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<document id="86BE3403AB0ABB97BAD0883AE5025ED3" ID-CLB-Dataset="294473" ID-DOI="10.26879/1312" ID-GBIF-Dataset="8d351e9f-9cd6-44e9-bfc1-5f5cb767a157" ID-ISSN="1094-8074" ID-Zenodo-Dep="10962373" ID-ZooBank="71FEB770-6EE3-4621-B752-872A1CBDB605" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1712860417308" checkinUser="felipe" docAuthor="Gašparič, Rok, Hyžný, Matúš, Hitij, Tomaž &amp; Šoster, Aleš" docDate="2024" docId="03FF1D383664CE38127CED0B397FE9BA" docLanguage="en" docName="PalaeontolElectron.27.1.a4.pdf" docOrigin="Palaeontologia Electronica (a 4) 27 (1)" docSource="http://dx.doi.org/10.26879/1312" docStyle="DocumentStyle:322FC57CC99045555090E8A84CFB029A.3:PalaeontolElectron.2018-.journal_article" docStyleId="322FC57CC99045555090E8A84CFB029A" docStyleName="PalaeontolElectron.2018-.journal_article" docStyleVersion="3" docTitle="Halicarcinus popeius Gašparič &amp; Hyžný &amp; Hitij &amp; Šoster 2024, sp. nov." docType="treatment" docVersion="4" lastPageNumber="8" masterDocId="FFC665403667CE3F133CEA0B3867EC7D" masterDocTitle="Late Oligocene decapod crustaceans from the Trbovlje Formation of Slovenia, with a description of two new species of hymenosomatid crabs" masterLastPageNumber="16" masterPageNumber="1" pageNumber="4" updateTime="1713793964816" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-SA-4.0">
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<mods:title id="E3C398CCC1086A1AF8762C0016D08F18">Late Oligocene decapod crustaceans from the Trbovlje Formation of Slovenia, with a description of two new species of hymenosomatid crabs</mods:title>
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<mods:namePart id="BC0C3EA851CC0E7DF49D5EDC54FD44E6">Gašparič, Rok</mods:namePart>
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<mods:date id="56B8F83AC6B0E522C702C87C505B4BFA">2024</mods:date>
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<treatment id="03FF1D383664CE38127CED0B397FE9BA" ID-DOI="http://doi.org/10.5281/zenodo.11033742" ID-Zenodo-Dep="11033742" LSID="urn:lsid:plazi:treatment:03FF1D383664CE38127CED0B397FE9BA" httpUri="http://treatment.plazi.org/id/03FF1D383664CE38127CED0B397FE9BA" lastPageId="7" lastPageNumber="8" pageId="3" pageNumber="4">
<subSubSection id="C34CFFA53664CE3C127CED0B3AFEEB64" box="[320,665,1792,1817]" pageId="3" pageNumber="4" type="nomenclature">
<paragraph id="8BE9AC2E3664CE3C127CED0B3AFEEB64" blockId="3.[320,665,1792,1849]" box="[320,665,1792,1817]" pageId="3" pageNumber="4">
<heading id="D0A11B423664CE3C127CED0B3AFEEB64" box="[320,665,1792,1817]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="4" reason="7">
<taxonomicName id="4C56D7AD3664CE3C127CED0B3A52EB64" ID-CoL="C34YV" authority="Gašparič &amp; Hyžný &amp; Hitij &amp; Šoster, 2024" authorityName="Gašparič &amp; Hyžný &amp; Hitij &amp; Šoster" authorityYear="2024" box="[320,565,1792,1817]" class="Malacostraca" family="Hymenosomatidae" genus="Halicarcinus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="popeius" status="sp. nov.">
<emphasis id="B922703C3664CE3C127CED0B3A52EB64" box="[320,565,1792,1817]" italics="true" pageId="3" pageNumber="4">Halicarcinus popeius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A211CD473664CE3C1101ED0B3AFEEB64" box="[573,665,1792,1817]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="C34CFFA53664CE3C1255ED2B3CECEAF3" pageId="3" pageNumber="4" type="description">
<paragraph id="8BE9AC2E3664CE3C1255ED2B3A08EB44" blockId="3.[320,665,1792,1849]" box="[361,623,1823,1849]" pageId="3" pageNumber="4">
<figureCitation id="136DB0AB3664CE3C1255ED2B39BCEB44" box="[361,475,1824,1849]" captionStart="FIGURE 3" captionStartId="4.[197,288,858,881]" captionTargetBox="[190,1437,163,833]" captionTargetId="figure-410@4.[190,1437,163,833]" captionTargetPageId="4" captionText="FIGURE 3. Halicarcinus popeius sp. nov., schematic reconstruction. A. Carapace in dorsal view; B. Detailed view of second type of rostrum and robust chelipeds (male); C. Third pereopod (P3) with detailed view of dactylus." figureDoi="http://doi.org/10.5281/zenodo.10962379" httpUri="https://zenodo.org/record/10962379/files/figure.png" pageId="3" pageNumber="4">Figures 3</figureCitation>
,
<figureCitation id="136DB0AB3664CE3C12DBED2B3991EB44" box="[487,502,1824,1849]" captionStart="FIGURE 4" captionStartId="5.[193,284,1453,1476]" captionTargetBox="[190,1436,167,1438]" captionTargetId="figure-121@5.[190,1436,167,1438]" captionTargetPageId="5" captionText="FIGURE 4. Halicarcinus popeius sp. nov. (AD) and Lucascinus trifailensis sp. nov. (E, F): A. RGA/SMNH 6377, dorsal, (holotype), visible hymenosomian rim (hr) and coloration patterns; B. RGA/SMNH 6373, dorsal, (paratype), male with well-preserved chelipeds and pereopods (P2-P5); C. RGA/SMNH 6385, dorsal, (paratype), male with bulbous chelipeds; D. RGA/SMNH 2041, ventral, (paratype), female with missing sternum; E. RGA/SMNH 6384, dorsal, (holotype), indicated rostrum and hymenosomian rim (hr); F. RGA/SMNH 6391, dorsal, (paratype), elongated chelipeds and well preserved pereopods (P2-P5). All scale bars equal 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10962381" httpUri="https://zenodo.org/record/10962381/files/figure.png" pageId="3" pageNumber="4">4</figureCitation>
A-D, 5A-C
</paragraph>
<paragraph id="8BE9AC2E3664CE3C13FCED413B65EB23" blockId="3.[192,770,1866,1886]" box="[192,770,1866,1886]" pageId="3" pageNumber="4">
<emphasis id="B922703C3664CE3C13FCED413B65EB23" bold="true" box="[192,770,1866,1886]" pageId="3" pageNumber="4">
zoobank.org/
<uuid id="FFF096FB3664CE3C1277ED413B65EB23" box="[331,770,1866,1886]" pageId="3" pageNumber="4">BC3F122F-89A8-4BDB-A174-664B834FA1D1</uuid>
</emphasis>
</paragraph>
<paragraph id="8BE9AC2E3664CE3C1074EC5E3CECEAF3" blockId="3.[840,1441,1621,1875]" pageId="3" pageNumber="4">
2023
<taxonomicName id="4C56D7AD3664CE3C1094EC5E3C73EA13" authorityName="Samouelle" authorityYear="1819" box="[936,1044,1621,1646]" pageId="3" pageNumber="2" rank="superFamily" superFamily="Majoidea">Majoidea</taxonomicName>
gen. et sp. indet. Kovalchuk et al., p. 5, figs. 2E, F.
</paragraph>
</subSubSection>
<subSubSection id="C34CFFA53664CE3C1074EC913BCCEB2E" pageId="3" pageNumber="4" type="etymology">
<paragraph id="8BE9AC2E3664CE3C1074EC913BCCEB2E" blockId="3.[840,1441,1621,1875]" pageId="3" pageNumber="4">
<emphasis id="B922703C3664CE3C1074EC913BB1EACE" bold="true" box="[840,982,1690,1715]" pageId="3" pageNumber="4">Etymology.</emphasis>
The species name originates from a Latinised name of the cartoon character Popeye, the sailor man. A fictional character who, despite the diminutive stature, possesses strong, bulging forearms, reminiscent of the chelipeds of the new species.
</paragraph>
</subSubSection>
<caption id="DF29FCA63663CE3B13F9E9513D4AEFF2" ID-DOI="http://doi.org/10.5281/zenodo.10962379" ID-Zenodo-Dep="10962379" httpUri="https://zenodo.org/record/10962379/files/figure.png" pageId="4" pageNumber="5" startId="4.[197,288,858,881]" targetBox="[190,1437,163,833]" targetPageId="4" targetType="figure">
<paragraph id="8BE9AC2E3663CE3B13F9E9513D4AEFF2" blockId="4.[197,1438,858,911]" pageId="4" pageNumber="5">
<emphasis id="B922703C3663CE3B13F9E951395CEF0F" bold="true" box="[197,315,858,882]" pageId="4" pageNumber="5">FIGURE 3.</emphasis>
<taxonomicName id="4C56D7AD3663CE3B127FE9503A46EF0F" authority="Gašparič &amp; Hyžný &amp; Hitij &amp; Šoster, 2024" authorityName="Gašparič &amp; Hyžný &amp; Hitij &amp; Šoster" authorityYear="2024" box="[323,545,859,882]" class="Malacostraca" family="Hymenosomatidae" genus="Halicarcinus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="popeius" status="sp. nov.">
<emphasis id="B922703C3663CE3B127FE9503A46EF0F" box="[323,545,859,882]" italics="true" pageId="4" pageNumber="5">Halicarcinus popeius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A211CD473663CE3B1115E9503AE6EF0F" box="[553,641,859,882]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
, schematic reconstruction. A. Carapace in dorsal view; B. Detailed view of second type of rostrum and robust chelipeds (male); C. Third pereopod (P3) with detailed view of dactylus.
</paragraph>
</caption>
<subSubSection id="C34CFFA53663CE3B13FCE9CD3AA5E982" pageId="4" pageNumber="5" type="diagnosis">
<paragraph id="8BE9AC2E3663CE3B13FCE9CD3AA5E982" blockId="4.[192,793,966,1896]" pageId="4" pageNumber="5">
<emphasis id="B922703C3663CE3B13FCE9CD3920EFA2" bold="true" box="[192,327,966,991]" pageId="4" pageNumber="5">Diagnosis.</emphasis>
Carapace oval, wider than long, surrounded by a wide rim. Trilobate to trapezoid rostrum; length not extending as far as eyes. Eyes on short, stout eyestalks; extending beyond rostrum. Anterolateral corners forming a rounded postocular projection; either as rounded lobe or a pair of forward pointing teeth. Lateral side of carapace rounded; almost vertical posteriorly; with a concavity at posterolateral margin. Dorsal carapace with continuous gastro-cardiac groove, forming a hexagonal cardiac region. Chelipeds massive in males, sub-equal, homomorph, with swollen barrel-like propodus; more slender in females. Both fingers long, occluding with a gape; no dentation of the occlusal margin. Pereopods very long and slender. Dactylus broad, distinctively curved, tip composed of two developed, sub-equal, teeth; ventral edge armed by a row of blunt tubercles.
</paragraph>
</subSubSection>
<subSubSection id="C34CFFA53663CE3B13FCEC053CA8E835" pageId="4" pageNumber="5" type="materials_examined">
<paragraph id="8BE9AC2E3663CE3B13FCEC053B39E862" blockId="4.[192,793,966,1896]" lastBlockId="4.[840,1441,966,1911]" pageId="4" pageNumber="5">
<emphasis id="B922703C3663CE3B13FCEC053917EA5A" bold="true" box="[192,368,1550,1575]" pageId="4" pageNumber="5">Type material.</emphasis>
<materialsCitation id="3B3EA6733663CE3B1244EC053A05EA3A" pageId="4" pageNumber="5">
<typeStatus id="54ED128C3663CE3B1244EC053986EA5A" box="[376,481,1550,1575]" pageId="4" pageNumber="5" type="holotype">Holotype</typeStatus>
is
<collectionCode id="ED4734EB3663CE3B113EEC053A5CEA55" box="[514,571,1550,1576]" pageId="4" pageNumber="5">RGA</collectionCode>
/
<specimenCode id="DBF004553663CE3B117FEC053B7FEA5A" box="[579,792,1550,1576]" collectionCode="SMNH" country="Sweden" httpUri="http://grbio.org/cool/8z6v-f8is" name="Department of Paleozoology, Swedish Museum of Natural History" pageId="4" pageNumber="5">SMNH 6377-6378</specimenCode>
(part and counterpart) (
<figureCitation id="136DB0AB3663CE3B12E7EC253A3DEA3A" box="[475,602,1582,1607]" captionStart="FIGURE 4" captionStartId="5.[193,284,1453,1476]" captionTargetBox="[190,1436,167,1438]" captionTargetId="figure-121@5.[190,1436,167,1438]" captionTargetPageId="5" captionText="FIGURE 4. Halicarcinus popeius sp. nov. (AD) and Lucascinus trifailensis sp. nov. (E, F): A. RGA/SMNH 6377, dorsal, (holotype), visible hymenosomian rim (hr) and coloration patterns; B. RGA/SMNH 6373, dorsal, (paratype), male with well-preserved chelipeds and pereopods (P2-P5); C. RGA/SMNH 6385, dorsal, (paratype), male with bulbous chelipeds; D. RGA/SMNH 2041, ventral, (paratype), female with missing sternum; E. RGA/SMNH 6384, dorsal, (holotype), indicated rostrum and hymenosomian rim (hr); F. RGA/SMNH 6391, dorsal, (paratype), elongated chelipeds and well preserved pereopods (P2-P5). All scale bars equal 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10962381" httpUri="https://zenodo.org/record/10962381/files/figure.png" pageId="4" pageNumber="5">Figure 4A</figureCitation>
)
</materialsCitation>
,
<materialsCitation id="3B3EA6733663CE3B114DEC2539ACEB7A" pageId="4" pageNumber="5">
<typeStatus id="54ED128C3663CE3B114DEC253A81EA3A" box="[625,742,1582,1607]" pageId="4" pageNumber="5" type="paratype">paratypes</typeStatus>
are
<collectionCode id="ED4734EB3663CE3B13FCEC45389EEA15" box="[192,249,1614,1640]" pageId="4" pageNumber="5">RGA</collectionCode>
/
<specimenCode id="DBF004553663CE3B123DEC453983EA1A" box="[257,484,1614,1640]" collectionCode="SMNH" country="Sweden" httpUri="http://grbio.org/cool/8z6v-f8is" name="Department of Paleozoology, Swedish Museum of Natural History" pageId="4" pageNumber="5">SMNH 6373-6374</specimenCode>
(part and counterpart) (
<figureCitation id="136DB0AB3663CE3B13FAEC653925EAFA" box="[198,322,1646,1671]" captionStart="FIGURE 4" captionStartId="5.[193,284,1453,1476]" captionTargetBox="[190,1436,167,1438]" captionTargetId="figure-121@5.[190,1436,167,1438]" captionTargetPageId="5" captionText="FIGURE 4. Halicarcinus popeius sp. nov. (AD) and Lucascinus trifailensis sp. nov. (E, F): A. RGA/SMNH 6377, dorsal, (holotype), visible hymenosomian rim (hr) and coloration patterns; B. RGA/SMNH 6373, dorsal, (paratype), male with well-preserved chelipeds and pereopods (P2-P5); C. RGA/SMNH 6385, dorsal, (paratype), male with bulbous chelipeds; D. RGA/SMNH 2041, ventral, (paratype), female with missing sternum; E. RGA/SMNH 6384, dorsal, (holotype), indicated rostrum and hymenosomian rim (hr); F. RGA/SMNH 6391, dorsal, (paratype), elongated chelipeds and well preserved pereopods (P2-P5). All scale bars equal 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10962381" httpUri="https://zenodo.org/record/10962381/files/figure.png" pageId="4" pageNumber="5">Figure 4B</figureCitation>
,
<figureCitation id="136DB0AB3663CE3B1273EC653911EAF5" box="[335,374,1646,1672]" captionStart="FIGURE 6" captionStartId="7.[195,286,862,885]" captionTargetBox="[190,1438,167,837]" captionTargetId="figure-377@7.[190,1438,167,837]" captionTargetPageId="7" captionText="FIGURE 6. Lucascinus trifailensis sp. nov., schematic reconstruction. A. Carapace in dorsal view; B. Detailed view of male pleon." figureDoi="http://doi.org/10.5281/zenodo.10962385" httpUri="https://zenodo.org/record/10962385/files/figure.png" pageId="4" pageNumber="5">6C</figureCitation>
),
<collectionCode id="ED4734EB3663CE3B12B7EC6539A3EAF5" box="[395,452,1646,1672]" pageId="4" pageNumber="5">RGA</collectionCode>
/
<specimenCode id="DBF004553663CE3B12F7EC653AC5EAFA" box="[459,674,1646,1672]" collectionCode="SMNH" country="Sweden" httpUri="http://grbio.org/cool/8z6v-f8is" name="Department of Paleozoology, Swedish Museum of Natural History" pageId="4" pageNumber="5">SMNH 6371-6372</specimenCode>
(part and counterpart),
<collectionCode id="ED4734EB3663CE3B1257EC8539C3EAD5" box="[363,420,1678,1704]" pageId="4" pageNumber="5">RGA</collectionCode>
/
<specimenCode id="DBF004553663CE3B1290EC853AEAEADA" box="[428,653,1678,1704]" collectionCode="SMNH" country="Sweden" httpUri="http://grbio.org/cool/8z6v-f8is" name="Department of Paleozoology, Swedish Museum of Natural History" pageId="4" pageNumber="5">SMNH 6369-6370</specimenCode>
(part and counterpart),
<collectionCode id="ED4734EB3663CE3B1257ECA539C3EAB5" box="[363,420,1710,1736]" pageId="4" pageNumber="5">RGA</collectionCode>
/
<specimenCode id="DBF004553663CE3B1290ECA53AEAEABA" box="[428,653,1710,1736]" collectionCode="SMNH" country="Sweden" httpUri="http://grbio.org/cool/8z6v-f8is" name="Department of Paleozoology, Swedish Museum of Natural History" pageId="4" pageNumber="5">SMNH 6385-6386</specimenCode>
(part and counterpart) (
<figureCitation id="136DB0AB3663CE3B125BECC5399DEA95" box="[359,506,1742,1768]" captionStart="FIGURE 4" captionStartId="5.[193,284,1453,1476]" captionTargetBox="[190,1436,167,1438]" captionTargetId="figure-121@5.[190,1436,167,1438]" captionTargetPageId="5" captionText="FIGURE 4. Halicarcinus popeius sp. nov. (AD) and Lucascinus trifailensis sp. nov. (E, F): A. RGA/SMNH 6377, dorsal, (holotype), visible hymenosomian rim (hr) and coloration patterns; B. RGA/SMNH 6373, dorsal, (paratype), male with well-preserved chelipeds and pereopods (P2-P5); C. RGA/SMNH 6385, dorsal, (paratype), male with bulbous chelipeds; D. RGA/SMNH 2041, ventral, (paratype), female with missing sternum; E. RGA/SMNH 6384, dorsal, (holotype), indicated rostrum and hymenosomian rim (hr); F. RGA/SMNH 6391, dorsal, (paratype), elongated chelipeds and well preserved pereopods (P2-P5). All scale bars equal 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10962381" httpUri="https://zenodo.org/record/10962381/files/figure.png" pageId="4" pageNumber="5">Figures 4C</figureCitation>
,
<figureCitation id="136DB0AB3663CE3B1133ECC53A55EA9A" box="[527,562,1742,1767]" captionStart="FIGURE 5" captionStartId="6.[192,283,1448,1471]" captionTargetBox="[190,1434,163,1432]" captionTargetId="figure-124@6.[190,1435,163,1432]" captionTargetPageId="6" captionText="FIGURE 5. Scanning electron photomicrographs of Halicarcinus popeius sp. nov. (AC) and Lucascinus trifailensis sp. nov. (D): A. RGA/SMNH 6385, details of anterior and left lateral border in dorsal view, pereopods (P2-P5), (composite of several photomicrographs); B. RGA/SMNH 2041, ventral side with missing sternum, details of anterior margin and buccal cavity, coxa of pereopods (cx2-cx5), pereopods (P2-P5), (composite of several photomicrographs); C. RGA/ SMNH 6373, detailed view of pereopodal (P3) dactylus; D. RGA/SMNH 6387, ventral side with male pleon, pereopods (P2-P5), abdominal somites (s1-s5), pleotelson (pt), sternites (st4-st8), sternal shield (s), third maxilliped (mxp3). Scale bars indicated on the photomicrographs." figureDoi="http://doi.org/10.5281/zenodo.10962383" httpUri="https://zenodo.org/record/10962383/files/figure.png" pageId="4" pageNumber="5">5A</figureCitation>
) and
<collectionCode id="ED4734EB3663CE3B11B4ECC53AA6EA95" box="[648,705,1742,1768]" pageId="4" pageNumber="5">RGA</collectionCode>
/
<specimenCode id="DBF004553663CE3B11F5ECC5389CEB7A" collectionCode="SMNH" country="Sweden" httpUri="http://grbio.org/cool/8z6v-f8is" name="Department of Paleozoology, Swedish Museum of Natural History" pageId="4" pageNumber="5">SMNH 2041</specimenCode>
(
<figureCitation id="136DB0AB3663CE3B1235ECE539F5EB7A" box="[265,402,1774,1799]" captionStart="FIGURE 4" captionStartId="5.[193,284,1453,1476]" captionTargetBox="[190,1436,167,1438]" captionTargetId="figure-121@5.[190,1436,167,1438]" captionTargetPageId="5" captionText="FIGURE 4. Halicarcinus popeius sp. nov. (AD) and Lucascinus trifailensis sp. nov. (E, F): A. RGA/SMNH 6377, dorsal, (holotype), visible hymenosomian rim (hr) and coloration patterns; B. RGA/SMNH 6373, dorsal, (paratype), male with well-preserved chelipeds and pereopods (P2-P5); C. RGA/SMNH 6385, dorsal, (paratype), male with bulbous chelipeds; D. RGA/SMNH 2041, ventral, (paratype), female with missing sternum; E. RGA/SMNH 6384, dorsal, (holotype), indicated rostrum and hymenosomian rim (hr); F. RGA/SMNH 6391, dorsal, (paratype), elongated chelipeds and well preserved pereopods (P2-P5). All scale bars equal 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10962381" httpUri="https://zenodo.org/record/10962381/files/figure.png" pageId="4" pageNumber="5">Figures 4D</figureCitation>
,
<figureCitation id="136DB0AB3663CE3B12A2ECE539A4EB7A" box="[414,451,1774,1799]" captionStart="FIGURE 5" captionStartId="6.[192,283,1448,1471]" captionTargetBox="[190,1434,163,1432]" captionTargetId="figure-124@6.[190,1435,163,1432]" captionTargetPageId="6" captionText="FIGURE 5. Scanning electron photomicrographs of Halicarcinus popeius sp. nov. (AC) and Lucascinus trifailensis sp. nov. (D): A. RGA/SMNH 6385, details of anterior and left lateral border in dorsal view, pereopods (P2-P5), (composite of several photomicrographs); B. RGA/SMNH 2041, ventral side with missing sternum, details of anterior margin and buccal cavity, coxa of pereopods (cx2-cx5), pereopods (P2-P5), (composite of several photomicrographs); C. RGA/ SMNH 6373, detailed view of pereopodal (P3) dactylus; D. RGA/SMNH 6387, ventral side with male pleon, pereopods (P2-P5), abdominal somites (s1-s5), pleotelson (pt), sternites (st4-st8), sternal shield (s), third maxilliped (mxp3). Scale bars indicated on the photomicrographs." figureDoi="http://doi.org/10.5281/zenodo.10962383" httpUri="https://zenodo.org/record/10962383/files/figure.png" pageId="4" pageNumber="5">5B</figureCitation>
)
</materialsCitation>
.
<materialsCitation id="3B3EA6733663CE3B12EBECE53B3DE862" collectionCode="RGA" pageId="4" pageNumber="5" specimenCode="SMNH 2039-2040, SMNH 6375-6376, SMNH 6381-6382, SMNH 6389-6390" specimenCount="1">
Other specimens are
<collectionCode id="ED4734EB3663CE3B11EBECE53B77EB75" box="[727,784,1774,1800]" pageId="4" pageNumber="5">RGA</collectionCode>
/
<specimenCode id="DBF004553663CE3B13FCED0539FAEB5A" box="[192,413,1806,1832]" collectionCode="SMNH" country="Sweden" httpUri="http://grbio.org/cool/8z6v-f8is" name="Department of Paleozoology, Swedish Museum of Natural History" pageId="4" pageNumber="5">SMNH 2039-2040</specimenCode>
(part and counterpart),
<collectionCode id="ED4734EB3663CE3B11EBED053B77EB55" box="[727,784,1806,1832]" pageId="4" pageNumber="5">RGA</collectionCode>
/
<specimenCode id="DBF004553663CE3B13FCED2539FAEB3A" box="[192,413,1838,1864]" collectionCode="SMNH" country="Sweden" httpUri="http://grbio.org/cool/8z6v-f8is" name="Department of Paleozoology, Swedish Museum of Natural History" pageId="4" pageNumber="5">SMNH 6375-6376</specimenCode>
(part and counterpart),
<collectionCode id="ED4734EB3663CE3B11EBED253B77EB35" box="[727,784,1838,1864]" pageId="4" pageNumber="5">RGA</collectionCode>
/
<specimenCode id="DBF004553663CE3B13FCED4539F3EB1A" box="[192,404,1870,1896]" collectionCode="SMNH" country="Sweden" httpUri="http://grbio.org/cool/8z6v-f8is" name="Department of Paleozoology, Swedish Museum of Natural History" pageId="4" pageNumber="5">SMNH 6381-6382</specimenCode>
(part and counterpart) and
<collectionCode id="ED4734EB3663CE3B11EBED453B77EB15" box="[727,784,1870,1896]" pageId="4" pageNumber="5">RGA</collectionCode>
/
<specimenCode id="DBF004553663CE3B1074E9CD3C41EFA2" box="[840,1062,966,992]" collectionCode="SMNH" country="Sweden" httpUri="http://grbio.org/cool/8z6v-f8is" name="Department of Paleozoology, Swedish Museum of Natural History" pageId="4" pageNumber="5">SMNH 6389-6390</specimenCode>
(part and counterpart). Measurements are given in
<figureCitation id="136DB0AB3663CE3B1762E9ED3CA4EF82" box="[1118,1219,998,1023]" captionStart="FIGURE 7" captionStartId="8.[192,283,1078,1101]" captionTargetBox="[297,1337,169,1053]" captionTargetId="figure-13@8.[289,1343,165,1056]" captionTargetPageId="8" captionText="FIGURE 7. Carapace length vs. maximum carapace width of hymenosomatid specimens from the “Neža” locality. Halicarcinus popeius sp. nov. are marked in blue triangles (circles when trapezoid rostrum), Lucascinus trifailensis sp. nov. specimens are marked with orange squares. All measurements in mm." figureDoi="http://doi.org/10.5281/zenodo.10962387" httpUri="https://zenodo.org/record/10962387/files/figure.png" pageId="4" pageNumber="5">Figure 7</figureCitation>
,
<tableCitation id="C6D499953663CE3B17F3E9ED3D42EF82" box="[1231,1317,998,1023]" captionStart="TABLE 1" captionStartId="8.[188,268,1207,1230]" captionText="TABLE 1. Carapace length (CL) and carapace width (CW) measurements of hymenosomatid specimens from the “Neža” locality. All measurements in mm." pageId="4" pageNumber="5">Table 1</tableCitation>
and
<tableCitation id="C6D499953663CE3B165CE9ED3B3DE862" captionStart="TABLE 2" captionStartId="9.[192,270,165,188]" captionText="TABLE 2. Pereopod measurements of hymenosomatid specimens from the “Neža” locality. Cheliped (P1), Pereopod (P2-P5), dactylus (dac), propodus (pro), carpus (car), merus (mer), ischium (isc). All measurements in mm." pageId="4" pageNumber="5">Table 2</tableCitation>
</materialsCitation>
.
</paragraph>
<paragraph id="8BE9AC2E3663CE3B1074EE253CA8E835" blockId="4.[840,1441,966,1911]" box="[840,1231,1070,1096]" pageId="4" pageNumber="5">
<materialsCitation id="3B3EA6733663CE3B1074EE253CACE835" box="[840,1227,1070,1096]" country="Slovenia" location="Trbovlje" pageId="4" pageNumber="5" specimenCount="1" typeStatus="holotype">
<emphasis id="B922703C3663CE3B1074EE253B8AE83A" bold="true" box="[840,1005,1070,1095]" pageId="4" pageNumber="5">
<typeStatus id="54ED128C3663CE3B1074EE253BE3E83A" box="[840,900,1070,1095]" pageId="4" pageNumber="5">Type</typeStatus>
locality.
</emphasis>
<location id="8E89FAF53663CE3B10C8EE253C31E83A" LSID="urn:lsid:plazi:treatment:03FF1D383664CE38127CED0B397FE9BA:8E89FAF53663CE3B10C8EE253C31E83A" box="[1012,1110,1070,1095]" country="Slovenia" name="Trbovlje" pageId="4" pageNumber="5">Trbovlje</location>
,
<collectingCountry id="F341ECBE3663CE3B175DEE253CACE835" box="[1121,1227,1070,1096]" name="Slovenia" pageId="4" pageNumber="5">Slovenia</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34CFFA53663CE391074EE5D3AEAEB07" lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5" type="description">
<paragraph id="8BE9AC2E3663CE3B1074EE5D3C47EA32" blockId="4.[840,1441,966,1911]" pageId="4" pageNumber="5">
<emphasis id="B922703C3663CE3B1074EE5D3B85E812" bold="true" box="[840,994,1110,1135]" pageId="4" pageNumber="5">Description.</emphasis>
Carapace oval, wider than long (CL/ CW = 0.820.93) (
<tableCitation id="C6D499953663CE3B171AEE7D3C18E8F2" box="[1062,1151,1142,1167]" captionStart="TABLE 1" captionStartId="8.[188,268,1207,1230]" captionText="TABLE 1. Carapace length (CL) and carapace width (CW) measurements of hymenosomatid specimens from the “Neža” locality. All measurements in mm." pageId="4" pageNumber="5">Table 1</tableCitation>
), surrounded by a wide hymenosomian rim. Trilobate to trapezoid rostrum; wide and short; width approx. 0.10.2 times carapace width; length not extending as far as eyes (
<figureCitation id="136DB0AB3663CE3B1072EEFD3BB6E972" box="[846,977,1270,1295]" captionStart="FIGURE 5" captionStartId="6.[192,283,1448,1471]" captionTargetBox="[190,1434,163,1432]" captionTargetId="figure-124@6.[190,1435,163,1432]" captionTargetPageId="6" captionText="FIGURE 5. Scanning electron photomicrographs of Halicarcinus popeius sp. nov. (AC) and Lucascinus trifailensis sp. nov. (D): A. RGA/SMNH 6385, details of anterior and left lateral border in dorsal view, pereopods (P2-P5), (composite of several photomicrographs); B. RGA/SMNH 2041, ventral side with missing sternum, details of anterior margin and buccal cavity, coxa of pereopods (cx2-cx5), pereopods (P2-P5), (composite of several photomicrographs); C. RGA/ SMNH 6373, detailed view of pereopodal (P3) dactylus; D. RGA/SMNH 6387, ventral side with male pleon, pereopods (P2-P5), abdominal somites (s1-s5), pleotelson (pt), sternites (st4-st8), sternal shield (s), third maxilliped (mxp3). Scale bars indicated on the photomicrographs." figureDoi="http://doi.org/10.5281/zenodo.10962383" httpUri="https://zenodo.org/record/10962383/files/figure.png" pageId="4" pageNumber="5">Figure 5A</figureCitation>
). Lobes close together; sub equal in length; median lobe slightly bigger than the other two, sometimes not differentiated and forming a single trapezoidal rostrum (
<figureCitation id="136DB0AB3663CE3B17A0EF5D3D78E912" box="[1180,1311,1366,1391]" captionStart="FIGURE 4" captionStartId="5.[193,284,1453,1476]" captionTargetBox="[190,1436,167,1438]" captionTargetId="figure-121@5.[190,1436,167,1438]" captionTargetPageId="5" captionText="FIGURE 4. Halicarcinus popeius sp. nov. (AD) and Lucascinus trifailensis sp. nov. (E, F): A. RGA/SMNH 6377, dorsal, (holotype), visible hymenosomian rim (hr) and coloration patterns; B. RGA/SMNH 6373, dorsal, (paratype), male with well-preserved chelipeds and pereopods (P2-P5); C. RGA/SMNH 6385, dorsal, (paratype), male with bulbous chelipeds; D. RGA/SMNH 2041, ventral, (paratype), female with missing sternum; E. RGA/SMNH 6384, dorsal, (holotype), indicated rostrum and hymenosomian rim (hr); F. RGA/SMNH 6391, dorsal, (paratype), elongated chelipeds and well preserved pereopods (P2-P5). All scale bars equal 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10962381" httpUri="https://zenodo.org/record/10962381/files/figure.png" pageId="4" pageNumber="5">Figure 4B</figureCitation>
). Eyes on short, stout eyestalks; extending beyond rostrum (
<figureCitation id="136DB0AB3663CE3B1072EF9D3BA8E9D2" box="[846,975,1430,1455]" captionStart="FIGURE 5" captionStartId="6.[192,283,1448,1471]" captionTargetBox="[190,1434,163,1432]" captionTargetId="figure-124@6.[190,1435,163,1432]" captionTargetPageId="6" captionText="FIGURE 5. Scanning electron photomicrographs of Halicarcinus popeius sp. nov. (AC) and Lucascinus trifailensis sp. nov. (D): A. RGA/SMNH 6385, details of anterior and left lateral border in dorsal view, pereopods (P2-P5), (composite of several photomicrographs); B. RGA/SMNH 2041, ventral side with missing sternum, details of anterior margin and buccal cavity, coxa of pereopods (cx2-cx5), pereopods (P2-P5), (composite of several photomicrographs); C. RGA/ SMNH 6373, detailed view of pereopodal (P3) dactylus; D. RGA/SMNH 6387, ventral side with male pleon, pereopods (P2-P5), abdominal somites (s1-s5), pleotelson (pt), sternites (st4-st8), sternal shield (s), third maxilliped (mxp3). Scale bars indicated on the photomicrographs." figureDoi="http://doi.org/10.5281/zenodo.10962383" httpUri="https://zenodo.org/record/10962383/files/figure.png" pageId="4" pageNumber="5">Figure 5A</figureCitation>
). Anterolateral corners rounded, forming a rounded postocular projection; either as rounded lobe or a pair of forward pointing teeth. Lateral side of carapace rounded; almost vertical posteriorly; with a concavity at posterolateral corner (
<figureCitation id="136DB0AB3663CE3B1040EC3D3B86EA32" box="[892,993,1590,1615]" captionStart="FIGURE 4" captionStartId="5.[193,284,1453,1476]" captionTargetBox="[190,1436,167,1438]" captionTargetId="figure-121@5.[190,1436,167,1438]" captionTargetPageId="5" captionText="FIGURE 4. Halicarcinus popeius sp. nov. (AD) and Lucascinus trifailensis sp. nov. (E, F): A. RGA/SMNH 6377, dorsal, (holotype), visible hymenosomian rim (hr) and coloration patterns; B. RGA/SMNH 6373, dorsal, (paratype), male with well-preserved chelipeds and pereopods (P2-P5); C. RGA/SMNH 6385, dorsal, (paratype), male with bulbous chelipeds; D. RGA/SMNH 2041, ventral, (paratype), female with missing sternum; E. RGA/SMNH 6384, dorsal, (holotype), indicated rostrum and hymenosomian rim (hr); F. RGA/SMNH 6391, dorsal, (paratype), elongated chelipeds and well preserved pereopods (P2-P5). All scale bars equal 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10962381" httpUri="https://zenodo.org/record/10962381/files/figure.png" pageId="4" pageNumber="5">Figure 4</figureCitation>
A-D).
</paragraph>
<paragraph id="8BE9AC2E3663CE3A10BEEC553B6AEB7F" blockId="4.[840,1441,966,1911]" lastBlockId="5.[192,793,1673,1890]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
Dorsal carapace with continuous gastro-cardiac groove, extending concavely from one side of hymenosomian rim to the other; forming a hexagonal cardiac region (
<figureCitation id="136DB0AB3663CE3B170BECB53CB9EAAA" box="[1079,1246,1726,1751]" captionStart="FIGURE 4" captionStartId="5.[193,284,1453,1476]" captionTargetBox="[190,1436,167,1438]" captionTargetId="figure-121@5.[190,1436,167,1438]" captionTargetPageId="5" captionText="FIGURE 4. Halicarcinus popeius sp. nov. (AD) and Lucascinus trifailensis sp. nov. (E, F): A. RGA/SMNH 6377, dorsal, (holotype), visible hymenosomian rim (hr) and coloration patterns; B. RGA/SMNH 6373, dorsal, (paratype), male with well-preserved chelipeds and pereopods (P2-P5); C. RGA/SMNH 6385, dorsal, (paratype), male with bulbous chelipeds; D. RGA/SMNH 2041, ventral, (paratype), female with missing sternum; E. RGA/SMNH 6384, dorsal, (holotype), indicated rostrum and hymenosomian rim (hr); F. RGA/SMNH 6391, dorsal, (paratype), elongated chelipeds and well preserved pereopods (P2-P5). All scale bars equal 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10962381" httpUri="https://zenodo.org/record/10962381/files/figure.png" pageId="4" pageNumber="5">Figure 4A, D</figureCitation>
). Frontal region trapezoidal; not differentiated from gastric region posteriorly. Hepatic and prebranchial regions small; triangular. Postbranchial region square; separated from cardiac region by the thoracic groove. Cardiac region ornamented by three longitudinally oval areas of brighter colour (
<figureCitation id="136DB0AB3662CE3A12CCEC823A08EADF" box="[496,623,1673,1698]" captionStart="FIGURE 4" captionStartId="5.[193,284,1453,1476]" captionTargetBox="[190,1436,167,1438]" captionTargetId="figure-121@5.[190,1436,167,1438]" captionTargetPageId="5" captionText="FIGURE 4. Halicarcinus popeius sp. nov. (AD) and Lucascinus trifailensis sp. nov. (E, F): A. RGA/SMNH 6377, dorsal, (holotype), visible hymenosomian rim (hr) and coloration patterns; B. RGA/SMNH 6373, dorsal, (paratype), male with well-preserved chelipeds and pereopods (P2-P5); C. RGA/SMNH 6385, dorsal, (paratype), male with bulbous chelipeds; D. RGA/SMNH 2041, ventral, (paratype), female with missing sternum; E. RGA/SMNH 6384, dorsal, (holotype), indicated rostrum and hymenosomian rim (hr); F. RGA/SMNH 6391, dorsal, (paratype), elongated chelipeds and well preserved pereopods (P2-P5). All scale bars equal 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10962381" httpUri="https://zenodo.org/record/10962381/files/figure.png" pageId="5" pageNumber="6">Figure 4A</figureCitation>
); smaller oval coloration of dorsal carapace also in gastric region, anteriorly of gastro-cardiac groove; crescentshaped coloration of carapace at posterior border.
</paragraph>
<caption id="DF29FCA63662CE3A13FDEFA63BE5EA2A" ID-DOI="http://doi.org/10.5281/zenodo.10962381" ID-Zenodo-Dep="10962381" httpUri="https://zenodo.org/record/10962381/files/figure.png" pageId="5" pageNumber="6" startId="5.[193,284,1453,1476]" targetBox="[190,1436,167,1438]" targetPageId="5" targetType="figure">
<paragraph id="8BE9AC2E3662CE3A13FDEFA63BE5EA2A" blockId="5.[193,1435,1453,1623]" pageId="5" pageNumber="6">
<emphasis id="B922703C3662CE3A13FDEFA63950E9B8" bold="true" box="[193,311,1453,1477]" pageId="5" pageNumber="6">FIGURE 4.</emphasis>
<taxonomicName id="4C56D7AD3662CE3A1202EFA53A7BE9B8" authority="Gašparič &amp; Hyžný &amp; Hitij &amp; Šoster, 2024" authorityName="Gašparič &amp; Hyžný &amp; Hitij &amp; Šoster" authorityYear="2024" box="[318,540,1454,1477]" class="Malacostraca" family="Hymenosomatidae" genus="Halicarcinus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="popeius" status="sp. nov.">
<emphasis id="B922703C3662CE3A1202EFA53A7BE9B8" box="[318,540,1454,1477]" italics="true" pageId="5" pageNumber="6">Halicarcinus popeius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A211CD473662CE3A111FEFA53A10E9B8" box="[547,631,1454,1477]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
(AD) and
<taxonomicName id="4C56D7AD3662CE3A11CEEFA53BBBE9B9" authority="Gašparič &amp; Hyžný &amp; Hitij &amp; Šoster, 2024" authorityName="Gašparič &amp; Hyžný &amp; Hitij &amp; Šoster" authorityYear="2024" box="[754,988,1453,1477]" class="Malacostraca" family="Hymenosomatidae" genus="Lucascinus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="trifailensis" status="sp. nov.">
<emphasis id="B922703C3662CE3A11CEEFA53BBBE9B9" box="[754,988,1453,1477]" italics="true" pageId="5" pageNumber="6">Lucascinus trifailensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A211CD473662CE3A10DFEFA53C50E9B8" box="[995,1079,1454,1477]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
(E, F): A. RGA/SMNH 6377, dorsal, (holotype), visible hymenosomian rim (hr) and coloration patterns; B. RGA/SMNH 6373, dorsal, (paratype), male with well-preserved chelipeds and pereopods (P2-P5); C. RGA/SMNH 6385, dorsal, (paratype), male with bulbous chelipeds; D. RGA/SMNH 2041, ventral, (paratype), female with missing sternum; E. RGA/SMNH 6384, dorsal, (holotype), indicated rostrum and hymenosomian rim (hr); F. RGA/SMNH 6391, dorsal, (paratype), elongated chelipeds and well preserved pereopods (P2-P5). All scale bars equal 5 mm.
</paragraph>
</caption>
<paragraph id="8BE9AC2E3662CE3913C6ED033AEAEB07" blockId="5.[192,793,1673,1890]" lastBlockId="6.[192,793,1696,1914]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
Chelipeds massive, with swollen barrel-like propodus (
<figureCitation id="136DB0AB3662CE3A127FED22398AEB3F" box="[323,493,1832,1858]" captionStart="FIGURE 4" captionStartId="5.[193,284,1453,1476]" captionTargetBox="[190,1436,167,1438]" captionTargetId="figure-121@5.[190,1436,167,1438]" captionTargetPageId="5" captionText="FIGURE 4. Halicarcinus popeius sp. nov. (AD) and Lucascinus trifailensis sp. nov. (E, F): A. RGA/SMNH 6377, dorsal, (holotype), visible hymenosomian rim (hr) and coloration patterns; B. RGA/SMNH 6373, dorsal, (paratype), male with well-preserved chelipeds and pereopods (P2-P5); C. RGA/SMNH 6385, dorsal, (paratype), male with bulbous chelipeds; D. RGA/SMNH 2041, ventral, (paratype), female with missing sternum; E. RGA/SMNH 6384, dorsal, (holotype), indicated rostrum and hymenosomian rim (hr); F. RGA/SMNH 6391, dorsal, (paratype), elongated chelipeds and well preserved pereopods (P2-P5). All scale bars equal 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10962381" httpUri="https://zenodo.org/record/10962381/files/figure.png" pageId="5" pageNumber="6">Figure 4A, C</figureCitation>
); length approx. 1.1-1.7 times of carapace length; slighter in females. Ischium short; merus slightly longer than wide; carpus longer than merus, swollen with rounded margins, almost as broad as long; manus greatly inflated, margins rounded and convex. Both fingers long, occluding with a gape; no dentation of the occlusal margin observable. Pereopods very long and slender (
<figureCitation id="136DB0AB3662CE3A10CCED423C15EB1F" box="[1008,1138,1865,1890]" captionStart="FIGURE 4" captionStartId="5.[193,284,1453,1476]" captionTargetBox="[190,1436,167,1438]" captionTargetId="figure-121@5.[190,1436,167,1438]" captionTargetPageId="5" captionText="FIGURE 4. Halicarcinus popeius sp. nov. (AD) and Lucascinus trifailensis sp. nov. (E, F): A. RGA/SMNH 6377, dorsal, (holotype), visible hymenosomian rim (hr) and coloration patterns; B. RGA/SMNH 6373, dorsal, (paratype), male with well-preserved chelipeds and pereopods (P2-P5); C. RGA/SMNH 6385, dorsal, (paratype), male with bulbous chelipeds; D. RGA/SMNH 2041, ventral, (paratype), female with missing sternum; E. RGA/SMNH 6384, dorsal, (holotype), indicated rostrum and hymenosomian rim (hr); F. RGA/SMNH 6391, dorsal, (paratype), elongated chelipeds and well preserved pereopods (P2-P5). All scale bars equal 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10962381" httpUri="https://zenodo.org/record/10962381/files/figure.png" pageId="5" pageNumber="6">Figure 4B</figureCitation>
,
<tableCitation id="C6D499953662CE3A17B8ED423C81EB1F" box="[1156,1254,1865,1890]" captionStart="TABLE 2" captionStartId="9.[192,270,165,188]" captionText="TABLE 2. Pereopod measurements of hymenosomatid specimens from the “Neža” locality. Cheliped (P1), Pereopod (P2-P5), dactylus (dac), propodus (pro), carpus (car), merus (mer), ischium (isc). All measurements in mm." pageId="5" pageNumber="6">Table 2</tableCitation>
), almost same length, approx. 2.0-3.5 times of carapace length. Ischium very short; merus longest; carpus about one-thirds as long as merus; propodus almost as long as merus. Dactylus shorter than the propodus, broad, distinctively curved, tip composed of two developed, sub-equal, teeth; ventral edge armed by a row of blunt tubercles (
<figureCitation id="136DB0AB3661CE391139ED6B3AE6EB07" box="[517,641,1888,1914]" captionStart="FIGURE 5" captionStartId="6.[192,283,1448,1471]" captionTargetBox="[190,1434,163,1432]" captionTargetId="figure-124@6.[190,1435,163,1432]" captionTargetPageId="6" captionText="FIGURE 5. Scanning electron photomicrographs of Halicarcinus popeius sp. nov. (AC) and Lucascinus trifailensis sp. nov. (D): A. RGA/SMNH 6385, details of anterior and left lateral border in dorsal view, pereopods (P2-P5), (composite of several photomicrographs); B. RGA/SMNH 2041, ventral side with missing sternum, details of anterior margin and buccal cavity, coxa of pereopods (cx2-cx5), pereopods (P2-P5), (composite of several photomicrographs); C. RGA/ SMNH 6373, detailed view of pereopodal (P3) dactylus; D. RGA/SMNH 6387, ventral side with male pleon, pereopods (P2-P5), abdominal somites (s1-s5), pleotelson (pt), sternites (st4-st8), sternal shield (s), third maxilliped (mxp3). Scale bars indicated on the photomicrographs." figureDoi="http://doi.org/10.5281/zenodo.10962383" httpUri="https://zenodo.org/record/10962383/files/figure.png" pageId="6" pageNumber="7">Figure 5C</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF29FCA63661CE3913FCEFA33AC8EA0D" ID-DOI="http://doi.org/10.5281/zenodo.10962383" ID-Zenodo-Dep="10962383" httpUri="https://zenodo.org/record/10962383/files/figure.png" pageId="6" pageNumber="7" startId="6.[192,283,1448,1471]" targetBox="[190,1434,163,1432]" targetPageId="6" targetType="figure">
<paragraph id="8BE9AC2E3661CE3913FCEFA33AC8EA0D" blockId="6.[192,1438,1448,1648]" pageId="6" pageNumber="7">
<emphasis id="B922703C3661CE3913FCEFA33952E9BD" bold="true" box="[192,309,1448,1472]" pageId="6" pageNumber="7">FIGURE 5.</emphasis>
Scanning electron photomicrographs of
<taxonomicName id="4C56D7AD3661CE3911DCEFA23BDAE9BD" authority="Gašparič &amp; Hyžný &amp; Hitij &amp; Šoster, 2024" authorityName="Gašparič &amp; Hyžný &amp; Hitij &amp; Šoster" authorityYear="2024" box="[736,957,1449,1472]" class="Malacostraca" family="Hymenosomatidae" genus="Halicarcinus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="popeius" status="sp. nov.">
<emphasis id="B922703C3661CE3911DCEFA23BDAE9BD" box="[736,957,1449,1472]" italics="true" pageId="6" pageNumber="7">Halicarcinus popeius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A211CD473661CE3910FFEFA23C71E9BD" box="[963,1046,1449,1472]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
(AC) and
<taxonomicName id="4C56D7AD3661CE3917B2EFA23D10E9C2" authority="Gašparič &amp; Hyžný &amp; Hitij &amp; Šoster, 2024" authorityName="Gašparič &amp; Hyžný &amp; Hitij &amp; Šoster" authorityYear="2024" box="[1166,1399,1448,1472]" class="Malacostraca" family="Hymenosomatidae" genus="Lucascinus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="trifailensis" status="sp. nov.">
<emphasis id="B922703C3661CE3917B2EFA23D10E9C2" box="[1166,1399,1448,1472]" italics="true" pageId="6" pageNumber="7">Lucascinus trifailensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A211CD473661CE391641EFA2388BE9A0" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
(D): A. RGA/SMNH 6385, details of anterior and left lateral border in dorsal view, pereopods (P2-P5), (composite of several photomicrographs); B. RGA/SMNH 2041, ventral side with missing sternum, details of anterior margin and buccal cavity, coxa of pereopods (cx2-cx5), pereopods (P2-P5), (composite of several photomicrographs); C. RGA/ SMNH 6373, detailed view of pereopodal (P3) dactylus; D. RGA/SMNH 6387, ventral side with male pleon, pereopods (P2-P5), abdominal somites (s1-s5), pleotelson (pt), sternites (st4-st8), sternal shield (s), third maxilliped (mxp3). Scale bars indicated on the photomicrographs.
</paragraph>
</caption>
<subSubSection id="C34CFFA53661CE381074ECAB397FE9BA" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="discussion">
<paragraph id="8BE9AC2E3661CE381074ECAB3960E92A" blockId="6.[840,1441,1696,1913]" lastBlockId="7.[192,793,958,1479]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<emphasis id="B922703C3661CE391074ECAB3BA7EAC4" bold="true" box="[840,960,1696,1721]" pageId="6" pageNumber="7">Remarks.</emphasis>
<taxonomicName id="4C56D7AD3661CE3910F6ECAB3CA5EAC4" authority="Gašparič &amp; Hyžný &amp; Hitij &amp; Šoster, 2024" authorityName="Gašparič &amp; Hyžný &amp; Hitij &amp; Šoster" authorityYear="2024" box="[970,1218,1696,1721]" class="Malacostraca" family="Hymenosomatidae" genus="Halicarcinus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="popeius" status="sp. nov.">
<emphasis id="B922703C3661CE3910F6ECAB3CA5EAC4" box="[970,1218,1696,1721]" italics="true" pageId="6" pageNumber="7">Halicarcinus popeius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A211CD473661CE3917F0ECAB3D47EAC4" box="[1228,1312,1696,1721]" pageId="6" pageNumber="7" rank="species">sp.nov.</taxonomicNameLabel>
is the first fossil representative of the genus. Thus, by the general habitus it would be most similar to
<taxonomicName id="4C56D7AD3661CE391679ECEB3D61EB64" authority="(H. Milne Edwards, 1853)" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1853" class="Malacostraca" family="Hymenosomatidae" genus="Halicarcinus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="quoyi">
<emphasis id="B922703C3661CE391679ECEB3BA9EB64" italics="true" pageId="6" pageNumber="7">Halicarcinus quoyi</emphasis>
(H.
<bibRefCitation id="EFC7D1DF3661CE39173CED0B3C99EB64" author="Milne Edwards, H." box="[1024,1278,1792,1817]" pageId="6" pageNumber="7" refId="ref8169" refString="Milne Edwards, H. 1853. Memoire sur la famille des Ocypodiens. Annales des Sciences Naturelles (ser. 3), 20: 163 - 228." year="1853">Milne Edwards, 1853</bibRefCitation>
)
</taxonomicName>
, with which it shares a wider than long carapace with broad rim, a trilobed rostrum, shape of pereopodal dactyli and strongly swollen, bulbous propodi of male chelipeds. The new fossil species differs from
<taxonomicName id="4C56D7AD3660CE381188E9B53B7FEFAA" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1853" box="[692,792,958,983]" class="Malacostraca" family="Hymenosomatidae" genus="Halicarcinus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="quoyi">
<emphasis id="B922703C3660CE381188E9B53B7FEFAA" box="[692,792,958,983]" italics="true" pageId="7" pageNumber="8">H. quoyi</emphasis>
</taxonomicName>
in possessing postorbital projections at anterolateral corners of carapace and in development of dorsal carapace regions, especially wide hexagonal cardiac region. Additionally, we recognize heterogeneity in development of rostra within the fossil material, which can be explained as intraspecific variations or possible sexual dimorphism.
<bibRefCitation id="EFC7D1DF3660CE3813FCEEB5393BE8AA" author="Lucas, J. S." box="[192,348,1214,1239]" pageId="7" pageNumber="8" refId="ref7874" refString="Lucas, J. S. 1981. Spider crabs of the family Hymenosomatidae (Crustacea; Brachyura) with particular reference to Australian species: systematics and biology. Records of the Australian Museum, 33: 148 - 247. https: // doi. org / 10.3853 / j. 0067 - 1975.33.1980.199" year="1981">Lucas (1981)</bibRefCitation>
previously recognized that the shape and relative length of the rostrum within
<taxonomicName id="4C56D7AD3660CE381195EED5388CE96A" authorityName="White" authorityYear="1846" class="Malacostraca" family="Hymenosomatidae" genus="Halicarcinus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B922703C3660CE381195EED5388CE96A" italics="true" pageId="7" pageNumber="8">Halicarcinus</emphasis>
</taxonomicName>
are sometimes sexually dimorphic, although such variations were not reported in
<taxonomicName id="4C56D7AD3660CE3811BBEF153967E92A" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1853" class="Malacostraca" family="Hymenosomatidae" genus="Halicarcinus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="quoyi">
<emphasis id="B922703C3660CE3811BBEF153967E92A" italics="true" pageId="7" pageNumber="8">Halicarcinus quoyi</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="DF29FCA63660CE3813FFE9553927EFEF" ID-DOI="http://doi.org/10.5281/zenodo.10962385" ID-Zenodo-Dep="10962385" httpUri="https://zenodo.org/record/10962385/files/figure.png" pageId="7" pageNumber="8" startId="7.[195,286,862,885]" targetBox="[190,1438,167,837]" targetPageId="7" targetType="figure">
<paragraph id="8BE9AC2E3660CE3813FFE9553927EFEF" blockId="7.[195,1435,862,914]" pageId="7" pageNumber="8">
<emphasis id="B922703C3660CE3813FFE955395EEF08" bold="true" box="[195,313,862,885]" pageId="7" pageNumber="8">FIGURE 6.</emphasis>
<taxonomicName id="4C56D7AD3660CE38127CE9553A4EEF08" authority="Gašparič &amp; Hyžný &amp; Hitij &amp; Šoster, 2024" authorityName="Gašparič &amp; Hyžný &amp; Hitij &amp; Šoster" authorityYear="2024" box="[320,553,862,885]" class="Malacostraca" family="Hymenosomatidae" genus="Lucascinus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="trifailensis" status="sp. nov.">
<emphasis id="B922703C3660CE38127CE9553A4EEF08" box="[320,553,862,885]" italics="true" pageId="7" pageNumber="8">Lucascinus trifailensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A211CD473660CE38110CE9553AE0EF08" box="[560,647,862,885]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
, schematic reconstruction. A. Carapace in dorsal view; B. Detailed view of male pleon.
</paragraph>
</caption>
<paragraph id="8BE9AC2E3660CE3813FCEF6D3A3FE902" blockId="7.[192,793,958,1479]" box="[192,600,1381,1407]" pageId="7" pageNumber="8">
<emphasis id="B922703C3660CE3813FCEF6D397EE902" bold="true" box="[192,281,1382,1407]" pageId="7" pageNumber="8">Range.</emphasis>
Late Oligocene (Chattian).
</paragraph>
<paragraph id="8BE9AC2E3660CE3813FCEF86397FE9BA" blockId="7.[192,793,958,1479]" pageId="7" pageNumber="8">
<emphasis id="B922703C3660CE3813FCEF86393CE9DA" bold="true" box="[192,347,1421,1447]" pageId="7" pageNumber="8">Occurrence.</emphasis>
To date, known only from the
<typeStatus id="54ED128C3660CE3811DAEF853B7FE9DA" box="[742,792,1422,1447]" pageId="7" pageNumber="8">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>