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<document id="6D59A4E5C89004C5D2847A321EDE3E90" ID-CLB-Dataset="32789" ID-DOI="10.11646/zootaxa.4291.3.2" ID-GBIF-Dataset="ffc831a0-51a8-4aac-ab00-55297fc91dd9" ID-ISSN="1175-5326" ID-Zenodo-Dep="827606" ID-ZooBank="887FBB09-E370-4F46-837F-5E7F24F4CE42" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1500105344825" checkinUser="plazi" docAuthor="He, Liang &amp; Zhou, Hong-Zhang" docDate="2017" docId="03FC87F45B678566FF406AB789477B23" docLanguage="en" docName="zootaxa.4291.3.2.pdf" docOrigin="Zootaxa 4291 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Protocypus admirabilis He &amp; Zhou, 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="466" masterDocId="FFC5FF8C5B62856EFFD76F7B8B2B7E13" masterDocTitle="Taxonomy of the genus Protocypus J. Müller, 1923 (Coleoptera: Staphylinidae: Staphylinini) and two new species from China" masterLastPageNumber="480" masterPageNumber="458" pageNumber="463" updateTime="1698464906243" updateUser="ExternalLinkService">
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<mods:title id="561CE57138393136AE6688D153476087">Taxonomy of the genus Protocypus J. Müller, 1923 (Coleoptera: Staphylinidae: Staphylinini) and two new species from China</mods:title>
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<mods:namePart id="55C5A9BA8501D95839005D9FF2126605">He, Liang</mods:namePart>
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<mods:namePart id="C3AE0D8ED8E6FE57F003A22F8B0AE6BE">Zhou, Hong-Zhang</mods:namePart>
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<mods:date id="8D457E8696F6711FE0791429955D68B9">2017</mods:date>
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<treatment id="03FC87F45B678566FF406AB789477B23" ID-DOI="http://doi.org/10.5281/zenodo.6022964" ID-GBIF-Taxon="132548142" ID-Zenodo-Dep="6022964" LSID="urn:lsid:plazi:treatment:03FC87F45B678566FF406AB789477B23" httpUri="http://treatment.plazi.org/id/03FC87F45B678566FF406AB789477B23" lastPageId="8" lastPageNumber="466" pageId="5" pageNumber="463">
<subSubSection id="C34F65695B67856BFF406AB7891B7BF5" box="[151,560,1484,1510]" pageId="5" pageNumber="463" type="nomenclature">
<paragraph id="8BEA36E25B67856BFF406AB7891B7BF5" blockId="5.[151,560,1484,1543]" box="[151,560,1484,1510]" pageId="5" pageNumber="463">
<heading id="D0A2818E5B67856BFF406AB7891B7BF5" bold="true" box="[151,560,1484,1510]" fontSize="11" level="1" pageId="5" pageNumber="463" reason="1">
<emphasis id="B921EAF05B67856BFF406AB7891B7BF5" bold="true" box="[151,560,1484,1510]" pageId="5" pageNumber="463">
2.
<taxonomicName id="4C554D615B67856BFF636AB78AE77BF5" ID-CoL="4N528" box="[180,460,1484,1510]" class="Insecta" family="Staphylinidae" genus="Protocypus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="463" phylum="Arthropoda" rank="species" species="admirabilis" status="sp. nov.">
<emphasis id="B921EAF05B67856BFF636AB78AE77BF5" bold="true" box="[180,460,1484,1510]" italics="true" pageId="5" pageNumber="463">Protocypus admirabilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A212578B5B67856BFE046AB7891B7BF5" box="[467,560,1484,1510]" pageId="5" pageNumber="463" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34F65695B67856BFF406A958A6A7814" box="[151,321,1518,1543]" pageId="5" pageNumber="463" type="description">
<paragraph id="8BEA36E25B67856BFF406A958A6A7814" blockId="5.[151,560,1484,1543]" box="[151,321,1518,1543]" pageId="5" pageNumber="463">
(
<figureCitation id="136E2A675B67856BFF486A958A137814" box="[159,312,1518,1543]" captionStart="FIGURE 1 2" captionStartId="4.[151,250,1945,1967]" captionTargetBox="[165,1429,193,1925]" captionTargetId="figure@4.[165,1429,193,1925]" captionTargetPageId="4" captionText="FIGURE 1 2. Protocypus taibaiensis sp. nov.: A. male tergite 10; B and D. aedeagus, lateral view; C. aedeagus, ventral view; E. female tergite 10; F. apical portion of median lobe, ventral view, paramere removed; G. underside of apical portion of paramere; H. male sternite 9; I. second gonocoxite of female genital segment; J. male sternite 8 (Scale bars = 0.50 mm)." httpUri="https://zenodo.org/record/827612/files/figure.png" pageId="5" pageNumber="463">Figs. 2-1, 2-2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34F65695B67856BFF40694D89A07910" pageId="5" pageNumber="463" type="materials_examined">
<paragraph id="8BEA36E25B67856BFF40694D89BD785C" blockId="5.[151,1436,1590,2011]" box="[151,662,1590,1615]" pageId="5" pageNumber="463">
<materialsCitation id="3B3D3CBF5B67856BFF40694D89BD785C" ID-GBIF-Occurrence="1572323547" box="[151,662,1590,1615]" country="China" location="Hubei" pageId="5" pageNumber="463" specimenCount="1" stateProvince="Hubei" typeStatus="holotype">
<emphasis id="B921EAF05B67856BFF40694D8AD0785C" bold="true" box="[151,507,1590,1615]" pageId="5" pageNumber="463">
<typeStatus id="54EE88405B67856BFF40694D8BF9785C" box="[151,210,1590,1615]" pageId="5" pageNumber="463">Type</typeStatus>
locality:
<collectingCountry id="F34276725B67856BFE97694D8AB4785C" box="[320,415,1590,1615]" name="China" pageId="5" pageNumber="463">CHINA</collectingCountry>
,
<collectingRegion id="4991F8005B67856BFE7C694D8ADD785C" box="[427,502,1590,1615]" country="China" name="Hubei" pageId="5" pageNumber="463">Hubei</collectingRegion>
,
</emphasis>
Shennongjia.
</materialsCitation>
</paragraph>
<paragraph id="8BEA36E25B67856BFF10692189A07910" blockId="5.[151,1436,1590,2011]" pageId="5" pageNumber="463">
<materialsCitation id="3B3D3CBF5B67856BFF106921882A7884" ID-GBIF-Occurrence="1572323550" collectingDate="1998-07-28" collectingDateMax="1998-08-10" collectingDateMin="1998-07-28" collectorName="Hongzhang Zhou" country="China" elevation="1870" location="Hongshitougou" municipality="Shennongjia" pageId="5" pageNumber="463" specimenCount="2" stateProvince="Hubei Province" typeStatus="holotype">
<emphasis id="B921EAF05B67856BFF1069218AC37860" bold="true" box="[199,488,1626,1651]" pageId="5" pageNumber="463">
<typeStatus id="54EE88405B67856BFF1069218A297860" box="[199,258,1626,1651]" pageId="5" pageNumber="463">Type</typeStatus>
material.
<typeStatus id="54EE88405B67856BFEAC69218AC37860" box="[379,488,1626,1651]" pageId="5" pageNumber="463" type="holotype">Holotype</typeStatus>
</emphasis>
male,
<emphasis id="B921EAF05B67856BFDE5692189C37860" bold="true" box="[562,744,1626,1651]" pageId="5" pageNumber="463">
<collectingCountry id="F34276725B67856BFDE5692189B87860" box="[562,659,1626,1651]" name="China" pageId="5" pageNumber="463">CHINA</collectingCountry>
:
<collectingRegion id="4991F8005B67856BFD48692189C37860" box="[671,744,1626,1651]" country="China" name="Hubei" pageId="5" pageNumber="463">Hubei</collectingRegion>
</emphasis>
Province,
<collectingMunicipality id="6B8EAC985B67856BFC88692188C57860" box="[863,1006,1626,1651]" pageId="5" pageNumber="463">Shennongjia</collectingMunicipality>
,
<location id="8E8A60395B67856BFC2E69218F827860" LSID="urn:lsid:plazi:treatment:03FC87F45B678566FF406AB789477B23:8E8A60395B67856BFC2E69218F827860" box="[1017,1193,1626,1651]" country="China" municipality="Shennongjia" name="Hongshitougou" pageId="5" pageNumber="463" stateProvince="Hubei Province">Hongshitougou</location>
,
<elevation id="0078D1D15B67856BFB6469208E217861" box="[1203,1290,1627,1651]" metricMagnitude="3" metricUnit="m" metricValue="1.8699999999999999" pageId="5" pageNumber="463" unit="m" value="1870.0">
<quantity id="4CAD9B075B67856BFB6469208E217861" box="[1203,1290,1627,1651]" metricMagnitude="3" metricUnit="m" metricValue="1.8699999999999999" pageId="5" pageNumber="463" unit="m" value="1870.0">1870 m</quantity>
</elevation>
,
<collectingMethod id="52144EF55B67856BFAC269218EB37861" box="[1301,1432,1626,1651]" pageId="5" pageNumber="463">pitfall traps</collectingMethod>
,
<collectingDate id="EFAFE9CA5B67856BFF4069048AA97884" box="[151,386,1663,1687]" pageId="5" pageNumber="463" value="1998-07-28" valueMax="1998-08-10" valueMin="1998-07-28">
<date id="FFEB10225B67856BFF4069048AA97884" box="[151,386,1663,1687]" pageId="5" pageNumber="463" value="1998-07-28" valueMax="1998-08-10" valueMin="1998-07-28">28.VII10.VIII.1998</date>
</collectingDate>
, coll.
<collectorName id="26A053345B67856BFE12690589A27884" box="[453,649,1662,1687]" pageId="5" pageNumber="463">Hongzhang Zhou</collectorName>
(IZ-CAS)
</materialsCitation>
.
<materialsCitation id="3B3D3CBF5B67856BFCDB6905891D78A8" ID-GBIF-Occurrence="1572323537" collectingDate="1998-07-28" collectingDateMax="1998-08-10" collectingDateMin="1998-07-28" collectorName="Hongzhang Zhou" country="China" elevation="1870" location="Hongshitougou" municipality="Shennongjia" pageId="5" pageNumber="463" specimenCount="11" specimenCount-female="7" specimenCount-male="4" stateProvince="Hubei Province" typeStatus="paratype">
<emphasis id="B921EAF05B67856BFCDB690588AF7884" bold="true" box="[780,900,1662,1687]" pageId="5" pageNumber="463">
<typeStatus id="54EE88405B67856BFCDB690588AF7884" box="[780,900,1662,1687]" pageId="5" pageNumber="463" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<emphasis id="B921EAF05B67856BFC4569058F627884" bold="true" box="[914,1097,1662,1687]" pageId="5" pageNumber="463">
<collectingCountry id="F34276725B67856BFC45690588DF7884" box="[914,1012,1662,1687]" name="China" pageId="5" pageNumber="463">CHINA</collectingCountry>
:
<collectingRegion id="4991F8005B67856BFBD769058F627884" box="[1024,1097,1662,1687]" country="China" name="Hubei" pageId="5" pageNumber="463">Hubei</collectingRegion>
</emphasis>
Province:
<specimenCount id="9D53FD6B5B67856BFB1469048E367884" box="[1219,1309,1662,1687]" pageId="5" pageNumber="463" type="male">4 males</specimenCount>
,
<specimenCount id="9D53FD6B5B67856BFAFF69048EB37884" box="[1320,1432,1662,1687]" pageId="5" pageNumber="463" type="female">7 females</specimenCount>
, same data as holotype (IZ-CAS)
</materialsCitation>
;
<materialsCitation id="3B3D3CBF5B67856BFD9969D889F978CC" ID-GBIF-Occurrence="1572323573" collectingDate="1998-07-24" collectingDateMax="1998-08-08" collectingDateMin="1998-07-24" collectorName="Xiaodong Yu" country="China" elevation="1920" location="Jiuhuping" municipality="Shennongjia" pageId="5" pageNumber="463" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Hubei Province" typeStatus="paratype">
<specimenCount id="9D53FD6B5B67856BFD9969D8899878A8" box="[590,691,1698,1723]" pageId="5" pageNumber="463" type="male">3 males</specimenCount>
,
<specimenCount id="9D53FD6B5B67856BFD1C69D8886D78A8" box="[715,838,1698,1723]" pageId="5" pageNumber="463" type="female">2 females</specimenCount>
,
<collectingMunicipality id="6B8EAC985B67856BFC8969D988C678A8" box="[862,1005,1698,1723]" pageId="5" pageNumber="463">Shennongjia</collectingMunicipality>
,
<location id="8E8A60395B67856BFBD369D98F5E78A8" LSID="urn:lsid:plazi:treatment:03FC87F45B678566FF406AB789477B23:8E8A60395B67856BFBD369D98F5E78A8" box="[1028,1141,1698,1723]" country="China" municipality="Shennongjia" name="Jiuhuping" pageId="5" pageNumber="463" stateProvince="Hubei Province">Jiuhuping</location>
,
<elevation id="0078D1D15B67856BFB5A69D88FDB78A9" box="[1165,1264,1699,1723]" metricMagnitude="3" metricUnit="m" metricValue="1.92" pageId="5" pageNumber="463" unit="m" value="1920.0">
<quantity id="4CAD9B075B67856BFB5A69D88FDB78A9" box="[1165,1264,1699,1723]" metricMagnitude="3" metricUnit="m" metricValue="1.92" pageId="5" pageNumber="463" unit="m" value="1920.0">1920 m</quantity>
</elevation>
,
<collectingMethod id="52144EF55B67856BFADF69D98EB378A9" box="[1288,1432,1698,1723]" pageId="5" pageNumber="463">pitfall traps</collectingMethod>
,
<collectingDate id="EFAFE9CA5B67856BFF4069BC8A5878CC" box="[151,371,1735,1759]" pageId="5" pageNumber="463" value="1998-07-24" valueMax="1998-08-08" valueMin="1998-07-24">
<date id="FFEB10225B67856BFF4069BC8A5878CC" box="[151,371,1735,1759]" pageId="5" pageNumber="463" value="1998-07-24" valueMax="1998-08-08" valueMin="1998-07-24">24.VII8.VIII.1998</date>
</collectingDate>
, coll.
<collectorName id="26A053345B67856BFE6969BD897D78CD" box="[446,598,1734,1759]" pageId="5" pageNumber="463">Xiaodong Yu</collectorName>
(IZ-CAS)
</materialsCitation>
;
<materialsCitation id="3B3D3CBF5B67856BFD3769BC89A07910" ID-GBIF-Occurrence="1572323559" collectingDate="1998-08-01" collectingDateMax="1998-08-08" collectingDateMin="1998-08-01" collectorName="Xiaodong Yu" country="China" elevation="1610" location="Pingqiangangou" municipality="Shennongjia" pageId="5" pageNumber="463" specimenCount="1" specimenCount-male="1" stateProvince="Hubei Province" typeStatus="paratype">
<specimenCount id="9D53FD6B5B67856BFD3769BC881978CC" box="[736,818,1734,1759]" pageId="5" pageNumber="463" type="male">1 male</specimenCount>
,
<collectingMunicipality id="6B8EAC985B67856BFC9669BD88FB78CC" box="[833,976,1734,1759]" pageId="5" pageNumber="463">Shennongjia</collectingMunicipality>
,
<location id="8E8A60395B67856BFC0869BD8FB378CC" LSID="urn:lsid:plazi:treatment:03FC87F45B678566FF406AB789477B23:8E8A60395B67856BFC0869BD8FB378CC" box="[991,1176,1734,1759]" country="China" municipality="Shennongjia" name="Pingqiangangou" pageId="5" pageNumber="463" stateProvince="Hubei Province">Pingqiangangou</location>
,
<elevation id="0078D1D15B67856BFB7069BD8E2A78CD" box="[1191,1281,1734,1759]" metricMagnitude="3" metricUnit="m" metricValue="1.61" pageId="5" pageNumber="463" unit="m" value="1610.0">
<quantity id="4CAD9B075B67856BFB7069BD8E2A78CD" box="[1191,1281,1734,1759]" metricMagnitude="3" metricUnit="m" metricValue="1.61" pageId="5" pageNumber="463" unit="m" value="1610.0">1610 m</quantity>
</elevation>
,
<collectingMethod id="52144EF55B67856BFAC669BD8EB378CD" box="[1297,1432,1734,1759]" pageId="5" pageNumber="463">pitfall traps</collectingMethod>
,
<collectingDate id="EFAFE9CA5B67856BFF4069908A1C7910" box="[151,311,1771,1795]" pageId="5" pageNumber="463" value="1998-08-01" valueMax="1998-08-08" valueMin="1998-08-01">
<date id="FFEB10225B67856BFF4069908A1C7910" box="[151,311,1771,1795]" pageId="5" pageNumber="463" value="1998-08-01" valueMax="1998-08-08" valueMin="1998-08-01">18.VIII.1998</date>
</collectingDate>
, coll.
<collectorName id="26A053345B67856BFEAC699189247911" box="[379,527,1770,1795]" pageId="5" pageNumber="463">Xiaodong Yu</collectorName>
(IZ-CAS).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34F65695B678566FF1068758EB77DFF" lastPageId="8" lastPageNumber="466" pageId="5" pageNumber="463" type="description">
<paragraph id="8BEA36E25B67856BFF1068758AC37958" blockId="5.[151,1436,1590,2011]" pageId="5" pageNumber="463">
<emphasis id="B921EAF05B67856BFF1068758A567934" bold="true" box="[199,381,1806,1831]" pageId="5" pageNumber="463">Measurements.</emphasis>
Body length:
<quantity id="4CAD9B075B67856BFDC9687489EE7935" box="[542,709,1807,1831]" metricMagnitude="-2" metricUnit="m" metricValue="2.175" metricValueMax="2.42" metricValueMin="1.93" pageId="5" pageNumber="463" unit="mm" value="21.75" valueMax="24.2" valueMin="19.3">19.324.2 mm</quantity>
. CL: 0.76; EL: 3.09; ELS: 1.27; EW: 3.91; HL: 3.05; HW: 3.79; PO: 1.42; PL: 3.86; PW: 3.61.
</paragraph>
<paragraph id="8BEA36E25B67856BFF10682D890279C9" blockId="5.[151,1436,1590,2011]" pageId="5" pageNumber="463">
<emphasis id="B921EAF05B67856BFF10682D8A72797C" bold="true" box="[199,345,1878,1903]" pageId="5" pageNumber="463">Description.</emphasis>
Large sized species. Body relatively robust, black to brownish black; maxillary and labial palpi dark brunneous; antennae brownish black; legs brownish black, except protibia much paler and tends to be yellowish-brown; pubescence of dorsal side of head and pronotum brownish black, and that of elytra and abdominal tergites almost the same.
</paragraph>
<caption id="DF2A666A5B648568FF406814896979D1" httpUri="https://zenodo.org/record/827614/files/figure.png" pageId="6" pageNumber="464" targetBox="[175,1424,225,1838]" targetPageId="6">
<paragraph id="8BEA36E25B648568FF406814896979D1" blockId="6.[151,1436,1903,1986]" pageId="6" pageNumber="464">
<emphasis id="B921EAF05B648568FF4068148A007996" bold="true" box="[151,299,1903,1925]" pageId="6" pageNumber="464">FIGURE 2-1.</emphasis>
<taxonomicName id="4C554D615B648568FEE1680B89037996" box="[310,552,1903,1925]" class="Insecta" family="Staphylinidae" genus="Protocypus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="464" phylum="Arthropoda" rank="species" species="admirabilis" status="sp. nov.">
<emphasis id="B921EAF05B648568FEE1680B89037996" box="[310,552,1903,1925]" italics="true" pageId="6" pageNumber="464">Protocypus admirabilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A212578B5B648568FDE5681489AD7996" box="[562,646,1903,1925]" pageId="6" pageNumber="464" rank="species">
<emphasis id="B921EAF05B648568FDE5681489AD7996" bold="true" box="[562,646,1903,1925]" pageId="6" pageNumber="464">sp. nov.</emphasis>
</taxonomicNameLabel>
: A. habitus of species (2.0 mm); B. aedeagus, lateral view; (0.5 mm); C. aedeagus, ventral view (0.5 mm); D. underside of apical portion of paramere (0.5 mm); E. second gonocoxite of female genital segment (0.5 mm). (Scale in parentheses).
</paragraph>
</caption>
<caption id="DF2A666A5B658569FF4068ED8E7D79F9" httpUri="https://zenodo.org/record/827616/files/figure.png" pageId="7" pageNumber="465" targetBox="[162,1426,193,1925]" targetPageId="7">
<paragraph id="8BEA36E25B658569FF4068ED8E7D79F9" blockId="7.[151,1436,1942,2026]" pageId="7" pageNumber="465">
<emphasis id="B921EAF05B658569FF4068ED8A0379BF" bold="true" box="[151,296,1942,1964]" pageId="7" pageNumber="465">FIGURE 2-2.</emphasis>
<taxonomicName id="4C554D615B658569FEF968E3893079BF" box="[302,539,1942,1965]" class="Insecta" family="Staphylinidae" genus="Protocypus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="465" phylum="Arthropoda" rank="species" species="admirabilis" status="sp. nov.">
<emphasis id="B921EAF05B658569FEF968E3893079BF" box="[302,539,1942,1965]" italics="true" pageId="7" pageNumber="465">Protocypus admirabilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A212578B5B658569FDF668ED895A79BF" box="[545,625,1942,1964]" pageId="7" pageNumber="465" rank="species">
<emphasis id="B921EAF05B658569FDF668ED895A79BF" bold="true" box="[545,625,1942,1964]" pageId="7" pageNumber="465">sp. nov.</emphasis>
</taxonomicNameLabel>
: A. male tergite 10; B and D. aedeagus, lateral view; C. aedeagus, ventral view; E. female tergite 10; F. apical portion of median lobe, ventral view, paramere removed; G. underside of apical portion of paramere; H. male sternite 9; I. second gonocoxite of female genital segment; J. male sternite 8 (Scale bars = 0.50 mm).
</paragraph>
</caption>
<paragraph id="8BEA36E25B6A8566FF106FEC8A617CE3" blockId="8.[151,1437,151,1328]" pageId="8" pageNumber="466">Head of transversely oval shape, with curved lateral sides, slightly dilated posteriad and broadest at about posterior third, distinctly wider than long (ratio 1.24); eyes small and flat, considerably shorter than tempora (ratio 0.54) in dorsal view; punctation and pubescence on dorsal surface of head dense and fine, punctation gradually becoming sparser toward clypeus; puncture intervals smaller than diameters of punctures, with fine and dense microsculpture; no impunctate midline present. Antennae moderately long, segment 3 longer than segment 2 (ratio 1.22), segments 4 to 9 distinctly longer than wide (ratio greater than 1.20), segment 10 vaguely longer than wide, last segment short and wide. Pronotum vaguely longer than wide (ratio 1.07), with almost parallel sides and widely rounded posterior angles; narrow marginal groove disappearing downwards at anterior third of pronotal length; impunctate midline present only on posterior half of pronotal disc; punctation and pubescence on disc about same as that on dorsal surface of head. Pronotal hypomeron with only a few setae. Scutellum finely punctate and setose on entire surface except central basal parts, with microsculpture on entire surface. Elytra short, vaguely dilated posteriad and distinctly depressed at base; elytral length shorter than pronotum along midline, with a ratio of 0.33 along suture and a ratio of 0.80 along sides; elytral punctation fine and dense, combined with dense granulose microsculpture. Wings markedly reduced and budlike. Abdominal tergite 2 densely and finely punctate and pubescent on entire surface; all visible abdominal tergites finely and densely punctate and pubescent, punctuation gradually becoming sparser and coarser toward apex of abdomen; no yellowish tomentose pubescence on visible tergites 4 and 5.</paragraph>
<paragraph id="8BEA36E25B6A8566FF106D878EAB7DDB" blockId="8.[151,1437,151,1328]" pageId="8" pageNumber="466">
<emphasis id="B921EAF05B6A8566FF106D878A207D06" bold="true" box="[199,267,764,789]" pageId="8" pageNumber="466">Male.</emphasis>
Abdominal sternite 8 with triangular medioapical emargination moderately wide and obtuse (
<figureCitation id="136E2A675B6A8566FAF66D878E517D07" box="[1313,1402,764,789]" captionStart="FIGURE 1 2" captionStartId="4.[151,250,1945,1967]" captionTargetBox="[165,1429,193,1925]" captionTargetId="figure@4.[165,1429,193,1925]" captionTargetPageId="4" captionText="FIGURE 1 2. Protocypus taibaiensis sp. nov.: A. male tergite 10; B and D. aedeagus, lateral view; C. aedeagus, ventral view; E. female tergite 10; F. apical portion of median lobe, ventral view, paramere removed; G. underside of apical portion of paramere; H. male sternite 9; I. second gonocoxite of female genital segment; J. male sternite 8 (Scale bars = 0.50 mm)." httpUri="https://zenodo.org/record/827612/files/figure.png" pageId="8" pageNumber="466">Fig. 2-2</figureCitation>
J). Sternite 9 with moderately deep, obtuse apical emargination, tapered basal portion rather wide and long (
<figureCitation id="136E2A675B6A8566FA966C648EB77D2B" box="[1345,1436,799,824]" captionStart="FIGURE 1 2" captionStartId="4.[151,250,1945,1967]" captionTargetBox="[165,1429,193,1925]" captionTargetId="figure@4.[165,1429,193,1925]" captionTargetPageId="4" captionText="FIGURE 1 2. Protocypus taibaiensis sp. nov.: A. male tergite 10; B and D. aedeagus, lateral view; C. aedeagus, ventral view; E. female tergite 10; F. apical portion of median lobe, ventral view, paramere removed; G. underside of apical portion of paramere; H. male sternite 9; I. second gonocoxite of female genital segment; J. male sternite 8 (Scale bars = 0.50 mm)." httpUri="https://zenodo.org/record/827612/files/figure.png" pageId="8" pageNumber="466">Fig. 2-2</figureCitation>
H). Tergite 10 relatively narrow, evenly narrowed toward widely arcuate apex, sparsely setose (
<figureCitation id="136E2A675B6A8566FADB6C3F8E407D4F" box="[1292,1387,836,861]" captionStart="FIGURE 1 2" captionStartId="4.[151,250,1945,1967]" captionTargetBox="[165,1429,193,1925]" captionTargetId="figure@4.[165,1429,193,1925]" captionTargetPageId="4" captionText="FIGURE 1 2. Protocypus taibaiensis sp. nov.: A. male tergite 10; B and D. aedeagus, lateral view; C. aedeagus, ventral view; E. female tergite 10; F. apical portion of median lobe, ventral view, paramere removed; G. underside of apical portion of paramere; H. male sternite 9; I. second gonocoxite of female genital segment; J. male sternite 8 (Scale bars = 0.50 mm)." httpUri="https://zenodo.org/record/827612/files/figure.png" pageId="8" pageNumber="466">Fig. 2-2</figureCitation>
A). Aedeagus relatively long and simple (
<figureCitation id="136E2A675B6A8566FD9D6C1C898D7D93" box="[586,678,871,896]" captionStart="FIGURE 1 2" captionStartId="4.[151,250,1945,1967]" captionTargetBox="[165,1429,193,1925]" captionTargetId="figure@4.[165,1429,193,1925]" captionTargetPageId="4" captionText="FIGURE 1 2. Protocypus taibaiensis sp. nov.: A. male tergite 10; B and D. aedeagus, lateral view; C. aedeagus, ventral view; E. female tergite 10; F. apical portion of median lobe, ventral view, paramere removed; G. underside of apical portion of paramere; H. male sternite 9; I. second gonocoxite of female genital segment; J. male sternite 8 (Scale bars = 0.50 mm)." httpUri="https://zenodo.org/record/827612/files/figure.png" pageId="8" pageNumber="466">Fig. 2-2</figureCitation>
B, C &amp; D); median lobe distinctly asymmetrical as in
<figureCitation id="136E2A675B6A8566FAF66C1C8E577D93" box="[1313,1404,871,896]" captionStart="FIGURE 1 2" captionStartId="4.[151,250,1945,1967]" captionTargetBox="[165,1429,193,1925]" captionTargetId="figure@4.[165,1429,193,1925]" captionTargetPageId="4" captionText="FIGURE 1 2. Protocypus taibaiensis sp. nov.: A. male tergite 10; B and D. aedeagus, lateral view; C. aedeagus, ventral view; E. female tergite 10; F. apical portion of median lobe, ventral view, paramere removed; G. underside of apical portion of paramere; H. male sternite 9; I. second gonocoxite of female genital segment; J. male sternite 8 (Scale bars = 0.50 mm)." httpUri="https://zenodo.org/record/827612/files/figure.png" pageId="8" pageNumber="466">Fig. 2-2</figureCitation>
F; apical portion of paramere tongue-shaped, with sensory peg setae almost symmetrically arranged along lateral sides (
<figureCitation id="136E2A675B6A8566FF096CD48A137DDB" box="[222,312,943,968]" captionStart="FIGURE 1 2" captionStartId="4.[151,250,1945,1967]" captionTargetBox="[165,1429,193,1925]" captionTargetId="figure@4.[165,1429,193,1925]" captionTargetPageId="4" captionText="FIGURE 1 2. Protocypus taibaiensis sp. nov.: A. male tergite 10; B and D. aedeagus, lateral view; C. aedeagus, ventral view; E. female tergite 10; F. apical portion of median lobe, ventral view, paramere removed; G. underside of apical portion of paramere; H. male sternite 9; I. second gonocoxite of female genital segment; J. male sternite 8 (Scale bars = 0.50 mm)." httpUri="https://zenodo.org/record/827612/files/figure.png" pageId="8" pageNumber="466">Fig. 2-2</figureCitation>
G); lateral view of paramere not closely cling to median lobe near apex (
<figureCitation id="136E2A675B6A8566FBBB6CD48FFA7DDB" box="[1132,1233,943,968]" captionStart="FIGURE 1 2" captionStartId="4.[151,250,1945,1967]" captionTargetBox="[165,1429,193,1925]" captionTargetId="figure@4.[165,1429,193,1925]" captionTargetPageId="4" captionText="FIGURE 1 2. Protocypus taibaiensis sp. nov.: A. male tergite 10; B and D. aedeagus, lateral view; C. aedeagus, ventral view; E. female tergite 10; F. apical portion of median lobe, ventral view, paramere removed; G. underside of apical portion of paramere; H. male sternite 9; I. second gonocoxite of female genital segment; J. male sternite 8 (Scale bars = 0.50 mm)." httpUri="https://zenodo.org/record/827612/files/figure.png" pageId="8" pageNumber="466">Figs. 2-1</figureCitation>
B, 2-2 B &amp; D).
</paragraph>
<paragraph id="8BEA36E25B6A8566FF106CAF8EB77DFF" blockId="8.[151,1437,151,1328]" box="[199,1436,980,1005]" pageId="8" pageNumber="466">
<emphasis id="B921EAF05B6A8566FF106CAF8A0E7DFE" bold="true" box="[199,293,980,1005]" pageId="8" pageNumber="466">Female.</emphasis>
Second gonocoxite robust, shaped as in
<figureCitation id="136E2A675B6A8566FD3C6CAF886F7DFF" box="[747,836,980,1005]" captionStart="FIGURE 1 2" captionStartId="4.[151,250,1945,1967]" captionTargetBox="[165,1429,193,1925]" captionTargetId="figure@4.[165,1429,193,1925]" captionTargetPageId="4" captionText="FIGURE 1 2. Protocypus taibaiensis sp. nov.: A. male tergite 10; B and D. aedeagus, lateral view; C. aedeagus, ventral view; E. female tergite 10; F. apical portion of median lobe, ventral view, paramere removed; G. underside of apical portion of paramere; H. male sternite 9; I. second gonocoxite of female genital segment; J. male sternite 8 (Scale bars = 0.50 mm)." httpUri="https://zenodo.org/record/827612/files/figure.png" pageId="8" pageNumber="466">Fig. 2-1</figureCitation>
E. Tergite 10 relatively wide, shaped as in
<figureCitation id="136E2A675B6A8566FAF26CAF8E557DFF" box="[1317,1406,980,1005]" captionStart="FIGURE 1 2" captionStartId="4.[151,250,1945,1967]" captionTargetBox="[165,1429,193,1925]" captionTargetId="figure@4.[165,1429,193,1925]" captionTargetPageId="4" captionText="FIGURE 1 2. Protocypus taibaiensis sp. nov.: A. male tergite 10; B and D. aedeagus, lateral view; C. aedeagus, ventral view; E. female tergite 10; F. apical portion of median lobe, ventral view, paramere removed; G. underside of apical portion of paramere; H. male sternite 9; I. second gonocoxite of female genital segment; J. male sternite 8 (Scale bars = 0.50 mm)." httpUri="https://zenodo.org/record/827612/files/figure.png" pageId="8" pageNumber="466">Fig. 2-2</figureCitation>
E.
</paragraph>
</subSubSection>
<subSubSection id="C34F65695B6A8566FF106C8C8A0B7AFB" pageId="8" pageNumber="466" type="distribution">
<paragraph id="8BEA36E25B6A8566FF106C8C8A297A26" blockId="8.[151,1437,151,1328]" pageId="8" pageNumber="466">
<emphasis id="B921EAF05B6A8566FF106C8C8A4B7A03" bold="true" box="[199,352,1015,1040]" pageId="8" pageNumber="466">Distribution.</emphasis>
<taxonomicName id="4C554D615B6A8566FEBE6C8289597A03" box="[361,626,1016,1040]" class="Insecta" family="Staphylinidae" genus="Protocypus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="466" phylum="Arthropoda" rank="species" species="admirabilis" status="sp. nov.">
<emphasis id="B921EAF05B6A8566FEBE6C8289597A03" box="[361,626,1016,1040]" italics="true" pageId="8" pageNumber="466">Protocypus admirabilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A212578B5B6A8566FDAC6C8C89FC7A03" box="[635,727,1015,1040]" pageId="8" pageNumber="466" rank="species">
<emphasis id="B921EAF05B6A8566FDAC6C8C89FC7A03" bold="true" box="[635,727,1015,1040]" pageId="8" pageNumber="466">sp. nov.</emphasis>
</taxonomicNameLabel>
is at present known only from Shennongjia in western
<collectingRegion id="4991F8005B6A8566FA8F6C8C8BD57A26" country="China" name="Hubei" pageId="8" pageNumber="466">Hubei Province</collectingRegion>
.
</paragraph>
<paragraph id="8BEA36E25B6A8566FF106B448A0B7AFB" blockId="8.[151,1437,151,1328]" pageId="8" pageNumber="466">
<emphasis id="B921EAF05B6A8566FF106B448A3A7A4B" bold="true" box="[199,273,1087,1112]" pageId="8" pageNumber="466">Notes.</emphasis>
Most specimens of
<taxonomicName id="4C554D615B6A8566FE276B3A89DD7A4B" box="[496,758,1088,1112]" class="Insecta" family="Staphylinidae" genus="Protocypus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="466" phylum="Arthropoda" rank="species" species="admirabilis" status="sp. nov.">
<emphasis id="B921EAF05B6A8566FE276B3A89DD7A4B" box="[496,758,1088,1112]" italics="true" pageId="8" pageNumber="466">Protocypus admirabilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A212578B5B6A8566FD2B6B44887D7A4B" box="[764,854,1087,1112]" pageId="8" pageNumber="466" rank="species">
<emphasis id="B921EAF05B6A8566FD2B6B44887D7A4B" bold="true" box="[764,854,1087,1112]" pageId="8" pageNumber="466">sp. nov.</emphasis>
</taxonomicNameLabel>
were taken from the same pitfall traps together with specimens of
<taxonomicName id="4C554D615B6A8566FEE16B1E898D7A6E" authority="Smetana, 2005" authorityName="Smetana" authorityYear="2005" box="[310,678,1124,1149]" class="Insecta" family="Staphylinidae" genus="Protocypus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="466" phylum="Arthropoda" rank="species" species="latro">
<emphasis id="B921EAF05B6A8566FEE16B1E8AD87A6F" box="[310,499,1124,1148]" italics="true" pageId="8" pageNumber="466">Protocypus latro</emphasis>
<bibRefCitation id="EFC44B135B6A8566FE2A6B1F898D7A6E" author="Smetana" box="[509,678,1124,1149]" pageId="8" pageNumber="466" refString="Smetana, A. (2005) Contributions to the knowledge of the genera of the &quot; Staphylinus - complex &quot; (Coleoptera: Staphylinidae) of China Part. 5. The genus Protocypus J. Muller, 1923. Elytra, 33 (1), 269 - 301." type="journal article" year="2005">Smetana, 2005</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C554D615B6A8566FD356B1E8F4F7A6E" authority="Smetana, 2005" authorityName="Smetana" authorityYear="2005" box="[738,1124,1124,1149]" class="Insecta" family="Staphylinidae" genus="Protocypus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="466" phylum="Arthropoda" rank="species" species="pilifer">
<emphasis id="B921EAF05B6A8566FD356B1E88857A6F" box="[738,942,1124,1148]" italics="true" pageId="8" pageNumber="466">Protocypus pilifer</emphasis>
<bibRefCitation id="EFC44B135B6A8566FC6F6B1F8F4F7A6E" author="Smetana" box="[952,1124,1124,1149]" pageId="8" pageNumber="466" refString="Smetana, A. (2005) Contributions to the knowledge of the genera of the &quot; Staphylinus - complex &quot; (Coleoptera: Staphylinidae) of China Part. 5. The genus Protocypus J. Muller, 1923. Elytra, 33 (1), 269 - 301." type="journal article" year="2005">Smetana, 2005</bibRefCitation>
</taxonomicName>
, so it is possible that these three species share the same habitat.
<taxonomicName id="4C554D615B6A8566FDEE6BF2886B7AB3" box="[569,832,1160,1184]" class="Insecta" family="Staphylinidae" genus="Protocypus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="466" phylum="Arthropoda" rank="species" species="admirabilis" status="sp. nov.">
<emphasis id="B921EAF05B6A8566FDEE6BF2886B7AB3" box="[569,832,1160,1184]" italics="true" pageId="8" pageNumber="466">Protocypus admirabilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A212578B5B6A8566FC9E6BFC88887AB3" box="[841,931,1159,1184]" pageId="8" pageNumber="466" rank="species">
<emphasis id="B921EAF05B6A8566FC9E6BFC88887AB3" bold="true" box="[841,931,1159,1184]" pageId="8" pageNumber="466">sp. nov.</emphasis>
</taxonomicNameLabel>
can be distinguished from these two species and other species of genus
<taxonomicName id="4C554D615B6A8566FE066BD689647AD7" box="[465,591,1197,1220]" class="Insecta" family="Staphylinidae" genus="Protocypus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="466" phylum="Arthropoda" rank="genus">
<emphasis id="B921EAF05B6A8566FE066BD689647AD7" box="[465,591,1197,1220]" italics="true" pageId="8" pageNumber="466">Protocypus</emphasis>
</taxonomicName>
by the shape of the aedeagus, especially by the shape of the paramere in lateral view.
</paragraph>
</subSubSection>
<subSubSection id="C34F65695B6A8566FF106B8F89477B23" pageId="8" pageNumber="466" type="etymology">
<paragraph id="8BEA36E25B6A8566FF106B8F89477B23" blockId="8.[151,1437,151,1328]" pageId="8" pageNumber="466">
<emphasis id="B921EAF05B6A8566FF106B8F8A657B1E" bold="true" box="[199,334,1268,1293]" pageId="8" pageNumber="466">Etymology.</emphasis>
The specific epithet is the Latin adjective
<taxonomicName id="4C554D615B6A8566FCEC6B8F88977B1F" box="[827,956,1268,1292]" class="Insecta" family="Staphylinidae" genus="Protocypus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="466" phylum="Arthropoda" rank="species" species="admirabilis">
<emphasis id="B921EAF05B6A8566FCEC6B8F88977B1F" box="[827,956,1268,1292]" italics="true" pageId="8" pageNumber="466">admirabilis</emphasis>
</taxonomicName>
(very special, unexpected) to indicate the special form of the paramere of aedeagus.
</paragraph>
</subSubSection>
</treatment>
</document>