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<document id="51BC12A23D2D28FA8A076FAAD0EE6838" ID-CLB-Dataset="45975" ID-DOI="10.11646/zootaxa.3790.1.4" ID-GBIF-Dataset="c7663905-b87e-40e6-a2d7-b8e3a1819299" ID-ISSN="1175-5326" ID-Zenodo-Dep="252378" ID-ZooBank="45DFDC6F-6B3B-4706-B19E-F3661852CC99" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460410258542" checkinUser="plazi" docAuthor="Michelsen, Verner" docDate="2014" docId="03FC87FACB39AB73FF36FF061C8C6FE8" docLanguage="en" docName="zt03790p102.pdf" docOrigin="Zootaxa 3790 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ringdahlia Michelsen, 2014, gen. n." docType="treatment" docVersion="9" lastPageNumber="97" masterDocId="FFC5FF82CB32AB78FFA1FF9E1C666B0A" masterDocTitle="Taxonomic assessment of Chirosiomima Hennig (Diptera: Anthomyiidae), with proposal of a new genus for Hylemyia curtigena Ringdahl" masterLastPageNumber="102" masterPageNumber="86" pageNumber="97" updateTime="1698300401450" updateUser="ExternalLinkService">
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<mods:title id="B7493A14052419E6500337C5B80D4FEB">Taxonomic assessment of Chirosiomima Hennig (Diptera: Anthomyiidae), with proposal of a new genus for Hylemyia curtigena Ringdahl</mods:title>
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<treatment id="03FC87FACB39AB73FF36FF061C8C6FE8" ID-DOI="http://doi.org/10.5281/zenodo.6126230" ID-GBIF-Taxon="119423204" ID-Zenodo-Dep="6126230" LSID="urn:lsid:plazi:treatment:03FC87FACB39AB73FF36FF061C8C6FE8" httpUri="http://treatment.plazi.org/id/03FC87FACB39AB73FF36FF061C8C6FE8" lastPageNumber="97" pageId="11" pageNumber="97">
<subSubSection id="C34F6567CB39AB73FF36FF061DA96BB8" box="[151,463,151,178]" pageId="11" pageNumber="97" type="nomenclature">
<paragraph id="8BEA36ECCB39AB73FF36FF061DA96BB8" blockId="11.[151,463,151,178]" box="[151,463,151,178]" pageId="11" pageNumber="97">
<heading id="D0A28180CB39AB73FF36FF061DA96BB8" bold="true" box="[151,463,151,178]" fontSize="11" level="1" pageId="11" pageNumber="97" reason="1">
<emphasis id="B921EAFECB39AB73FF36FF061DA96BB8" bold="true" box="[151,463,151,178]" pageId="11" pageNumber="97">
Genus
<taxonomicName id="4C554D6FCB39AB73FF4FFF091D136BBB" ID-CoL="89RDH" box="[238,373,151,177]" class="Insecta" family="Anthomyiidae" genus="Ringdahlia" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="97" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B921EAFECB39AB73FF4FFF091D136BBB" bold="true" box="[238,373,151,177]" italics="true" pageId="11" pageNumber="97">Ringdahlia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2125785CB39AB73FEDDFF061DA96BB8" box="[380,463,152,178]" pageId="11" pageNumber="97" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34F6567CB39AB73FF36FF411EFF6BFE" box="[151,665,222,244]" pageId="11" pageNumber="97" type="materials_examined">
<paragraph id="8BEA36ECCB39AB73FF36FF411EFF6BFE" blockId="11.[151,665,222,244]" box="[151,665,222,244]" pageId="11" pageNumber="97">
<typeStatus id="54EE884ECB39AB73FF36FF411CAF6BFE" box="[151,201,223,244]" pageId="11" pageNumber="97">Type</typeStatus>
species:
<taxonomicName id="4C554D6FCB39AB73FE86FF411EF36BFE" authority="Ringdahl, 1935" authorityName="Ringdahl" authorityYear="1935" box="[295,661,222,244]" class="Insecta" family="Anthomyiidae" genus="Hylemyia" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="97" phylum="Arthropoda" rank="species" species="curtigena">
<emphasis id="B921EAFECB39AB73FE86FF411D896BFE" box="[295,495,223,244]" italics="true" pageId="11" pageNumber="97">Hylemyia curtigena</emphasis>
<bibRefCitation id="EFC44B1DCB39AB73FE54FF401EF36BFE" author="Ringdahl" box="[501,661,222,244]" pageId="11" pageNumber="97" refString="Ringdahl, O. (1935) Neue fennoskandische Musciden. Notulae entomologicae, 15, 26 - 31." type="journal article" year="1935">Ringdahl, 1935</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C34F6567CB39AB73FF36FEBF1F0E6AC0" pageId="11" pageNumber="97" type="etymology">
<paragraph id="8BEA36ECCB39AB73FF36FEBF1F0E6AC0" blockId="11.[151,1437,289,1250]" pageId="11" pageNumber="97">
<emphasis id="B921EAFECB39AB73FF36FEBF1D786A30" bold="true" box="[151,286,289,314]" pageId="11" pageNumber="97">Etymology.</emphasis>
The new genus is named after the Swedish dipterist Oscar Ringdahl (
<date id="FFEB102CCB39AB73FB9BFEBF18DB6A30" box="[1082,1213,289,314]" pageId="11" pageNumber="97" value="1885" valueMax="1966">18851966</date>
), author of the
<typeStatus id="54EE884ECB39AB73FACDFEBC19FA6A30" box="[1388,1436,290,314]" pageId="11" pageNumber="97">type</typeStatus>
species. He was a dedicated collector of
<taxonomicName id="4C554D6FCB39AB73FDFFFEDB1F5E6A54" authority="Brachycera" authorityName="Brachycera" box="[606,824,325,350]" kingdom="Plantae" order="Diptera" pageId="11" pageNumber="97" rank="order">Diptera Brachycera</taxonomicName>
from all parts of
<collectingCountry id="F342767CCB39AB73FC5CFEDB183F6A54" box="[1021,1113,325,350]" name="Sweden" pageId="11" pageNumber="97">Sweden</collectingCountry>
. His early acquaintance with the
<collectingCountry id="F342767CCB39AB73FF64FEF71D796A8B" box="[197,287,361,385]" name="Germany" pageId="11" pageNumber="97">German</collectingCountry>
dipterist Paul Stein (
<date id="FFEB102CCB39AB73FDBCFEF71EF86A88" box="[541,670,361,386]" pageId="11" pageNumber="97" value="1852" valueMax="1921">18521921</date>
) may have given him the impetus to take special interest in the muscoid families (
<taxonomicName id="4C554D6FCB39AB73FED3FE131D856AAC" box="[370,483,397,422]" class="Insecta" family="Muscidae" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="97" phylum="Arthropoda" rank="family">Muscidae</taxonomicName>
,
<taxonomicName id="4C554D6FCB39AB73FE54FE131E056AAC" box="[501,611,397,422]" class="Insecta" family="Fanniidae" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="97" phylum="Arthropoda" rank="family">Fanniidae</taxonomicName>
and
<taxonomicName id="4C554D6FCB39AB73FD06FE131F2B6AAC" box="[679,845,397,422]" class="Insecta" family="Anthomyiidae" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="97" phylum="Arthropoda" rank="family">Anthomyiidae</taxonomicName>
). He soon became the foremost authority on this important but notoriously difficult group of flies in Scandinavia.
</paragraph>
</subSubSection>
<subSubSection id="C34F6567CB39AB73FF66FE4B1C8C6FE8" pageId="11" pageNumber="97" type="diagnosis">
<paragraph id="8BEA36ECCB39AB73FF66FE4B1F63685F" blockId="11.[151,1437,289,1250]" pageId="11" pageNumber="97">
<emphasis id="B921EAFECB39AB73FF66FE4B1D5C6AE4" bold="true" box="[199,314,469,494]" pageId="11" pageNumber="97">Diagnosis</emphasis>
. The following combination of characters may help to separate male and female
<taxonomicName id="4C554D6FCB39AB73FB41FE4B193B6AE7" box="[1248,1373,469,493]" class="Insecta" family="Anthomyiidae" genus="Ringdahlia" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="97" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B921EAFECB39AB73FB41FE4B193B6AE7" box="[1248,1373,469,493]" italics="true" pageId="11" pageNumber="97">Ringdahlia</emphasis>
</taxonomicName>
from other anthomyiid genera: medium sized, sexually dimorphic, body and appendages extensively dark-coloured; arista short-pubescent; male head (
<figureCitation id="136E2A69CB39AB73FD8CFD831EE4693C" box="[557,642,541,566]" captionStart="FIGURES 27 30" captionStartId="12.[151,264,1623,1646]" captionTargetBox="[174,1405,219,1602]" captionTargetId="figure@12.[151,1436,193,1602]" captionTargetPageId="12" captionText="FIGURES 27 30. Ringdahlia curtigena. 27, 28. Male from Finland, habitus and head, lateral views. 29. Female from Finland, labella. 30. Female from Finland, head and anterior thorax, anterodorsal view. Scale bars: 1 mm (27); 0.5 mm (28, 30); 0.1 mm (29)." httpUri="https://zenodo.org/record/252385/files/figure.png" pageId="11" pageNumber="97">Fig. 28</figureCitation>
) in lateral view with antenna inserted beneath the middle and gena narrow, only half as wide as postpedicel; female frons relatively narrow, on lower part only one-third of total width of head (
<figureCitation id="136E2A69CB39AB73FF5DFDFB1D346974" box="[252,338,613,638]" captionStart="FIGURES 27 30" captionStartId="12.[151,264,1623,1646]" captionTargetBox="[174,1405,219,1602]" captionTargetId="figure@12.[151,1436,193,1602]" captionTargetPageId="12" captionText="FIGURES 27 30. Ringdahlia curtigena. 27, 28. Male from Finland, habitus and head, lateral views. 29. Female from Finland, labella. 30. Female from Finland, head and anterior thorax, anterodorsal view. Scale bars: 1 mm (27); 0.5 mm (28, 30); 0.1 mm (29)." httpUri="https://zenodo.org/record/252385/files/figure.png" pageId="11" pageNumber="97">Fig. 30</figureCitation>
); setulae on upper occiput and anterior declivity of thorax rather coarse and blunt (
<figureCitation id="136E2A69CB39AB73FAA2FDFB19E96974" box="[1283,1423,613,638]" captionStart="FIGURES 27 30" captionStartId="12.[151,264,1623,1646]" captionTargetBox="[174,1405,219,1602]" captionTargetId="figure@12.[151,1436,193,1602]" captionTargetPageId="12" captionText="FIGURES 27 30. Ringdahlia curtigena. 27, 28. Male from Finland, habitus and head, lateral views. 29. Female from Finland, labella. 30. Female from Finland, head and anterior thorax, anterodorsal view. Scale bars: 1 mm (27); 0.5 mm (28, 30); 0.1 mm (29)." httpUri="https://zenodo.org/record/252385/files/figure.png" pageId="11" pageNumber="97">Figs. 28, 30</figureCitation>
); presutural rows of acrostichals wider apart from each other than their distance to dorsocentral rows and with setulae present between rows (
<figureCitation id="136E2A69CB39AB73FE4CFD331E2769CC" box="[493,577,685,710]" captionStart="FIGURES 27 30" captionStartId="12.[151,264,1623,1646]" captionTargetBox="[174,1405,219,1602]" captionTargetId="figure@12.[151,1436,193,1602]" captionTargetPageId="12" captionText="FIGURES 27 30. Ringdahlia curtigena. 27, 28. Male from Finland, habitus and head, lateral views. 29. Female from Finland, labella. 30. Female from Finland, head and anterior thorax, anterodorsal view. Scale bars: 1 mm (27); 0.5 mm (28, 30); 0.1 mm (29)." httpUri="https://zenodo.org/record/252385/files/figure.png" pageId="11" pageNumber="97">Fig. 30</figureCitation>
); vein C with
<emphasis id="B921EAFECB39AB73FD7FFD301E8C69CF" box="[734,746,686,709]" italics="true" pageId="11" pageNumber="97">v</emphasis>
setulae; mid tibia with 0
<emphasis id="B921EAFECB39AB73FBA7FD30187869CF" box="[1030,1054,686,709]" italics="true" pageId="11" pageNumber="97">av</emphasis>
, 1
<emphasis id="B921EAFECB39AB73FBE0FD33183B69CF" box="[1089,1117,685,709]" italics="true" pageId="11" pageNumber="97">ad</emphasis>
, 46
<emphasis id="B921EAFECB39AB73FB3DFD3318DE69CF" box="[1180,1208,685,709]" italics="true" pageId="11" pageNumber="97">pd</emphasis>
and 0
<emphasis id="B921EAFECB39AB73FAA3FD30197B69CF" box="[1282,1309,686,709]" italics="true" pageId="11" pageNumber="97">pv</emphasis>
setae; hind tibia with 34
<emphasis id="B921EAFECB39AB73FE9BFD4F1D3069E3" box="[314,342,721,745]" italics="true" pageId="11" pageNumber="97">pd</emphasis>
setae, without
<emphasis id="B921EAFECB39AB73FE5EFD4C1E6B69E3" box="[511,525,722,745]" italics="true" pageId="11" pageNumber="97">p</emphasis>
or
<emphasis id="B921EAFECB39AB73FD92FD4C1E2869E3" box="[563,590,722,745]" italics="true" pageId="11" pageNumber="97">pv</emphasis>
setae, apically without
<emphasis id="B921EAFECB39AB73FCF6FD4F1F1569E3" box="[855,883,721,745]" italics="true" pageId="11" pageNumber="97">pd</emphasis>
and
<emphasis id="B921EAFECB39AB73FC0BFD4C1FA369E3" box="[938,965,722,745]" italics="true" pageId="11" pageNumber="97">pv</emphasis>
setae; male terminalia of unique structure in respect to epandrium, cercal plate and surstyli (
<figureCitation id="136E2A69CB39AB73FD6AFD6B1F336804" box="[715,853,757,782]" captionStart="FIGURES 31 42" captionStartId="13.[151,264,1864,1887]" captionTargetBox="[235,1334,212,1814]" captionTargetId="figure@13.[206,1359,193,1828]" captionTargetPageId="13" captionText="FIGURES 31 42. Ringdahlia curtigena, males from Finland. 31. Tergosternites VI VIII and sternites IV and V, laterodorsal view. 32. Sternite V, lateral view. 33. Hypopygium, posterior view. 34. Same, after removal of cerci. 35. Hypopygium, lateral view. 36 39. Cerci, dorsal, anterior, anteroventral and lateral views (x = ventral sclerite complex; y = bowed sclerite rod. 40. Pregonite (left) and postgonite, lateral view, 41. Phallapodeme, phallus and ejaculatory pump, lateral view. 42. Phallapodeme and phallus, anterolateral view. Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/252386/files/figure.png" pageId="11" pageNumber="97">Figs. 3339</figureCitation>
); female oviscapt (
<figureCitation id="136E2A69CB39AB73FB8AFD6B181B6804" box="[1067,1149,757,782]" captionStart="FIGURE 43" captionStartId="14.[151,250,1402,1424]" captionTargetBox="[223,1356,210,1348]" captionTargetId="figure@14.[216,1371,193,1381]" captionTargetPageId="14" captionText="FIGURE 43. Ringdahlia curtigena, female from Finland, oviscapt, flat mount. Scale bar: 0.25 mm." httpUri="https://zenodo.org/record/252387/files/figure.png" pageId="11" pageNumber="97">Fig. 43</figureCitation>
) short, basally thick with large tergites VI and VII partly exposed and covered in grey dusting; field of enlarged spiniform scales developed in ventral “intersegmental” membrane of segment VIII.
</paragraph>
<paragraph id="8BEA36ECCB39AB73FF66FCFF1C8C6FE8" blockId="11.[151,1437,289,1250]" pageId="11" pageNumber="97">
<emphasis id="B921EAFECB39AB73FF66FCFF1D166870" bold="true" box="[199,368,865,890]" pageId="11" pageNumber="97">Relationships.</emphasis>
<bibRefCitation id="EFC44B1DCB39AB73FED6FCFF1E706870" author="Hennig" box="[375,534,865,890]" pageId="11" pageNumber="97" refString="Hennig, W. (1972) Anthomyiidae [part]. In: Lindner, E. (Ed.), Die Fliegen der palaearktischen Region, 7 (1), pp. 425 - 472." type="journal article" year="1972">Hennig (1972</bibRefCitation>
,
<bibRefCitation id="EFC44B1DCB39AB73FD83FCFF1E016870" author="Hennig" box="[546,615,865,890]" pageId="11" pageNumber="97" refString="Hennig, W. (1976 a) Anthomyiidae [part]. In: Lindner, E. (Ed.), Die Fliegen der palaearktischen Region, 7 (1), pp. 921 - 974." type="journal article" year="1976" yearSuffix="a">1976a</bibRefCitation>
) admitted that his attempts to classify
<taxonomicName id="4C554D6FCB39AB73FBBCFCFF190C6870" authority="Ringdahl" authorityName="Ringdahl" box="[1053,1386,865,890]" class="Insecta" family="Anthomyiidae" genus="Hylemyia" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="97" phylum="Arthropoda" rank="species" species="curtigena">
<emphasis id="B921EAFECB39AB73FBBCFCFF189D6873" box="[1053,1275,865,889]" italics="true" pageId="11" pageNumber="97">Hylemyia curtigena</emphasis>
Ringdahl
</taxonomicName>
in a “modern” genus were either preliminary or poorly substantiated. As shown in the following, morphology presents ample justification for classifying the present species in a new genus,
<taxonomicName id="4C554D6FCB39AB73FC0DFC37184F68CB" box="[940,1065,937,961]" class="Insecta" family="Anthomyiidae" genus="Ringdahlia" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="97" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B921EAFECB39AB73FC0DFC37184F68CB" box="[940,1065,937,961]" italics="true" pageId="11" pageNumber="97">Ringdahlia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2125785CB39AB73FB91FC3418F168CB" box="[1072,1175,938,962]" pageId="11" pageNumber="97" rank="genus">gen. nov.</taxonomicNameLabel>
At first sight, the male of
<taxonomicName id="4C554D6FCB39AB73FF18FC501D2168EF" box="[185,327,974,997]" class="Insecta" family="Anthomyiidae" genus="Hylemyia" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="97" phylum="Arthropoda" rank="species" species="curtigena">
<emphasis id="B921EAFECB39AB73FF18FC501D2168EF" box="[185,327,974,997]" italics="true" pageId="11" pageNumber="97">R. curtigena</emphasis>
</taxonomicName>
(
<figureCitation id="136E2A69CB39AB73FEFAFC531DD668EC" box="[347,432,973,998]" captionStart="FIGURES 27 30" captionStartId="12.[151,264,1623,1646]" captionTargetBox="[174,1405,219,1602]" captionTargetId="figure@12.[151,1436,193,1602]" captionTargetPageId="12" captionText="FIGURES 27 30. Ringdahlia curtigena. 27, 28. Male from Finland, habitus and head, lateral views. 29. Female from Finland, labella. 30. Female from Finland, head and anterior thorax, anterodorsal view. Scale bars: 1 mm (27); 0.5 mm (28, 30); 0.1 mm (29)." httpUri="https://zenodo.org/record/252385/files/figure.png" pageId="11" pageNumber="97">Fig. 27</figureCitation>
) looks unremarkable with the typical appearance of an aerial anthomyiid fly, perhaps suggestive of a
<taxonomicName id="4C554D6FCB39AB73FEECFC6F1DBE6F03" box="[333,472,1009,1033]" class="Insecta" family="Anthomyiidae" genus="Botanophila" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="97" phylum="Arthropoda" rank="genus">
<emphasis id="B921EAFECB39AB73FEECFC6F1DBE6F03" box="[333,472,1009,1033]" italics="true" pageId="11" pageNumber="97">Botanophila</emphasis>
</taxonomicName>
or
<taxonomicName id="4C554D6FCB39AB73FDA3FC6C1E116F03" box="[514,631,1010,1033]" class="Insecta" family="Anthomyiidae" genus="Lasiomma" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="97" phylum="Arthropoda" rank="genus">
<emphasis id="B921EAFECB39AB73FDA3FC6C1E116F03" box="[514,631,1010,1033]" italics="true" pageId="11" pageNumber="97">Lasiomma</emphasis>
</taxonomicName>
species. On closer inspection, the male terminalia exhibit a wealth of autapomorphic details presenting no obvious hints on relationships to existing anthomyiid genera. The small, extensively membranized distiphallus is superficially suggestive of
<taxonomicName id="4C554D6FCB39AB73FC16FBA718246F5B" box="[951,1090,1081,1105]" class="Insecta" family="Anthomyiidae" genus="Botanophila" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="97" phylum="Arthropoda" rank="genus">
<emphasis id="B921EAFECB39AB73FC16FBA718246F5B" box="[951,1090,1081,1105]" italics="true" pageId="11" pageNumber="97">Botanophila</emphasis>
</taxonomicName>
, but such relationship is not supported by the slender, articulated pregonites. The female, that has now become available for study, has an oviscapt (
<figureCitation id="136E2A69CB39AB73FEAAFB1F1D056F90" box="[267,355,1153,1178]" captionStart="FIGURE 43" captionStartId="14.[151,250,1402,1424]" captionTargetBox="[223,1356,210,1348]" captionTargetId="figure@14.[216,1371,193,1381]" captionTargetPageId="14" captionText="FIGURE 43. Ringdahlia curtigena, female from Finland, oviscapt, flat mount. Scale bar: 0.25 mm." httpUri="https://zenodo.org/record/252387/files/figure.png" pageId="11" pageNumber="97">Fig. 43</figureCitation>
) without any of the defining characteristics of
<taxonomicName id="4C554D6FCB39AB73FC08FB1F182E6F93" box="[937,1096,1153,1177]" class="Insecta" family="Anthomyiidae" genus="Chirosiomima" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="97" phylum="Arthropoda" rank="genus">
<emphasis id="B921EAFECB39AB73FC08FB1F182E6F93" box="[937,1096,1153,1177]" italics="true" pageId="11" pageNumber="97">Chirosiomima</emphasis>
</taxonomicName>
(
<figureCitation id="136E2A69CB39AB73FBFEFB1F18DC6F90" box="[1119,1210,1153,1178]" captionStart="FIGURES 10 12" captionStartId="6.[151,264,1932,1955]" captionTargetBox="[176,1380,202,1871]" captionTargetId="figure@6.[170,1406,193,1899]" captionTargetPageId="6" captionText="FIGURES 10 12. Chirosiomima spp. 10. C. obscurinervis, male from Iran, sternites IV and V. 11. C. obscurinervis, lectotype male from Ghana, hypopygium, posterolateral view. 12. C. gestroi, female from Tunisia, oviscapt, flat mount (s = sternite; sp = spiracle; t = tergite). Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/252381/files/figure.png" pageId="11" pageNumber="97">Fig. 12</figureCitation>
). Besides that, the morphology of the female presents no evidence in support of a closest relationship to any known genus or group of genera.
</paragraph>
</subSubSection>
</treatment>
</document>