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<mods:title id="86C9E74828352D11E697B6F44389C75D">Taxonomy of the genus Protocypus J. Müller, 1923 (Coleoptera: Staphylinidae: Staphylinini) and two new species from China</mods:title>
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<mods:namePart id="CD2EFD88636986705A9D76E59064C0B4">He, Liang</mods:namePart>
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<mods:namePart id="FD751C4F9056E52B65EC282DB775EE72">Zhou, Hong-Zhang</mods:namePart>
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<mods:date id="5D96B00C312E2A597D091A81F206CBA3">2017</mods:date>
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<treatment id="03FC87F45B6F8563FF406FE38FE27CD3" ID-DOI="http://doi.org/10.5281/zenodo.6022972" ID-GBIF-Taxon="132548138" ID-Zenodo-Dep="6022972" LSID="urn:lsid:plazi:treatment:03FC87F45B6F8563FF406FE38FE27CD3" httpUri="http://treatment.plazi.org/id/03FC87F45B6F8563FF406FE38FE27CD3" lastPageNumber="471" pageId="13" pageNumber="471">
<subSubSection id="C34F65695B6F8563FF406FE389127EA1" box="[151,569,151,178]" pageId="13" pageNumber="471" type="nomenclature">
<paragraph id="8BEA36E25B6F8563FF406FE389127EA1" blockId="13.[151,569,151,211]" box="[151,569,151,178]" pageId="13" pageNumber="471">
<heading id="D0A2818E5B6F8563FF406FE389127EA1" bold="true" box="[151,569,151,178]" fontSize="11" level="1" pageId="13" pageNumber="471" reason="1">
<emphasis id="B921EAF05B6F8563FF406FE389127EA1" bold="true" box="[151,569,151,178]" pageId="13" pageNumber="471">
4.
<taxonomicName id="4C554D615B6F8563FF636FEC89127EA1" ID-CoL="4N52R" authority="Smetana, 2005" authorityName="Smetana" authorityYear="2005" box="[180,569,151,178]" class="Insecta" family="Staphylinidae" genus="Protocypus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="471" phylum="Arthropoda" rank="species" species="latro">
<emphasis id="B921EAF05B6F8563FF636FEC8A577EA2" bold="true" box="[180,380,151,177]" italics="true" pageId="13" pageNumber="471">Protocypus latro</emphasis>
<bibRefCitation id="EFC44B135B6F8563FE546FE389127EA1" author="Smetana" box="[387,569,152,178]" pageId="13" pageNumber="471" refString="Smetana, A. (2005) Contributions to the knowledge of the genera of the &quot; Staphylinus - complex &quot; (Coleoptera: Staphylinidae) of China Part. 5. The genus Protocypus J. Muller, 1923. Elytra, 33 (1), 269 - 301." type="journal article" year="2005">Smetana, 2005</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C34F65695B6F8563FF406FC1892E7F24" pageId="13" pageNumber="471" type="description">
<paragraph id="8BEA36E25B6F8563FF406FC18A1D7EC0" blockId="13.[151,569,151,211]" box="[151,310,186,211]" pageId="13" pageNumber="471">(Fig. 4-1, 4-2)</paragraph>
<paragraph id="8BEA36E25B6F8563FF406E79892E7F24" blockId="13.[151,1436,258,311]" pageId="13" pageNumber="471">
<bibRefCitation id="EFC44B135B6F8563FF406E798A197F0B" author="Smetana" box="[151,306,258,280]" pageId="13" pageNumber="471" refString="Smetana, A. (2005) Contributions to the knowledge of the genera of the &quot; Staphylinus - complex &quot; (Coleoptera: Staphylinidae) of China Part. 5. The genus Protocypus J. Muller, 1923. Elytra, 33 (1), 269 - 301." type="journal article" year="2005">Smetana, 2005</bibRefCitation>
: 294 (
<taxonomicName id="4C554D615B6F8563FEA56E788ACF7F0B" box="[370,484,259,280]" class="Insecta" family="Staphylinidae" genus="Protocypus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="471" phylum="Arthropoda" rank="genus">
<emphasis id="B921EAF05B6F8563FEA56E788ACF7F0B" box="[370,484,259,280]" italics="true" pageId="13" pageNumber="471">Protocypus</emphasis>
</taxonomicName>
; Type locality: China, Hubei, Dashennongjia);
<bibRefCitation id="EFC44B135B6F8563FC1E6E798FFD7F0B" author="Schulke" box="[969,1238,258,280]" pageId="13" pageNumber="471" refString="Schulke, M. &amp; Smetana, A. (2015) Staphylinidae. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Fol. 2. Hydrophiloidea - Staphylinoidea. Revised and Updated Edition. Brill, Leiden &amp; Boston, pp. 901 - 1134." type="journal article" year="2015">Schülke &amp; Smetana, 2015</bibRefCitation>
: 1100 (
<taxonomicName id="4C554D615B6F8563FAF46E788EBE7F0B" box="[1315,1429,259,280]" class="Insecta" family="Staphylinidae" genus="Protocypus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="471" phylum="Arthropoda" rank="genus">
<emphasis id="B921EAF05B6F8563FAF46E788EBE7F0B" box="[1315,1429,259,280]" italics="true" pageId="13" pageNumber="471">Protocypus</emphasis>
</taxonomicName>
; Catalog of Palaearctic Region).
</paragraph>
</subSubSection>
<subSubSection id="C34F65695B6F8563FF406E1F89927C47" pageId="13" pageNumber="471" type="materials_examined">
<paragraph id="8BEA36E25B6F8563FF406E1F89927C47" blockId="13.[151,1436,356,704]" pageId="13" pageNumber="471">
<materialsCitation id="3B3D3CBF5B6F8563FF406E1F88677FB3" ID-GBIF-Occurrence="1572323577" collectingDate="1998-07-23" collectingDateMax="1998-07-30" collectingDateMin="1998-07-23" collectorName="Xiaodong Yu" country="China" county="Material" elevation="1450" location="Guanmen Shan" municipality="Shennongjia" pageId="13" pageNumber="471" specimenCount="58" specimenCount-female="30" specimenCount-male="28" stateProvince="Hubei Province">
<emphasis id="B921EAF05B6F8563FF406E1F896F7F6E" bold="true" box="[151,580,356,381]" pageId="13" pageNumber="471">
<collectingCounty id="628B4E6E5B6F8563FF406E1F8BD47F6E" box="[151,255,356,381]" pageId="13" pageNumber="471">Material</collectingCounty>
examined.
<collectingCountry id="F34276725B6F8563FE5C6E1F8AC67F6E" box="[395,493,356,381]" name="China" pageId="13" pageNumber="471">CHINA</collectingCountry>
:
<collectingRegion id="4991F8005B6F8563FE2C6E1F896F7F6E" box="[507,580,356,381]" country="China" name="Hubei" pageId="13" pageNumber="471">Hubei</collectingRegion>
</emphasis>
Province:
<specimenCount id="9D53FD6B5B6F8563FD166E1F88017F6E" box="[705,810,356,381]" pageId="13" pageNumber="471" type="male">28 males</specimenCount>
,
<specimenCount id="9D53FD6B5B6F8563FCE06E1F889C7F6E" box="[823,951,356,381]" pageId="13" pageNumber="471" type="female">30 females</specimenCount>
,
<collectingMunicipality id="6B8EAC985B6F8563FC136E1F8F787F6E" box="[964,1107,356,381]" pageId="13" pageNumber="471">Shennongjia</collectingMunicipality>
,
<location id="8E8A60395B6F8563FBB76E1F8E3A7F6E" LSID="urn:lsid:plazi:treatment:03FC87F45B6F8563FF406FE38FE27CD3:8E8A60395B6F8563FBB76E1F8E3A7F6E" box="[1120,1297,356,381]" country="China" county="Material" municipality="Shennongjia" name="Guanmen Shan" pageId="13" pageNumber="471" stateProvince="Hubei Province">Guanmen Shan</location>
,
<quantity id="4CAD9B075B6F8563FAC96E1F8B847F8C" metricMagnitude="3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="13" pageNumber="471" unit="m" value="1450.0" valueMax="1500.0" valueMin="1400.0">
<elevation id="0078D1D15B6F8563FAC96E1F8B847F8C" metricMagnitude="3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="13" pageNumber="471" unit="m" value="1450.0" valueMax="1500.0" valueMin="1400.0">1400–1500 m</elevation>
</quantity>
,
<collectingMethod id="52144EF55B6F8563FF6D6EFC8A167F8C" box="[186,317,391,416]" pageId="13" pageNumber="471">pitfall traps</collectingMethod>
,
<collectingDate id="EFAFE9CA5B6F8563FE9F6EF38AD67FB3" box="[328,509,392,416]" pageId="13" pageNumber="471" value="1998-07-23" valueMax="1998-07-30" valueMin="1998-07-23">
<date id="FFEB10225B6F8563FE9F6EF38AD67FB3" box="[328,509,392,416]" pageId="13" pageNumber="471" value="1998-07-23" valueMax="1998-07-30" valueMin="1998-07-23">23–30.VII.1998</date>
</collectingDate>
, coll.
<collectorName id="26A053345B6F8563FD976EFC89FE7F8C" box="[576,725,391,416]" pageId="13" pageNumber="471">Xiaodong Yu</collectorName>
(IZ-CAS)
</materialsCitation>
;
<materialsCitation id="3B3D3CBF5B6F8563FC806EF38F927FD7" ID-GBIF-Occurrence="1572323572" collectingDate="1998-07-30" collectingDateMax="1998-08-11" collectingDateMin="1998-07-30" collectorName="Haisheng Zhou &amp; Junjian He" country="China" elevation="1450" location="Shennongjia" pageId="13" pageNumber="471" specimenCount="85" specimenCount-female="51" specimenCount-male="34" stateProvince="Shan">
<specimenCount id="9D53FD6B5B6F8563FC806EF388967FB3" box="[855,957,391,416]" pageId="13" pageNumber="471" type="male">34 males</specimenCount>
,
<specimenCount id="9D53FD6B5B6F8563FC1F6EFC8F6E7FB3" box="[968,1093,391,416]" pageId="13" pageNumber="471" type="female">51 females</specimenCount>
,
<location id="8E8A60395B6F8563FB876EFC8FF47FB3" LSID="urn:lsid:plazi:treatment:03FC87F45B6F8563FF406FE38FE27CD3:8E8A60395B6F8563FB876EFC8FF47FB3" box="[1104,1247,391,416]" country="China" name="Shennongjia" pageId="13" pageNumber="471" stateProvince="Shan">Shennongjia</location>
, Guanmen
<collectingRegion id="4991F8005B6F8563FA8A6EFC8EB37FB3" box="[1373,1432,391,416]" country="Myanmar" name="Shan" pageId="13" pageNumber="471">Shan</collectingRegion>
,
<elevation id="0078D1D15B6F8563FF406ED78A1F7FD7" box="[151,308,428,453]" metricMagnitude="3" metricUnit="m" metricValue="1.45" metricValueMax="1.46" metricValueMin="1.44" pageId="13" pageNumber="471" unit="m" value="1450.0" valueMax="1460.0" valueMin="1440.0">
<quantity id="4CAD9B075B6F8563FF406ED78A1F7FD7" box="[151,308,428,453]" metricMagnitude="3" metricUnit="m" metricValue="1.45" metricValueMax="1.46" metricValueMin="1.44" pageId="13" pageNumber="471" unit="m" value="1450.0" valueMax="1460.0" valueMin="1440.0">1440–1460 m</quantity>
</elevation>
,
<collectingMethod id="52144EF55B6F8563FEE86ED78AE97FD7" box="[319,450,428,453]" pageId="13" pageNumber="471">pitfall traps</collectingMethod>
,
<collectingDate id="EFAFE9CA5B6F8563FE196ED7899C7FD7" box="[462,695,428,452]" pageId="13" pageNumber="471" value="1998-07-30" valueMax="1998-08-11" valueMin="1998-07-30">
<date id="FFEB10225B6F8563FE196ED7899C7FD7" box="[462,695,428,452]" pageId="13" pageNumber="471" value="1998-07-30" valueMax="1998-08-11" valueMin="1998-07-30">30.VII–11.VIII.1998</date>
</collectingDate>
, coll.
<collectorName id="26A053345B6F8563FD2D6ED7888D7FD6" box="[762,934,428,453]" pageId="13" pageNumber="471">Haisheng Zhou</collectorName>
&amp;
<collectorName id="26A053345B6F8563FC1D6ED78F697FD7" box="[970,1090,428,453]" pageId="13" pageNumber="471">Junjian He</collectorName>
(IZ-CAS)
</materialsCitation>
;
<materialsCitation id="3B3D3CBF5B6F8563FB126ED78A177C1F" ID-GBIF-Occurrence="1572323594" collectingDate="2002-08-03" collectingDateMax="2002-08-06" collectingDateMin="2002-08-03" collectorName="Xiaodong Yu &amp; Jie Wu" country="China" elevation="1550" location="Shennongjia" pageId="13" pageNumber="471" specimenCount="9" specimenCount-female="6" specimenCount-male="3" stateProvince="Shan">
<specimenCount id="9D53FD6B5B6F8563FB126ED78E367FD6" box="[1221,1309,428,453]" pageId="13" pageNumber="471" type="male">3 males</specimenCount>
,
<specimenCount id="9D53FD6B5B6F8563FAFE6ED78EB37FD6" box="[1321,1432,428,453]" pageId="13" pageNumber="471" type="female">6 females</specimenCount>
,
<location id="8E8A60395B6F8563FF406EB48A0D7FFB" LSID="urn:lsid:plazi:treatment:03FC87F45B6F8563FF406FE38FE27CD3:8E8A60395B6F8563FF406EB48A0D7FFB" box="[151,294,463,488]" country="China" name="Shennongjia" pageId="13" pageNumber="471" stateProvince="Shan">Shennongjia</location>
, Guanmen
<collectingRegion id="4991F8005B6F8563FE716EB48ACA7FFB" box="[422,481,463,488]" country="Myanmar" name="Shan" pageId="13" pageNumber="471">Shan</collectingRegion>
,
<elevation id="0078D1D15B6F8563FE3B6EB4896F7FF4" box="[492,580,463,488]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="13" pageNumber="471" unit="m" value="1550.0">
<quantity id="4CAD9B075B6F8563FE3B6EB4896F7FF4" box="[492,580,463,488]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="13" pageNumber="471" unit="m" value="1550.0">1550 m</quantity>
</elevation>
, Japanese cedar forest,
<collectingMethod id="52144EF55B6F8563FC9A6EB488FB7FF4" box="[845,976,463,488]" pageId="13" pageNumber="471">pitfall traps</collectingMethod>
,
<collectingDate id="EFAFE9CA5B6F8563FC0B6EB48F557FFB" box="[988,1150,463,488]" pageId="13" pageNumber="471" value="2002-08-03" valueMax="2002-08-06" valueMin="2002-08-03">
<date id="FFEB10225B6F8563FC0B6EB48F557FFB" box="[988,1150,463,488]" pageId="13" pageNumber="471" value="2002-08-03" valueMax="2002-08-06" valueMin="2002-08-03">3–6.VIII.2002</date>
</collectingDate>
, coll.
<collectorName id="26A053345B6F8563FB146EB48E737FF4" box="[1219,1368,463,488]" pageId="13" pageNumber="471">Xiaodong Yu</collectorName>
&amp;
<collectorName id="26A053345B6F8563FAAA6EB48B957C1F" pageId="13" pageNumber="471">Jie Wu</collectorName>
(IZ-CAS)
</materialsCitation>
;
<materialsCitation id="3B3D3CBF5B6F8563FE996E8F8A967C23" ID-GBIF-Occurrence="1572323583" collectingDate="1998-07-24" collectingDateMax="1998-08-08" collectingDateMin="1998-07-24" collectorName="Xiaodong Yu" country="China" elevation="2180" location="Dalongtan" municipality="Shennongjia" pageId="13" pageNumber="471" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Shan">
<specimenCount id="9D53FD6B5B6F8563FE996E8F8A857C1E" box="[334,430,500,525]" pageId="13" pageNumber="471" type="male">3 males</specimenCount>
,
<specimenCount id="9D53FD6B5B6F8563FE176E8F891E7C1E" box="[448,565,500,525]" pageId="13" pageNumber="471" type="female">2 females</specimenCount>
,
<collectingMunicipality id="6B8EAC985B6F8563FD906E8F89FD7C1E" box="[583,726,500,525]" pageId="13" pageNumber="471">Shennongjia</collectingMunicipality>
,
<location id="8E8A60395B6F8563FD306E8F88747C1E" LSID="urn:lsid:plazi:treatment:03FC87F45B6F8563FF406FE38FE27CD3:8E8A60395B6F8563FD306E8F88747C1E" box="[743,863,500,525]" country="China" municipality="Shennongjia" name="Dalongtan" pageId="13" pageNumber="471" stateProvince="Shan">Dalongtan</location>
,
<elevation id="0078D1D15B6F8563FCA66E8F88E47C1F" box="[881,975,500,524]" metricMagnitude="3" metricUnit="m" metricValue="2.18" pageId="13" pageNumber="471" unit="m" value="2180.0">
<quantity id="4CAD9B075B6F8563FCA66E8F88E47C1F" box="[881,975,500,524]" metricMagnitude="3" metricUnit="m" metricValue="2.18" pageId="13" pageNumber="471" unit="m" value="2180.0">2180 m</quantity>
</elevation>
,
<collectingMethod id="52144EF55B6F8563FC366E8F8F417C1F" box="[993,1130,500,525]" pageId="13" pageNumber="471">pitfall traps</collectingMethod>
,
<collectingDate id="EFAFE9CA5B6F8563FBAB6E8F8E737C1F" box="[1148,1368,500,524]" pageId="13" pageNumber="471" value="1998-07-24" valueMax="1998-08-08" valueMin="1998-07-24">
<date id="FFEB10225B6F8563FBAB6E8F8E737C1F" box="[1148,1368,500,524]" pageId="13" pageNumber="471" value="1998-07-24" valueMax="1998-08-08" valueMin="1998-07-24">24.VII–8.VIII.1998</date>
</collectingDate>
, coll.
<collectorName id="26A053345B6F8563FF406D6C8A137C3C" box="[151,312,535,560]" pageId="13" pageNumber="471">Xiaodong Yu</collectorName>
(IZ-CAS)
</materialsCitation>
;
<materialsCitation id="3B3D3CBF5B6F8563FE026D6C89927C47" ID-GBIF-Occurrence="1572323584" collectingDate="1998-07-27" collectingDateMax="1998-08-10" collectingDateMin="1998-07-27" collectorName="Junjian He" country="China" county="Shennongjia" elevation="1870" location="Chegou" municipality="Jiuhuping" pageId="13" pageNumber="471" specimenCount="11" specimenCount-female="5" specimenCount-male="6" stateProvince="Shan">
<specimenCount id="9D53FD6B5B6F8563FE026D6C89107C23" box="[469,571,535,560]" pageId="13" pageNumber="471" type="male">6 males</specimenCount>
,
<specimenCount id="9D53FD6B5B6F8563FD836D6C89FB7C23" box="[596,720,535,560]" pageId="13" pageNumber="471" type="female">5 females</specimenCount>
,
<collectingCounty id="628B4E6E5B6F8563FD3F6D6C885C7C23" box="[744,887,535,560]" pageId="13" pageNumber="471">Shennongjia</collectingCounty>
,
<collectingMunicipality id="6B8EAC985B6F8563FC586D6C8F2B7C23" box="[911,1024,535,560]" pageId="13" pageNumber="471">Jiuhuping</collectingMunicipality>
,
<location id="8E8A60395B6F8563FBCE6D6C8F587C23" LSID="urn:lsid:plazi:treatment:03FC87F45B6F8563FF406FE38FE27CD3:8E8A60395B6F8563FBCE6D6C8F587C23" box="[1049,1139,535,560]" country="China" county="Shennongjia" municipality="Jiuhuping" name="Chegou" pageId="13" pageNumber="471" stateProvince="Shan">Chegou</location>
,
<elevation id="0078D1D15B6F8563FB5C6D638FC47C3C" box="[1163,1263,536,560]" metricMagnitude="3" metricUnit="m" metricValue="1.8699999999999999" pageId="13" pageNumber="471" unit="m" value="1870.0">
<quantity id="4CAD9B075B6F8563FB5C6D638FC47C3C" box="[1163,1263,536,560]" metricMagnitude="3" metricUnit="m" metricValue="1.8699999999999999" pageId="13" pageNumber="471" unit="m" value="1870.0">1870 m</quantity>
</elevation>
,
<collectingMethod id="52144EF55B6F8563FADF6D6C8EB37C3C" box="[1288,1432,535,560]" pageId="13" pageNumber="471">pitfall traps</collectingMethod>
,
<collectingDate id="EFAFE9CA5B6F8563FF406D478AA97C47" box="[151,386,572,596]" pageId="13" pageNumber="471" value="1998-07-27" valueMax="1998-08-10" valueMin="1998-07-27">
<date id="FFEB10225B6F8563FF406D478AA97C47" box="[151,386,572,596]" pageId="13" pageNumber="471" value="1998-07-27" valueMax="1998-08-10" valueMin="1998-07-27">27.VII–10.VIII.1998</date>
</collectingDate>
, coll.
<collectorName id="26A053345B6F8563FE126D4789157C47" box="[453,574,572,597]" pageId="13" pageNumber="471">Junjian He</collectorName>
(IZ-CAS).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34F65695B6F8563FF106D248AC27C8E" pageId="13" pageNumber="471" type="description">
<paragraph id="8BEA36E25B6F8563FF106D248AC27C8E" blockId="13.[151,1436,356,704]" pageId="13" pageNumber="471">
<emphasis id="B921EAF05B6F8563FF106D248A567C6B" bold="true" box="[199,381,607,632]" pageId="13" pageNumber="471">Measurements.</emphasis>
Body length:
<quantity id="4CAD9B075B6F8563FDC96D2489EE7C64" box="[542,709,607,632]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" metricValueMax="2.58" metricValueMin="2.02" pageId="13" pageNumber="471" unit="mm" value="23.0" valueMax="25.8" valueMin="20.2">20.2–25.8 mm</quantity>
. CL: 0.78; EL: 3.17; ELS: 1.38; EW: 3.99; HL: 3.17; HW: 3.94; PO: 1.54; PL: 3.80; PW: 3.64.
</paragraph>
</subSubSection>
<subSubSection id="C34F65695B6F8563FF106DDC8FE27CD3" box="[199,1225,679,704]" pageId="13" pageNumber="471" type="distribution">
<paragraph id="8BEA36E25B6F8563FF106DDC8FE27CD3" blockId="13.[151,1436,356,704]" box="[199,1225,679,704]" pageId="13" pageNumber="471">
<emphasis id="B921EAF05B6F8563FF106DDC8A4B7CD3" bold="true" box="[199,352,679,704]" pageId="13" pageNumber="471">Distribution.</emphasis>
<taxonomicName id="4C554D615B6F8563FEB06DD289097CD3" box="[359,546,680,704]" class="Insecta" family="Staphylinidae" genus="Protocypus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="471" phylum="Arthropoda" rank="species" species="latro">
<emphasis id="B921EAF05B6F8563FEB06DD289097CD3" box="[359,546,680,704]" italics="true" pageId="13" pageNumber="471">Protocypus latro</emphasis>
</taxonomicName>
is at present known from Dashennongjia in
<collectingRegion id="4991F8005B6F8563FBC56DDC8FEE7CD3" box="[1042,1221,679,704]" country="China" name="Hubei" pageId="13" pageNumber="471">Hubei Province</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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