF8DFE15EC61FD9EFAD9" box="[443,542,1328,1353]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="22" phylum="Chordata" rank="species" species="cervinus">
<emphasis id="B9225E428E1DFF8DFE15EC61FD9EFAD9" box="[443,542,1328,1353]" italics="true" pageId="10" pageNumber="22">cervinus</emphasis>
</taxonomicName>
) where-
<taxonomicName id="4C56F9D38E1DFF8DFF2CEC03FF1AFAFC" box="[130,154,1362,1388]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="22" phylum="Chordata" rank="species" species="muscias">
<emphasis id="B9225E428E1DFF8DFF2CEC03FF1AFAFC" box="[130,154,1362,1388]" italics="true" pageId="10" pageNumber="22">as</emphasis>
</taxonomicName>
those of
<taxonomicName id="4C56F9D38E1DFF8DFEA9EC03FE03FAFB" authorityName="Bates &amp; Rossiter &amp; Suyanto &amp; Kingston" authorityYear="2007" box="[263,387,1361,1387]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="22" phylum="Chordata" rank="species" species="boeadii">
<emphasis id="B9225E428E1DFF8DFEA9EC03FE03FAFB" box="[263,387,1361,1387]" italics="true" pageId="10" pageNumber="22">H. boeadii</emphasis>
</taxonomicName>
are not enlarged. The taxon
<taxonomicName id="4C56F9D38E1DFF8DFF7EEC22FED6FA1D" box="[208,342,1395,1421]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="22" phylum="Chordata" rank="species" species="schneidersi">
<emphasis id="B9225E428E1DFF8DFF7EEC22FED6FA1D" box="[208,342,1395,1421]" italics="true" pageId="10" pageNumber="22">schneidersi</emphasis>
</taxonomicName>
was included as a syn- onym of
<emphasis id="B9225E428E1DFF8DFF5CECC7FE44FA20" box="[242,452,1430,1456]" italics="true" pageId="10" pageNumber="22">H. c. labuanensis</emphasis>
by Jenkins and Hill (1981). Although it is typical of
<emphasis id="B9225E428E1DFF8DFD9EECE8FF40FA64" italics="true" pageId="10" pageNumber="22">H. cer- vinus</emphasis>
in the morphology of the noseleaf and rostrum, it differs in having a more robust skull with significantly broader zygomata (116.1% of mastoid width), a character shared with
<taxonomicName id="4C56F9D38E1DFF8DFEB5EF35FE1CF9ED" authorityName="Bates &amp; Rossiter &amp; Suyanto &amp; Kingston" authorityYear="2007" box="[283,412,1635,1661]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="22" phylum="Chordata" rank="species" species="boeadii">
<emphasis id="B9225E428E1DFF8DFEB5EF35FE1CF9ED" box="[283,412,1635,1661]" italics="true" pageId="10" pageNumber="22">H. boeadii</emphasis>
</taxonomicName>
. These differences in the skull suggest that
<bibRefCitation id="EFC7FFA18E1DFF8DFE19EFD7FDC4F930" author="Tate" box="[439,580,1670,1696]" firstAuthor="Tate" pageId="10" pageNumber="22" pagination="353 - 393" refId="ref8389" refString="TATE, G. H. H. 1941. A review of the genus Hippo- and Magazine of Natural History, London, 12: sideros with special reference to Indo-Australian 212 - 217. species. Bulletin of the American Museum of THOMAS, O., and G. DORIA. 1886. Note intorno ad Natural History, 78: 353 - 393. alcune Chirotteri. Annali del Museo Civico di" type="journal article" year="1941">Tate (1941)</bibRefCitation>
may have been correct in treating
<taxonomicName id="4C56F9D38E1DFF8DFE77EFF8FD03F952" box="[473,643,1704,1730]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="22" phylum="Chordata" rank="species" species="schneidersi">
<emphasis id="B9225E428E1DFF8DFE77EFF8FD03F952" box="[473,643,1704,1730]" italics="true" pageId="10" pageNumber="22">H. schneidersi</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BE982508E1DFF8DFD00E9F5FB2EFE62" blockId="10.[686,1199,163,498]" pageId="10" pageNumber="22">
as a distinct species. The taxon
<emphasis id="B9225E428E1DFF8DFB83E9F2FB2EFF2D" box="[1069,1198,163,189]" italics="true" pageId="10" pageNumber="22">batchianus</emphasis>
is poorly known (
<bibRefCitation id="EFC7FFA18E1DFF8DFC2EE997FC7EFF71" author="HILL, J. E." box="[896,1022,198,225]" journalOrPublisher="Bulletin of the British Museum (Natural History), Zoology" pageId="10" pageNumber="22" pagination="1 - 129" part="11" refId="ref8178" refString="HILL, J. E. 1963. A revison of the genus Hipposideros. Bulletin of the British Museum (Natural History), Zoology, 11: 1 - 129" title="A revison of the genus Hipposideros" type="journal article" year="1963">Hill, 1963</bibRefCitation>
). It was exam- ined by
<bibRefCitation id="EFC7FFA18E1DFF8DFCBEE9B8FC1BFE92" author="Tate" box="[784,923,232,259]" firstAuthor="Tate" pageId="10" pageNumber="22" pagination="353 - 393" refId="ref8389" refString="TATE, G. H. H. 1941. A review of the genus Hippo- and Magazine of Natural History, London, 12: sideros with special reference to Indo-Australian 212 - 217. species. Bulletin of the American Museum of THOMAS, O., and G. DORIA. 1886. Note intorno ad Natural History, 78: 353 - 393. alcune Chirotteri. Annali del Museo Civico di" type="journal article" year="1941">Tate (1941)</bibRefCitation>
who included it in the
<taxonomicName id="4C56F9D38E1DFF8DFD00E85BFC96FEB4" authorityName="Cantor" authorityYear="1846" box="[686,790,266,292]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="22" phylum="Chordata" rank="species" species="galeritus">
<emphasis id="B9225E428E1DFF8DFD00E85BFC96FEB4" box="[686,790,266,292]" italics="true" pageId="10" pageNumber="22">galeritus</emphasis>
</taxonomicName>
group but was subsequently re- ferred to
<taxonomicName id="4C56F9D38E1DFF8DFC85E87CFD6DFEF9" authority="(Jenkins and Hill" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="22" phylum="Chordata" rank="species" species="cervinus">
<emphasis id="B9225E428E1DFF8DFC85E87CFC3BFED6" box="[811,955,301,326]" italics="true" pageId="10" pageNumber="22">H. cervinus</emphasis>
(Jenkins and Hill, 1981
</taxonomicName>
). Its forearm, tail and upper toothrow lengths of
<quantity id="4CAE2FB58E1DFF8DFC86E820FC13FE1B" box="[808,915,369,396]" metricMagnitude="-2" metricUnit="m" metricValue="4.75" pageId="10" pageNumber="22" unit="mm" value="47.5">47.5 mm</quantity>
,
<quantity id="4CAE2FB58E1DFF8DFC33E820FC6FFE1B" box="[925,1007,369,396]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="10" pageNumber="22" unit="mm" value="25.0">25 mm</quantity>
and
<quantity id="4CAE2FB58E1DFF8DFB89E820FB03FE1B" box="[1063,1155,369,395]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="10" pageNumber="22" unit="mm" value="6.4">6.4 mm</quantity>
, re- spectively, suggest that it is a typical
<emphasis id="B9225E428E1DFF8DFBF2E8C5FD73FE5F" italics="true" pageId="10" pageNumber="22">H. cer- vinus.</emphasis>
The taxon
<emphasis id="B9225E428E1DFF8DFC26E8E7FB84FE5F" box="[904,1028,438,463]" italics="true" pageId="10" pageNumber="22">misorensis</emphasis>
is also poor- ly known, with all the current informa-
</paragraph>
<paragraph id="8BE982508E1DFF8DFD00E8ABFCFEFD71" blockId="10.[686,1199,506,1730]" pageId="10" pageNumber="22">
tion based on a plate published by
<bibRefCitation id="EFC7FFA18E1DFF8DFBC8E8ABFC82FDA7" author="PETERS, W." bookContentInfo="70 plates" journalOrPublisher="Berlin" pageId="10" pageNumber="22" refId="ref8026" refString="PETERS, W. 1906. Museum Zoologicum Berolinense (Plates). Berlin, 70 plates." title="Museum Zoologicum Berolinense (Plates)" type="book" year="1906">Peters (1906)</bibRefCitation>
. However, according to Jenkins and Hill (1981), the noseleaf closely resem- bles that of
<emphasis id="B9225E428E1DFF8DFCE8EB30FC5AFDEA" box="[838,986,609,634]" italics="true" pageId="10" pageNumber="22">
<taxonomicName id="4C56F9D38E1DFF8DFCE8EB30FC56FDEA" box="[838,982,609,634]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="22" phylum="Chordata" rank="species" species="cervinus">H. cervinus</taxonomicName>
.
</emphasis>
Jenkins and Hill (1981) included
<emphasis id="B9225E428E1DFF8DFCC1EBD2FC68FD0D" box="[879,1000,643,669]" italics="true" pageId="10" pageNumber="22">celebensis</emphasis>
as a synonym of
<emphasis id="B9225E428E1DFF8DFD00EBF7FCD0FD2F" box="[686,848,678,703]" italics="true" pageId="10" pageNumber="22">
H. c.
<taxonomicName id="4C56F9D38E1DFF8DFD43EBF7FCD0FD2F" box="[749,848,678,703]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="22" phylum="Chordata" rank="species" species="cervinus">cervinus</taxonomicName>
</emphasis>
and the two taxa appear to be indistinguishable.
</paragraph>
<paragraph id="8BE982508E1DFF8DFD7DEBBBFBC2FAB7" blockId="10.[686,1199,506,1730]" pageId="10" pageNumber="22">
In addition to
<taxonomicName id="4C56F9D38E1DFF8DFCD7EBBBFB80FC93" box="[889,1024,746,771]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="22" phylum="Chordata" rank="species" species="cervinus">
<emphasis id="B9225E428E1DFF8DFCD7EBBBFB80FC93" box="[889,1024,746,771]" italics="true" pageId="10" pageNumber="22">H. cervinus</emphasis>
</taxonomicName>
and H.
<emphasis id="B9225E428E1DFF8DFBF0EBB8FD50FCB5" italics="true" pageId="10" pageNumber="22">galeritus</emphasis>
, there are six species which not only have two supplementary leaflets but also have the inner leaflet shorter than the outer, a character shared with
<emphasis id="B9225E428E1DFF8DFC67EA22FBCEFC1C" box="[969,1102,882,908]" italics="true" pageId="10" pageNumber="22">
<taxonomicName id="4C56F9D38E1DFF8DFC67EA22FBCAFC1C" authorityName="Bates &amp; Rossiter &amp; Suyanto &amp; Kingston" authorityYear="2007" box="[969,1098,882,908]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="22" phylum="Chordata" rank="species" species="boeadii">H. boeadii</taxonomicName>
.
</emphasis>
Four of these,
<taxonomicName id="4C56F9D38E1DFF8DFD50EAC4FB2EFC3E" authority="Thomas and Doria, 1886" authorityName="Thomas and Doria" authorityYear="1886" box="[766,1198,916,943]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="22" phylum="Chordata" rank="species" species="muscinus">
<emphasis id="B9225E428E1DFF8DFD50EAC4FCEBFC3E" box="[766,875,917,942]" italics="true" pageId="10" pageNumber="22">muscinus</emphasis>
Thomas and Doria, 1886
</taxonomicName>
(described from Fly River,
<collectingRegion id="49924CB28E1DFF8DFBB2EAE6FBE4FC41" box="[1052,1124,951,977]" country="Indonesia" name="Papua Barat" pageId="10" pageNumber="22">Papua</collectingRegion>
New
<collectingCountry id="F341C2C08E1DFF8DFD00EA88FC87FC64" box="[686,775,985,1012]" name="Guinea-Bissau" pageId="10" pageNumber="22">Guinea</collectingCountry>
),
<taxonomicName id="4C56F9D38E1DFF8DFC8AEA88FBE0FC63" authority="Thomas, 1913" authorityName="Thomas" authorityYear="1913" box="[804,1120,985,1011]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="22" phylum="Chordata" rank="species" species="wollastoni">
<emphasis id="B9225E428E1DFF8DFC8AEA88FC1EFC63" box="[804,926,985,1011]" italics="true" pageId="10" pageNumber="22">wollastoni</emphasis>
<bibRefCitation id="EFC7FFA18E1DFF8DFC00EA88FBE0FC63" author="THOMAS, O." box="[942,1120,985,1011]" journalOrPublisher="Annals and Magazine of Natural History, London" pageId="10" pageNumber="22" pagination="212 - 217" part="12" refId="ref8215" refString="THOMAS, O. 1913. On new mammals obtained by the Utkawa Expedition to Dutch New Guinea. Annals and Magazine of Natural History, London, 12: 212 - 217." title="On new mammals obtained by the Utkawa Expedition to Dutch New Guinea" type="journal article" year="1913">Thomas, 1913</bibRefCitation>
</taxonomicName>
(Uta- kawa River, Irian Jaya),
<taxonomicName id="4C56F9D38E1DFF8DFC73EAADFD69FBA8" authority="Matschie, 1903" authorityName="Matschie" authorityYear="1903" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="22" phylum="Chordata" rank="species" species="semoni">
<emphasis id="B9225E428E1DFF8DFC73EAADFBB0FB85" box="[989,1072,1020,1045]" italics="true" pageId="10" pageNumber="22">semoni</emphasis>
<bibRefCitation id="EFC7FFA18E1DFF8DFB95EAAAFD69FBA8" author="MATSCHIE, P." journalOrPublisher="Denkschriften der MedizinischNaturwissenschaftlichen Gesellschaft zu Jena, 8: 771" pageId="10" pageNumber="22" pagination="129 - 136" part="6" refId="ref7769" refString="MATSCHIE, P. 1903. Die Chiropteren, Insectivoren und Muriden der Semon'schen Forschungsreise (R. W. Semon, Zoologische Forschungsreisen in Australien und dem Malayischen Archipel. 1891 - 1893). Denkschriften der MedizinischNaturwissenschaftlichen Gesellschaft zu Jena, 8: 771 - 778 [Heft 6: 129 - 136]." title="Die Chiropteren, Insectivoren und Muriden der Semon'schen Forschungsreise (R. W. Semon, Zoologische Forschungsreisen in Australien und dem Malayischen Archipel. 1891 - 1893)" type="journal article" year="1903">Matschie, 1903</bibRefCitation>
</taxonomicName>
(
<collectingRegion id="49924CB28E1DFF8DFD55ED4CFC0AFBA7" box="[763,906,1053,1079]" country="Australia" name="Queensland" pageId="10" pageNumber="22">Queensland</collectingRegion>
,
<collectingCountry id="F341C2C08E1DFF8DFC36ED4CFB88FBA8" box="[920,1032,1053,1080]" name="Australia" pageId="10" pageNumber="22">Australia</collectingCountry>
), and
<emphasis id="B9225E428E1DFF8DFBFAED4FFB2EFBA7" box="[1108,1198,1054,1079]" italics="true" pageId="10" pageNumber="22">stenotis</emphasis>
<bibRefCitation id="EFC7FFA18E1DFF8DFD00ED11FCD7FBCA" author="THOMAS, O." box="[686,855,1088,1114]" journalOrPublisher="Annals and Magazine of Natural History, London" pageId="10" pageNumber="22" pagination="212 - 217" part="12" refId="ref8215" refString="THOMAS, O. 1913. On new mammals obtained by the Utkawa Expedition to Dutch New Guinea. Annals and Magazine of Natural History, London, 12: 212 - 217." title="On new mammals obtained by the Utkawa Expedition to Dutch New Guinea" type="journal article" year="1913">Thomas, 1913</bibRefCitation>
(
<collectingRegion id="49924CB28E1DFF8DFCC8ED11FBC3FBCA" box="[870,1091,1088,1114]" country="Australia" name="Northern Territory" pageId="10" pageNumber="22">Northern Territory</collectingRegion>
, Australia) were included in the
<emphasis id="B9225E428E1DFF8DFC70ED33FBB6FBEC" box="[990,1078,1122,1148]" italics="true" pageId="10" pageNumber="22">cyclops</emphasis>
group by
<bibRefCitation id="EFC7FFA18E1DFF8DFD00EDD5FCB8FB0F" author="HILL, J. E." box="[686,824,1156,1183]" journalOrPublisher="Bulletin of the British Museum (Natural History), Zoology" pageId="10" pageNumber="22" pagination="1 - 129" part="11" refId="ref8178" refString="HILL, J. E. 1963. A revison of the genus Hipposideros. Bulletin of the British Museum (Natural History), Zoology, 11: 1 - 129" title="A revison of the genus Hipposideros" type="journal article" year="1963">Hill (1963)</bibRefCitation>
. Two others,
<taxonomicName id="4C56F9D38E1DFF8DFC4EEDD4FCA8FB51" authority="Hill, 1985" authorityName="Hill" authorityYear="1985" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="22" phylum="Chordata" rank="species" species="corynophyllus">
<emphasis id="B9225E428E1DFF8DFC4EEDD4FB2EFB0E" box="[992,1198,1156,1182]" italics="true" pageId="10" pageNumber="22">H. corynophyllus</emphasis>
<bibRefCitation id="EFC7FFA18E1DFF8DFD00EDF7FCA8FB51" author="HILL, J. E." box="[686,808,1190,1217]" journalOrPublisher="Mammalia" pageId="10" pageNumber="22" pagination="525 - 535" part="49" refId="ref7174" refString="HILL, J. E. 1985. Records of bats (Chiroptera) from New Guinea, with the description of a new Hipposideros (Hipposideridae). Mammalia, 49: 525 - 535." title="Records of bats (Chiroptera) from New Guinea, with the description of a new Hipposideros (Hipposideridae)" type="journal article" year="1985">Hill, 1985</bibRefCitation>
</taxonomicName>
(West Sepik, Papua New Guinea) and
<taxonomicName id="4C56F9D38E1DFF8DFCBEED98FBBBFB73" authority="Flannery" authorityName="Flannery" box="[784,1083,1225,1251]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="22" phylum="Chordata" rank="species" species="edwardshilli">
<emphasis id="B9225E428E1DFF8DFCBEED98FC49FB73" box="[784,969,1225,1251]" italics="true" pageId="10" pageNumber="22">H. edwardshilli</emphasis>
Flannery
</taxonomicName>
and Col- gan 1993 (Bewani Mountains, Papua New
<collectingCountry id="F341C2C08E1DFF8DFD00EC5CFC85FAB8" box="[686,773,1293,1320]" name="Guinea-Bissau" pageId="10" pageNumber="22">Guinea</collectingCountry>
) were added subsequently.
</paragraph>
<paragraph id="8BE982508E1DFF8DFD7DEC61FB2EF952" blockId="10.[686,1199,506,1730]" pageId="10" pageNumber="22">
In comparison to
<taxonomicName id="4C56F9D38E1DFF8DFC07EC61FBA8FAD9" authorityName="Bates &amp; Rossiter &amp; Suyanto &amp; Kingston" authorityYear="2007" box="[937,1064,1327,1353]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="22" phylum="Chordata" rank="species" species="boeadii">
<emphasis id="B9225E428E1DFF8DFC07EC61FBA8FAD9" box="[937,1064,1327,1353]" italics="true" pageId="10" pageNumber="22">H. boeadii</emphasis>
</taxonomicName>
, H.
<emphasis id="B9225E428E1DFF8DFBCEEC61FD57FAFB" italics="true" pageId="10" pageNumber="22">muscinus</emphasis>
has larger, more inflated anterior chambers of the rostrum, larger cochleae, which are more closely situated together, and with the first upper premolar (P
<superScript id="7C232F188E1DFF8DFC57ECE2FB83FA56" attach="left" box="[1017,1027,1459,1478]" fontSize="8" pageId="10" pageNumber="22">2</superScript>
) less reduced and situated in the toothrow. The posterior leaf of the noseleaf has a small tubercle which is not present in
<emphasis id="B9225E428E1DFF8DFC5DEF71FBF8F9A9" box="[1011,1144,1567,1593]" italics="true" pageId="10" pageNumber="22">
<taxonomicName id="4C56F9D38E1DFF8DFC5DEF71FBF4F9A9" authorityName="Bates &amp; Rossiter &amp; Suyanto &amp; Kingston" authorityYear="2007" box="[1011,1140,1567,1593]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="22" phylum="Chordata" rank="species" species="boeadii">H. boeadii</taxonomicName>
.
</emphasis>
The taxon
<taxonomicName id="4C56F9D38E1DFF8DFD56EF10FCF2F9CB" authorityName="Thomas" authorityYear="1913" box="[760,882,1601,1627]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="22" phylum="Chordata" rank="species" species="wollastoni">
<emphasis id="B9225E428E1DFF8DFD56EF10FCF2F9CB" box="[760,882,1601,1627]" italics="true" pageId="10" pageNumber="22">wollastoni</emphasis>
</taxonomicName>
has a distinctive noseleaf. The intermediate leaf has a large projecting median eminence and the upper edge of the leaf has two lateral elevations. In addition,
</paragraph>
<paragraph id="8BE982508E1CFF8CFF2CE9F2FDE9FD0E" blockId="11.[130,646,163,1730]" pageId="11" pageNumber="23">
there is a transverse supplementary struc- ture which is developed from the posterior face of the posterior leaf and is almost equal in height to the posterior leaf. There are also significant differences in the skull, most notably in the greatly expanded cochleae.
<taxonomicName id="4C56F9D38E1CFF8CFF2CE820FEFCFE1B" authorityName="Matschie" authorityYear="1903" box="[130,380,369,395]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="23" phylum="Chordata" rank="species" species="semoni">
<emphasis id="B9225E428E1CFF8CFF2CE820FEFCFE1B" box="[130,380,369,395]" italics="true" pageId="11" pageNumber="23">Hipposideros semoni</emphasis>
</taxonomicName>
,
<taxonomicName id="4C56F9D38E1CFF8CFE20E823FD8FFE1B" box="[398,527,370,395]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="23" phylum="Chordata" rank="species" species="stenotis">
<emphasis id="B9225E428E1CFF8CFE20E823FD8FFE1B" box="[398,527,370,395]" italics="true" pageId="11" pageNumber="23">H. stenotis</emphasis>
</taxonomicName>
,
<emphasis id="B9225E428E1CFF8CFD8EE823FD05FE1B" box="[544,645,370,395]" italics="true" pageId="11" pageNumber="23">H. cory-</emphasis>
nophyllus and
<taxonomicName id="4C56F9D38E1CFF8CFE98E8C5FE71FE3D" box="[310,497,403,429]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="23" phylum="Chordata" rank="species" species="edwardshilli">
<emphasis id="B9225E428E1CFF8CFE98E8C5FE71FE3D" box="[310,497,403,429]" italics="true" pageId="11" pageNumber="23">H. edwardshilli</emphasis>
</taxonomicName>
can be dis- tinguished externally from
<taxonomicName id="4C56F9D38E1CFF8CFE78E8E7FDD9FE5F" authorityName="Bates &amp; Rossiter &amp; Suyanto &amp; Kingston" authorityYear="2007" box="[470,601,437,463]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="23" phylum="Chordata" rank="species" species="boeadii">
<emphasis id="B9225E428E1CFF8CFE78E8E7FDD9FE5F" box="[470,601,437,463]" italics="true" pageId="11" pageNumber="23">H. boeadii</emphasis>
</taxonomicName>
by well developed club-shaped projections on the intermediate and posterior leaves of the noseleaf. In the skulls of
<taxonomicName id="4C56F9D38E1CFF8CFE76EB4CFDD0FDA6" authorityName="Matschie" authorityYear="1903" box="[472,592,541,566]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="23" phylum="Chordata" rank="species" species="semoni">
<emphasis id="B9225E428E1CFF8CFE76EB4CFDD0FDA6" box="[472,592,541,566]" italics="true" pageId="11" pageNumber="23">H. semoni</emphasis>
</taxonomicName>
and
<taxonomicName id="4C56F9D38E1CFF8CFF2CEB6EFE82FDC8" box="[130,258,575,600]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="23" phylum="Chordata" rank="species" species="stenotis">
<emphasis id="B9225E428E1CFF8CFF2CEB6EFE82FDC8" box="[130,258,575,600]" italics="true" pageId="11" pageNumber="23">H. stenotis</emphasis>
</taxonomicName>
, the cochleae are enormous and almost touching; the posterior commissure of M
<superScript id="7C232F188E1CFF8CFF13EB2FFF47FD01" attach="left" box="[189,199,638,657]" fontSize="8" pageId="11" pageNumber="23">3</superScript>
is not greatly reduced (
<bibRefCitation id="EFC7FFA18E1CFF8CFE71EBD2FDDCFD0E" author="HILL, J. E." box="[479,604,643,670]" journalOrPublisher="Bulletin of the British Museum (Natural History), Zoology" pageId="11" pageNumber="23" pagination="1 - 129" part="11" refId="ref8178" refString="HILL, J. E. 1963. A revison of the genus Hipposideros. Bulletin of the British Museum (Natural History), Zoology, 11: 1 - 129" title="A revison of the genus Hipposideros" type="journal article" year="1963">Hill, 1963</bibRefCitation>
).
</paragraph>
<paragraph id="8BE982508E1CFF8CFF06EBF4FE0BFB50" blockId="11.[130,646,163,1730]" pageId="11" pageNumber="23">
The taxon
<taxonomicName id="4C56F9D38E1CFF8CFE8BEBF7FE64FD2F" baseAuthorityName="Lesueur and Petit" baseAuthorityYear="1807" box="[293,484,677,703]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="23" phylum="Chordata" rank="species" species="crumeniferus">
<emphasis id="B9225E428E1CFF8CFE8BEBF7FE64FD2F" box="[293,484,677,703]" italics="true" pageId="11" pageNumber="23">H. crumeniferus</emphasis>
</taxonomicName>
(Lesueur and Petit, 1807) (described from Timor) was listed as incertae sedis by Corbet and Hill (1992), pending the rediscovery of the hol- otype or the collection of topotypes. Pre- viously,
<bibRefCitation id="EFC7FFA18E1CFF8CFF48EA00FEEFFCFA" author="Tate" box="[230,367,848,875]" firstAuthor="Tate" pageId="11" pageNumber="23" pagination="353 - 393" refId="ref8389" refString="TATE, G. H. H. 1941. A review of the genus Hippo- and Magazine of Natural History, London, 12: sideros with special reference to Indo-Australian 212 - 217. species. Bulletin of the American Museum of THOMAS, O., and G. DORIA. 1886. Note intorno ad Natural History, 78: 353 - 393. alcune Chirotteri. Annali del Museo Civico di" type="journal article" year="1941">Tate (1941)</bibRefCitation>
suggested that it might have affinities to
<taxonomicName id="4C56F9D38E1CFF8CFECCEA22FE56FC1C" box="[354,470,883,908]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="23" phylum="Chordata" rank="species" species="papua">
<emphasis id="B9225E428E1CFF8CFECCEA22FE56FC1C" box="[354,470,883,908]" italics="true" pageId="11" pageNumber="23">H. papua</emphasis>
</taxonomicName>
(Thomas and Doria, 1886) (Biak Island, Papua Province). However, based on the plate in the Atlas of Lesueur and Petit (1807),
<bibRefCitation id="EFC7FFA18E1CFF8CFE19EA88FDBCFC64" author="HILL, J. E." box="[439,572,985,1012]" journalOrPublisher="Bulletin of the British Museum (Natural History), Zoology" pageId="11" pageNumber="23" pagination="1 - 129" part="11" refId="ref8178" refString="HILL, J. E. 1963. A revison of the genus Hipposideros. Bulletin of the British Museum (Natural History), Zoology, 11: 1 - 129" title="A revison of the genus Hipposideros" type="journal article" year="1963">Hill (1963)</bibRefCitation>
noted that unlike
<taxonomicName id="4C56F9D38E1CFF8CFEA4EAADFEF8FB85" box="[266,376,1020,1045]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="23" phylum="Chordata" rank="species" species="papua">
<emphasis id="B9225E428E1CFF8CFEA4EAADFEF8FB85" box="[266,376,1020,1045]" italics="true" pageId="11" pageNumber="23">H. papua</emphasis>
</taxonomicName>
the posterior (second) leaflet does not extend beneath the anterior part of the horseshoe; this would also dis- tinguish it from
<emphasis id="B9225E428E1CFF8CFE91ED32FE42FBEC" box="[319,450,1122,1148]" italics="true" pageId="11" pageNumber="23">
<taxonomicName id="4C56F9D38E1CFF8CFE91ED32FE3EFBEC" authorityName="Bates &amp; Rossiter &amp; Suyanto &amp; Kingston" authorityYear="2007" box="[319,446,1122,1148]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="23" phylum="Chordata" rank="species" species="boeadii">H. boeadii</taxonomicName>
.
</emphasis>
<bibRefCitation id="EFC7FFA18E1CFF8CFE67ED33FDCCFBED" author="HILL, J. E." box="[457,588,1122,1149]" journalOrPublisher="Bulletin of the British Museum (Natural History), Zoology" pageId="11" pageNumber="23" pagination="1 - 129" part="11" refId="ref8178" refString="HILL, J. E. 1963. A revison of the genus Hipposideros. Bulletin of the British Museum (Natural History), Zoology, 11: 1 - 129" title="A revison of the genus Hipposideros" type="journal article" year="1963">Hill (1963)</bibRefCitation>
sug- gested that it was referable to
<taxonomicName id="4C56F9D38E1CFF8CFE71EDD4FDE5FB0E" box="[479,613,1157,1182]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="23" phylum="Chordata" rank="species" species="cervinus">
<emphasis id="B9225E428E1CFF8CFE71EDD4FDE5FB0E" box="[479,613,1157,1182]" italics="true" pageId="11" pageNumber="23">H. cervinus</emphasis>
</taxonomicName>
or a closely related form.
</paragraph>
<paragraph id="8BE982508E1CFF8CFF06ED98FB2FFCB3" blockId="11.[130,646,163,1730]" lastBlockId="11.[683,1199,163,1383]" pageId="11" pageNumber="23">
<taxonomicName id="4C56F9D38E1CFF8CFF06ED98FE25FB73" authorityName="Bates &amp; Rossiter &amp; Suyanto &amp; Kingston" authorityYear="2007" box="[168,421,1225,1251]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="23" phylum="Chordata" rank="species" species="boeadii">
<emphasis id="B9225E428E1CFF8CFF06ED98FE25FB73" box="[168,421,1225,1251]" italics="true" pageId="11" pageNumber="23">Hipposideros boeadii</emphasis>
</taxonomicName>
appears most sim- ilar to
<emphasis id="B9225E428E1CFF8CFF76EDBDFED1FA95" box="[216,337,1260,1285]" italics="true" pageId="11" pageNumber="23">
<taxonomicName id="4C56F9D38E1CFF8CFF76EDBDFECDFA95" box="[216,333,1260,1285]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="23" phylum="Chordata" rank="species" species="papua">H. papua</taxonomicName>
.
</emphasis>
This taxon has not been recorded from Sulawesi but is known from the Molucca Islands and western New
<collectingCountry id="F341C2C08E1CFF8CFF2CEC00FF57FAFB" box="[130,215,1361,1387]" name="Guinea-Bissau" pageId="11" pageNumber="23">Guinea</collectingCountry>
(
<bibRefCitation id="EFC7FFA18E1CFF8CFF43EC00FE27FAFC" author="FLANNERY, T. F." bookContentInfo="464 pp." box="[237,423,1361,1388]" journalOrPublisher="Reed Books, Aus- tralia" pageId="11" pageNumber="23" refId="ref7659" refString="FLANNERY, T. F. 1995. Mammals of the South-West Pacific and Moluccan Islands. Reed Books, Aus- tralia, 464 pp." title="Mammals of the South-West Pacific and Moluccan Islands" type="book" year="1995">Flannery, 1995</bibRefCitation>
). With a forearm length of
<quantity id="4CAE2FB58E1CFF8CFEADEC25FE36FA1E" box="[259,438,1396,1423]" metricMagnitude="-2" metricUnit="m" metricValue="5.16" metricValueMax="5.36" metricValueMin="4.96" pageId="11" pageNumber="23" unit="mm" value="51.6" valueMax="53.6" valueMin="49.6">49.653.6 mm</quantity>
and a condylocanine length of
<quantity id="4CAE2FB58E1CFF8CFEFDECC7FD84FA21" box="[339,516,1430,1457]" metricMagnitude="-2" metricUnit="m" metricValue="1.7149999999999999" metricValueMax="1.76" metricValueMin="1.67" pageId="11" pageNumber="23" unit="mm" value="17.15" valueMax="17.6" valueMin="16.7">16.717.6 mm</quantity>
, H. papua considerably exceeds
<taxonomicName id="4C56F9D38E1CFF8CFE2DECE8FE7FFA42" authorityName="Bates &amp; Rossiter &amp; Suyanto &amp; Kingston" authorityYear="2007" box="[387,511,1464,1490]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="23" phylum="Chordata" rank="species" species="boeadii">
<emphasis id="B9225E428E1CFF8CFE2DECE8FE7FFA42" box="[387,511,1464,1490]" italics="true" pageId="11" pageNumber="23">H. boeadii</emphasis>
</taxonomicName>
(
<quantity id="4CAE2FB58E1CFF8CFDA0ECE9FF30FA65" metricMagnitude="-2" metricUnit="m" metricValue="4.16" metricValueMax="4.27" metricValueMin="4.05" pageId="11" pageNumber="23" unit="mm" value="41.6" valueMax="42.7" valueMin="40.5">40.542.7 mm</quantity>
and
<quantity id="4CAE2FB58E1CFF8CFF47EC8BFE13FA65" box="[233,403,1498,1525]" metricMagnitude="-2" metricUnit="m" metricValue="1.535" metricValueMax="1.55" metricValueMin="1.52" pageId="11" pageNumber="23" unit="mm" value="15.35" valueMax="15.5" valueMin="15.2">15.215.5 mm</quantity>
respectively) in size (
<tableCitation id="C6D4B7EB8E1CFF8CFF24ECACFF72F987" box="[138,242,1533,1559]" captionStart="TABLE" captionStartId="4.[86,99,647,674]" captionTargetPageId="4" captionText="TABLE 1. External measurements (in mm) and body mass (in g) of four species of Hipposideros recently collected in Sulawesi. Used abbreviations: FA — forearm length; 5MET, 4MET, 3MET — fifth, fourth, and third metacarpal lengths; 5MET/FA× 100 — % length of fifth metacarpal relative to forearm length; 3D1P, 3D2P — length of first and second phalanges of the third digit; 3D1P/3MET × 100 — % length of first phalanx of the third digit relative to third metacarpal length; TAIL — tail length; HF — foot; TIBIA — tibia; E — ear and WGT — body mass. Sample sizes differing from those reported under n are given in parentheses. Mean and SD are included when sample size is ≥ 3" httpUri="http://table.plazi.org/id/DF29D2D88E13FF83FFF8EBD6FD8AFC83" pageId="11" pageNumber="23" tableUuid="DF29D2D88E13FF83FFF8EBD6FD8AFC83">Tables 1</tableCitation>
and
<tableCitation id="C6D4B7EB8E1CFF8CFE9CECACFEC3F987" box="[306,323,1533,1559]" captionStart="TABLE" captionStartId="9.[89,102,641,668]" captionText="TABLE 2. Cranial and dental measurements (in mm) of four species of Hipposideros recently collected in Sulawesi. Used abbreviations: GTL — greatest length of skull; CBL — condylobasal length; CCL — condylo-canine length; ZB — zygomatic breadth; BB — breadth of braincase; MW — mastoid width; ZB/MW × 100 — % breadth of zygomata relative to mastoid breadth; RW — rostral width; PC — postorbital constriction; CM3 — upper toothrow length; CM3/CCL × 100 — % length of upper toothrow relative to condylo-canine length; M3M3 — palatal breadth; CM — lower toothrow length; M — mandible length. Sample sizes 3 differing from those reported under n are given in parentheses. Mean and SD are included when sample size is ≥ 3" pageId="11" pageNumber="23">2</tableCitation>
). In addition, in
<taxonomicName id="4C56F9D38E1CFF8CFDBAECACFD05F986" box="[532,645,1533,1558]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="23" phylum="Chordata" rank="species" species="papua">
<emphasis id="B9225E428E1CFF8CFDBAECACFD05F986" box="[532,645,1533,1558]" italics="true" pageId="11" pageNumber="23">H. papua</emphasis>
</taxonomicName>
there is usually (but not always) a very small third supplementary leaflet to the noseleaf (Hill and Rozendaal, 1989); this was not detected in the four specimens of
<taxonomicName id="4C56F9D38E1CFF8CFF2CEFF8FF7FF952" authorityName="Bates &amp; Rossiter &amp; Suyanto &amp; Kingston" authorityYear="2007" box="[130,255,1704,1730]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="23" phylum="Chordata" rank="species" species="boeadii">
<emphasis id="B9225E428E1CFF8CFF2CEFF8FF7FF952" box="[130,255,1704,1730]" italics="true" pageId="11" pageNumber="23">H. boeadii</emphasis>
</taxonomicName>
currently known from Sulawesi. The wing morphology also appears to differ slightly between the two species (although current sample sizes are small). In
<emphasis id="B9225E428E1CFF8CFBFDE9B8FD5BFEB4" italics="true" pageId="11" pageNumber="23">
<taxonomicName id="4C56F9D38E1CFF8CFBFDE9B8FB2EFE92" box="[1107,1198,232,258]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="23" phylum="Chordata" rank="species" species="boein">H. boe-</taxonomicName>
adii
</emphasis>
, the fifth metacarpal is relatively longer, 70.574.8% (
<emphasis id="B9225E428E1CFF8CFC16E87CFC47FED6" box="[952,967,301,326]" italics="true" pageId="11" pageNumber="23">n</emphasis>
= 4) of forearm length; in
<taxonomicName id="4C56F9D38E1CFF8CFC89E81EFC16FEF8" box="[807,918,335,360]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="23" phylum="Chordata" rank="species" species="papua">
<emphasis id="B9225E428E1CFF8CFC89E81EFC16FEF8" box="[807,918,335,360]" italics="true" pageId="11" pageNumber="23">H. papua</emphasis>
</taxonomicName>
, the comparable figure is 65.1 65.6% (
<emphasis id="B9225E428E1CFF8CFCDEE820FCFFFE1A" box="[880,895,369,394]" italics="true" pageId="11" pageNumber="23">n</emphasis>
= 2). The skull of
<emphasis id="B9225E428E1CFF8CFBCCE820FD66FE3C" italics="true" pageId="11" pageNumber="23">
<taxonomicName id="4C56F9D38E1CFF8CFBCCE820FB2EFE1A" box="[1122,1198,368,394]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="23" phylum="Chordata" rank="species" species="bonophyllus">H. bo-</taxonomicName>
eadii
</emphasis>
is also long in comparison to fore- arm length. In
<taxonomicName id="4C56F9D38E1CFF8CFCF3E8E4FC5AFE5E" authorityName="Bates &amp; Rossiter &amp; Suyanto &amp; Kingston" authorityYear="2007" box="[861,986,436,462]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="23" phylum="Chordata" rank="species" species="boeadii">
<emphasis id="B9225E428E1CFF8CFCF3E8E4FC5AFE5E" box="[861,986,436,462]" italics="true" pageId="11" pageNumber="23">H. boeadii</emphasis>
</taxonomicName>
, it is 36.238.2% (
<emphasis id="B9225E428E1CFF8CFD1BE886FD44FE60" box="[693,708,471,496]" italics="true" pageId="11" pageNumber="23">n</emphasis>
= 4) (CCL/FA × 100), whereas in
<emphasis id="B9225E428E1CFF8CFBCCE886FD57FD82" italics="true" pageId="11" pageNumber="23">
<taxonomicName id="4C56F9D38E1CFF8CFBCCE886FB2EFE60" box="[1122,1198,471,496]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="23" phylum="Chordata" rank="species" species="pathe">H. pa-</taxonomicName>
pua
</emphasis>
, it is 33.234.3% (
<emphasis id="B9225E428E1CFF8CFC19E8A8FC46FD82" box="[951,966,505,530]" italics="true" pageId="11" pageNumber="23">n</emphasis>
= 4). The morphol- ogy of the skulls and dentition is otherwise similar between the two species with the possible exception of the first upper premo- lar (P
<superScript id="7C232F188E1CFF8CFD42EB2DFD76FD1F" attach="left" box="[748,758,636,655]" fontSize="8" pageId="11" pageNumber="23">2</superScript>
), which is minute and extruded from the toothrow in
<taxonomicName id="4C56F9D38E1CFF8CFCC9EBF5FC63FD2D" authorityName="Bates &amp; Rossiter &amp; Suyanto &amp; Kingston" authorityYear="2007" box="[871,995,675,701]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="23" phylum="Chordata" rank="species" species="boeadii">
<emphasis id="B9225E428E1CFF8CFCC9EBF5FC63FD2D" box="[871,995,675,701]" italics="true" pageId="11" pageNumber="23">H. boeadii</emphasis>
</taxonomicName>
and is larger and situated in the toothrow in
<taxonomicName id="4C56F9D38E1CFF8CFC54EB97FBEAFD4F" box="[1018,1130,710,735]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="23" phylum="Chordata" rank="species" species="papua">
<emphasis id="B9225E428E1CFF8CFC54EB97FBEAFD4F" box="[1018,1130,710,735]" italics="true" pageId="11" pageNumber="23">H. papua</emphasis>
</taxonomicName>
(Fig. 3), although further specimens may show this to be an individually variable character.
</paragraph>
<paragraph id="8BE982508E1CFF8CFD7FEA7AFC69FAF7" blockId="11.[683,1199,163,1383]" pageId="11" pageNumber="23">
<taxonomicName id="4C56F9D38E1CFF8CFD7FEA7AFC4CFCD5" authorityName="Bates &amp; Rossiter &amp; Suyanto &amp; Kingston" authorityYear="2007" box="[721,972,811,837]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="23" phylum="Chordata" rank="species" species="boeadii">
<emphasis id="B9225E428E1CFF8CFD7FEA7AFC4CFCD5" box="[721,972,811,837]" italics="true" pageId="11" pageNumber="23">Hipposideros boeadii</emphasis>
</taxonomicName>
is morphologically discrete from
<taxonomicName id="4C56F9D38E1CFF8CFCFAEA1FFB2EFCF7" authority="(Schneider, 1800)" baseAuthorityName="Schneider" baseAuthorityYear="1800" box="[852,1198,845,871]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="23" phylum="Chordata" rank="species" species="speoris">
<emphasis id="B9225E428E1CFF8CFCFAEA1FFC50FCF7" box="[852,976,846,871]" italics="true" pageId="11" pageNumber="23">H. speoris</emphasis>
(
<bibRefCitation id="EFC7FFA18E1CFF8CFC4CEA1CFB26FCF7" author="SCHNEIDER, J. G." box="[994,1190,845,871]" journalOrPublisher="Leipzig" pageId="11" pageNumber="23" refId="ref8211" refString="SCHNEIDER, J. G. 1800. In Schreber, J. C. D. Die Sau- gethiere 1774 - 1855. Leipzig." title="In Schreber, J. C. D. Die Sau- gethiere 1774" type="book" year="1800">Schneider, 1800</bibRefCitation>
)
</taxonomicName>
(described from
<collectingRegion id="49924CB28E1CFF8CFC2EEA3EFB98FC19" box="[896,1048,879,905]" country="India" name="Tamil Nadu" pageId="11" pageNumber="23">Tamil Nadu</collectingRegion>
,
<collectingCountry id="F341C2C08E1CFF8CFB83EA3EFBEBFC1A" box="[1069,1131,879,906]" name="India" pageId="11" pageNumber="23">India</collectingCountry>
) and
<taxonomicName id="4C56F9D38E1CFF8CFD05EAC3FBF0FC3C" authority="(Horsfield, 1823) (Java)" baseAuthorityName="Horsfield" baseAuthorityYear="1823" box="[683,1136,913,940]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="23" phylum="Chordata" rank="species" species="larvatus">
<emphasis id="B9225E428E1CFF8CFD05EAC3FCB6FC3B" box="[683,822,913,939]" italics="true" pageId="11" pageNumber="23">H. larvatus</emphasis>
(
<bibRefCitation id="EFC7FFA18E1CFF8CFCE2EAC0FB8EFC3C" author="HORSFIELD, T." box="[844,1038,913,940]" journalOrPublisher="London, unpaginated" pageId="11" pageNumber="23" refId="ref7283" refString="HORSFIELD, T. 1823. Zoological researches in Java and the neighbouring islands. Kingsbury, Parbury and Allen, London, unpaginated." title="Zoological researches in Java and the neighbouring islands. Kingsbury, Parbury and Allen" type="book" year="1823">Horsfield, 1823</bibRefCitation>
) (Java)
</taxonomicName>
, two species with which
<taxonomicName id="4C56F9D38E1CFF8CFC37EAE5FB89FC5D" box="[921,1033,948,973]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="23" phylum="Chordata" rank="species" species="papua">
<emphasis id="B9225E428E1CFF8CFC37EAE5FB89FC5D" box="[921,1033,948,973]" italics="true" pageId="11" pageNumber="23">H. papua</emphasis>
</taxonomicName>
is sometimes linked (for full discussion see below). Unlike
<taxonomicName id="4C56F9D38E1CFF8CFCA2EAA9FC12FB82" authorityName="Bates &amp; Rossiter &amp; Suyanto &amp; Kingston" authorityYear="2007" box="[780,914,1016,1042]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="23" phylum="Chordata" rank="species" species="boeadii">
<emphasis id="B9225E428E1CFF8CFCA2EAA9FC12FB82" box="[780,914,1016,1042]" italics="true" pageId="11" pageNumber="23">H. boeadii</emphasis>
</taxonomicName>
, both
<taxonomicName id="4C56F9D38E1CFF8CFC5EEAA9FBF3FB81" baseAuthorityName="Schneider" baseAuthorityYear="1800" box="[1008,1139,1016,1041]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="23" phylum="Chordata" rank="species" species="speoris">
<emphasis id="B9225E428E1CFF8CFC5EEAA9FBF3FB81" box="[1008,1139,1016,1041]" italics="true" pageId="11" pageNumber="23">H. speoris</emphasis>
</taxonomicName>
and
<taxonomicName id="4C56F9D38E1CFF8CFD05ED4AFCB1FBA4" baseAuthorityName="Horsfield" baseAuthorityYear="1823" box="[683,817,1050,1076]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="23" phylum="Chordata" rank="species" species="larvatus">
<emphasis id="B9225E428E1CFF8CFD05ED4AFCB1FBA4" box="[683,817,1050,1076]" italics="true" pageId="11" pageNumber="23">H. larvatus</emphasis>
</taxonomicName>
have three supplementary leaf- lets, with the first leaflet exceeding or about equal in length to the second.
<emphasis id="B9225E428E1CFF8CFB82ED0FFCB3FB0A" italics="true" pageId="11" pageNumber="23">
Hipposide- ros
<taxonomicName id="4C56F9D38E1CFF8CFD75EDD1FCB3FB0A" authorityName="Bates &amp; Rossiter &amp; Suyanto &amp; Kingston" authorityYear="2007" box="[731,819,1152,1178]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="23" phylum="Chordata" rank="species" species="boeadii">boeadii</taxonomicName>
</emphasis>
also differs in its rostral mor- phology and smaller size. The forearm and condylo-canine lengths of these latter two taxa are respectively: FA: 45.654.0 mm and
<quantity id="4CAE2FB58E1CFF8CFD4AEC59FC19FAB3" box="[740,921,1288,1315]" metricMagnitude="-2" metricUnit="m" metricValue="6.3" metricValueMax="6.4799999999999995" metricValueMin="6.12" pageId="11" pageNumber="23" unit="mm" value="63.0" valueMax="64.8" valueMin="61.2">61.264.8 mm</quantity>
; CCL:
<quantity id="4CAE2FB58E1CFF8CFC52EC59FB2EFAB3" box="[1020,1198,1288,1315]" metricMagnitude="-2" metricUnit="m" metricValue="1.67" metricValueMax="1.75" metricValueMin="1.59" pageId="11" pageNumber="23" unit="mm" value="16.7" valueMax="17.5" valueMin="15.9">15.917.5 mm</quantity>
and
<quantity id="4CAE2FB58E1CFF8CFD46EC7AFC1EFAD5" box="[744,926,1323,1349]" metricMagnitude="-2" metricUnit="m" metricValue="2.11" metricValueMax="2.1399999999999997" metricValueMin="2.08" pageId="11" pageNumber="23" unit="mm" value="21.1" valueMax="21.4" valueMin="20.8">20.821.4 mm</quantity>
(measurements from Bates and Harrison, 1997).
</paragraph>
</subSubSection>
<subSubSection id="C34CD1DB8E1CFF8BFD05ECC6FEDFFEF7" lastPageId="12" lastPageNumber="24" pageId="11" pageNumber="23" type="description">
<paragraph id="8BE982508E1CFF8CFD05ECC6FC85FA21" blockId="11.[683,773,1431,1457]" box="[683,773,1431,1457]" pageId="11" pageNumber="23">
<taxonomicName id="4C56F9D38E1CFF8CFD05ECC6FC85FA21" authority="Habitat" authorityName="Habitat" box="[683,773,1431,1457]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="23" phylum="Chordata" rank="species" species="papua">
<emphasis id="B9225E428E1CFF8CFD05ECC6FC85FA21" box="[683,773,1431,1457]" italics="true" pageId="11" pageNumber="23">Habitat</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BE982508E1CFF8CFD7FEC8AFB2EF952" blockId="11.[683,1199,1499,1730]" pageId="11" pageNumber="23">
All individuals of
<taxonomicName id="4C56F9D38E1CFF8CFC1EEC8DFBAEFA65" authorityName="Bates &amp; Rossiter &amp; Suyanto &amp; Kingston" authorityYear="2007" box="[944,1070,1499,1525]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="23" phylum="Chordata" rank="species" species="boeadii">
<emphasis id="B9225E428E1CFF8CFC1EEC8DFBAEFA65" box="[944,1070,1499,1525]" italics="true" pageId="11" pageNumber="23">H. boeadii</emphasis>
</taxonomicName>
were cap- tured in lowland rainforest between one and four kilometres from the adjacent sa- vannah. The forest was disturbed, and veg- etation near to the edge was characterised by a high incidence of bamboo and rattan. Other bat species recorded at this site, in
</paragraph>
<paragraph id="8BE982508E1BFF8BFF2CE9F2FEDFFEF7" blockId="12.[130,646,163,359]" pageId="12" pageNumber="24">
order of relative abundance, were
<emphasis id="B9225E428E1BFF8BFDBEE9F2FF7EFF4F" italics="true" pageId="12" pageNumber="24">
<taxonomicName id="4C56F9D38E1BFF8BFDBEE9F2FD05FF2D" authorityName="Gray" authorityYear="1842" box="[528,645,163,189]" class="Mammalia" family="Vespertilionidae" genus="Kerivoula" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="24" phylum="Chordata" rank="genus">Kerivoula</taxonomicName>
hardwickii
</emphasis>
, Rhinolophus
<taxonomicName id="4C56F9D38E1BFF8BFE16E997FD96FF4F" box="[440,534,198,223]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="24" phylum="Chordata" rank="species" species="euryotis">
<emphasis id="B9225E428E1BFF8BFE16E997FD96FF4F" box="[440,534,198,223]" italics="true" pageId="12" pageNumber="24">euryotis</emphasis>
</taxonomicName>
,
<emphasis id="B9225E428E1BFF8BFD83E997FF58FE91" italics="true" pageId="12" pageNumber="24">
<taxonomicName id="4C56F9D38E1BFF8BFD83E997FD05FF4F" box="[557,645,198,223]" class="Mammalia" family="Vespertilionidae" genus="Murina" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="24" phylum="Chordata" rank="genus">Murina</taxonomicName>
florium
</emphasis>
,
<taxonomicName id="4C56F9D38E1BFF8BFF44E9B9FEFCFE91" baseAuthorityName="Temminck" baseAuthorityYear="1840" box="[234,380,231,257]" class="Mammalia" family="Vespertilionidae" genus="Kerivoula" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="24" phylum="Chordata" rank="species" species="papillosa">
<emphasis id="B9225E428E1BFF8BFF44E9B9FEFCFE91" box="[234,380,231,257]" italics="true" pageId="12" pageNumber="24">K. papillosa</emphasis>
</taxonomicName>
,
<taxonomicName id="4C56F9D38E1BFF8BFE23E9B9FD97FE91" box="[397,535,232,257]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="24" phylum="Chordata" rank="species" species="cervinus">
<emphasis id="B9225E428E1BFF8BFE23E9B9FD97FE91" box="[397,535,232,257]" italics="true" pageId="12" pageNumber="24">H. cervinus</emphasis>
</taxonomicName>
, R. cele-
<emphasis id="B9225E428E1BFF8BFF2CE858FF4BFEB3" box="[130,203,265,291]" italics="true" pageId="12" pageNumber="24">bensis</emphasis>
, Phoniscus
<taxonomicName id="4C56F9D38E1BFF8BFEC8E858FE36FEB2" baseAuthorityName="Peters" baseAuthorityYear="1866" box="[358,438,265,290]" class="Mammalia" family="Vespertilionidae" genus="Phoniscus" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="24" phylum="Chordata" rank="species" species="jagorii">
<emphasis id="B9225E428E1BFF8BFEC8E858FE36FEB2" box="[358,438,265,290]" italics="true" pageId="12" pageNumber="24">jagorii</emphasis>
</taxonomicName>
, and an uniden- tified
<taxonomicName id="4C56F9D38E1BFF8BFF64E87BFEE7FED4" box="[202,359,298,324]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="24" phylum="Chordata" rank="genus">
<emphasis id="B9225E428E1BFF8BFF64E87BFEE7FED4" box="[202,359,298,324]" italics="true" pageId="12" pageNumber="24">Hipposideros</emphasis>
</taxonomicName>
species morphological- ly close to
<taxonomicName id="4C56F9D38E1BFF8BFEAAE81CFED8FEF6" authorityName="Templeton" authorityYear="1848" box="[260,344,333,358]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="24" phylum="Chordata" rank="species" species="ater">
<emphasis id="B9225E428E1BFF8BFEAAE81CFED8FEF6" box="[260,344,333,358]" italics="true" pageId="12" pageNumber="24">H. ater</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C34CD1DB8E1BFF8BFF2CE8C2FED7FDDF" pageId="12" pageNumber="24" type="distribution">
<paragraph id="8BE982508E1BFF8BFF2CE8C2FE91FE3D" blockId="12.[130,273,403,429]" box="[130,273,403,429]" pageId="12" pageNumber="24">
<emphasis id="B9225E428E1BFF8BFF2CE8C2FE91FE3D" box="[130,273,403,429]" italics="true" pageId="12" pageNumber="24">Distribution</emphasis>
</paragraph>
<paragraph id="8BE982508E1BFF8BFF06E881FED7FDDF" blockId="12.[130,646,463,591]" pageId="12" pageNumber="24">
To date,
<taxonomicName id="4C56F9D38E1BFF8BFEBFE881FE0FFE79" authorityName="Bates &amp; Rossiter &amp; Suyanto &amp; Kingston" authorityYear="2007" box="[273,399,463,489]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="24" phylum="Chordata" rank="species" species="boeadii">
<emphasis id="B9225E428E1BFF8BFEBFE881FE0FFE79" box="[273,399,463,489]" italics="true" pageId="12" pageNumber="24">H. boeadii</emphasis>
</taxonomicName>
is only known from the
<typeStatus id="54ED3CF28E1BFF8BFF03E8A3FF60FD9C" box="[173,224,498,524]" pageId="12" pageNumber="24">type</typeStatus>
locality. Future studies may show it to be more widespread elsewhere in Indo- nesia and beyond.
</paragraph>
</subSubSection>
<subSubSection id="C34CD1DB8E1BFF8BFF2CEB2DFF71FBF6" pageId="12" pageNumber="24" type="biology_ecology">
<paragraph id="8BE982508E1BFF8BFF2CEB2DFE00FD06" blockId="12.[130,384,636,662]" box="[130,384,636,662]" pageId="12" pageNumber="24">
Reproductive
<emphasis id="B9225E428E1BFF8BFE8BEB2DFE00FD06" box="[293,384,636,662]" italics="true" pageId="12" pageNumber="24">Biology</emphasis>
</paragraph>
<paragraph id="8BE982508E1BFF8BFF06EBE9FF71FBF6" blockId="12.[130,646,696,1126]" pageId="12" pageNumber="24">
The nipples and pubic nipples of eight female
<taxonomicName id="4C56F9D38E1BFF8BFF75EB8AFED8FD64" authorityName="Bates &amp; Rossiter &amp; Suyanto &amp; Kingston" authorityYear="2007" box="[219,344,730,756]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="24" phylum="Chordata" rank="species" species="boeadii">
<emphasis id="B9225E428E1BFF8BFF75EB8AFED8FD64" box="[219,344,730,756]" italics="true" pageId="12" pageNumber="24">H. boeadii</emphasis>
</taxonomicName>
were examined. Four in- dividuals were parous (including the two fe- male voucher specimens) and four nul- liparous. Pregnancy was not detected by either external abdominal palpation of the released individuals, or dissection of the two female specimens. However, vaginal plugs were found in three individuals (in one of the parous specimens and in two of the nulliparous females released at the site), indicating that mating had recently occurred.
</paragraph>
</subSubSection>
<subSubSection id="C34CD1DB8E1BFF8BFF2CEDC5FCE7FBE7" pageId="12" pageNumber="24" type="description">
<paragraph id="8BE982508E1BFF8BFF2CEDC5FECDFB3D" blockId="12.[130,333,1172,1197]" box="[130,333,1172,1197]" pageId="12" pageNumber="24">
<emphasis id="B9225E428E1BFF8BFF2CEDC5FECDFB3D" box="[130,333,1172,1197]" italics="true" pageId="12" pageNumber="24">Taxonomic Notes</emphasis>
</paragraph>
<paragraph id="8BE982508E1BFF8BFF06ED81FD05F952" blockId="12.[130,646,1232,1730]" pageId="12" pageNumber="24">
Based on external and cranial morpholo- gy
<taxonomicName id="4C56F9D38E1BFF8BFF09EDA2FEA4FA9C" authorityName="Bates &amp; Rossiter &amp; Suyanto &amp; Kingston" authorityYear="2007" box="[167,292,1266,1292]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="24" phylum="Chordata" rank="species" species="boeadii">
<emphasis id="B9225E428E1BFF8BFF09EDA2FEA4FA9C" box="[167,292,1266,1292]" italics="true" pageId="12" pageNumber="24">H. boeadii</emphasis>
</taxonomicName>
appears to have strong affini- ties to
<taxonomicName id="4C56F9D38E1BFF8BFF7CEC45FEC0FABD" box="[210,320,1300,1325]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="24" phylum="Chordata" rank="species" species="papua">
<emphasis id="B9225E428E1BFF8BFF7CEC45FEC0FABD" box="[210,320,1300,1325]" italics="true" pageId="12" pageNumber="24">H. papua</emphasis>
</taxonomicName>
, a species whose taxonom- ic position is uncertain.
<bibRefCitation id="EFC7FFA18E1BFF8BFE30EC67FDA7FADF" author="Tate" box="[414,551,1333,1360]" firstAuthor="Tate" pageId="12" pageNumber="24" pagination="353 - 393" refId="ref8389" refString="TATE, G. H. H. 1941. A review of the genus Hippo- and Magazine of Natural History, London, 12: sideros with special reference to Indo-Australian 212 - 217. species. Bulletin of the American Museum of THOMAS, O., and G. DORIA. 1886. Note intorno ad Natural History, 78: 353 - 393. alcune Chirotteri. Annali del Museo Civico di" type="journal article" year="1941">Tate (1941)</bibRefCitation>
consid- ered
<taxonomicName id="4C56F9D38E1BFF8BFF10EC09FEAEFAE1" box="[190,302,1368,1393]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="24" phylum="Chordata" rank="species" species="papua">
<emphasis id="B9225E428E1BFF8BFF10EC09FEAEFAE1" box="[190,302,1368,1393]" italics="true" pageId="12" pageNumber="24">H. papua</emphasis>
</taxonomicName>
to be a perplexing species which may have been derived from near the base of the
<taxonomicName id="4C56F9D38E1BFF8BFE99ECCAFE0CFA24" baseAuthorityName="Schneider" baseAuthorityYear="1800" box="[311,396,1435,1460]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="24" phylum="Chordata" rank="species" species="speoris">
<emphasis id="B9225E428E1BFF8BFE99ECCAFE0CFA24" box="[311,396,1435,1460]" italics="true" pageId="12" pageNumber="24">speoris</emphasis>
</taxonomicName>
and
<taxonomicName id="4C56F9D38E1BFF8BFE68ECCBFDAEFA24" authorityName="Cantor" authorityYear="1846" box="[454,558,1434,1460]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="24" phylum="Chordata" rank="species" species="galeritus">
<emphasis id="B9225E428E1BFF8BFE68ECCBFDAEFA24" box="[454,558,1434,1460]" italics="true" pageId="12" pageNumber="24">galeritus</emphasis>
</taxonomicName>
(
<emphasis id="B9225E428E1BFF8BFDEFECCAFF2BFA46" italics="true" pageId="12" pageNumber="24">cervi- nus</emphasis>
) stems. According to
<bibRefCitation id="EFC7FFA18E1BFF8BFE69ECEDFDD1FA47" author="HILL, J. E." box="[455,593,1468,1495]" journalOrPublisher="Bulletin of the British Museum (Natural History), Zoology" pageId="12" pageNumber="24" pagination="1 - 129" part="11" refId="ref8178" refString="HILL, J. E. 1963. A revison of the genus Hipposideros. Bulletin of the British Museum (Natural History), Zoology, 11: 1 - 129" title="A revison of the genus Hipposideros" type="journal article" year="1963">Hill (1963)</bibRefCitation>
, the ears and cranial characters of
<taxonomicName id="4C56F9D38E1BFF8BFE70EC8FFDCBFA67" box="[478,587,1502,1527]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="24" phylum="Chordata" rank="species" species="papua">
<emphasis id="B9225E428E1BFF8BFE70EC8FFDCBFA67" box="[478,587,1502,1527]" italics="true" pageId="12" pageNumber="24">H. papua</emphasis>
</taxonomicName>
sug- gest affinity with the
<taxonomicName id="4C56F9D38E1BFF8BFE2CECAEFE57F989" box="[386,471,1535,1561]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="24" phylum="Chordata" rank="species" species="bicolor">
<emphasis id="B9225E428E1BFF8BFE2CECAEFE57F989" box="[386,471,1535,1561]" italics="true" pageId="12" pageNumber="24">bicolor</emphasis>
</taxonomicName>
group, where- as its specialised noseleaf indicates no close connection with any other members. Some affinity to the
<emphasis id="B9225E428E1BFF8BFE86EF35FE00F9EE" box="[296,384,1636,1662]" italics="true" pageId="12" pageNumber="24">cyclops</emphasis>
group is suggested by the extension of the second leaflet beneath the anterior leaf, a feature otherwise pecu-
</paragraph>
<paragraph id="8BE982508E1BFF8BFD05E9F2FCE7FBE7" blockId="12.[683,1199,163,1143]" pageId="12" pageNumber="24">
liar to
<emphasis id="B9225E428E1BFF8BFD50E9F2FCD6FF2D" box="[766,854,163,189]" italics="true" pageId="12" pageNumber="24">cyclops</emphasis>
<bibRefCitation id="EFC7FFA18E1BFF8BFCCEE9F2FBAAFF2E" author="Temminck" box="[864,1066,163,190]" firstAuthor="Temminck" pageId="12" pageNumber="24" refId="ref8465" refString="TEMMINCK, C. J. 1853. Esquisses zoologiques sur la Storia Naturale (di Genova) Giacamo Doria, 4: Cote de Guinee. 1 re partie, les mammif P res. E. J. 201 - 207. Brill, Leiden, 256 pp. TOMES, R. F. 1859. Notice of five species of bats in" type="book" year="1853">Temminck, 1853</bibRefCitation>
(described from
<collectingCountry id="F341C2C08E1BFF8BFD42E994FCBCFF4F" box="[748,828,197,223]" name="Ghana" pageId="12" pageNumber="24">Ghana</collectingCountry>
, West Africa) and its allies, but
<taxonomicName id="4C56F9D38E1BFF8BFD05E9B9FC9FFE91" box="[683,799,232,257]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="24" phylum="Chordata" rank="species" species="papua">
<emphasis id="B9225E428E1BFF8BFD05E9B9FC9FFE91" box="[683,799,232,257]" italics="true" pageId="12" pageNumber="24">H. papua</emphasis>
</taxonomicName>
differs from these in its broad ears, the presence of an incipient third leaflet, unenlarged bullae and convergent toothrows.
<bibRefCitation id="EFC7FFA18E1BFF8BFCECE81FFC59FEF8" author="HILL, J. E." box="[834,985,333,360]" journalOrPublisher="Bulletin of the British Museum (Natural History), Zoology" pageId="12" pageNumber="24" pagination="1 - 129" part="11" refId="ref8178" refString="HILL, J. E. 1963. A revison of the genus Hipposideros. Bulletin of the British Museum (Natural History), Zoology, 11: 1 - 129" title="A revison of the genus Hipposideros" type="journal article" year="1963">Hill (1963)</bibRefCitation>
concluded that
<taxonomicName id="4C56F9D38E1BFF8BFD05E821FC98FE19" box="[683,792,368,393]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="24" phylum="Chordata" rank="species" species="papua">
<emphasis id="B9225E428E1BFF8BFD05E821FC98FE19" box="[683,792,368,393]" italics="true" pageId="12" pageNumber="24">H. papua</emphasis>
</taxonomicName>
should be regarded as a species of remote origin within the
<taxonomicName id="4C56F9D38E1BFF8BFC76E8C3FBADFE3C" box="[984,1069,402,428]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="24" phylum="Chordata" rank="species" species="bicolor">
<emphasis id="B9225E428E1BFF8BFC76E8C3FBADFE3C" box="[984,1069,402,428]" italics="true" pageId="12" pageNumber="24">bicolor</emphasis>
</taxonomicName>
group, as- sociated probably with the line leading to the
<taxonomicName id="4C56F9D38E1BFF8BFD74E887FCC7FE7F" authorityName="Thomas and Doria" authorityYear="1886" box="[730,839,470,495]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="24" phylum="Chordata" rank="species" species="muscinus">
<emphasis id="B9225E428E1BFF8BFD74E887FCC7FE7F" box="[730,839,470,495]" italics="true" pageId="12" pageNumber="24">muscinus</emphasis>
</taxonomicName>
group and possibly with that leading to the
<taxonomicName id="4C56F9D38E1BFF8BFCC0E8A9FC43FD81" baseAuthorityName="Schneider" baseAuthorityYear="1800" box="[878,963,504,529]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="24" phylum="Chordata" rank="species" species="speoris">
<emphasis id="B9225E428E1BFF8BFCC0E8A9FC43FD81" box="[878,963,504,529]" italics="true" pageId="12" pageNumber="24">speoris</emphasis>
</taxonomicName>
group. This view was followed by Corbet and Hill (1992) and
<bibRefCitation id="EFC7FFA18E1BFF8BFD05EB6DFC10FDC6" author="KOOPMAN, K. F." bookContentInfo="217 pp." box="[683,912,572,598]" journalOrPublisher="Walter de Gruyter, Berlin" pageId="12" pageNumber="24" refId="ref7464" refString="KOOPMAN, K. F. 1994. Chiroptera: systematics. Pp. 1 - 217, in Handbook of Zoology, Vol. 8. Mammalia, Part 60. Walter de Gruyter, Berlin, vii + 217 pp." title="Chiroptera: systematics. Pp. 1 - 217, in Handbook of Zoology, Vol. 8. Mammalia, Part 60" type="book" year="1994">Koopman (1994)</bibRefCitation>
, who also assigned
<taxonomicName id="4C56F9D38E1BFF8BFD05EB0FFC9DFDE7" box="[683,797,606,631]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="24" phylum="Chordata" rank="species" species="papua">
<emphasis id="B9225E428E1BFF8BFD05EB0FFC9DFDE7" box="[683,797,606,631]" italics="true" pageId="12" pageNumber="24">H. papua</emphasis>
</taxonomicName>
to the
<taxonomicName id="4C56F9D38E1BFF8BFCD1EB0FFC54FDE8" box="[895,980,606,632]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="24" phylum="Chordata" rank="species" species="bicolor">
<emphasis id="B9225E428E1BFF8BFCD1EB0FFC54FDE8" box="[895,980,606,632]" italics="true" pageId="12" pageNumber="24">bicolor</emphasis>
</taxonomicName>
group, which in- cluded
<taxonomicName id="4C56F9D38E1BFF8BFCACEBD1FCEAFD0A" authorityName="Cantor" authorityYear="1846" box="[770,874,640,666]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="24" phylum="Chordata" rank="species" species="galeritus">
<emphasis id="B9225E428E1BFF8BFCACEBD1FCEAFD0A" box="[770,874,640,666]" italics="true" pageId="12" pageNumber="24">galeritus</emphasis>
</taxonomicName>
but omitted
<taxonomicName id="4C56F9D38E1BFF8BFBA8EBD1FBDBFD09" baseAuthorityName="Schneider" baseAuthorityYear="1800" box="[1030,1115,640,665]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="24" phylum="Chordata" rank="species" species="speoris">
<emphasis id="B9225E428E1BFF8BFBA8EBD1FBDBFD09" box="[1030,1115,640,665]" italics="true" pageId="12" pageNumber="24">speoris</emphasis>
</taxonomicName>
. How- ever, it was not followed by Hand and Kirsch (1998) or Bogdanowicz and Owen (1998). The former found no close affinities between
<taxonomicName id="4C56F9D38E1BFF8BFCBDEA58FCFFFCB2" box="[787,895,777,802]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="24" phylum="Chordata" rank="species" species="papua">
<emphasis id="B9225E428E1BFF8BFCBDEA58FCFFFCB2" box="[787,895,777,802]" italics="true" pageId="12" pageNumber="24">H. papua</emphasis>
</taxonomicName>
and other members of the genus, whilst the latter tentatively suggested that
<taxonomicName id="4C56F9D38E1BFF8BFD4CEA1CFCD4FCF6" box="[738,852,845,870]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="24" phylum="Chordata" rank="species" species="papua">
<emphasis id="B9225E428E1BFF8BFD4CEA1CFCD4FCF6" box="[738,852,845,870]" italics="true" pageId="12" pageNumber="24">H. papua</emphasis>
</taxonomicName>
was most closely related to
<taxonomicName id="4C56F9D38E1BFF8BFD05EA3EFCB1FC18" baseAuthorityName="Horsfield" baseAuthorityYear="1823" box="[683,817,878,904]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="24" phylum="Chordata" rank="species" species="larvatus">
<emphasis id="B9225E428E1BFF8BFD05EA3EFCB1FC18" box="[683,817,878,904]" italics="true" pageId="12" pageNumber="24">H. larvatus</emphasis>
</taxonomicName>
, which is a member of the
<emphasis id="B9225E428E1BFF8BFBD3EA3EFD59FC3A" italics="true" pageId="12" pageNumber="24">spe- oris</emphasis>
group (sensu
<bibRefCitation id="EFC7FFA18E1BFF8BFC36EAC1FB9CFC3B" author="HILL, J. E." box="[920,1052,912,939]" journalOrPublisher="Bulletin of the British Museum (Natural History), Zoology" pageId="12" pageNumber="24" pagination="1 - 129" part="11" refId="ref8178" refString="HILL, J. E. 1963. A revison of the genus Hipposideros. Bulletin of the British Museum (Natural History), Zoology, 11: 1 - 129" title="A revison of the genus Hipposideros" type="journal article" year="1963">Hill, 1963</bibRefCitation>
). Based on preliminary phylogenetic comparisons,
<taxonomicName id="4C56F9D38E1BFF8BFD05EA84FCA7FC7E" authorityName="Bates &amp; Rossiter &amp; Suyanto &amp; Kingston" authorityYear="2007" box="[683,807,980,1006]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="24" phylum="Chordata" rank="species" species="boeadii">
<emphasis id="B9225E428E1BFF8BFD05EA84FCA7FC7E" box="[683,807,980,1006]" italics="true" pageId="12" pageNumber="24">H. boeadii</emphasis>
</taxonomicName>
appears to be more closely relat- ed to
<taxonomicName id="4C56F9D38E1BFF8BFD5CEAA6FBF4FB80" authority="(E. Geoffroy, 1813)" baseAuthorityName="E. Geoffroy" baseAuthorityYear="1813" box="[754,1140,1014,1040]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="24" phylum="Chordata" rank="species" species="diadema">
<emphasis id="B9225E428E1BFF8BFD5CEAA6FCFEFB80" box="[754,894,1014,1040]" italics="true" pageId="12" pageNumber="24">H. diadema</emphasis>
(E.
<bibRefCitation id="EFC7FFA18E1BFF8BFC1BEAA7FBECFB80" author="GEOFFROY, E." box="[949,1132,1014,1040]" journalOrPublisher="Annales du Mu- seum National d'Histoire Naturelle (Paris)" pageId="12" pageNumber="24" pagination="254 - 266" part="20" refId="ref7859" refString="GEOFFROY, E. 1813. Sur un genre de chauve-souris sous le nom de Rhinolophes (1). Annales du Mu- seum National d'Histoire Naturelle (Paris), 20: 254 - 266." title="Sur un genre de chauve-souris sous le nom de Rhinolophes" type="journal article" year="1813">Geoffroy, 1813</bibRefCitation>
)
</taxonomicName>
(de- scribed from Timor) and
<taxonomicName id="4C56F9D38E1BFF8BFC7CED48FBD6FBA2" baseAuthorityName="Horsfield" baseAuthorityYear="1823" box="[978,1110,1048,1074]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="24" phylum="Chordata" rank="species" species="larvatus">
<emphasis id="B9225E428E1BFF8BFC7CED48FBD6FBA2" box="[978,1110,1048,1074]" italics="true" pageId="12" pageNumber="24">H. larvatus</emphasis>
</taxonomicName>
than to
<taxonomicName id="4C56F9D38E1BFF8BFD05ED6AFCB6FBC4" authorityName="Cantor" authorityYear="1846" box="[683,822,1082,1108]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="24" phylum="Chordata" rank="species" species="galeritus">
<emphasis id="B9225E428E1BFF8BFD05ED6AFCB6FBC4" box="[683,822,1082,1108]" italics="true" pageId="12" pageNumber="24">H. galeritus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C56F9D38E1BFF8BFCC0ED6AFC74FBC4" box="[878,1012,1083,1108]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="24" phylum="Chordata" rank="species" species="cervinus">
<emphasis id="B9225E428E1BFF8BFCC0ED6AFC74FBC4" box="[878,1012,1083,1108]" italics="true" pageId="12" pageNumber="24">H. cervinus</emphasis>
</taxonomicName>
(S. Murray, un- published data).
</paragraph>
</subSubSection>
</treatment>
</document>